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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.745.23685" ID-PMC="PMC5904534" ID-Pensoft-Pub="1313-2970-745-101" ID-Pensoft-UUID="047DFFCAFFA5F32EA97C873F4708943F" ID-PubMed="29670449" ID-Zenodo-Dep="1222435" ID-ZooBank="8785DC6BC2A244FD94B6243EB07C717F" ModsDocID="1313-2970-745-101" checkinTime="1521799180220" checkinUser="pensoft" docAuthor="Moore, Matthew R., Cave, Ronald D. &amp; Branham, Marc A." docDate="2018" docId="49007555752243B2687BAD201CF946EA" docLanguage="en" docName="ZooKeys 745: 101-378" docOrigin="ZooKeys 745" docPubDate="2018-03-22" docSource="http://dx.doi.org/10.3897/zookeys.745.23685" docTitle="Cyclocephala variipenis Dechambre 1999" docType="treatment" docVersion="6" id="047DFFCAFFA5F32EA97C873F4708943F" lastPageNumber="223" masterDocId="047DFFCAFFA5F32EA97C873F4708943F" masterDocTitle="Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)" masterLastPageNumber="378" masterPageNumber="101" pageNumber="223" updateTime="1668165599527" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moore, Matthew R.</mods:namePart>
<mods:affiliation>Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
<mods:nameIdentifier type="email">cyclocephala@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cave, Ronald D.</mods:namePart>
<mods:affiliation>Department of Entomology and Nematology, University of Florida, Indian River Research and Education Center, 2199 South Rock Road, Fort Pierce, FL 34945, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Branham, Marc A.</mods:namePart>
<mods:affiliation>Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2018</mods:date>
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<mods:number>2018-03-22</mods:number>
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<treatment ID-GBIF-Taxon="142480449" LSID="urn:lsid:plazi:treatment:49007555752243B2687BAD201CF946EA" httpUri="http://treatment.plazi.org/id/49007555752243B2687BAD201CF946EA" lastPageNumber="223" pageId="122" pageNumber="223">
<subSubSection pageId="122" pageNumber="223" type="nomenclature">
<paragraph pageId="122" pageNumber="223">
<taxonomicName LSID="49007555-7522-43B2-687B-AD201CF946EA" authority="Dechambre, 1999" authorityName="Dechambre" authorityYear="1999" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala variipenis" order="Coleoptera" pageId="122" pageNumber="223" phylum="Arthropoda" rank="species" species="variipenis">Cyclocephala variipenis Dechambre, 1999</taxonomicName>
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<paragraph pageId="122" pageNumber="223">
<taxonomicName authorityName="Dechambre" authorityYear="1999" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala variipenis" order="Coleoptera" pageId="122" pageNumber="223" phylum="Arthropoda" rank="species" species="variipenis">Cyclocephala variipenis</taxonomicName>
Dechambre, 1999: 19-20 [original combination].
</paragraph>
</subSubSection>
<subSubSection pageId="122" pageNumber="223" type="types">
<paragraph pageId="122" pageNumber="223">Types.</paragraph>
<paragraph pageId="122" pageNumber="223">
Holotype ♂ at MNHN (
<bibRefCitation author="Dechambre, R-P" pageId="173" pageNumber="274" refId="B239" refString="Dechambre, R-P, 1999. Vingt nouvelles especes et une nouvelle sous-especes de Cyclocephala Burmeister, 1847. Les Coleopteres du Monde 14, Supplement 1. Dynastidae americains. Hillside Books, Canterbury, 24 pp. + 1 pls." title="Vingt nouvelles especes et une nouvelle sous-especes de Cyclocephala Burmeister, 1847. Les Coleopteres du Monde 14, Supplement 1. Dynastidae americains. Hillside Books, Canterbury, 24 pp. + 1 pls." year="1999">Dechambre 1999</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="122" pageNumber="223" type="distribution">
<paragraph pageId="122" pageNumber="223">Distribution.</paragraph>
<paragraph pageId="122" pageNumber="223">BOLIVIA: La Paz.</paragraph>
</subSubSection>
<subSubSection pageId="122" pageNumber="223" type="references">
<paragraph pageId="122" pageNumber="223">References.</paragraph>
<paragraph pageId="122" pageNumber="223">
<bibRefCitation author="Dechambre, R-P" pageId="173" pageNumber="274" refId="B239" refString="Dechambre, R-P, 1999. Vingt nouvelles especes et une nouvelle sous-especes de Cyclocephala Burmeister, 1847. Les Coleopteres du Monde 14, Supplement 1. Dynastidae americains. Hillside Books, Canterbury, 24 pp. + 1 pls." title="Vingt nouvelles especes et une nouvelle sous-especes de Cyclocephala Burmeister, 1847. Les Coleopteres du Monde 14, Supplement 1. Dynastidae americains. Hillside Books, Canterbury, 24 pp. + 1 pls." year="1999">Dechambre 1999</bibRefCitation>
,
<bibRefCitation author="Krajcik, M" pageId="200" pageNumber="301" refId="B658" refString="Krajcik, M, 2005. Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma.X, Supplement No. 2, Pilsen, 122 pp." title="Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma. X, Supplement No. 2, Pilsen, 122 pp." year="2005">Krajcik 2005</bibRefCitation>
,
<bibRefCitation author="Krajcik, M" pageId="200" pageNumber="301" refId="B659" refString="Krajcik, M, 2012. Checklist of the world Scarabaeoidea. Animma.X Supplement 5, Pilsen, 278 pp." title="Checklist of the world Scarabaeoidea. Animma. X Supplement 5, Pilsen, 278 pp." year="2012">2012</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<heading id="9A5E81C9FFA36D3EFF1FFF23583EFF4F" bold="true" box="[151,568,151,176]" fontSize="10" level="2" pageId="30" pageNumber="131" reason="2">
<taxonomicName id="06A94D26FFA36D3EFF1FFF23583EFF4F" ID-CoL="3HL3F" authority="Jousseaume, 1888" authorityName="Jousseaume" authorityYear="1888" box="[151,568,151,176]" class="Gastropoda" family="Turbinidae" genus="Guildfordia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="30" pageNumber="131" phylum="Mollusca" rank="species" species="yoka">
<emphasis id="F3DDEAB7FFA36D3EFF1FFF23583EFF4F" bold="true" box="[151,568,151,176]" pageId="30" pageNumber="131">
<emphasis id="F3DDEAB7FFA36D3EFF1FFF235B5DFF4F" bold="true" box="[151,347,151,176]" italics="true" pageId="30" pageNumber="131">Guildfordia yoka</emphasis>
Jousseaume, 1888
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<paragraph id="C11636A5FFA36D3EFF1FFF085B4CFF2A" blockId="30.[151,568,151,213]" box="[151,330,188,213]" pageId="30" pageNumber="131">
(
<figureCitation id="59922A20FFA36D3EFF16FF085B45FF2A" box="[158,323,188,213]" captionStart="FIGURES 69 – 73" captionStartId="29.[151,264,1843,1865]" captionTargetBox="[243,1351,195,1825]" captionTargetId="figure@29.[241,1354,193,1828]" captionTargetPageId="29" captionText="FIGURES 69 – 73. Turbinidae. 69. Guildfordia yoka Jousseaume, 1888. RGM 608.376. Locality Anda 1. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 70. Guildfordia yoka Jousseaume, 1888. RGM 608.377. Locality Anda 1. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 71. Guildfordia yoka Jousseaume, 1888. RGM 608.378. Locality Anda 2. (a) basal view, (b) apical view. 72. Guildfordia yoka Jousseaume, 1888. RGM 608.379. Locality Anda 1. (a) apertural view, (b) basal view, (c) apical view. 73. Guildfordia yoka Jousseaume, 1888. RGM 608.380. Locality Anda 3. (a) SEM detail of protoconch." httpUri="https://zenodo.org/record/250974/files/figure.png" pageId="30" pageNumber="131">Figures 69–73</figureCitation>
)
</paragraph>
<paragraph id="C11636A5FFA36D3EFF1FFEB0595BFEE5" blockId="30.[151,1117,260,312]" box="[151,861,260,282]" pageId="30" pageNumber="131">
2000
<taxonomicName id="06A94D26FFA36D3EFF7DFEB05859FEE5" authority="Jousseaume, 1888" authorityName="Jousseaume" authorityYear="1888" box="[245,607,260,282]" class="Gastropoda" family="Turbinidae" genus="Guildfordia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="30" pageNumber="131" phylum="Mollusca" rank="species" species="yoka">
<emphasis id="F3DDEAB7FFA36D3EFF7DFEB05BA4FEE5" box="[245,418,260,282]" italics="true" pageId="30" pageNumber="131">Guildfordia yoka</emphasis>
Jousseaume, 1888
</taxonomicName>
—Sasaki: p. 101, fig. 51.
</paragraph>
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<paragraph id="C11636A5FFA36D3EFF1FFE965E5BFEC7" blockId="30.[151,1117,260,312]" box="[151,1117,290,312]" pageId="30" pageNumber="131">
2008
<taxonomicName id="06A94D26FFA36D3EFF7DFE9658E2FEC7" ID-CoL="3HL34" authority="Habe &amp; Okutani, 1983" authorityName="Habe &amp; Okutani" authorityYear="1983" box="[245,740,290,312]" class="Gastropoda" family="Turbinidae" genus="Guildfordia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="30" pageNumber="131" phylum="Mollusca" rank="subSpecies" species="yoka" subSpecies="delicata">
<emphasis id="F3DDEAB7FFA36D3EFF7DFE965BFCFEC7" box="[245,506,290,312]" italics="true" pageId="30" pageNumber="131">Guildfordia yoka delicata</emphasis>
<bibRefCitation id="A5384B54FFA36D3EFD88FE9658E2FEC7" author="Habe" box="[512,740,290,312]" pageId="30" pageNumber="131" refString="Habe, T. &amp; Okutani, T. (1983) A new vesicomyid bivalve and two new gastropods of the genera Guildfordia and Xenophora from the Philippines. Venus, 42, 1 - 7." type="journal article" year="1983">Habe &amp; Okutani, 1983</bibRefCitation>
</taxonomicName>
—Kreipl &amp; Alf: p. 250, pl. 70, fig, 5.
</paragraph>
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<subSubSection id="89B3652EFFA36D3EFF1FFED159EDFDCD" pageId="30" pageNumber="131" type="materials_examined">
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<emphasis id="F3DDEAB7FFA36D3EFF1FFED15B00FE81" bold="true" box="[151,262,357,382]" pageId="30" pageNumber="131">Material.</emphasis>
Anda 1 (42); Anda 2 (38); Anda 3 (24); Anda 4 (18); Anda 6 (4); AndaDeVos (4); AndaClif1 (1); AndaClif3 (4); Roxas (1).
</paragraph>
<paragraph id="C11636A5FFA36D3EFF4FFE1959EDFDCD" blockId="30.[151,1436,357,670]" pageId="30" pageNumber="131">
<emphasis id="F3DDEAB7FFA36D3EFF4FFE195B9CFE39" bold="true" box="[199,410,429,454]" pageId="30" pageNumber="131">Characterization.</emphasis>
Shell almost planispiral, with reddish surface, H
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,
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W
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mm; P planispiral, 1.1 whorls, two discontinuous spiral ribs and granulate surface in between, DN
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; slightly elevated terminal varix; T 7–9 long spines per whorl, initially triangular, thin hollow, on later whorls elongate and massive; rows of knobs below the suture; umbilicus covered with white, shiny callus.
</paragraph>
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<emphasis id="F3DDEAB7FFA36D3EFF4FFD895B66FDA9" bold="true" box="[199,352,573,598]" pageId="30" pageNumber="131">Distribution.</emphasis>
Today the species occurs in Central southern
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and southward to Indo-West Pacific, at
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depth on sand bottom (
<bibRefCitation id="A5384B54FFA36D3EFE78FDD55887FD85" author="Sasaki" box="[496,641,609,634]" pageId="30" pageNumber="131" refString="Sasaki, T. (2000) Family Turbinidae. In: Okutani, T. (Ed.), Marine Mollusks in Japan. Tokai University Press, Tokyo, pp. 100 - 101." type="book chapter" year="2000">Sasaki 2000</bibRefCitation>
). It has also been reported from the
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, Aliguay
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, from
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depth (
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).
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105
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<emphasis id="F3DDEAB7FFA56D38FF1FFA485B3EF9E9" bold="true" box="[151,312,1532,1558]" italics="true" pageId="24" pageNumber="125">Archiminolia</emphasis>
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spec. 1
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(
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<emphasis id="F3DDEAB7FFA56D38FF1FF9D25B00F980" bold="true" box="[151,262,1638,1663]" pageId="24" pageNumber="125">Material.</emphasis>
AndaDeVos (4).
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<emphasis id="F3DDEAB7FFA56D38FF4FF93E5B9CF95C" bold="true" box="[199,410,1674,1699]" pageId="24" pageNumber="125">Characterization.</emphasis>
Shell low, H
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W
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mm; P 1.1 whorls, with two spiral ribs, DN
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; P/T boundary thin, slightly raised varix; first three T whorls with low spiral and axial ribs and axially arranged microscopic granules; fourth whorl smooth, coronate.
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<emphasis id="F3DDEAB7FFA56D38FF4FF9425B3DF8F0" bold="true" box="[199,315,1782,1807]" pageId="24" pageNumber="125">Remarks.</emphasis>
This species is tentatively assigned to the genus
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<emphasis id="F3DDEAB7FFA56D38FCFCF9425E01F8F0" box="[884,1031,1782,1807]" italics="true" pageId="24" pageNumber="125">Archiminolia</emphasis>
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<emphasis id="F3DDEAB7FFA56D38FE73F8AE5888F8CC" box="[507,654,1818,1843]" italics="true" pageId="24" pageNumber="125">Archiminolia</emphasis>
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species collected from the
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in the MNHN collection (STW, pers. obs.).
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79
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Genus
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species.
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(by original designation); Recent, Indo-Pacific.
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<emphasis id="F3DDEAB7FFA56D38FF1FFBB85B1CFBD9" bold="true" box="[151,282,1036,1062]" italics="true" pageId="24" pageNumber="125">Spectamen</emphasis>
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spec. 1
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(
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)
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<emphasis id="F3DDEAB7FFA56D38FF1FFBC25B00FB70" bold="true" box="[151,262,1142,1167]" pageId="24" pageNumber="125">Material.</emphasis>
AndaDeVos (1); Tiep 3 (1).
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<emphasis id="F3DDEAB7FFA56D38FF4FFB2E5B9CFB4C" bold="true" box="[199,410,1178,1203]" pageId="24" pageNumber="125">Characterization.</emphasis>
Material probably consists of juveniles; shell low, H
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; P/T boundary thin slightly raised axial rib; T with 6 primary low rounded spiral ribs; body whorl with 3 keels; aperture trapezoid; umbilical ridge beaded; umbilicus wide.
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Genus
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<bibRefCitation id="A5384B54FFA56D38FEF1FCEA581DFC87" author="Iredale" box="[377,539,862,888]" pageId="24" pageNumber="125" refString="Iredale, T. (1924) Results from Roy Bell's molluscan collections. Proceedings of the Linnean Society of New South Wales, 49, 179 - 279." type="journal article" year="1924">Iredale, 1924</bibRefCitation>
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species.
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<taxonomicName id="06A94D26FFA56D38FEA1FC11588AFC44" authority="Watson, 1880" authorityName="Watson" authorityYear="1880" box="[297,652,933,955]" class="Gastropoda" family="Trochidae" genus="Trochus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="24" pageNumber="125" phylum="Mollusca" rank="species" species="philippensis">
<emphasis id="F3DDEAB7FFA56D38FEA1FC115BFCFC44" box="[297,506,933,955]" italics="true" pageId="24" pageNumber="125">Trochus philippensis</emphasis>
<bibRefCitation id="A5384B54FFA56D38FD89FC12588AFC44" author="Watson" box="[513,652,933,955]" pageId="24" pageNumber="125" refString="Watson, R. B. (1880) Mollusca of the H. M. S. ' Challenger' Expedition, part v. Journal of the Linnean Society, 15, 87 - 126. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1880. tb 00346. x" type="journal article" year="1880">Watson, 1880</bibRefCitation>
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(by original designation); Recent, Indo-Pacific.
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<heading id="9A5E81C9FFA56D38FF1FFF2C59B4FF4D" bold="true" box="[151,946,152,178]" fontSize="11" level="1" pageId="24" pageNumber="125" reason="1">
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<taxonomicName id="06A94D26FFA56D38FF1FFF2C5923FF4D" ID-CoL="4Y4XN" authority="Poppe, Tagaro &amp; Dekker, 2006" authorityName="Poppe, Tagaro &amp; Dekker" authorityYear="2006" box="[151,805,152,178]" class="Gastropoda" family="Solariellidae" genus="Solariella" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="24" pageNumber="125" phylum="Mollusca" rank="species" species="segersi">
<emphasis id="F3DDEAB7FFA56D38FF1FFF2C5B0AFF4D" bold="true" box="[151,268,152,178]" italics="true" pageId="24" pageNumber="125">Solariella</emphasis>
cf.
<emphasis id="F3DDEAB7FFA56D38FEB1FF2C5B8AFF4D" bold="true" box="[313,396,152,178]" italics="true" pageId="24" pageNumber="125">segersi</emphasis>
(
<bibRefCitation id="A5384B54FFA56D38FE13FF2C591BFF4D" author="Poppe" box="[411,797,152,178]" pageId="24" pageNumber="125" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter" year="2006">Poppe, Tagaro &amp; Dekker, 2006</bibRefCitation>
)
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nov. comb.
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<paragraph id="C11636A5FFA56D38FF1FFF0E5B4CFF2C" blockId="24.[151,946,152,211]" box="[151,330,186,211]" pageId="24" pageNumber="125">
(
<figureCitation id="59922A20FFA56D38FF16FF0E5B45FF2C" box="[158,323,186,211]" captionStart="FIGURES 56 – 58" captionStartId="25.[151,264,1887,1909]" captionTargetBox="[228,1363,195,1865]" captionTargetId="figure@25.[226,1366,193,1868]" captionTargetPageId="25" captionText="FIGURES 56 – 58. Solariellidae. 56. Solariella cf. segersi (Poppe, Tagaro &amp; Dekker, 2006). RGM 608.348. Locality Anda 5. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 57. Solariella cf. segersi (Poppe, Tagaro &amp; Dekker, 2006). RGM 608.349. Locality Anda 5. (a) apertural view, (b) basal view, (c) apical view. 58. Spectamen spec. 1. RGM 608.350. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view, (e) SEM, side view, (f) SEM, oblique view, scale bar in a – f 500 µm, (g) SEM detail of protoconch." httpUri="https://zenodo.org/record/250971/files/figure.png" pageId="24" pageNumber="125">Figures 56–57</figureCitation>
)
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cf. 2006
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<emphasis id="F3DDEAB7FFA56D38FF75FEB65B9DFEE7" box="[253,411,258,280]" italics="true" pageId="24" pageNumber="125">Minolia segersi</emphasis>
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nov. spec.—Poppe
<emphasis id="F3DDEAB7FFA56D38FDECFEB7589EFEE7" box="[612,664,258,280]" italics="true" pageId="24" pageNumber="125">et al.</emphasis>
: p. 132–133, pl. 72, figs 1–3. cf. 2009
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<emphasis id="F3DDEAB7FFA56D38FF75FE905B9DFEC5" box="[253,411,292,314]" italics="true" pageId="24" pageNumber="125">Minolia segersi</emphasis>
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—Vilvens: p. 94, figs 93–100.
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<emphasis id="F3DDEAB7FFA56D38FF1FFED25B00FE80" bold="true" box="[151,262,358,383]" pageId="24" pageNumber="125">Material.</emphasis>
Anda 5 (2).
</paragraph>
<paragraph id="C11636A5FFA56D38FF4FFE3E5E62FE14" blockId="24.[151,1436,358,779]" pageId="24" pageNumber="125">
<emphasis id="F3DDEAB7FFA56D38FF4FFE3E5B9CFE5C" bold="true" box="[199,410,394,419]" pageId="24" pageNumber="125">Characterization.</emphasis>
Shell conical, thick-walled, H
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W
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mm; protoconch damaged or missing in studied material; sculpture of spiral and axial ribs with low knobs; deep sutural depression; base of body whorl with spiral ribs; umbilicus open, encircled by grooves and with axial and spiral chords.
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<emphasis id="F3DDEAB7FFA56D38FF4FFE425B66FDF0" bold="true" box="[199,352,502,527]" pageId="24" pageNumber="125">Distribution.</emphasis>
Apart from the studied material,
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<emphasis id="F3DDEAB7FFA56D38FD6CFE4259ADFDF1" box="[740,939,502,527]" italics="true" pageId="24" pageNumber="125">Solariella segersi</emphasis>
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has been found in the
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between
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depth and has been reported from the East
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Sea at
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depth (
<bibRefCitation id="A5384B54FFA56D38FBA8FDAF5EE0FDCC" author="Poppe" box="[1056,1254,538,563]" pageId="24" pageNumber="125" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter" year="2006">
Poppe
<emphasis id="F3DDEAB7FFA56D38FBE6FDAF5EA0FDCC" box="[1134,1190,538,563]" italics="true" pageId="24" pageNumber="125">et al.</emphasis>
2006
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) as well as from Eastern
<collectingCountry id="B9BE7635FFA56D38FF7AFD8A5B59FDA8" box="[242,351,574,599]" name="Indonesia" pageId="24" pageNumber="125">Indonesia</collectingCountry>
from
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depth (
<bibRefCitation id="A5384B54FFA56D38FDFEFD8A5917FDA8" author="Vilvens" box="[630,785,574,599]" pageId="24" pageNumber="125" refString="Vilvens, C. (2009) New species and new records of Solariellidae (Gastropoda: Trochoidea) from Indonesia and Taiwan. Novapex: Trimestriel de la Societe Belge de Malacologie, 10 (3), 69 - 96." type="journal article" year="2009">Vilvens 2009</bibRefCitation>
).
</paragraph>
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<subSubSection id="89B3652EFFA56D38FF4FFDD65B6DFCF4" pageId="24" pageNumber="125" type="discussion">
<paragraph id="C11636A5FFA56D38FF4FFDD65B6DFCF4" blockId="24.[151,1436,358,779]" pageId="24" pageNumber="125">
<emphasis id="F3DDEAB7FFA56D38FF4FFDD65B3DFD84" bold="true" box="[199,315,610,635]" pageId="24" pageNumber="125">Remarks.</emphasis>
The species has been assigned to
<taxonomicName id="06A94D26FFA56D38FD5BFDD65947FD84" box="[723,833,610,635]" class="Gastropoda" family="Solariellidae" genus="Solariella" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="24" pageNumber="125" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA56D38FD5BFDD65947FD84" box="[723,833,610,635]" italics="true" pageId="24" pageNumber="125">Solariella</emphasis>
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on the basis of its moderately tall spire, turriculate outline, spiral sculpture, deep umbilicus and complete peristome, as well as genetic data (
<bibRefCitation id="A5384B54FFA56D38FB17FD325F97FD60" author="Williams" box="[1183,1425,646,671]" pageId="24" pageNumber="125" refString="Williams, S. T., Smith, L., Herbert, D. Marshall, B., Waren, A., Kiel, S., Dyal, P., Linse, K., Vilvens, C., &amp; Kano, Y. (2013) Cenozoic climate change and diversification on the continental shelf and slope: evolution of gastropod diversity in the family Solariellidae (Trochoidea). Ecology and Evolution, 3, 887 - 917. http: // dx. doi. org / 10.1002 / ece 3.513" type="journal article" year="2013">
Williams
<emphasis id="F3DDEAB7FFA56D38FA98FD335F4DFD60" box="[1296,1355,646,671]" italics="true" pageId="24" pageNumber="125">et al.</emphasis>
2013
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). The shells in the studied material do resemble
<taxonomicName id="06A94D26FFA56D38FD22FD1E5976FD3D" box="[682,880,682,707]" class="Gastropoda" family="Solariellidae" genus="Solariella" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="24" pageNumber="125" phylum="Mollusca" rank="species" species="segersi">
<emphasis id="F3DDEAB7FFA56D38FD22FD1E5976FD3D" box="[682,880,682,707]" italics="true" pageId="24" pageNumber="125">Solariella segersi</emphasis>
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, but the vertical arrangement of the spiral ribs is slightly different: the lower spiral rib is situated slightly lower in the fossil material so that it becomes visible only on the body whorl.
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Genus
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<emphasis id="F3DDEAB7FFA76D3AFF62F8EF5B77F88B" bold="true" box="[234,369,1883,1908]" italics="true" pageId="26" pageNumber="127">Guildfordia</emphasis>
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<paragraph id="C11636A5FFA76D3AFF1FF8105EFAF842" blockId="26.[151,1276,1956,1981]" box="[151,1276,1956,1981]" pageId="26" pageNumber="127">
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<typeStatus id="1E128807FFA76D3AFF1FF8105AD4F842" box="[151,210,1956,1981]" pageId="26" pageNumber="127">Type</typeStatus>
species.
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<taxonomicName id="06A94D26FFA76D3AFEB0F81058B8F843" authority="Philippi, 1841" authorityName="Philippi" authorityYear="1841" box="[312,702,1956,1981]" class="Gastropoda" family="Trochidae" genus="Trochus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="26" pageNumber="127" phylum="Mollusca" rank="species" species="triumphans">
<emphasis id="F3DDEAB7FFA76D3AFEB0F810581FF842" box="[312,537,1956,1981]" italics="true" pageId="26" pageNumber="127">Trochus triumphans</emphasis>
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(by subsequent designation); Recent, Indo-Pacific.
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<paragraph id="C11636A5FFA76D3AFF1FFBDA5885FB77" blockId="26.[151,643,1134,1194]" box="[151,643,1134,1160]" pageId="26" pageNumber="127">
<heading id="9A5E81C9FFA76D3AFF1FFBDA5885FB77" bold="true" box="[151,643,1134,1160]" fontSize="11" level="1" pageId="26" pageNumber="127" reason="1">
<emphasis id="F3DDEAB7FFA76D3AFF1FFBDA5885FB77" bold="true" box="[151,643,1134,1160]" pageId="26" pageNumber="127">
<taxonomicName id="06A94D26FFA76D3AFF1FFBDA5BF3FB77" authority="Vilvens, 2009" authorityName="Vilvens" authorityYear="2009" box="[151,501,1134,1160]" class="Gastropoda" family="Trochidae" genus="Ilanga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="26" pageNumber="127" phylum="Mollusca" rank="species" species="konos">
<emphasis id="F3DDEAB7FFA76D3AFF1FFBDA5B30FB77" bold="true" box="[151,310,1134,1160]" italics="true" pageId="26" pageNumber="127">Ilanga konos</emphasis>
(
<bibRefCitation id="A5384B54FFA76D3AFECDFBDA5BEBFB77" author="Vilvens" box="[325,493,1134,1160]" pageId="26" pageNumber="127" refString="Vilvens, C. (2009) New species and new records of Solariellidae (Gastropoda: Trochoidea) from Indonesia and Taiwan. Novapex: Trimestriel de la Societe Belge de Malacologie, 10 (3), 69 - 96." type="journal article" year="2009">Vilvens, 2009</bibRefCitation>
)
</taxonomicName>
nov. comb.
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<paragraph id="C11636A5FFA76D3AFF1FFB255B4CFB55" blockId="26.[151,643,1134,1194]" box="[151,330,1169,1194]" pageId="26" pageNumber="127">
(
<figureCitation id="59922A20FFA76D3AFF16FB255B45FB55" box="[158,323,1169,1194]" captionStart="FIGURES 64 – 68" captionStartId="28.[151,264,1845,1867]" captionTargetBox="[240,1348,195,1825]" captionTargetId="figure@28.[238,1351,193,1828]" captionTargetPageId="28" captionText="FIGURES 64 – 68. Solariellidae. 64. Ilanga konos (Vilvens, 2009). RGM 608.366. Locality Anda 1. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 65. Ilanga konos (Vilvens, 2009). RGM 608.367. Locality Tiep 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 66. Ilanga konos (Vilvens, 2009). RGM 608.368. Locality Tiep 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 67. Ilanga konos (Vilvens, 2009). RGM 608.369. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 68. Ilanga konos (Vilvens, 2009). RGM 608.370. Locality Anda 1. SEM detail of protoconch." httpUri="https://zenodo.org/record/250973/files/figure.png" pageId="26" pageNumber="127">Figures 64–68</figureCitation>
)
</paragraph>
<paragraph id="C11636A5FFA76D3AFF1FFB6D5932FB10" blockId="26.[151,820,1241,1263]" box="[151,820,1241,1263]" pageId="26" pageNumber="127">
2009
<taxonomicName id="06A94D26FFA76D3AFF7DFB6E5BA5FB10" box="[245,419,1241,1263]" class="Gastropoda" family="Solariellidae" genus="Microgaza" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="26" pageNumber="127" phylum="Mollusca" rank="species" species="konos">
<emphasis id="F3DDEAB7FFA76D3AFF7DFB6E5BA5FB10" box="[245,419,1241,1263]" italics="true" pageId="26" pageNumber="127">Microgaza konos</emphasis>
</taxonomicName>
nov. spec.—Vilvens: p. 88, figs 64–66.
</paragraph>
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<subSubSection id="89B3652EFFA76D3AFF1FFAA85F60FA5F" pageId="26" pageNumber="127" type="materials_examined">
<paragraph id="C11636A5FFA76D3AFF1FFAA859DEFACB" blockId="26.[151,1436,1308,1656]" box="[151,984,1308,1333]" pageId="26" pageNumber="127">
<emphasis id="F3DDEAB7FFA76D3AFF1FFAA85B00FACA" bold="true" box="[151,262,1308,1333]" pageId="26" pageNumber="127">Material.</emphasis>
Anda 1 (5); Anda 2 (2); Anda 6 (1); AndaDeVos (9); Tiep 2 (3).
</paragraph>
<paragraph id="C11636A5FFA76D3AFF4FFA8B5F60FA5F" blockId="26.[151,1436,1308,1656]" pageId="26" pageNumber="127">
<emphasis id="F3DDEAB7FFA76D3AFF4FFA8B5B9CFAA7" bold="true" box="[199,410,1343,1368]" pageId="26" pageNumber="127">Characterization.</emphasis>
Shell broad and umbilicate H
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,
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W
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mm; P with distinct spiral ribs and axially arranged granulate microsculpture, DN
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; P/T boundary sharp at 1.1 whorls; four spiral ribs on early T whorls, later whorls smooth; pinkish blotches; umbilical ridge narrow, four spiral ribs or more in umbilicus.
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<paragraph id="C11636A5FFA76D3AFF4FFA185956FA17" blockId="26.[151,1436,1308,1656]" pageId="26" pageNumber="127">
<emphasis id="F3DDEAB7FFA76D3AFF4FFA185B66FA3A" bold="true" box="[199,352,1452,1477]" pageId="26" pageNumber="127">Distribution.</emphasis>
Apart from the studied material,
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<emphasis id="F3DDEAB7FFA76D3AFD7EFA18598AFA3A" box="[758,908,1452,1477]" italics="true" pageId="26" pageNumber="127">Ilanga konos</emphasis>
</taxonomicName>
is known only from the
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locality Bashi channel, south
<collectingCountry id="B9BE7635FFA76D3AFEB7FA7B5B92FA17" box="[319,404,1487,1512]" name="Taiwan" pageId="26" pageNumber="127">Taiwan</collectingCountry>
, from a depth of
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(
<bibRefCitation id="A5384B54FFA76D3AFD21FA7B5942FA17" author="Vilvens" box="[681,836,1487,1512]" pageId="26" pageNumber="127" refString="Vilvens, C. (2009) New species and new records of Solariellidae (Gastropoda: Trochoidea) from Indonesia and Taiwan. Novapex: Trimestriel de la Societe Belge de Malacologie, 10 (3), 69 - 96." type="journal article" year="2009">Vilvens 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFA76D3AFF4FFA405EA2F987" pageId="26" pageNumber="127" type="discussion">
<paragraph id="C11636A5FFA76D3AFF4FFA405EA2F987" blockId="26.[151,1436,1308,1656]" pageId="26" pageNumber="127">
<emphasis id="F3DDEAB7FFA76D3AFF4FFA405B3DF9F2" bold="true" box="[199,315,1524,1549]" pageId="26" pageNumber="127">Remarks.</emphasis>
This species has been reassigned to
<taxonomicName id="06A94D26FFA76D3AFD55FA405920F9F2" box="[733,806,1524,1549]" class="Gastropoda" family="Trochidae" genus="Ilanga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="26" pageNumber="127" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA76D3AFD55FA405920F9F2" box="[733,806,1524,1549]" italics="true" pageId="26" pageNumber="127">Ilanga</emphasis>
</taxonomicName>
on the basis of genetic data that shows that all western Pacific and Indian Ocean ‘
<emphasis id="F3DDEAB7FFA76D3AFE48F9AC5847F9D0" box="[448,577,1560,1583]" italics="true" pageId="26" pageNumber="127">Microgaza’</emphasis>
like species, such as
<taxonomicName id="06A94D26FFA76D3AFCB9F9AD59F7F9CF" box="[817,1009,1559,1584]" class="Gastropoda" family="Solariellidae" genus="Microgaza" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="26" pageNumber="127" phylum="Mollusca" rank="species" species="konos">
<emphasis id="F3DDEAB7FFA76D3AFCB9F9AD59F7F9CF" box="[817,1009,1559,1584]" italics="true" pageId="26" pageNumber="127">Microgaza konos</emphasis>
</taxonomicName>
, fall into a clade with the
<typeStatus id="1E128807FFA76D3AFA9CF9AC5F42F9CF" box="[1300,1348,1560,1584]" pageId="26" pageNumber="127">type</typeStatus>
species of
<taxonomicName id="06A94D26FFA76D3AFF30F9885808F9AB" authority="Williams et al. 2013" authorityName="Williams et al." authorityYear="2013" box="[184,526,1596,1621]" class="Gastropoda" family="Trochidae" genus="Ilanga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="26" pageNumber="127" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA76D3AFF30F9885B07F9AA" box="[184,257,1596,1621]" italics="true" pageId="26" pageNumber="127">Ilanga</emphasis>
(
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Williams
<emphasis id="F3DDEAB7FFA76D3AFE0CF9895BB9F9AA" box="[388,447,1596,1621]" italics="true" pageId="26" pageNumber="127">et al.</emphasis>
2013
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)
</taxonomicName>
. Both
<bibRefCitation id="A5384B54FFA76D3AFDD5F9885901F9AB" author="Herbert" box="[605,775,1596,1621]" pageId="26" pageNumber="127" refString="Herbert, D. G. (1987) Revision of the Solariellinae (Mollusca: Prosobranchia: Trochidae) in southern Africa. Annals of the Natal Museum, 28, 283 - 382." type="journal article" year="1987">Herbert (1987)</bibRefCitation>
and
<bibRefCitation id="A5384B54FFA76D3AFCCBF98859FDF9AB" author="Marshall" box="[835,1019,1596,1621]" pageId="26" pageNumber="127" refString="Marshall, B. A. (1999) A revision of the recent Solariellinae (Gastropoda: Trochoidea) of the New Zealand Region. The Nautilus, 113, 4 - 42." type="journal article" year="1999">
<collectingCountry id="B9BE7635FFA76D3AFCCBF98859A0F9AA" box="[835,934,1596,1621]" name="Marshall Islands" pageId="26" pageNumber="127">Marshall</collectingCountry>
(1999)
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recognize
<taxonomicName id="06A94D26FFA76D3AFBF4F9885EC3F9AA" box="[1148,1221,1596,1621]" class="Gastropoda" family="Trochidae" genus="Ilanga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="26" pageNumber="127" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA76D3AFBF4F9885EC3F9AA" box="[1148,1221,1596,1621]" italics="true" pageId="26" pageNumber="127">Ilanga</emphasis>
</taxonomicName>
and
<taxonomicName id="06A94D26FFA76D3AFA8AF9895F7CF9AB" box="[1282,1402,1597,1620]" class="Gastropoda" family="Solariellidae" genus="Microgaza" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="26" pageNumber="127" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA76D3AFA8AF9895F7CF9AB" box="[1282,1402,1597,1620]" italics="true" pageId="26" pageNumber="127">Microgaza</emphasis>
</taxonomicName>
as distinct genera, but additional genetic studies are needed to confirm the delimitation of each.
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<emphasis id="F3DDEAB7FFA76D3AFEA1FBBC5BF2FBE1" box="[297,500,1032,1054]" italics="true" pageId="26" pageNumber="127">Trochus laevissimus</emphasis>
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von
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(by original desination); Recent,
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(
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nov. comb.
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(
<figureCitation id="59922A20FFA76D3AFF16FED35B44FE7F" box="[158,322,359,384]" captionStart="FIGURES 59 – 63" captionStartId="27.[151,264,1860,1882]" captionTargetBox="[233,1351,197,1843]" captionTargetId="figure@27.[231,1354,195,1845]" captionTargetPageId="27" captionText="FIGURES 59 – 63. Solariellidae. 59. Archiminolia spec. 1. RGM 608.352. Locality AndaDeVos. (a) rear view, (b) basal view, (c) apical view. 60. Archiminolia spec. 1. RGM 608.353. Locality AndaDeVos. SEM detail of protoconch. 61. Zetela tabakotanii (Poppe, Tagaro &amp; Dekker, 2006). RGM 608.355. Locality Anda 1. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 62. Zetela tabakotanii (Poppe, Tagaro &amp; Dekker, 2006). RGM 608.356. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 63. Zetela tabakotanii (Poppe, Tagaro &amp; Dekker, 2006). RGM 608.357. Locality Anda 1. (a) SEM, apical view, (b) SEM detail of protoconch." httpUri="https://zenodo.org/record/250972/files/figure.png" pageId="26" pageNumber="127">Figures 61–63</figureCitation>
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2006
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<emphasis id="F3DDEAB7FFA76D3AFF79FE045BBCFE39" box="[241,442,432,454]" italics="true" pageId="26" pageNumber="127">Minolia tabakotanii</emphasis>
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nov. spec.—Poppe
<emphasis id="F3DDEAB7FFA76D3AFD0CFE0558B1FE39" box="[644,695,432,454]" italics="true" pageId="26" pageNumber="127">et al.</emphasis>
: p. 133, pl. 68, fig. 1. 2008d
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<emphasis id="F3DDEAB7FFA76D3AFF79FE7A5BBCFE1B" box="[241,442,462,484]" italics="true" pageId="26" pageNumber="127">Minolia tabakotanii</emphasis>
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—Poppe &amp; Tagaro: p. 224, pl. 57, fig. 2.
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<emphasis id="F3DDEAB7FFA76D3AFF1FFDA55B00FDD5" bold="true" box="[151,262,529,554]" pageId="26" pageNumber="127">Material.</emphasis>
Anda 1 (12); Anda 2 (14); Anda 3 (7); Anda 4 (26); Anda 5 (3); Anda 6 (2); AndaDeVos (4); AndaClif1 (3).
</paragraph>
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<emphasis id="F3DDEAB7FFA76D3AFF4FFDED5B9CFD8D" bold="true" box="[199,410,601,626]" pageId="26" pageNumber="127">Characterization.</emphasis>
Small turbiniform shell with rounded whorls, H
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,
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W
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mm); P smooth, globular, DN 0.10–0.11; P/T boundary at 1.2 whorls marked by indistinct rib; T reticulate ornamentation with axial lamellae and microscopic granules in interspaces; aperture circular; narrow umbilicus.
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<emphasis id="F3DDEAB7FFA76D3AFF4FFD715B66FD21" bold="true" box="[199,352,709,734]" pageId="26" pageNumber="127">Distribution.</emphasis>
Apart from the studied material, known only from the
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locality offshore Mactan
<collectingCountry id="B9BE7635FFA76D3AFAD8FD715F91FD21" box="[1360,1431,709,734]" name="Iceland" pageId="26" pageNumber="127">Island</collectingCountry>
,
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from
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depth (
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Poppe
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2006
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;
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).
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<emphasis id="F3DDEAB7FFA76D3AFF4FFCB95B3DFCD9" bold="true" box="[199,315,781,806]" pageId="26" pageNumber="127">Remarks.</emphasis>
This species is assigned to
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<emphasis id="F3DDEAB7FFA76D3AFDF4FCB958C4FCD9" box="[636,706,781,806]" italics="true" pageId="26" pageNumber="127">Zetela</emphasis>
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on the basis of the similarity of its sculpture to that of the
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species
<taxonomicName id="06A94D26FFA76D3AFF67FC865884FCB5" authority="Murdoch &amp; Suter, 1906" authorityName="Murdoch &amp; Suter" authorityYear="1906" box="[239,642,817,842]" class="Gastropoda" family="Solariellidae" genus="Minolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="26" pageNumber="127" phylum="Mollusca" rank="species" species="textilis">
<emphasis id="F3DDEAB7FFA76D3AFF67FC865B5EFCB5" box="[239,344,817,842]" italics="true" pageId="26" pageNumber="127">Z. textilis</emphasis>
(
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. It differs from that species by having a much smaller umbilicus and a more continuous peristome.
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<emphasis id="F3DDEAB7FFA96D34FF1FF8A45827F8D5" bold="true" box="[151,545,1808,1834]" pageId="20" pageNumber="121">
Genus
<taxonomicName id="06A94D26FFA96D34FF66F8A45827F8D5" ID-CoL="7NZ6R" authority="Iredale, 1924" authorityName="Iredale" authorityYear="1924" box="[238,545,1808,1834]" class="Gastropoda" family="Trochidae" genus="Ethminolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA96D34FF66F8A45B71F8D5" bold="true" box="[238,375,1808,1834]" italics="true" pageId="20" pageNumber="121">Ethminolia</emphasis>
<bibRefCitation id="A5384B54FFA96D34FEF6F8A45827F8D5" author="Iredale" box="[382,545,1808,1834]" pageId="20" pageNumber="121" refString="Iredale, T. (1924) Results from Roy Bell's molluscan collections. Proceedings of the Linnean Society of New South Wales, 49, 179 - 279." type="journal article" year="1924">Iredale, 1924</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFA96D34FF1FF8E25E13F893" box="[151,1045,1878,1900]" pageId="20" pageNumber="121" type="materials_examined">
<paragraph id="C11636A5FFA96D34FF1FF8E25E13F893" blockId="20.[151,1045,1878,1900]" box="[151,1045,1878,1900]" pageId="20" pageNumber="121">
<emphasis id="F3DDEAB7FFA96D34FF1FF8E25B24F893" bold="true" box="[151,290,1878,1900]" pageId="20" pageNumber="121">
<typeStatus id="1E128807FFA96D34FF1FF8E25ACAF893" box="[151,204,1878,1900]" pageId="20" pageNumber="121">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="06A94D26FFA96D34FEA1F8E25895F893" authority="Iredale, 1924" authorityName="Iredale" authorityYear="1924" box="[297,659,1878,1900]" class="Gastropoda" family="Trochidae" genus="Ethminolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="probabilis">
<emphasis id="F3DDEAB7FFA96D34FEA1F8E25801F893" box="[297,519,1878,1900]" italics="true" pageId="20" pageNumber="121">Ethminolia probabilis</emphasis>
<bibRefCitation id="A5384B54FFA96D34FD85F8E25895F893" author="Iredale" box="[525,659,1878,1900]" pageId="20" pageNumber="121" refString="Iredale, T. (1924) Results from Roy Bell's molluscan collections. Proceedings of the Linnean Society of New South Wales, 49, 179 - 279." type="journal article" year="1924">Iredale, 1924</bibRefCitation>
</taxonomicName>
(by monotypy); Recent, Indo-Pacific.
</paragraph>
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243
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<mods:title id="C4B3ECF8B811A3C2C67A34D84192A62C">On some Vetigastropoda (Mollusca, Gastropoda) from the Plio-Pleistocene of the Philippines with descriptions of three new species</mods:title>
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<paragraph id="C11636A5FFA96D34FF1FFCD4583CFC85" blockId="20.[151,570,864,923]" box="[151,570,864,890]" pageId="20" pageNumber="121">
<heading id="9A5E81C9FFA96D34FF1FFCD4583CFC85" bold="true" box="[151,570,864,890]" fontSize="11" level="1" pageId="20" pageNumber="121" reason="1">
<taxonomicName id="06A94D26FFA96D34FF1FFCD4583CFC85" ID-CoL="XTRJ" authority="Gould, 1861" authorityName="Gould" authorityYear="1861" box="[151,570,864,890]" class="Gastropoda" family="Trochidae" genus="Conotalopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="musiva">
<emphasis id="F3DDEAB7FFA96D34FF1FFCD4583CFC85" bold="true" box="[151,570,864,890]" pageId="20" pageNumber="121">
<emphasis id="F3DDEAB7FFA96D34FF1FFCD45B8EFC85" bold="true" box="[151,392,864,890]" italics="true" pageId="20" pageNumber="121">Conotalopia musiva</emphasis>
(
<bibRefCitation id="A5384B54FFA96D34FE1FFCD45834FC85" author="Gould" box="[407,562,864,890]" pageId="20" pageNumber="121" refString="Gould, A. A. (1861) Descriptions of shells collected by the North Pacific Exploring Expedition. Proceedings of the Boston Society of Natural History, 8, 14 - 40." type="journal article" year="1861">Gould, 1861</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C11636A5FFA96D34FF1FFC365B4DFC64" blockId="20.[151,570,864,923]" box="[151,331,898,923]" pageId="20" pageNumber="121">
(
<figureCitation id="59922A20FFA96D34FF16FC365B44FC64" box="[158,322,898,923]" captionStart="FIGURES 45 – 52" captionStartId="21.[151,264,1857,1879]" captionTargetBox="[232,1350,195,1840]" captionTargetId="figure@21.[230,1353,193,1843]" captionTargetPageId="21" captionText="FIGURES 45 – 52. Trochidae. 45. Pseudotalopia taiwanensis Chen, 2006. RGM 608.316. Locality Anda 3. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 46. Pseudotalopia taiwanensis Chen, 2006. RGM 608.317. Locality Tiep 3. (a) rear view, (b) apertural view, (c) basal view, (d) apical view, (e) detail protoconch. 47. Pseudotalopia taiwanensis Chen, 2006. RGM 608.318. Locality Tiep 2. (a) rear view, (b) detail protoconch. 48. Conotalopia musiva (Gould, 1861). RGM 608.319. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 49. Conotalopia musiva (Gould, 1861). RGM 608.320. Locality AndaDeVos. (a) SEM, side view, (b) SEM detail of protoconch." httpUri="https://zenodo.org/record/250969/files/figure.png" pageId="20" pageNumber="121">Figures 48–49</figureCitation>
)
</paragraph>
<paragraph id="C11636A5FFA96D34FF1FFC7E58B5FC1F" blockId="20.[151,980,970,1115]" box="[151,691,970,992]" pageId="20" pageNumber="121">
1861
<taxonomicName id="06A94D26FFA96D34FF7BFC7F5BACFC1F" box="[243,426,971,992]" class="Gastropoda" family="Trochidae" genus="Margarita" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="musiva">
<emphasis id="F3DDEAB7FFA96D34FF7BFC7F5BACFC1F" box="[243,426,971,992]" italics="true" pageId="20" pageNumber="121">Margarita musiva</emphasis>
</taxonomicName>
nov. spec.—Gould: p. 15.
</paragraph>
<paragraph id="C11636A5FFA96D34FF1FFC5D588DFBE1" blockId="20.[151,980,970,1115]" pageId="20" pageNumber="121">
1989
<taxonomicName id="06A94D26FFA96D34FF7BFC5D5BE7FC00" box="[243,481,1001,1023]" class="Gastropoda" family="Solariellidae" genus="Minolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="holdsworthana">
<emphasis id="F3DDEAB7FFA96D34FF7BFC5D5BE7FC00" box="[243,481,1001,1023]" italics="true" pageId="20" pageNumber="121">Minolia holdsworthana</emphasis>
</taxonomicName>
(G. &amp;
<bibRefCitation id="A5384B54FFA96D34FDAEFC5E58C7FC00" author="Nevill" box="[550,705,1001,1023]" pageId="20" pageNumber="121" refString="Nevill, G. &amp; Nevill, H. (1871) Descriptions of new Mollusca from the eastern regions. Journal of the Asiatic Society of Bengal, 40, 1 - 11." type="journal article" year="1871">H. Nevill, 1871</bibRefCitation>
)—Herbert: p. 370, fig. 1h. 1999
<taxonomicName id="06A94D26FFA96D34FF7BFBBC5BC6FBE1" box="[243,448,1032,1054]" class="Gastropoda" family="Trochidae" genus="Conotalopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="musiva">
<emphasis id="F3DDEAB7FFA96D34FF7BFBBC5BC6FBE1" box="[243,448,1032,1054]" italics="true" pageId="20" pageNumber="121">Conotalopia musiva</emphasis>
</taxonomicName>
—Higo
<emphasis id="F3DDEAB7FFA96D34FD9AFBBD5843FBE1" box="[530,581,1032,1054]" italics="true" pageId="20" pageNumber="121">et al.</emphasis>
: p. 67.
</paragraph>
<paragraph id="C11636A5FFA96D34FF1FFB925975FBA4" blockId="20.[151,980,970,1115]" pageId="20" pageNumber="121">
2006
<taxonomicName id="06A94D26FFA96D34FF7BFB925BDFFBC2" box="[243,473,1062,1085]" class="Gastropoda" family="Trochidae" genus="Pseudominolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="musiva">
<emphasis id="F3DDEAB7FFA96D34FF7BFB925BDFFBC2" box="[243,473,1062,1085]" italics="true" pageId="20" pageNumber="121">Pseudominolia musiva</emphasis>
</taxonomicName>
—Poppe
<emphasis id="F3DDEAB7FFA96D34FDB0FB9C586DFBC3" box="[568,619,1062,1085]" italics="true" pageId="20" pageNumber="121">et al.</emphasis>
: p. 109, pl. 58, fig. 3. 2008c
<taxonomicName id="06A94D26FFA96D34FF7BFBF15BDFFBA4" box="[243,473,1093,1115]" class="Gastropoda" family="Trochidae" genus="Pseudominolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="musiva">
<emphasis id="F3DDEAB7FFA96D34FF7BFBF15BDFFBA4" box="[243,473,1093,1115]" italics="true" pageId="20" pageNumber="121">Pseudominolia musiva</emphasis>
</taxonomicName>
—Poppe &amp; Tagaro: p. 200, pl. 45, fig. 2.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFA96D34FF1FFB335E95FAF2" pageId="20" pageNumber="121" type="materials_examined">
<paragraph id="C11636A5FFA96D34FF1FFB335BC4FB5F" blockId="20.[151,1436,1159,1725]" box="[151,450,1159,1184]" pageId="20" pageNumber="121">
<emphasis id="F3DDEAB7FFA96D34FF1FFB335B00FB5F" bold="true" box="[151,262,1159,1184]" pageId="20" pageNumber="121">Material.</emphasis>
AndaDeVos (2).
</paragraph>
<paragraph id="C11636A5FFA96D34FF4FFB185E95FAF2" blockId="20.[151,1436,1159,1725]" pageId="20" pageNumber="121">
<emphasis id="F3DDEAB7FFA96D34FF4FFB185B9CFB3A" bold="true" box="[199,410,1196,1221]" pageId="20" pageNumber="121">Characterization.</emphasis>
Shell minute, H
<quantity id="06519B40FFA96D34FDD3FB1858B5FB3B" box="[603,691,1196,1220]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="20" pageNumber="121" unit="mm" value="0.7">0.7 mm</quantity>
,
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W
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mm; DN
<quantity id="06519B40FFA96D34FCFFFB1859D9FB3B" box="[887,991,1196,1220]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="121" unit="mm" value="0.09">0.09 mm</quantity>
; P terminated by well defined, rounded varix at 1.1. whorls; T whorls with a sharp keel and covered by fine, axially arranged granules; body whorl with 3 sharp spiral ribs; base broad and flat; umbilicus narrow, umbilical ridge weakly granulated.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFA96D34FF4FFAA35E73FAAA" pageId="20" pageNumber="121" type="distribution">
<paragraph id="C11636A5FFA96D34FF4FFAA35E73FAAA" blockId="20.[151,1436,1159,1725]" pageId="20" pageNumber="121">
<emphasis id="F3DDEAB7FFA96D34FF4FFAA35B66FACF" bold="true" box="[199,352,1303,1328]" pageId="20" pageNumber="121">Distribution.</emphasis>
Reported from Bohol,
<collectingCountry id="B9BE7635FFA96D34FDECFAA358E3FACF" box="[612,741,1303,1328]" name="Philippines" pageId="20" pageNumber="121">Philippines</collectingCountry>
,
<quantity id="06519B40FFA96D34FD78FAAC592FFAD0" box="[752,809,1304,1328]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="121" unit="m" value="38.0">38 m</quantity>
depth, Wakasa Bay,
<collectingCountry id="B9BE7635FFA96D34FB9AFAAC5E53FAD0" box="[1042,1109,1304,1327]" name="Japan" pageId="20" pageNumber="121">Japan</collectingCountry>
,
<quantity id="06519B40FFA96D34FBD7FAA35ED6FAD0" box="[1119,1232,1303,1328]" metricMagnitude="2" metricUnit="m" metricValue="1.005" metricValueMax="1.61" metricValueMin="0.4" pageId="20" pageNumber="121" unit="m" value="100.5" valueMax="161.0" valueMin="40.0">40–161 m</quantity>
depth (
<bibRefCitation id="A5384B54FFA96D34FAACFAA35AD5FAAB" author="Higo" pageId="20" pageNumber="121" refString="Higo, S., Callomon, P. &amp; Goto, Y. (1999) Catalogue and bibliography of the marine shell-bearing Mollusca of Japan. Elle Scientific Publications, Osaka, 749 pp." type="book" year="1999">
Higo
<emphasis id="F3DDEAB7FFA96D34FAECFAAD5F9AFACF" box="[1380,1436,1303,1328]" italics="true" pageId="20" pageNumber="121">et al.</emphasis>
1999
</bibRefCitation>
), and
<collectingCountry id="B9BE7635FFA96D34FE9DFA895B9AFAAA" box="[277,412,1341,1365]" name="Hong Kong" pageId="20" pageNumber="121">Hong Kong</collectingCountry>
,
<collectingCountry id="B9BE7635FFA96D34FE2FFA88580FFAAA" box="[423,521,1340,1365]" name="Pakistan" pageId="20" pageNumber="121">Pakistan</collectingCountry>
,
<collectingCountry id="B9BE7635FFA96D34FD9CFA885883FAAA" box="[532,645,1340,1365]" name="Sri Lanka" pageId="20" pageNumber="121">Sri Lanka</collectingCountry>
,
<collectingCountry id="B9BE7635FFA96D34FD07FA8858F0FAAA" box="[655,758,1340,1365]" name="Malaysia" pageId="20" pageNumber="121">Malaysia</collectingCountry>
and
<collectingCountry id="B9BE7635FFA96D34FCA4FA885996FAAA" box="[812,912,1340,1365]" name="Thailand" pageId="20" pageNumber="121">Thailand</collectingCountry>
(
<bibRefCitation id="A5384B54FFA96D34FC16FA885E6FFAAA" author="Poppe" box="[926,1129,1340,1365]" pageId="20" pageNumber="121" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter" year="2006">
Poppe
<emphasis id="F3DDEAB7FFA96D34FC65FA895E20FAAA" box="[1005,1062,1340,1365]" italics="true" pageId="20" pageNumber="121">et al.</emphasis>
2006
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFA96D34FF4FFAEB5958F942" pageId="20" pageNumber="121" type="discussion">
<paragraph id="C11636A5FFA96D34FF4FFAEB5958F942" blockId="20.[151,1436,1159,1725]" pageId="20" pageNumber="121">
<emphasis id="F3DDEAB7FFA96D34FF4FFAEB5B3DFA87" bold="true" box="[199,315,1375,1400]" pageId="20" pageNumber="121">Remarks.</emphasis>
<taxonomicName id="06A94D26FFA96D34FECDFAEB5857FA87" box="[325,593,1375,1400]" class="Gastropoda" family="Solariellidae" genus="Minolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="holdsworthana">
<emphasis id="F3DDEAB7FFA96D34FECDFAEB5857FA87" box="[325,593,1375,1400]" italics="true" pageId="20" pageNumber="121">Minolia holdsworthana</emphasis>
</taxonomicName>
(G. &amp;
<bibRefCitation id="A5384B54FFA96D34FD21FAD45966FA87" author="Nevill" box="[681,864,1375,1400]" pageId="20" pageNumber="121" refString="Nevill, G. &amp; Nevill, H. (1871) Descriptions of new Mollusca from the eastern regions. Journal of the Asiatic Society of Bengal, 40, 1 - 11." type="journal article" year="1871">H. Nevill, 1871</bibRefCitation>
) was synonymized with
<taxonomicName id="06A94D26FFA96D34FB0FFAD55AC9FA63" authority="Melvill, 1918" authorityName="Melvill" authorityYear="1918" class="Gastropoda" family="Solariellidae" genus="Minolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="charmosyne">
<emphasis id="F3DDEAB7FFA96D34FB0FFAD55F31FA87" box="[1159,1335,1375,1400]" italics="true" pageId="20" pageNumber="121">M. charmosyne</emphasis>
<bibRefCitation id="A5384B54FFA96D34FACAFAEB5AC9FA63" author="Melvill" pageId="20" pageNumber="121" refString="Melvill, J. C. (1918) Descriptions of thirty-four species of marine mollusca from the Persian Gulf, Gulf of Oman and Arabian Sea, collected by Mr. F. W. Townsend. Annals and Magazine of Natural History, 1, 137 - 158." type="journal article" year="1918">Melvill, 1918</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="A5384B54FFA96D34FF72FA305BA3FA63" author="Herbert" box="[250,421,1412,1437]" pageId="20" pageNumber="121" refString="Herbert, D. G. (1989) Pagodatrochus, a new genus for Minolia variabilis H. Adams, 1873 (Gastropoda: Trochidae). Journal of Molluscan Studies, 55, 365 - 372. http: // dx. doi. org / 10.1093 / mollus / 55.3.365" type="journal article" year="1989">Herbert (1989)</bibRefCitation>
. These two species were synonymized with
<taxonomicName id="06A94D26FFA96D34FC12FA315EFFFA62" authority="Gould, 1861" authorityName="Gould" authorityYear="1861" box="[922,1273,1412,1437]" class="Gastropoda" family="Trochidae" genus="Margarita" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="musiva">
<emphasis id="F3DDEAB7FFA96D34FC12FA315E62FA63" box="[922,1124,1413,1436]" italics="true" pageId="20" pageNumber="121">Margarita musiva</emphasis>
<bibRefCitation id="A5384B54FFA96D34FBE4FA305EFFFA62" author="Gould" box="[1132,1273,1412,1437]" pageId="20" pageNumber="121" refString="Gould, A. A. (1861) Descriptions of shells collected by the North Pacific Exploring Expedition. Proceedings of the Boston Society of Natural History, 8, 14 - 40." type="journal article" year="1861">Gould, 1861</bibRefCitation>
</taxonomicName>
and attributed to
<taxonomicName id="06A94D26FFA96D34FF3EFA135B58FA3F" box="[182,350,1447,1472]" class="Gastropoda" family="Trochidae" genus="Pseudominolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA96D34FF3EFA135B58FA3F" box="[182,350,1447,1472]" italics="true" pageId="20" pageNumber="121">Pseudominolia</emphasis>
</taxonomicName>
by
<bibRefCitation id="A5384B54FFA96D34FE05FA1C586BFA3F" box="[397,621,1447,1472]" pageId="20" pageNumber="121" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter">
Poppe
<emphasis id="F3DDEAB7FFA96D34FE55FA1D581FFA3F" box="[477,537,1447,1472]" italics="true" pageId="20" pageNumber="121">et al.</emphasis>
(2006)
</bibRefCitation>
. The synonymy is considered here to be justified, based on the strong conchological similarity, but the allocation to
<taxonomicName id="06A94D26FFA96D34FD20FA785956FA1A" box="[680,848,1484,1509]" class="Gastropoda" family="Trochidae" genus="Pseudominolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA96D34FD20FA785956FA1A" box="[680,848,1484,1509]" italics="true" pageId="20" pageNumber="121">Pseudominolia</emphasis>
</taxonomicName>
by
<bibRefCitation id="A5384B54FFA96D34FC08FA795E58FA1A" box="[896,1118,1484,1509]" pageId="20" pageNumber="121" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter">
Poppe
<emphasis id="F3DDEAB7FFA96D34FC47FA795E0CFA1A" box="[975,1034,1484,1509]" italics="true" pageId="20" pageNumber="121">et al.</emphasis>
(2006)
</bibRefCitation>
is superfluous, because the species is already allocated to
<taxonomicName id="06A94D26FFA96D34FE67FA5B587DF9F7" box="[495,635,1519,1544]" class="Gastropoda" family="Trochidae" genus="Conotalopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA96D34FE67FA5B587DF9F7" box="[495,635,1519,1544]" italics="true" pageId="20" pageNumber="121">Conotalopia</emphasis>
</taxonomicName>
, according to
<bibRefCitation id="A5384B54FFA96D34FC96FA5B59EBF9F7" box="[798,1005,1519,1544]" pageId="20" pageNumber="121" refString="Higo, S., Callomon, P. &amp; Goto, Y. (1999) Catalogue and bibliography of the marine shell-bearing Mollusca of Japan. Elle Scientific Publications, Osaka, 749 pp." type="book">
Higo
<emphasis id="F3DDEAB7FFA96D34FCD6FA455991F9F7" box="[862,919,1519,1544]" italics="true" pageId="20" pageNumber="121">et al.</emphasis>
(1999)
</bibRefCitation>
. The allocation to
<taxonomicName id="06A94D26FFA96D34FB48FA5B5F4AF9F7" box="[1216,1356,1519,1544]" class="Gastropoda" family="Trochidae" genus="Conotalopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA96D34FB48FA5B5F4AF9F7" box="[1216,1356,1519,1544]" italics="true" pageId="20" pageNumber="121">Conotalopia</emphasis>
</taxonomicName>
is also preferred here, because
<taxonomicName id="06A94D26FFA96D34FE2FF9A1581EF9D3" box="[423,536,1557,1580]" class="Gastropoda" family="Trochidae" genus="Margarita" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="musiva">
<emphasis id="F3DDEAB7FFA96D34FE2FF9A1581EF9D3" box="[423,536,1557,1580]" italics="true" pageId="20" pageNumber="121">C. musiva</emphasis>
</taxonomicName>
shares similarities with other species of
<taxonomicName id="06A94D26FFA96D34FC63F9A05E71F9D2" box="[1003,1143,1556,1581]" class="Gastropoda" family="Trochidae" genus="Conotalopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA96D34FC63F9A05E71F9D2" box="[1003,1143,1556,1581]" italics="true" pageId="20" pageNumber="121">Conotalopia</emphasis>
</taxonomicName>
, such as a relatively high spire and strong spiral ribs. Furthermore, the apical beak of the protoconch, which is a diagnostic feature for
<taxonomicName id="06A94D26FFA96D34FF1FF9E85BF7F98B" authority="Herbert 1992" authorityName="Herbert" authorityYear="1992" box="[151,497,1628,1653]" class="Gastropoda" family="Trochidae" genus="Pseudominolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA96D34FF1FF9E85B39F98A" box="[151,319,1628,1653]" italics="true" pageId="20" pageNumber="121">Pseudominolia</emphasis>
(
<bibRefCitation id="A5384B54FFA96D34FEC6F9E85BEFF98B" author="Herbert" box="[334,489,1628,1653]" pageId="20" pageNumber="121" refString="Herbert, D. G. (1992) Revision of the Umboniinae in southern Africa and Mozambique (Mollusca: Prosobranchia: Trochidae). Annals of the Natal Museum, 33, 379 - 459." type="journal article" year="1992">Herbert 1992</bibRefCitation>
)
</taxonomicName>
, is very poorly developed in the studied material. Unfortunately, there are no SEM pictures of the protoconch of
<typeStatus id="1E128807FFA96D34FE70F934582EF967" box="[504,552,1664,1688]" pageId="20" pageNumber="121">type</typeStatus>
material of
<taxonomicName id="06A94D26FFA96D34FD36F9CB5E76F967" authority="Gould, 1861" authorityName="Gould" authorityYear="1861" box="[702,1136,1663,1688]" class="Gastropoda" family="Trochidae" genus="Pseudominolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="musiva">
<emphasis id="F3DDEAB7FFA96D34FD36F9CB59C7F967" box="[702,961,1663,1688]" italics="true" pageId="20" pageNumber="121">Pseudominolia musiva</emphasis>
(
<bibRefCitation id="A5384B54FFA96D34FC5DF9CB5E6EF967" author="Gould" box="[981,1128,1663,1688]" pageId="20" pageNumber="121" refString="Gould, A. A. (1861) Descriptions of shells collected by the North Pacific Exploring Expedition. Proceedings of the Boston Society of Natural History, 8, 14 - 40." type="journal article" year="1861">Gould, 1861</bibRefCitation>
)
</taxonomicName>
or the synonymized taxa available to verify the absence of the apical beak in the species.
</paragraph>
</subSubSection>
</treatment>
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81
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<paragraph id="C11636A5FFA96D34FF1FFD06582AFD33" blockId="20.[151,556,690,716]" box="[151,556,690,716]" pageId="20" pageNumber="121">
<heading id="9A5E81C9FFA96D34FF1FFD06582AFD33" bold="true" box="[151,556,690,716]" fontSize="11" level="1" pageId="20" pageNumber="121" reason="1">
<emphasis id="F3DDEAB7FFA96D34FF1FFD06582AFD33" bold="true" box="[151,556,690,716]" pageId="20" pageNumber="121">
Genus
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<emphasis id="F3DDEAB7FFA96D34FF66FD065B84FD33" bold="true" box="[238,386,690,716]" italics="true" pageId="20" pageNumber="121">Conotalopia</emphasis>
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</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFA96D34FF1FFD4D5E35FCF0" box="[151,1075,761,783]" pageId="20" pageNumber="121" type="materials_examined">
<paragraph id="C11636A5FFA96D34FF1FFD4D5E35FCF0" blockId="20.[151,1075,761,783]" box="[151,1075,761,783]" pageId="20" pageNumber="121">
<emphasis id="F3DDEAB7FFA96D34FF1FFD4D5B24FCF0" bold="true" box="[151,290,761,783]" pageId="20" pageNumber="121">
<typeStatus id="1E128807FFA96D34FF1FFD4D5ACAFCF0" box="[151,204,761,783]" pageId="20" pageNumber="121">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="06A94D26FFA96D34FEA1FD4D5868FCF0" authority="Melvill, 1891" authorityName="Melvill" authorityYear="1891" box="[297,622,761,783]" class="Gastropoda" family="Solariellidae" genus="Minolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="henniana">
<emphasis id="F3DDEAB7FFA96D34FEA1FD4D5BD8FCF0" box="[297,478,761,783]" italics="true" pageId="20" pageNumber="121">Minolia henniana</emphasis>
<bibRefCitation id="A5384B54FFA96D34FE6CFD4D5868FCF0" author="Melvill" box="[484,622,761,783]" pageId="20" pageNumber="121" refString="Melvill, J. C. (1891) Descriptions of eleven new species belonging to the genera Columbarium, Pisania, Minolia, Liotia and Solarium. Journal of Conchology, 6, 405 - 411." type="journal article" year="1891">Melvill, 1891</bibRefCitation>
</taxonomicName>
(by original designation); Recent,
<collectingCountry id="B9BE7635FFA96D34FC47FD4D5E29FCF0" box="[975,1071,761,783]" name="Australia" pageId="20" pageNumber="121">Australia</collectingCountry>
.
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<emphasis id="F3DDEAB7FFAB6D36FF1FFA775B6BFA22" bold="true" box="[151,365,1475,1501]" italics="true" pageId="22" pageNumber="123">Solariella chodon</emphasis>
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2009
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<emphasis id="F3DDEAB7FFAB6D36FF79F9995BA0F9BC" box="[241,422,1581,1603]" italics="true" pageId="22" pageNumber="123">Solariella chodon</emphasis>
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nov. spec.—Vilvens: p. 74–75, figs 11–16.
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<emphasis id="F3DDEAB7FFAB6D36FF1FF9DB5B00F977" bold="true" box="[151,262,1647,1672]" pageId="22" pageNumber="123">Material.</emphasis>
Anda 1 (19); Anda 2 (13); Anda 3 (7); Anda 4 (2); AndaClif 1 (1); Roxas (1); Tiep 1 (1).
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<emphasis id="F3DDEAB7FFAB6D36FF4FF9205B9CF952" bold="true" box="[199,410,1684,1709]" pageId="22" pageNumber="123">Characterization.</emphasis>
Shell thin, cyrtoconoid, glossy, H
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,
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mm; whorls shouldered and keeled; protoconch with six spiral ribs, DN
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; terminal varix very fine at 1.0 whorls; axials and spirals on first two T whorls with microscopic granules in between; later whorls with distinctly beaded subsutural ridge and strong beaded basal spiral rib, shell smooth in between; strong sutural depression; base of body whorl almost smooth; five narrow spiral chords with low spines in umbilicus that is encircled by beads.
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<emphasis id="F3DDEAB7FFAB6D36FF4FF8F35B66F89F" bold="true" box="[199,352,1863,1888]" pageId="22" pageNumber="123">Distribution.</emphasis>
Apart from the studied material, known only from the
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locality, the Kai Islands in
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, from
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depth (
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Genus
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species.
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(by monotypy); Pliocene,
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Family
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Solariellidae
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<emphasis id="F3DDEAB7FFAB6D36FF1FFB315B0DFB61" bold="true" box="[151,267,1157,1182]" pageId="22" pageNumber="123">Remarks.</emphasis>
This family was considered a subfamily of
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Trochidae (
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Bouchet
<emphasis id="F3DDEAB7FFAB6D36FC7FFB325E34FB61" box="[1015,1074,1157,1182]" italics="true" pageId="22" pageNumber="123">et al.</emphasis>
2005
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)
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, but has since then been raised to family level (
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Williams
<emphasis id="F3DDEAB7FFAB6D36FD89FB1E583FFB3D" box="[513,569,1193,1218]" italics="true" pageId="22" pageNumber="123">et al.</emphasis>
2008
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).
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<heading id="9A5E81C9FFAB6D36FF1FFF2C5BFFFF4D" bold="true" box="[151,505,152,178]" fontSize="11" level="1" pageId="22" pageNumber="123" reason="1">
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<taxonomicName id="06A94D26FFAB6D36FF1FFF2C5B7CFF4D" ID-CoL="3BLMR" box="[151,378,152,178]" class="Gastropoda" family="Trochidae" genus="Ethminolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="123" phylum="Mollusca" rank="species" species="wareni">
<emphasis id="F3DDEAB7FFAB6D36FF1FFF2C5B7CFF4D" bold="true" box="[151,378,152,178]" italics="true" pageId="22" pageNumber="123">Ethminolia wareni</emphasis>
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nov. spec.
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(
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)
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<emphasis id="F3DDEAB7FFAB6D36FF1FFEB65B43FEE4" bold="true" box="[151,325,258,283]" pageId="22" pageNumber="123">
<typeStatus id="1E128807FFAB6D36FF1FFEB65AD4FEE4" box="[151,210,258,283]" pageId="22" pageNumber="123">Type</typeStatus>
material.
</emphasis>
<typeStatus id="1E128807FFAB6D36FEC4FEB65BB2FEE4" box="[332,436,258,283]" pageId="22" pageNumber="123" type="holotype">Holotype</typeStatus>
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608.321,
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<collectionCode id="A7B8AE60FFAB6D36FD55FEB7591BFEE4" box="[733,797,259,283]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="123">RGM</collectionCode>
608.322 and
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608.323 (Anda 4).
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<emphasis id="F3DDEAB7FFAB6D36FF4FFE925B63FEC0" bold="true" box="[199,357,294,319]" pageId="22" pageNumber="123">
<typeStatus id="1E128807FFAB6D36FF4FFE925B04FEC0" box="[199,258,294,319]" pageId="22" pageNumber="123">Type</typeStatus>
locality.
</emphasis>
Anda 2, Bolinao,
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, Santa Cruz Formation, Pliocene or Early Pleistocene.
</paragraph>
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<emphasis id="F3DDEAB7FFAB6D36FF4FFEFE5BAEFE9C" bold="true" box="[199,424,330,355]" pageId="22" pageNumber="123">Derivatio nominis.</emphasis>
Named after Anders Warén (Swedish Museum of Natural History), a prominent deepwater malacologist.
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<paragraph id="C11636A5FFAB6D36FF4FFE265EC5FE30" blockId="22.[151,1436,258,1003]" pageId="22" pageNumber="123">
<emphasis id="F3DDEAB7FFAB6D36FF4FFE265B85FE54" bold="true" box="[199,387,402,427]" pageId="22" pageNumber="123">Other material.</emphasis>
Anda 1 (74); Anda 2 (66); Anda 3 (28); Anda 4 (42); Anda 5 (3); Anda 6 (11); AndaDeVos (27); AndaClif 1 (1); AndaClif 3 (4); Roxas (3); Tiep 1 (1); Tiep 3 (18); Tiep 4 (2); Tiep 5 (27).
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<emphasis id="F3DDEAB7FFAB6D36FF4FFE6E5B47FE0C" bold="true" box="[199,321,474,499]" pageId="22" pageNumber="123">Diagnosis.</emphasis>
Shell low conical, H 3.0 mm,
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W
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mm; DN
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; early teleoconch whorls angular; later whorls with subsutural row of knobs; two peripheral ribs; many weak ribs on base; open umbilicus with furrowed umbilical ridge.
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<emphasis id="F3DDEAB7FFAB6D36FF4FFDF25B5FFDA0" bold="true" box="[199,345,582,607]" pageId="22" pageNumber="123">Description.</emphasis>
This shell is conical with a low spire (H/W ratio 0.67). Larger specimens are covered with brownish blotches and a nacreous shine. The protoconch is small, smooth and consists of 1.1 whorls. The P/T boundary is marked by an adnate varix. After the boundary, three spiral cords are present. The upper rib becomes more prominent and shouldered, while the lower ribs gradually disappear after two whorls. On the third whorl, a spiral row of knobs is gradually formed directly below the suture. The rib and the shoulder it delimits gradually disappear on the fourth whorl. Growth lines are microscopic. The body whorl has two spiral ribs on the periphery and many weak spiral ribs on the base. The aperture is broken in all specimens, but it appears to be subquadrate, with a straight columellar lip. The umbilicus is deep and staircase-like, because of the straight angle of the columella and the base. The edge of the umbilicus has many narrow furrows.
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<emphasis id="F3DDEAB7FFAB6D36FF4FFC3E5B79FC5C" bold="true" box="[199,383,906,931]" pageId="22" pageNumber="123">Differentiation.</emphasis>
This species differs from
<taxonomicName id="06A94D26FFAB6D36FD2EFC3859EBFC5C" authority="Okutani, 1961" authorityName="Okutani" authorityYear="1961" box="[678,1005,906,931]" class="Gastropoda" family="Trochidae" genus="Ethminolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="123" phylum="Mollusca" rank="species" species="nektonica">
<emphasis id="F3DDEAB7FFAB6D36FD2EFC385935FC5C" box="[678,819,906,931]" italics="true" pageId="22" pageNumber="123">E. nektonica</emphasis>
(
<bibRefCitation id="A5384B54FFAB6D36FCCAFC3E59E3FC5C" author="Okutani" box="[834,997,906,931]" pageId="22" pageNumber="123" refString="Okutani, T. (1961) Description of Solariella nektonica, sp. nov. with special reference to its swimming behavior. Ve nus, 23, 72 - 90." type="journal article" year="1961">Okutani, 1961</bibRefCitation>
)
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by the lack of spiral ribs on the upper side of the later whorls, while
<taxonomicName id="06A94D26FFAB6D36FE66FC1B5933FC38" authority="Okutani, 1961" authorityName="Okutani" authorityYear="1961" box="[494,821,942,967]" class="Gastropoda" family="Trochidae" genus="Ethminolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="123" phylum="Mollusca" rank="species" species="nektonica">
<emphasis id="F3DDEAB7FFAB6D36FE66FC1B587CFC38" box="[494,634,942,967]" italics="true" pageId="22" pageNumber="123">E. nektonica</emphasis>
(
<bibRefCitation id="A5384B54FFAB6D36FD01FC1A592BFC38" author="Okutani" box="[649,813,942,967]" pageId="22" pageNumber="123" refString="Okutani, T. (1961) Description of Solariella nektonica, sp. nov. with special reference to its swimming behavior. Ve nus, 23, 72 - 90." type="journal article" year="1961">Okutani, 1961</bibRefCitation>
)
</taxonomicName>
has rather strong spiral ribs on the apical side of these whorls. The subsutural row of knobs is more pronounced in
<taxonomicName id="06A94D26FFAB6D36FCB3FC6059A0FC14" box="[827,934,980,1003]" class="Gastropoda" family="Trochidae" genus="Ethminolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="123" phylum="Mollusca" rank="species" species="wareni">
<emphasis id="F3DDEAB7FFAB6D36FCB3FC6059A0FC14" box="[827,934,980,1003]" italics="true" pageId="22" pageNumber="123">E. wareni</emphasis>
</taxonomicName>
.
</paragraph>
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</treatment>
</document>

113
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<heading id="9A5E81C9FFAD6D30FF1FFB445B79FAF5" bold="true" box="[151,383,1264,1290]" fontSize="11" level="1" pageId="16" pageNumber="117" reason="1">
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<emphasis id="F3DDEAB7FFAD6D30FF1FFB445B23FAF5" bold="true" box="[151,293,1264,1290]" italics="true" pageId="16" pageNumber="117">Calliotropis</emphasis>
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spec. 1
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<paragraph id="C11636A5FFAD6D30FF1FFAA65B4CFAD4" blockId="16.[151,383,1264,1323]" box="[151,330,1298,1323]" pageId="16" pageNumber="117">
(
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<emphasis id="F3DDEAB7FFAD6D30FF1FFAEE5B00FA8C" bold="true" box="[151,262,1370,1395]" pageId="16" pageNumber="117">Material.</emphasis>
Anda 1 (1); Anda 2 (2); Anda 4 (2).
</paragraph>
<paragraph id="C11636A5FFAD6D30FF4FFACA583CF9FC" blockId="16.[151,1436,1370,1683]" pageId="16" pageNumber="117">
<emphasis id="F3DDEAB7FFAD6D30FF4FFACA5B9CFA68" bold="true" box="[199,410,1406,1431]" pageId="16" pageNumber="117">Characterization.</emphasis>
Shell with rounded whorls, H
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,
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W
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mm; P large, bulbous, granulate, 1.1 whorls, DN
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; P/T boundary distinct rib; T with reticulate ornamentation; weak spines on intersections of spiral and axial ribs; microsculpture of discontinuous granulate spiral threads; body whorl with four spiral ribs; aperture rounded; umbilicus moderately wide.
</paragraph>
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<subSubSection id="89B3652EFFAD6D30FF4FF9BA5E28F96C" pageId="16" pageNumber="117" type="discussion">
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<emphasis id="F3DDEAB7FFAD6D30FF4FF9BA5B3DF9D8" bold="true" box="[199,315,1550,1575]" pageId="16" pageNumber="117">Remarks.</emphasis>
These shells are probably all juveniles of a single species that could not be matched with any of the other
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<emphasis id="F3DDEAB7FFAD6D30FF5FF9865B5BF9B4" box="[215,349,1586,1611]" italics="true" pageId="16" pageNumber="117">Calliotropis</emphasis>
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species in the studied material. It is most similar to
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<emphasis id="F3DDEAB7FFAD6D30FC2AF9875E2FF9B4" box="[930,1065,1586,1611]" italics="true" pageId="16" pageNumber="117">C. limbifera</emphasis>
(
<bibRefCitation id="A5384B54FFAD6D30FBBFF9865EF1F9B4" author="Schepman" box="[1079,1271,1586,1611]" pageId="16" pageNumber="117" refString="Schepman, M. M. (1908) The Prosobranchia of the Siboga Expedition-Part I Rhipidoglossa and Docoglossa with an appendix by Prof. R. Bergh. Siboga-Expeditie Monographie, 49, 1 - 98." type="journal article" year="1908">Schepman, 1908</bibRefCitation>
)
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, but the spiral ribs are placed lower on the whorls in these specimens than in
<taxonomicName id="06A94D26FFAD6D30FCFBF9E35ED1F990" authority="Schepman, 1908" authorityName="Schepman" authorityYear="1908" box="[883,1239,1622,1647]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="16" pageNumber="117" phylum="Mollusca" rank="species" species="limbifera">
<emphasis id="F3DDEAB7FFAD6D30FCFBF9E359FBF990" box="[883,1021,1622,1647]" italics="true" pageId="16" pageNumber="117">C. limbifera</emphasis>
(
<bibRefCitation id="A5384B54FFAD6D30FB86F9E25EC8F990" author="Schepman" box="[1038,1230,1622,1647]" pageId="16" pageNumber="117" refString="Schepman, M. M. (1908) The Prosobranchia of the Siboga Expedition-Part I Rhipidoglossa and Docoglossa with an appendix by Prof. R. Bergh. Siboga-Expeditie Monographie, 49, 1 - 98." type="journal article" year="1908">Schepman, 1908</bibRefCitation>
)
</taxonomicName>
, the microscopic threads are spirally instead of axially arranged and the protoconch is less inclined.
</paragraph>
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130
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<emphasis id="F3DDEAB7FFAD6D30FF1FFD655B9DFD14" bold="true" box="[151,411,721,747]" italics="true" pageId="16" pageNumber="117">Calliotropis limbifera</emphasis>
(
<bibRefCitation id="A5384B54FFAD6D30FE22FD65587EFD14" author="Schepman" box="[426,632,721,747]" pageId="16" pageNumber="117" refString="Schepman, M. M. (1908) The Prosobranchia of the Siboga Expedition-Part I Rhipidoglossa and Docoglossa with an appendix by Prof. R. Bergh. Siboga-Expeditie Monographie, 49, 1 - 98." type="journal article" year="1908">Schepman, 1908</bibRefCitation>
)
</emphasis>
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(
<figureCitation id="59922A20FFAD6D30FF16FD405B45FCF3" box="[158,323,756,781]" captionStart="FIGURES 33 – 38" captionStartId="17.[151,264,1857,1879]" captionTargetBox="[235,1353,195,1840]" captionTargetId="figure@17.[233,1356,193,1843]" captionTargetPageId="17" captionText="FIGURES 33 – 38. Calliotropidae. 33. Calliotropis limbifera (Schepman, 1908). RGM 608.292. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) apical view, (d) spire detail. 34. Calliotropis limbifera (Schepman, 1908). RGM 608.293. Locality AndaDeVos. SEM detail of protoconch. 35. Calliotropis spec. 1. RGM 608.296. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 36. Calliotropis spec. 1. RGM 608.297. Locality Anda 4. SEM detail of protoconch. 37. Calliotropis spec. 2. RGM 608.301. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 38. Calliotropis spec. 2. RGM 608.302. Locality Anda 3. SEM detail of protoconch." httpUri="https://zenodo.org/record/250967/files/figure.png" pageId="16" pageNumber="117">Figures 33–34</figureCitation>
)
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<paragraph id="C11636A5FFAD6D30FF1FFC88589EFC8F" blockId="16.[151,963,828,911]" pageId="16" pageNumber="117">
1908
<taxonomicName id="06A94D26FFAD6D30FF7DFC885BE5FCAD" box="[245,483,828,850]" class="Gastropoda" family="Eucyclidae" genus="Solariellopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="16" pageNumber="117" phylum="Mollusca" rank="species" species="limbifera">
<emphasis id="F3DDEAB7FFAD6D30FF7DFC885BE5FCAD" box="[245,483,828,850]" italics="true" pageId="16" pageNumber="117">Solariellopsis limbifera</emphasis>
</taxonomicName>
nov. spec.—Schepman: p. 54–55, pl. IV, fig. 3. 1979
<taxonomicName id="06A94D26FFAD6D30FF7DFCEE5BD7FC8F" box="[245,465,858,880]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="16" pageNumber="117" phylum="Mollusca" rank="species" species="limbifera">
<emphasis id="F3DDEAB7FFAD6D30FF7DFCEE5BD7FC8F" box="[245,465,858,880]" italics="true" pageId="16" pageNumber="117">Calliotropis limbifera</emphasis>
</taxonomicName>
—
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: p. 526.
</paragraph>
<paragraph id="C11636A5FFAD6D30FF1FFCCD5911FC70" blockId="16.[151,963,828,911]" box="[151,791,889,911]" pageId="16" pageNumber="117">
2007
<taxonomicName id="06A94D26FFAD6D30FF7DFCCD5BD7FC70" box="[245,465,889,911]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="16" pageNumber="117" phylum="Mollusca" rank="species" species="limbifera">
<emphasis id="F3DDEAB7FFAD6D30FF7DFCCD5BD7FC70" box="[245,465,889,911]" italics="true" pageId="16" pageNumber="117">Calliotropis limbifera</emphasis>
</taxonomicName>
—Vilvens: p. 50, figs 172–175.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFAD6D30FF1FFC0258A1FBA0" pageId="16" pageNumber="117" type="materials_examined">
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<emphasis id="F3DDEAB7FFAD6D30FF1FFC025B00FC30" bold="true" box="[151,262,950,975]" pageId="16" pageNumber="117">Material.</emphasis>
AndaDeVos (
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, damaged;
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, damaged); AndaClif3 (
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, damaged).
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<emphasis id="F3DDEAB7FFAD6D30FF4FFC6E5B9CFC0C" bold="true" box="[199,410,986,1011]" pageId="16" pageNumber="117">Characterization.</emphasis>
Shell broadly conical, keeled, H
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,
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mm; P large, bulbous, weakly granulate, 1.1 whorls, DN
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; P/T boundary weak rib; T microsculpture of discontinuous axial threads; reticulate ornamentation on first 2.5 T whorls; ornamentation changes to rows of knobs, rows of scales with axial lirae in between and a keel ornamented with axial ribs.
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<subSubSection id="89B3652EFFAD6D30FF4FFBDE5ECFFB58" pageId="16" pageNumber="117" type="distribution">
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<emphasis id="F3DDEAB7FFAD6D30FF4FFBDE5B66FB7C" bold="true" box="[199,352,1130,1155]" pageId="16" pageNumber="117">Distribution.</emphasis>
Apart from the studied material the species is known from the Sulu Archipelago,
<collectingCountry id="B9BE7635FFAD6D30FA9FFBDE5F9EFB7C" box="[1303,1432,1130,1155]" name="Philippines" pageId="16" pageNumber="117">Philippines</collectingCountry>
,
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depth (
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) and the south-western Pacific,
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depth (
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).
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120
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<emphasis id="F3DDEAB7FFAD6D30FF1FFF2C5B23FF4D" bold="true" box="[151,293,152,178]" italics="true" pageId="16" pageNumber="117">Calliotropis</emphasis>
cf.
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<paragraph id="C11636A5FFAD6D30FF1FFF0E5B4CFF2C" blockId="16.[151,661,152,211]" box="[151,330,186,211]" pageId="16" pageNumber="117">
(
<figureCitation id="59922A20FFAD6D30FF16FF0E5B44FF2C" box="[158,322,186,211]" captionStart="FIGURES 26 – 32" captionStartId="15.[151,264,1793,1815]" captionTargetBox="[259,1325,197,1767]" captionTargetId="figure@15.[257,1328,195,1768]" captionTargetPageId="15" captionText="FIGURES 26 – 32. Calliotropidae. 26. Calliotropis arenosa nov. spec. RGM 608.272 (holotype). Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 27. Calliotropis arenosa nov. spec. RGM 608.273 (paratype). Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 28. Calliotropis arenosa nov. spec. RGM 608.274 (paratype). Locality Anda 3. (a) rear view, (b) apertural view, (c) apical view. 29. Calliotropis arenosa nov. spec. RGM 608.275 (paratype). Locality AndaDeVos. SEM detail of protoconch. 30. Calliotropis cf. pyramoeides Vilvens, 2007. RGM 608.281. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 31. Calliotropis cf. pyramoeides Vilvens, 2007. RGM 608.282. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 32. Calliotropis cf. pyramoeides Vilvens, 2007. RGM 608.283. Locality Anda 1. (a) SEM, side view, (b) SEM detail of protoconch." httpUri="https://zenodo.org/record/250966/files/figure.png" pageId="16" pageNumber="117">Figures 30–32</figureCitation>
)
</paragraph>
<paragraph id="C11636A5FFAD6D30FF1FFEB65998FEE7" blockId="16.[151,926,258,280]" box="[151,926,258,280]" pageId="16" pageNumber="117">
cf. 2007
<taxonomicName id="06A94D26FFAD6D30FF7DFEB65BF0FEE7" box="[245,502,258,280]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="16" pageNumber="117" phylum="Mollusca" rank="species" species="pyramoeides">
<emphasis id="F3DDEAB7FFAD6D30FF7DFEB65BF0FEE7" box="[245,502,258,280]" italics="true" pageId="16" pageNumber="117">Calliotropis pyramoeides</emphasis>
</taxonomicName>
nov. spec.—Vilvens: p. 36, figs 112–115.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFAD6D30FF1FFEF15978FDED" pageId="16" pageNumber="117" type="materials_examined">
<paragraph id="C11636A5FFAD6D30FF1FFEF15AB9FE7E" blockId="16.[151,1436,325,638]" pageId="16" pageNumber="117">
<emphasis id="F3DDEAB7FFAD6D30FF1FFEF15B00FEA1" bold="true" box="[151,262,325,350]" pageId="16" pageNumber="117">Material.</emphasis>
Anda 1 (20); Anda 2 (19); Anda 3 (11); Anda 5 (4); Anda 6 (5); AndaDeVos (4); AndaClif3 (3); Tiep 1 (1).
</paragraph>
<paragraph id="C11636A5FFAD6D30FF4FFE395978FDED" blockId="16.[151,1436,325,638]" pageId="16" pageNumber="117">
<emphasis id="F3DDEAB7FFAD6D30FF4FFE395B9CFE59" bold="true" box="[199,410,397,422]" pageId="16" pageNumber="117">Characterization.</emphasis>
Shell broadly conical, H
<quantity id="06519B40FFAD6D30FD5CFE3A5937FE5A" box="[724,817,398,422]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="117" unit="mm" value="2.8">2.8 mm</quantity>
,
<geoCoordinate id="A49D5062FFAD6D30FCCBFE3A598BFE59" box="[835,909,398,422]" direction="west" orientation="longitude" pageId="16" pageNumber="117" precision="5555" value="-2.9">
W
<quantity id="06519B40FFAD6D30FCE2FE3A598BFE59" box="[874,909,398,422]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="16" pageNumber="117" unit="mm" value="2.9">2.9</quantity>
</geoCoordinate>
mm; P globular, finely granulate, with four discontinuous spiral ribs and zigzag patterns, 1.1 whorls, DN
<quantity id="06519B40FFAD6D30FCC4FE0559F4FE36" box="[844,1010,433,457]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="1.7999999999999998" metricValueMin="1.7" pageId="16" pageNumber="117" unit="mm" value="0.175" valueMax="0.18" valueMin="0.17">0.17–0.18 mm</quantity>
; P/T boundary thin rib; T ornamented with spiral and axial ribs with spines at intersections; spines lower on whorl develop into C-shaped lamellae; umbilicus broad, encircled by rib, with spiral and axial ribs inside.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFAD6D30FF4FFDA958F4FDA5" pageId="16" pageNumber="117" type="distribution">
<paragraph id="C11636A5FFAD6D30FF4FFDA958F4FDA5" blockId="16.[151,1436,325,638]" pageId="16" pageNumber="117">
<emphasis id="F3DDEAB7FFAD6D30FF4FFDA95B66FDC9" bold="true" box="[199,352,541,566]" pageId="16" pageNumber="117">Distribution.</emphasis>
South-western Pacific (
<collectingCountry id="B9BE7635FFAD6D30FDF8FDAA5927FDC9" box="[624,801,541,566]" name="New Caledonia" pageId="16" pageNumber="117">New Caledonia</collectingCountry>
area), alive at
<quantity id="06519B40FFAD6D30FC43FDAA5E15FDCA" box="[971,1043,542,566]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="117" unit="m" value="200.0">200 m</quantity>
depth, empty shells at
<quantity id="06519B40FFAD6D30FA94FDA95F9AFDCA" box="[1308,1436,541,566]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="16" pageNumber="117" unit="m" value="300.0" valueMax="350.0" valueMin="250.0">250–350 m</quantity>
depth (
<bibRefCitation id="A5384B54FFAD6D30FF6BFDF55B78FDA6" author="Vilvens" box="[227,382,577,602]" pageId="16" pageNumber="117" refString="Vilvens, C. (2007) New species and new records of Calliotropis (Gastropoda: Chilodontidae: Calliotropinae) from Indo- Pacific. Novapex: Trimestriel de la Societe Belge de Malacologie, 8 (Hors Serie 5), 1 - 72." type="journal article" year="2007">Vilvens 2007</bibRefCitation>
), apart from the studied material.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFAD6D30FF4FFDD15EB0FD81" box="[199,1206,613,638]" pageId="16" pageNumber="117" type="discussion">
<paragraph id="C11636A5FFAD6D30FF4FFDD15EB0FD81" blockId="16.[151,1436,325,638]" box="[199,1206,613,638]" pageId="16" pageNumber="117">
<emphasis id="F3DDEAB7FFAD6D30FF4FFDD15B3DFD81" bold="true" box="[199,315,613,638]" pageId="16" pageNumber="117">Remarks.</emphasis>
The base of our shells appears to be taller than those figured in
<bibRefCitation id="A5384B54FFAD6D30FB8FFDD15EB4FD81" author="Vilvens" box="[1031,1202,613,638]" pageId="16" pageNumber="117" refString="Vilvens, C. (2007) New species and new records of Calliotropis (Gastropoda: Chilodontidae: Calliotropinae) from Indo- Pacific. Novapex: Trimestriel de la Societe Belge de Malacologie, 8 (Hors Serie 5), 1 - 72." type="journal article" year="2007">Vilvens (2007)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

149
data/49/00/87/490087B3FFAD6D32FF1FF9685F87FF2A.xml Normal file
View file

@ -0,0 +1,149 @@
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<treatment id="490087B3FFAD6D32FF1FF9685F87FF2A" ID-DOI="http://doi.org/10.5281/zenodo.5671289" ID-GBIF-Taxon="119412419" ID-Zenodo-Dep="5671289" LSID="urn:lsid:plazi:treatment:490087B3FFAD6D32FF1FF9685F87FF2A" httpUri="http://treatment.plazi.org/id/490087B3FFAD6D32FF1FF9685F87FF2A" lastPageId="18" lastPageNumber="119" pageId="16" pageNumber="117">
<subSubSection id="89B3652EFFAD6D30FF1FF9685B4CF8E8" pageId="16" pageNumber="117" type="nomenclature">
<paragraph id="C11636A5FFAD6D30FF1FF9685B79F909" blockId="16.[151,383,1756,1815]" box="[151,383,1756,1782]" pageId="16" pageNumber="117">
<heading id="9A5E81C9FFAD6D30FF1FF9685B79F909" bold="true" box="[151,383,1756,1782]" fontSize="11" level="1" pageId="16" pageNumber="117" reason="1">
<emphasis id="F3DDEAB7FFAD6D30FF1FF9685B79F909" bold="true" box="[151,383,1756,1782]" pageId="16" pageNumber="117">
<taxonomicName id="06A94D26FFAD6D30FF1FF9685B23F909" ID-CoL="5c283633-8a11-41f9-a0b0-ee438c32f58a" box="[151,293,1756,1782]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="16" pageNumber="117" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFAD6D30FF1FF9685B23F909" bold="true" box="[151,293,1756,1782]" italics="true" pageId="16" pageNumber="117">Calliotropis</emphasis>
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spec. 2
</emphasis>
</heading>
</paragraph>
<paragraph id="C11636A5FFAD6D30FF1FF94A5B4CF8E8" blockId="16.[151,383,1756,1815]" box="[151,330,1790,1815]" pageId="16" pageNumber="117">
(
<figureCitation id="59922A20FFAD6D30FF16F94A5B45F8E8" box="[158,323,1790,1815]" captionStart="FIGURES 33 – 38" captionStartId="17.[151,264,1857,1879]" captionTargetBox="[235,1353,195,1840]" captionTargetId="figure@17.[233,1356,193,1843]" captionTargetPageId="17" captionText="FIGURES 33 – 38. Calliotropidae. 33. Calliotropis limbifera (Schepman, 1908). RGM 608.292. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) apical view, (d) spire detail. 34. Calliotropis limbifera (Schepman, 1908). RGM 608.293. Locality AndaDeVos. SEM detail of protoconch. 35. Calliotropis spec. 1. RGM 608.296. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 36. Calliotropis spec. 1. RGM 608.297. Locality Anda 4. SEM detail of protoconch. 37. Calliotropis spec. 2. RGM 608.301. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 38. Calliotropis spec. 2. RGM 608.302. Locality Anda 3. SEM detail of protoconch." httpUri="https://zenodo.org/record/250967/files/figure.png" pageId="16" pageNumber="117">Figures 37–38</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFAD6D30FF1FF8F25E18F810" pageId="16" pageNumber="117" type="materials_examined">
<paragraph id="C11636A5FFAD6D30FF1FF8F25898F8A0" blockId="16.[151,1436,1862,2031]" box="[151,670,1862,1887]" pageId="16" pageNumber="117">
<emphasis id="F3DDEAB7FFAD6D30FF1FF8F25B00F8A0" bold="true" box="[151,262,1862,1887]" pageId="16" pageNumber="117">Material.</emphasis>
Anda 1 (1); Anda 2 (1); Anda 3 (1).
</paragraph>
<paragraph id="C11636A5FFAD6D30FF4FF8DE5E18F810" blockId="16.[151,1436,1862,2031]" pageId="16" pageNumber="117">
<emphasis id="F3DDEAB7FFAD6D30FF4FF8DE5B9CF87C" bold="true" box="[199,410,1898,1923]" pageId="16" pageNumber="117">Characterization.</emphasis>
Shell low trochiform, H 2.0 mm,
<geoCoordinate id="A49D5062FFAD6D30FC94F8DF5966F87C" box="[796,864,1899,1923]" direction="west" orientation="longitude" pageId="16" pageNumber="117" precision="5555" value="-2.8">
W
<quantity id="06519B40FFAD6D30FCB5F8DF5966F87C" box="[829,864,1899,1923]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="117" unit="mm" value="2.8">2.8</quantity>
</geoCoordinate>
mm; P globular, slightly elevated, 1.2 whorls, DN 0.18; P/T boundary weak rib; sculpture of axial ribs and two weak spiral ribs, with spines at intersections which become stronger and hollow; upper spiral rib becomes acute; many small axial ribs below lower spiral rib; umbilicus wide, with distinct umbilical ridge, axial ribs with small spines inside.
</paragraph>
</subSubSection>
<caption id="95D6662DFFAC6D31FF1FF8F559BCF80F" httpUri="https://zenodo.org/record/250967/files/figure.png" pageId="17" pageNumber="118" targetBox="[235,1353,195,1840]" targetPageId="17">
<paragraph id="C11636A5FFAC6D31FF1FF8F559BCF80F" blockId="17.[151,1436,1857,2032]" pageId="17" pageNumber="118">
<emphasis id="F3DDEAB7FFAC6D31FF1FF8F55B53F8A8" bold="true" box="[151,341,1857,1879]" pageId="17" pageNumber="118">FIGURES 33–38.</emphasis>
<taxonomicName id="06A94D26FFAC6D31FED4F8F55BF6F8A8" box="[348,496,1857,1879]" class="Gastropoda" family="Calliotropidae" kingdom="Animalia" pageId="17" pageNumber="118" phylum="Mollusca" rank="family">Calliotropidae</taxonomicName>
.
<emphasis id="F3DDEAB7FFAC6D31FE72F8F5581FF8A8" bold="true" box="[506,537,1857,1879]" pageId="17" pageNumber="118">33.</emphasis>
<taxonomicName id="06A94D26FFAC6D31FDA9F8F559C5F8A8" authority="Schepman, 1908" authorityName="Schepman" authorityYear="1908" box="[545,963,1857,1879]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="17" pageNumber="118" phylum="Mollusca" rank="species" species="limbifera">
<emphasis id="F3DDEAB7FFAC6D31FDA9F8F558F8F8A8" box="[545,766,1857,1879]" italics="true" pageId="17" pageNumber="118">Calliotropis limbifera</emphasis>
(Schepman, 1908)
</taxonomicName>
. RGM 608.292. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) apical view, (d) spire detail.
<emphasis id="F3DDEAB7FFAC6D31FD50F8EB58F1F88A" bold="true" box="[728,759,1887,1909]" pageId="17" pageNumber="118">34.</emphasis>
<taxonomicName id="06A94D26FFAC6D31FD76F8EB5E99F88A" authority="Schepman, 1908" authorityName="Schepman" authorityYear="1908" box="[766,1183,1887,1909]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="17" pageNumber="118" phylum="Mollusca" rank="species" species="limbifera">
<emphasis id="F3DDEAB7FFAC6D31FD76F8EB59DDF88A" box="[766,987,1887,1909]" italics="true" pageId="17" pageNumber="118">Calliotropis limbifera</emphasis>
(Schepman, 1908)
</taxonomicName>
. RGM 608.293. Locality AndaDeVos. SEM detail of protoconch.
<emphasis id="F3DDEAB7FFAC6D31FDB4F8CA585AF86B" bold="true" box="[572,604,1918,1940]" pageId="17" pageNumber="118">35.</emphasis>
<taxonomicName id="06A94D26FFAC6D31FDECF8CA58D8F86B" box="[612,734,1918,1940]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="17" pageNumber="118" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFAC6D31FDECF8CA58D8F86B" box="[612,734,1918,1940]" italics="true" pageId="17" pageNumber="118">Calliotropis</emphasis>
</taxonomicName>
spec.1. RGM 608.296. Locality Anda2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view.
<emphasis id="F3DDEAB7FFAC6D31FD93F829583CF84C" bold="true" box="[539,570,1949,1971]" pageId="17" pageNumber="118">36.</emphasis>
<taxonomicName id="06A94D26FFAC6D31FDCAF82958BAF84C" box="[578,700,1949,1971]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="17" pageNumber="118" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFAC6D31FDCAF82958BAF84C" box="[578,700,1949,1971]" italics="true" pageId="17" pageNumber="118">Calliotropis</emphasis>
</taxonomicName>
spec.1. RGM 608.297. Locality Anda4. SEM detail of protoconch.
<emphasis id="F3DDEAB7FFAC6D31FAF4F8295F9AF84C" bold="true" box="[1404,1436,1949,1971]" pageId="17" pageNumber="118">37.</emphasis>
<taxonomicName id="06A94D26FFAC6D31FF1FF8085B17F82D" box="[151,273,1980,2002]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="17" pageNumber="118" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFAC6D31FF1FF8085B17F82D" box="[151,273,1980,2002]" italics="true" pageId="17" pageNumber="118">Calliotropis</emphasis>
</taxonomicName>
spec.2. RGM 608.301. Locality Anda2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view.
<emphasis id="F3DDEAB7FFAC6D31FAF4F8085F9AF82D" bold="true" box="[1404,1436,1980,2002]" pageId="17" pageNumber="118">38.</emphasis>
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spec.2. RGM 608.302. Locality Anda3. SEM detail of protoconch.
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<emphasis id="F3DDEAB7FFAF6D32FF4FFF235B3DFF4F" bold="true" box="[199,315,151,176]" pageId="18" pageNumber="119">Remarks.</emphasis>
The studied specimens differ from
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<emphasis id="F3DDEAB7FFAF6D32FD5FFF2C5940FF4F" box="[727,838,151,176]" italics="true" pageId="18" pageNumber="119">C. bucina</emphasis>
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(
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and
<collectingCountry id="B9BE7635FFAF6D32FB60FF2C5F43FF4F" box="[1256,1349,152,176]" name="Mayotte" pageId="18" pageNumber="119">Mayotte</collectingCountry>
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) and
<taxonomicName id="06A94D26FFAF6D32FF4EFF095948FF2A" authority="Poppe, Tagaro &amp; Dekker, 2006" authorityName="Poppe, Tagaro &amp; Dekker" authorityYear="2006" box="[198,846,188,213]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="18" pageNumber="119" phylum="Mollusca" rank="species" species="sagarinoi">
<emphasis id="F3DDEAB7FFAF6D32FF4EFF095B54FF2B" box="[198,338,189,212]" italics="true" pageId="18" pageNumber="119">C. sagarinoi</emphasis>
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(
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)
</taxonomicName>
by having a more angular keel and higher whorls.
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79
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(by monotypy); Recent, Indo-Pacific.
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(
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<emphasis id="F3DDEAB7FFAF6D32FF1FFBCB5B00FB67" bold="true" box="[151,262,1151,1176]" pageId="18" pageNumber="119">Material.</emphasis>
Anda 2 (
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and 2 partially preserved subadults); Anda 3 (
<specimenCount id="D7AFFD2CFFAF6D32FB97FB345EA5FB67" box="[1055,1187,1151,1176]" pageId="18" pageNumber="119" type="juvenile">2 juveniles</specimenCount>
); Anda 4 (
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partially preserved); Anda 6 (
<specimenCount id="D7AFFD2CFFAF6D32FE69FB105867FB42" box="[481,609,1188,1213]" pageId="18" pageNumber="119" type="juvenile">2 juveniles</specimenCount>
); AndaDeVos (
<specimenCount id="D7AFFD2CFFAF6D32FC82FB10598CFB42" box="[778,906,1188,1213]" pageId="18" pageNumber="119" type="juvenile">2 juveniles</specimenCount>
); AndaClif3 (
<specimenCount id="D7AFFD2CFFAF6D32FBAEFB105E9FFB42" box="[1062,1177,1188,1213]" pageId="18" pageNumber="119" type="juvenile">1 juvenile</specimenCount>
).
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<emphasis id="F3DDEAB7FFAF6D32FF4FFB735B9CFB1F" bold="true" box="[199,410,1223,1248]" pageId="18" pageNumber="119">Characterization.</emphasis>
Shell conical, H
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,
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W
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mm; DN
<quantity id="06519B40FFAF6D32FCFFFB7C5E1BFB20" box="[887,1053,1224,1248]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.7999999999999998" pageId="18" pageNumber="119" unit="mm" value="0.19" valueMax="0.2" valueMin="0.18">0.18–0.20 mm</quantity>
; distinct terminal varix marks P/T boundary at 1.1 whorls; spiral and axial sculpture with spines at intersections; lower primary rib develops into peripheral row of spines; sutural ramp progressively concave; base of studied material damaged; umbilicus probably open, encircled by two rows of granules.
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<emphasis id="F3DDEAB7FFAF6D32FF1FFC895B00FCA9" bold="true" box="[151,262,829,854]" pageId="18" pageNumber="119">Material.</emphasis>
Anda 4 (1).
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<emphasis id="F3DDEAB7FFAF6D32FF4FFCD55B9CFC85" bold="true" box="[199,410,865,890]" pageId="18" pageNumber="119">Characterization.</emphasis>
Shell conical, H
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mm; DN 0.21; P/T boundary at 1.0 whorls characterized by sudden onset of teleoconch ornamentation of axial and radial ribs with knobs at intersections; base of body whorl with spiral ribs; umbilicus apparently closed.
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Genus
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<emphasis id="F3DDEAB7FFAF6D32FF62FD975B73FDC3" bold="true" box="[234,373,547,572]" italics="true" pageId="18" pageNumber="119">Calliostoma</emphasis>
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species.
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<taxonomicName id="06A94D26FFAF6D32FEA1FDD8588BFD7D" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[297,653,620,642]" class="Gastropoda" family="Trochidae" genus="Trochus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="18" pageNumber="119" phylum="Mollusca" rank="species" species="zizyphinus">
<emphasis id="F3DDEAB7FFAF6D32FEA1FDD85BEFFD7D" box="[297,489,620,642]" italics="true" pageId="18" pageNumber="119">Trochus zizyphinus</emphasis>
<bibRefCitation id="A5384B54FFAF6D32FE78FDD8588BFD7D" author="Linnaeus" box="[496,653,620,642]" pageId="18" pageNumber="119" refString="Linnaeus, C. (1758) Tomus I. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Laurentii Salvii, Holmiae, 1 - 824. http: // dx. doi. org / 10.5962 / bhl. title. 542" type="book chapter" year="1758">Linnaeus, 1758</bibRefCitation>
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(by monotypy); Recent, Europe.
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<emphasis id="F3DDEAB7FFAF6D32FF1FFE035B0DFE2F" bold="true" box="[151,267,439,464]" pageId="18" pageNumber="119">Remarks.</emphasis>
Most calliostomatid species have a characteristic protoconch showing a honeycomb pattern.
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<heading id="9A5E81C9FFAF6D32FF1FF96A58E8F908" bold="true" box="[151,750,1758,1783]" fontSize="10" level="2" pageId="18" pageNumber="119" reason="2">
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<taxonomicName id="06A94D26FFAF6D32FF1FF96A5861F908" ID-CoL="4PCP4" authority="Chen, 2006" authorityName="Chen" authorityYear="2006" box="[151,615,1758,1783]" class="Gastropoda" family="Trochidae" genus="Pseudotalopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="18" pageNumber="119" phylum="Mollusca" rank="species" species="taiwanensis">
<emphasis id="F3DDEAB7FFAF6D32FF1FF96A5BCEF908" bold="true" box="[151,456,1758,1783]" italics="true" pageId="18" pageNumber="119">Pseudotalopia taiwanensis</emphasis>
(
<bibRefCitation id="A5384B54FFAF6D32FE5FF96A5866F908" author="Chen" box="[471,608,1758,1783]" pageId="18" pageNumber="119" refString="Chen, W. D. (2006) A new species of Gibbula (Gastropoda: Trochidae) from Taiwan. Bulletin of Malacology Taiwan, 30, 21 - 25." type="journal article" year="2006">Chen, 2006</bibRefCitation>
)
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nov. comb.
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(
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)
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<paragraph id="C11636A5FFAF6D32FF1FF8FE58DEF880" blockId="18.[151,775,1866,1919]" pageId="18" pageNumber="119">
2006
<taxonomicName id="06A94D26FFAF6D32FF7DF8FE5BC0F89F" box="[245,454,1866,1888]" class="Gastropoda" family="Trochidae" genus="Gibbula" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="18" pageNumber="119" phylum="Mollusca" rank="species" species="taiwanensis">
<emphasis id="F3DDEAB7FFAF6D32FF7DF8FE5BC0F89F" box="[245,454,1866,1888]" italics="true" pageId="18" pageNumber="119">Gibbula taiwanensis</emphasis>
</taxonomicName>
nov. spec.—Chen: p. 21, fig. 1. 2007
<taxonomicName id="06A94D26FFAF6D32FF7DF8DD5BC0F880" box="[245,454,1897,1919]" class="Gastropoda" family="Trochidae" genus="Gibbula" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="18" pageNumber="119" phylum="Mollusca" rank="species" species="taiwanensis">
<emphasis id="F3DDEAB7FFAF6D32FF7DF8DD5BC0F880" box="[245,454,1897,1919]" italics="true" pageId="18" pageNumber="119">Gibbula taiwanensis</emphasis>
</taxonomicName>
—Chen &amp; Fu: p. 67, fig. 2.
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<emphasis id="F3DDEAB7FFAF6D32FF1FF8185B00F83A" bold="true" box="[151,262,1964,1989]" pageId="18" pageNumber="119">Material.</emphasis>
Anda 3 (
<specimenCount id="D7AFFD2CFFAF6D32FEF9F8185BC7F83A" box="[369,449,1964,1989]" pageId="18" pageNumber="119" type="adult">1 adult</specimenCount>
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<specimenCount id="D7AFFD2CFFAF6D32FC00F81859FDF83A" box="[904,1019,1964,1989]" pageId="18" pageNumber="119" type="juvenile">1 juvenile</specimenCount>
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</paragraph>
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<paragraph id="C11636A5FFAE6D33FF1FF8F559BBF80F" blockId="19.[151,1436,1857,2032]" pageId="19" pageNumber="120">
<emphasis id="F3DDEAB7FFAE6D33FF1FF8F55B52F8A8" bold="true" box="[151,340,1857,1879]" pageId="19" pageNumber="120">FIGURES 39–44.</emphasis>
<taxonomicName id="06A94D26FFAE6D33FED2F8F5580FF8A8" box="[346,521,1857,1879]" class="Gastropoda" family="Calliostomatidae" kingdom="Animalia" pageId="19" pageNumber="120" phylum="Mollusca" rank="family">Calliostomatidae</taxonomicName>
.
<emphasis id="F3DDEAB7FFAE6D33FD9BF8F55835F8A8" bold="true" box="[531,563,1857,1879]" pageId="19" pageNumber="120">39.</emphasis>
<taxonomicName id="06A94D26FFAE6D33FDB1F8F558B2F8A8" box="[569,692,1857,1879]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="19" pageNumber="120" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFAE6D33FDB1F8F558B2F8A8" box="[569,692,1857,1879]" italics="true" pageId="19" pageNumber="120">Calliostoma</emphasis>
</taxonomicName>
spec.1. RGM 608.304. Locality Anda4. (a) rear view, (b) apertural view, (c) basal view, (d) apical view, (e) detail protoconch.
<emphasis id="F3DDEAB7FFAE6D33FD3BF8EB58D5F88A" bold="true" box="[691,723,1887,1909]" pageId="19" pageNumber="120">40.</emphasis>
<taxonomicName id="06A94D26FFAE6D33FD51F8EB5953F88A" box="[729,853,1887,1909]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="19" pageNumber="120" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFAE6D33FD51F8EB5953F88A" box="[729,853,1887,1909]" italics="true" pageId="19" pageNumber="120">Calliostoma</emphasis>
</taxonomicName>
spec.2. RGM 608.305. Locality Anda3. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view.
<emphasis id="F3DDEAB7FFAE6D33FC82F8CA592FF86B" bold="true" box="[778,809,1918,1940]" pageId="19" pageNumber="120">41.</emphasis>
<taxonomicName id="06A94D26FFAE6D33FCB9F8CA59ABF86B" box="[817,941,1918,1940]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="19" pageNumber="120" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFAE6D33FCB9F8CA59ABF86B" box="[817,941,1918,1940]" italics="true" pageId="19" pageNumber="120">Calliostoma</emphasis>
</taxonomicName>
spec.2. RGM 608.306. Locality Anda6. (a) rear view, (b) apertural view, (c) basal view.
<emphasis id="F3DDEAB7FFAE6D33FDA3F829584CF84C" bold="true" box="[555,586,1949,1971]" pageId="19" pageNumber="120">42.</emphasis>
<taxonomicName id="06A94D26FFAE6D33FDD8F82958CAF84C" box="[592,716,1949,1971]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="19" pageNumber="120" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFAE6D33FDD8F82958CAF84C" box="[592,716,1949,1971]" italics="true" pageId="19" pageNumber="120">Calliostoma</emphasis>
</taxonomicName>
spec.2. RGM 608.307. Locality Anda2. (a) basal view, (b) apical view, (c) rear view.
<emphasis id="F3DDEAB7FFAE6D33FEA2F8085B4CF82D" bold="true" box="[298,330,1980,2002]" pageId="19" pageNumber="120">43.</emphasis>
<taxonomicName id="06A94D26FFAE6D33FEDBF8085BC8F82D" box="[339,462,1980,2002]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="19" pageNumber="120" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFAE6D33FEDBF8085BC8F82D" box="[339,462,1980,2002]" italics="true" pageId="19" pageNumber="120">Calliostoma</emphasis>
</taxonomicName>
spec.2. RGM 608.308. Locality Anda2. (a) rear view, (b) apertural view, (c) apical view.
<emphasis id="F3DDEAB7FFAE6D33FAF4F8085F9AF82D" bold="true" box="[1404,1436,1980,2002]" pageId="19" pageNumber="120">44.</emphasis>
<taxonomicName id="06A94D26FFAE6D33FF1FF86E5B15F80F" box="[151,275,2010,2032]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="19" pageNumber="120" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFAE6D33FF1FF86E5B15F80F" box="[151,275,2010,2032]" italics="true" pageId="19" pageNumber="120">Calliostoma</emphasis>
</taxonomicName>
spec.2. RGM 608.309. Locality Anda3. SEM detail of protoconch.
</paragraph>
</caption>
<paragraph id="C11636A5FFA96D34FF4FFF235823FEE2" blockId="20.[151,1436,151,536]" pageId="20" pageNumber="121">
<emphasis id="F3DDEAB7FFA96D34FF4FFF235B9CFF4F" bold="true" box="[199,410,151,176]" pageId="20" pageNumber="121">Characterization.</emphasis>
Domed, thick trochiform shell with narrow spiral ribs, a well-developed basal keel and color blotches on a pinkish background color, H
<quantity id="06519B40FFA96D34FD31FF085917FF2B" box="[697,785,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="20" pageNumber="121" unit="mm" value="5.9">5.9 mm</quantity>
,
<geoCoordinate id="A49D5062FFA96D34FC94FF095967FF2B" box="[796,865,188,212]" direction="west" orientation="longitude" pageId="20" pageNumber="121" precision="5555" value="-7.3">
W
<quantity id="06519B40FFA96D34FCB6FF085967FF2B" box="[830,865,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="20" pageNumber="121" unit="mm" value="7.3">7.3</quantity>
</geoCoordinate>
mm; protoconch glossy white, DN
<quantity id="06519B40FFA96D34FB7AFF085F9EFF2B" box="[1266,1432,188,212]" metricMagnitude="-5" metricUnit="m" metricValue="8.499999999999998" metricValueMax="9.0" metricValueMin="8.0" pageId="20" pageNumber="121" unit="mm" value="0.08499999999999999" valueMax="0.09" valueMin="0.08">0.08–0.09 mm</quantity>
; P/T boundary poorly defined at 1.2 whorls; inner lip white, glossy, columellar lip twisted; deep, narrow umbilicus with spiral cord and weak lamellae.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFA96D34FF4FFE935B8BFE9A" pageId="20" pageNumber="121" type="distribution">
<paragraph id="C11636A5FFA96D34FF4FFE935B8BFE9A" blockId="20.[151,1436,151,536]" pageId="20" pageNumber="121">
<emphasis id="F3DDEAB7FFA96D34FF4FFE935B66FEBF" bold="true" box="[199,352,295,320]" pageId="20" pageNumber="121">Distribution.</emphasis>
Hitherto the species has only been reported from waters off
<collectingCountry id="B9BE7635FFA96D34FB8EFE935E5DFEBF" box="[1030,1115,295,320]" name="Taiwan" pageId="20" pageNumber="121">Taiwan</collectingCountry>
, on sandy bottoms,
<quantity id="06519B40FFA96D34FAB2FE935AABFE9B" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="20" pageNumber="121" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">150–200 m</quantity>
depth (
<bibRefCitation id="A5384B54FFA96D34FE88FEF85B87FE9A" author="Chen" box="[256,385,332,357]" pageId="20" pageNumber="121" refString="Chen, W. D. (2006) A new species of Gibbula (Gastropoda: Trochidae) from Taiwan. Bulletin of Malacology Taiwan, 30, 21 - 25." type="journal article" year="2006">Chen 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFA96D34FF4FFEDB5F3FFDE7" pageId="20" pageNumber="121" type="discussion">
<paragraph id="C11636A5FFA96D34FF4FFEDB5F3FFDE7" blockId="20.[151,1436,151,536]" pageId="20" pageNumber="121">
<emphasis id="F3DDEAB7FFA96D34FF4FFEDB5B3DFE77" bold="true" box="[199,315,367,392]" pageId="20" pageNumber="121">Remarks.</emphasis>
<bibRefCitation id="A5384B54FFA96D34FECCFEDB5BD5FE77" author="Chen" box="[324,467,367,392]" pageId="20" pageNumber="121" refString="Chen, W. D. (2006) A new species of Gibbula (Gastropoda: Trochidae) from Taiwan. Bulletin of Malacology Taiwan, 30, 21 - 25." type="journal article" year="2006">Chen (2006)</bibRefCitation>
found a specimen of this species inside the stomach of a starfish, so apparently this species is predated on by starfishes. The deep-water Indo-West Pacific species currently assigned to
<taxonomicName id="06A94D26FFA96D34FA9DFE205F77FE52" box="[1301,1393,404,429]" class="Gastropoda" family="Trochidae" genus="Gibbula" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA96D34FA9DFE205F77FE52" box="[1301,1393,404,429]" italics="true" pageId="20" pageNumber="121">Gibbula</emphasis>
</taxonomicName>
are likely unrelated to that European genus of intertidal species (STW unpub. genetic data). We tentatively assign ‘
<emphasis id="F3DDEAB7FFA96D34FF28FE685B94FE0B" box="[160,402,476,501]" italics="true" pageId="20" pageNumber="121">
Gibbula’
<taxonomicName id="06A94D26FFA96D34FE86FE695B94FE0B" box="[270,402,477,500]" class="Gastropoda" family="Trochidae" genus="Gibbula" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="species" species="taiwanensis">taiwanensis</taxonomicName>
</emphasis>
to the deep-sea genus
<taxonomicName id="06A94D26FFA96D34FD10FE68593CFE0A" box="[664,826,476,501]" class="Gastropoda" family="Trochidae" genus="Pseudotalopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA96D34FD10FE68593CFE0A" box="[664,826,476,501]" italics="true" pageId="20" pageNumber="121">Pseudotalopia</emphasis>
</taxonomicName>
based on preliminary genetic data (STW). Molecular data show that
<taxonomicName id="06A94D26FFA96D34FEB7FE4B5BE7FDE7" box="[319,481,511,536]" class="Gastropoda" family="Trochidae" genus="Pseudotalopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="20" pageNumber="121" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFA96D34FEB7FE4B5BE7FDE7" box="[319,481,511,536]" italics="true" pageId="20" pageNumber="121">Pseudotalopia</emphasis>
</taxonomicName>
should be assigned to the trochid subfamily
<taxonomicName id="06A94D26FFA96D34FC5EFE4B5F33FDE7" authority="(Williams 2012)" authorityName="Gray" authorityYear="1857" baseAuthorityName="Williams" baseAuthorityYear="2012" box="[982,1333,511,536]" class="Gastropoda" family="Trochidae" kingdom="Animalia" pageId="20" pageNumber="121" phylum="Mollusca" rank="subFamily" subFamily="Cantharidinae">
Cantharidinae (
<bibRefCitation id="A5384B54FFA96D34FB0AFE4B5F2BFDE7" author="Williams" box="[1154,1325,511,536]" pageId="20" pageNumber="121" refString="Williams, S. T. (2012) Advances in molecular systematics of the vetigastropod superfamily Trochoidea. Zoologica Scripta, 41, 571 - 595. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.2012.00552. x" type="journal article" year="2012">Williams 2012</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

107
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<subSubSection id="89B3652EFFB06D2DFF1FFAC85B4CFA48" pageId="13" pageNumber="114" type="nomenclature">
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<heading id="9A5E81C9FFB06D2DFF1FFAC85B84FA69" bold="true" box="[151,386,1404,1430]" fontSize="11" level="1" pageId="13" pageNumber="114" reason="1">
<emphasis id="F3DDEAB7FFB06D2DFF1FFAC85B84FA69" bold="true" box="[151,386,1404,1430]" pageId="13" pageNumber="114">
<emphasis id="F3DDEAB7FFB06D2DFF1FFAC85B21FA69" bold="true" box="[151,295,1404,1430]" italics="true" pageId="13" pageNumber="114">
<taxonomicName id="06A94D26FFB06D2DFF1FFAC85B1AFA69" box="[151,284,1404,1430]" class="Gastropoda" family="Seguenziidae" genus="Thelyssina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="genus">Thelyssina</taxonomicName>
?
</emphasis>
spec. 2
</emphasis>
</heading>
</paragraph>
<paragraph id="C11636A5FFB06D2DFF1FFA2A5B4CFA48" blockId="13.[151,386,1404,1463]" box="[151,330,1438,1463]" pageId="13" pageNumber="114">
(
<figureCitation id="59922A20FFB06D2DFF16FA2A5B44FA48" box="[158,322,1438,1463]" captionStart="FIGURES 18 – 25" captionStartId="12.[151,264,1787,1809]" captionTargetBox="[262,1331,195,1768]" captionTargetId="figure@12.[260,1334,193,1771]" captionTargetPageId="12" captionText="FIGURES 18 – 25. Seguenziidae. 18. Calliobasis lapulapui Poppe, Tagaro &amp; Dekker, 2006. RGM 608.251. Locality Anda 1. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 19. Calliobasis lapulapui Poppe, Tagaro &amp; Dekker, 2006. RGM 608.252. Locality Anda 3. (a) SEM, side view, (b) SEM detail of protoconch. 20. Thelyssina? spec. 1. RGM 608.263. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 21. Thelyssina? spec. 1. RGM 608.264. Locality Anda 2. (a) SEM detail of protoconch, (b) SEM, side view. 22. Thelyssina? spec. 1. RGM 608.265. Locality Anda 2. SEM, basal view. 23. Thelyssina? spec. 2. RGM 608.268. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 24. Thelyssina? spec. 2. RGM 608.269. Locality AndaDeVos. SEM detail of protoconch. 25. Thelyssina? spec. 2. RGM 608.270 Locality AndaDeVos. SEM, basal view." httpUri="https://zenodo.org/record/250965/files/figure.png" pageId="13" pageNumber="114">Figures 23–25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB06D2DFF1FFA52591BF970" pageId="13" pageNumber="114" type="materials_examined">
<paragraph id="C11636A5FFB06D2DFF1FFA525BD6FA00" blockId="13.[151,1436,1510,1859]" box="[151,464,1510,1535]" pageId="13" pageNumber="114">
<emphasis id="F3DDEAB7FFB06D2DFF1FFA525B00FA00" bold="true" box="[151,262,1510,1535]" pageId="13" pageNumber="114">Material.</emphasis>
AndaDeVos (10).
</paragraph>
<paragraph id="C11636A5FFB06D2DFF4FF9BE591BF970" blockId="13.[151,1436,1510,1859]" pageId="13" pageNumber="114">
<emphasis id="F3DDEAB7FFB06D2DFF4FF9BE5B9CF9DC" bold="true" box="[199,410,1546,1571]" pageId="13" pageNumber="114">Characterization.</emphasis>
Shell minute, H
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,
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W
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mm, trochiform, turreted; DN
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; P/T boundary thin varix at 1.1 whorls; second varix at 1.6 whorls; T whorls shouldered, with axial lirae and minute pits; body whorl with rounded shoulder and low spiral rib; aperture subquadrate; very thin nacre observed at the damaged outer lip of one specimen; umbilicus narrow, with spiral rib inside.
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<emphasis id="F3DDEAB7FFB06D2DFF4FF92E5B3DF94C" bold="true" box="[199,315,1690,1715]" pageId="13" pageNumber="114">Remarks.</emphasis>
This seguenzioid skeneimorph species is also provisionally placed in
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<emphasis id="F3DDEAB7FFB06D2DFBF4F92E5EF2F94C" box="[1148,1268,1690,1715]" italics="true" pageId="13" pageNumber="114">Thelyssina</emphasis>
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, based on the presence of minute pits, a well defined angular shoulder, a basal spiral rib, an umbilical spiral rib and a small protoconch with anastomosing sculpture and a shape similar to the protoconch of the
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species for the genus. However, this assignment is open to further scrutiny as
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<taxonomicName id="06A94D26FFB06D2DFC95F8B2599EF8E0" box="[797,920,1798,1823]" class="Gastropoda" family="Seguenziidae" genus="Thelyssina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="genus">Thelyssina</taxonomicName>
?
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spec. 2 is relatively tall and lacks dendritic sculpture.
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?
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spec. 1
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(
<figureCitation id="59922A20FFB06D2DFF16FC885B44FCAB" box="[158,322,828,853]" captionStart="FIGURES 18 – 25" captionStartId="12.[151,264,1787,1809]" captionTargetBox="[262,1331,195,1768]" captionTargetId="figure@12.[260,1334,193,1771]" captionTargetPageId="12" captionText="FIGURES 18 – 25. Seguenziidae. 18. Calliobasis lapulapui Poppe, Tagaro &amp; Dekker, 2006. RGM 608.251. Locality Anda 1. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 19. Calliobasis lapulapui Poppe, Tagaro &amp; Dekker, 2006. RGM 608.252. Locality Anda 3. (a) SEM, side view, (b) SEM detail of protoconch. 20. Thelyssina? spec. 1. RGM 608.263. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 21. Thelyssina? spec. 1. RGM 608.264. Locality Anda 2. (a) SEM detail of protoconch, (b) SEM, side view. 22. Thelyssina? spec. 1. RGM 608.265. Locality Anda 2. SEM, basal view. 23. Thelyssina? spec. 2. RGM 608.268. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 24. Thelyssina? spec. 2. RGM 608.269. Locality AndaDeVos. SEM detail of protoconch. 25. Thelyssina? spec. 2. RGM 608.270 Locality AndaDeVos. SEM, basal view." httpUri="https://zenodo.org/record/250965/files/figure.png" pageId="13" pageNumber="114">Figures 20–22</figureCitation>
)
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<emphasis id="F3DDEAB7FFB06D2DFF1FFC305B00FC62" bold="true" box="[151,262,900,925]" pageId="13" pageNumber="114">Material.</emphasis>
Anda 2 (5); AndaDeVos (2).
</paragraph>
<paragraph id="C11636A5FFB06D2DFF4FFC135F53FBF7" blockId="13.[151,1436,900,1320]" pageId="13" pageNumber="114">
<emphasis id="F3DDEAB7FFB06D2DFF4FFC135B9CFC3F" bold="true" box="[199,410,935,960]" pageId="13" pageNumber="114">Characterization.</emphasis>
Shell minute, H
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mm; P 1.1 whorls, with fine anastomosing sculpture, DN
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; P/T boundary rounded varix; second varix at 1.6 whorls; T whorls with a sharp keel; microsculpture granulate; body whorl with two sharp spiral ribs; aperture rounded; umbilicus narrow; umbilical ridge weak.
</paragraph>
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<subSubSection id="89B3652EFFB06D2DFF4FFBA05EB5FAD7" pageId="13" pageNumber="114" type="discussion">
<paragraph id="C11636A5FFB06D2DFF4FFBA05EB5FAD7" blockId="13.[151,1436,900,1320]" pageId="13" pageNumber="114">
<emphasis id="F3DDEAB7FFB06D2DFF4FFBA05B3DFBD2" bold="true" box="[199,315,1044,1069]" pageId="13" pageNumber="114">Remarks.</emphasis>
This seguenzioid skeneimorph species is provisionally placed in
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<emphasis id="F3DDEAB7FFB06D2DFB96FBA05E90FBD2" box="[1054,1174,1044,1069]" italics="true" pageId="13" pageNumber="114">Thelyssina</emphasis>
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, based on the presence of dendritic teleoconch sculpture and a keeled shoulder. At the highest magnifications available, we observed very brilliantly shiny shell material at the inner side of the outer lip that we interpret as nacre. The small protoconch is also similar to the protoconch of the
<typeStatus id="1E128807FFB06D2DFDCDFB345873FB67" box="[581,629,1152,1176]" pageId="13" pageNumber="114">type</typeStatus>
species for the genus,
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<emphasis id="F3DDEAB7FFB06D2DFC09FB3559E9FB67" box="[897,1007,1151,1176]" italics="true" pageId="13" pageNumber="114">T. sterrha</emphasis>
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, in the anastomosing sculpture and characteristic shape. However, this assignment is uncertain, because the fossil species is relatively tall and more strongly granulate and keeled.
<emphasis id="F3DDEAB7FFB06D2DFD28FB735920FB1F" box="[672,806,1223,1248]" italics="true" pageId="13" pageNumber="114">
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?
</emphasis>
spec. 1 also resembles
<taxonomicName id="06A94D26FFB06D2DFBB5FB735ADCFAFA" authority="Golikov, 1967" authorityName="Golikov" authorityYear="1967" class="Gastropoda" family="Trochidae" genus="Conotalopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="minima">
<emphasis id="F3DDEAB7FFB06D2DFBB5FB735F2EFB1F" box="[1085,1320,1223,1248]" italics="true" pageId="13" pageNumber="114">Conotalopia minima</emphasis>
(
<bibRefCitation id="A5384B54FFB06D2DFAB1FB735AD5FAFA" author="Golikov" pageId="13" pageNumber="114" refString="Golikov, A. N. (1967) in Golikov, A. N. &amp; Scarlato, O. A. Molluscs of the Possiet Bay (Sea of Japan) and their ecology. Trudy Zoologicheskogo Instituta Leningrad, 42, 5 - 154." type="journal article" year="1967">Golikov, 1967</bibRefCitation>
)
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, a Recent trochid species from
<collectingCountry id="B9BE7635FFB06D2DFDCDFB59588EFAFB" box="[581,648,1261,1284]" name="Japan" pageId="13" pageNumber="114">Japan</collectingCountry>
, in general shape, but it has a smaller protoconch relative to the first teleoconch whorl, a dendritic granulate sculpture on the teleoconch and a narrower umbilicus.
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Genus
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<emphasis id="F3DDEAB7FFB06D2DFF66FDD85B76FD79" bold="true" box="[238,368,620,646]" italics="true" pageId="13" pageNumber="114">Thelyssina</emphasis>
<bibRefCitation id="A5384B54FFB06D2DFEFFFDD85834FD79" author="Marshall" box="[375,562,620,646]" pageId="13" pageNumber="114" refString="Marshall, B. A. (1983) Recent and Tertiary Seguenziidae (Mollusca: Gastropoda) from the New Zealand region. New Zealand Journal of Zoology, 10, 235 - 262. http: // dx. doi. org / 10.1080 / 03014223.1983.10423911" type="journal article" year="1983">
<collectingCountry id="B9BE7635FFB06D2DFEFFFDD85BEDFD79" box="[375,491,620,646]" name="Marshall Islands" pageId="13" pageNumber="114">Marshall</collectingCountry>
, 1983
</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB06D2DFF1FFD065E72FD37" box="[151,1140,690,712]" pageId="13" pageNumber="114" type="materials_examined">
<paragraph id="C11636A5FFB06D2DFF1FFD065E72FD37" blockId="13.[151,1140,690,712]" box="[151,1140,690,712]" pageId="13" pageNumber="114">
<emphasis id="F3DDEAB7FFB06D2DFF1FFD065B24FD37" bold="true" box="[151,290,690,712]" pageId="13" pageNumber="114">
<typeStatus id="1E128807FFB06D2DFF1FFD065ACAFD37" box="[151,204,690,712]" pageId="13" pageNumber="114">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="06A94D26FFB06D2DFEA1FD065883FD37" authority="Marshall, 1983" authorityName="Marshall" authorityYear="1983" box="[297,645,690,712]" class="Gastropoda" family="Seguenziidae" genus="Thelyssina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="sterrha">
<emphasis id="F3DDEAB7FFB06D2DFEA1FD065BE3FD37" box="[297,485,690,712]" italics="true" pageId="13" pageNumber="114">Thelyssina sterrha</emphasis>
<bibRefCitation id="A5384B54FFB06D2DFE63FD065883FD37" author="Marshall" box="[491,645,690,712]" pageId="13" pageNumber="114" refString="Marshall, B. A. (1983) Recent and Tertiary Seguenziidae (Mollusca: Gastropoda) from the New Zealand region. New Zealand Journal of Zoology, 10, 235 - 262. http: // dx. doi. org / 10.1080 / 03014223.1983.10423911" type="journal article" year="1983">
<collectingCountry id="B9BE7635FFB06D2DFE63FD06584EFD37" box="[491,584,690,712]" name="Marshall Islands" pageId="13" pageNumber="114">Marshall</collectingCountry>
, 1983
</bibRefCitation>
</taxonomicName>
(by original designation); Recent,
<collectingCountry id="B9BE7635FFB06D2DFC6DFD075E76FD37" box="[997,1136,690,712]" name="New Zealand" pageId="13" pageNumber="114">New Zealand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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170
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<mods:title id="90388B9657F1D584D502F8205A41E0DD">On some Vetigastropoda (Mollusca, Gastropoda) from the Plio-Pleistocene of the Philippines with descriptions of three new species</mods:title>
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<paragraph id="C11636A5FFB36D2EFF1FFDA8580DFDC9" blockId="14.[151,523,540,599]" box="[151,523,540,566]" pageId="14" pageNumber="115">
<heading id="9A5E81C9FFB36D2EFF1FFDA8580DFDC9" bold="true" box="[151,523,540,566]" fontSize="11" level="1" pageId="14" pageNumber="115" reason="1">
<emphasis id="F3DDEAB7FFB36D2EFF1FFDA8580DFDC9" bold="true" box="[151,523,540,566]" pageId="14" pageNumber="115">
<taxonomicName id="06A94D26FFB36D2EFF1FFDA85B8AFDC9" ID-CoL="dc3d7180-fb3a-4bd0-bfd3-f33cc950bf1b" box="[151,396,540,566]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="arenosa">
<emphasis id="F3DDEAB7FFB36D2EFF1FFDA85B8AFDC9" bold="true" box="[151,396,540,566]" italics="true" pageId="14" pageNumber="115">Calliotropis arenosa</emphasis>
</taxonomicName>
nov. spec.
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</paragraph>
<paragraph id="C11636A5FFB36D2EFF1FFD8A5B4DFDA8" blockId="14.[151,523,540,599]" box="[151,331,574,599]" pageId="14" pageNumber="115">
(
<figureCitation id="59922A20FFB36D2EFF16FD8A5B45FDA8" box="[158,323,574,599]" captionStart="FIGURES 26 – 32" captionStartId="15.[151,264,1793,1815]" captionTargetBox="[259,1325,197,1767]" captionTargetId="figure@15.[257,1328,195,1768]" captionTargetPageId="15" captionText="FIGURES 26 – 32. Calliotropidae. 26. Calliotropis arenosa nov. spec. RGM 608.272 (holotype). Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 27. Calliotropis arenosa nov. spec. RGM 608.273 (paratype). Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 28. Calliotropis arenosa nov. spec. RGM 608.274 (paratype). Locality Anda 3. (a) rear view, (b) apertural view, (c) apical view. 29. Calliotropis arenosa nov. spec. RGM 608.275 (paratype). Locality AndaDeVos. SEM detail of protoconch. 30. Calliotropis cf. pyramoeides Vilvens, 2007. RGM 608.281. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 31. Calliotropis cf. pyramoeides Vilvens, 2007. RGM 608.282. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 32. Calliotropis cf. pyramoeides Vilvens, 2007. RGM 608.283. Locality Anda 1. (a) SEM, side view, (b) SEM detail of protoconch." httpUri="https://zenodo.org/record/250966/files/figure.png" pageId="14" pageNumber="115">Figures 26–29</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB36D2EFF1FFD325B18FCF4" pageId="14" pageNumber="115" type="materials_examined">
<paragraph id="C11636A5FFB36D2EFF1FFD325B62FD3C" blockId="14.[151,1436,646,1931]" pageId="14" pageNumber="115">
<emphasis id="F3DDEAB7FFB36D2EFF1FFD325B42FD60" bold="true" box="[151,324,646,671]" pageId="14" pageNumber="115">
<typeStatus id="1E128807FFB36D2EFF1FFD325AD4FD60" box="[151,210,646,671]" pageId="14" pageNumber="115">Type</typeStatus>
material.
</emphasis>
<typeStatus id="1E128807FFB36D2EFEC3FD325BABFD60" box="[331,429,646,671]" pageId="14" pageNumber="115" type="holotype">holotype</typeStatus>
<collectionCode id="A7B8AE60FFB36D2EFE3BFD335BF5FD60" box="[435,499,647,671]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="115">RGM</collectionCode>
608.272.
<typeStatus id="1E128807FFB36D2EFDEAFD3358C2FD60" box="[610,708,647,671]" pageId="14" pageNumber="115" type="paratype">Paratype</typeStatus>
<collectionCode id="A7B8AE60FFB36D2EFD43FD33590DFD60" box="[715,779,647,671]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="115">RGM</collectionCode>
608.273 and
<collectionCode id="A7B8AE60FFB36D2EFC2AFD3359E4FD60" box="[930,994,647,671]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="115">RGM</collectionCode>
608.275 (
<typeStatus id="1E128807FFB36D2EFBD9FD335E85FD60" box="[1105,1155,647,671]" pageId="14" pageNumber="115">type</typeStatus>
locality);
<typeStatus id="1E128807FFB36D2EFB7DFD335F53FD60" box="[1269,1365,647,671]" pageId="14" pageNumber="115" type="paratype">paratype</typeStatus>
<collectionCode id="A7B8AE60FFB36D2EFAD4FD335F9AFD60" box="[1372,1436,647,671]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="115">RGM</collectionCode>
608.274 (Anda 3).
</paragraph>
<paragraph id="C11636A5FFB36D2EFF4FFD7A5B18FCF4" blockId="14.[151,1436,646,1931]" pageId="14" pageNumber="115">
<emphasis id="F3DDEAB7FFB36D2EFF4FFD7A5B76FD18" bold="true" box="[199,368,718,743]" pageId="14" pageNumber="115">
<typeStatus id="1E128807FFB36D2EFF4FFD7A5B04FD18" box="[199,258,718,743]" pageId="14" pageNumber="115">Type</typeStatus>
locality.
</emphasis>
AndaDeVos, Bolinao province,
<collectingCountry id="B9BE7635FFB36D2EFC8FFD7A598EFD18" box="[775,904,718,743]" name="Philippines" pageId="14" pageNumber="115">Philippines</collectingCountry>
, Santa Cruz Formation, Pliocene or Early Pleistocene.
</paragraph>
</subSubSection>
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<paragraph id="C11636A5FFB36D2EFF4FFCA259C7FCD0" blockId="14.[151,1436,646,1931]" box="[199,961,790,815]" pageId="14" pageNumber="115">
<emphasis id="F3DDEAB7FFB36D2EFF4FFCA25BA2FCD0" bold="true" box="[199,420,790,815]" pageId="14" pageNumber="115">Derivatio nominis.</emphasis>
Named after the sediment covering adult shells.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB36D2EFF4FFC8E5EBCFCAC" box="[199,1210,826,851]" pageId="14" pageNumber="115" type="materials_examined">
<paragraph id="C11636A5FFB36D2EFF4FFC8E5EBCFCAC" blockId="14.[151,1436,646,1931]" box="[199,1210,826,851]" pageId="14" pageNumber="115">
<emphasis id="F3DDEAB7FFB36D2EFF4FFC8E5B87FCAC" bold="true" box="[199,385,826,851]" pageId="14" pageNumber="115">Other material.</emphasis>
Anda 2 (1); Anda 5 (1); Anda 6 (49); AndaDeVos (102); AndaClif3 (18).
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB36D2EFF4FFCEA58F6FC41" pageId="14" pageNumber="115" type="diagnosis">
<paragraph id="C11636A5FFB36D2EFF4FFCEA58F6FC41" blockId="14.[151,1436,646,1931]" pageId="14" pageNumber="115">
<emphasis id="F3DDEAB7FFB36D2EFF4FFCEA5B47FC88" bold="true" box="[199,321,862,887]" pageId="14" pageNumber="115">Diagnosis.</emphasis>
Shell conical, covered in sediment, H
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,
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W
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mm; P large, DN
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); T sculpture of multiple axial ribs and two spiral ribs, with granules at intersections; suture canaliculate; aperture with three lamellae and a columellar denticle; umbilicus narrow.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB36D2EFF4FFC7E5919F970" pageId="14" pageNumber="115" type="description">
<paragraph id="C11636A5FFB36D2EFF4FFC7E59B2FA25" blockId="14.[151,1436,646,1931]" pageId="14" pageNumber="115">
<emphasis id="F3DDEAB7FFB36D2EFF4FFC7E5B5FFC1C" bold="true" box="[199,345,970,995]" pageId="14" pageNumber="115">Description.</emphasis>
This species has a conical-shaped shell that is covered with a thin, even layer of fine sand from the early teleoconch onwards. This sand is most likely agglutinated by the live animal. The protoconch has a large, globose nucleus (DN 185–192 µm) and it is finely granular and consists of 1.0 whorls. The P/T boundary is marked by a thin varix. The teleoconch bears microscopic pustules. The first teleoconch whorl is sculpted with 15 axial ribs. Their interspaces are about three times as wide as their width. Sediment cover develops within the first 0.5 teleoconch whorl. Two spiral ribs appear on the second teleoconch whorl. The axial ribs develop granules at intersections with these spiral ribs. The whorl profile changes from round to angular as the spiral ribs become more pronounced. The area between the spiral ribs becomes slightly concave. The suture is canaliculate. The body whorl bears four additional spiral ribs. The area between the middle rib and basal rib is also slightly concave. The three basal ribs are slightly weaker than the three ribs on the preceding whorls. The base is slightly convex. The aperture is weakly nacreous and subcircular to subquadrate. The parietal margin is slightly convex and contains a thin callus and a weak knob on its upper part. The general shape of the outer lip is rounded, but the spiral ribs produce local angulations. The inner side of the outer lip contains two weak lamellae and a stronger basal lamella. The columellar lip is also rounded but the columella is nearly straight. A weak denticle is present at midpoint of columella in
<typeStatus id="1E128807FFB36D2EFF1FFA765AFAFA24" box="[151,252,1474,1499]" pageId="14" pageNumber="115" type="holotype">holotype</typeStatus>
, but absent in most other specimens. The umbilicus is narrow.
</paragraph>
<paragraph id="C11636A5FFB36D2EFF4FFA525919F970" blockId="14.[151,1436,646,1931]" pageId="14" pageNumber="115">
<emphasis id="F3DDEAB7FFB36D2EFF4FFA525B79FA00" bold="true" box="[199,383,1510,1535]" pageId="14" pageNumber="115">Differentiation.</emphasis>
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<emphasis id="F3DDEAB7FFB36D2EFE04FA52587FFA01" box="[396,633,1510,1535]" italics="true" pageId="14" pageNumber="115">Calliotropis arenosa</emphasis>
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is most similar to the slightly taller
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<emphasis id="F3DDEAB7FFB36D2EFBCAFA535ED8FA00" box="[1090,1246,1510,1535]" italics="true" pageId="14" pageNumber="115">C. acherontis</emphasis>
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, 1979
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(Recent, southwest-Pacific), but the latter species does not agglutinate sand to the shell. Furthermore, the specimens figured by
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(1979)
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and
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do not appear to have lamellae or a denticle in the aperture, while the larger specimens in the studied material, in which the aperture is not filled with sediment, always have lamellae and sometimes also have a denticle.
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<emphasis id="F3DDEAB7FFB36D2EFF4FF92E5B3DF94C" bold="true" box="[199,315,1690,1715]" pageId="14" pageNumber="115">Remarks.</emphasis>
The layer of sediment on the adults of this species is considered here as a characteristic of the species, because it is present in all adult material and its appearance, thickness and evenness is constant and similar in all adults. Furthermore, such a characteristic layer is not present in any other species of shells in the studied material. The origin of this layer of sediment remains speculative, but is perhaps facilitated by periostracal fluids, as it is in the Caenogastropod genus
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<emphasis id="F3DDEAB7FFB36D2EFDC9F89E589BF8BC" box="[577,669,1834,1859]" italics="true" pageId="14" pageNumber="115">Scaliola</emphasis>
Adams, 1860 (
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)
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. One specimen in the studied material has grown wider whorls after a break in the shell, so injuries to snails of this species may sometimes alter shell development.
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were previously regarded as a subfamily of
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by
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Bouchet
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. However,
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Kano
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regarded the
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(
<figureCitation id="59922A20FFB46D29FF16FF0E5B44FF2C" box="[158,322,186,211]" captionStart="FIGURES 8 – 12" captionStartId="8.[151,264,1849,1871]" captionTargetBox="[243,1351,195,1825]" captionTargetId="figure@8.[241,1354,193,1828]" captionTargetPageId="8" captionText="FIGURES 8 – 12. Seguenziidae. 8. Seguenzia donaldi Ladd, 1982. RGM 608.215. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 9. Seguenzia donaldi Ladd, 1982. RGM 608.216. Locality Anda 3. (a) rear view, (b) apertural view. 10. Seguenzia cf. elegantissima Poppe, Tagaro &amp; Dekker, 2006. RGM 608.221. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 11. Seguenzia cf. elegantissima Poppe, Tagaro &amp; Dekker, 2006. RGM 608.222. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) spire. 12. Seguenzia cf. elegantissima Poppe, Tagaro &amp; Dekker, 2006. RGM 608.223. Locality AndaDeVos. SEM detail of protoconch." httpUri="https://zenodo.org/record/250963/files/figure.png" pageId="9" pageNumber="110">Figures 10–12</figureCitation>
)
</paragraph>
<paragraph id="C11636A5FFB46D29FF1FFEB65981FEC8" blockId="9.[151,1071,258,311]" pageId="9" pageNumber="110">
2006
<taxonomicName id="06A94D26FFB46D29FF7DFEB75BEBFEE7" box="[245,493,258,280]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="9" pageNumber="110" phylum="Mollusca" rank="species" species="elegantissima">
<emphasis id="F3DDEAB7FFB46D29FF7DFEB75BEBFEE7" box="[245,493,258,280]" italics="true" pageId="9" pageNumber="110">Seguenzia elegantissima</emphasis>
</taxonomicName>
nov. spec.—Poppe, Tagaro &amp; Dekker: p. 26, pl. 3, fig. 2. 2008a
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<emphasis id="F3DDEAB7FFB46D29FF7DFE965BEBFEC8" box="[245,493,289,311]" italics="true" pageId="9" pageNumber="110">Seguenzia elegantissima</emphasis>
</taxonomicName>
—Poppe &amp; Tagaro: p. 162, pl. 26, fig. 4.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB46D29FF1FFED05801FDCF" pageId="9" pageNumber="110" type="materials_examined">
<paragraph id="C11636A5FFB46D29FF1FFED05985FE83" blockId="9.[151,1436,356,885]" box="[151,899,356,381]" pageId="9" pageNumber="110">
<emphasis id="F3DDEAB7FFB46D29FF1FFED05B00FE82" bold="true" box="[151,262,356,381]" pageId="9" pageNumber="110">Material.</emphasis>
Anda 6 (6); AndaDeVos (18); AndaClif3 (7); Tiep 1 (1).
</paragraph>
<paragraph id="C11636A5FFB46D29FF4FFE335801FDCF" blockId="9.[151,1436,356,885]" pageId="9" pageNumber="110">
<emphasis id="F3DDEAB7FFB46D29FF4FFE335B9CFE5F" bold="true" box="[199,410,391,416]" pageId="9" pageNumber="110">Characterization.</emphasis>
High spired, H
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,
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W
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mm; P 1.1 whorls, with erect nucleus and two discontinuous spiral ribs, DN
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; T with fine but regular and well developed axial lirae on early whorls, fading on later whorls; slightly erect upper keel; markedly beaded subsutural cord on later T whorls; body whorl with three peripheral spiral ribs and four basal spiral ribs; columellar lip with low basal fold or broadly folded; upper apertural sinus pronounced and thickened.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB46D29FF4FFD885983FD87" pageId="9" pageNumber="110" type="distribution">
<paragraph id="C11636A5FFB46D29FF4FFD885983FD87" blockId="9.[151,1436,356,885]" pageId="9" pageNumber="110">
<emphasis id="F3DDEAB7FFB46D29FF4FFD885B66FDAC" bold="true" box="[199,352,572,597]" pageId="9" pageNumber="110">Distribution.</emphasis>
Apart from the studied material, this species is known only from the
<typeStatus id="1E128807FFB46D29FBF2FD895EACFDAA" box="[1146,1194,573,597]" pageId="9" pageNumber="110">type</typeStatus>
locality in the Bohol Sea,
<collectingCountry id="B9BE7635FFB46D29FF45FDEB5B48FD87" box="[205,334,607,632]" name="Philippines" pageId="9" pageNumber="110">Philippines</collectingCountry>
, from
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water depth (
<bibRefCitation id="A5384B54FFB46D29FD26FDD4597FFD87" author="Poppe" box="[686,889,607,632]" pageId="9" pageNumber="110" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter" year="2006">
Poppe
<emphasis id="F3DDEAB7FFB46D29FD75FDD55933FD87" box="[765,821,607,632]" italics="true" pageId="9" pageNumber="110">et al.</emphasis>
2006
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB46D29FF4FFD305F39FC8A" pageId="9" pageNumber="110" type="discussion">
<paragraph id="C11636A5FFB46D29FF4FFD305F39FC8A" blockId="9.[151,1436,356,885]" pageId="9" pageNumber="110">
<emphasis id="F3DDEAB7FFB46D29FF4FFD305B3DFD62" bold="true" box="[199,315,644,669]" pageId="9" pageNumber="110">Remarks.</emphasis>
The elongate shape, thin and regular axials, the beaded suprasutural ridge and the folded columellar lip distinguish this species from the two species treated above. The assignment of the present material to
<taxonomicName id="06A94D26FFB46D29FA0FFD1C5B37FD1A" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="9" pageNumber="110" phylum="Mollusca" rank="species" species="elegantissima">
<emphasis id="F3DDEAB7FFB46D29FA0FFD1C5B37FD1A" italics="true" pageId="9" pageNumber="110">S. elegantissima</emphasis>
</taxonomicName>
is uncertain, because the outer lip is severely damaged in all specimens, making a complete comparison with the material described by
<bibRefCitation id="A5384B54FFB46D29FDF5FD445953FCF7" box="[637,853,751,776]" pageId="9" pageNumber="110" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter">
Poppe
<emphasis id="F3DDEAB7FFB46D29FD43FD455902FCF7" box="[715,772,751,776]" italics="true" pageId="9" pageNumber="110">et al.</emphasis>
(2006)
</bibRefCitation>
impossible. Furthermore the subsutural spiral ridge appears to be smooth to marginally knobby in the recent species, whereas it is distinctly beaded in the fossil material studied here.
<taxonomicName id="06A94D26FFB46D29FE18FC8C5919FCAF" authority="Marshall, 1991" authorityName="Marshall" authorityYear="1991" box="[400,799,823,848]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="9" pageNumber="110" phylum="Mollusca" rank="species" species="praeceps">
<emphasis id="F3DDEAB7FFB46D29FE18FC8C586BFCAF" box="[400,621,824,848]" italics="true" pageId="9" pageNumber="110">Seguenzia praeceps</emphasis>
<bibRefCitation id="A5384B54FFB46D29FDFCFC835919FCAF" author="Marshall" box="[628,799,823,848]" pageId="9" pageNumber="110" refString="Marshall, B. A. (1991) Mollusca Gastropoda: Seguenziidae from New Caledonia and the Loyalty Islands. Memoires du Museum Nationale d'Histoire Naturelle Serie A Zoologie, 150, 41 - 109." type="journal article" year="1991">
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, 1991
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="06A94D26FFB46D29FCA4FC8C5E4EFCAF" authority="Marshall, 1983" authorityName="Marshall" authorityYear="1983" box="[812,1096,823,848]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="9" pageNumber="110" phylum="Mollusca" rank="species" species="fulgida">
<emphasis id="F3DDEAB7FFB46D29FCA4FC8C5991FCAF" box="[812,919,823,848]" italics="true" pageId="9" pageNumber="110">S. fulgida</emphasis>
<bibRefCitation id="A5384B54FFB46D29FC16FC835E4EFCAF" author="Marshall" box="[926,1096,823,848]" pageId="9" pageNumber="110" refString="Marshall, B. A. (1983) Recent and Tertiary Seguenziidae (Mollusca: Gastropoda) from the New Zealand region. New Zealand Journal of Zoology, 10, 235 - 262. http: // dx. doi. org / 10.1080 / 03014223.1983.10423911" type="journal article" year="1983">
<collectingCountry id="B9BE7635FFB46D29FC16FC835E02FCAF" box="[926,1028,823,848]" name="Marshall Islands" pageId="9" pageNumber="110">Marshall</collectingCountry>
, 1983
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="06A94D26FFB46D29FBF6FC8C5F9AFCAF" authority="Marshall, 1983" authorityName="Marshall" authorityYear="1983" box="[1150,1436,823,848]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="9" pageNumber="110" phylum="Mollusca" rank="species" species="serrata">
<emphasis id="F3DDEAB7FFB46D29FBF6FC8C5EEAFCAF" box="[1150,1260,824,848]" italics="true" pageId="9" pageNumber="110">S. serrata</emphasis>
<bibRefCitation id="A5384B54FFB46D29FB7AFC835F9AFCAF" author="Marshall" box="[1266,1436,823,848]" pageId="9" pageNumber="110" refString="Marshall, B. A. (1983) Recent and Tertiary Seguenziidae (Mollusca: Gastropoda) from the New Zealand region. New Zealand Journal of Zoology, 10, 235 - 262. http: // dx. doi. org / 10.1080 / 03014223.1983.10423911" type="journal article" year="1983">
<collectingCountry id="B9BE7635FFB46D29FB7AFC835F5EFCAF" box="[1266,1368,823,848]" name="Marshall Islands" pageId="9" pageNumber="110">Marshall</collectingCountry>
, 1983
</bibRefCitation>
</taxonomicName>
are quite similar in shape, but their subsutural ridge is not as strongly crenulated as in the studied material.
</paragraph>
</subSubSection>
</treatment>
</document>

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<heading id="9A5E81C9FFB46D29FF1FFBC25BF9FB6F" bold="true" box="[151,511,1142,1168]" fontSize="11" level="1" pageId="9" pageNumber="110" reason="1">
<emphasis id="F3DDEAB7FFB46D29FF1FFBC25BF9FB6F" bold="true" box="[151,511,1142,1168]" pageId="9" pageNumber="110">
<taxonomicName id="06A94D26FFB46D29FF1FFBC25B86FB6F" ID-CoL="3JGWT" box="[151,384,1142,1168]" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="9" pageNumber="110" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="F3DDEAB7FFB46D29FF1FFBC25B86FB6F" bold="true" box="[151,384,1142,1168]" italics="true" pageId="9" pageNumber="110">Halystina conoidea</emphasis>
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nov. spec.
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<paragraph id="C11636A5FFB46D29FF1FFB2D5B4CFB4D" blockId="9.[151,511,1142,1202]" box="[151,330,1177,1202]" pageId="9" pageNumber="110">
(
<figureCitation id="59922A20FFB46D29FF16FB2D5B44FB4D" box="[158,322,1177,1202]" captionStart="FIGURES 13 – 17" captionStartId="11.[151,264,1855,1877]" captionTargetBox="[235,1343,195,1825]" captionTargetId="figure@11.[233,1346,193,1828]" captionTargetPageId="11" captionText="FIGURES 13 – 17. Seguenziidae. 13. Halystina conoidea nov. spec. RGM 608.229 (holotype). Locality AndaDeVos. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 14. Halystina conoidea nov. spec. RGM 608.230 (paratype). Locality Anda 6. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 15. Halystina conoidea nov. spec. RGM 608.231 (paratype). Locality AndaDeVos. SEM detail of protoconch. 16. Fluxinella membranacea Marshall, 1991. RGM 608.239. Locality Anda 1. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 17. Fluxinella membranacea Marshall, 1991. RGM 608.240. Locality Anda 1. (a) apical view, (b) SEM detail of surface sculpture." httpUri="https://zenodo.org/record/250964/files/figure.png" pageId="9" pageNumber="110">Figures 13–15</figureCitation>
)
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<emphasis id="F3DDEAB7FFB46D29FF1FFB555B49FB05" bold="true" box="[151,335,1249,1274]" pageId="9" pageNumber="110">
<typeStatus id="1E128807FFB46D29FF1FFB555AD4FB05" box="[151,210,1249,1274]" pageId="9" pageNumber="110">Type</typeStatus>
material.
</emphasis>
<typeStatus id="1E128807FFB46D29FEE8FB555BCEFB05" box="[352,456,1249,1274]" pageId="9" pageNumber="110" type="holotype">Holotype</typeStatus>
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608.229.
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608.230 (Anda 6); 7
<typeStatus id="1E128807FFB46D29FBFCFB565ED9FB05" box="[1140,1247,1250,1274]" pageId="9" pageNumber="110" type="paratype">paratypes</typeStatus>
<collectionCode id="A7B8AE60FFB46D29FB78FB565F36FB05" box="[1264,1328,1250,1274]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="110">RGM</collectionCode>
608.235 (AndaDeVos).
</paragraph>
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<emphasis id="F3DDEAB7FFB46D29FF4FFA9D5B63FABD" bold="true" box="[199,357,1321,1346]" pageId="9" pageNumber="110">
<typeStatus id="1E128807FFB46D29FF4FFA9D5B04FABD" box="[199,258,1321,1346]" pageId="9" pageNumber="110">Type</typeStatus>
locality.
</emphasis>
AndaDeVos, Bolinao,
<collectingCountry id="B9BE7635FFB46D29FDEFFA9D58EEFABD" box="[615,744,1321,1346]" name="Philippines" pageId="9" pageNumber="110">Philippines</collectingCountry>
, Santa Cruz Formation, Pliocene or Early Pleistocene.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB46D29FF4FFAF95E32FA99" box="[199,1076,1357,1382]" pageId="9" pageNumber="110" type="etymology">
<paragraph id="C11636A5FFB46D29FF4FFAF95E32FA99" blockId="9.[151,1436,1249,2030]" box="[199,1076,1357,1382]" pageId="9" pageNumber="110">
<emphasis id="F3DDEAB7FFB46D29FF4FFAF95BA2FA99" bold="true" box="[199,420,1357,1382]" pageId="9" pageNumber="110">Derivatio nominis.</emphasis>
Named after the conical outline of this species. Adjective.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB46D29FF4FFAC55E7EFA75" box="[199,1144,1393,1418]" pageId="9" pageNumber="110" type="materials_examined">
<paragraph id="C11636A5FFB46D29FF4FFAC55E7EFA75" blockId="9.[151,1436,1249,2030]" box="[199,1144,1393,1418]" pageId="9" pageNumber="110">
<emphasis id="F3DDEAB7FFB46D29FF4FFAC55B87FA75" bold="true" box="[199,385,1393,1418]" pageId="9" pageNumber="110">Other material.</emphasis>
Anda 5 (1); Anda 6 (2); AndaDeVos (7); AndaClif3 (4); Tiep 1 (1).
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB46D29FF4FFA2159CBF9E5" pageId="9" pageNumber="110" type="diagnosis">
<paragraph id="C11636A5FFB46D29FF4FFA2159CBF9E5" blockId="9.[151,1436,1249,2030]" pageId="9" pageNumber="110">
<emphasis id="F3DDEAB7FFB46D29FF4FFA215B47FA51" bold="true" box="[199,321,1429,1454]" pageId="9" pageNumber="110">Diagnosis.</emphasis>
Shell conoid, H
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,
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W
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mm, DN
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, low spiral ribs, thin axial lirae; axial lirae gradually disappearing; two slightly thicker but low spiral ribs at middle of whorl, just above suture; after second teleoconch whorl, third spiral rib appears just below suture; body whorl with additional rib on periphery; 11 spiral ribs on the convex base; base of aperture with broad short siphonal canal.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB46D2AFF4FF9915EAEFD7A" lastPageId="10" lastPageNumber="111" pageId="9" pageNumber="110" type="description">
<paragraph id="C11636A5FFB46D2AFF4FF9915B73FE77" blockId="9.[151,1436,1249,2030]" lastBlockId="10.[151,1436,151,645]" lastPageId="10" lastPageNumber="111" pageId="9" pageNumber="110">
<emphasis id="F3DDEAB7FFB46D29FF4FF9915B5FF9C1" bold="true" box="[199,345,1573,1598]" pageId="9" pageNumber="110">Description.</emphasis>
The protoconch is paucispiral and has rounded whorls and a relatively dull white surface. The protoconch surface in the studied specimens is corroded, but two spiral ribs appear to be present. The nucleus is slightly inclined. The P/T boundary is located at about 1.1 whorls. The teleoconch ornamentation starts with thin axial riblets that are low and narrow but well delimited. They have a reverse sigmoid shape and are most pronounced at the upper quarter of the whorl. Within 0.2 whorls from the P/T boundary, a very low and rounded spiral rib develops at half the height of the whorl that subsequently moves to two-thirds of the whorl height within half a whorl. This poorly defined rib delimits a low shoulder on the first and second teleoconch whorl. Already on the first teleoconch whorl, a second spiral develops at approximately one quarter of the whorl height. On the second teleoconch whorl, a narrow subsutural ridge develops that forces the suture into a narrow and shallow depression. The whorl profile becomes gradually flatter with the initial spiral rib becoming even lower and located at 0.6 of the whorl height and the basal rib at 0.2. In between the spiral ribs, the profile is straight or slightly concave. The axials (approximately 55–60 on the second teleoconch whorl) gradually disappear over most parts of the whorls, but can remain visible at the top of the whorls. The suprasutural spiral forms a low and broad median keel on the body whorl, together with a spiral rib on the periphery. At the slightly convex base of the body whorl, 11 more or less regularly spaced low spiral ribs occur, whose interspaces are about twice as wide as the ribs. The aperture is damaged and/or partially covered in all specimens, but is subtrapezoidal in juveniles and subquadrate in adults, where the outer lip is expanded basally. The outer lip also has a retraction in the upper part, between the suture and the periphery. The columellar lip is barely thickened, appears slightly twisted and grades into a broad and shallow basal sinus. A single specimen contains a very subtle columellar fold on the base of the columella. The umbilicus is rimate.
</paragraph>
<paragraph id="C11636A5FFB76D2AFF4FFE205EAEFD7A" blockId="10.[151,1436,151,645]" pageId="10" pageNumber="111">
<emphasis id="F3DDEAB7FFB76D2AFF4FFE205B79FE52" bold="true" box="[199,383,404,429]" pageId="10" pageNumber="111">Differentiation.</emphasis>
<taxonomicName id="06A94D26FFB76D2AFE02FE20586CFE52" box="[394,618,404,429]" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="F3DDEAB7FFB76D2AFE02FE20586CFE52" box="[394,618,404,429]" italics="true" pageId="10" pageNumber="111">Halystina conoidea</emphasis>
</taxonomicName>
differs from
<taxonomicName id="06A94D26FFB76D2AFC83FE215E42FE53" authority="Marshall, 1991" authorityName="Marshall" authorityYear="1991" box="[779,1092,404,429]" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="vaubani">
<emphasis id="F3DDEAB7FFB76D2AFC83FE21598DFE52" box="[779,907,404,429]" italics="true" pageId="10" pageNumber="111">H. vaubani</emphasis>
<bibRefCitation id="A5384B54FFB76D2AFC1FFE205E42FE53" author="Marshall" box="[919,1092,404,429]" pageId="10" pageNumber="111" refString="Marshall, B. A. (1991) Mollusca Gastropoda: Seguenziidae from New Caledonia and the Loyalty Islands. Memoires du Museum Nationale d'Histoire Naturelle Serie A Zoologie, 150, 41 - 109." type="journal article" year="1991">
<collectingCountry id="B9BE7635FFB76D2AFC1FFE2059FBFE52" box="[919,1021,404,429]" name="Marshall Islands" pageId="10" pageNumber="111">Marshall</collectingCountry>
, 1991
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="06A94D26FFB76D2AFB0CFE215AD4FE2F" authority="Marshall, 1991" authorityName="Marshall" authorityYear="1991" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="caledonica">
<emphasis id="F3DDEAB7FFB76D2AFB0CFE215F20FE52" box="[1156,1318,404,429]" italics="true" pageId="10" pageNumber="111">H. caledonica</emphasis>
<bibRefCitation id="A5384B54FFB76D2AFAB9FE205AD4FE2F" author="Marshall" pageId="10" pageNumber="111" refString="Marshall, B. A. (1991) Mollusca Gastropoda: Seguenziidae from New Caledonia and the Loyalty Islands. Memoires du Museum Nationale d'Histoire Naturelle Serie A Zoologie, 150, 41 - 109." type="journal article" year="1991">
<collectingCountry id="B9BE7635FFB76D2AFAB9FE205F91FE52" box="[1329,1431,404,429]" name="Marshall Islands" pageId="10" pageNumber="111">Marshall</collectingCountry>
, 1991
</bibRefCitation>
</taxonomicName>
, both Recent species from
<collectingCountry id="B9BE7635FFB76D2AFD83FE0C58BBFE2F" box="[523,701,439,464]" name="New Caledonia" pageId="10" pageNumber="111">New Caledonia</collectingCountry>
, in having axial ribs that become obsolete instead of being well developed throughout the teleoconch. It differs from
<taxonomicName id="06A94D26FFB76D2AFD6FFE695E1CFE0B" authority="Marshall, 1991" authorityName="Marshall" authorityYear="1991" box="[743,1050,476,501]" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="carinata">
<emphasis id="F3DDEAB7FFB76D2AFD6FFE69596EFE0B" box="[743,872,477,500]" italics="true" pageId="10" pageNumber="111">H. carinata</emphasis>
<bibRefCitation id="A5384B54FFB76D2AFCE6FE685E1CFE0B" author="Marshall" box="[878,1050,476,501]" pageId="10" pageNumber="111" refString="Marshall, B. A. (1991) Mollusca Gastropoda: Seguenziidae from New Caledonia and the Loyalty Islands. Memoires du Museum Nationale d'Histoire Naturelle Serie A Zoologie, 150, 41 - 109." type="journal article" year="1991">
<collectingCountry id="B9BE7635FFB76D2AFCE6FE6859D2FE0A" box="[878,980,476,501]" name="Marshall Islands" pageId="10" pageNumber="111">Marshall</collectingCountry>
, 1991
</bibRefCitation>
</taxonomicName>
, also from
<collectingCountry id="B9BE7635FFB76D2AFB1EFE695F40FE0A" box="[1174,1350,476,501]" name="New Caledonia" pageId="10" pageNumber="111">New Caledonia</collectingCountry>
, and
<taxonomicName id="06A94D26FFB76D2AFA09FE695BA9FDE7" authority="Barnard, 1963" authorityName="Barnard" authorityYear="1963" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="F3DDEAB7FFB76D2AFA09FE695AEAFDE7" italics="true" pageId="10" pageNumber="111">H. simplex</emphasis>
(
<bibRefCitation id="A5384B54FFB76D2AFF77FE4B5BA1FDE7" author="Barnard" box="[255,423,511,536]" pageId="10" pageNumber="111" refString="Barnard, K. H. (1963) Contribution to the knowledge of South African marine Mollusca. Part 4. Gastropoda: Prosobranchiata: Rhipidoglossa, Docoglossa. Tectibranchiata. Polyplacophora. Solenogastres. Scaphopoda. Annals of the South African museum, 47, 201 - 360." type="journal article" year="1963">Barnard, 1963</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="B9BE7635FFB76D2AFE74FE4B5896FDE7" box="[508,656,511,536]" name="South Africa" pageId="10" pageNumber="111">South Africa</collectingCountry>
by a more subtle shoulder. It resembles most closely
<taxonomicName id="06A94D26FFB76D2AFA9DFDB55BF0FDC2" authority="Poppe, Tagaro &amp; Dekker, 2006" authorityName="Poppe, Tagaro &amp; Dekker" authorityYear="2006" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="globulus">
<emphasis id="F3DDEAB7FFB76D2AFA9DFDB55F9AFDE7" box="[1301,1436,511,536]" italics="true" pageId="10" pageNumber="111">H. globulus</emphasis>
<bibRefCitation id="A5384B54FFB76D2AFF1FFD915BF0FDC2" author="Poppe" box="[151,502,548,573]" pageId="10" pageNumber="111" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter" year="2006">Poppe, Tagaro &amp; Dekker, 2006</bibRefCitation>
</taxonomicName>
, from the
<collectingCountry id="B9BE7635FFB76D2AFDEFFD9058EEFDC2" box="[615,744,548,573]" name="Philippines" pageId="10" pageNumber="111">Philippines</collectingCountry>
, but differs in the almost entirely closed umbilicus (instead of open).
<taxonomicName id="06A94D26FFB76D2AFF60FDF35BC1FD9F" box="[232,455,583,608]" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="F3DDEAB7FFB76D2AFF60FDF35BC1FD9F" box="[232,455,583,608]" italics="true" pageId="10" pageNumber="111">Halystina conoidea</emphasis>
</taxonomicName>
differs from
<taxonomicName id="06A94D26FFB76D2AFDEEFDFD5E06FD9F" authority="Bertolaso &amp; Palazzi, 1999" authorityName="Bertolaso &amp; Palazzi" authorityYear="1999" box="[614,1024,583,608]" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="edax">
<emphasis id="F3DDEAB7FFB76D2AFDEEFDFD58C7FD9F" box="[614,705,583,608]" italics="true" pageId="10" pageNumber="111">H. edax</emphasis>
<bibRefCitation id="A5384B54FFB76D2AFD44FDF35E06FD9F" author="Bertolaso" box="[716,1024,583,608]" pageId="10" pageNumber="111" refString="Bertolaso, L. &amp; Palazzi, S. (1999) A new representative of Pliocene Seguenziidae from the Emilia-Romagno Appenine region. Bollettino Malacologico, 34, 105 - 107." type="journal article" year="1999">Bertolaso &amp; Palazzi, 1999</bibRefCitation>
</taxonomicName>
, from the Pliocene of
<collectingCountry id="B9BE7635FFB76D2AFA82FDF35F38FD9F" box="[1290,1342,583,608]" name="Italy" pageId="10" pageNumber="111">Italy</collectingCountry>
, and
<taxonomicName id="06A94D26FFB76D2AFA09FDFD5BC0FD7A" authority="Thiele, 1925" authorityName="Thiele" authorityYear="1925" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="siberutensis">
<emphasis id="F3DDEAB7FFB76D2AFA09FDFD5B1BFD7A" italics="true" pageId="10" pageNumber="111">H. siberutensis</emphasis>
(
<bibRefCitation id="A5384B54FFB76D2AFEA3FDD85BB8FD7A" author="Thiele" box="[299,446,620,645]" pageId="10" pageNumber="111" refString="Thiele, J. (1925) Gastropoda der Deutschen Tiefsee-Expedition, II. Wissenschaftliche Ergebnisse der Deutschen Tiefsee- Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899, 17, 35 - 382. http: // dx. doi. org / 10.1038 / 078267 a 0" type="journal article" year="1925">Thiele, 1925</bibRefCitation>
)
</taxonomicName>
, a Recent species from
<collectingCountry id="B9BE7635FFB76D2AFD47FDD85939FD7A" box="[719,831,620,645]" name="Indonesia" pageId="10" pageNumber="111">Indonesia</collectingCountry>
, in having an angular periphery.
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<emphasis id="F3DDEAB7FFB76D2AFF1FF9AB5822F9C7" bold="true" box="[151,548,1567,1592]" pageId="10" pageNumber="111">
Genus
<taxonomicName id="06A94D26FFB76D2AFF62F9AB5822F9C7" ID-CoL="7NRKX" authority="Marshall, 1983" authorityName="Marshall" authorityYear="1983" box="[234,548,1567,1592]" class="Gastropoda" family="Seguenziidae" genus="Calliobasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFB76D2AFF62F9AB5B6CF9C7" bold="true" box="[234,362,1567,1592]" italics="true" pageId="10" pageNumber="111">Calliobasis</emphasis>
<bibRefCitation id="A5384B54FFB76D2AFEF9F9AB5822F9C7" author="Marshall" box="[369,548,1567,1592]" pageId="10" pageNumber="111" refString="Marshall, B. A. (1983) Recent and Tertiary Seguenziidae (Mollusca: Gastropoda) from the New Zealand region. New Zealand Journal of Zoology, 10, 235 - 262. http: // dx. doi. org / 10.1080 / 03014223.1983.10423911" type="journal article" year="1983">
<collectingCountry id="B9BE7635FFB76D2AFEF9F9AB5BE6F9C7" box="[369,480,1567,1592]" name="Marshall Islands" pageId="10" pageNumber="111">Marshall</collectingCountry>
, 1983
</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB76D2AFF1FF9DC5EFDF981" box="[151,1275,1640,1662]" pageId="10" pageNumber="111" type="materials_examined">
<paragraph id="C11636A5FFB76D2AFF1FF9DC5EFDF981" blockId="10.[151,1275,1640,1662]" box="[151,1275,1640,1662]" pageId="10" pageNumber="111">
<emphasis id="F3DDEAB7FFB76D2AFF1FF9DC5B24F981" bold="true" box="[151,290,1640,1662]" pageId="10" pageNumber="111">
<typeStatus id="1E128807FFB76D2AFF1FF9DC5ACAF981" box="[151,204,1640,1662]" pageId="10" pageNumber="111">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="06A94D26FFB76D2AFEA1F9DC58DFF981" authority="Cotton &amp; Godfrey, 1938" authorityName="Cotton &amp; Godfrey" authorityYear="1938" box="[297,729,1640,1662]" class="Gastropoda" family="Seguenziidae" genus="Basilissa" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="bombax">
<emphasis id="F3DDEAB7FFB76D2AFEA1F9DC5BDDF981" box="[297,475,1640,1662]" italics="true" pageId="10" pageNumber="111">Basilissa bombax</emphasis>
<bibRefCitation id="A5384B54FFB76D2AFE69F9DC58DFF981" author="Cotton" box="[481,729,1640,1662]" pageId="10" pageNumber="111" refString="Cotton, B. C. &amp; Godfrey, F. K. (1938) New species of South Australian Gastropoda. Records of the South Australian Museum, 6, 199 - 206." type="journal article" year="1938">Cotton &amp; Godfrey, 1938</bibRefCitation>
</taxonomicName>
(by original designation); Recent, southern
<collectingCountry id="B9BE7635FFB76D2AFB1EF9DC5EF0F981" box="[1174,1270,1640,1662]" name="Australia" pageId="10" pageNumber="111">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

171
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<subSubSection id="89B3652EFFB76D2AFF1FFC31598BFBBC" pageId="10" pageNumber="111" type="nomenclature">
<paragraph id="C11636A5FFB76D2AFF1FFC31588DFC60" blockId="10.[151,651,901,960]" box="[151,651,901,927]" pageId="10" pageNumber="111">
<heading id="9A5E81C9FFB76D2AFF1FFC31588DFC60" bold="true" box="[151,651,901,927]" fontSize="11" level="1" pageId="10" pageNumber="111" reason="1">
<taxonomicName id="06A94D26FFB76D2AFF1FFC31588DFC60" ID-CoL="6JBYD" authority="Marshall, 1991" authorityName="Marshall" authorityYear="1991" box="[151,651,901,927]" class="Gastropoda" family="Seguenziidae" genus="Fluxinella" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="membranacea">
<emphasis id="F3DDEAB7FFB76D2AFF1FFC31588DFC60" bold="true" box="[151,651,901,927]" pageId="10" pageNumber="111">
<emphasis id="F3DDEAB7FFB76D2AFF1FFC315BCEFC60" bold="true" box="[151,456,901,927]" italics="true" pageId="10" pageNumber="111">Fluxinella membranacea</emphasis>
<bibRefCitation id="A5384B54FFB76D2AFE47FC31588DFC60" author="Marshall" box="[463,651,901,927]" pageId="10" pageNumber="111" refString="Marshall, B. A. (1991) Mollusca Gastropoda: Seguenziidae from New Caledonia and the Loyalty Islands. Memoires du Museum Nationale d'Histoire Naturelle Serie A Zoologie, 150, 41 - 109." type="journal article" year="1991">
<collectingCountry id="B9BE7635FFB76D2AFE47FC315845FC60" box="[463,579,901,927]" name="Marshall Islands" pageId="10" pageNumber="111">Marshall</collectingCountry>
, 1991
</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C11636A5FFB76D2AFF1FFC135B4CFC3F" blockId="10.[151,651,901,960]" box="[151,330,935,960]" pageId="10" pageNumber="111">
(
<figureCitation id="59922A20FFB76D2AFF16FC135B44FC3F" box="[158,322,935,960]" captionStart="FIGURES 13 – 17" captionStartId="11.[151,264,1855,1877]" captionTargetBox="[235,1343,195,1825]" captionTargetId="figure@11.[233,1346,193,1828]" captionTargetPageId="11" captionText="FIGURES 13 – 17. Seguenziidae. 13. Halystina conoidea nov. spec. RGM 608.229 (holotype). Locality AndaDeVos. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 14. Halystina conoidea nov. spec. RGM 608.230 (paratype). Locality Anda 6. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 15. Halystina conoidea nov. spec. RGM 608.231 (paratype). Locality AndaDeVos. SEM detail of protoconch. 16. Fluxinella membranacea Marshall, 1991. RGM 608.239. Locality Anda 1. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 17. Fluxinella membranacea Marshall, 1991. RGM 608.240. Locality Anda 1. (a) apical view, (b) SEM detail of surface sculpture." httpUri="https://zenodo.org/record/250964/files/figure.png" pageId="10" pageNumber="111">Figures 16–17</figureCitation>
)
</paragraph>
<paragraph id="C11636A5FFB76D2AFF1FFC445940FBDB" blockId="10.[151,990,1008,1091]" pageId="10" pageNumber="111">
1991
<taxonomicName id="06A94D26FFB76D2AFF7DFC445BF4FBF9" box="[245,498,1008,1030]" class="Gastropoda" family="Seguenziidae" genus="Fluxinella" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="membranacea" status="sp. nov.">
<emphasis id="F3DDEAB7FFB76D2AFF7DFC445BF4FBF9" box="[245,498,1008,1030]" italics="true" pageId="10" pageNumber="111">Fluxinella membranacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE57CCFFB76D2AFE71FC455840FBF9" box="[505,582,1008,1030]" pageId="10" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
—
<bibRefCitation id="A5384B54FFB76D2AFDE8FC4458FBFBF9" author="Marshall" box="[608,765,1008,1030]" pageId="10" pageNumber="111" refString="Marshall, B. A. (1991) Mollusca Gastropoda: Seguenziidae from New Caledonia and the Loyalty Islands. Memoires du Museum Nationale d'Histoire Naturelle Serie A Zoologie, 150, 41 - 109." type="journal article" year="1991">
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, 1991
</bibRefCitation>
: p. 68, figs 91, 93, 94. 2006
<taxonomicName id="06A94D26FFB76D2AFF7DFBBA5BF4FBDB" box="[245,498,1038,1060]" class="Gastropoda" family="Seguenziidae" genus="Fluxinella" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="membranacea">
<emphasis id="F3DDEAB7FFB76D2AFF7DFBBA5BF4FBDB" box="[245,498,1038,1060]" italics="true" pageId="10" pageNumber="111">Fluxinella membranacea</emphasis>
</taxonomicName>
—Poppe
<emphasis id="F3DDEAB7FFB76D2AFDD9FBA45882FBDB" box="[593,644,1038,1061]" italics="true" pageId="10" pageNumber="111">et al.</emphasis>
: p. 23, pl. 2, fig. 4.
</paragraph>
<paragraph id="C11636A5FFB76D2AFF1FFB99598BFBBC" blockId="10.[151,990,1008,1091]" box="[151,909,1069,1091]" pageId="10" pageNumber="111">
2008a
<taxonomicName id="06A94D26FFB76D2AFF7DFB995BF4FBBC" box="[245,498,1069,1091]" class="Gastropoda" family="Seguenziidae" genus="Fluxinella" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="membranacea">
<emphasis id="F3DDEAB7FFB76D2AFF7DFB995BF4FBBC" box="[245,498,1069,1091]" italics="true" pageId="10" pageNumber="111">Fluxinella membranacea</emphasis>
</taxonomicName>
—Poppe &amp; Tagaro: p. 160, pl. 25, fig. 4.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB76D2AFF1FFBDB5F9AFB0A" pageId="10" pageNumber="111" type="materials_examined">
<paragraph id="C11636A5FFB76D2AFF1FFBDB5862FB53" blockId="10.[151,1436,1135,1485]" pageId="10" pageNumber="111">
<emphasis id="F3DDEAB7FFB76D2AFF1FFBDB5B00FB77" bold="true" box="[151,262,1135,1160]" pageId="10" pageNumber="111">Material.</emphasis>
Anda 1 (28); Anda 2 (41); Anda 3 (26); Anda 4 (37); Anda 5 (2); Anda 6 (5); AndaDeVos (15); AndaClif1 (1); AndaClif3 (2); Roxas (3).
</paragraph>
<paragraph id="C11636A5FFB76D2AFF4FFB035F9AFB0A" blockId="10.[151,1436,1135,1485]" pageId="10" pageNumber="111">
<emphasis id="F3DDEAB7FFB76D2AFF4FFB035B9CFB2F" bold="true" box="[199,410,1207,1232]" pageId="10" pageNumber="111">Characterization.</emphasis>
Shell very low domed, H
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,
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W
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mm, disk-shaped, smooth and shiny; P small, smooth, 1.1 whorls, DN
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; T growth lines very fine, sigmoid; prominent keel on periphery; umbilicus wide.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB76D2AFF4FFB4B5F8AFAC2" pageId="10" pageNumber="111" type="distribution">
<paragraph id="C11636A5FFB76D2AFF4FFB4B5F8AFAC2" blockId="10.[151,1436,1135,1485]" pageId="10" pageNumber="111">
<emphasis id="F3DDEAB7FFB76D2AFF4FFB4B5B66FAE7" bold="true" box="[199,352,1279,1304]" pageId="10" pageNumber="111">Distribution.</emphasis>
Shells of
<taxonomicName id="06A94D26FFB76D2AFE5CFB4B58E9FAE7" box="[468,751,1279,1304]" class="Gastropoda" family="Seguenziidae" genus="Fluxinella" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="membranacea">
<emphasis id="F3DDEAB7FFB76D2AFE5CFB4B58E9FAE7" box="[468,751,1279,1304]" italics="true" pageId="10" pageNumber="111">Fluxinella membranacea</emphasis>
</taxonomicName>
have been reported from off southern
<collectingCountry id="B9BE7635FFB76D2AFB2EFAB45F51FAE7" box="[1190,1367,1279,1304]" name="New Caledonia" pageId="10" pageNumber="111">New Caledonia</collectingCountry>
,
<quantity id="06519B40FFB76D2AFAECFB4B5AD8FAC3" metricMagnitude="2" metricUnit="m" metricValue="3.45" metricValueMax="4.4" metricValueMin="2.5" pageId="10" pageNumber="111" unit="m" value="345.0" valueMax="440.0" valueMin="250.0">250– 440 m</quantity>
depth (
<bibRefCitation id="A5384B54FFB76D2AFEB9FA905BD1FAC3" author="Marshall" box="[305,471,1316,1341]" pageId="10" pageNumber="111" refString="Marshall, B. A. (1991) Mollusca Gastropoda: Seguenziidae from New Caledonia and the Loyalty Islands. Memoires du Museum Nationale d'Histoire Naturelle Serie A Zoologie, 150, 41 - 109." type="journal article" year="1991">
<collectingCountry id="B9BE7635FFB76D2AFEB9FA905B90FAC2" box="[305,406,1316,1341]" name="Marshall Islands" pageId="10" pageNumber="111">Marshall</collectingCountry>
1991
</bibRefCitation>
) and from Balicasag
<collectingCountry id="B9BE7635FFB76D2AFD4FFA905908FAC2" box="[711,782,1316,1341]" name="Iceland" pageId="10" pageNumber="111">Island</collectingCountry>
,
<collectingCountry id="B9BE7635FFB76D2AFC91FA90599CFAC2" box="[793,922,1316,1341]" name="Philippines" pageId="10" pageNumber="111">Philippines</collectingCountry>
, from
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depth (
<bibRefCitation id="A5384B54FFB76D2AFB3DFA905F86FAC2" author="Poppe" box="[1205,1408,1316,1341]" pageId="10" pageNumber="111" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter" year="2006">
Poppe
<emphasis id="F3DDEAB7FFB76D2AFA8CFA915F3AFAC2" box="[1284,1340,1316,1341]" italics="true" pageId="10" pageNumber="111">et al.</emphasis>
2006
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB76D2AFF4FFAF35EE1FA32" pageId="10" pageNumber="111" type="discussion">
<paragraph id="C11636A5FFB76D2AFF4FFAF35EE1FA32" blockId="10.[151,1436,1135,1485]" pageId="10" pageNumber="111">
<emphasis id="F3DDEAB7FFB76D2AFF4FFAF35B3DFA9F" bold="true" box="[199,315,1351,1376]" pageId="10" pageNumber="111">Remarks.</emphasis>
The specimen shown by
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is similar to the shells in the studied material. The specimen shown by
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Poppe
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(2006
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, pl. 2, fig. 4) has a more pronounced folded columellar lip, a more pronounced shoulder and median keel than the fossil material as well as the material shown by
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. More material is required to investigate whether these differences represent intraspecific variation.
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(
<figureCitation id="59922A20FFB76D2AFF16F9525B47F900" box="[158,321,1766,1791]" captionStart="FIGURES 18 – 25" captionStartId="12.[151,264,1787,1809]" captionTargetBox="[262,1331,195,1768]" captionTargetId="figure@12.[260,1334,193,1771]" captionTargetPageId="12" captionText="FIGURES 18 – 25. Seguenziidae. 18. Calliobasis lapulapui Poppe, Tagaro &amp; Dekker, 2006. RGM 608.251. Locality Anda 1. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 19. Calliobasis lapulapui Poppe, Tagaro &amp; Dekker, 2006. RGM 608.252. Locality Anda 3. (a) SEM, side view, (b) SEM detail of protoconch. 20. Thelyssina? spec. 1. RGM 608.263. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 21. Thelyssina? spec. 1. RGM 608.264. Locality Anda 2. (a) SEM detail of protoconch, (b) SEM, side view. 22. Thelyssina? spec. 1. RGM 608.265. Locality Anda 2. SEM, basal view. 23. Thelyssina? spec. 2. RGM 608.268. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 24. Thelyssina? spec. 2. RGM 608.269. Locality AndaDeVos. SEM detail of protoconch. 25. Thelyssina? spec. 2. RGM 608.270 Locality AndaDeVos. SEM, basal view." httpUri="https://zenodo.org/record/250965/files/figure.png" pageId="10" pageNumber="111">Figures 18–19</figureCitation>
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<paragraph id="C11636A5FFB76D2AFF1FF89A596FF899" blockId="10.[151,909,1838,1894]" pageId="10" pageNumber="111">
2006
<taxonomicName id="06A94D26FFB76D2AFF7DF89A5BC9F8BB" box="[245,463,1838,1860]" class="Gastropoda" family="Seguenziidae" genus="Calliobasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="lapulapui">
<emphasis id="F3DDEAB7FFB76D2AFF7DF89A5BC9F8BB" box="[245,463,1838,1860]" italics="true" pageId="10" pageNumber="111">Calliobasis lapulapui</emphasis>
</taxonomicName>
nov. spec.—Poppe
<emphasis id="F3DDEAB7FFB76D2AFD11F88458CAF8BB" box="[665,716,1838,1861]" italics="true" pageId="10" pageNumber="111">et al.</emphasis>
: p. 21, pl. 1, fig. 2. 2008a
<taxonomicName id="06A94D26FFB76D2AFF7DF8E45BC9F899" box="[245,463,1872,1894]" class="Gastropoda" family="Seguenziidae" genus="Calliobasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="lapulapui">
<emphasis id="F3DDEAB7FFB76D2AFF7DF8E45BC9F899" box="[245,463,1872,1894]" italics="true" pageId="10" pageNumber="111">Calliobasis lapulapui</emphasis>
</taxonomicName>
—Poppe &amp; Tagaro: p. 160, pl. 25, fig. 1.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB76D2DFF1FF8265B13FEE2" lastPageId="13" lastPageNumber="114" pageId="10" pageNumber="111" type="materials_examined">
<paragraph id="C11636A5FFB76D2AFF1FF8265B8CF830" blockId="10.[151,1436,1938,1999]" pageId="10" pageNumber="111">
<emphasis id="F3DDEAB7FFB76D2AFF1FF8265B00F854" bold="true" box="[151,262,1938,1963]" pageId="10" pageNumber="111">Material.</emphasis>
Anda 1 (72); Anda 2 (81); Anda 3 (44); Anda 4 (83); Anda 5 (1); Anda 6 (2); AndaDeVos (3); Roxas (1); Tiep 3 (3); Tiep 5 (5).
</paragraph>
<caption id="95D6662DFFB66D2BFF1FF88B5F2EF811" httpUri="https://zenodo.org/record/250964/files/figure.png" pageId="11" pageNumber="112" targetBox="[235,1343,195,1825]" targetPageId="11">
<paragraph id="C11636A5FFB66D2BFF1FF88B5F2EF811" blockId="11.[151,1436,1855,2030]" pageId="11" pageNumber="112">
<emphasis id="F3DDEAB7FFB66D2BFF1FF88B5B53F8AA" bold="true" box="[151,341,1855,1877]" pageId="11" pageNumber="112">FIGURES 13–17.</emphasis>
<taxonomicName id="06A94D26FFB66D2BFED5F88B5BE1F8AA" box="[349,487,1855,1877]" class="Gastropoda" family="Seguenziidae" kingdom="Animalia" pageId="11" pageNumber="112" phylum="Mollusca" rank="family">Seguenziidae</taxonomicName>
.
<emphasis id="F3DDEAB7FFB66D2BFE7AF88B5814F8AA" bold="true" box="[498,530,1855,1877]" pageId="11" pageNumber="112">13.</emphasis>
<taxonomicName id="06A94D26FFB66D2BFD92F88B58E4F8AA" box="[538,738,1855,1877]" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="F3DDEAB7FFB66D2BFD92F88B58E4F8AA" box="[538,738,1855,1877]" italics="true" pageId="11" pageNumber="112">Halystina conoidea</emphasis>
</taxonomicName>
nov. spec. RGM 608.229 (holotype). Locality AndaDeVos. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view.
<emphasis id="F3DDEAB7FFB66D2BFC1CF8E959B5F88C" bold="true" box="[916,947,1885,1907]" pageId="11" pageNumber="112">14.</emphasis>
<taxonomicName id="06A94D26FFB66D2BFC36F8E95E8EF88C" box="[958,1160,1885,1907]" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="F3DDEAB7FFB66D2BFC36F8E95E8EF88C" box="[958,1160,1885,1907]" italics="true" pageId="11" pageNumber="112">Halystina conoidea</emphasis>
</taxonomicName>
nov. spec. RGM 608.230 (paratype). Locality Anda6. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view.
<emphasis id="F3DDEAB7FFB66D2BFA98F8C85F29F86D" bold="true" box="[1296,1327,1916,1938]" pageId="11" pageNumber="112">15.</emphasis>
<taxonomicName id="06A94D26FFB66D2BFAB1F8C85AF2F84E" class="Gastropoda" family="Seguenziidae" genus="Halystina" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="F3DDEAB7FFB66D2BFAB1F8C85AF2F84E" italics="true" pageId="11" pageNumber="112">Halystina conoidea</emphasis>
</taxonomicName>
nov. spec. RGM 608.231 (paratype). Locality AndaDeVos. SEM detail of protoconch.
<emphasis id="F3DDEAB7FFB66D2BFBFDF82F5E93F84E" bold="true" box="[1141,1173,1947,1969]" pageId="11" pageNumber="112">16.</emphasis>
<taxonomicName id="06A94D26FFB66D2BFB15F82F5B35F830" authority="Marshall, 1991" authorityName="Marshall" authorityYear="1991" class="Gastropoda" family="Seguenziidae" genus="Fluxinella" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="membranacea">
<emphasis id="F3DDEAB7FFB66D2BFB15F82F5F9DF84E" box="[1181,1435,1947,1969]" italics="true" pageId="11" pageNumber="112">Fluxinella membranacea</emphasis>
Marshall, 1991
</taxonomicName>
. RGM 608.239. Locality Anda1. (a) rear view, (b) apertural view, (c) basal view, (d) apical view.
<emphasis id="F3DDEAB7FFB66D2BFA86F80D5F21F830" bold="true" box="[1294,1319,1977,1999]" pageId="11" pageNumber="112">17</emphasis>
.
<taxonomicName id="06A94D26FFB66D2BFABCF80D5BCFF811" authority="Marshall, 1991" authorityName="Marshall" authorityYear="1991" class="Gastropoda" family="Seguenziidae" genus="Fluxinella" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="membranacea">
<emphasis id="F3DDEAB7FFB66D2BFABCF80D5B20F811" italics="true" pageId="11" pageNumber="112">Fluxinella membranacea</emphasis>
Marshall, 1991
</taxonomicName>
. RGM 608.240. Locality Anda1. (a) apical view, (b) SEM detail of surface sculpture.
</paragraph>
</caption>
<caption id="95D6662DFFB16D2CFF1FF94F5988F818" httpUri="https://zenodo.org/record/250965/files/figure.png" pageId="12" pageNumber="113" targetBox="[262,1331,195,1768]" targetPageId="12">
<paragraph id="C11636A5FFB16D2CFF1FF94F5988F818" blockId="12.[151,1436,1787,2023]" pageId="12" pageNumber="113">
<emphasis id="F3DDEAB7FFB16D2CFF1FF94F5B55F8EE" bold="true" box="[151,339,1787,1809]" pageId="12" pageNumber="113">FIGURES 18–25.</emphasis>
<taxonomicName id="06A94D26FFB16D2CFED1F94F5BE5F8EE" box="[345,483,1787,1809]" class="Gastropoda" family="Seguenziidae" kingdom="Animalia" pageId="12" pageNumber="113" phylum="Mollusca" rank="family">Seguenziidae</taxonomicName>
.
<emphasis id="F3DDEAB7FFB16D2CFE64F94F580DF8EE" bold="true" box="[492,523,1787,1809]" pageId="12" pageNumber="113">18.</emphasis>
<taxonomicName id="06A94D26FFB16D2CFD9AF94F5E29F8EE" authority="Poppe, Tagaro &amp; Dekker, 2006" authorityName="Poppe, Tagaro &amp; Dekker" authorityYear="2006" box="[530,1071,1787,1809]" class="Gastropoda" family="Seguenziidae" genus="Calliobasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="12" pageNumber="113" phylum="Mollusca" rank="species" species="lapulapui">
<emphasis id="F3DDEAB7FFB16D2CFD9AF94F58EAF8EE" box="[530,748,1787,1809]" italics="true" pageId="12" pageNumber="113">Calliobasis lapulapui</emphasis>
Poppe, Tagaro &amp; Dekker, 2006
</taxonomicName>
. RGM 608.251. Locality Anda1. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view.
<emphasis id="F3DDEAB7FFB16D2CFC18F8AD59B6F8D0" bold="true" box="[912,944,1817,1839]" pageId="12" pageNumber="113">19.</emphasis>
<taxonomicName id="06A94D26FFB16D2CFC3FF8AD5ACBF8B1" authority="Poppe, Tagaro &amp; Dekker, 2006" authorityName="Poppe, Tagaro &amp; Dekker" authorityYear="2006" class="Gastropoda" family="Seguenziidae" genus="Calliobasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="12" pageNumber="113" phylum="Mollusca" rank="species" species="lapulapui">
<emphasis id="F3DDEAB7FFB16D2CFC3FF8AD5E94F8D0" box="[951,1170,1817,1839]" italics="true" pageId="12" pageNumber="113">Calliobasis lapulapui</emphasis>
Poppe, Tagaro &amp; Dekker, 2006
</taxonomicName>
. RGM 608.252. Locality Anda3. (a) SEM, side view, (b) SEM detail of protoconch.
<emphasis id="F3DDEAB7FFB16D2CFBEBF88C5E85F8B1" bold="true" box="[1123,1155,1848,1870]" pageId="12" pageNumber="113">20.</emphasis>
<emphasis id="F3DDEAB7FFB16D2CFB05F88C5F00F8B1" box="[1165,1286,1848,1870]" italics="true" pageId="12" pageNumber="113">
<taxonomicName id="06A94D26FFB16D2CFB05F88C5EFAF8B1" box="[1165,1276,1848,1870]" class="Gastropoda" family="Seguenziidae" genus="Thelyssina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="12" pageNumber="113" phylum="Mollusca" rank="genus">Thelyssina</taxonomicName>
?
</emphasis>
spec.1. RGM 608.263. Locality Anda2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view.
<emphasis id="F3DDEAB7FFB16D2CFBECF8E35E82F892" bold="true" box="[1124,1156,1879,1901]" pageId="12" pageNumber="113">21.</emphasis>
<emphasis id="F3DDEAB7FFB16D2CFB06F8E35F01F892" box="[1166,1287,1879,1901]" italics="true" pageId="12" pageNumber="113">
<taxonomicName id="06A94D26FFB16D2CFB06F8E35EFBF892" box="[1166,1277,1879,1901]" class="Gastropoda" family="Seguenziidae" genus="Thelyssina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="12" pageNumber="113" phylum="Mollusca" rank="genus">Thelyssina</taxonomicName>
?
</emphasis>
spec.1. RGM 608.264. Locality Anda2. (a) SEM detail of protoconch, (b) SEM, side view.
<emphasis id="F3DDEAB7FFB16D2CFC3AF8C159D4F874" bold="true" box="[946,978,1909,1931]" pageId="12" pageNumber="113">22.</emphasis>
<emphasis id="F3DDEAB7FFB16D2CFC51F8C15E57F874" box="[985,1105,1909,1931]" italics="true" pageId="12" pageNumber="113">
<taxonomicName id="06A94D26FFB16D2CFC51F8C15E4EF874" box="[985,1096,1909,1931]" class="Gastropoda" family="Seguenziidae" genus="Thelyssina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="12" pageNumber="113" phylum="Mollusca" rank="genus">Thelyssina</taxonomicName>
?
</emphasis>
spec.1. RGM 608.265. Locality Anda2. SEM, basal view.
<emphasis id="F3DDEAB7FFB16D2CFE2EF8205BC0F855" bold="true" box="[422,454,1940,1962]" pageId="12" pageNumber="113">23.</emphasis>
<emphasis id="F3DDEAB7FFB16D2CFE46F8205841F855" box="[462,583,1940,1962]" italics="true" pageId="12" pageNumber="113">
<taxonomicName id="06A94D26FFB16D2CFE46F820583BF855" box="[462,573,1940,1962]" class="Gastropoda" family="Seguenziidae" genus="Thelyssina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="12" pageNumber="113" phylum="Mollusca" rank="genus">Thelyssina</taxonomicName>
?
</emphasis>
spec.2. RGM 608.268. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view.
<emphasis id="F3DDEAB7FFB16D2CFE4EF8075BE0F836" bold="true" box="[454,486,1971,1993]" pageId="12" pageNumber="113">24.</emphasis>
<emphasis id="F3DDEAB7FFB16D2CFE79F807586FF836" box="[497,617,1971,1993]" italics="true" pageId="12" pageNumber="113">
<taxonomicName id="06A94D26FFB16D2CFE79F8075866F836" box="[497,608,1971,1993]" class="Gastropoda" family="Seguenziidae" genus="Thelyssina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="12" pageNumber="113" phylum="Mollusca" rank="genus">Thelyssina</taxonomicName>
?
</emphasis>
spec.2. RGM 608.269. Locality AndaDeVos. SEM detail of protoconch.
<emphasis id="F3DDEAB7FFB16D2CFAF4F8075F9AF836" bold="true" box="[1404,1436,1971,1993]" pageId="12" pageNumber="113">25.</emphasis>
<emphasis id="F3DDEAB7FFB16D2CFF1FF8655B09F818" box="[151,271,2001,2023]" italics="true" pageId="12" pageNumber="113">
<taxonomicName id="06A94D26FFB16D2CFF1FF8655B00F818" box="[151,262,2001,2023]" class="Gastropoda" family="Seguenziidae" genus="Thelyssina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeogastropoda" pageId="12" pageNumber="113" phylum="Mollusca" rank="genus">Thelyssina</taxonomicName>
?
</emphasis>
spec.2. RGM 608.270 Locality AndaDeVos. SEM, basal view.
</paragraph>
</caption>
<paragraph id="C11636A5FFB06D2DFF4FFF235B13FEE2" blockId="13.[151,1436,151,464]" pageId="13" pageNumber="114">
<emphasis id="F3DDEAB7FFB06D2DFF4FFF235B9CFF4F" bold="true" box="[199,410,151,176]" pageId="13" pageNumber="114">Characterization.</emphasis>
Conical shell with regularly-spaced, well-defined erect sigmoid axial ribs, H
<quantity id="06519B40FFB06D2DFA91FF235F75FF50" box="[1305,1395,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="114" unit="mm" value="2.5">2.5 mm</quantity>
,
<geoCoordinate id="A49D5062FFB06D2DFA09FF2C5ABCFF2A" direction="west" orientation="longitude" pageId="13" pageNumber="114" precision="5555" value="-2.6">
W
<quantity id="06519B40FFB06D2DFF1FFF085ABCFF2A" box="[151,186,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="114" unit="mm" value="2.6">2.6</quantity>
</geoCoordinate>
mm; P globular, 1.1 whorls, DN
<quantity id="06519B40FFB06D2DFDB3FF0858A2FF2B" box="[571,676,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="114" unit="mm" value="0.1">0.10 mm</quantity>
; one or two spiral ribs on lower T whorls; body whorl with strong keel; base with nine spiral ribs; plane of aperture tilted downwards; outer lip with strong upper and lower retractions.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB06D2DFF4FFE935991FE9A" pageId="13" pageNumber="114" type="distribution">
<paragraph id="C11636A5FFB06D2DFF4FFE935991FE9A" blockId="13.[151,1436,151,464]" pageId="13" pageNumber="114">
<emphasis id="F3DDEAB7FFB06D2DFF4FFE935B66FEBF" bold="true" box="[199,352,295,320]" pageId="13" pageNumber="114">Distribution.</emphasis>
Apart from the studied material,
<taxonomicName id="06A94D26FFB06D2DFD50FE9359CCFEBF" box="[728,970,295,320]" class="Gastropoda" family="Seguenziidae" genus="Calliobasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="lapulapui">
<emphasis id="F3DDEAB7FFB06D2DFD50FE9359CCFEBF" box="[728,970,295,320]" italics="true" pageId="13" pageNumber="114">Calliobasis lapulapui</emphasis>
</taxonomicName>
has been reported from the Dipolog Bay, Mindanao
<collectingCountry id="B9BE7635FFB06D2DFE87FEF85B50FE9A" box="[271,342,332,357]" name="Iceland" pageId="13" pageNumber="114">Island</collectingCountry>
,
<collectingCountry id="B9BE7635FFB06D2DFEEAFEF85BE5FE9A" box="[354,483,332,357]" name="Philippines" pageId="13" pageNumber="114">Philippines</collectingCountry>
,
<quantity id="06519B40FFB06D2DFE66FEF8586BFE9B" box="[494,621,332,357]" metricMagnitude="2" metricUnit="m" metricValue="1.735" metricValueMax="1.75" metricValueMin="1.72" pageId="13" pageNumber="114" unit="m" value="173.5" valueMax="175.0" valueMin="172.0">172–175 m</quantity>
depth (
<bibRefCitation id="A5384B54FFB06D2DFD48FEF8598DFE9A" author="Poppe" box="[704,907,332,357]" pageId="13" pageNumber="114" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter" year="2006">
Poppe
<emphasis id="F3DDEAB7FFB06D2DFC87FEF95941FE9A" box="[783,839,332,357]" italics="true" pageId="13" pageNumber="114">et al.</emphasis>
2006
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB06D2DFF4FFEDB5866FDC3" pageId="13" pageNumber="114" type="discussion">
<paragraph id="C11636A5FFB06D2DFF4FFEDB59F7FE2F" blockId="13.[151,1436,151,464]" pageId="13" pageNumber="114">
<emphasis id="F3DDEAB7FFB06D2DFF4FFEDB5B3DFE77" bold="true" box="[199,315,367,392]" pageId="13" pageNumber="114">Remarks.</emphasis>
The specimen shown in
<bibRefCitation id="A5384B54FFB06D2DFDD8FEC45920FE77" box="[592,806,367,392]" pageId="13" pageNumber="114" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter">
Poppe
<emphasis id="F3DDEAB7FFB06D2DFD15FEC558D3FE77" box="[669,725,367,392]" italics="true" pageId="13" pageNumber="114">et al.</emphasis>
(2006)
</bibRefCitation>
does not have the elaborate outer lip organisation, but is likely not fully grown. It is smaller than the fossil material studied here (H
<quantity id="06519B40FFB06D2DFC6DFE205E3AFE53" box="[997,1084,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="114" unit="mm" value="2.2">2.2 mm</quantity>
versus H
<quantity id="06519B40FFB06D2DFB25FE205F10FE53" box="[1197,1302,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.53" pageId="13" pageNumber="114" unit="mm" value="2.53">2.53 mm</quantity>
). The shape of the shell and the ornamentation are the same as in the fossils studied here.
</paragraph>
<paragraph id="C11636A5FFB06D2DFF1FFD905866FDC3" blockId="13.[151,608,548,573]" box="[151,608,548,573]" pageId="13" pageNumber="114">
<emphasis id="F3DDEAB7FFB06D2DFF1FFD905B14FDC2" bold="true" box="[151,274,548,573]" pageId="13" pageNumber="114">Subfamily</emphasis>
<taxonomicName id="06A94D26FFB06D2DFE91FD905866FDC3" authority="Marshall, 1991" authorityName="Marshall" authorityYear="1991" box="[281,608,548,573]" class="Gastropoda" family="Seguenziidae" kingdom="Animalia" pageId="13" pageNumber="114" phylum="Mollusca" rank="subFamily" subFamily="Asthelysinae">
<emphasis id="F3DDEAB7FFB06D2DFE91FD905BB6FDC2" bold="true" box="[281,432,548,573]" pageId="13" pageNumber="114">Asthelysinae</emphasis>
<bibRefCitation id="A5384B54FFB06D2DFE3EFD905866FDC3" author="Marshall" box="[438,608,548,573]" pageId="13" pageNumber="114" refString="Marshall, B. A. (1991) Mollusca Gastropoda: Seguenziidae from New Caledonia and the Loyalty Islands. Memoires du Museum Nationale d'Histoire Naturelle Serie A Zoologie, 150, 41 - 109." type="journal article" year="1991">
<collectingCountry id="B9BE7635FFB06D2DFE3EFD90581AFDC2" box="[438,540,548,573]" name="Marshall Islands" pageId="13" pageNumber="114">Marshall</collectingCountry>
, 1991
</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="B1EB690457149C6FF33797B1C8D6A8C4" ID-CLB-Dataset="46615" ID-DOI="10.11646/zootaxa.3755.2.1" ID-GBIF-Dataset="e02bbd39-b7ee-4ddf-8109-3407dd6d2ea2" ID-ISSN="1175-5326" ID-Zenodo-Dep="250959" ID-ZooBank="1872ECAB-3C5C-4D76-93A0-A8626F75B96E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460372737068" checkinUser="plazi" docAuthor="Helwerda, Renate Ariane, Wesselingh, Frank Pieter &amp; Williams, Suzanne T." docDate="2014" docId="490087B3FFB86D25FF1FFBA65F62F8DE" docLanguage="en" docName="zt03755p135.pdf" docOrigin="Zootaxa 3755 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Homalopoma tosaense Habe 1953" docType="treatment" docVersion="10" lastPageNumber="106" masterDocId="B539FFCBFFBD6D20FF88FFB45A06FFFF" masterDocTitle="On some Vetigastropoda (Mollusca, Gastropoda) from the Plio-Pleistocene of the Philippines with descriptions of three new species" masterLastPageNumber="135" masterPageNumber="101" pageNumber="106" updateTime="1698295924955" updateUser="ExternalLinkService">
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<mods:title id="E57E6B02E6553352C914F93CEFD0A062">On some Vetigastropoda (Mollusca, Gastropoda) from the Plio-Pleistocene of the Philippines with descriptions of three new species</mods:title>
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<mods:namePart id="07A5BD407BA220D29EA71E83B762B76E">Helwerda, Renate Ariane</mods:namePart>
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<treatment id="490087B3FFB86D25FF1FFBA65F62F8DE" ID-DOI="http://doi.org/10.5281/zenodo.5671247" ID-GBIF-Taxon="119412409" ID-Zenodo-Dep="5671247" LSID="urn:lsid:plazi:treatment:490087B3FFB86D25FF1FFBA65F62F8DE" httpUri="http://treatment.plazi.org/id/490087B3FFB86D25FF1FFBA65F62F8DE" lastPageNumber="106" pageId="5" pageNumber="106">
<subSubSection id="89B3652EFFB86D25FF1FFBA6597AFB6D" pageId="5" pageNumber="106" type="nomenclature">
<paragraph id="C11636A5FFB86D25FF1FFBA6583BFBD3" blockId="5.[151,573,1042,1101]" box="[151,573,1042,1068]" pageId="5" pageNumber="106">
<heading id="9A5E81C9FFB86D25FF1FFBA6583BFBD3" bold="true" box="[151,573,1042,1068]" fontSize="11" level="1" pageId="5" pageNumber="106" reason="1">
<taxonomicName id="06A94D26FFB86D25FF1FFBA6583BFBD3" ID-CoL="3MDTB" authority="Habe, 1953" authorityName="Habe" authorityYear="1953" box="[151,573,1042,1068]" class="Gastropoda" family="Colloniidae" genus="Homalopoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="species" species="tosaense">
<emphasis id="F3DDEAB7FFB86D25FF1FFBA6583BFBD3" bold="true" box="[151,573,1042,1068]" pageId="5" pageNumber="106">
<emphasis id="F3DDEAB7FFB86D25FF1FFBA65BAEFBD3" bold="true" box="[151,424,1042,1068]" italics="true" pageId="5" pageNumber="106">Homalopoma tosaense</emphasis>
<bibRefCitation id="A5384B54FFB86D25FE38FBA6583BFBD3" author="Habe" box="[432,573,1042,1068]" pageId="5" pageNumber="106" refString="Habe, T. (1953) Descriptions of twelve new Japanese shells. Venus, 17, 130 - 144." type="journal article" year="1953">Habe, 1953</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C11636A5FFB86D25FF1FFB805B28FBB2" blockId="5.[151,573,1042,1101]" box="[151,302,1076,1101]" pageId="5" pageNumber="106">
(
<figureCitation id="59922A20FFB86D25FF16FB805B23FBB2" box="[158,293,1076,1101]" captionStart="FIGURES 3 – 5" captionStartId="6.[151,264,1797,1819]" captionTargetBox="[262,1331,195,1769]" captionTargetId="figure@6.[260,1334,193,1771]" captionTargetPageId="6" captionText="FIGURES 3 – 5. Colloniidae. 3. Homalopoma tosaense Habe, 1953. RGM 608.201. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. For this and remaining figures all material derives from the Piacenzian or Gelasian Santa Cruz Formation of Cabarruyan Island and nearby Luzon island, Philippines. Locality data in Table 1. Scale bars shells = 1 mm, scale bars SEM = 100 µm. 4. Homalopoma tosaense Habe, 1953. RGM 608.202. Locality Anda 2. (a) rear view, (b) apertural view. 5. Homalopoma tosaense Habe, 1953. RGM 608.203. Locality AndaDeVos, SEM detail of protoconch. FIGURES 6 – 7. Seguenziidae. 6. Seguenzia cf. keikoae Poppe, Tagaro &amp; Dekker, 2006. RGM 608.207. Locality Roxas. (a) rear view, (b) side view, (c) apertural view, (d) basal view, (e) apical view, (f) detail of protoconch, scale bar 100 µm. 7. Seguenzia cf. keikoae Poppe, Tagaro &amp; Dekker, 2006. RGM 608.208. Locality Anda 3. (a) rear view, (b) apertural view." httpUri="https://zenodo.org/record/250962/files/figure.png" pageId="5" pageNumber="106">Figures 3–5</figureCitation>
)
</paragraph>
<paragraph id="C11636A5FFB86D25FF1FFBC8597AFB6D" blockId="5.[151,1344,1148,1324]" box="[151,892,1148,1170]" pageId="5" pageNumber="106">
1982
<taxonomicName id="06A94D26FFB86D25FF6DFBC85BB5FB6D" box="[229,435,1148,1170]" class="Gastropoda" family="Colloniidae" genus="Homalopoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="species" species="druidi">
<emphasis id="F3DDEAB7FFB86D25FF6DFBC85BB5FB6D" box="[229,435,1148,1170]" italics="true" pageId="5" pageNumber="106">Homalopoma druidi</emphasis>
</taxonomicName>
nov. spec.—Ladd: p. 27–28, pl. 36, figs 5–8.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB86D25FF1FFB2F5F46FAD3" pageId="5" pageNumber="106" type="reference_group">
<paragraph id="C11636A5FFB86D25FF1FFB2F592CFB4E" blockId="5.[151,1344,1148,1324]" box="[151,810,1179,1201]" pageId="5" pageNumber="106">
1999
<taxonomicName id="06A94D26FFB86D25FF6DFB2F585DFB4E" authority="Habe, 1953" authorityName="Habe" authorityYear="1953" box="[229,603,1179,1201]" class="Gastropoda" family="Colloniidae" genus="Homalopoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="species" species="tosaensis">
<emphasis id="F3DDEAB7FFB86D25FF6DFB2F5BD4FB4E" box="[229,466,1179,1201]" italics="true" pageId="5" pageNumber="106">Homalopoma tosaensis</emphasis>
(
<bibRefCitation id="A5384B54FFB86D25FE57FB2F5852FB4E" author="Habe" box="[479,596,1179,1201]" pageId="5" pageNumber="106" refString="Habe, T. (1953) Descriptions of twelve new Japanese shells. Venus, 17, 130 - 144." type="journal article" year="1953">Habe, 1953</bibRefCitation>
)
</taxonomicName>
—Higo
<emphasis id="F3DDEAB7FFB86D25FD38FB2858E5FB4E" box="[688,739,1179,1201]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
: p. 46.
</paragraph>
<paragraph id="C11636A5FFB86D25FF1FFB0E597AFB2F" blockId="5.[151,1344,1148,1324]" box="[151,892,1210,1232]" pageId="5" pageNumber="106">
2001
<taxonomicName id="06A94D26FFB86D25FF6DFB0E5846FB2F" ID-CoL="9GDR9" authority="Habe, 1953" authorityName="Habe" authorityYear="1953" box="[229,576,1210,1232]" class="Gastropoda" family="Colloniidae" genus="Leptothyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="species" species="tosaensis">
<emphasis id="F3DDEAB7FFB86D25FF6DFB0E5BB1FB2F" box="[229,439,1210,1232]" italics="true" pageId="5" pageNumber="106">Leptothyra tosaensis</emphasis>
(
<bibRefCitation id="A5384B54FFB86D25FE4DFB0E583FFB2F" author="Habe" box="[453,569,1210,1232]" pageId="5" pageNumber="106" refString="Habe, T. (1953) Descriptions of twelve new Japanese shells. Venus, 17, 130 - 144." type="journal article" year="1953">Habe, 1953</bibRefCitation>
)
</taxonomicName>
—Higo
<emphasis id="F3DDEAB7FFB86D25FD1DFB0F58CEFB2F" box="[661,712,1210,1232]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
: p. 17, fig. G198.
</paragraph>
<paragraph id="C11636A5FFB86D25FF1FFB6C5905FB11" blockId="5.[151,1344,1148,1324]" box="[151,771,1240,1262]" pageId="5" pageNumber="106">
2013
<taxonomicName id="06A94D26FFB86D25FF6DFB6C5839FB11" ID-CoL="9GDR9" authority="Habe, 1953" authorityName="Habe" authorityYear="1953" box="[229,575,1240,1262]" class="Gastropoda" family="Colloniidae" genus="Leptothyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="species" species="tosaensis">
<emphasis id="F3DDEAB7FFB86D25FF6DFB6C5BB1FB11" box="[229,439,1240,1262]" italics="true" pageId="5" pageNumber="106">Leptothyra tosaensis</emphasis>
(
<bibRefCitation id="A5384B54FFB86D25FE4DFB6C583EFB11" author="Habe" box="[453,568,1240,1262]" pageId="5" pageNumber="106" refString="Habe, T. (1953) Descriptions of twelve new Japanese shells. Venus, 17, 130 - 144." type="journal article" year="1953">Habe, 1953</bibRefCitation>
)
</taxonomicName>
—Poppe &amp; Poppe:
</paragraph>
<paragraph id="C11636A5FFB86D25FF6DFB435F46FAD3" blockId="5.[151,1344,1148,1324]" pageId="5" pageNumber="106">
http://www.conchology.be/?t=27&amp;family=
<taxonomicName id="06A94D26FFB86D25FD06FB435946FAF2" ID-CoL="6c408ed9-d677-46a6-8b54-251504fea780" box="[654,832,1271,1293]" class="Gastropoda" family="Colloniidae" kingdom="Animalia" pageId="5" pageNumber="106" phylum="Mollusca" rank="family">COLLONIIDAE</taxonomicName>
&amp;species=
<taxonomicName id="06A94D26FFB86D25FC21FB435E1AFAF2" ID-CoL="5DG6" box="[937,1052,1271,1293]" class="Gastropoda" family="Colloniidae" genus="Leptothyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="134" phylum="Mollusca" rank="genus">Leptothyra</taxonomicName>
%20
<taxonomicName id="06A94D26FFB86D25FBC0FB435ACCFAD3" ID-CoL="9GDR9" class="Gastropoda" family="Colloniidae" genus="Leptothyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="species" species="tosaensis">tosaensis 2013</taxonomicName>
<taxonomicName id="06A94D26FFB86D25FF6DFAA25843FAD3" ID-CoL="3MDTB" authority="Habe, 1953" authorityName="Habe" authorityYear="1953" box="[229,581,1302,1324]" class="Gastropoda" family="Colloniidae" genus="Homalopoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="species" species="tosaense">
<emphasis id="F3DDEAB7FFB86D25FF6DFAA25BCAFAD3" box="[229,460,1302,1324]" italics="true" pageId="5" pageNumber="106">Homalopoma tosaense</emphasis>
<bibRefCitation id="A5384B54FFB86D25FE5AFAA25843FAD3" author="Habe" box="[466,581,1302,1324]" pageId="5" pageNumber="106" refString="Habe, T. (1953) Descriptions of twelve new Japanese shells. Venus, 17, 130 - 144." type="journal article" year="1953">Habe, 1953</bibRefCitation>
</taxonomicName>
—Rosenberg: http://marinespecies.org/aphia.php?p=taxdetails&amp;id=737892
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB86D25FF1FFAEC5941F9FF" pageId="5" pageNumber="106" type="materials_examined">
<paragraph id="C11636A5FFB86D25FF1FFAEC58D4FA8E" blockId="5.[151,1436,1368,1825]" box="[151,722,1368,1393]" pageId="5" pageNumber="106">
<emphasis id="F3DDEAB7FFB86D25FF1FFAEC5B00FA8E" bold="true" box="[151,262,1368,1393]" pageId="5" pageNumber="106">Material.</emphasis>
Anda 2 (1); Anda 6 (1); AndaDeVos (4).
</paragraph>
<paragraph id="C11636A5FFB86D25FF4FFAC85941F9FF" blockId="5.[151,1436,1368,1825]" pageId="5" pageNumber="106">
<emphasis id="F3DDEAB7FFB86D25FF4FFAC85B9CFA6B" bold="true" box="[199,410,1404,1429]" pageId="5" pageNumber="106">Characterization.</emphasis>
Shell turbiniform, robust, H
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,
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W
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mm; P1 smooth, 1.0 whorls, DN
<quantity id="06519B40FFB86D25FAA4FAC85AC0FA47" metricMagnitude="-4" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="5" pageNumber="106" unit="mm" value="0.16" valueMax="0.17" valueMin="0.15">0.15–0.17 mm</quantity>
; P2, 0.3 whorls, with two robust spiral ribs on median-upper part and third basal spiral rib; P/T boundary distinct terminal varix at 1.3 whorls; T sculpture of spiral ribs; additional spiral ribs between uppermost spiral and suture; spiral and axial lirae in interspaces; umbilicus narrow.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB86D25FF4FF9B85F12F9B6" pageId="5" pageNumber="106" type="distribution">
<paragraph id="C11636A5FFB86D25FF4FF9B85F12F9B6" blockId="5.[151,1436,1368,1825]" pageId="5" pageNumber="106">
<emphasis id="F3DDEAB7FFB86D25FF4FF9B85B66F9DA" bold="true" box="[199,352,1548,1573]" pageId="5" pageNumber="106">Distribution.</emphasis>
Apart from the studied material, this species is known from Tosa Bay and southern Kyushu (
<collectingCountry id="B9BE7635FFB86D25FF17F9855AE2F9B6" box="[159,228,1585,1609]" name="Japan" pageId="5" pageNumber="106">Japan</collectingCountry>
), East
<collectingCountry id="B9BE7635FFB86D25FEA6F9845B77F9B6" box="[302,369,1584,1609]" name="China" pageId="5" pageNumber="106">China</collectingCountry>
Sea (
<bibRefCitation id="A5384B54FFB86D25FE27F984586DF9B6" author="Higo" box="[431,619,1584,1609]" pageId="5" pageNumber="106" refString="Higo, S., Callomon, P. &amp; Goto, Y. (1999) Catalogue and bibliography of the marine shell-bearing Mollusca of Japan. Elle Scientific Publications, Osaka, 749 pp." type="book" year="1999">
Higo
<emphasis id="F3DDEAB7FFB86D25FE78F986582EF9B6" box="[496,552,1584,1609]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
1999
</bibRefCitation>
), as well as Pliocene deposits of
<collectingCountry id="B9BE7635FFB86D25FC53F9845E03F9B6" box="[987,1029,1584,1609]" name="Fiji" pageId="5" pageNumber="106">Viti</collectingCountry>
Levu,
<collectingCountry id="B9BE7635FFB86D25FBDBF9845E7CF9B6" box="[1107,1146,1584,1609]" name="Fiji" pageId="5" pageNumber="106">Fiji</collectingCountry>
(
<bibRefCitation id="A5384B54FFB86D25FB00F9845F01F9B6" author="Ladd" box="[1160,1287,1584,1609]" pageId="5" pageNumber="106" refString="Ladd, H. S. (1982) Cenozoic fossil mollusks from Western Pacific islands; Gastropods (Eulimidae and Volutidae through Terebridae). Geological Survey Professional Paper, 1171, 1 - 100." type="journal article" year="1982">Ladd 1982</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFB86D25FF4FF9E05F62F8DE" pageId="5" pageNumber="106" type="discussion">
<paragraph id="C11636A5FFB86D25FF4FF9E05F62F8DE" blockId="5.[151,1436,1368,1825]" pageId="5" pageNumber="106">
<emphasis id="F3DDEAB7FFB86D25FF4FF9E05B3DF992" bold="true" box="[199,315,1620,1645]" pageId="5" pageNumber="106">Remarks.</emphasis>
<taxonomicName id="06A94D26FFB86D25FEC9F9E058BBF992" authority="Ladd, 1982" authorityName="Ladd" authorityYear="1982" box="[321,701,1620,1645]" class="Gastropoda" family="Colloniidae" genus="Homalopoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="species" species="druidi">
<emphasis id="F3DDEAB7FFB86D25FEC9F9E05823F992" box="[321,549,1620,1645]" italics="true" pageId="5" pageNumber="106">Homalopoma druidi</emphasis>
(
<bibRefCitation id="A5384B54FFB86D25FDBBF9E058B2F992" author="Ladd" box="[563,692,1620,1645]" pageId="5" pageNumber="106" refString="Ladd, H. S. (1982) Cenozoic fossil mollusks from Western Pacific islands; Gastropods (Eulimidae and Volutidae through Terebridae). Geological Survey Professional Paper, 1171, 1 - 100." type="journal article" year="1982">Ladd, 1982</bibRefCitation>
)
</taxonomicName>
is synonymized here with
<taxonomicName id="06A94D26FFB86D25FC6FF9E05F6BF992" authority="Habe, 1953" authorityName="Habe" authorityYear="1953" box="[999,1389,1620,1645]" class="Gastropoda" family="Colloniidae" genus="Homalopoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="species" species="tosaense">
<emphasis id="F3DDEAB7FFB86D25FC6FF9E05EE0F992" box="[999,1254,1620,1645]" italics="true" pageId="5" pageNumber="106">Homalopoma tosaense</emphasis>
<bibRefCitation id="A5384B54FFB86D25FB64F9E05F6BF992" author="Habe" box="[1260,1389,1620,1645]" pageId="5" pageNumber="106" refString="Habe, T. (1953) Descriptions of twelve new Japanese shells. Venus, 17, 130 - 144." type="journal article" year="1953">Habe, 1953</bibRefCitation>
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and
<taxonomicName id="06A94D26FFB86D25FF1FF9CC583CF96E" authority="Habe, 1953" authorityName="Habe" authorityYear="1953" box="[151,570,1656,1681]" class="Gastropoda" family="Colloniidae" genus="Leptothyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="species" species="tosaensis">
<emphasis id="F3DDEAB7FFB86D25FF1FF9CC5B8CF96E" box="[151,394,1656,1681]" italics="true" pageId="5" pageNumber="106">Leptothyra tosaensis</emphasis>
(
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)
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, because of its overall similar conchology. Although the
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of
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<emphasis id="F3DDEAB7FFB86D25FF1FF9285B7AF94A" box="[151,380,1692,1717]" italics="true" pageId="5" pageNumber="106">Homalopoma druidi</emphasis>
(
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)
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has a rather high spire when compared to the
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of
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<emphasis id="F3DDEAB7FFB86D25FB3AF9285F9AF94A" box="[1202,1436,1692,1717]" italics="true" pageId="5" pageNumber="106">Leptothyra tosaensis</emphasis>
(
<bibRefCitation id="A5384B54FFB86D25FF17F9745B23F926" author="Habe" box="[159,293,1728,1753]" pageId="5" pageNumber="106" refString="Habe, T. (1953) Descriptions of twelve new Japanese shells. Venus, 17, 130 - 144." type="journal article" year="1953">Habe, 1953</bibRefCitation>
)
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, its spire height falls well within the range of intraspecific variation observed on the website: http:// www.conchology.be/?t=27&amp;family=
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&amp;species=
<taxonomicName id="06A94D26FFB86D25FCEFF95059E0F902" box="[871,998,1764,1789]" class="Gastropoda" family="Colloniidae" genus="Leptothyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="134" phylum="Mollusca" rank="genus">Leptothyra</taxonomicName>
%20
<taxonomicName id="06A94D26FFB86D25FB9FF9505E7BF902" box="[1047,1149,1764,1789]" class="Gastropoda" family="Colloniidae" genus="Leptothyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="species" species="tosaensis">tosaensis</taxonomicName>
. It remains unclear whether the species should be assigned to
<taxonomicName id="06A94D26FFB86D25FDF8F8BC58EDF8DE" box="[624,747,1800,1825]" class="Gastropoda" family="Colloniidae" genus="Leptothyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFB86D25FDF8F8BC58EDF8DE" box="[624,747,1800,1825]" italics="true" pageId="5" pageNumber="106">Leptothyra</emphasis>
</taxonomicName>
or
<taxonomicName id="06A94D26FFB86D25FC98F8BC59AFF8DE" box="[784,937,1800,1825]" class="Gastropoda" family="Colloniidae" genus="Homalopoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="106" phylum="Mollusca" rank="genus">
<emphasis id="F3DDEAB7FFB86D25FC98F8BC59AFF8DE" box="[784,937,1800,1825]" italics="true" pageId="5" pageNumber="106">Homalopoma</emphasis>
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, as these genera are not well delimited.
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79
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<taxonomicName id="06A94D26FFBA6D27FF1FFAB25801FADF" ID-CoL="6YB3N" authority="Ladd, 1982" authorityName="Ladd" authorityYear="1982" box="[151,519,1286,1312]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="donaldi">
<emphasis id="F3DDEAB7FFBA6D27FF1FFAB25801FADF" bold="true" box="[151,519,1286,1312]" pageId="7" pageNumber="108">
<emphasis id="F3DDEAB7FFBA6D27FF1FFAB25B73FADF" bold="true" box="[151,373,1286,1312]" italics="true" pageId="7" pageNumber="108">Seguenzia donaldi</emphasis>
<bibRefCitation id="A5384B54FFBA6D27FEF4FAB25801FADF" author="Ladd" box="[380,519,1286,1312]" pageId="7" pageNumber="108" refString="Ladd, H. S. (1982) Cenozoic fossil mollusks from Western Pacific islands; Gastropods (Eulimidae and Volutidae through Terebridae). Geological Survey Professional Paper, 1171, 1 - 100." type="journal article" year="1982">Ladd, 1982</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C11636A5FFBA6D27FF1FFA9D5B28FABD" blockId="7.[151,519,1286,1346]" box="[151,302,1321,1346]" pageId="7" pageNumber="108">
(
<figureCitation id="59922A20FFBA6D27FF16FA9D5B20FABD" box="[158,294,1321,1346]" captionStart="FIGURES 8 – 12" captionStartId="8.[151,264,1849,1871]" captionTargetBox="[243,1351,195,1825]" captionTargetId="figure@8.[241,1354,193,1828]" captionTargetPageId="8" captionText="FIGURES 8 – 12. Seguenziidae. 8. Seguenzia donaldi Ladd, 1982. RGM 608.215. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 9. Seguenzia donaldi Ladd, 1982. RGM 608.216. Locality Anda 3. (a) rear view, (b) apertural view. 10. Seguenzia cf. elegantissima Poppe, Tagaro &amp; Dekker, 2006. RGM 608.221. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) side view, (d) basal view, (e) apical view. 11. Seguenzia cf. elegantissima Poppe, Tagaro &amp; Dekker, 2006. RGM 608.222. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) spire. 12. Seguenzia cf. elegantissima Poppe, Tagaro &amp; Dekker, 2006. RGM 608.223. Locality AndaDeVos. SEM detail of protoconch." httpUri="https://zenodo.org/record/250963/files/figure.png" pageId="7" pageNumber="108">Figures 8–9</figureCitation>
)
</paragraph>
<paragraph id="C11636A5FFBA6D27FF1FFAC55984FA78" blockId="7.[151,898,1393,1415]" box="[151,898,1393,1415]" pageId="7" pageNumber="108">
1982
<taxonomicName id="06A94D26FFBA6D27FF79FAC65BADFA78" box="[241,427,1393,1415]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="donaldi">
<emphasis id="F3DDEAB7FFBA6D27FF79FAC65BADFA78" box="[241,427,1393,1415]" italics="true" pageId="7" pageNumber="108">Seguenzia donaldi</emphasis>
</taxonomicName>
nov. spec.—Ladd: p. 32–33, pl. 35, figs 9–15.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFBA6D27FF1FFA005B67F980" pageId="7" pageNumber="108" type="materials_examined">
<paragraph id="C11636A5FFBA6D27FF1FFA005915FA33" blockId="7.[151,1436,1460,1952]" box="[151,787,1460,1485]" pageId="7" pageNumber="108">
<emphasis id="F3DDEAB7FFBA6D27FF1FFA005B00FA32" bold="true" box="[151,262,1460,1485]" pageId="7" pageNumber="108">Material.</emphasis>
Anda 6 (6); AndaDeVos (29); AndaClif3 (10).
</paragraph>
<paragraph id="C11636A5FFBA6D27FF4FFA635B67F980" blockId="7.[151,1436,1460,1952]" pageId="7" pageNumber="108">
<emphasis id="F3DDEAB7FFBA6D27FF4FFA635B9CFA0F" bold="true" box="[199,410,1495,1520]" pageId="7" pageNumber="108">Characterization.</emphasis>
Shell relatively broad conical to trochiform, H
<quantity id="06519B40FFBA6D27FC49FA6C5E1DFA10" box="[961,1051,1496,1520]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="108" unit="mm" value="1.9">1.9 mm</quantity>
,
<geoCoordinate id="A49D5062FFBA6D27FBA1FA6C5E76FA0F" box="[1065,1136,1496,1520]" direction="west" orientation="longitude" pageId="7" pageNumber="108" precision="5555" value="-1.9">
W
<quantity id="06519B40FFBA6D27FBC5FA6C5E76FA0F" box="[1101,1136,1496,1520]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="108" unit="mm" value="1.9">1.9</quantity>
</geoCoordinate>
mm; P small, 1.1 whorls; nucleus weakly inclined, DN
<quantity id="06519B40FFBA6D27FE60FA485889F9EB" box="[488,655,1532,1556]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="7" pageNumber="108" unit="mm" value="0.125" valueMax="0.14" valueMin="0.11">0.11–0.14 mm</quantity>
; P/T boundary weak; T with numerous sigmoid axial lirae fading on later T whorls; relatively wide body whorl with three peripheral spiral ribs and five to seven basal spiral ribs; upper apertural retraction deep, upper projection strongly expanding forwards; basal columellar fold or tooth present; umbilicus narrow.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFBA6D27FF4FF9385ADBF85F" pageId="7" pageNumber="108" type="distribution">
<paragraph id="C11636A5FFBA6D27FF4FF9385B2FF937" blockId="7.[151,1436,1460,1952]" pageId="7" pageNumber="108">
<emphasis id="F3DDEAB7FFBA6D27FF4FF9385B66F95A" bold="true" box="[199,352,1676,1701]" pageId="7" pageNumber="108">Distribution.</emphasis>
Apart from the studied material, this species is known from Pleistocene deposits of
<collectingCountry id="B9BE7635FFBA6D27FAC8F9395F9AF95B" box="[1344,1436,1677,1700]" name="Vanuatu" pageId="7" pageNumber="108">Vanuatu</collectingCountry>
(
<bibRefCitation id="A5384B54FFBA6D27FF16F91B5B1BF937" author="Ladd" box="[158,285,1711,1736]" pageId="7" pageNumber="108" refString="Ladd, H. S. (1982) Cenozoic fossil mollusks from Western Pacific islands; Gastropods (Eulimidae and Volutidae through Terebridae). Geological Survey Professional Paper, 1171, 1 - 100." type="journal article" year="1982">Ladd 1982</bibRefCitation>
).
</paragraph>
<paragraph id="C11636A5FFBA6D27FF4FF9605ADBF85F" blockId="7.[151,1436,1460,1952]" pageId="7" pageNumber="108">
<emphasis id="F3DDEAB7FFBA6D27FF4FF9605B79F912" bold="true" box="[199,383,1748,1773]" pageId="7" pageNumber="108">Differentiation.</emphasis>
This species is rather similar to
<taxonomicName id="06A94D26FFBA6D27FD7FF9615F08F912" authority="Poppe, Tagaro &amp; Dekker, 2006" authorityName="Poppe, Tagaro &amp; Dekker" authorityYear="2006" box="[759,1294,1748,1773]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="trochiformis">
<emphasis id="F3DDEAB7FFBA6D27FD7FF9615999F912" box="[759,927,1748,1773]" italics="true" pageId="7" pageNumber="108">S. trochiformis</emphasis>
<bibRefCitation id="A5384B54FFBA6D27FC2FF9615F08F912" author="Poppe" box="[935,1294,1748,1773]" pageId="7" pageNumber="108" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter" year="2006">Poppe, Tagaro &amp; Dekker, 2006</bibRefCitation>
</taxonomicName>
, in having a strong tooth on the columella and an aperture with projections and retractions. However,
<taxonomicName id="06A94D26FFBA6D27FB24F94C5F53F8EF" box="[1196,1365,1783,1808]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="trochiformis">
<emphasis id="F3DDEAB7FFBA6D27FB24F94C5F53F8EF" box="[1196,1365,1783,1808]" italics="true" pageId="7" pageNumber="108">S. trochiformis</emphasis>
</taxonomicName>
has a higher spire (H/W ratio of
<typeStatus id="1E128807FFBA6D27FE49F8A85825F8CA" box="[449,547,1820,1845]" pageId="7" pageNumber="108" type="holotype">holotype</typeStatus>
<bibRefCitation id="A5384B54FFBA6D27FDA2F8A958FFF8CA" author="Poppe" box="[554,761,1820,1845]" pageId="7" pageNumber="108" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter" year="2006">
Poppe
<emphasis id="F3DDEAB7FFBA6D27FDFEF8A958A8F8CA" box="[630,686,1820,1845]" italics="true" pageId="7" pageNumber="108">et al.</emphasis>
, 2006
</bibRefCitation>
: 1.10; H/W ratio of
<taxonomicName id="06A94D26FFBA6D27FC5FF8A95E4EF8CA" box="[983,1096,1820,1845]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="donaldi">
<emphasis id="F3DDEAB7FFBA6D27FC5FF8A95E4EF8CA" box="[983,1096,1820,1845]" italics="true" pageId="7" pageNumber="108">S. donaldi</emphasis>
</taxonomicName>
1.02-1.07 (table 2)) and a less swollen body whorl.
<taxonomicName id="06A94D26FFBA6D27FE01F8F4585FF8A7" box="[393,601,1855,1880]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="donaldi">
<emphasis id="F3DDEAB7FFBA6D27FE01F8F4585FF8A7" box="[393,601,1855,1880]" italics="true" pageId="7" pageNumber="108">Seguenzia donaldi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="06A94D26FFBA6D27FD7BF8F4598DF8A7" box="[755,907,1855,1880]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="keikoae">
<emphasis id="F3DDEAB7FFBA6D27FD7BF8F4590EF8A8" box="[755,776,1856,1879]" italics="true" pageId="7" pageNumber="108">S.</emphasis>
cf.
<emphasis id="F3DDEAB7FFBA6D27FCBEF88B598DF8A7" box="[822,907,1855,1880]" italics="true" pageId="7" pageNumber="108">keikoae</emphasis>
</taxonomicName>
by its wider whorls. The keel in
<taxonomicName id="06A94D26FFBA6D27FA85F8F45F87F8A7" box="[1293,1409,1855,1880]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="donaldi">
<emphasis id="F3DDEAB7FFBA6D27FA85F8F45F87F8A7" box="[1293,1409,1855,1880]" italics="true" pageId="7" pageNumber="108">S. donaldi</emphasis>
</taxonomicName>
is stronger, its axial lirae are weaker, the protoconch less inclined, the base of the body whorl flatter and the umbilicus wider.
</paragraph>
</subSubSection>
</treatment>
</document>

134
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<subSubSection id="89B3652EFFBA6D27FF1FFD1A5951FCB1" pageId="7" pageNumber="108" type="nomenclature">
<paragraph id="C11636A5FFBA6D27FF1FFD1A5925FD37" blockId="7.[151,803,686,746]" box="[151,803,686,712]" pageId="7" pageNumber="108">
<heading id="9A5E81C9FFBA6D27FF1FFD1A5925FD37" bold="true" box="[151,803,686,712]" fontSize="11" level="1" pageId="7" pageNumber="108" reason="1">
<taxonomicName id="06A94D26FFBA6D27FF1FFD1A5925FD37" ID-CoL="6YB2F" authority="Poppe, Tagaro &amp; Dekker, 2006" authorityName="Poppe, Tagaro &amp; Dekker" authorityYear="2006" box="[151,803,686,712]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="keikoae">
<emphasis id="F3DDEAB7FFBA6D27FF1FFD1A5925FD37" bold="true" box="[151,803,686,712]" pageId="7" pageNumber="108">
<emphasis id="F3DDEAB7FFBA6D27FF1FFD1A5B15FD37" bold="true" box="[151,275,686,712]" italics="true" pageId="7" pageNumber="108">Seguenzia</emphasis>
cf.
<emphasis id="F3DDEAB7FFBA6D27FEB7FD1A5B9AFD37" bold="true" box="[319,412,686,712]" italics="true" pageId="7" pageNumber="108">keikoae</emphasis>
<bibRefCitation id="A5384B54FFBA6D27FE2BFD1A5925FD37" author="Poppe" box="[419,803,686,712]" pageId="7" pageNumber="108" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter" year="2006">Poppe, Tagaro &amp; Dekker, 2006</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C11636A5FFBA6D27FF1FFD655B28FD15" blockId="7.[151,803,686,746]" box="[151,302,721,746]" pageId="7" pageNumber="108">(Figures 6–7)</paragraph>
<paragraph id="C11636A5FFBA6D27FF1FFCAD5951FCB1" blockId="7.[151,1023,793,846]" pageId="7" pageNumber="108">
cf. 2006
<taxonomicName id="06A94D26FFBA6D27FE8BFCAE5BBBFCD0" box="[259,445,793,815]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="keikoae">
<emphasis id="F3DDEAB7FFBA6D27FE8BFCAE5BBBFCD0" box="[259,445,793,815]" italics="true" pageId="7" pageNumber="108">Seguenzia keikoae</emphasis>
</taxonomicName>
nov. spec.—Poppe, Tagaro &amp; Dekker: p. 27, pl. 5, fig. 3. cf. 2008a
<taxonomicName id="06A94D26FFBA6D27FE8BFC8D5BBBFCB1" box="[259,445,824,846]" class="Gastropoda" family="Seguenziidae" genus="Seguenzia" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="keikoae">
<emphasis id="F3DDEAB7FFBA6D27FE8BFC8D5BBBFCB1" box="[259,445,824,846]" italics="true" pageId="7" pageNumber="108">Seguenzia keikoae</emphasis>
</taxonomicName>
—Poppe &amp; Tagaro: p. 162, pl. 26, fig. 3.
</paragraph>
</subSubSection>
<subSubSection id="89B3652EFFBA6D27FF1FFCCE5B92FC00" pageId="7" pageNumber="108" type="materials_examined">
<paragraph id="C11636A5FFBA6D27FF1FFCCE5F0DFC6C" blockId="7.[151,1436,890,1203]" box="[151,1291,890,915]" pageId="7" pageNumber="108">
<emphasis id="F3DDEAB7FFBA6D27FF1FFCCE5B00FC6C" bold="true" box="[151,262,890,915]" pageId="7" pageNumber="108">Material.</emphasis>
Anda 2 (1); Anda 3 (1); Anda 4 (1); Anda 6 (1); AndaDeVos (2); AndaClif3 (1); Roxas (6).
</paragraph>
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<emphasis id="F3DDEAB7FFBA6D27FF4FFC2A5B9CFC48" bold="true" box="[199,410,926,951]" pageId="7" pageNumber="108">Characterization.</emphasis>
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<emphasis id="F3DDEAB7FFBA6D27FE28FC2B5814FC49" box="[416,530,927,950]" italics="true" pageId="7" pageNumber="108">Seguenzia</emphasis>
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with numerous low, thin more or less regularly spaced sigmoid axial lirae; H
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,
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W
<quantity id="06519B40FFBA6D27FF7CFC775B11FC24" box="[244,279,963,987]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="108" unit="mm" value="1.7">1.7</quantity>
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mm; DN
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; two strong spiral ribs on shoulder, five on base; columella denticulate; adult shell with closed umbilicus.
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<emphasis id="F3DDEAB7FFBA6D27FF4FFBBE5B66FBDC" bold="true" box="[199,352,1034,1059]" pageId="7" pageNumber="108">Distribution.</emphasis>
Apart from the studied material,
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<emphasis id="F3DDEAB7FFBA6D27FD61FBBF59BCFBDC" box="[745,954,1034,1059]" italics="true" pageId="7" pageNumber="108">Seguenzia keikoae</emphasis>
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has been reported from Bohol, off Pamilacan
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,
<collectingCountry id="B9BE7635FFBA6D27FD08FB9A5907FBB8" box="[640,769,1070,1095]" name="Philippines" pageId="7" pageNumber="108">Philippines</collectingCountry>
, in a sample from between
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water depth.
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<emphasis id="F3DDEAB7FFBA6D27FF4FFBE65B3DFB95" bold="true" box="[199,315,1106,1131]" pageId="7" pageNumber="108">Remarks.</emphasis>
The attribution to
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<emphasis id="F3DDEAB7FFBA6D27FD9BFBE75883FB94" box="[531,645,1106,1131]" italics="true" pageId="7" pageNumber="108">S. keikoae</emphasis>
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is uncertain despite the overall similar shell morphology because the adult specimens in our material have a closed umbilicus. Specimens shown by
<bibRefCitation id="A5384B54FFBA6D27FB9DFBC35EE8FB70" box="[1045,1262,1142,1167]" pageId="7" pageNumber="108" refString="Poppe, G. T., Tagaro, S. P. &amp; Dekker, H. (2006) The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2, 3 - 228." type="book chapter">
Poppe
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(2006)
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have a slit-like umbilicus, but the latter specimens might be juveniles.
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Family
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<emphasis id="F3DDEAB7FFBA6D27FF1FFE935B0DFEBF" bold="true" box="[151,267,295,320]" pageId="7" pageNumber="108">Remarks.</emphasis>
Protoconchs were examined across a range of
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material. Apart from
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cf.
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, protoconchs were invariably deeply corroded and are not illustrated here.
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<paragraph id="D9FBB6F05EE51B8341A29C74944E0515" pageId="0" pageNumber="10948">Distribution</paragraph>
<paragraph id="E87E57760497EEDFC6B0D1D62421DB6B" pageId="0" pageNumber="10948">FLO*; FAI; PIC; SJG*; TER; SMG; SMR</paragraph>
</subSubSection>
<subSubSection id="03787C856AACC2BAAED9DE73DA7FCC98" pageId="0" pageNumber="10948" type="notes">
<paragraph id="CC6A0455E98FFEA2F85336D189287010" pageId="0" pageNumber="10948">Notes</paragraph>
<paragraph id="7077DF985DD08C89B52A3174F6640E8A" pageId="0" pageNumber="10948">Biogeographical Realm: Western Palearctic (Macaronesia)</paragraph>
</subSubSection>
</treatment>
</document>

204
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A-G)
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<caption id="95D64094FF81CF2ADFCAF8F5FCF3A6B0" ID-DOI="http://doi.org/10.5281/zenodo.5404088" ID-Zenodo-Dep="5404088" httpUri="https://zenodo.org/record/5404088/files/figure.png" pageId="5" pageNumber="155" startId="5.[159,171,1888,1907]" targetBox="[228,1333,325,1834]" targetPageId="5">
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FIGURE 2:
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<emphasis id="F3DDCC0EFF81CF2ADE50F8F5FE71A64C" box="[261,415,1888,1907]" italics="true" pageId="5" pageNumber="155">Nilotonia bisetosa</emphasis>
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, female: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – genital field; (D) – palp; (E) – I-Leg-4-6; (F) - IV-Leg-6. Scale bars = 200 µm (A-B), 50 µm (C-F).
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FIGURE 3:
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<emphasis id="F3DDCC0EFF82CF29DE51F8F5FE49A64C" box="[260,423,1888,1907]" italics="true" pageId="6" pageNumber="156">Nilotonia longiseta</emphasis>
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(A-C, E-G: male, D: female): (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – I-Leg-4-6; (D-E) – genital field; (F) – palp; (G) – IV-Leg-4-6. Scale bars = 200 µm (A-B), 50 µm (C-G).
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<paragraph id="C116101CFF83CF28DF90FE13FE4DA372" blockId="7.[159,762,311,1948]" pageId="7" pageNumber="157">Diagnosis — Male with a large area (L 106, W 104) of secondary sclerotization posterior to genital field; secondary sclerotization in female posterior to genital field smaller; pregenital sclerite of female extended anteriorly; setae of D2 and D3 long, posterior dorsal plate large.</paragraph>
<paragraph id="C116101CFF83CF28DF90FDF4FEA0A346" blockId="7.[159,762,311,1948]" box="[197,334,609,633]" pageId="7" pageNumber="157">Description</paragraph>
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Male — Idiosoma soft, dorsally L 551 (530-591), W 389 (421-429), ventrally L 608 (559-664). Dorsum with one large, posterior plate, L 160 (180- 218), W 112 (130-148). Setae of glandularia D2 and D3 (sensu
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) long. Cx-I separated medially, with triangular apodemes; Cx-I posteriorly with narrow secondary sclerotization. Cx-III with a blunt corner medially, anteriorly of genital field. Genital field (
<figureCitation id="59920C99FF83CF28DE15FC33FE7BA281" box="[320,405,934,958]" captionStart="FIGURE" captionStartId="6.[159,171,1888,1907]" captionTargetBox="[213,1358,313,1835]" captionTargetId="figure-7@6.[199,1389,307,1857]" captionTargetPageId="6" captionText="FIGURE 3: Nilotonia longiseta n. sp. (A-C, E-G: male, D: female): (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – I-Leg-4-6; (D-E) – genital field; (F) – palp; (G) – IV-Leg-4-6. Scale bars = 200 µm (A-B), 50 µm (C-G)." figureDoi="http://doi.org/10.5281/zenodo.5404090" httpUri="https://zenodo.org/record/5404090/files/figure.png" pageId="7" pageNumber="157">Fig. 3E</figureCitation>
) with three pairs of elongated acetabula, L 106, W 76. Pregenital sclerite large, bowed, posteriorly of genital field a somewhat triangular secondary sclerotization. Chelicere L 247, cheliceral claw L 74. Palp (
<figureCitation id="59920C99FF83CF28DE83FBA7FDC4A575" box="[470,554,1074,1098]" captionStart="FIGURE" captionStartId="6.[159,171,1888,1907]" captionTargetBox="[213,1358,313,1835]" captionTargetId="figure-7@6.[199,1389,307,1857]" captionTargetPageId="6" captionText="FIGURE 3: Nilotonia longiseta n. sp. (A-C, E-G: male, D: female): (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – I-Leg-4-6; (D-E) – genital field; (F) – palp; (G) – IV-Leg-4-6. Scale bars = 200 µm (A-B), 50 µm (C-G)." figureDoi="http://doi.org/10.5281/zenodo.5404090" httpUri="https://zenodo.org/record/5404090/files/figure.png" pageId="7" pageNumber="157">Fig. 3F</figureCitation>
): total L 316; L: P- 1, 22; P-2, 96; P-3, 62; P-4, 108; P-5, 28; P-2 with a 72 long ventral seta, which is inserted somewhat medially; ventral margin of P-2 distally with 5-6 denticles of variable size; P-3 with a long medial seta which extends beyond posterior margin of P-4; P-4 with a short setal tubercle, ventral margin with three setae various in length, of which the seta on the setal tubercle is the longest. L of I-Leg-4-6: 101, 126, 96. I- Leg-4 with four heavy distal setae, two longer, two shorter (
<figureCitation id="59920C99FF83CF28DE57FA04FEB5A496" box="[258,347,1425,1449]" captionStart="FIGURE" captionStartId="6.[159,171,1888,1907]" captionTargetBox="[213,1358,313,1835]" captionTargetId="figure-7@6.[199,1389,307,1857]" captionTargetPageId="6" captionText="FIGURE 3: Nilotonia longiseta n. sp. (A-C, E-G: male, D: female): (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – I-Leg-4-6; (D-E) – genital field; (F) – palp; (G) – IV-Leg-4-6. Scale bars = 200 µm (A-B), 50 µm (C-G)." figureDoi="http://doi.org/10.5281/zenodo.5404090" httpUri="https://zenodo.org/record/5404090/files/figure.png" pageId="7" pageNumber="157">Fig. 3C</figureCitation>
). L of IV-Leg-4-6: 178, 202, 182. IV- Leg-6 with a 90 long distal seta. IV-Leg-5 with three heavy distal setae, at least the longest of these serrated. IV-Leg-4 with five heavy distal setae, some of these (or all?) serrated. Claws of legs I-III with a comb and a clawlet. Excretory pore smooth.
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Female — Idiosoma soft, dorsally L 656 (559- 802), W 502 (421-591), ventrally L 713 (616-850). Dorsum with one large, posterior plate, L 194 (150- 196), W 124 (106-128), and a pair of smaller more anteriorly located plates. Setae of glandularia D2 and D3 (sensu
<bibRefCitation id="A5386DEDFF83CF28DE02F88EFE02A60C" author="Wiles, P. R." box="[343,492,1819,1843]" pageId="7" pageNumber="157" pagination="1237 - 1251" refId="ref18532" refString="Wiles, P. R. 1997 a - The homology of glands and glandularia in the water mites (Acari: Hydrachnidia) - J. Natur. Hist., 31: 1237 - 1251." type="journal article" year="1997">Wiles, 1997a</bibRefCitation>
) long. Cx-I separated medially, with triangular apodemes; Cx-I posteriorly with narrow secondary sclerotization. Cx-III with a blunt corner medially, anteriorly of genital field. Chelicerae of mounted female not measurable. Genital field with three pairs of elongated acetabula, L 146, W 82. Pregenital sclerite larger and more extended anteriorly compared to male (
<figureCitation id="59920C99FF83CF28DA3DFE35FCB0A0E4" captionStart="FIGURE" captionStartId="6.[159,171,1888,1907]" captionTargetBox="[213,1358,313,1835]" captionTargetId="figure-7@6.[199,1389,307,1857]" captionTargetPageId="6" captionText="FIGURE 3: Nilotonia longiseta n. sp. (A-C, E-G: male, D: female): (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – I-Leg-4-6; (D-E) – genital field; (F) – palp; (G) – IV-Leg-4-6. Scale bars = 200 µm (A-B), 50 µm (C-G)." figureDoi="http://doi.org/10.5281/zenodo.5404090" httpUri="https://zenodo.org/record/5404090/files/figure.png" pageId="7" pageNumber="157">Fig. 3D</figureCitation>
). Secondary sclerotization posterior to genital field as in male, but shorter. Palp: total L 420; L: P-1, 32; P-2, 136; P-3, 78; P-4, 140; P-5, 34; palp as in male, but ventral margin of P-2 with numerous (15-20) denticles. L of I-Leg-4-6: 120, 140, 104. L of IV-Leg-4-6: 206, 244, 236. Distal seta of IV-Leg-6 100 long. Legs as in male. Excretory pore smooth.
</paragraph>
<paragraph id="C116101CFF83CF28DC35FD54FC73A3C3" blockId="7.[826,1429,311,912]" pageId="7" pageNumber="157">Etymology — Named after the long setae on the dorsum.</paragraph>
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Remarks — The combination of long setae of D2 and D3, the typical shape of the female pregenital sclerite and a large dorsal plate separates this species from other
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Material examined — 0/1/0, Ayn Razat,
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Remarks — Previously reported from
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<emphasis id="F3DDCC0EFF80CF2BDFA3FB90FDB7A522" bold="true" box="[246,601,1029,1053]" italics="true" pageId="4" pageNumber="154">Nilotonia (Dartiella) bisetosa</emphasis>
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<paragraph id="C116101CFF80CF2BDE2DFBAAFDCEA568" blockId="4.[376,544,1087,1111]" box="[376,544,1087,1111]" pageId="4" pageNumber="154">
(
<figureCitation id="59920C99FF80CF2BDE2AFBAAFE05A568" box="[383,491,1087,1111]" captionStart="FIGURE" captionStartId="5.[159,171,1888,1907]" captionTargetBox="[228,1333,325,1834]" captionTargetId="figure-4@5.[213,1374,307,1857]" captionTargetPageId="5" captionText="FIGURE 2: Nilotonia bisetosa n. sp., female: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – genital field; (D) – palp; (E) – I-Leg-4-6; (F) - IV-Leg-6. Scale bars = 200 µm (A-B), 50 µm (C-F)." figureDoi="http://doi.org/10.5281/zenodo.5404088" httpUri="https://zenodo.org/record/5404088/files/figure.png" pageId="4" pageNumber="154">Figures 2</figureCitation>
A-F)
</paragraph>
<paragraph id="C116101CFF80CF2BDF90FB12FD9EA415" blockId="4.[159,762,1158,1949]" pageId="4" pageNumber="154">
Type series —
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<typeStatus id="1E12AEBEFF80CF2BDEC7FB13FDEFA5A1" box="[402,513,1158,1182]" pageId="4" pageNumber="154" type="holotype">Holotype</typeStatus>
female, spring
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, crossing road to
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border,
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.
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: 0/1/0, same data as holotype. Other material. 0/0/4, same data as holotype.
</paragraph>
<paragraph id="C116101CFF80CF2BDF90FAAAFF26A445" blockId="4.[159,762,1158,1949]" pageId="4" pageNumber="154">Diagnosis — IV-Leg-6 with two long, distal setae.</paragraph>
<paragraph id="C116101CFF80CF2BDF90FA1AFEA0A498" blockId="4.[159,762,1158,1949]" box="[197,334,1423,1447]" pageId="4" pageNumber="154">Description</paragraph>
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Female — Idiosoma soft, lineated, dorsally L 786 (1142), W 608 (891), ventrally L 842 (1102). Dorsum with one large, posterior plate, L 138 (113), W 94 (81). Setae associated with pre-antennal glandularia large. Cx-I separated medially, with triangular apodemes. Cx-III with a blunt corner medially, anteriorly of genital field. Genital field (
<figureCitation id="59920C99FF80CF2BDDC5F91AFD1CA798" box="[656,754,1679,1703]" captionStart="FIGURE" captionStartId="5.[159,171,1888,1907]" captionTargetBox="[228,1333,325,1834]" captionTargetId="figure-4@5.[213,1374,307,1857]" captionTargetPageId="5" captionText="FIGURE 2: Nilotonia bisetosa n. sp., female: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – genital field; (D) – palp; (E) – I-Leg-4-6; (F) - IV-Leg-6. Scale bars = 200 µm (A-B), 50 µm (C-F)." figureDoi="http://doi.org/10.5281/zenodo.5404088" httpUri="https://zenodo.org/record/5404088/files/figure.png" pageId="4" pageNumber="154">Fig. 2C</figureCitation>
) with three pairs of elongated acetabula, L 208, W 154. Pre-genital sclerite of moderate size, bowed, post-genital sclerite straight and narrow. Chelicera L 413, cheliceral claw L 100. Palp (
<figureCitation id="59920C99FF80CF2BDD19F88EFD41A60C" box="[588,687,1819,1843]" captionStart="FIGURE" captionStartId="5.[159,171,1888,1907]" captionTargetBox="[228,1333,325,1834]" captionTargetId="figure-4@5.[213,1374,307,1857]" captionTargetPageId="5" captionText="FIGURE 2: Nilotonia bisetosa n. sp., female: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – genital field; (D) – palp; (E) – I-Leg-4-6; (F) - IV-Leg-6. Scale bars = 200 µm (A-B), 50 µm (C-F)." figureDoi="http://doi.org/10.5281/zenodo.5404088" httpUri="https://zenodo.org/record/5404088/files/figure.png" pageId="4" pageNumber="154">Fig. 2D</figureCitation>
): total L 527; L: P-1, 32; P-2, 170, P-3, 94, P-4, 180, P-5, 51; P-2 ventrally with a long seta, ventral margin with numerous small denticles; P-4 with two setal tubercles, each with a seta, one of which is much longer than the other; ventral margin of P-4 with three more distal setae; P-5 with three heavy, distal setae. L of I-Leg-4-6 (
<figureCitation id="59920C99FF80CF2BDB61FE35FB65A087" box="[1076,1163,416,440]" captionStart="FIGURE" captionStartId="5.[159,171,1888,1907]" captionTargetBox="[228,1333,325,1834]" captionTargetId="figure-4@5.[213,1374,307,1857]" captionTargetPageId="5" captionText="FIGURE 2: Nilotonia bisetosa n. sp., female: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – genital field; (D) – palp; (E) – I-Leg-4-6; (F) - IV-Leg-6. Scale bars = 200 µm (A-B), 50 µm (C-F)." figureDoi="http://doi.org/10.5281/zenodo.5404088" httpUri="https://zenodo.org/record/5404088/files/figure.png" pageId="4" pageNumber="154">Fig. 2E</figureCitation>
): 176, 190, 140. I-Leg-5 anteroventrally with three setae, I-Leg-4 anteroventrally with three heavy, serrated setae, more medially two more serrated setae. L of IV-Leg-4-6: 259, 284, 243. IV-Leg-6 (
<figureCitation id="59920C99FF80CF2BDB7DFDB9FB6CA37B" box="[1064,1154,556,580]" captionStart="FIGURE" captionStartId="5.[159,171,1888,1907]" captionTargetBox="[228,1333,325,1834]" captionTargetId="figure-4@5.[213,1374,307,1857]" captionTargetPageId="5" captionText="FIGURE 2: Nilotonia bisetosa n. sp., female: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – genital field; (D) – palp; (E) – I-Leg-4-6; (F) - IV-Leg-6. Scale bars = 200 µm (A-B), 50 µm (C-F)." figureDoi="http://doi.org/10.5281/zenodo.5404088" httpUri="https://zenodo.org/record/5404088/files/figure.png" pageId="4" pageNumber="154">Fig. 2F</figureCitation>
) with two distal setae, the distal one 132 long, the proximal 123 long. Ventral margin of IV-Leg-5 with six setae, ventral margin of IV-Leg-4 with eight setae. IV-Leg-5 distally with five heavy setae of various lengths, IV-Leg-4 distally with seven heavy setae of various lengths, three of these long and serrate. Claws of legs I-III with comb. Excretory pore slit-like, with an indistinct ring.
</paragraph>
<paragraph id="C116101CFF80CF2BDC35FCE5FBAEA2B7" blockId="4.[826,1429,311,1489]" box="[864,1088,880,904]" pageId="4" pageNumber="154">Male — Unknown.</paragraph>
<paragraph id="C116101CFF80CF2BDC35FC0EFB95A500" blockId="4.[826,1429,311,1489]" pageId="4" pageNumber="154">Deutonymph — The nymphs collected together with the adults are provisionally assigned to the new species, IV-Leg-6 of the nymph has only one long seta. P-2 is ventrally without a seta, but the many denticles are present.</paragraph>
<paragraph id="C116101CFF80CF2BDC35FBC7FC48A5B2" blockId="4.[826,1429,311,1489]" pageId="4" pageNumber="154">Etymology — Named for the two long setae of IV-Leg-6.</paragraph>
<paragraph id="C116101CFF80CF2BDC35FB35FAF3A4EF" blockId="4.[826,1429,311,1489]" pageId="4" pageNumber="154">
Remarks — The subgenus
<taxonomicName id="06A96B9FFF80CF2BDBF7FB34FAEDA587" authorityName="Viets" authorityYear="1929" box="[1186,1283,1185,1208]" class="Arachnida" family="Anisitsiellidae" genus="Dartonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="F3DDCC0EFF80CF2BDBF7FB34FAEDA587" box="[1186,1283,1185,1208]" italics="true" pageId="4" pageNumber="154">Dartonia</emphasis>
</taxonomicName>
has IV-Leg- 6 with more than one heavy distal setae, but is also characterized by P-2 with more than eight setae and the claw of III-Leg has heavy clawlets, leading to a bidentate or tridentate appearance (
<bibRefCitation id="A5386DEDFF80CF2BDA67FAB8FC9AA457" author="Panesar, A. R." pageId="4" pageNumber="154" pagination="1 - 144" refId="ref18080" refString="Panesar, A. R. 2004 - Evolution in water mites (Hydrachnellae, Actinedida, Acari). A revision of the Anisitsiellidae Koenike, 1910 - Bonner Zool. Monogr., 52: 1 - 144." type="journal article" year="2004">Panesar, 2004</bibRefCitation>
). The latter two characters are absent in the new species, and therefore it is placed in the subgenus
<taxonomicName id="06A96B9FFF80CF2BDCDCFA03FC07A491" box="[905,1001,1430,1454]" class="Arachnida" family="Anisitsiellidae" genus="Nilotonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="154" phylum="Arthropoda" rank="subGenus" subGenus="Dartiella">
<emphasis id="F3DDCC0EFF80CF2BDCDCFA03FC07A491" box="[905,1001,1430,1454]" italics="true" pageId="4" pageNumber="154">Dartiella</emphasis>
</taxonomicName>
. No other species of this subgenus has a IV-Leg-6 with two distal long setae.
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(
<figureCitation id="59920C99FF83CF28DCA1FBF0FB8FA541" box="[1012,1121,1125,1150]" captionStart="FIGURE" captionStartId="8.[159,171,1888,1907]" captionTargetBox="[253,1361,311,1857]" captionTargetId="figure-7@8.[227,1361,307,1857]" captionTargetPageId="8" captionText="FIGURE 4: Torrenticola arabica n. sp., male (A-D, F holotype; E, G specimen from Wadi Ban Auy): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view; (D-E) – palp, lateral view; (F) – capitulum; (G) – ejaculatory complex. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404092" httpUri="https://zenodo.org/record/5404092/files/figure.png" pageId="7" pageNumber="157">Figures 4</figureCitation>
A-G, 5A-B)
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Type series —
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<typeStatus id="1E12AEBEFF83CF28DB4AFB33FB7FA581" box="[1055,1169,1190,1214]" pageId="7" pageNumber="157" type="holotype">Holotype</typeStatus>
: male, stream
<location id="C47646C7FF83CF28DA02FB33FC96A5DE" LSID="urn:lsid:plazi:treatment:4900A10AFF83CF24DC22FBB8FE22A5AD:C47646C7FF83CF28DA02FB33FC96A5DE" country="Oman" latitude="23.171618" longLatPrecision="1" longitude="57.199932" name="Wadi Ghul" pageId="7" pageNumber="157">Wadi Ghul</location>
,
<collectingCountry id="B9BE508CFF83CF28DCD3FB5CFC3FA5DE" box="[902,977,1225,1249]" name="Oman" pageId="7" pageNumber="157">Oman</collectingCountry>
,
<geoCoordinate id="A49D76DBFF83CF28DCB5FB5CFB80A5DF" box="[992,1134,1225,1249]" degrees="23" direction="north" minutes="10.297" orientation="latitude" pageId="7" pageNumber="157" precision="1" value="23.171618">23°10.297 N</geoCoordinate>
<geoCoordinate id="A49D76DBFF83CF28DB2DFB5CFAECA5DF" box="[1144,1282,1225,1249]" degrees="57" direction="east" minutes="11.996" orientation="longitude" pageId="7" pageNumber="157" precision="1" value="57.199932">57°11.996 E</geoCoordinate>
, alt.
<quantity id="0651BDF9FF83CF28DA10FB5CFA7EA5DF" box="[1349,1424,1225,1249]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="7" pageNumber="157" unit="m" value="805.0">
<elevation id="4A84F72FFF83CF28DA10FB5CFA7EA5DF" box="[1349,1424,1225,1249]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="7" pageNumber="157" unit="m" value="805.0">805 m</elevation>
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,
<date id="B51736DCFF83CF28DC6FFB79FC5AA43B" box="[826,948,1260,1284]" pageId="7" pageNumber="157" value="2008-11-12">
<collectingDate id="A553CF34FF83CF28DC6FFB79FC5AA43B" box="[826,948,1260,1284]" pageId="7" pageNumber="157" value="2008-11-12">12-xi-2008</collectingDate>
</date>
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.
<typeStatus id="1E12AEBEFF83CF28DC9CFB79FBAFA43B" box="[969,1089,1260,1284]" pageId="7" pageNumber="157" type="paratype">Paratypes</typeStatus>
: 20/26/0 (0/1/0 mounted), same data as holotype.
</paragraph>
<paragraph id="C116101CFF83CF28DC35FAAFFBBBA4A6" blockId="7.[826,1429,1190,1949]" pageId="7" pageNumber="157">
<materialsCitation id="71C11A41FF83CF28DC35FAAFFBBFA4A6" collectingDate="2008-09-13" country="Oman" elevation="655" latitude="23.278316" location="Further" longLatPrecision="1" longitude="57.4615" municipality="Wadi Ban Auf" pageId="7" pageNumber="157" specimenCount="1">
<location id="C47646C7FF83CF28DC35FAAFFC57A46D" LSID="urn:lsid:plazi:treatment:4900A10AFF83CF24DC22FBB8FE22A5AD:C47646C7FF83CF28DC35FAAFFC57A46D" box="[864,953,1338,1362]" country="Oman" latitude="23.278316" longLatPrecision="1" longitude="57.4615" municipality="Wadi Ban Auf" name="Further" pageId="7" pageNumber="157">Further</location>
records — 9/12/0 (2/1/0 mounted), stream
<collectingMunicipality id="21728A66FF83CF28DCDAFAC8FBD8A44A" box="[911,1078,1373,1397]" pageId="7" pageNumber="157">Wadi Ban Auf</collectingMunicipality>
,
<collectingCountry id="B9BE508CFF83CF28DB15FACBFB65A449" box="[1088,1163,1374,1398]" name="Oman" pageId="7" pageNumber="157">Oman</collectingCountry>
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<geoCoordinate id="A49D76DBFF83CF28DBC3FAC8FACEA44A" box="[1174,1312,1373,1397]" degrees="23" direction="north" minutes="16.699" orientation="latitude" pageId="7" pageNumber="157" precision="1" value="23.278316">23°16.699 N</geoCoordinate>
<geoCoordinate id="A49D76DBFF83CF28DA73FAC8FCA2A4A7" degrees="57" direction="east" minutes="27.690" orientation="longitude" pageId="7" pageNumber="157" precision="1" value="57.4615">57°27.690 E</geoCoordinate>
, alt.
<quantity id="0651BDF9FF83CF28DCD1FA15FC25A4A7" box="[900,971,1408,1432]" metricMagnitude="2" metricUnit="m" metricValue="6.55" pageId="7" pageNumber="157" unit="m" value="655.0">
<elevation id="4A84F72FFF83CF28DCD1FA15FC25A4A7" box="[900,971,1408,1432]" metricMagnitude="2" metricUnit="m" metricValue="6.55" pageId="7" pageNumber="157" unit="m" value="655.0">655 m</elevation>
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,
<date id="B51736DCFF83CF28DC82FA14FBBFA4A6" box="[983,1105,1409,1433]" pageId="7" pageNumber="157" value="2008-09-13">
<collectingDate id="A553CF34FF83CF28DC82FA14FBBFA4A6" box="[983,1105,1409,1433]" pageId="7" pageNumber="157" value="2008-09-13">13-ix-2008</collectingDate>
</date>
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.
</paragraph>
<paragraph id="C116101CFF83CF28DC35FA39FC50A7A9" blockId="7.[826,1429,1190,1949]" pageId="7" pageNumber="157">
Diagnosis — Median suture line of Cx-II+III short; Cx-IV extended posterior to genital field in both sexes; genital field L/W ratio
<quantity id="0651BDF9FF83CF28DA5CFA67FA7AA735" box="[1289,1428,1522,1546]" metricMagnitude="-2" metricUnit="m" metricValue="2.7685999999999997" metricValueMax="2.8701999999999996" metricValueMin="2.6670000000000003" pageId="7" pageNumber="157" unit="in" value="1.0899999999999999" valueMax="1.13" valueMin="1.05">1.05-1.13 in</quantity>
male; P-2 ventral projection curved distally; capitulum with a long rostrum; postgenital area in the male well developed, ratio postgenital L/idiosoma L 28.8-29.0.
</paragraph>
<paragraph id="C116101CFF83CF28DC35F93FFC07A7FD" blockId="7.[826,1429,1190,1949]" box="[864,1001,1706,1730]" pageId="7" pageNumber="157">Description</paragraph>
<paragraph id="C116101CFF83CF26DC35F940FDC4A6A8" blockId="7.[826,1429,1190,1949]" lastBlockId="9.[159,762,1323,1943]" lastPageId="9" lastPageNumber="159" pageId="7" pageNumber="157">
Male — (
<typeStatus id="1E12AEBEFF83CF28DC92F940FBDCA7D2" box="[967,1074,1749,1773]" pageId="7" pageNumber="157" type="holotype">holotype</typeStatus>
, in parentheses measurements of specimens from Wadi Ban Auf, n=2). Idiosoma (ventral view:
<figureCitation id="59920C99FF83CF28DCB2F88EFBD2A60C" box="[999,1084,1819,1843]" captionStart="FIGURE" captionStartId="8.[159,171,1888,1907]" captionTargetBox="[253,1361,311,1857]" captionTargetId="figure-7@8.[227,1361,307,1857]" captionTargetPageId="8" captionText="FIGURE 4: Torrenticola arabica n. sp., male (A-D, F holotype; E, G specimen from Wadi Ban Auy): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view; (D-E) – palp, lateral view; (F) – capitulum; (G) – ejaculatory complex. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404092" httpUri="https://zenodo.org/record/5404092/files/figure.png" pageId="7" pageNumber="157">Fig. 4B</figureCitation>
) L 759 (694-747), W 588 (556- 637); dorsal shield (
<figureCitation id="59920C99FF83CF28DB74F8ABFB92A668" box="[1057,1148,1854,1879]" captionStart="FIGURE" captionStartId="8.[159,171,1888,1907]" captionTargetBox="[253,1361,311,1857]" captionTargetId="figure-7@8.[227,1361,307,1857]" captionTargetPageId="8" captionText="FIGURE 4: Torrenticola arabica n. sp., male (A-D, F holotype; E, G specimen from Wadi Ban Auy): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view; (D-E) – palp, lateral view; (F) – capitulum; (G) – ejaculatory complex. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404092" httpUri="https://zenodo.org/record/5404092/files/figure.png" pageId="7" pageNumber="157">Fig. 4A</figureCitation>
) L 634 (606-675), W 456 (425-475), L/W ratio 1.39 (1.42-1.43); dorsal plate 606 (581-644); shoulder plate L 194 (184-201), W 63 (59-66), L/W ratio 3.1 (3.1); frontal plate L 133 (131- 136), W 53 (44-50), L/W ratio 2.5 (2.7-3.0); shoul- der/frontal plate L ratio 1.46 (1.4-1.5); gnathosomal bay L 167 (135-159), Cx-I total L 314 (256-291), Cx-I medial L 147 (119-128), Cx-II+III medial 86 (86-92); ratio Cx-I L/Cx-II+III medial L 3.65 (2.8-3.4); Cx- I medial L/Cx-II+III medial L 1.7 (1.3-1.5); genital field L/W 141 (139-144)/133 (127-128), L/W ratio 1.05 (1.1-1.13), ejaculatory complex (
<figureCitation id="59920C99FF8DCF26DD18F9D1FD44A763" box="[589,682,1604,1628]" captionStart="FIGURE" captionStartId="8.[159,171,1888,1907]" captionTargetBox="[253,1361,311,1857]" captionTargetId="figure-7@8.[227,1361,307,1857]" captionTargetPageId="8" captionText="FIGURE 4: Torrenticola arabica n. sp., male (A-D, F holotype; E, G specimen from Wadi Ban Auy): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view; (D-E) – palp, lateral view; (F) – capitulum; (G) – ejaculatory complex. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404092" httpUri="https://zenodo.org/record/5404092/files/figure.png" pageId="9" pageNumber="159">Fig. 4G</figureCitation>
) L 197 (184-197); distance genital field-excretory pore 186 (153-166), genital field-caudal idiosoma margin 219 (200-216); capitulum (
<figureCitation id="59920C99FF8DCF26DEF2F938FDEAA7FA" box="[423,516,1709,1733]" captionStart="FIGURE" captionStartId="8.[159,171,1888,1907]" captionTargetBox="[253,1361,311,1857]" captionTargetId="figure-7@8.[227,1361,307,1857]" captionTargetPageId="8" captionText="FIGURE 4: Torrenticola arabica n. sp., male (A-D, F holotype; E, G specimen from Wadi Ban Auy): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view; (D-E) – palp, lateral view; (F) – capitulum; (G) – ejaculatory complex. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404092" httpUri="https://zenodo.org/record/5404092/files/figure.png" pageId="9" pageNumber="159">Fig. 4F</figureCitation>
) ventral L 325 (306- 317); palp (
<figureCitation id="59920C99FF8DCF26DE76F945FE9AA7D7" box="[291,372,1744,1768]" captionStart="FIGURE" captionStartId="8.[159,171,1888,1907]" captionTargetBox="[253,1361,311,1857]" captionTargetId="figure-7@8.[227,1361,307,1857]" captionTargetPageId="8" captionText="FIGURE 4: Torrenticola arabica n. sp., male (A-D, F holotype; E, G specimen from Wadi Ban Auy): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view; (D-E) – palp, lateral view; (F) – capitulum; (G) – ejaculatory complex. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404092" httpUri="https://zenodo.org/record/5404092/files/figure.png" pageId="9" pageNumber="159">Figs. 4</figureCitation>
C-E) total L 346 (332-343), L: P-1, 32 (31-32); P-2, 123 (112-122); P-3, 63 (62-63); P-4, 108 (105-106); P-5, 20 (20-22); %L: P-1, 9.2 (9.3); P-2, 35.6 (33.8-35.6); P-3, 18.2 (18.4-18.7); P-4, 31.2 (30.9-31.6); P-5, 5.8 (5.8-6.6); L P-2/P-4 ratio 1.14 (1.07-1.15); P-2 ventral projection curved distally.
</paragraph>
<caption id="95D64094FF8CCF27DFCAF8F5FA98A6B0" ID-DOI="http://doi.org/10.5281/zenodo.5404092" ID-Zenodo-Dep="5404092" httpUri="https://zenodo.org/record/5404092/files/figure.png" pageId="8" pageNumber="158" startId="8.[159,171,1888,1907]" targetBox="[253,1361,311,1857]" targetPageId="8">
<paragraph id="C116101CFF8CCF27DFCAF8F5FA98A6B0" blockId="8.[159,1429,1888,1935]" pageId="8" pageNumber="158">
FIGURE 4:
<taxonomicName id="06A96B9FFF8CCF27DE5CF8F5FE5DA64C" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[265,435,1888,1907]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="158" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="F3DDCC0EFF8CCF27DE5CF8F5FE5DA64C" box="[265,435,1888,1907]" italics="true" pageId="8" pageNumber="158">Torrenticola arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF8CCF27DEEFF8F5FE1AA64C" box="[442,500,1888,1907]" pageId="8" pageNumber="158" rank="species">n. sp.</taxonomicNameLabel>
, male (A-D, F holotype; E, G specimen from Wadi Ban Auy): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view; (D-E) – palp, lateral view; (F) – capitulum; (G) – ejaculatory complex. Scale bars = 100 µm.
</paragraph>
</caption>
<caption id="95D64094FF8DCF26DE52FB42FAC0A5D5" ID-DOI="http://doi.org/10.5281/zenodo.5404094" ID-Zenodo-Dep="5404094" box="[263,1326,1239,1258]" httpUri="https://zenodo.org/record/5404094/files/figure.png" pageId="9" pageNumber="159" startId="9.[263,275,1239,1258]" targetBox="[198,1355,322,1178]" targetPageId="9">
<paragraph id="C116101CFF8DCF26DE52FB42FAC0A5D5" blockId="9.[263,1326,1239,1258]" box="[263,1326,1239,1258]" pageId="9" pageNumber="159">
FIGURE 5:
<taxonomicName id="06A96B9FFF8DCF26DE39FB42FDFBA5D5" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[364,533,1239,1258]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="159" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="F3DDCC0EFF8DCF26DE39FB42FDFBA5D5" box="[364,533,1239,1258]" italics="true" pageId="9" pageNumber="159">Torrenticola arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF8DCF26DD4EFB42FDBEA5D5" box="[539,592,1239,1258]" pageId="9" pageNumber="159" rank="species">n. sp.</taxonomicNameLabel>
, female: (A) – dorsal shield; (B) – idiosoma, ventral view. Scale bars = 100 µm.
</paragraph>
</caption>
<paragraph id="C116101CFF8DCF24DC35FABEFD6FA04D" blockId="9.[826,1429,1323,1943]" lastBlockId="11.[159,762,311,1170]" lastPageId="11" lastPageNumber="161" pageId="9" pageNumber="159">
Female — (
<typeStatus id="1E12AEBEFF8DCF26DCBAFABEFBB5A47C" box="[1007,1115,1323,1347]" pageId="9" pageNumber="159" type="paratype">paratype</typeStatus>
, in parentheses measurements of specimen from Wadi Ban Auf) similar to the male. Ventral shield (
<figureCitation id="59920C99FF8DCF26DB26FAE4FB3FA4B6" box="[1139,1233,1393,1417]" captionStart="FIGURE" captionStartId="9.[263,275,1239,1258]" captionTargetBox="[198,1355,322,1178]" captionTargetId="figure-4@9.[189,1399,302,1207]" captionTargetPageId="9" captionText="FIGURE 5: Torrenticola arabica n. sp., female: (A) – dorsal shield; (B) – idiosoma, ventral view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404094" httpUri="https://zenodo.org/record/5404094/files/figure.png" pageId="9" pageNumber="159">Fig. 5B</figureCitation>
) L 856 (813), W 669 (675); dorsal shield (
<figureCitation id="59920C99FF8DCF26DB02FA01FB41A493" box="[1111,1199,1428,1452]" captionStart="FIGURE" captionStartId="9.[263,275,1239,1258]" captionTargetBox="[198,1355,322,1178]" captionTargetId="figure-4@9.[189,1399,302,1207]" captionTargetPageId="9" captionText="FIGURE 5: Torrenticola arabica n. sp., female: (A) – dorsal shield; (B) – idiosoma, ventral view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404094" httpUri="https://zenodo.org/record/5404094/files/figure.png" pageId="9" pageNumber="159">Fig. 5A</figureCitation>
) L 772 (719), W 525 (522), L/W ratio 1.47 (1.38); dorsal plate 734 (688); shoulder plate L 216 (222), W 75 (73), L/W ratio 2.9 (3.0); frontal plate L 166 (153), W 63 (56), L/W ratio 2.6 (2.7); shoulder/frontal plate L ratio 1.3 (1.45); gnathosomal bay L 184 (169), Cx-I total L 328 (293), Cx-I medial L 142 (125), Cx-II+III medial L 44 (56); ratio Cx-I L/Cx-II+III medial L 7.5 (5.2); Cx-I medial L/Cx-II+III medial L 3.2 (2.2); genital field L/W 164 (159)/159 (163), L/W ratio 1.03 (0.98); distance genital field-excretory pore 247 (230), genital fieldcaudal idiosoma margin 316 (292); capitulum ventral L 363 (364); palp total L 384 (391), L: P-1, 35 (34); P-2, 132 (139); P-3, 71 (71); P-4, 123 (122); P- 5, 23 (25); %L: P-1, 9.1 (8.7); P-2, 34.4 (35.6); P-3 18.5, (18.2); P-4, 32.0 (31.2); P-5, 6.0 (6.4); L P-2/P-4 ratio 1.07 (1.14); shape and setation as in male.
</paragraph>
<caption id="95D64094FF8ECF25DFCAF8F5FD5AA6B0" ID-DOI="http://doi.org/10.5281/zenodo.5404096" ID-Zenodo-Dep="5404096" httpUri="https://zenodo.org/record/5404096/files/figure.png" pageId="10" pageNumber="160" startId="10.[159,171,1888,1907]" targetBox="[272,1292,313,1849]" targetPageId="10">
<paragraph id="C116101CFF8ECF25DFCAF8F5FD5AA6B0" blockId="10.[159,1429,1888,1935]" pageId="10" pageNumber="160">
FIGURE 6:
<taxonomicName id="06A96B9FFF8ECF25DE51F8F5FE24A64D" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[260,458,1888,1907]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="160" phylum="Arthropoda" rank="species" species="omanensis" status="sp. nov.">
<emphasis id="F3DDCC0EFF8ECF25DE51F8F5FE24A64D" box="[260,458,1888,1907]" italics="true" pageId="10" pageNumber="160">Torrenticola omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF8ECF25DE9BF8F5FDEDA64C" box="[462,515,1888,1907]" pageId="10" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
, male: (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view; (D) – palp, lateral view; (E) – ejaculatory complex. Scale bars = 100 µm.
</paragraph>
</caption>
<paragraph id="C116101CFF8FCF24DF90FE13FDBFA529" blockId="11.[159,762,311,1170]" pageId="11" pageNumber="161">
Remarks — Due to a ventral projection on P- 2 which curves distally, one (of 4) P-4 ventral setae long, capitulum with a long rostrum, similarly shaped ejaculatory complex and Cx-IV posteriorly extended well beyond genital field in both sexes,
<taxonomicName id="06A96B9FFF8FCF24DFBBFDA3FE2AA371" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[238,452,566,590]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="161" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="F3DDCC0EFF8FCF24DFBBFDA3FE2AA371" box="[238,452,566,590]" italics="true" pageId="11" pageNumber="161">Torrenticola arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF8FCF24DE9AFDA3FDF4A372" box="[463,538,566,589]" pageId="11" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
, closely resembles
<emphasis id="F3DDCC0EFF8FCF24DFCAFDCFFEF6A34E" box="[159,280,602,625]" italics="true" pageId="11" pageNumber="161">T. semisuta</emphasis>
(Halík, 1930), a species widespread from the Indian Himalayas to
<collectingCountry id="B9BE508CFF8FCF24DE8FFDE9FDAAA3AB" box="[474,580,636,660]" name="Malaysia" pageId="11" pageNumber="161">Malaysia</collectingCountry>
(
<bibRefCitation id="A5386DEDFF8FCF24DD0CFDE9FD1BA3AB" author="Wiles, P. R." box="[601,757,636,660]" pageId="11" pageNumber="161" pagination="191 - 236" refId="ref18570" refString="Wiles, P. R. 1997 b - Asian and Oriental Torrenticolidae Piersig, 1902 (Acari: Hydrachnidia: Lebertioidea): a revision of the family and descriptions of new species of Torrenticola Piersig and Pseudotorrenticola Walter, from Southeast Asia - J. Natur. Hist., 31: 191 - 236." type="journal article" year="1997">Wiles, 1997b</bibRefCitation>
; Pe˘si´c
<emphasis id="F3DDCC0EFF8FCF24DFB7FD35FEF6A388" box="[226,280,671,695]" italics="true" pageId="11" pageNumber="161">et al.</emphasis>
, 2007; Pe˘si´c and
<bibRefCitation id="A5386DEDFF8FCF24DEB6FD0AFD8CA388" author="Smit H." box="[483,610,671,695]" pageId="11" pageNumber="161" pagination="38 - 62" refId="ref18229" refString="Pe ˘ si'c, V., Smit H. 2009 - Water mites of the family Torrenticolidae Piersig, 1902 (Acari: Hydrachnidia) from Thailand, Part I. The genera Torrenticola Piersig, 1896, Neoatractides Lundblad, 1941 and Pseudotorrenticola Walter, 1906 - Zootaxa, 1982: 38 - 62." type="journal article" year="2009">Smit, 2009</bibRefCitation>
).
<emphasis id="F3DDCC0EFF8FCF24DD2CFD0AFF13A3E5" italics="true" pageId="11" pageNumber="161">
<taxonomicName id="06A96B9FFF8FCF24DD2CFD0AFD17A388" authorityName="Piersig" authorityYear="1896" box="[633,761,671,695]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="161" phylum="Arthropoda" rank="genus">Torrenticola</taxonomicName>
semisuta
</emphasis>
(for an analysis of diagnostic characters of this species see Pe˘si´c and
<bibRefCitation id="A5386DEDFF8FCF24DE84FD70FDA2A3C2" author="Smit H." box="[465,588,741,765]" pageId="11" pageNumber="161" pagination="38 - 62" refId="ref18229" refString="Pe ˘ si'c, V., Smit H. 2009 - Water mites of the family Torrenticolidae Piersig, 1902 (Acari: Hydrachnidia) from Thailand, Part I. The genera Torrenticola Piersig, 1896, Neoatractides Lundblad, 1941 and Pseudotorrenticola Walter, 1906 - Zootaxa, 1982: 38 - 62." type="journal article" year="2009">Smit, 2009</bibRefCitation>
) can be distinguished from
<taxonomicName id="06A96B9FFF8FCF24DE10FC9CFE5EA21F" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[325,432,776,800]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="161" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="F3DDCC0EFF8FCF24DE10FC9CFEBBA21F" box="[325,341,777,800]" italics="true" pageId="11" pageNumber="161">T</emphasis>
.
<emphasis id="F3DDCC0EFF8FCF24DE31FC9DFE5EA21F" box="[356,432,776,800]" italics="true" pageId="11" pageNumber="161">arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF8FCF24DEEDFC9CFE12A21F" box="[440,508,777,800]" pageId="11" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
, in the following features: Cxgl-4 located sub-apically (apically in
<taxonomicName id="06A96B9FFF8FCF24DDE1FCB9FF27A259" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="161" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="F3DDCC0EFF8FCF24DDE1FCB9FF27A259" italics="true" pageId="11" pageNumber="161">T. arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF8FCF24DF85FCDAFEFEA259" box="[208,272,846,870]" pageId="11" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
), male genital field more elongated (L/W 1.23-1.25;
<quantity id="0651BDF9FF8FCF24DE42FCE4FE70A2B6" box="[279,414,881,905]" metricMagnitude="-2" metricUnit="m" metricValue="2.7685999999999997" metricValueMax="2.8701999999999996" metricValueMin="2.6670000000000003" pageId="11" pageNumber="161" unit="in" value="1.0899999999999999" valueMax="1.13" valueMin="1.05">1.05-1.13 in</quantity>
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<emphasis id="F3DDCC0EFF8FCF24DEFCFCE7FDFAA2B6" box="[425,532,881,905]" italics="true" pageId="11" pageNumber="161">T. arabica</emphasis>
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), the excretory pore and Vgl-2 slightly shifted from the line of primary sclerotization in both sexes, and the shortened male postgenital area (18.8-22% of the total idiosoma L; in
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<emphasis id="F3DDCC0EFF8FCF24DE60FC6BFE73A529" box="[309,413,1022,1046]" italics="true" pageId="11" pageNumber="161">T. arabica</emphasis>
</taxonomicName>
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28-29%).
</paragraph>
<paragraph id="C116101CFF8FCF24DF90FBBFFE0DA55A" blockId="11.[159,762,311,1170]" pageId="11" pageNumber="161">Etymology — The species is named for its occurrence in Arabian Peninsula.</paragraph>
<paragraph id="C116101CFF8FCF24DF90FBEFFE22A5AD" blockId="11.[159,762,311,1170]" box="[197,460,1146,1170]" pageId="11" pageNumber="161">
Distribution —
<collectingCountry id="B9BE508CFF8FCF24DE29FBEFFE26A5AD" box="[380,456,1146,1170]" name="Oman" pageId="11" pageNumber="161">Oman</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

139
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<emphasis id="F3DDCC0EFF86CF2DDCCEF8BEFB44A67C" bold="true" box="[923,1194,1835,1859]" italics="true" pageId="2" pageNumber="152">Hydrodroma rheophila</emphasis>
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examined — 1/2/0,
<collectingMunicipality id="21728A66FF86CF2DDB92F8F7FABAA646" box="[1223,1364,1889,1914]" pageId="2" pageNumber="152">Ayn Tabraq</collectingMunicipality>
, captured spring,
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,
<date id="B51736DCFF86CF2DDA71F811FA7EA6A3" box="[1316,1424,1924,1948]" pageId="2" pageNumber="152" value="2008-11-07">
<collectingDate id="A553CF34FF86CF2DDA71F811FA7EA6A3" box="[1316,1424,1924,1948]" pageId="2" pageNumber="152" value="2008-11-07">7-xi-2008</collectingDate>
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;
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5/2/0,
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, stream,
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; 1/0/0, stream
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, alt.
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0/1/0,
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,
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FIGURE 1:
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<emphasis id="F3DDCC0EFF87CF2CDE50F8F5FE53A64C" box="[261,445,1888,1907]" italics="true" pageId="3" pageNumber="153">Bharatavolzia arabica</emphasis>
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, female: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – palp; (D) – I-Leg-4-6; (E) – IV-Leg-4-6. Scale bars = 200 µm (A-B), 50 µm (C-E).
</paragraph>
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<paragraph id="C116101CFF80CF2BDF90FE58FF01A337" blockId="4.[159,762,311,520]" pageId="4" pageNumber="154">
Distribution —
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,
<collectingCountry id="B9BE508CFF80CF2BDE98FE58FDADA0DA" box="[461,579,461,485]" name="Indonesia" pageId="4" pageNumber="154">Indonesia</collectingCountry>
,
<collectingCountry id="B9BE508CFF80CF2BDD04FE5BFD6AA0DA" box="[593,644,462,485]" name="Iran" pageId="4" pageNumber="154">Iran</collectingCountry>
. New for
<collectingCountry id="B9BE508CFF80CF2BDFCAFE65FF05A337" box="[159,235,496,520]" name="Oman" pageId="4" pageNumber="154">Oman</collectingCountry>
.
</paragraph>
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129
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<emphasis id="F3DDCC0EFF86CF2DDFE6FAC8FD4DA44A" bold="true" box="[179,675,1373,1397]" italics="true" pageId="2" pageNumber="152">Bharatavolzia (Bharatavolziella) arabica</emphasis>
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n. sp
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(
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A-E)
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Type series —
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<paragraph id="C116101CFF86CF2DDF90F9C3FE28A78B" blockId="2.[159,762,1506,1948]" pageId="2" pageNumber="152">Diagnosis — Gnathosoma slender in ventral view; P-4, in addition to the medial seta, with a heavy anteroventral seta.</paragraph>
<paragraph id="C116101CFF86CF2DDF90F95FFEA0A7DD" blockId="2.[159,762,1506,1948]" box="[197,334,1738,1762]" pageId="2" pageNumber="152">Description</paragraph>
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Female — Idiosoma dorsally L 802, W 520, with a large anteromedial and a large posteromedial plate and five pairs of smaller lateral plates. Eyes absent. Anteromedial plate L 263, W 336, anteriorly with a rounded extension. Posteromedial plate L 482, W 300. Large anterolateral plate L 243, W 102; large posterolateral plate L 251, W 70. All dorsal plates and platelets without glandularia, but the associated setae present. Idiosoma ventrally L 867, W 502. Suture lines of coxal plates indistinct. Genital field placed between first coxae, L 57, W 94. Gnathosoma slender in ventral view. Palp (
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): total L 236; L: P-1, 8; P-2, 84; P-3, 52; P-4, 60; P-5, 32; P-4 anteroventrally with a heavy seta, medial side of P-4 also with a stout seta. L of I-Leg-4-6: 76, 96, 84 (120 to tip of segment). I-Leg-6 anterodorsally with a heavy seta, I-Leg-4 and I-Leg-5 anteroventrally with 1-3 setae. L of IV-Leg-4-6: 96, 106, 88 (140 to tip of segment). IV-Leg-6 anterodorsally with two heavy setae close to each other. IV-Leg-4 and -5 anteroventrally with a group of setae, one of these setae of IV-Leg-5 with large pectinations. All legs with large claws.
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<paragraph id="C116101CFF86CF2DDC35FC4AFC9FA525" blockId="2.[826,1429,311,1688]" pageId="2" pageNumber="152">Etymology — Named after the Arabian Peninsula.</paragraph>
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Remarks — This is the fourth known species of the genus
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, two species are known from
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(
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), while one species is known from
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(
<bibRefCitation id="A5386DEDFF86CF2DDC91FB00FA67A592" author="Schwoerbel, J. &amp; Sepasgosarian H." box="[964,1417,1173,1197]" pageId="2" pageNumber="152" pagination="293 - 301" refId="ref18352" refString="Schwoerbel, J., Sepasgosarian H. 1980 - Wassermilben (Acari, Prostigmata, Hydrachnellae) aus dem Iran - Entom. Mitt. Zool. Mus. Hamburg, 6 (107): 293 - 301." type="journal article" year="1980">Schwoerbel and Sepasgosarian, 1980</bibRefCitation>
).
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erected the subgenus
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for species without eye capsules. The new species differs from the two other known species of this subgenus in the slender gnathosoma and the presence of two heavy setae on P-4 instead of one or none. Moreover, compared to the Indian
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Cook the new species has a larger anterior extension of the anteromedial dorsal plate, and compared to
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Schwoerbel and Sepasgosarian from
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the large anterolateral plate is less slender. The large posterolateral platelet of the new species is much longer than the smaller platelet posterior to it, but in
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these platelets are much less different in size.
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<subSubSection id="89B34397FF88CF23DFAFF972FD70A7C0" box="[250,670,1767,1791]" pageId="12" pageNumber="162" type="nomenclature">
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<taxonomicName id="06A96B9FFF88CF23DFAFF972FD70A7C0" authority="Gerecke, 2004" authorityName="Gerecke" authorityYear="2004" box="[250,670,1767,1791]" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="162" phylum="Arthropoda" rank="species" species="oman">
<emphasis id="F3DDCC0EFF88CF23DFAFF972FD70A7C0" bold="true" box="[250,670,1767,1791]" pageId="12" pageNumber="162">
<emphasis id="F3DDCC0EFF88CF23DFAFF972FE1FA7C1" bold="true" box="[250,497,1767,1791]" italics="true" pageId="12" pageNumber="162">Monatractides oman</emphasis>
<bibRefCitation id="A5386DEDFF88CF23DEADF972FD70A7C0" author="Gerecke, R." box="[504,670,1767,1791]" pageId="12" pageNumber="162" pagination="2833 - 2849" refId="ref17893" refString="Gerecke, R. 2004 - Water mites from spring- and groundwater habitats in the Sultanate of Oman - J. Nat. Hist., 38: 2833 - 2849." type="journal article" year="2004">Gerecke, 2004</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89B34397FF88CF22DE20F88AFB89A337" lastPageId="13" lastPageNumber="163" pageId="12" pageNumber="162" type="description">
<paragraph id="C116101CFF88CF23DE20F88AFDCDA608" blockId="12.[373,547,1823,1847]" box="[373,547,1823,1847]" pageId="12" pageNumber="162">
(
<figureCitation id="59920C99FF88CF23DE28F88AFE06A608" box="[381,488,1823,1847]" captionStart="FIGURE" captionStartId="12.[159,171,1523,1542]" captionTargetBox="[189,1399,302,1491]" captionTargetId="figure-7@12.[189,1399,302,1491]" captionTargetPageId="12" captionText="FIGURE 8: Monatractides oman Gerecke, 2004, female: (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404100" httpUri="https://zenodo.org/record/5404100/files/figure.png" pageId="12" pageNumber="162">Figures 8</figureCitation>
A-C)
</paragraph>
<paragraph id="C116101CFF88CF22DF90F8F4FDC1A087" blockId="12.[159,761,1889,1948]" lastBlockId="13.[159,762,311,1949]" lastPageId="13" lastPageNumber="163" pageId="12" pageNumber="162">
<materialsCitation id="71C11A41FF88CF23DF90F8F4FBBAA744" collectingDate="2008-11-13" elevation="571" latitude="23.278316" location="Material" longLatPrecision="1" longitude="57.4615" municipality="Wadi Bani Auf" pageId="12" pageNumber="162" specimenCount="1">
<location id="C47646C7FF88CF23DF90F8F4FEC6A646" LSID="urn:lsid:plazi:treatment:4900A10AFF88CF22DFAFF972FB89A337:C47646C7FF88CF23DF90F8F4FEC6A646" box="[197,296,1889,1913]" latitude="23.278316" longLatPrecision="1" longitude="57.4615" municipality="Wadi Bani Auf" name="Material" pageId="12" pageNumber="162">Material</location>
examined — 4/5/1 (1/1/0 mounted), stream
<collectingMunicipality id="21728A66FF88CF23DFAEF811FE59A6A3" box="[251,439,1924,1948]" pageId="12" pageNumber="162">Wadi Bani Auf</collectingMunicipality>
,
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<geoCoordinate id="A49D76DBFF88CF23DD3DF811FD1BA6A3" box="[616,757,1924,1948]" degrees="57" direction="east" minutes="27.690" orientation="longitude" pageId="12" pageNumber="162" precision="1" value="57.4615">57°27.690 E</geoCoordinate>
, alt.
<quantity id="0651BDF9FF88CF23DC2CF9F6FC29A745" box="[889,967,1635,1659]" metricMagnitude="2" metricUnit="m" metricValue="5.71" pageId="12" pageNumber="162" unit="m" value="571.0">
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,
<date id="B51736DCFF88CF23DC8FF9F6FBBAA744" box="[986,1108,1635,1659]" pageId="12" pageNumber="162" value="2008-11-13">
<collectingDate id="A553CF34FF88CF23DC8FF9F6FBBAA744" box="[986,1108,1635,1659]" pageId="12" pageNumber="162" value="2008-11-13">13-xi-2008</collectingDate>
</date>
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;
<materialsCitation id="71C11A41FF88CF23DB3DF9F6FC28A7FE" collectingDate="2008-11-07" latitude="17.100567" location="Ayn Tabraq" longLatPrecision="1" longitude="54.32665" pageId="12" pageNumber="162" specimenCount="1">
0/2/0 (0/1/0 mounted),
<location id="C47646C7FF88CF23DC6FF913FC2AA7A1" LSID="urn:lsid:plazi:treatment:4900A10AFF88CF22DFAFF972FB89A337:C47646C7FF88CF23DC6FF913FC2AA7A1" box="[826,964,1670,1694]" latitude="17.100567" longLatPrecision="1" longitude="54.32665" name="Ayn Tabraq" pageId="12" pageNumber="162">Ayn Tabraq</location>
, captured spring,
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<geoCoordinate id="A49D76DBFF88CF23DA73F913FCA2A7FF" degrees="54" direction="east" minutes="19.599" orientation="longitude" pageId="12" pageNumber="162" precision="1" value="54.32665">54°19.599 E</geoCoordinate>
,
<date id="B51736DCFF88CF23DC0FF93CFC28A7FE" box="[858,966,1705,1729]" pageId="12" pageNumber="162" value="2008-11-07">
<collectingDate id="A553CF34FF88CF23DC0FF93CFC28A7FE" box="[858,966,1705,1729]" pageId="12" pageNumber="162" value="2008-11-07">7-xi-2008</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71C11A41FF88CF23DC81F93CFAD9A7DB" collectingDate="2008-11-07" latitude="17.100567" location="Ayn Tabraq" longLatPrecision="1" longitude="54.32665" pageId="12" pageNumber="162" specimenCount="1">
3/1/0 (1/0/0 mounted),
<location id="C47646C7FF88CF23DA51F93CFA7FA7FE" LSID="urn:lsid:plazi:treatment:4900A10AFF88CF22DFAFF972FB89A337:C47646C7FF88CF23DA51F93CFA7FA7FE" box="[1284,1425,1705,1729]" latitude="17.100567" longLatPrecision="1" longitude="54.32665" name="Ayn Tabraq" pageId="12" pageNumber="162">Ayn Tabraq</location>
, stream,
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<geoCoordinate id="A49D76DBFF88CF23DB67F959FB52A7DC" box="[1074,1212,1740,1764]" degrees="54" direction="east" minutes="19.599" orientation="longitude" pageId="12" pageNumber="162" precision="1" value="54.32665">54°19.599 E</geoCoordinate>
,
<date id="B51736DCFF88CF23DB9EF959FAD9A7DB" box="[1227,1335,1740,1764]" pageId="12" pageNumber="162" value="2008-11-07">
<collectingDate id="A553CF34FF88CF23DB9EF959FAD9A7DB" box="[1227,1335,1740,1764]" pageId="12" pageNumber="162" value="2008-11-07">7-xi-2008</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71C11A41FF88CF23DA13F959FBD1A615" collectingDate="2008-11-03" latitude="17.097366" location="Ayn Hamran" longLatPrecision="1" longitude="54.281433" pageId="12" pageNumber="162" specimenCount="1">
9/9/1,
<location id="C47646C7FF88CF23DC6FF97AFC09A638" LSID="urn:lsid:plazi:treatment:4900A10AFF88CF22DFAFF972FB89A337:C47646C7FF88CF23DC6FF97AFC09A638" box="[826,999,1775,1799]" latitude="17.097366" longLatPrecision="1" longitude="54.281433" name="Ayn Hamran" pageId="12" pageNumber="162">Ayn Hamran</location>
, captured spring,
<geoCoordinate id="A49D76DBFF88CF23DBADF97AFA7BA638" box="[1272,1429,1775,1799]" degrees="17" direction="north" minutes="05.842" orientation="latitude" pageId="12" pageNumber="162" precision="1" value="17.097366">17°05.842 N</geoCoordinate>
<geoCoordinate id="A49D76DBFF88CF23DC6FF887FC2AA615" box="[826,964,1810,1834]" degrees="54" direction="east" minutes="16.886" orientation="longitude" pageId="12" pageNumber="162" precision="1" value="54.281433">54°16.886 E</geoCoordinate>
,
<date id="B51736DCFF88CF23DC86F887FBD1A615" box="[979,1087,1810,1834]" pageId="12" pageNumber="162" value="2008-11-03">
<collectingDate id="A553CF34FF88CF23DC86F887FBD1A615" box="[979,1087,1810,1834]" pageId="12" pageNumber="162" value="2008-11-03">3-xi-2008</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71C11A41FF88CF23DB1AF887FA7FA672" collectingDate="2008-11-15" elevation="421" latitude="24.686083" location="Hatta Pools" longLatPrecision="1" longitude="56.159267" pageId="12" pageNumber="162" specimenCount="1">
0/1/0, stream
<location id="C47646C7FF88CF23DA57F886FA7EA615" LSID="urn:lsid:plazi:treatment:4900A10AFF88CF22DFAFF972FB89A337:C47646C7FF88CF23DA57F886FA7EA615" box="[1282,1424,1810,1834]" latitude="24.686083" longLatPrecision="1" longitude="56.159267" name="Hatta Pools" pageId="12" pageNumber="162">Hatta Pools</location>
,
<geoCoordinate id="A49D76DBFF88CF23DC6FF8A0FC25A672" box="[826,971,1845,1869]" degrees="24" direction="north" minutes="41.165" orientation="latitude" pageId="12" pageNumber="162" precision="1" value="24.686083">24°41.165 N</geoCoordinate>
<geoCoordinate id="A49D76DBFF88CF23DC8DF8A0FB8BA672" box="[984,1125,1845,1869]" degrees="56" direction="east" minutes="09.556" orientation="longitude" pageId="12" pageNumber="162" precision="1" value="56.159267">56°09.556 E</geoCoordinate>
, alt.
<quantity id="0651BDF9FF88CF23DBE3F8A0FAEAA672" box="[1206,1284,1845,1869]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="12" pageNumber="162" unit="m" value="421.0">
<elevation id="4A84F72FFF88CF23DBE3F8A0FAEAA672" box="[1206,1284,1845,1869]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="12" pageNumber="162" unit="m" value="421.0">421 m</elevation>
</quantity>
,
<date id="B51736DCFF88CF23DA42F8A0FA7FA672" box="[1303,1425,1845,1869]" pageId="12" pageNumber="162" value="2008-11-15">
<collectingDate id="A553CF34FF88CF23DA42F8A0FA7FA672" box="[1303,1425,1845,1869]" pageId="12" pageNumber="162" value="2008-11-15">15-xi-2008</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71C11A41FF88CF23DC6FF8CDFBEFA6AC" collectingDate="2008-11-04" latitude="17.0706" location="Wadi Hanna" longLatPrecision="1" longitude="54.60815" pageId="12" pageNumber="162" specimenCount="1" specimenCount-juvenile="1">
2/0/0,
<location id="C47646C7FF88CF23DCC4F8CDFBC7A64F" LSID="urn:lsid:plazi:treatment:4900A10AFF88CF22DFAFF972FB89A337:C47646C7FF88CF23DCC4F8CDFBC7A64F" box="[913,1065,1880,1904]" latitude="17.0706" longLatPrecision="1" longitude="54.60815" name="Wadi Hanna" pageId="12" pageNumber="162">Wadi Hanna</location>
,
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<geoCoordinate id="A49D76DBFF88CF23DB9EF8CDFABDA64F" box="[1227,1363,1880,1904]" degrees="54" direction="east" minutes="36.489" orientation="longitude" pageId="12" pageNumber="162" precision="1" value="54.60815">54°36.489 E</geoCoordinate>
,
<date id="B51736DCFF88CF23DA35F8CDFC9CA6AC" pageId="12" pageNumber="162" value="2008-11-04">
<collectingDate id="A553CF34FF88CF23DA35F8CDFC9CA6AC" pageId="12" pageNumber="162" value="2008-11-04">4-xi- 2008</collectingDate>
</date>
;
<specimenCount id="D7AFDB95FF88CF23DC2AF8EEFBEFA6AC" box="[895,1025,1915,1939]" pageId="12" pageNumber="162" type="juvenile">1 juvenile</specimenCount>
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/
<materialsCitation id="71C11A41FF88CF22DB5EF8EEFD71A070" collectingDate="2008-11-12" lastPageId="13" lastPageNumber="163" latitude="23.1617" location="Wadi Ghul" longLatPrecision="1" longitude="57.20252" pageId="12" pageNumber="162" specimenCount="1">
0/1, small cascading stream
<location id="C47646C7FF88CF22DA02F8EEFF33A070" LSID="urn:lsid:plazi:treatment:4900A10AFF88CF22DFAFF972FB89A337:C47646C7FF88CF22DA02F8EEFF33A070" lastPageId="13" lastPageNumber="163" latitude="23.1617" longLatPrecision="1" longitude="57.20252" name="Wadi Ghul" pageId="12" pageNumber="162">Wadi Ghul</location>
,
<geoCoordinate id="A49D76DBFF89CF22DFBBFEA2FE90A071" box="[238,382,311,335]" degrees="23" direction="north" minutes="09.702" orientation="latitude" pageId="13" pageNumber="163" precision="1" value="23.1617">23°09.702 N</geoCoordinate>
<geoCoordinate id="A49D76DBFF89CF22DEDCFEA2FDFBA071" box="[393,533,311,335]" degrees="57" direction="east" minutes="12.151" orientation="longitude" pageId="13" pageNumber="163" precision="1" value="57.20252">57°12.151 E</geoCoordinate>
,
<date id="B51736DCFF89CF22DD73FEA2FD71A070" box="[550,671,311,335]" pageId="13" pageNumber="163" value="2008-11-12">
<collectingDate id="A553CF34FF89CF22DD73FEA2FD71A070" box="[550,671,311,335]" pageId="13" pageNumber="163" value="2008-11-12">12-xi-2008</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71C11A41FF89CF22DDE4FEA2FEC7A0AA" collectingDate="2008-11-03" latitude="17.130117" location="Ayn Razat" longLatPrecision="1" longitude="54.237183" pageId="13" pageNumber="163" specimenCount="1">
6/3/0 (1/0/0 mounted),
<location id="C47646C7FF89CF22DE23FECFFE1CA04E" LSID="urn:lsid:plazi:treatment:4900A10AFF88CF22DFAFF972FB89A337:C47646C7FF89CF22DE23FECFFE1CA04E" box="[374,498,346,370]" latitude="17.130117" longLatPrecision="1" longitude="54.237183" name="Ayn Razat" pageId="13" pageNumber="163">Ayn Razat</location>
,
<geoCoordinate id="A49D76DBFF89CF22DEAEFECFFD6BA04E" box="[507,645,346,370]" degrees="17" direction="north" minutes="07.807" orientation="latitude" pageId="13" pageNumber="163" precision="1" value="17.130117">17°07.807 N</geoCoordinate>
<geoCoordinate id="A49D76DBFF89CF22DDDEFECFFF5FA0AB" degrees="54" direction="east" minutes="14.231" orientation="longitude" pageId="13" pageNumber="163" precision="1" value="54.237183">54°14.231 E</geoCoordinate>
,
<date id="B51736DCFF89CF22DFE8FEE8FEC7A0AA" box="[189,297,381,405]" pageId="13" pageNumber="163" value="2008-11-03">
<collectingDate id="A553CF34FF89CF22DFE8FEE8FEC7A0AA" box="[189,297,381,405]" pageId="13" pageNumber="163" value="2008-11-03">3-xi-2008</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71C11A41FF89CF22DE61FEE8FDC5A087" collectingDate="2008-11-12" elevation="805" latitude="23.171618" location="Wadi Ghul" longLatPrecision="1" longitude="57.199932" pageId="13" pageNumber="163" specimenCount="1">
3/8/0, stream
<location id="C47646C7FF89CF22DEB5FEE8FD8DA0AA" LSID="urn:lsid:plazi:treatment:4900A10AFF88CF22DFAFF972FB89A337:C47646C7FF89CF22DEB5FEE8FD8DA0AA" box="[480,611,381,405]" latitude="23.171618" longLatPrecision="1" longitude="57.199932" name="Wadi Ghul" pageId="13" pageNumber="163">Wadi Ghul</location>
,
<geoCoordinate id="A49D76DBFF89CF22DD3BFEE8FD17A0AB" box="[622,761,381,405]" degrees="23" direction="north" minutes="10.297" orientation="latitude" pageId="13" pageNumber="163" precision="1" value="23.171618">23°10.297 N</geoCoordinate>
<geoCoordinate id="A49D76DBFF89CF22DFCAFE35FEC8A088" box="[159,294,416,440]" degrees="57" direction="east" minutes="11.996" orientation="longitude" pageId="13" pageNumber="163" precision="1" value="57.199932">57°11.996 E</geoCoordinate>
, alt.
<quantity id="0651BDF9FF89CF22DE0BFE35FE4BA088" box="[350,421,416,440]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="13" pageNumber="163" unit="m" value="805.0">
<elevation id="4A84F72FFF89CF22DE0BFE35FE4BA088" box="[350,421,416,440]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="13" pageNumber="163" unit="m" value="805.0">805 m</elevation>
</quantity>
,
<date id="B51736DCFF89CF22DEE4FE35FDC5A087" box="[433,555,416,440]" pageId="13" pageNumber="163" value="2008-11-12">
<collectingDate id="A553CF34FF89CF22DEE4FE35FDC5A087" box="[433,555,416,440]" pageId="13" pageNumber="163" value="2008-11-12">12-xi-2008</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="C116101CFF89CF22DF90FE59FEA0A0DB" blockId="13.[159,762,311,1949]" box="[197,334,460,484]" pageId="13" pageNumber="163">Description</paragraph>
<paragraph id="C116101CFF89CF22DF90FE6DFD5CA5F2" blockId="13.[159,762,311,1949]" pageId="13" pageNumber="163">Male — (from Wadi Bani Auf, in parentheses specimen from Ayn Tabraq). Idiosoma L 881 (963), W 650 (788); dorsal shield L 734 (794), W 500 (619), L/W ratio 1.47 (1.28); dorsal plate L 675 (719); shoulder plate L 239 (255), W 84 (94), L/W ratio 2.9 (2.7); frontal plate L 126 (119), W 81 (94) ratio 1.6 (1.3); shoulder/frontal plate L ratio 1.9 (2.1); capitular bay L 164 (189); total L 267 (319), Cx-I medial L 103 (131), Cx-II+III medial 128 (125); ratio Cx-I L/Cx-II+III medial L 2.1 (2.6); Cx-I medial L/Cx-II+III medial L 0.8 (1.05); genital field L/W 144 (164)/120 (141), L/W ratio 1.2 (1.16); ejaculatory complex L 173 (219); distance genital field-excretory pore 188 (205), genital field-caudal idiosoma margin 328 (344); capitulum ventral L 205 (218); chelicera L 235 (270), basal segment L/H (222/33), claw L (49), basal segment/claw ratio (4.5), L/H ratio (6.7); palp: total L 227 (250), L: P-1, 26 (29); P-2, 72 (83); P-3, 43 (47); P-4, 58 (63); P-5, 28 (28); %L: P-1, 11.5 (11.6); P-2, 31.7 (33.2); P-3, 18.9 (18.8); P-4, 25.6 (25.2); P-5, 12.3 (11.2); P-2/P-4 ratio 1.24 (1.3).</paragraph>
<paragraph id="C116101CFF89CF22DF90FB74FD17A7DB" blockId="13.[159,762,311,1949]" pageId="13" pageNumber="163">
Female — (from Wadi Bani Auf). Idiosoma (ventral view:
<figureCitation id="59920C99FF89CF22DE4CFA91FE80A423" box="[281,366,1284,1308]" captionStart="FIGURE" captionStartId="12.[159,171,1523,1542]" captionTargetBox="[189,1399,302,1491]" captionTargetId="figure-7@12.[189,1399,302,1491]" captionTargetPageId="12" captionText="FIGURE 8: Monatractides oman Gerecke, 2004, female: (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404100" httpUri="https://zenodo.org/record/5404100/files/figure.png" pageId="13" pageNumber="163">Fig. 8B</figureCitation>
) L 938, W 681; dorsal shield (
<figureCitation id="59920C99FF89CF22DD98FA91FF2FA47F" captionStart="FIGURE" captionStartId="12.[159,171,1523,1542]" captionTargetBox="[189,1399,302,1491]" captionTargetId="figure-7@12.[189,1399,302,1491]" captionTargetPageId="12" captionText="FIGURE 8: Monatractides oman Gerecke, 2004, female: (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404100" httpUri="https://zenodo.org/record/5404100/files/figure.png" pageId="13" pageNumber="163">Fig. 8A</figureCitation>
) L 744, W 538, L/W ratio 1.38; dorsal plate L 694; shoulder plate L 237, W 75, L/W ratio 3.16; frontal plate L 126, W 78, L/W ratio 1.6; shoulder/frontal plate L ratio 1.9; capitular bay L 175; Cx-I total L 280, Cx-I medial L 104, Cx-II+III medial 100; ratio Cx- I L/Cx-II+III medial L 2.8; Cx-I medial L/Cx-II+III medial L 1.04; genital field L/W 165/159, L/W ratio 1.04; egg maximum diameter L 206; distance genital field-excretory pore 213, genital field-caudal idiosoma margin 366; capitulum ventral L 209; chelicera L 243; palp (
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): total L 237, L and %L (in parentheses): P-1, 29 (12.2); P-2, 75 (31.6); P-3, 43 (18.1); P-4, 62 (26.2); P-5, 28 (11.8); P-2/P-4 ratio 1.2.
</paragraph>
<paragraph id="C116101CFF89CF22DF90F96DFC81A0E4" blockId="13.[159,762,311,1949]" lastBlockId="13.[826,1429,311,520]" pageId="13" pageNumber="163">
Remarks — The original description (
<bibRefCitation id="A5386DEDFF89CF22DDC6F96DFF38A60C" author="Gerecke, R." pageId="13" pageNumber="163" pagination="2833 - 2849" refId="ref17893" refString="Gerecke, R. 2004 - Water mites from spring- and groundwater habitats in the Sultanate of Oman - J. Nat. Hist., 38: 2833 - 2849." type="journal article" year="2004">Gerecke, 2004</bibRefCitation>
) of this species was based on a single male and
<specimenCount id="D7AFDB95FF89CF22DFCAF8AAFEDDA669" box="[159,307,1854,1878]" pageId="13" pageNumber="163" type="juvenile">two juvenile</specimenCount>
female specimens from Rustaq, Wadi Al Fara,
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(
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).
<bibRefCitation id="A5386DEDFF89CF22DDCEF8F4FF0BA6A2" author="Gerecke, R." pageId="13" pageNumber="163" pagination="2833 - 2849" refId="ref17893" refString="Gerecke, R. 2004 - Water mites from spring- and groundwater habitats in the Sultanate of Oman - J. Nat. Hist., 38: 2833 - 2849." type="journal article" year="2004">Gerecke (2004)</bibRefCitation>
stated that the juvenile females are probably atypical in idiosoma shape and therefore not included in the
<typeStatus id="1E12AEBEFF89CF22DC82FECFFBE5A04D" box="[983,1035,346,370]" pageId="13" pageNumber="163">type</typeStatus>
series. As no illustration has been given of the mature female we give additional figures here based on the specimen from Wadi Bani Auf.
</paragraph>
<paragraph id="C116101CFF89CF22DC35FE65FB89A337" blockId="13.[826,1429,311,520]" box="[864,1127,496,520]" pageId="13" pageNumber="163">
Distribution —
<collectingCountry id="B9BE508CFF89CF22DB42FE65FB8DA337" box="[1047,1123,496,520]" name="Oman" pageId="13" pageNumber="163">Oman</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

142
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<heading id="9A5EA770FF89CF22DCF3FD64FAC7A236" bold="true" box="[934,1321,753,777]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="163" reason="2">
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<taxonomicName id="06A96B9FFF89CF22DCF3FD64FB0EA236" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[934,1248,753,777]" class="Arachnida" family="Limnesiidae" genus="Limnesia" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="163" phylum="Arthropoda" rank="species" species="kochi" status="sp. nov." subGenus="Limnesia">
<emphasis id="F3DDCC0EFF89CF22DCF3FD64FB0EA236" bold="true" box="[934,1248,753,777]" italics="true" pageId="13" pageNumber="163">Limnesia (Limnesia) kochi</emphasis>
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<paragraph id="C116101CFF89CF22DB47FCBFFB52A27D" blockId="13.[1042,1212,810,834]" box="[1042,1212,810,834]" pageId="13" pageNumber="163">
(
<figureCitation id="59920C99FF89CF22DB4FFCBFFB68A27D" box="[1050,1158,810,834]" captionStart="FIGURE" captionStartId="14.[159,171,1888,1907]" captionTargetBox="[276,1336,326,1824]" captionTargetId="figure-7@14.[252,1336,307,1857]" captionTargetPageId="14" captionText="FIGURE 9: Limnesia kochi n. sp. (A-D: male, E: female): (A, E) – coxal and genital field; (B) – palp; (C) – I-Leg-5-6; (D) – IV-Leg-5-6. Scale bars = 50 µm (A-D)." figureDoi="http://doi.org/10.5281/zenodo.5404102" httpUri="https://zenodo.org/record/5404102/files/figure.png" pageId="13" pageNumber="163">Figures 9</figureCitation>
A-E)
</paragraph>
<paragraph id="C116101CFF89CF22DC35FCE5FBA4A509" blockId="13.[826,1429,879,1948]" pageId="13" pageNumber="163">
Type material —
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<typeStatus id="1E12AEBEFF89CF22DB18FCFAFB52A2B8" box="[1101,1212,879,903]" pageId="13" pageNumber="163" type="holotype">Holotype</typeStatus>
male, unnamed stream crossing road to
<location id="C47646C7FF89CF22DB30FC07FB59A294" LSID="urn:lsid:plazi:treatment:4900A10AFF89CF20DCF3FD64FE43A55C:C47646C7FF89CF22DB30FC07FB59A294" box="[1125,1207,914,939]" country="Oman" latitude="24.64505" longLatPrecision="1" longitude="56.112667" municipality="Ray" name="Al Fay" pageId="13" pageNumber="163">Al Fay</location>
, near
<collectingMunicipality id="21728A66FF89CF22DA52FC06FADAA294" box="[1287,1332,915,939]" pageId="13" pageNumber="163">Ray</collectingMunicipality>
,
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,
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, alt.
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,
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<collectingDate id="A553CF34FF89CF22DA42FC20FA7FA2F2" box="[1303,1425,949,973]" pageId="13" pageNumber="163" value="2008-11-15">15-xi-2008</collectingDate>
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.
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<typeStatus id="1E12AEBEFF89CF22DC6FFC4CFC5CA2CE" box="[826,946,985,1009]" pageId="13" pageNumber="163" type="paratype">Paratypes</typeStatus>
: 2/1/3, same data as holotype; 2/0/0, stream
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<paragraph id="C116101CFF89CF22DC35FBDEFBE5A5F3" blockId="13.[826,1429,879,1948]" pageId="13" pageNumber="163">Diagnosis — Ventral setal tubercle of P-2 located far anteriorly, Glandula Limnesiae located near anterior margin of Cx-III; genital field of male indented anteriorly.</paragraph>
<paragraph id="C116101CFF89CF22DC35FB74FC07A5C6" blockId="13.[826,1429,879,1948]" box="[864,1001,1249,1273]" pageId="13" pageNumber="163">Description</paragraph>
<paragraph id="C116101CFF89CF22DC35FA98FB2BA6A3" blockId="13.[826,1429,879,1948]" pageId="13" pageNumber="163">
Male — Idiosoma soft, L 494 (462-616), W 401 (365-494). Integument lineated. Dorsum with a small posterior platelet,
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diameter. Cx-I with short apodemes, not fused medially. Cx-III medially with a narrow, secondary sclerotization. Glandulae Limnesiae located close to anterior margin of Cx-III. Genital field with three pairs of acetabula and 10 small setae, indented anteriorly and posteriorly, L 106, W 98, gonopore L 62, W 26. Palp (
<figureCitation id="59920C99FF89CF22DC17F9DCFC4DA75E" box="[834,931,1609,1633]" captionStart="FIGURE" captionStartId="14.[159,171,1888,1907]" captionTargetBox="[276,1336,326,1824]" captionTargetId="figure-7@14.[252,1336,307,1857]" captionTargetPageId="14" captionText="FIGURE 9: Limnesia kochi n. sp. (A-D: male, E: female): (A, E) – coxal and genital field; (B) – palp; (C) – I-Leg-5-6; (D) – IV-Leg-5-6. Scale bars = 50 µm (A-D)." figureDoi="http://doi.org/10.5281/zenodo.5404102" httpUri="https://zenodo.org/record/5404102/files/figure.png" pageId="13" pageNumber="163">Fig. 9B</figureCitation>
): total L 286; L: P-1, 18; P-2, 66; P-3, 66; P-4, 104; P-5, 32; P-2 with a ventral setal tubercle located far anteriorly, peg-like seta on tubercle small; P-4 with three small setal tubercles, two larger and one smaller. L of I-Leg-4-6: 76, 86, 72. L of IV- Leg-4-6: 124, 136, 152; IV-Leg-6 with a large distal seta and two smaller setae (
<figureCitation id="59920C99FF89CF22DBD7F88EFB31A60C" box="[1154,1247,1819,1843]" captionStart="FIGURE" captionStartId="14.[159,171,1888,1907]" captionTargetBox="[276,1336,326,1824]" captionTargetId="figure-7@14.[252,1336,307,1857]" captionTargetPageId="14" captionText="FIGURE 9: Limnesia kochi n. sp. (A-D: male, E: female): (A, E) – coxal and genital field; (B) – palp; (C) – I-Leg-5-6; (D) – IV-Leg-5-6. Scale bars = 50 µm (A-D)." figureDoi="http://doi.org/10.5281/zenodo.5404102" httpUri="https://zenodo.org/record/5404102/files/figure.png" pageId="13" pageNumber="163">Fig. 9D</figureCitation>
). IV-Leg-5 with four swimming setae, IV-Leg-4 with three swimming setae, III-Leg-4 and III-Leg-5 with three and two swimming setae respectively.
</paragraph>
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<paragraph id="C116101CFF8ACF21DFCAF8F5FE90A6B0" blockId="14.[159,1429,1888,1935]" pageId="14" pageNumber="164">
FIGURE 9:
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<emphasis id="F3DDCC0EFF8ACF21DE50F8F5FE6BA64C" box="[261,389,1888,1907]" italics="true" pageId="14" pageNumber="164">Limnesia kochi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF8ACF21DEDFF8F5FE51A64C" box="[394,447,1888,1907]" pageId="14" pageNumber="164" rank="species">n. sp.</taxonomicNameLabel>
(A-D: male, E: female): (A, E) – coxal and genital field; (B) – palp; (C) – I-Leg-5-6; (D) – IV-Leg-5-6. Scale bars = 50 µm (A-D).
</paragraph>
</caption>
<paragraph id="C116101CFF8BCF20DF90FEA2FD4EA228" blockId="15.[159,762,311,1123]" pageId="15" pageNumber="165">Female — Idiosoma soft, L 429, W 324. Female juvenile, no platelet visible. Cx-I with short apodemes, not fused medially. Cx-III medially with a narrow, secondary sclerotization. Glandulae Limnesiae located close to anterior margin of Cx-III. Genital field L 158, W 110, with three pairs of acetabula and 8-9 small setae, first pair of acetabula distanced from two posterior pairs. Pregenital sclerite slender. Palp: total L 324; L: P-1, 18; P-2, 90; P-3, 74; P-4, 110; P-5, 32; palp as in male. L of I-Leg-4-6: 78, 88, 66 (93 to tip). L of IV-Leg-4-6: 120, 140, 130. IV-Leg-5 with five swimming setae, IV-Leg-4 with two swimming setae, III-Leg-4 and III-Leg-5 with three and two swimming setae respectively.</paragraph>
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Etymology — Named after the German acarologist C.L. Koch (1778-1857), who erected the genus
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<emphasis id="F3DDCC0EFF8BCF20DFCAFCE4FEEEA2B7" box="[159,256,881,904]" italics="true" pageId="15" pageNumber="165">Limnesia</emphasis>
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.
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<paragraph id="C116101CFF8BCF20DF90FC0EFE43A55C" blockId="15.[159,762,311,1123]" pageId="15" pageNumber="165">Remarks — The location of the Glandulae Limnesiae is unusual within the genus. Its normal location is medial of the suture line of the Cx-III and Cx-IV. Moreover, the location of the setal tubercle of P-2 is also unusual, as it is normally located in the middle of the segment.</paragraph>
</subSubSection>
</treatment>
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A-C, E-G, 11A-C)
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0/5/0, &quot;
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, open well, diesel pump, UTM 0605256/2607586 (
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<geoCoordinate id="A49D76DBFF8BCF20DD66F9EAFD5BA7A8" box="[563,693,1663,1687]" degrees="58" direction="east" minutes="01" orientation="longitude" pageId="15" pageNumber="165" precision="15" seconds="53" value="58.031387">58°01’53”E</geoCoordinate>
),
<date id="B51736DCFF8BCF20DD83F9EAFF1AA785" pageId="15" pageNumber="165" value="1996-03-28">28- iii.1996</date>
, leg.
<collectorName id="6C5C75CAFF8BCF20DE64F937FE9DA785" box="[305,371,1698,1722]" pageId="15" pageNumber="165">Stock</collectorName>
,
<collectorName id="6C5C75CAFF8BCF20DE2BF937FE10A785" box="[382,510,1698,1722]" pageId="15" pageNumber="165">Vermeulen</collectorName>
&amp;
<collectorName id="6C5C75CAFF8BCF20DD75F937FD68A785" box="[544,646,1698,1722]" pageId="15" pageNumber="165">Al Nofli.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C116101CFF8BCF20DF90F958FE8AA671" blockId="15.[159,762,1277,1948]" pageId="15" pageNumber="165">
Other material —
<materialsCitation id="71C11A41FF8BCF20DEF1F958FE8AA671" collectingDate="2006-08" collectorName="Kumar." country="India" county="Alaknanda River" location="Uttaranchal" municipality="Dhundeshwargad" pageId="15" pageNumber="165" specimenCount="1">
1/0/0 (1/0/0 mounted),
<collectingCountry id="B9BE508CFF8BCF20DD8DF95BFF28A637" name="India" pageId="15" pageNumber="165">India</collectingCountry>
,
<location id="C47646C7FF8BCF20DF83F965FE8DA637" LSID="urn:lsid:plazi:treatment:4900A10AFF8BCF20DFF0FB12FAA2A3C8:C47646C7FF8BCF20DF83F965FE8DA637" box="[214,355,1776,1800]" country="India" county="Alaknanda River" municipality="Dhundeshwargad" name="Uttaranchal" pageId="15" pageNumber="165">Uttaranchal</location>
,
<location id="C47646C7FF8BCF20DE26F965FD8BA637" LSID="urn:lsid:plazi:treatment:4900A10AFF8BCF20DFF0FB12FAA2A3C8:C47646C7FF8BCF20DE26F965FD8BA637" box="[371,613,1776,1800]" country="India" county="Alaknanda River" municipality="Dhundeshwargad" name="Garhwal Himalayas" pageId="15" pageNumber="165">Garhwal Himalayas</location>
,
<collectingMunicipality id="21728A66FF8BCF20DD20F965FF17A614" pageId="15" pageNumber="165">Dhundeshwargad</collectingMunicipality>
stream, a tributary of
<collectingCounty id="28776890FF8BCF20DEADF886FD2EA614" box="[504,704,1811,1835]" pageId="15" pageNumber="165">Alaknanda River</collectingCounty>
,
<date id="B51736DCFF8BCF20DD9FF886FF3AA671" pageId="15" pageNumber="165" value="2006-08">
<collectingDate id="A553CF34FF8BCF20DD9FF886FF3AA671" pageId="15" pageNumber="165" value="2006-08">viii- 2006</collectingDate>
</date>
leg.
<collectorName id="6C5C75CAFF8BCF20DE58F8A2FE8AA671" box="[269,356,1847,1870]" pageId="15" pageNumber="165">Kumar.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C116101CFF8BCF20DF90F8F4FA84A358" blockId="15.[159,762,1277,1948]" lastBlockId="15.[826,1429,311,694]" pageId="15" pageNumber="165">
Remarks — The specimens from
<collectingCountry id="B9BE508CFF8BCF20DD08F8F7FD48A645" box="[605,678,1890,1914]" name="Oman" pageId="15" pageNumber="165">Oman</collectingCountry>
fit the description of
<taxonomicName id="06A96B9FFF8BCF20DE1FF810FDD7A6A3" authorityName="Cook" authorityYear="1967" box="[330,569,1924,1948]" class="Arachnida" family="Limnesiidae" genus="Limnesia" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="165" phylum="Arthropoda" rank="species" species="pinguipalpis">
<emphasis id="F3DDCC0EFF8BCF20DE1FF810FDD7A6A3" box="[330,569,1924,1948]" italics="true" pageId="15" pageNumber="165">Limnesia pinguipalpis</emphasis>
</taxonomicName>
, described from the Oriental part of
<collectingCountry id="B9BE508CFF8BCF20DB76FEA2FB8FA070" box="[1059,1121,311,335]" name="India" pageId="15" pageNumber="165">India</collectingCountry>
(
<collectingRegion id="036DDEFEFF8BCF20DB25FEA2FAE7A070" box="[1136,1289,311,335]" country="India" name="Maharashtra" pageId="15" pageNumber="165">Maharashtra</collectingRegion>
,
<collectingRegion id="036DDEFEFF8BCF20DA41FEA2FA7EA070" box="[1300,1424,311,335]" country="India" name="Karnataka" pageId="15" pageNumber="165">Karnataka</collectingRegion>
,
<collectingRegion id="036DDEFEFF8BCF20DC6FFECFFC27A04D" box="[826,969,346,370]" country="India" name="Tamil Nadu" pageId="15" pageNumber="165">Tamil Nadu</collectingRegion>
-
<bibRefCitation id="A5386DEDFF8BCF20DCB0FECFFB83A04D" author="Cook, D. R." box="[997,1133,346,370]" pageId="15" pageNumber="165" pagination="1 - 411" refId="ref17836" refString="Cook, D. R. 1967 - Water mites from India - Mem. Amer. Ent. Inst., 9: 1 - 411." type="journal article" year="1967">Cook, 1967</bibRefCitation>
). The only non-Oriental record from
<collectingCountry id="B9BE508CFF8BCF20DC9BFEE8FBE2A0AA" box="[974,1036,381,405]" name="India" pageId="15" pageNumber="165">India</collectingCountry>
comes from Garhwal Himalayas (see above).
<taxonomicName id="06A96B9FFF8BCF20DCB4FE34FB3AA087" authorityName="Cook" authorityYear="1967" box="[993,1236,416,440]" class="Arachnida" family="Limnesiidae" genus="Limnesia" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="165" phylum="Arthropoda" rank="species" species="pinguipalpis">
<emphasis id="F3DDCC0EFF8BCF20DCB4FE34FB3AA087" box="[993,1236,416,440]" italics="true" pageId="15" pageNumber="165">Limnesia pinguipalpis</emphasis>
</taxonomicName>
is similar to
<emphasis id="F3DDCC0EFF8BCF20DA2AFE34FC64A0E4" italics="true" pageId="15" pageNumber="165">L. monodi</emphasis>
but differs in the more extensive secondary sclerotization associated with the posterior end of the male genital field (
<figureCitation id="59920C99FF8BCF20DB6EFD9CFB7BA31E" box="[1083,1173,521,545]" captionStart="FIGURE" captionStartId="16.[159,171,1884,1903]" captionTargetBox="[228,1348,326,1823]" captionTargetId="figure-7@16.[219,1368,302,1852]" captionTargetPageId="16" captionText="FIGURE 10: Limnesia pinguipalpis Cook, male (A-C, F-G: specimens from Oman, E: specimen from India): (A) – idiosoma, dorsal view; (B) – genital field; (C, E) – palp; (F) – I-Leg-5-6; (G) – IV-Leg-5-6. D. Limnesia monodi, Zaire, lake Kivu, male: (D) – palp. Scale bars = 200 µm (A), 50 µm (B-G)." figureDoi="http://doi.org/10.5281/zenodo.5404104" httpUri="https://zenodo.org/record/5404104/files/figure.png" pageId="15" pageNumber="165">Fig.10B</figureCitation>
), and the ventral margin of P-2 more or less straight (
<figureCitation id="59920C99FF8BCF20DBEAFDB9FA8DA37B" box="[1215,1379,556,580]" captionStart="FIGURE" captionStartId="16.[159,171,1884,1903]" captionTargetBox="[228,1348,326,1823]" captionTargetId="figure-7@16.[219,1368,302,1852]" captionTargetPageId="16" captionText="FIGURE 10: Limnesia pinguipalpis Cook, male (A-C, F-G: specimens from Oman, E: specimen from India): (A) – idiosoma, dorsal view; (B) – genital field; (C, E) – palp; (F) – I-Leg-5-6; (G) – IV-Leg-5-6. D. Limnesia monodi, Zaire, lake Kivu, male: (D) – palp. Scale bars = 200 µm (A), 50 µm (B-G)." figureDoi="http://doi.org/10.5281/zenodo.5404104" httpUri="https://zenodo.org/record/5404104/files/figure.png" pageId="15" pageNumber="165">Fig. 10C, 10E</figureCitation>
) vs. convexly protruding in
<emphasis id="F3DDCC0EFF8BCF20DB05FDC5FB52A358" box="[1104,1212,591,615]" italics="true" pageId="15" pageNumber="165">L. monodi</emphasis>
(see
<figureCitation id="59920C99FF8BCF20DBA2FDDAFAB0A358" box="[1271,1374,591,615]" captionStart="FIGURE" captionStartId="16.[159,171,1884,1903]" captionTargetBox="[228,1348,326,1823]" captionTargetId="figure-7@16.[219,1368,302,1852]" captionTargetPageId="16" captionText="FIGURE 10: Limnesia pinguipalpis Cook, male (A-C, F-G: specimens from Oman, E: specimen from India): (A) – idiosoma, dorsal view; (B) – genital field; (C, E) – palp; (F) – I-Leg-5-6; (G) – IV-Leg-5-6. D. Limnesia monodi, Zaire, lake Kivu, male: (D) – palp. Scale bars = 200 µm (A), 50 µm (B-G)." figureDoi="http://doi.org/10.5281/zenodo.5404104" httpUri="https://zenodo.org/record/5404104/files/figure.png" pageId="15" pageNumber="165">Fig. 10D</figureCitation>
).
</paragraph>
<paragraph id="C116101CFF8BCF20DC35FDEEFB8AA389" blockId="15.[826,1429,311,694]" pageId="15" pageNumber="165">
Distribution —
<collectingCountry id="B9BE508CFF8BCF20DB43FDEEFBB8A3AC" box="[1046,1110,635,659]" name="India" pageId="15" pageNumber="165">India</collectingCountry>
;
<collectingCountry id="B9BE508CFF8BCF20DB35FDEEFB47A3AC" box="[1120,1193,635,659]" name="Oman" pageId="15" pageNumber="165">Oman</collectingCountry>
(&quot;
<taxonomicName id="06A96B9FFF8BCF20DB97FDE9FACDA3AC" authorityName="Koch" authorityYear="1836" box="[1218,1315,636,659]" class="Arachnida" family="Limnesiidae" genus="Limnesia" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="F3DDCC0EFF8BCF20DB97FDE9FACDA3AC" box="[1218,1315,636,659]" italics="true" pageId="15" pageNumber="165">Limnesia</emphasis>
</taxonomicName>
[
<emphasis id="F3DDCC0EFF8BCF20DA67FDEEFC9FA389" italics="true" pageId="15" pageNumber="165">Tetralimnesia</emphasis>
] sp.&quot;
<bibRefCitation id="A5386DEDFF8BCF20DCE7FD0BFBB7A389" author="Gerecke, R." box="[946,1113,670,694]" pageId="15" pageNumber="165" pagination="2833 - 2849" refId="ref17893" refString="Gerecke, R. 2004 - Water mites from spring- and groundwater habitats in the Sultanate of Oman - J. Nat. Hist., 38: 2833 - 2849." type="journal article" year="2004">Gerecke, 2004</bibRefCitation>
).
</paragraph>
<paragraph id="C116101CFF8BCF20DCD6FD4AFAA2A3C8" blockId="15.[899,1356,735,759]" box="[899,1356,735,759]" pageId="15" pageNumber="165">
Subfamily
<taxonomicName id="06A96B9FFF8BCF20DB57FD4AFAA2A3C8" authority="Viets, 1940" authorityName="Viets" authorityYear="1940" box="[1026,1356,735,759]" class="Arachnida" family="Limnesiidae" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="165" phylum="Arthropoda" rank="subFamily" subFamily="Protolimnesiinae">Protolimnesiinae Viets, 1940</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

151
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<heading id="9A5EA770FF8FCF24DF99FB2DFD22A5EE" bold="true" box="[204,716,1208,1233]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="161" reason="2">
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<taxonomicName id="06A96B9FFF8FCF24DF99FB2DFD6DA5EE" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[204,643,1208,1233]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="161" phylum="Arthropoda" rank="species" species="omanensis" status="sp. nov." subGenus="Megapalpis">
<emphasis id="F3DDCC0EFF8FCF24DF99FB2DFD6DA5EE" bold="true" box="[204,643,1208,1233]" italics="true" pageId="11" pageNumber="161">Torrenticola (Megapalpis) omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF8FCF24DDDFFB2CFD22A5EE" box="[650,716,1209,1233]" pageId="11" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
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(
<figureCitation id="59920C99FF8FCF24DE0FFB67FE28A435" box="[346,454,1266,1290]" captionStart="FIGURE" captionStartId="10.[159,171,1888,1907]" captionTargetBox="[272,1292,313,1849]" captionTargetId="figure-7@10.[254,1334,307,1857]" captionTargetPageId="10" captionText="FIGURE 6: Torrenticola omanensis n. sp., male: (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view; (D) – palp, lateral view; (E) – ejaculatory complex. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404096" httpUri="https://zenodo.org/record/5404096/files/figure.png" pageId="11" pageNumber="161">Figures 6</figureCitation>
A-E, 7A-B)
</paragraph>
<paragraph id="C116101CFF8FCF24DF90FAA2FF34A486" blockId="11.[159,762,1335,1949]" pageId="11" pageNumber="161">
Type series —
<materialsCitation id="71C11A41FF8FCF24DED7FAA2FEF7A4AA" collectingDate="2008-11-15" country="Oman" elevation="421" latitude="24.686083" location="Hatta Pools" longLatPrecision="1" longitude="56.159267" pageId="11" pageNumber="161" specimenCount="1" typeStatus="holotype">
<typeStatus id="1E12AEBEFF8FCF24DED7FAA2FE1AA470" box="[386,500,1335,1359]" pageId="11" pageNumber="161" type="holotype">Holotype</typeStatus>
: male, stream
<location id="C47646C7FF8FCF24DDEDFAA2FF0FA44D" LSID="urn:lsid:plazi:treatment:4900A10AFF8FCF23DF99FB2DFE22A744:C47646C7FF8FCF24DDEDFAA2FF0FA44D" country="Oman" latitude="24.686083" longLatPrecision="1" longitude="56.159267" name="Hatta Pools" pageId="11" pageNumber="161">Hatta Pools</location>
,
<collectingCountry id="B9BE508CFF8FCF24DFBAFACFFED4A44D" box="[239,314,1370,1394]" name="Oman" pageId="11" pageNumber="161">Oman</collectingCountry>
,
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<geoCoordinate id="A49D76DBFF8FCF24DE8AFACFFD87A44E" box="[479,617,1370,1394]" degrees="56" direction="east" minutes="09.556" orientation="longitude" pageId="11" pageNumber="161" precision="1" value="56.159267">56°09.556 E</geoCoordinate>
, alt.
<quantity id="0651BDF9FF8FCF24DDFFFACFFD1AA44E" box="[682,756,1370,1394]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="11" pageNumber="161" unit="m" value="421.0">
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,
<date id="B51736DCFF8FCF24DFCAFAE8FEF7A4AA" box="[159,281,1405,1429]" pageId="11" pageNumber="161" value="2008-11-15">
<collectingDate id="A553CF34FF8FCF24DFCAFAE8FEF7A4AA" box="[159,281,1405,1429]" pageId="11" pageNumber="161" value="2008-11-15">15-xi-2008</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="1E12AEBEFF8FCF24DE7BFAE8FE74A4AA" box="[302,410,1405,1429]" pageId="11" pageNumber="161" type="paratype">Paratype</typeStatus>
: (1/0/0), same data as holotype.
</paragraph>
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FIGURE 7:
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<emphasis id="F3DDCC0EFF8FCF24DCF0FACBFB83A44F" box="[933,1133,1374,1393]" italics="true" pageId="11" pageNumber="161">Torrenticola omanensis</emphasis>
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, male: (A) – capitulum; (B) – chelicera. Scale bar = 100 µm.
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<paragraph id="C116101CFF8FCF24DF90FA58FE25A7E5" blockId="11.[159,762,1335,1949]" pageId="11" pageNumber="161">Diagnosis — Males (female unknown). Cx-IV extended strongly posterior to genital field; P-2 long (L ratio P-2/P-4 2.5) and with strong ventral denticles distally to slightly pronounced ventral projection, P-3 with serrulate margin distally from small pointed ventral projection; capitulum with a very long rostrum; cheliceral claw relatively short (basal segment/claw ratio 12.0).</paragraph>
<paragraph id="C116101CFF8FCF24DF90F97AFEA0A638" blockId="11.[159,762,1335,1949]" box="[197,334,1775,1799]" pageId="11" pageNumber="161">Description</paragraph>
<paragraph id="C116101CFF8FCF24DF90F88EFAE9A2F9" blockId="11.[159,762,1335,1949]" lastBlockId="11.[826,1429,311,966]" pageId="11" pageNumber="161">
Male — Idiosoma (ventral view:
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) L 763, W 563; dorsal shield (
<figureCitation id="59920C99FF8FCF24DEF0F8ABFDECA669" box="[421,514,1854,1878]" captionStart="FIGURE" captionStartId="10.[159,171,1888,1907]" captionTargetBox="[272,1292,313,1849]" captionTargetId="figure-7@10.[254,1334,307,1857]" captionTargetPageId="10" captionText="FIGURE 6: Torrenticola omanensis n. sp., male: (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view; (D) – palp, lateral view; (E) – ejaculatory complex. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404096" httpUri="https://zenodo.org/record/5404096/files/figure.png" pageId="11" pageNumber="161">Fig. 6A</figureCitation>
) L 675, W 466, L/W ratio 1.45; dorsal plate 634; shoulder plate L 209, W 67, L/W ratio 3.1; frontal plate L 138, W 53, L/W ratio 2.6; shoulder/frontal plate L ratio 1.5; gnathosomal bay L 166, Cx-I total L 300, Cx-I medial L 133, Cx-II+III medial 67; ratio Cx-I L/Cx-II+III medial L 4.5; Cx-I medial L/Cx-II+III medial L 2.0; Cx- IV extended posterior to genital field; Cxgl-4 anterior to Cxgl-2 and located adjacent to Leg-II socket; genital field L/W 144/125, L/W ratio 1.15, ejaculatory complex (
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) L 162; distance genital fieldexcretory pore 199, genital field-caudal idiosoma margin 241; capitulum (
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) with long rostrum, ventral L 628; chelicera (
<figureCitation id="59920C99FF8FCF24DB37FD00FB53A391" box="[1122,1213,661,686]" captionStart="FIGURE" captionStartId="11.[826,838,1374,1393]" captionTargetBox="[832,1417,1010,1318]" captionTargetId="figure-499@11.[832,1422,1002,1342]" captionTargetPageId="11" captionText="FIGURE 7: Torrenticola omanensis n. sp., male: (A) – capitulum; (B) – chelicera. Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404098" httpUri="https://zenodo.org/record/5404098/files/figure.png" pageId="11" pageNumber="161">Fig. 7B</figureCitation>
) L 663, basal segment L 634, claw L 53, basal segment/claw ratio L 12.0; palp (
<figureCitation id="59920C99FF8FCF24DC96FD49FBF5A3CB" box="[963,1051,732,756]" captionStart="FIGURE" captionStartId="10.[159,171,1888,1907]" captionTargetBox="[272,1292,313,1849]" captionTargetId="figure-7@10.[254,1334,307,1857]" captionTargetPageId="10" captionText="FIGURE 6: Torrenticola omanensis n. sp., male: (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view; (D) – palp, lateral view; (E) – ejaculatory complex. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404096" httpUri="https://zenodo.org/record/5404096/files/figure.png" pageId="11" pageNumber="161">Figs. 6</figureCitation>
C-D) total L 620, L and %L (in parentheses): P-1, 88 (14.2); P-2, 297 (47.9); P-3, 91 (14.7); P-4, 119 (19.2); P-5, 25 (4.0); P-2/P-4 ratio 2.5; P-2 with well developed ventral denticles distally to slightly pronounced ventral projection bearing long seta, P-3 with serrulate margin distally from small pointed ventral projection bearing seta.
</paragraph>
<paragraph id="C116101CFF8FCF24DC35FA5EFBBBA4DC" blockId="11.[826,1429,1483,1949]" box="[864,1109,1483,1507]" pageId="11" pageNumber="161">Female — unknown.</paragraph>
<paragraph id="C116101CFF8FCF24DC35FA6DFB21A673" blockId="11.[826,1429,1483,1949]" pageId="11" pageNumber="161">
Remarks. The new species can be distinguished from other members of the subgenus
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Halbert,
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in the combination of the characteristic shape of the palp (P-2 very long and with ventral denticles distally to slightly pronounced ventral projection, P-3 with serrulate margin distally from small pointed ventral projection), capitulum with very long rostrum, a relatively short cheliceral claw (basal segment/claw ratio 12.0) and Cx-IV greatly extended posterior to genital field.
</paragraph>
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Etymology — The species is named for its occurrence in
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<paragraph id="C116101CFF88CF23DFCAFF6FFEB3A032" blockId="12.[159,349,250,269]" box="[159,349,250,269]" pageId="12" pageNumber="162">Smit H. and Pe˘si´c V.</paragraph>
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FIGURE 8:
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, female: (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, medial view. Scale bars = 100 µm.
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Distribution —
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A-D, 15A-E)
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Type material —
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<collectingCountry id="B9BE508CFF90CF3BDD5BFBACFDB7A56E" box="[526,601,1081,1105]" name="Oman" pageId="20" pageNumber="170">Oman</collectingCountry>
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<collectingDate id="A553CF34FF90CF3BDE99FBC9FDABA54B" box="[460,581,1116,1140]" pageId="20" pageNumber="170" value="2008-11-12">12-xi-2008</collectingDate>
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;
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1/0/0 (1/0/0 mounted), stream
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<collectingCountry id="B9BE508CFF90CF3BDD41FBEAFDB1A5A8" box="[532,607,1151,1175]" name="Oman" pageId="20" pageNumber="170">Oman</collectingCountry>
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<geoCoordinate id="A49D76DBFF90CF3BDFCAFB37FEC8A585" box="[159,294,1186,1210]" degrees="56" direction="east" minutes="09.556" orientation="longitude" pageId="20" pageNumber="170" precision="1" value="56.159267">56°09.556 E</geoCoordinate>
, alt.
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<collectingDate id="A553CF34FF90CF3BDEE4FB37FDC5A585" box="[433,555,1186,1210]" pageId="20" pageNumber="170" value="2008-11-15">15-xi-2008</collectingDate>
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<paragraph id="C116101CFF90CF3BDF90FB44FE40A4A6" blockId="20.[159,762,940,1948]" pageId="20" pageNumber="170">
Diagnosis — Genital field in the male with anterior margin concavely indented and Ac-3 distanced from anterior two acetabula on each side, I-Leg-5 with S-1 and -2 far distanced (&gt;
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both sexes), S-1 curved, with blunt tip, S-2 with strongly developed dorsal projection.
</paragraph>
<paragraph id="C116101CFF90CF3BDF90FA25FEA0A4F7" blockId="20.[159,762,940,1948]" box="[197,334,1456,1480]" pageId="20" pageNumber="170">Description</paragraph>
<paragraph id="C116101CFF90CF3BDF90FA75FD26A6A3" blockId="20.[159,762,940,1948]" pageId="20" pageNumber="170">General features — Dorsal integument: striated; muscle attachment plates smooth. Coxal field: mediocaudal margin of Cx-I+II with a large, slightly indented area between the laterally directed apodemes of Cx-II. Genital field: Ac in a weakly curved line. Palp: weak sexual dimorphism, P- 2 ventral margin slightly convex, P-3 straight, P- 4 sword seta near distal hair. I-Leg: I-Leg-5 with S-1 and -2 strongly heteromorphic and distanced, ventral seta close to insertion of S-1, curved, with blunt tip, S-2 with strongly developed dorsal projection, bluntly pointed, I-Leg-6 long and slender, only slightly narrowed from the base to the tip.</paragraph>
<paragraph id="C116101CFF90CF3BDC35FEA2FABAA229" blockId="20.[826,1429,311,1948]" pageId="20" pageNumber="170">
Male — (
<typeStatus id="1E12AEBEFF90CF3BDC92FEA2FBDCA070" box="[967,1074,311,335]" pageId="20" pageNumber="170" type="holotype">holotype</typeStatus>
, in parentheses measurements of specimen from stream Hatta Pools). Idiosoma L/W 553 (566)/366 (381). Coxal field (
<figureCitation id="59920C99FF90CF3BDA4CFEE8FA67A0AA" box="[1305,1417,381,405]" captionStart="FIGURE" captionStartId="21.[159,171,1888,1907]" captionTargetBox="[248,1308,325,1844]" captionTargetId="figure-4@21.[248,1340,307,1857]" captionTargetPageId="21" captionText="FIGURE 14: Atractides arabicus n. sp., male (A-B, D holotype, C paratype): (A) – I-Leg- 5 and - 6; (B) – idiosoma, ventral view; (C) – genital field; (D) – palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404112" httpUri="https://zenodo.org/record/5404112/files/figure.png" pageId="20" pageNumber="170">Fig. 14B</figureCitation>
): L 303 (316), Cx-III W 319 (325), Cx-I+II mL 71 (72), Cx-I+2 lL 169 (178). Palp (
<figureCitation id="59920C99FF90CF3BDBD5FE56FB1FA0E4" box="[1152,1265,451,475]" captionStart="FIGURE" captionStartId="21.[159,171,1888,1907]" captionTargetBox="[248,1308,325,1844]" captionTargetId="figure-4@21.[248,1340,307,1857]" captionTargetPageId="21" captionText="FIGURE 14: Atractides arabicus n. sp., male (A-B, D holotype, C paratype): (A) – I-Leg- 5 and - 6; (B) – idiosoma, ventral view; (C) – genital field; (D) – palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404112" httpUri="https://zenodo.org/record/5404112/files/figure.png" pageId="20" pageNumber="170">Fig. 14D</figureCitation>
): palp total L 302 (317), dL: P-1, 29 (30); P-2, 66 (71); P-3, 71 (77); P-4, 102 (105); P-5, 34 (34); %L: P-1, 9.6 (9.5); P- 2, 21.9 (23.1); P-3, 23.5 (24.3); P-4, 33.8 (33.1); P-5, 11.3 (10.7); L P-2/P-4 0.65 (0.68); P-4 ventral hairs long. Genital field (
<figureCitation id="59920C99FF90CF3BDB76FDE7FB62A3B4" box="[1059,1164,626,651]" captionStart="FIGURE" captionStartId="21.[159,171,1888,1907]" captionTargetBox="[248,1308,325,1844]" captionTargetId="figure-4@21.[248,1340,307,1857]" captionTargetPageId="21" captionText="FIGURE 14: Atractides arabicus n. sp., male (A-B, D holotype, C paratype): (A) – I-Leg- 5 and - 6; (B) – idiosoma, ventral view; (C) – genital field; (D) – palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404112" httpUri="https://zenodo.org/record/5404112/files/figure.png" pageId="20" pageNumber="170">Fig. 14C</figureCitation>
): anterior margin concavely indented, lateral margin weakly indented between Ac-2 and Ac-3, L/W 89 (95)/94 (102), Ac-3 rather small, distanced from anterior two acetabula on each side, L Ac-1-3: 26 (26), 28 (28), 22 (25).
</paragraph>
<paragraph id="C116101CFF90CF3BDC35FCBEFB29A2CD" blockId="20.[826,1429,311,1948]" pageId="20" pageNumber="170">
I-Leg (
<figureCitation id="59920C99FF90CF3BDCE5FCBEFBCAA27C" box="[944,1060,811,835]" captionStart="FIGURE" captionStartId="21.[159,171,1888,1907]" captionTargetBox="[248,1308,325,1844]" captionTargetId="figure-4@21.[248,1340,307,1857]" captionTargetPageId="21" captionText="FIGURE 14: Atractides arabicus n. sp., male (A-B, D holotype, C paratype): (A) – I-Leg- 5 and - 6; (B) – idiosoma, ventral view; (C) – genital field; (D) – palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404112" httpUri="https://zenodo.org/record/5404112/files/figure.png" pageId="20" pageNumber="170">Fig. 14A</figureCitation>
): I-Leg-5 dL 212 (218), vL 114 (124), dL/Vl 1.86 (1.76), HB 50 (50), dL/HB 4.2 (4.4), S-1 L 100 (103), L/W 8.6 (8.9), S-2 L 63 (67), L/W 5.4 (7.3), distance of sword setae at I-Leg-5 43 (43), L S- 1/2 1.59 (1.54); I-Leg-6 L 159 (157), HB 15 (15), L/HB 10.3 (10.2); L I-Leg-5/6 1.34 (1.39).
</paragraph>
<paragraph id="C116101CFF90CF3BDC35FB93FB71A4FC" blockId="20.[826,1429,311,1948]" pageId="20" pageNumber="170">
Female — (from stream Wadi Ghul, in parentheses measurement of specimen from small cascading stream Wadi Ghul). Idiosoma L/W 944 (700)/663 (519). Coxal field (
<figureCitation id="59920C99FF90CF3BDB4FFBFAFB68A5B8" box="[1050,1158,1135,1159]" captionStart="FIGURE" captionStartId="22.[159,171,1888,1907]" captionTargetBox="[257,1318,322,1842]" captionTargetId="figure-7@22.[249,1338,307,1857]" captionTargetPageId="22" captionText="FIGURE 15: Atractides arabicus n. sp., female: (A-B) – I-Leg- 5 and - 6; (C) – palp, lateral view; (D) – coxal field; (E) – genital field. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404114" httpUri="https://zenodo.org/record/5404114/files/figure.png" pageId="20" pageNumber="170">Fig. 15D</figureCitation>
): L 400 (378), Cx-III W 448 (389), Cx-I+II mL 78 (79), Cx-I+II lL 213 (204); apodemes Cx-II directed caudolaterally. Palp (
<figureCitation id="59920C99FF90CF3BDA3DFB20FC85A5CF" captionStart="FIGURE" captionStartId="22.[159,171,1888,1907]" captionTargetBox="[257,1318,322,1842]" captionTargetId="figure-7@22.[249,1338,307,1857]" captionTargetPageId="22" captionText="FIGURE 15: Atractides arabicus n. sp., female: (A-B) – I-Leg- 5 and - 6; (C) – palp, lateral view; (D) – coxal field; (E) – genital field. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404114" httpUri="https://zenodo.org/record/5404114/files/figure.png" pageId="20" pageNumber="170">Fig. 15C</figureCitation>
): palp total L 386 (373), dL: P-1, 34 (34); P-2, 85 (82); P-3, 102 (99); P-4, 128 (122); P-5, 37 (36); %L: P- 1, 8.8 (9.1); P-2, 22.0 (22.0); P-3, 26.4 (26.5); P-4, 33.2 (32.7); P-5, 9.6 (9.7); L P-2/P-4 0.66 (0.67). Genital field (
<figureCitation id="59920C99FF90CF3BDCD5FAF0FC04A443" box="[896,1002,1380,1405]" captionStart="FIGURE" captionStartId="22.[159,171,1888,1907]" captionTargetBox="[257,1318,322,1842]" captionTargetId="figure-7@22.[249,1338,307,1857]" captionTargetPageId="22" captionText="FIGURE 15: Atractides arabicus n. sp., female: (A-B) – I-Leg- 5 and - 6; (C) – palp, lateral view; (D) – coxal field; (E) – genital field. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404114" httpUri="https://zenodo.org/record/5404114/files/figure.png" pageId="20" pageNumber="170">Fig. 15E</figureCitation>
): L 163 (135), genital plates slightly indented between Ac, genital plate L 109 (100), L Ac-1-3: 35 (33), 34 (32), 35 (34).
</paragraph>
<paragraph id="C116101CFF90CF3BDC35FA42FB54A7A1" blockId="20.[826,1429,311,1948]" pageId="20" pageNumber="170">
I-Leg (
<figureCitation id="59920C99FF90CF3BDCFBFA42FBEFA4D0" box="[942,1025,1495,1519]" captionStart="FIGURE" captionStartId="22.[159,171,1888,1907]" captionTargetBox="[257,1318,322,1842]" captionTargetId="figure-7@22.[249,1338,307,1857]" captionTargetPageId="22" captionText="FIGURE 15: Atractides arabicus n. sp., female: (A-B) – I-Leg- 5 and - 6; (C) – palp, lateral view; (D) – coxal field; (E) – genital field. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404114" httpUri="https://zenodo.org/record/5404114/files/figure.png" pageId="20" pageNumber="170">Fig. 15</figureCitation>
A-B): I-Leg-5 dL 295 (294), vL 151 (154), dL/vL 2.0 (1.9), HB 76 (79), dL/HB 3.9 (3.7), S-1 L 119 (119), L/W 8.6 (7.4), S-2 L 74 (75), L/W 5.3 (5.4), distance of sword setae at I-Leg-5 77 (69), L S- 1/2 1.6 (1.6); I-Leg-6 L 209 (199), HB 18 (17), L/HB 11.8 (11.7); L I-Leg-5/6 1.4 (1.48).
</paragraph>
<paragraph id="C116101CFF90CF3BDC35F927FB97A6A3" blockId="20.[826,1429,311,1948]" pageId="20" pageNumber="170">
Remarks —
<taxonomicName id="06A96B9FFF90CF3BDCA1F927FB28A7F5" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[1012,1222,1714,1738]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="170" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="F3DDCC0EFF90CF3BDCA1F927FB28A7F5" box="[1012,1222,1714,1738]" italics="true" pageId="20" pageNumber="170">Atractides arabicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF90CF3BDB9AF926FAF8A7F5" box="[1231,1302,1715,1738]" pageId="20" pageNumber="170" rank="species">n. sp.</taxonomicNameLabel>
is unique within the genus due to the particular shape of S-2 (with a well developed dent on the outer margin of seta). In addition, the male is characterized by the anterior margin of genital field concavely indented, with rather small Ac-3 distanced from the two anterior acetabula on each side.
</paragraph>
<caption id="95D64094FF91CF3ADFCAF8F5FD70A6B0" ID-DOI="http://doi.org/10.5281/zenodo.5404112" ID-Zenodo-Dep="5404112" httpUri="https://zenodo.org/record/5404112/files/figure.png" pageId="21" pageNumber="171" startId="21.[159,171,1888,1907]" targetBox="[248,1308,325,1844]" targetPageId="21">
<paragraph id="C116101CFF91CF3ADFCAF8F5FD70A6B0" blockId="21.[159,1428,1888,1935]" pageId="21" pageNumber="171">
FIGURE 14:
<taxonomicName id="06A96B9FFF91CF3ADE5AF8F5FE5BA64C" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[271,437,1888,1907]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="171" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="F3DDCC0EFF91CF3ADE5AF8F5FE5BA64C" box="[271,437,1888,1907]" italics="true" pageId="21" pageNumber="171">Atractides arabicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF91CF3ADEECF8F5FE00A64C" box="[441,494,1888,1907]" pageId="21" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
, male (A-B, D holotype, C paratype): (A) – I-Leg- 5 and - 6; (B) – idiosoma, ventral view; (C) – genital field; (D) – palp, medial view. Scale bars = 100 µm.
</paragraph>
</caption>
<caption id="95D64094FF92CF39DFCAF8F5FEA3A6B0" ID-DOI="http://doi.org/10.5281/zenodo.5404114" ID-Zenodo-Dep="5404114" httpUri="https://zenodo.org/record/5404114/files/figure.png" pageId="22" pageNumber="172" startId="22.[159,171,1888,1907]" targetBox="[257,1318,322,1842]" targetPageId="22">
<paragraph id="C116101CFF92CF39DFCAF8F5FEA3A6B0" blockId="22.[159,1429,1888,1935]" pageId="22" pageNumber="172">
FIGURE 15:
<taxonomicName id="06A96B9FFF92CF39DE47F8F5FE57A64C" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[274,441,1888,1907]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="172" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="F3DDCC0EFF92CF39DE47F8F5FE57A64C" box="[274,441,1888,1907]" italics="true" pageId="22" pageNumber="172">Atractides arabicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF92CF39DEEBF8F5FE1BA64C" box="[446,501,1888,1907]" pageId="22" pageNumber="172" rank="species">n. sp.</taxonomicNameLabel>
, female: (A-B) – I-Leg- 5 and - 6; (C) – palp, lateral view; (D) – coxal field; (E) – genital field. Scale bars = 100 µm.
</paragraph>
</caption>
<paragraph id="C116101CFF93CF38DF90FEA2FDFBA04D" blockId="23.[159,762,311,415]" pageId="23" pageNumber="173">Etymology — The species is named for its occurrence on the Arabian peninsula.</paragraph>
<paragraph id="C116101CFF93CF38DF90FE12FE22A0A0" blockId="23.[159,762,311,415]" box="[197,460,391,415]" pageId="23" pageNumber="173">
Distribution —
<collectingCountry id="B9BE508CFF93CF38DE29FE12FE26A0A0" box="[380,456,391,415]" name="Oman" pageId="23" pageNumber="173">Oman</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="C116101CFF93CF38DFE4FE52FD06A0DF" blockId="23.[177,744,455,480]" box="[177,744,455,480]" pageId="23" pageNumber="173">
<heading id="9A5EA770FF93CF38DFE4FE52FD06A0DF" bold="true" box="[177,744,455,480]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="173" reason="2">
<emphasis id="F3DDCC0EFF93CF38DFE4FE52FD06A0DF" bold="true" box="[177,744,455,480]" pageId="23" pageNumber="173">
<taxonomicName id="06A96B9FFF93CF38DFE4FE52FD70A0E0" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[177,670,455,479]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="173" phylum="Arthropoda" rank="species" species="omanensis" status="sp. nov." subGenus="Tympanomegapus">
<emphasis id="F3DDCC0EFF93CF38DFE4FE52FD70A0E0" bold="true" box="[177,670,455,479]" italics="true" pageId="23" pageNumber="173">Atractides (Tympanomegapus) omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF93CF38DDF0FE5DFD06A0DF" box="[677,744,456,480]" pageId="23" pageNumber="173" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="89B34397FF93CF37DE4EFD94FEB3A032" lastPageId="24" lastPageNumber="174" pageId="23" pageNumber="173" type="description">
<paragraph id="C116101CFF93CF38DE4EFD94FD93A325" blockId="23.[283,637,513,538]" box="[283,637,513,538]" pageId="23" pageNumber="173">
(
<figureCitation id="59920C99FF93CF38DE76FD94FE72A326" box="[291,412,513,537]" captionStart="FIGURE" captionStartId="24.[159,171,1693,1712]" captionTargetBox="[189,1399,302,1661]" captionTargetId="figure-7@24.[189,1399,302,1661]" captionTargetPageId="24" captionText="FIGURE 16: Atractides omanensis n. sp., male (A-D holotype, E paratype): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, lateral view; (D-E) – I-Leg- 5 and - 6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404116" httpUri="https://zenodo.org/record/5404116/files/figure.png" pageId="23" pageNumber="173">Figures 16</figureCitation>
A-E, 17, 18, 19A-D)
</paragraph>
<paragraph id="C116101CFF93CF38DF90FDDFFEF3A20B" blockId="23.[159,762,586,1948]" pageId="23" pageNumber="173">
Type material —
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<typeStatus id="1E12AEBEFF93CF38DEFDFDDFFDF4A35D" box="[424,538,586,610]" pageId="23" pageNumber="173" type="holotype">Holotype</typeStatus>
: male, unnamed stream crossing road to
<location id="C47646C7FF93CF38DE9FFDF8FDF5A3BA" LSID="urn:lsid:plazi:treatment:4900A10AFF93CF37DFE4FE52FEB3A032:C47646C7FF93CF38DE9FFDF8FDF5A3BA" box="[458,539,621,645]" country="Oman" latitude="24.64505" longLatPrecision="1" longitude="56.112667" municipality="Ray" name="Al Fay" pageId="23" pageNumber="173">Al Fay</location>
, near
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,
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,
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, alt.
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,
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<collectingDate id="A553CF34FF93CF38DD29FD05FD18A397" box="[636,758,656,680]" pageId="23" pageNumber="173" value="2008-11-15">15-xi-2008</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="71C11A41FF93CF38DFCAFD26FEF7A20B" collectingDate="2008-11-12" country="Oman" elevation="805" latitude="23.171618" location="Wadi Ghul" longLatPrecision="1" longitude="57.199932" municipality="Ray" pageId="23" pageNumber="173" specimenCount="1" typeStatus="paratype">
<typeStatus id="1E12AEBEFF93CF38DFCAFD26FEF9A3F4" box="[159,279,691,715]" pageId="23" pageNumber="173" type="paratype">Paratypes</typeStatus>
: 0/1/0 (0/1/0 mounted), same data as holotype; 1/0/0 (1/0/0 mounted), stream
<location id="C47646C7FF93CF38DDE9FD43FF33A22E" LSID="urn:lsid:plazi:treatment:4900A10AFF93CF37DFE4FE52FEB3A032:C47646C7FF93CF38DDE9FD43FF33A22E" country="Oman" latitude="23.171618" longLatPrecision="1" longitude="57.199932" name="Wadi Ghul" pageId="23" pageNumber="173">Wadi Ghul</location>
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<geoCoordinate id="A49D76DBFF93CF38DE88FD6CFD88A22F" box="[477,614,761,785]" degrees="57" direction="east" minutes="11.996" orientation="longitude" pageId="23" pageNumber="173" precision="1" value="57.199932">57°11.996 E</geoCoordinate>
, alt.
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,
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<collectingDate id="A553CF34FF93CF38DFCAFC89FEF7A20B" box="[159,281,796,820]" pageId="23" pageNumber="173" value="2008-11-12">12-xi-2008</collectingDate>
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.
</paragraph>
<paragraph id="C116101CFF93CF38DF90FCDCFE14A52E" blockId="23.[159,762,586,1948]" pageId="23" pageNumber="173">Diagnosis — Muscle attachments sclerotized, cheliceral claw relatively strong, P-3 ventral margins with one seta, little enlarged, sword seta on P-4 enlarged, not hair-like, on the level of distoventral hair, P-5 without cheeks; excretory pore sclerotized; leg claws with dorsal clawlet.</paragraph>
<paragraph id="C116101CFF93CF38DF90FBB3FEA0A501" blockId="23.[159,762,586,1948]" box="[197,334,1062,1086]" pageId="23" pageNumber="173">Description</paragraph>
<paragraph id="C116101CFF93CF38DF90FBC1FECFA4BB" blockId="23.[159,762,586,1948]" pageId="23" pageNumber="173">General features — Dorsal integument: striated; muscle attachment plates: sexual dimorphism in the dorsal area. Genital field: Ac in triangular arrangement. Palp: weak sexual dimorphism, P-3 ventral margin with one seta, P-4 sword seta enlarged, inserted on level of the distoventral hair, P-5 without cheeks. I-Leg: setae S-1 and -2 inserted on the same level, distally truncate; leg claws with dorsal clawlet.</paragraph>
<paragraph id="C116101CFF93CF38DF90FA0FFC83A04D" blockId="23.[159,762,586,1948]" lastBlockId="23.[826,1429,311,1949]" pageId="23" pageNumber="173">
Male — (
<typeStatus id="1E12AEBEFF93CF38DE64FA0FFE72A48D" box="[305,412,1434,1458]" pageId="23" pageNumber="173" type="holotype">holotype</typeStatus>
, in parentheses measurement of specimen from stream Wadi Ghul). Idiosoma L/W (422)/(288). Dorsal shield (
<figureCitation id="59920C99FF93CF38DD48FA75FD6FA4C7" box="[541,641,1504,1528]" captionStart="FIGURE" captionStartId="24.[159,171,1693,1712]" captionTargetBox="[189,1399,302,1661]" captionTargetId="figure-7@24.[189,1399,302,1661]" captionTargetPageId="24" captionText="FIGURE 16: Atractides omanensis n. sp., male (A-D holotype, E paratype): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, lateral view; (D-E) – I-Leg- 5 and - 6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404116" httpUri="https://zenodo.org/record/5404116/files/figure.png" pageId="23" pageNumber="173">Fig. 16A</figureCitation>
) L/W 381 (372)/247 (247) including postoc and Dgl-3-6; ventral shield (
<figureCitation id="59920C99FF93CF38DE7DF9B3FE7EA701" box="[296,400,1574,1598]" captionStart="FIGURE" captionStartId="24.[159,171,1693,1712]" captionTargetBox="[189,1399,302,1661]" captionTargetId="figure-7@24.[189,1399,302,1661]" captionTargetPageId="24" captionText="FIGURE 16: Atractides omanensis n. sp., male (A-D holotype, E paratype): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, lateral view; (D-E) – I-Leg- 5 and - 6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404116" httpUri="https://zenodo.org/record/5404116/files/figure.png" pageId="23" pageNumber="173">Fig. 16B</figureCitation>
) L/W 394 (400)/344 (331), including coxae, Vgl-3 and -4, genital field, excretory pore and Vgl-1 and -2; coxal field: completely fused: Cx-III W 255. Palp (
<figureCitation id="59920C99FF93CF38DEDEF91AFE1DA798" box="[395,499,1679,1703]" captionStart="FIGURE" captionStartId="24.[159,171,1693,1712]" captionTargetBox="[189,1399,302,1661]" captionTargetId="figure-7@24.[189,1399,302,1661]" captionTargetPageId="24" captionText="FIGURE 16: Atractides omanensis n. sp., male (A-D holotype, E paratype): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, lateral view; (D-E) – I-Leg- 5 and - 6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404116" httpUri="https://zenodo.org/record/5404116/files/figure.png" pageId="23" pageNumber="173">Fig. 16C</figureCitation>
): total L 189 (198), dL: P-1, 24 (25); P-2, 40 (45); P-3, 42 (42); P-4, 60 (63); P-5, 23 (23); %L: P-1, 12.7 (12.6); P-2, 21.2 (22.7); P-3, 22.2 (21.2); P-4, 31.8 (31.8); P-5, 12.2 (11.6); L P-2/P-4 0.67 (0.71); chelicera total L 142 (151), basal segment L 97 (108), claw L 49 (49), L ratio basal segment/claw 2.0 (2.2); capitulum ventral L 82 (83). Genital field: fused with the ventral shield, but suture lines still evident, W 89 (95), L Ac-1-3: 15 (13), 12 (12-14), 17 (15).
</paragraph>
<paragraph id="C116101CFF93CF38DC35FE16FBC0A375" blockId="23.[826,1429,311,1949]" pageId="23" pageNumber="173">
I-Leg (
<figureCitation id="59920C99FF93CF38DCF9FE16FC15A0A4" box="[940,1019,387,411]" captionStart="FIGURE" captionStartId="24.[159,171,1693,1712]" captionTargetBox="[189,1399,302,1661]" captionTargetId="figure-7@24.[189,1399,302,1661]" captionTargetPageId="24" captionText="FIGURE 16: Atractides omanensis n. sp., male (A-D holotype, E paratype): (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – palp, lateral view; (D-E) – I-Leg- 5 and - 6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404116" httpUri="https://zenodo.org/record/5404116/files/figure.png" pageId="23" pageNumber="173">Fig. 16</figureCitation>
D-E): I-Leg-5 dL 79 (82), vL 62 (63), dL/vL 1.3 (1.3), HB 27 (28), dL/HB 2.9 (2.9), S-1 L 32 (36), L/W 10.3 (7.8), S-2 L 32 (36), L/W 10.3 (7.8), distance of sword setae at I-Leg-5 0 (0), L S-1/2 1.0 (1.0); I-Leg-6 L 80 (82), HB 19 (21), L/HB 4.3 (3.8); L I-Leg-5/6 0.98 (1.03).
</paragraph>
<paragraph id="C116101CFF93CF38DC35FDCEFC08A5D5" blockId="23.[826,1429,311,1949]" pageId="23" pageNumber="173">
Female — Idiosoma L/W 694/475; muscle attachment plates (
<figureCitation id="59920C99FF93CF38DB59FDEBFB83A3A9" box="[1036,1133,638,662]" captionStart="FIGURE" captionStartId="25.[374,386,1902,1921]" captionTargetBox="[272,1321,343,1811]" captionTargetId="figure-4@25.[231,1357,320,1870]" captionTargetPageId="25" captionText="FIGURE 17: Atractides omanensis n. sp., female: idiosoma, dorsal view. Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404118" httpUri="https://zenodo.org/record/5404118/files/figure.png" pageId="23" pageNumber="173">Fig. 17</figureCitation>
): Postoc and D-1 fused to a longish platelet, Dgl-3-6 separate, D-2-5 sclerotized, D-2 and -5 as roundish platelets, D-3 and - 4 as longish platelets, V-1-3 sclerotized. Coxal field (
<figureCitation id="59920C99FF93CF38DA3DFD72FCB9A21C" captionStart="FIGURE" captionStartId="26.[370,382,1835,1854]" captionTargetBox="[230,1281,400,1774]" captionTargetId="figure-7@26.[189,1399,388,1803]" captionTargetPageId="26" captionText="FIGURE 18: Atractides omanensis n. sp., female: idiosoma, ventral view. Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404120" httpUri="https://zenodo.org/record/5404120/files/figure.png" pageId="23" pageNumber="173">Fig. 18</figureCitation>
): posterior margin of Cx-I truncated, apodemes of Cx-II forming an acute angle, coxae with borders of secondary sclerite; L 319, Cx-III W 380, Cx-I+II mL 134, Cx-I+II lL 233. Palp (
<figureCitation id="59920C99FF93CF38DBC8FCE1FAE1A2B3" box="[1181,1295,884,908]" captionStart="FIGURE" captionStartId="27.[159,171,1562,1581]" captionTargetBox="[189,1399,302,1530]" captionTargetId="figure-4@27.[189,1399,302,1530]" captionTargetPageId="27" captionText="FIGURE 19: Atractides omanensis n. sp., female: (A) – capitulum and palp, lateral view; (B) – palp, medial view; (C) – I-Leg- 5 and - 6; (D) – chelicera. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404122" httpUri="https://zenodo.org/record/5404122/files/figure.png" pageId="23" pageNumber="173">Figs. 19B</figureCitation>
): palp total L 274, dL and %L (in parentheses): P-1, 32 (11.7); P-2, 62 (22.6); P-3, 63 (23.0); P-4, 82 (29.9); P-5, 35 (12.8); L P-2/P-4 0.76; chelicera (
<figureCitation id="59920C99FF93CF38DB93FC48FADAA2CA" box="[1222,1332,989,1013]" captionStart="FIGURE" captionStartId="27.[159,171,1562,1581]" captionTargetBox="[189,1399,302,1530]" captionTargetId="figure-4@27.[189,1399,302,1530]" captionTargetPageId="27" captionText="FIGURE 19: Atractides omanensis n. sp., female: (A) – capitulum and palp, lateral view; (B) – palp, medial view; (C) – I-Leg- 5 and - 6; (D) – chelicera. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404122" httpUri="https://zenodo.org/record/5404122/files/figure.png" pageId="23" pageNumber="173">Fig. 19D</figureCitation>
) total L 292, basal segment L 195, claw L 108, L ratio basal segment/claw 1.8; capitulum (
<figureCitation id="59920C99FF93CF38DBFFFBB6FAF9A504" box="[1194,1303,1059,1083]" captionStart="FIGURE" captionStartId="27.[159,171,1562,1581]" captionTargetBox="[189,1399,302,1530]" captionTargetId="figure-4@27.[189,1399,302,1530]" captionTargetPageId="27" captionText="FIGURE 19: Atractides omanensis n. sp., female: (A) – capitulum and palp, lateral view; (B) – palp, medial view; (C) – I-Leg- 5 and - 6; (D) – chelicera. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404122" httpUri="https://zenodo.org/record/5404122/files/figure.png" pageId="23" pageNumber="173">Fig. 19A</figureCitation>
) ventral L 129. Genital field (
<figureCitation id="59920C99FF93CF38DB4CFBD3FB81A561" box="[1049,1135,1094,1118]" captionStart="FIGURE" captionStartId="26.[370,382,1835,1854]" captionTargetBox="[230,1281,400,1774]" captionTargetId="figure-7@26.[189,1399,388,1803]" captionTargetPageId="26" captionText="FIGURE 18: Atractides omanensis n. sp., female: idiosoma, ventral view. Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404120" httpUri="https://zenodo.org/record/5404120/files/figure.png" pageId="23" pageNumber="173">Fig. 18</figureCitation>
): L/W 137/153, pregenital robust, halfmoon-shaped, L 117, genital plate L 75, L Ac-1-3: 24, 24, 27; egg maximum diameter (n=6) 113-118; excretory pore sclerotized; Vgl-1 fused to Vgl-2.
</paragraph>
<paragraph id="C116101CFF93CF38DC35FB69FC9EA49F" blockId="23.[826,1429,311,1949]" pageId="23" pageNumber="173">
I-Leg (
<figureCitation id="59920C99FF93CF38DCFAFB6EFBF2A42B" box="[943,1052,1275,1300]" captionStart="FIGURE" captionStartId="27.[159,171,1562,1581]" captionTargetBox="[189,1399,302,1530]" captionTargetId="figure-4@27.[189,1399,302,1530]" captionTargetPageId="27" captionText="FIGURE 19: Atractides omanensis n. sp., female: (A) – capitulum and palp, lateral view; (B) – palp, medial view; (C) – I-Leg- 5 and - 6; (D) – chelicera. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404122" httpUri="https://zenodo.org/record/5404122/files/figure.png" pageId="23" pageNumber="173">Fig. 19C</figureCitation>
): I-Leg-5 dL 111, vL 86, dL/vL 1.29, HB 34, dL/HB 3.3, S-1 L 42, L/W 9.1, S-2 L 42, L/W 9.1, distance of sword setae at I-Leg-5 0, L S- 1/2 1.0; I-Leg-6 L 106, HB 23, L/HB 4.6; L I-Leg-5/6 1.05.
</paragraph>
<paragraph id="C116101CFF93CF38DC35FA24FA69A618" blockId="23.[826,1429,311,1949]" pageId="23" pageNumber="173">
Remarks — Due to the presence of dorsal shield and the ventral surface covered by an extended shield including coxae, Vgl-3 and -4, excretory pore and genital field in the male, and gnathosoma with elongated rostrum, the new species is similar to
<taxonomicName id="06A96B9FFF93CF38DC6FF9F5FB34A747" authority="Cook, 1967" authorityName="Cook" authorityYear="1967" box="[826,1242,1632,1656]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="173" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="F3DDCC0EFF93CF38DC6FF9F5FC05A747" box="[826,1003,1632,1656]" italics="true" pageId="23" pageNumber="173">Atractides yukii</emphasis>
<bibRefCitation id="A5386DEDFF93CF38DCA3F9F5FB93A747" author="Cook, D. R." box="[1014,1149,1632,1656]" pageId="23" pageNumber="173" pagination="1 - 411" refId="ref17836" refString="Cook, D. R. 1967 - Water mites from India - Mem. Amer. Ent. Inst., 9: 1 - 411." type="journal article" year="1967">Cook, 1967</bibRefCitation>
(
<collectingCountry id="B9BE508CFF93CF38DBC5F9F5FB3CA747" box="[1168,1234,1632,1656]" name="India" pageId="23" pageNumber="173">India</collectingCountry>
)
</taxonomicName>
. From the latter it can be distinguished in the following features: gnathosomal rostrum more slender and pointed, P- 3 ventral margins with one seta, sword seta on P-4 enlarged, not hair-like, setae S-1 and -2 inserted on the same level and presence of dorsal leg clawlets.
</paragraph>
<paragraph id="C116101CFF93CF38DC35F8ADFC03A64B" blockId="23.[826,1429,311,1949]" pageId="23" pageNumber="173">
Etymology — The species is named for its occurrence in
<collectingCountry id="B9BE508CFF93CF38DCC8F8C9FC07A64B" box="[925,1001,1884,1908]" name="Oman" pageId="23" pageNumber="173">Oman</collectingCountry>
.
</paragraph>
<paragraph id="C116101CFF93CF38DC35F811FB89A6A2" blockId="23.[826,1429,311,1949]" box="[864,1127,1924,1949]" pageId="23" pageNumber="173">
Distribution —
<collectingCountry id="B9BE508CFF93CF38DB42F810FB8DA6A2" box="[1047,1123,1925,1949]" name="Oman" pageId="23" pageNumber="173">Oman</collectingCountry>
.
</paragraph>
<paragraph id="C116101CFF9CCF37DFCAFF6FFEB3A032" blockId="24.[159,349,250,269]" box="[159,349,250,269]" pageId="24" pageNumber="174">Smit H. and Pe˘si´c V.</paragraph>
</subSubSection>
</treatment>
</document>

202
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<emphasis id="F3DDCC0EFF99CF32DFE9F8DEFED6A65C" bold="true" box="[188,312,1867,1891]" italics="true" pageId="29" pageNumber="179">Axonopsis</emphasis>
(
<emphasis id="F3DDCC0EFF99CF32DE1DF8DFFDE8A65D" bold="true" box="[328,518,1866,1890]" italics="true" pageId="29" pageNumber="179">Brachypodopsis</emphasis>
)
<emphasis id="F3DDCC0EFF99CF32DD43F8DEFD7DA65C" bold="true" box="[534,659,1867,1891]" italics="true" pageId="29" pageNumber="179">omanensis</emphasis>
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(
<figureCitation id="59920C99FF99CF32DE19F811FE2BA6A2" box="[332,453,1924,1949]" captionStart="FIGURE" captionStartId="30.[159,171,1888,1907]" captionTargetBox="[261,1333,334,1854]" captionTargetId="figure-7@30.[255,1333,307,1857]" captionTargetPageId="30" captionText="FIGURE 21: Axonopsis omanensis n. sp. (A-F male [A-B, D-F holotype, C paratype], G female): (A) – dorsal shield; (B, G) – idiosoma, ventral view; (C) – genital field; (D-E) – palp; (F) – I-Leg. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404126" httpUri="https://zenodo.org/record/5404126/files/figure.png" pageId="29" pageNumber="179">Figures 21</figureCitation>
A-G, 28B-C)
</paragraph>
<paragraph id="C116101CFF99CF32DC35FEA2FBA8A087" blockId="29.[826,1429,311,440]" pageId="29" pageNumber="179">
Type series —
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<typeStatus id="1E12AEBEFF99CF32DB48FEA2FB61A070" box="[1053,1167,311,335]" pageId="29" pageNumber="179" type="holotype">Holotype</typeStatus>
: male, stream
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,
<collectingCountry id="B9BE508CFF99CF32DCD9FECFFC39A04D" box="[908,983,346,370]" name="Oman" pageId="29" pageNumber="179">Oman</collectingCountry>
,
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, alt.
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: 2/2/0 (1/1/0 mounted), same data as holotype.
</paragraph>
<paragraph id="C116101CFF99CF32DC35FE40FCB6A36A" blockId="29.[826,1429,469,597]" pageId="29" pageNumber="179">
Other material —
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0/5/0 (0/1/0 mounted), unnamed stream crossing road to
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, near
<collectingMunicipality id="21728A66FF99CF32DA36FE6DFA7EA32F" box="[1379,1424,504,528]" pageId="29" pageNumber="179">Ray</collectingMunicipality>
,
<collectingCountry id="B9BE508CFF99CF32DC6FFD8EFC6BA30C" box="[826,901,539,563]" name="Oman" pageId="29" pageNumber="179">Oman</collectingCountry>
,
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<geoCoordinate id="A49D76DBFF99CF32DBF2FD8EFADEA30D" box="[1191,1328,539,563]" degrees="56" direction="east" minutes="06.760" orientation="longitude" pageId="29" pageNumber="179" precision="1" value="56.112667">56°06.760 E</geoCoordinate>
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<paragraph id="C116101CFF99CF32DC35FDE6FC27A262" blockId="29.[826,1429,627,861]" pageId="29" pageNumber="179">Diagnosis — Colour pattern consisting of a light yellowish central area; dorsal shield with an weakly defined ridge on each side extending anterolaterally to the region of the anterior muscle scars; second, third and fourth legs with swimming setae (III-Leg- 5 with 4 swimming setae; IV-Leg-5 with 3 swimming setae).</paragraph>
<paragraph id="C116101CFF99CF32DC35FCEFFC07A2AD" blockId="29.[864,1001,890,914]" box="[864,1001,890,914]" pageId="29" pageNumber="179">Description</paragraph>
<paragraph id="C116101CFF99CF30DC35FC3AFD5FA04D" blockId="29.[826,1429,943,1949]" lastBlockId="31.[159,762,311,1684]" lastPageId="31" lastPageNumber="181" pageId="29" pageNumber="179">
Male — Dorsal and ventral shields anteriorly fused; dorsal shield L 341 (341), W 263 (269); eye pigment well developed; dorsal shield with seven pairs of glandularia (the seventh pair of glandularia inconspicuous, flanking the excretory pore); postocularia well distanced from anterior margin; colour pattern consisting of a light yellowish central area (
<figureCitation id="59920C99FF99CF32DCFFFB31FBE1A583" box="[938,1039,1188,1212]" captionStart="FIGURE" captionStartId="39.[159,171,1329,1348]" captionTargetBox="[237,1351,302,1297]" captionTargetId="figure-4@39.[237,1351,302,1297]" captionTargetPageId="39" captionText="FIGURE 28: Dorsal shield: (A) – Axonopsis quadarramensis sensu Pe˘si´c &amp; Gerecke, 2003; (B) – A. omanensis n. sp., male; (C) – A. omanensis n. sp., female; (D) – A. arabica n. sp., female; (E) – A. balneatoris n. sp., male; (F) – Omanaxonopsis arabica n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.5404140" httpUri="https://zenodo.org/record/5404140/files/figure.png" pageId="29" pageNumber="179">Fig. 28B</figureCitation>
); dorsal shield with a weakly defined ridge on each side extending anterolaterally in the region of the anterior muscle scars (shown as broken lines on
<figureCitation id="59920C99FF99CF32DCA3FA98FBB0A41A" box="[1014,1118,1293,1317]" captionStart="FIGURE" captionStartId="30.[159,171,1888,1907]" captionTargetBox="[261,1333,334,1854]" captionTargetId="figure-7@30.[255,1333,307,1857]" captionTargetPageId="30" captionText="FIGURE 21: Axonopsis omanensis n. sp. (A-F male [A-B, D-F holotype, C paratype], G female): (A) – dorsal shield; (B, G) – idiosoma, ventral view; (C) – genital field; (D-E) – palp; (F) – I-Leg. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404126" httpUri="https://zenodo.org/record/5404126/files/figure.png" pageId="29" pageNumber="179">Fig. 21A</figureCitation>
); excretory pore located at posterior end of dorsal shield; ventral shield L 322 (326), W (322); capitular bay L 87 (82); two pairs of glandularia lying between the genital field and insertions of the fourth legs, these relatively close together; three pairs of acetabula, arranged in an arc; width between most lateral pair of acetabula 104 (111); gonopore W 13 (15); palp (
<figureCitation id="59920C99FF99CF32DB90F996FAF7A724" box="[1221,1305,1539,1563]" captionStart="FIGURE" captionStartId="30.[159,171,1888,1907]" captionTargetBox="[261,1333,334,1854]" captionTargetId="figure-7@30.[255,1333,307,1857]" captionTargetPageId="30" captionText="FIGURE 21: Axonopsis omanensis n. sp. (A-F male [A-B, D-F holotype, C paratype], G female): (A) – dorsal shield; (B, G) – idiosoma, ventral view; (C) – genital field; (D-E) – palp; (F) – I-Leg. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404126" httpUri="https://zenodo.org/record/5404126/files/figure.png" pageId="29" pageNumber="179">Fig. 21</figureCitation>
D-E): palp total L 184 (179), dL: P-1, 27 (28); P-2, 40 (39); P-3, 26 (25); P-4, 66 (64); P-5, 25 (23); %L: P-1, 14.7 (15.6); P-2, 21.7 (21.8); P-3, 14.1 (14.0); P-4, 35.9 (35.8); P-5, 13.6 (12.9); L P-2/P-4 ratio 0.61 (0.61); ventral margin of P-2 convex, distal margin of P-3 with well developed hyaline extensions, P-4 ventrally dilated near insertions of a pair of setae (one hair-like and one relatively heavy seta), and with a more robust spine near distal margin of the segment. Legs: L of I-Leg-3-6: 26 (28), 39 (45), 62 (62), 68 (66); L of IV-Leg-2-6: 65 (66), 44 (51), 68 (65), 79 (78), 69 (74); swimming setae distributed as follows: II-Leg-5 2, III-Leg-4 2, III-Leg-5 4, IV-Leg-4 2, IV-Leg-5 3.
</paragraph>
<caption id="95D64094FF9ACF31DFCAF8F5FC77A6B0" ID-DOI="http://doi.org/10.5281/zenodo.5404126" ID-Zenodo-Dep="5404126" httpUri="https://zenodo.org/record/5404126/files/figure.png" pageId="30" pageNumber="180" startId="30.[159,171,1888,1907]" targetBox="[261,1333,334,1854]" targetPageId="30">
<paragraph id="C116101CFF9ACF31DFCAF8F5FC77A6B0" blockId="30.[159,1429,1888,1935]" pageId="30" pageNumber="180">
FIGURE 21:
<taxonomicName id="06A96B9FFF9ACF31DE40F8F5FE21A64D" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[277,463,1888,1907]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="30" pageNumber="180" phylum="Arthropoda" rank="species" species="omanensis" status="sp. nov.">
<emphasis id="F3DDCC0EFF9ACF31DE40F8F5FE21A64D" box="[277,463,1888,1907]" italics="true" pageId="30" pageNumber="180">Axonopsis omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF9ACF31DE83F8F5FDE0A64C" box="[470,526,1888,1907]" pageId="30" pageNumber="180" rank="species">n. sp.</taxonomicNameLabel>
(A-F male [A-B, D-F holotype, C paratype], G female): (A) – dorsal shield; (B, G) – idiosoma, ventral view; (C) – genital field; (D-E) – palp; (F) – I-Leg. Scale bars = 100 µm.
</paragraph>
</caption>
<paragraph id="C116101CFF9BCF30DF90FE12FE46A3E4" blockId="31.[159,762,311,1684]" pageId="31" pageNumber="181">
Female — similar to male, except in shape of the genital field (
<figureCitation id="59920C99FF9BCF30DE3EFE3FFE39A0FD" box="[363,471,426,450]" captionStart="FIGURE" captionStartId="30.[159,171,1888,1907]" captionTargetBox="[261,1333,334,1854]" captionTargetId="figure-7@30.[255,1333,307,1857]" captionTargetPageId="30" captionText="FIGURE 21: Axonopsis omanensis n. sp. (A-F male [A-B, D-F holotype, C paratype], G female): (A) – dorsal shield; (B, G) – idiosoma, ventral view; (C) – genital field; (D-E) – palp; (F) – I-Leg. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404126" httpUri="https://zenodo.org/record/5404126/files/figure.png" pageId="31" pageNumber="181">Fig. 21G</figureCitation>
); dorsal shield L 366, W 266; ventral shield L 350, W 331; capitular bay L 83; width between most lateral pair of acetabula 109; gonopore W 38; palp: total L 179, dL and %L (in parentheses): P-1, 27 (15.1); P-2, 40 (22.3); P-3, 25 (14.0); P-4, 63 (35.2); P-5, 24 (13.4); L P-2/P-4 ratio 0.64. Legs: L of I-Leg-2-6: 30, 29, 45, 59, 60; L of IV- Leg-4-6: 61, 72, 71; number of swimming setae on the legs as in the male.
</paragraph>
<paragraph id="C116101CFF9BCF30DF90FD65FEBCA214" blockId="31.[159,762,311,1684]" pageId="31" pageNumber="181">
Etymology — The species is named for its occurrence in
<collectingCountry id="B9BE508CFF9BCF30DE57FC86FEA0A214" box="[258,334,787,811]" name="Oman" pageId="31" pageNumber="181">Oman</collectingCountry>
.
</paragraph>
<paragraph id="C116101CFF9BCF30DF90FCD4FD4DA41E" blockId="31.[159,762,311,1684]" pageId="31" pageNumber="181">
Remarks — Due to the presence of two pairs of glandularia lying between the genital field and insertions of the fourth legs and similar shape of the palp,
<taxonomicName id="06A96B9FFF9BCF30DE4EFC3FFDE6A2FD" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[283,520,938,962]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="181" phylum="Arthropoda" rank="species" species="omanensis" status="sp. nov.">
<emphasis id="F3DDCC0EFF9BCF30DE4EFC3FFDE6A2FD" box="[283,520,938,962]" italics="true" pageId="31" pageNumber="181">Axonopsis omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FF9BCF30DD43FC3FFD87A2FE" box="[534,617,938,961]" pageId="31" pageNumber="181" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="06A96B9FFF9BCF30DFCAFC58FDACA2DA" authority="Valdecasas, 1981" authorityName="Valdecasas" authorityYear="1981" box="[159,578,973,997]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="181" phylum="Arthropoda" rank="species" species="guadaramensis">
<emphasis id="F3DDCC0EFF9BCF30DFCAFC58FE86A2DA" box="[159,360,973,997]" italics="true" pageId="31" pageNumber="181">A. guadaramensis</emphasis>
<bibRefCitation id="A5386DEDFF9BCF30DE20FC58FDACA2DA" author="Valdecasas, A. G." box="[373,578,973,997]" pageId="31" pageNumber="181" pagination="254 - 258" refId="ref18472" refString="Valdecasas, A. G. 1981 - Axonopsis (Brachypodopsis) guadarramensis n. sp. from Central Spain (Acari, Parasitengona) - Arch. Hydrobiol., 91: 254 - 258." type="journal article" year="1981">Valdecasas, 1981</bibRefCitation>
</taxonomicName>
, a species described originally from Sierra de Guadarrama (Central Spain,
<bibRefCitation id="A5386DEDFF9BCF30DE7AFB86FE1BA514" author="Valdecasas, A. G." box="[303,501,1043,1067]" pageId="31" pageNumber="181" pagination="254 - 258" refId="ref18472" refString="Valdecasas, A. G. 1981 - Axonopsis (Brachypodopsis) guadarramensis n. sp. from Central Spain (Acari, Parasitengona) - Arch. Hydrobiol., 91: 254 - 258." type="journal article" year="1981">Valdecasas 1981</bibRefCitation>
), and later recorded from
<collectingCountry id="B9BE508CFF9BCF30DFB7FBA3FEDCA571" box="[226,306,1078,1102]" name="Greece" pageId="31" pageNumber="181">Greece</collectingCountry>
(Pe˘si´c and
<bibRefCitation id="A5386DEDFF9BCF30DE90FBA3FD9EA571" author="Gerecke R." box="[453,624,1078,1102]" pageId="31" pageNumber="181" pagination="563 - 576" refId="ref18121" refString="Pe ˘ si'c, V., Gerecke R. 2003 - Water mites of the genera Albaxona, Axonopsis, Barbaxonella and Erebaxonopsis (Acari, Hydrachnidia: Aturidae: Axonopsinae) from Central Europe and Mediterranean area - Arch. Hydrobiol., 139: 563 - 576." type="journal article" year="2003">Gerecke, 2003</bibRefCitation>
). The new species resembles specimens from
<collectingCountry id="B9BE508CFF9BCF30DD1EFBCCFD75A54E" box="[587,667,1113,1137]" name="Greece" pageId="31" pageNumber="181">Greece</collectingCountry>
also in the presence and distribution of swimming setae on the legs. However in
<taxonomicName id="06A96B9FFF9BCF30DEFAFB35FD9AA588" authorityName="Valdecasas" authorityYear="1981" box="[431,628,1183,1207]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="181" phylum="Arthropoda" rank="species" species="guadaramensis">
<emphasis id="F3DDCC0EFF9BCF30DEFAFB35FD9AA588" box="[431,628,1183,1207]" italics="true" pageId="31" pageNumber="181">A. guadaramensis</emphasis>
</taxonomicName>
, according to
<bibRefCitation id="A5386DEDFF9BCF30DFE9FB56FE65A5E4" author="Valdecasas, A. G." box="[188,395,1219,1243]" pageId="31" pageNumber="181" pagination="254 - 258" refId="ref18472" refString="Valdecasas, A. G. 1981 - Axonopsis (Brachypodopsis) guadarramensis n. sp. from Central Spain (Acari, Parasitengona) - Arch. Hydrobiol., 91: 254 - 258." type="journal article" year="1981">Valdecasas (1981)</bibRefCitation>
, all legs are without swimming setae. Most probably, the specimens from
<collectingCountry id="B9BE508CFF9BCF30DDFCFB73FD17A5C1" box="[681,761,1254,1278]" name="Greece" pageId="31" pageNumber="181">Greece</collectingCountry>
therefore represent a species new to science.
</paragraph>
<paragraph id="C116101CFF9BCF30DF90FAA3FDF9A758" blockId="31.[159,762,311,1684]" pageId="31" pageNumber="181">
The new species from
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differs from specimens from
<collectingCountry id="B9BE508CFF9BCF30DE74FACCFE9FA44E" box="[289,369,1369,1393]" name="Greece" pageId="31" pageNumber="181">Greece</collectingCountry>
in its characteristic colour pattern (compare
<figureCitation id="59920C99FF9BCF30DE42FAE9FE31A4AB" box="[279,479,1404,1428]" captionStart="FIGURE" captionStartId="39.[159,171,1329,1348]" captionTargetBox="[237,1351,302,1297]" captionTargetId="figure-4@39.[237,1351,302,1297]" captionTargetPageId="39" captionText="FIGURE 28: Dorsal shield: (A) – Axonopsis quadarramensis sensu Pe˘si´c &amp; Gerecke, 2003; (B) – A. omanensis n. sp., male; (C) – A. omanensis n. sp., female; (D) – A. arabica n. sp., female; (E) – A. balneatoris n. sp., male; (F) – Omanaxonopsis arabica n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.5404140" httpUri="https://zenodo.org/record/5404140/files/figure.png" pageId="31" pageNumber="181">Figs. 28A and B</figureCitation>
) and the ridge on each side of the dorsal shield in the region of the anterior muscle scars being less prominent (this ridge more prominent in the specimens from
<collectingCountry id="B9BE508CFF9BCF30DD70FA73FD94A4C1" box="[549,634,1510,1534]" name="Greece" pageId="31" pageNumber="181">Greece</collectingCountry>
). A further difference is found in the well delineated suture line visible between genital field and ventral shield in the male specimen from
<collectingCountry id="B9BE508CFF9BCF30DEEAF9DAFDFDA758" box="[447,531,1615,1639]" name="Greece" pageId="31" pageNumber="181">Greece</collectingCountry>
.
</paragraph>
<paragraph id="C116101CFF9BCF30DF90F9E9FE22A7AB" blockId="31.[159,762,311,1684]" box="[197,460,1660,1684]" pageId="31" pageNumber="181">
Distribution —
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.
</paragraph>
</subSubSection>
</treatment>
</document>

179
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<taxonomicName id="06A96B9FFF9BCF30DF9BF929FD6CA7EB" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[206,642,1724,1748]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="181" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov." subGenus="Brachypodopsis">
<emphasis id="F3DDCC0EFF9BCF30DF9BF929FEA4A7EB" bold="true" box="[206,330,1724,1748]" italics="true" pageId="31" pageNumber="181">Axonopsis</emphasis>
(
<emphasis id="F3DDCC0EFF9BCF30DE0FF929FDF6A7EB" bold="true" box="[346,536,1724,1748]" italics="true" pageId="31" pageNumber="181">Brachypodopsis</emphasis>
)
<emphasis id="F3DDCC0EFF9BCF30DD7DF929FD6CA7EB" bold="true" box="[552,642,1724,1748]" italics="true" pageId="31" pageNumber="181">arabica</emphasis>
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(
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A-E, 23A-B, 28D)
</paragraph>
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Type series —
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<typeStatus id="1E12AEBEFF9BCF30DE2CF8ABFE05A669" box="[377,491,1854,1878]" pageId="31" pageNumber="181" type="holotype">Holotype</typeStatus>
: female, stream
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,
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,
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,
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, alt.
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, near
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,
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, alt.
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.
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<paragraph id="C116101CFF9BCF30DC35FE3CFB26A389" blockId="31.[826,1429,311,1948]" pageId="31" pageNumber="181">Diagnosis — Two pairs of glandularia lying between the genital field and insertions of the fourth legs, the posterior pair closely approaching the genital field in the male; P-4 club-shaped, with a pair of hair like setae on its ventral margin; relatively large gonopore in female; II-leg-5 with two swimming setae, III-Leg-5 with two swimming setae, IV- Leg-5 with three swimming setae.</paragraph>
<paragraph id="C116101CFF9BCF30DC35FD5FFC07A3DD" blockId="31.[826,1429,311,1948]" box="[864,1001,714,738]" pageId="31" pageNumber="181">Description</paragraph>
<paragraph id="C116101CFF9BCF30DC35FD63FB28A4F3" blockId="31.[826,1429,311,1948]" pageId="31" pageNumber="181">
Female — Dorsal and ventral shields anteriorly fused; dorsal shield L 391, W 303, with seven pairs of glandularia (
<figureCitation id="59920C99FF9BCF30DCAFFCA9FB9DA26A" box="[1018,1139,828,853]" captionStart="FIGURE" captionStartId="32.[159,171,1888,1907]" captionTargetBox="[256,1315,319,1838]" captionTargetId="figure-7@32.[239,1347,307,1857]" captionTargetPageId="32" captionText="FIGURE 22: Axonopsis arabica n. sp., female: (A) – dorsal shield, (B) – idiosoma, ventral view; (C-D) – palp; (E) – I-Leg. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404128" httpUri="https://zenodo.org/record/5404128/files/figure.png" pageId="31" pageNumber="181">Fig. 22A</figureCitation>
); postocularia well distanced from anterior margin; dorsal shield without a colour pattern and without prominent ridges, eye pigment well developed; excretory pore located at posterior end of dorsal shield; ventral shield (
<figureCitation id="59920C99FF9BCF30DA3DFC5CFC88A53B" captionStart="FIGURE" captionStartId="32.[159,171,1888,1907]" captionTargetBox="[256,1315,319,1838]" captionTargetId="figure-7@32.[239,1347,307,1857]" captionTargetPageId="32" captionText="FIGURE 22: Axonopsis arabica n. sp., female: (A) – dorsal shield, (B) – idiosoma, ventral view; (C-D) – palp; (E) – I-Leg. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404128" httpUri="https://zenodo.org/record/5404128/files/figure.png" pageId="31" pageNumber="181">Fig. 22B</figureCitation>
) L 412, W 395; capitular bay L 111; two pairs of glandularia lying between the genital field and insertions of the fourth legs; three pairs of genital acetabula, these arranged in an arc; width between most lateral pair of acetabula 157; gonopore relatively large, W 101; palp (
<figureCitation id="59920C99FF9BCF30DB24FB0EFB24A58C" box="[1137,1226,1179,1203]" captionStart="FIGURE" captionStartId="32.[159,171,1888,1907]" captionTargetBox="[256,1315,319,1838]" captionTargetId="figure-7@32.[239,1347,307,1857]" captionTargetPageId="32" captionText="FIGURE 22: Axonopsis arabica n. sp., female: (A) – dorsal shield, (B) – idiosoma, ventral view; (C-D) – palp; (E) – I-Leg. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404128" httpUri="https://zenodo.org/record/5404128/files/figure.png" pageId="31" pageNumber="181">Fig. 22</figureCitation>
C-D): total L 217, dL and %L (in parentheses): P-1, 34 (15.7); P-2, 50 (23.0); P-3, 29 (13.4); P-4, 79 (36.4); P-5, 25 (11.5); L P-2/P-4 ratio 0.63; ventral margin of P-2 convex, P- 4 club-shaped, with a hair like setae on the ventral side. Legs: L of I-Leg-2-6 (
<figureCitation id="59920C99FF9BCF30DB2CFADFFB0CA45C" box="[1145,1250,1354,1379]" captionStart="FIGURE" captionStartId="32.[159,171,1888,1907]" captionTargetBox="[256,1315,319,1838]" captionTargetId="figure-7@32.[239,1347,307,1857]" captionTargetPageId="32" captionText="FIGURE 22: Axonopsis arabica n. sp., female: (A) – dorsal shield, (B) – idiosoma, ventral view; (C-D) – palp; (E) – I-Leg. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404128" httpUri="https://zenodo.org/record/5404128/files/figure.png" pageId="31" pageNumber="181">Fig. 22E</figureCitation>
): 36, 32, 55, 65, 62; L of IV-Leg: 52, 68, 59, 69, 85, 79; swimming setae numbers: II-Leg-5 2 (rather short), III-Leg-4 2, III-Leg-5 2, IV-Leg-4 2, IV-Leg-5 3.
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Male — similar to female, except in shape of the genital field (
<figureCitation id="59920C99FF9BCF30DC82F996FBAFA724" box="[983,1089,1539,1563]" captionStart="FIGURE" captionStartId="33.[311,323,1092,1111]" captionTargetBox="[217,1343,335,1029]" captionTargetId="figure-4@33.[189,1399,302,1060]" captionTargetPageId="33" captionText="FIGURE 23: Axonopsis arabica n. sp., male: (A) – idiosoma, ventral view; (B) – palp. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404130" httpUri="https://zenodo.org/record/5404130/files/figure.png" pageId="31" pageNumber="181">Fig. 23A</figureCitation>
); dorsal shield L 372, W 309; ventral shield L 372, W 378; capitular bay L 111; the posterior pair of glandularia lying between the genital field and insertions of the fourth legs closely approaching the genital field; width between most lateral pair of acetabula 112; gonopore W 23; palp (
<figureCitation id="59920C99FF9BCF30DA3DF927FC86A7D2" captionStart="FIGURE" captionStartId="33.[311,323,1092,1111]" captionTargetBox="[217,1343,335,1029]" captionTargetId="figure-4@33.[189,1399,302,1060]" captionTargetPageId="33" captionText="FIGURE 23: Axonopsis arabica n. sp., male: (A) – idiosoma, ventral view; (B) – palp. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404130" httpUri="https://zenodo.org/record/5404130/files/figure.png" pageId="31" pageNumber="181">Fig. 23B</figureCitation>
): total L 214, dL and %L (in parentheses): P-1, 36 (16.8); P-2, 49 (22.9); P-3, 28 (13.1); P-4, 78 (36.4); P-5, 23 (10.7); L P-2/P-4 ratio 0.63. Legs: L of I-Leg- 2-6: 37, 32, 57, 67, 63; L of IV-Leg: 54, 66, 63, 75, 88, 78; number of swimming setae of the legs as in the female.
</paragraph>
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FIGURE 22:
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<emphasis id="F3DDCC0EFFA4CF0FDE47F8F5FE40A64C" box="[274,430,1888,1907]" italics="true" pageId="32" pageNumber="182">Axonopsis arabica</emphasis>
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, female: (A) – dorsal shield, (B) – idiosoma, ventral view; (C-D) – palp; (E) – I-Leg. Scale bars = 100 µm.
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FIGURE 23:
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<emphasis id="F3DDCC0EFFA5CF0EDEFDFBD1FDAAA568" box="[424,580,1092,1111]" italics="true" pageId="33" pageNumber="183">Axonopsis arabica</emphasis>
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, male: (A) – idiosoma, ventral view; (B) – palp. Scale bars = 100 µm.
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Remarks — Due to the presence of two pairs of glandularia lying between the genital field and insertions of the fourth legs and swimming setae on the second, third and fourth legs,
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, resembles
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<emphasis id="F3DDCC0EFFA5CF0EDE22FAB0FDE4A403" box="[375,522,1317,1340]" italics="true" pageId="33" pageNumber="183">A. omanensis</emphasis>
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from which it can be distinguished in the following features: P- 4 club-shaped, not distally tapering, presence of a hair-like, not heavy seta on the ventral side of P-4, female gonopore relatively large and the posterior pair of glandularia lying between the genital field and insertions of the fourth legs shifted closely to the genital field in the male.
</paragraph>
<paragraph id="C116101CFFA5CF0EDF90F9DDFE0DA7BC" blockId="33.[159,762,1176,1714]" pageId="33" pageNumber="183">Etymology — The species is named for its occurrence in Arabian Peninsula.</paragraph>
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Distribution —
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.
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soari
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K. Viets, 1911
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<emphasis id="F3DDCC0EFF9FCF34DFB1F963FD85A632" bold="true" box="[228,619,1781,1806]" italics="true" pageId="27" pageNumber="177">Neumania (Neumania) indentata</emphasis>
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(
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A-E)
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Type material —
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male, unnamed stream crossing road tyo
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/0, same data as holotype; 0/2/0, polls
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<paragraph id="C116101CFF9FCF34DC35F8BFFBF2A65A" blockId="27.[826,1428,1834,1893]" pageId="27" pageNumber="177">Diagnosis — Male with IV-Legs modified in shape and setation.</paragraph>
<paragraph id="C116101CFF9FCF34DC35F811FC07A6A3" blockId="27.[864,1001,1924,1948]" box="[864,1001,1924,1948]" pageId="27" pageNumber="177">Description</paragraph>
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FIGURE 20:
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<emphasis id="F3DDCC0EFF98CF33DE45F8F5FE51A64C" box="[272,447,1888,1907]" italics="true" pageId="28" pageNumber="178">Neumania indentata</emphasis>
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(A-D: male, E: female): (A, E) – idiosoma, ventral view; (B) – palp; (C) – I-Leg-4-6; (D) – IV-Leg-4-6. Scale bars = 50 µm (A-E).
</paragraph>
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<paragraph id="C116101CFF99CF32DF90FEA2FD14A2F9" blockId="29.[159,762,311,1698]" pageId="29" pageNumber="179">
Male — Idiosoma dorsally soft, ventrally sclerotized; L ventrally 384, W 294, dorsal idiosoma L 373. Coxal plates reticulated, posteromedial corner of Cx-IV without such reticulation. Apodemes of Cx-I long, reaching onto Cx-IV (
<figureCitation id="59920C99FF99CF32DD47FE56FD94A0E4" box="[530,634,451,475]" captionStart="FIGURE" captionStartId="28.[159,171,1888,1907]" captionTargetBox="[237,1335,307,1857]" captionTargetId="figure-7@28.[224,1364,307,1857]" captionTargetPageId="28" captionText="FIGURE 20: Neumania indentata n. sp. (A-D: male, E: female): (A, E) – idiosoma, ventral view; (B) – palp; (C) – I-Leg-4-6; (D) – IV-Leg-4-6. Scale bars = 50 µm (A-E)." figureDoi="http://doi.org/10.5281/zenodo.5404124" httpUri="https://zenodo.org/record/5404124/files/figure.png" pageId="29" pageNumber="179">Fig. 20A</figureCitation>
). Cx-II and Cx-IV laterally with stout seta. Gonopore L 40. Genital field with seven pairs of acetabula. Palp (
<figureCitation id="59920C99FF99CF32DD98FD9CFF20A37B" captionStart="FIGURE" captionStartId="28.[159,171,1888,1907]" captionTargetBox="[237,1335,307,1857]" captionTargetId="figure-7@28.[224,1364,307,1857]" captionTargetPageId="28" captionText="FIGURE 20: Neumania indentata n. sp. (A-D: male, E: female): (A, E) – idiosoma, ventral view; (B) – palp; (C) – I-Leg-4-6; (D) – IV-Leg-4-6. Scale bars = 50 µm (A-E)." figureDoi="http://doi.org/10.5281/zenodo.5404124" httpUri="https://zenodo.org/record/5404124/files/figure.png" pageId="29" pageNumber="179">Fig. 20B</figureCitation>
): total L 203; L: P-1, 17; P-2, 60; P-3, 38; P-4, 64; P-5, 24; P-4 with a short anteroventral tubercle. L of I-Leg-4-6: 114, 130, 130. L of IV-Leg-4-6: 104, 136, 94. Ventral margin of IV-Leg-5 proximally with a triangular projection, in distal 2/3 concave, with five strong, broad setae, one of these very broad, with denticulated tip. Ventral margin of IV-Leg-6 proximally undulating, in the centre indented (
<figureCitation id="59920C99FF99CF32DDD3FCB7FD00A205" box="[646,750,802,826]" captionStart="FIGURE" captionStartId="28.[159,171,1888,1907]" captionTargetBox="[237,1335,307,1857]" captionTargetId="figure-7@28.[224,1364,307,1857]" captionTargetPageId="28" captionText="FIGURE 20: Neumania indentata n. sp. (A-D: male, E: female): (A, E) – idiosoma, ventral view; (B) – palp; (C) – I-Leg-4-6; (D) – IV-Leg-4-6. Scale bars = 50 µm (A-E)." figureDoi="http://doi.org/10.5281/zenodo.5404124" httpUri="https://zenodo.org/record/5404124/files/figure.png" pageId="29" pageNumber="179">Fig. 20D</figureCitation>
). First and second leg with long fluted or grooved setae. Swimming setae numbers: III-Leg-4 six, III- Leg-5 two, IV-Leg-3 one, IV-Leg-4 two and IV-Leg-5 three. Claws of legs without clawlet and claw blade.
</paragraph>
<paragraph id="C116101CFF99CF32DF90FC4EFDF8A4A7" blockId="29.[159,762,311,1698]" pageId="29" pageNumber="179">
Female — Idiosoma soft, ventrally L 458 (530- 608), W 348 (429-551), dorsally L 454 (532-660). Apodemes of Cx-I extending to middle of Cx-IV (
<figureCitation id="59920C99FF99CF32DFF2FBD1FEFFA562" box="[167,273,1092,1117]" captionStart="FIGURE" captionStartId="28.[159,171,1888,1907]" captionTargetBox="[237,1335,307,1857]" captionTargetId="figure-7@28.[224,1364,307,1857]" captionTargetPageId="28" captionText="FIGURE 20: Neumania indentata n. sp. (A-D: male, E: female): (A, E) – idiosoma, ventral view; (B) – palp; (C) – I-Leg-4-6; (D) – IV-Leg-4-6. Scale bars = 50 µm (A-E)." figureDoi="http://doi.org/10.5281/zenodo.5404124" httpUri="https://zenodo.org/record/5404124/files/figure.png" pageId="29" pageNumber="179">Fig. 20E</figureCitation>
). Posterolateral corners of Cx-III and Cx- IV with a pair of stout setae. Coxal plates reticulated as in male. Genital field with 10 pairs of acetabula. Palp: total L 290; L: P-1, 26; P-2, 92; P-3, 50; P-4, 90; P-5, 32; palp as in male. L of I-Leg-4-6: 170, 168, 142. L of IV-Leg-4-6: 134, 162, 140. First and second legs with long grooved or fluted setae. Swimming setae numbers: III-Leg-3 one, III-Leg-4 five, III-Leg-5 two, IV-Leg-3 two, IV-Leg-4 and -5 three. Claws of legs without clawlet and claw blade.
</paragraph>
<paragraph id="C116101CFF99CF32DF90FA38FDC1A4D7" blockId="29.[159,762,311,1698]" pageId="29" pageNumber="179">Etymology — Named for the indented sixth segment of the fourth leg of the male.</paragraph>
<paragraph id="C116101CFF99CF32DF90FA6BFEACA79D" blockId="29.[159,762,311,1698]" pageId="29" pageNumber="179">
Remarks — The shape and chaetotaxy of the fourth leg is not found in any other
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<emphasis id="F3DDCC0EFF99CF32DDDCF9B4FD17A707" box="[649,761,1569,1592]" italics="true" pageId="29" pageNumber="179">Neumania</emphasis>
species.
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The female is not characteristic, and very likely not separable from related species (e.g.
<emphasis id="F3DDCC0EFF99CF32DDFAF9FDFF35A79D" italics="true" pageId="29" pageNumber="179">N. ambigua</emphasis>
Piersig).
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<treatment id="4900A10AFFA0CF06DCCDF951FD97A3B4" LSID="urn:lsid:plazi:treatment:4900A10AFFA0CF06DCCDF951FD97A3B4" httpUri="http://treatment.plazi.org/id/4900A10AFFA0CF06DCCDF951FD97A3B4" lastPageId="41" lastPageNumber="189" pageId="36" pageNumber="186">
<subSubSection id="89B34397FFA0CF0BDCCDF951FAD9A7E3" box="[920,1335,1731,1756]" pageId="36" pageNumber="186" type="nomenclature">
<paragraph id="C116101CFFA0CF0BDCCDF951FAD9A7E3" blockId="36.[920,1335,1731,1756]" box="[920,1335,1731,1756]" pageId="36" pageNumber="186">
<heading id="9A5EA770FFA0CF0BDCCDF951FAD9A7E3" bold="true" box="[920,1335,1731,1756]" centered="true" fontSize="10" level="2" pageId="36" pageNumber="186" reason="2">
<emphasis id="F3DDCC0EFFA0CF0BDCCDF951FAD9A7E3" bold="true" box="[920,1335,1731,1756]" pageId="36" pageNumber="186">
<taxonomicName id="06A96B9FFFA0CF0BDCCDF951FB00A7E4" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[920,1262,1731,1756]" class="Arachnida" family="Arrenuridae" genus="Arrenurus" kingdom="Animalia" order="Trombidiformes" pageId="36" pageNumber="186" phylum="Arthropoda" rank="species" species="dugesi" status="sp. nov." subGenus="Arrenurus">
<emphasis id="F3DDCC0EFFA0CF0BDCCDF951FB00A7E4" bold="true" box="[920,1262,1731,1756]" italics="true" pageId="36" pageNumber="186">Arrenurus (Arrenurus) dugesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA0CF0BDBA1F951FAD9A7E3" box="[1268,1335,1732,1756]" pageId="36" pageNumber="186" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89B34397FFA0CF06DB59F969FD97A3B4" lastPageId="41" lastPageNumber="191" pageId="36" pageNumber="186" type="description">
<paragraph id="C116101CFFA0CF0BDB59F969FB2CA62B" blockId="36.[1036,1218,1788,1812]" box="[1036,1218,1788,1812]" pageId="36" pageNumber="186">
(
<figureCitation id="59920C99FFA0CF0BDB41F969FB60A62B" box="[1044,1166,1788,1812]" captionStart="FIGURE" captionStartId="40.[159,171,1888,1907]" captionTargetBox="[271,1305,316,1820]" captionTargetId="figure-7@40.[249,1338,307,1857]" captionTargetPageId="40" captionText="FIGURE 29: Arrenurus dugesi n. sp. (A-E: male, F: female): (A) – idiosoma, dorsal view; (B, F) – idiosoma, ventral view; (C) – petiole, ventral view; (D) – petiole, lateral view; (E) – palp. Scale bars = 200 µm (A-B, F), 50 µm (C-E)." figureDoi="http://doi.org/10.5281/zenodo.5404142" httpUri="https://zenodo.org/record/5404142/files/figure.png" pageId="36" pageNumber="186">Figures 29</figureCitation>
A-F)
</paragraph>
<paragraph id="C116101CFFA0CF0BDC35F8AAFAC2A6A3" blockId="36.[826,1429,1854,1949]" pageId="36" pageNumber="186">
Type material —
<materialsCitation id="71C11A41FFA0CF0BDB66F8ABFA7FA646" collectingDate="2008-11-04" country="Oman" latitude="17.053934" location="Wadi Hanna" longLatPrecision="1" longitude="54.60815" pageId="36" pageNumber="186" specimenCount="1" typeStatus="holotype">
<typeStatus id="1E12AEBEFFA0CF0BDB66F8ABFB4CA669" box="[1075,1186,1854,1878]" pageId="36" pageNumber="186" type="holotype">Holotype</typeStatus>
male,
<location id="C47646C7FFA0CF0BDBA2F8ABFA7EA669" LSID="urn:lsid:plazi:treatment:4900A10AFFA0CF06DCCDF951FD97A3B4:C47646C7FFA0CF0BDBA2F8ABFA7EA669" box="[1271,1424,1854,1878]" country="Oman" latitude="17.053934" longLatPrecision="1" longitude="54.60815" name="Wadi Hanna" pageId="36" pageNumber="186">Wadi Hanna</location>
,
<collectingCountry id="B9BE508CFFA0CF0BDC6FF8F7FC6BA645" box="[826,901,1890,1914]" name="Oman" pageId="36" pageNumber="186">Oman</collectingCountry>
,
<geoCoordinate id="A49D76DBFFA0CF0BDCFEF8F4FBA5A646" box="[939,1099,1889,1913]" degrees="17" direction="north" minutes="03.236" orientation="latitude" pageId="36" pageNumber="186" precision="1" value="17.053934">17°03.236 N</geoCoordinate>
<geoCoordinate id="A49D76DBFFA0CF0BDB32F8F4FAEAA646" box="[1127,1284,1889,1913]" degrees="54" direction="east" minutes="36.489" orientation="longitude" pageId="36" pageNumber="186" precision="1" value="54.60815">54°36.489 E</geoCoordinate>
,
<date id="B51736DCFFA0CF0BDA71F8F4FA7FA646" box="[1316,1425,1889,1913]" pageId="36" pageNumber="186" value="2008-11-04">
<collectingDate id="A553CF34FFA0CF0BDA71F8F4FA7FA646" box="[1316,1425,1889,1913]" pageId="36" pageNumber="186" value="2008-11-04">4-xi-2008</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="1E12AEBEFFA0CF0BDC6FF810FC5CA6A2" box="[826,946,1925,1949]" pageId="36" pageNumber="186" type="paratype">Paratypes</typeStatus>
: 12/8/0, same data as holotype.
</paragraph>
<caption id="95D64094FFA1CF0ADFCAF8F5FD63A6B0" ID-DOI="http://doi.org/10.5281/zenodo.5404136" ID-Zenodo-Dep="5404136" httpUri="https://zenodo.org/record/5404136/files/figure.png" pageId="37" pageNumber="187" startId="37.[159,171,1888,1907]" targetBox="[247,1346,321,1826]" targetPageId="37">
<paragraph id="C116101CFFA1CF0ADFCAF8F5FD63A6B0" blockId="37.[159,1429,1888,1935]" pageId="37" pageNumber="187">
FIGURE 26:
<taxonomicName id="06A96B9FFFA1CF0ADE46F8F5FE0EA64C" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[275,480,1888,1907]" class="Arachnida" family="Aturidae" genus="Omanaxonopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="37" pageNumber="187" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="F3DDCC0EFFA1CF0ADE46F8F5FE0EA64C" box="[275,480,1888,1907]" italics="true" pageId="37" pageNumber="187">Omanaxonopsis arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA1CF0ADEB3F8F5FDF0A64C" box="[486,542,1888,1907]" pageId="37" pageNumber="187" rank="species">n. sp.</taxonomicNameLabel>
, female: (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – genital field; (D) – capitulum; (E-F) – palp; (G) – chelicera. Scale bars = 100 µm.
</paragraph>
</caption>
<caption id="95D64094FFA2CF09DE36FAF1FB3CA448" ID-DOI="http://doi.org/10.5281/zenodo.5404138" ID-Zenodo-Dep="5404138" box="[355,1234,1379,1399]" httpUri="https://zenodo.org/record/5404138/files/figure.png" pageId="38" pageNumber="188" startId="38.[355,367,1380,1399]" targetBox="[192,1363,308,1302]" targetPageId="38">
<paragraph id="C116101CFFA2CF09DE36FAF1FB3CA448" blockId="38.[355,1234,1379,1399]" box="[355,1234,1379,1399]" pageId="38" pageNumber="188">
FIGURE 27:
<taxonomicName id="06A96B9FFFA2CF09DE81FAF1FD4FA448" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[468,673,1380,1399]" class="Arachnida" family="Aturidae" genus="Omanaxonopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="38" pageNumber="188" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="F3DDCC0EFFA2CF09DE81FAF1FD4FA448" box="[468,673,1380,1399]" italics="true" pageId="38" pageNumber="188">Omanaxonopsis arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA2CF09DDF3FAF1FD35A448" box="[678,731,1380,1399]" pageId="38" pageNumber="188" rank="species">n. sp.</taxonomicNameLabel>
, female: (A) – I-Leg; (B) – IV-Leg. Scale bar = 100 µm.
</paragraph>
</caption>
<paragraph id="C116101CFFA2CF09DF90FA2DFF15A79D" blockId="38.[159,762,1464,1698]" pageId="38" pageNumber="188">Diagnosis — Male: cauda short; pygal lobes distinct; hyaline membrane with pointed angles; petiole rounded posteriorly; ligulate process of petiole rounded, lying well distanced from posterior margin of petiole. Female: Genital plates short, tapering laterally, sloping towards posterolateral idiosoma margin.</paragraph>
<paragraph id="C116101CFFA2CF09DF90F954FEA0A7E6" blockId="38.[197,334,1729,1753]" box="[197,334,1729,1753]" pageId="38" pageNumber="188">Description</paragraph>
<paragraph id="C116101CFFA2CF09DF90F96DFA7AA6A8" blockId="38.[159,762,1784,1948]" lastBlockId="38.[826,1429,1464,1944]" pageId="38" pageNumber="188">
Male — Idiosoma greenish, L (including petiole) 1073 (932-1154), W 802 (701-850), posterior margin slightly concave. D1 on small humps. Dorsal shield (
<figureCitation id="59920C99FFA2CF09DFF2F8F4FEF5A645" box="[167,283,1889,1914]" captionStart="FIGURE" captionStartId="40.[159,171,1888,1907]" captionTargetBox="[271,1305,316,1820]" captionTargetId="figure-7@40.[249,1338,307,1857]" captionTargetPageId="40" captionText="FIGURE 29: Arrenurus dugesi n. sp. (A-E: male, F: female): (A) – idiosoma, dorsal view; (B, F) – idiosoma, ventral view; (C) – petiole, ventral view; (D) – petiole, lateral view; (E) – palp. Scale bars = 200 µm (A-B, F), 50 µm (C-E)." figureDoi="http://doi.org/10.5281/zenodo.5404142" httpUri="https://zenodo.org/record/5404142/files/figure.png" pageId="38" pageNumber="188">Fig. 29A</figureCitation>
) W 437 (397-486). Cauda short, pygal lobes distinct. Hyaline membrane with pointed angles, posterior margin concave. Petiole (
<figureCitation id="59920C99FFA2CF09DA77FA2DFA63A4EF" box="[1314,1421,1464,1488]" captionStart="FIGURE" captionStartId="40.[159,171,1888,1907]" captionTargetBox="[271,1305,316,1820]" captionTargetId="figure-7@40.[249,1338,307,1857]" captionTargetPageId="40" captionText="FIGURE 29: Arrenurus dugesi n. sp. (A-E: male, F: female): (A) – idiosoma, dorsal view; (B, F) – idiosoma, ventral view; (C) – petiole, ventral view; (D) – petiole, lateral view; (E) – palp. Scale bars = 200 µm (A-B, F), 50 µm (C-E)." figureDoi="http://doi.org/10.5281/zenodo.5404142" httpUri="https://zenodo.org/record/5404142/files/figure.png" pageId="38" pageNumber="188">Fig. 29C</figureCitation>
) rounded posteriorly, ligulate process rounded, well distanced from posterior margin of petiole, uplifted only slightly. Petiole ventrally with two lateral rows of irregularly shaped denticles. Setae associated with petiole bifurcated, these setae longer than petiole. Cx-I not extending to anterior idiosoma margin (
<figureCitation id="59920C99FFA2CF09DC17F938FC40A7F9" box="[834,942,1709,1734]" captionStart="FIGURE" captionStartId="40.[159,171,1888,1907]" captionTargetBox="[271,1305,316,1820]" captionTargetId="figure-7@40.[249,1338,307,1857]" captionTargetPageId="40" captionText="FIGURE 29: Arrenurus dugesi n. sp. (A-E: male, F: female): (A) – idiosoma, dorsal view; (B, F) – idiosoma, ventral view; (C) – petiole, ventral view; (D) – petiole, lateral view; (E) – palp. Scale bars = 200 µm (A-B, F), 50 µm (C-E)." figureDoi="http://doi.org/10.5281/zenodo.5404142" httpUri="https://zenodo.org/record/5404142/files/figure.png" pageId="38" pageNumber="188">Fig. 29B</figureCitation>
). Gonopore L 58. Genital plates narrow, extending to lateral idiosoma margin. Palp (
<figureCitation id="59920C99FFA2CF09DA3DF945FC87A633" captionStart="FIGURE" captionStartId="40.[159,171,1888,1907]" captionTargetBox="[271,1305,316,1820]" captionTargetId="figure-7@40.[249,1338,307,1857]" captionTargetPageId="40" captionText="FIGURE 29: Arrenurus dugesi n. sp. (A-E: male, F: female): (A) – idiosoma, dorsal view; (B, F) – idiosoma, ventral view; (C) – petiole, ventral view; (D) – petiole, lateral view; (E) – palp. Scale bars = 200 µm (A-B, F), 50 µm (C-E)." figureDoi="http://doi.org/10.5281/zenodo.5404142" httpUri="https://zenodo.org/record/5404142/files/figure.png" pageId="38" pageNumber="188">Fig. 29E</figureCitation>
): total L 364; L: P-1, 40; P-2, 94; P-3, 70; P-4, 100; P-5, 60; P-2 with two setae in anteroventral corner and two medial setae more anterodorsally. L of I-Leg-4-6: 190, 186, 176. L of IV-Leg-4-6: 316, 114, 124; IV-Leg-4 with a long spur. Claws of legs with a
</paragraph>
<paragraph id="C116101CFFA3CF08DB33FF6FFA7BA032" blockId="39.[1126,1429,250,269]" box="[1126,1429,250,269]" pageId="39" pageNumber="189">
<bibCitation id="14831870FFA3CF08DB33FF6FFA7BA032" box="[1126,1429,250,269]" issue="2" journal="Acarologia" page="151" pageId="39" pageNumber="189" score="5.405330260433823" volume="50" year="2010">Acarologia 50(2): 151–195 (2010)</bibCitation>
</paragraph>
<caption id="95D64094FFA3CF08DFCAFAA4FAE6A460" ID-DOI="http://doi.org/10.5281/zenodo.5404140" ID-Zenodo-Dep="5404140" httpUri="https://zenodo.org/record/5404140/files/figure.png" pageId="39" pageNumber="189" startId="39.[159,171,1329,1348]" targetBox="[237,1351,302,1297]" targetPageId="39">
<paragraph id="C116101CFFA3CF08DFCAFAA4FAE6A460" blockId="39.[159,1429,1328,1376]" pageId="39" pageNumber="189">
FIGURE 28: Dorsal shield: (A) –
<emphasis id="F3DDCC0EFFA3CF08DE9AFAA4FD02A47C" box="[463,748,1328,1348]" italics="true" pageId="39" pageNumber="189">
<taxonomicName id="06A96B9FFFA3CF08DE9AFAA4FDC7A47B" authorityName="Piersig" authorityYear="1893" box="[463,553,1329,1348]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="39" pageNumber="189" phylum="Arthropoda" rank="genus">Axonopsis</taxonomicName>
quadarramensis sensu
</emphasis>
Pe˘si´c &amp;
<bibRefCitation id="A5386DEDFFA3CF08DC6BFAA5FC2CA47B" author="Gerecke R." box="[830,962,1328,1348]" pageId="39" pageNumber="189" pagination="563 - 576" refId="ref18121" refString="Pe ˘ si'c, V., Gerecke R. 2003 - Water mites of the genera Albaxona, Axonopsis, Barbaxonella and Erebaxonopsis (Acari, Hydrachnidia: Aturidae: Axonopsinae) from Central Europe and Mediterranean area - Arch. Hydrobiol., 139: 563 - 576." type="journal article" year="2003">Gerecke, 2003</bibRefCitation>
; (B) –
<taxonomicName id="06A96B9FFFA3CF08DCA9FAA4FB9EA47C" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[1020,1136,1329,1348]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="39" pageNumber="189" phylum="Arthropoda" rank="species" species="omanensis" status="sp. nov.">
<emphasis id="F3DDCC0EFFA3CF08DCA9FAA4FB9EA47C" box="[1020,1136,1329,1348]" italics="true" pageId="39" pageNumber="189">A. omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA3CF08DB20FAA4FB45A47B" box="[1141,1195,1329,1348]" pageId="39" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
, male; (C) –
<taxonomicName id="06A96B9FFFA3CF08DA74FAA4FA7BA47C" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[1313,1429,1329,1348]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="39" pageNumber="189" phylum="Arthropoda" rank="species" species="omanensis" status="sp. nov.">
<emphasis id="F3DDCC0EFFA3CF08DA74FAA4FA7BA47C" box="[1313,1429,1329,1348]" italics="true" pageId="39" pageNumber="189">A. omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA3CF08DF94FAD8FF18A45F" box="[193,246,1357,1376]" pageId="39" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
, female; (D) –
<taxonomicName id="06A96B9FFFA3CF08DE29FAD9FE3DA460" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[380,467,1356,1375]" class="Arachnida" family="Aturidae" genus="Omanaxonopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="39" pageNumber="189" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="F3DDCC0EFFA3CF08DE29FAD9FE3DA460" box="[380,467,1356,1375]" italics="true" pageId="39" pageNumber="189">A. arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA3CF08DE8CFAD8FDE0A45F" box="[473,526,1357,1376]" pageId="39" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
, female; (E) –
<taxonomicName id="06A96B9FFFA3CF08DDC4FAD9FCE7A460" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[657,777,1356,1375]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="39" pageNumber="189" phylum="Arthropoda" rank="species" species="balneatoris" status="sp. nov.">
<emphasis id="F3DDCC0EFFA3CF08DDC4FAD9FCE7A460" box="[657,777,1356,1375]" italics="true" pageId="39" pageNumber="189">A. balneatoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA3CF08DC58FAD8FCADA45F" box="[781,835,1357,1376]" pageId="39" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
, male; (F) –
<taxonomicName id="06A96B9FFFA3CF08DCE6FAD9FB6EA460" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[947,1152,1356,1375]" class="Arachnida" family="Aturidae" genus="Omanaxonopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="39" pageNumber="189" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="F3DDCC0EFFA3CF08DCE6FAD9FB6EA460" box="[947,1152,1356,1375]" italics="true" pageId="39" pageNumber="189">Omanaxonopsis arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA3CF08DBD0FAD8FB54A45F" box="[1157,1210,1357,1376]" pageId="39" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
, female.
</paragraph>
</caption>
<paragraph id="C116101CFFA3CF08DFCAFA34FE0DA4E4" blockId="39.[159,762,1441,1500]" pageId="39" pageNumber="189">clawlet as large as claw, third and fourth legs with numerous swimming setae.</paragraph>
<paragraph id="C116101CFFA3CF08DF90F996FB55A7B5" blockId="39.[159,762,1539,1949]" lastBlockId="39.[826,1429,1441,1948]" pageId="39" pageNumber="189">
Females — Idiosoma greenish-brownish, L 1045 (948-1175), W 932 (850-1037). Anterior margin of idiosoma straight, posterolateral corners of idiosoma indistinct. Dorsal shield complete, L 664 (599-774), W 543 (494-616). D1 on small humps, setae of D4 long. Cx-I plates extending beyond anterior margin of idiosoma. Medial margin of Cx-IV longer than medial margin of Cx-III. Medial distance of Cx- IV shorter than width of one genital valve. Gonopore L 164, width of gonopore 186, genital valves with small sclerotized patches. Genital plates short, shorter than width of gonopore, sloping towards posterolateral idiosoma margin (
<figureCitation id="59920C99FFA3CF08DBEFFA34FAF0A486" box="[1210,1310,1441,1465]" captionStart="FIGURE" captionStartId="40.[159,171,1888,1907]" captionTargetBox="[271,1305,316,1820]" captionTargetId="figure-7@40.[249,1338,307,1857]" captionTargetPageId="40" captionText="FIGURE 29: Arrenurus dugesi n. sp. (A-E: male, F: female): (A) – idiosoma, dorsal view; (B, F) – idiosoma, ventral view; (C) – petiole, ventral view; (D) – petiole, lateral view; (E) – palp. Scale bars = 200 µm (A-B, F), 50 µm (C-E)." figureDoi="http://doi.org/10.5281/zenodo.5404142" httpUri="https://zenodo.org/record/5404142/files/figure.png" pageId="39" pageNumber="189">Fig. 29F</figureCitation>
). Palp: total L 330; L: P-1, 32; P-2, 92; P-3, 62; P-4, 92; P-5, 52; P-2 with two anteromedial setae and two setae located more anterodorsally. L of I-Leg-4-6: 195, 160, 148. L of IV-Leg-4-6: 230, 182, 160. Claws of legs with a clawlet as large as claw, third and fourth legs with numerous swimming setae.
</paragraph>
<paragraph id="C116101CFFA3CF08DC35F931FB2CA63D" blockId="39.[826,1429,1441,1948]" pageId="39" pageNumber="189">
Etymology — Named after the French acarologist Antoine-Louis Delsecautz DugŁs (1797-1838), who erected the genus
<taxonomicName id="06A96B9FFFA3CF08DB1CF97EFB52A63D" authorityName="DugLs" authorityYear="1834" box="[1097,1212,1771,1794]" class="Arachnida" family="Arrenuridae" genus="Arrenurus" kingdom="Animalia" order="Trombidiformes" pageId="39" pageNumber="189" phylum="Arthropoda" rank="genus">
<emphasis id="F3DDCC0EFFA3CF08DB1CF97EFB52A63D" box="[1097,1212,1771,1794]" italics="true" pageId="39" pageNumber="189">Arrenurus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C116101CFFA3CF06DC35F88EFD97A3B4" blockId="39.[826,1429,1441,1948]" lastBlockId="41.[159,762,311,651]" lastPageId="41" lastPageNumber="191" pageId="39" pageNumber="189">
Remarks — Two
<taxonomicName id="06A96B9FFFA3CF08DB64F889FAEFA60C" box="[1073,1281,1819,1843]" class="Arachnida" family="Arrenuridae" genus="Arrenurus" kingdom="Animalia" order="Trombidiformes" pageId="39" pageNumber="189" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="F3DDCC0EFFA3CF08DB64F889FB4AA60C" box="[1073,1188,1820,1843]" italics="true" pageId="39" pageNumber="189">Arrenurus</emphasis>
species
</taxonomicName>
having also a petiole with a row of ventral teeth differ as follows:
<emphasis id="F3DDCC0EFFA3CF08DCD2F8F7FBD6A646" box="[903,1080,1889,1913]" italics="true" pageId="39" pageNumber="189">A. glenifferensis</emphasis>
Lundblad, 1941 from southern Africa has a long cauda and a triangular ligulate process,
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Mota¸s, 1927 from the Western Palaearctic has also a long cauda and a more angular-shaped petiole and ligulate process. The assignment of the female is uncertain, as
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of the females were collected on the
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locality. The most common of these have been assigned to the new species, the less common to
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Smit
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(see below). Moreover, the females here assigned to
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Smit
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share with males of the same number and arrangement of medial setae on P-2.
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<emphasis id="F3DDCC0EFFA0CF0BDCE7FEA2FB3AA071" bold="true" box="[946,1236,310,335]" italics="true" pageId="36" pageNumber="186">Omanaxonopsis arabica</emphasis>
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(
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A-G, 27A-B, 28F)
</paragraph>
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Type series —
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: female, spring
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, crossing road to
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border,
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<paragraph id="C116101CFFA0CF0BDC35FDB6FB76A303" blockId="36.[826,1429,433,1624]" box="[864,1176,547,572]" pageId="36" pageNumber="186">Diagnosis — As for genus.</paragraph>
<paragraph id="C116101CFFA0CF0BDC35FDDAFC07A358" blockId="36.[826,1429,433,1624]" box="[864,1001,591,615]" pageId="36" pageNumber="186">Description</paragraph>
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Female — Dorsal and ventral shields anteriorly fused; dorsal shield L 364, W 270; eye pigment well developed; dorsal shield (
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) with four pairs of glandularia; postocularia well distanced from anterior margin; excretory pore located at posterior end of dorsal shield; ventral shield (
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) L 378, W 331; lateral margins of ventral shield with an irregular truncate projection on each side located considerably posterior to the insertions of the fourth leg; capitular bay L 91; one pair of glandularia lying between the genital field and insertions of the fourth legs; acetabula difficult to see in the rugose integument but 5-6 pairs appear to be present, width between most lateral pair of acetabula 104; gonopore relatively large, W 42; chelicera (
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) total L 118, basal segment L 84, claw L 38, L ratio basal segment/claw 2.2; capitulum (
<figureCitation id="59920C99FFA0CF0BDBBCFB39FABFA5FB" box="[1257,1361,1196,1220]" captionStart="FIGURE" captionStartId="37.[159,171,1888,1907]" captionTargetBox="[247,1346,321,1826]" captionTargetId="figure-4@37.[242,1346,307,1857]" captionTargetPageId="37" captionText="FIGURE 26: Omanaxonopsis arabica n. sp., female: (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – genital field; (D) – capitulum; (E-F) – palp; (G) – chelicera. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404136" httpUri="https://zenodo.org/record/5404136/files/figure.png" pageId="36" pageNumber="186">Fig. 26D</figureCitation>
) ventral L 62; palp (
<figureCitation id="59920C99FFA0CF0BDCA7FB5AFBA9A5D8" box="[1010,1095,1231,1255]" captionStart="FIGURE" captionStartId="37.[159,171,1888,1907]" captionTargetBox="[247,1346,321,1826]" captionTargetId="figure-4@37.[242,1346,307,1857]" captionTargetPageId="37" captionText="FIGURE 26: Omanaxonopsis arabica n. sp., female: (A) – dorsal shield; (B) – idiosoma, ventral view; (C) – genital field; (D) – capitulum; (E-F) – palp; (G) – chelicera. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404136" httpUri="https://zenodo.org/record/5404136/files/figure.png" pageId="36" pageNumber="186">Fig. 26</figureCitation>
E-F): total L 196, dL and %L (in parentheses): P-1, 29 (14.8); P-2, 46 (23.5); P-3, 26 (13.3); P-4, 72 (36.7); P-5, 23 (11.7); L P-2/P-4 ratio 0.64; distoventral portion of P-2 expanded. Legs: L of I-Leg (
<figureCitation id="59920C99FFA0CF0BDCF0FACEFBE3A44B" box="[933,1037,1371,1396]" captionStart="FIGURE" captionStartId="38.[355,367,1380,1399]" captionTargetBox="[192,1363,308,1302]" captionTargetId="figure-7@38.[189,1399,302,1348]" captionTargetPageId="38" captionText="FIGURE 27: Omanaxonopsis arabica n. sp., female: (A) – I-Leg; (B) – IV-Leg. Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404138" httpUri="https://zenodo.org/record/5404138/files/figure.png" pageId="36" pageNumber="186">Fig. 27A</figureCitation>
): 32, 37, 32, 46, 63, 66; L of IV-Leg (
<figureCitation id="59920C99FFA0CF0BDC17FAEAFC45A4A8" box="[834,939,1407,1431]" captionStart="FIGURE" captionStartId="38.[355,367,1380,1399]" captionTargetBox="[192,1363,308,1302]" captionTargetId="figure-7@38.[189,1399,302,1348]" captionTargetPageId="38" captionText="FIGURE 27: Omanaxonopsis arabica n. sp., female: (A) – I-Leg; (B) – IV-Leg. Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404138" httpUri="https://zenodo.org/record/5404138/files/figure.png" pageId="36" pageNumber="186">Fig. 27B</figureCitation>
): 54, 68, 62, 69, 93, 80; IV-Leg-5 with two swimming hairs; the claws with dorsal and ventral clawlets.
</paragraph>
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<paragraph id="C116101CFFA0CF0BDC35F989FBF5A767" blockId="36.[826,1429,433,1624]" pageId="36" pageNumber="186">Etymology — Named for its occurrence on the Arabian peninsula.</paragraph>
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<heading id="9A5EA770FFA0CF0BDE67FB02FD88A590" bold="true" box="[306,614,1175,1199]" centered="true" fontSize="10" level="2" pageId="36" pageNumber="186" reason="2">
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<emphasis id="F3DDCC0EFFA0CF0BDE67FB02FE1DA590" bold="true" box="[306,499,1175,1199]" italics="true" pageId="36" pageNumber="186">Omanaxonopsis</emphasis>
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Diagnosis — Characters of the
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and the subfamily
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. Dorsal and ventral shields anteriorly fused; dorsal shield with four pairs of glandularia and a pair of postocularia; no ridge on each side extending anteriorly from area of insertion of fourth coxae; lateral margins of ventral shield with an irregular truncate projection on each side located considerably posterior to the insertions of the fourth leg; one pair of glandularia lying between the genital field and insertions of the fourth legs; genital field with 5-6 pairs of acetabula, acetabular plates fused with the ventral shield; leg claws with dorsal and ventral clawlets.
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<typeStatus id="1E12AEBEFFA0CF0BDF90F932FF10A780" box="[197,254,1703,1727]" pageId="36" pageNumber="186">Type</typeStatus>
species —
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<emphasis id="F3DDCC0EFFA0CF0BDED5F932FD91A781" box="[384,639,1702,1726]" italics="true" pageId="36" pageNumber="186">Omanaxonopsis arabica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA0CF0BDDD3F932FD28A781" box="[646,710,1703,1726]" pageId="36" pageNumber="186" rank="species">n. sp.</taxonomicNameLabel>
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Remarks — The new genus appears to be close to
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<emphasis id="F3DDCC0EFFA0CF0BDFE9F96CFEC2A62F" box="[188,300,1785,1808]" italics="true" pageId="36" pageNumber="186">Axonopsis</emphasis>
Piersig
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, but differs from it in the genital field with 5-6 pairs of acetabula, lateral margins of ventral shield with an irregular truncate projection on each side and leg claws with dorsal and ventral clawlets.
</paragraph>
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</treatment>
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199
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)
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(
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A-F, 25A-B, 28E)
</paragraph>
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Type series —
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<typeStatus id="1E12AEBEFFA5CF0EDE3EF8F4FE33A646" box="[363,477,1889,1913]" pageId="33" pageNumber="183" type="holotype">Holotype</typeStatus>
: male, stream
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,
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,
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,
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, alt.
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, leg.
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:
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1/0/0 (1/0/0 mounted), same data as holotype; 0/1/0 (0/1/0 mounted), stream
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,
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<geoCoordinate id="A49D76DBFFA5CF0EDC02FA94FC30A426" box="[855,990,1281,1305]" degrees="57" direction="east" minutes="11.996" orientation="longitude" pageId="33" pageNumber="183" precision="1" value="57.199932">57°11.996 E</geoCoordinate>
, alt.
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, leg.
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Diagnosis — Idiosoma slender (L/W about
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both sexes) with subparallel lateral margins; IV- Leg-6 strongly inflated distally (L/H 2.7-2.8), with concave dorsal margin in its proximal part.
</paragraph>
<paragraph id="C116101CFFA5CF0EDC35FA45FC07A4D7" blockId="33.[864,1001,1488,1512]" box="[864,1001,1488,1512]" pageId="33" pageNumber="183">Description</paragraph>
<paragraph id="C116101CFFA5CF0CDC35F996FB15A494" blockId="33.[826,1429,1539,1948]" lastBlockId="35.[826,1429,1393,1452]" lastPageId="35" lastPageNumber="185" pageId="33" pageNumber="183">
Male — (
<typeStatus id="1E12AEBEFFA5CF0EDC92F996FBDCA724" box="[967,1074,1539,1563]" pageId="33" pageNumber="183" type="holotype">holotype</typeStatus>
, in parentheses measurements of
<typeStatus id="1E12AEBEFFA5CF0EDC0CF9B3FC28A701" box="[857,966,1574,1598]" pageId="33" pageNumber="183" type="paratype">paratype</typeStatus>
). Dorsal and ventral shields anteriorly fused; dorsal shield L 397 (397), W 262 (259); dorsal shield (
<figureCitation id="59920C99FFA5CF0EDCEAF9F9FBDEA7BB" box="[959,1072,1644,1668]" captionStart="FIGURE" captionStartId="34.[159,171,1888,1907]" captionTargetBox="[264,1294,308,1826]" captionTargetId="figure-7@34.[252,1335,307,1857]" captionTargetPageId="34" captionText="FIGURE 24: Axonopsis balneatoris n. sp., male (B-C, E-F holotype, A, D paratype): (A) – dorsal shield, (B) – idiosoma, ventral view; (C-D) – palp; (E) – I-Leg-4-6; (F) – IV-Leg-2-6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404132" httpUri="https://zenodo.org/record/5404132/files/figure.png" pageId="33" pageNumber="183">Fig. 24A</figureCitation>
) bearing six pairs of glandularia (the sixth pair inconspicuous, flanking the excretory pore); postocularia well distanced from anterior margin; dorsal shield with a central ridge and a lateral ridge on each side lateral to the glandularia; eye pigment well developed; excretory pore located at posterior end of dorsal shield; ventral shield (
<figureCitation id="59920C99FFA5CF0EDCC4F8F4FC19A645" box="[913,1015,1889,1914]" captionStart="FIGURE" captionStartId="34.[159,171,1888,1907]" captionTargetBox="[264,1294,308,1826]" captionTargetId="figure-7@34.[252,1335,307,1857]" captionTargetPageId="34" captionText="FIGURE 24: Axonopsis balneatoris n. sp., male (B-C, E-F holotype, A, D paratype): (A) – dorsal shield, (B) – idiosoma, ventral view; (C-D) – palp; (E) – I-Leg-4-6; (F) – IV-Leg-2-6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404132" httpUri="https://zenodo.org/record/5404132/files/figure.png" pageId="33" pageNumber="183">Fig. 24B</figureCitation>
) L 441 (445), W 291 (284), L/W ratio 1.52 (1.57) with subparallel lateral margins; capitular bay L 96 (100); one pair of glandularia lying between the genital field and insertions of the fourth legs, three pairs of acetabula, arranged in an arc; width between most lateral pair of acetabula 128 (128); gonopore W 15.5 (16); ejaculatory complex L 87 (89); palp (
<figureCitation id="59920C99FFA7CF0CDEC2F9B5FE00A707" box="[407,494,1568,1592]" captionStart="FIGURE" captionStartId="34.[159,171,1888,1907]" captionTargetBox="[264,1294,308,1826]" captionTargetId="figure-7@34.[252,1335,307,1857]" captionTargetPageId="34" captionText="FIGURE 24: Axonopsis balneatoris n. sp., male (B-C, E-F holotype, A, D paratype): (A) – dorsal shield, (B) – idiosoma, ventral view; (C-D) – palp; (E) – I-Leg-4-6; (F) – IV-Leg-2-6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404132" httpUri="https://zenodo.org/record/5404132/files/figure.png" pageId="35" pageNumber="185">Fig. 24</figureCitation>
C-D): total L 178 (185), dL: P-1, 25 (25); P-2, 41 (44); P-3, 29 (31); P-4, 62 (63); P-5, 21 (22); %L: P-1, 14.0 (13.5); P-2, 23.0 (23.8); P-3, 16.3 (16.8); P-4, 34.8 (34.1); P-5, 11.8 (11.9); L P-2/P-4 ratio 0.66 (0.7); distal margin of P-3 with well developed hyaline extensions, middle of ventral side of P-4 expanded, bearing a relatively heavy seta which lies on a small pointed tubercle. Legs: L of I-Leg-3-6 (
<figureCitation id="59920C99FFA7CF0CDFF2F8ADFEE1A66E" box="[167,271,1848,1873]" captionStart="FIGURE" captionStartId="34.[159,171,1888,1907]" captionTargetBox="[264,1294,308,1826]" captionTargetId="figure-7@34.[252,1335,307,1857]" captionTargetPageId="34" captionText="FIGURE 24: Axonopsis balneatoris n. sp., male (B-C, E-F holotype, A, D paratype): (A) – dorsal shield, (B) – idiosoma, ventral view; (C-D) – palp; (E) – I-Leg-4-6; (F) – IV-Leg-2-6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404132" httpUri="https://zenodo.org/record/5404132/files/figure.png" pageId="35" pageNumber="185">Fig. 24E</figureCitation>
): 41 (45), 50 (48), 60 (62); I-Leg-6 L/H 2.0 (2.1); L of IV-Leg-2-6 (
<figureCitation id="59920C99FFA7CF0CDEF0F8C9FDE0A64B" box="[421,526,1884,1908]" captionStart="FIGURE" captionStartId="33.[311,323,1092,1111]" captionTargetBox="[217,1343,335,1029]" captionTargetId="figure-4@33.[189,1399,302,1060]" captionTargetPageId="33" captionText="FIGURE 23: Axonopsis arabica n. sp., male: (A) – idiosoma, ventral view; (B) – palp. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404130" httpUri="https://zenodo.org/record/5404130/files/figure.png" pageId="35" pageNumber="185">Fig. 23E</figureCitation>
): 75 (74), 65 (62), 69 (65), 80 (82), 99 (99); IV-Leg-6 (
<figureCitation id="59920C99FFA7CF0CDD50F8EAFD8BA6A8" box="[517,613,1919,1943]" captionStart="FIGURE" captionStartId="34.[159,171,1888,1907]" captionTargetBox="[264,1294,308,1826]" captionTargetId="figure-7@34.[252,1335,307,1857]" captionTargetPageId="34" captionText="FIGURE 24: Axonopsis balneatoris n. sp., male (B-C, E-F holotype, A, D paratype): (A) – dorsal shield, (B) – idiosoma, ventral view; (C-D) – palp; (E) – I-Leg-4-6; (F) – IV-Leg-2-6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404132" httpUri="https://zenodo.org/record/5404132/files/figure.png" pageId="35" pageNumber="185">Fig. 24F</figureCitation>
) strongly elevated distally, L/H 2.7 (2.8), with concave dorsal margin; legs without swimming setae.
</paragraph>
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<paragraph id="C116101CFFA6CF0DDFCAF8F5FD1AA6B0" blockId="34.[159,1428,1888,1935]" pageId="34" pageNumber="184">
FIGURE 24:
<taxonomicName id="06A96B9FFFA6CF0DDE45F8F5FE20A64C" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[272,462,1888,1907]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="34" pageNumber="184" phylum="Arthropoda" rank="species" species="balneatoris" status="sp. nov.">
<emphasis id="F3DDCC0EFFA6CF0DDE45F8F5FE20A64C" box="[272,462,1888,1907]" italics="true" pageId="34" pageNumber="184">Axonopsis balneatoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA6CF0DDE86F8F5FDE6A64C" box="[467,520,1888,1907]" pageId="34" pageNumber="184" rank="species">n. sp.</taxonomicNameLabel>
, male (B-C, E-F holotype, A, D paratype): (A) – dorsal shield, (B) – idiosoma, ventral view; (C-D) – palp; (E) – I-Leg-4-6; (F) – IV-Leg-2-6. Scale bars = 100 µm.
</paragraph>
</caption>
<caption id="95D64094FFA7CF0CDFAEFA88FAD7A40F" ID-DOI="http://doi.org/10.5281/zenodo.5404134" ID-Zenodo-Dep="5404134" box="[251,1337,1308,1328]" httpUri="https://zenodo.org/record/5404134/files/figure.png" pageId="35" pageNumber="185" startId="35.[251,263,1309,1328]" targetBox="[190,1406,326,1241]" targetPageId="35">
<paragraph id="C116101CFFA7CF0CDFAEFA88FAD7A40F" blockId="35.[251,1337,1308,1328]" box="[251,1337,1308,1328]" pageId="35" pageNumber="185">
FIGURE 25:
<taxonomicName id="06A96B9FFFA7CF0CDE39FA88FDC7A410" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[364,553,1308,1328]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="35" pageNumber="185" phylum="Arthropoda" rank="species" species="balneatoris" status="sp. nov.">
<emphasis id="F3DDCC0EFFA7CF0CDE39FA88FDC7A410" box="[364,553,1308,1328]" italics="true" pageId="35" pageNumber="185">Axonopsis balneatoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA7CF0CDD7BFA88FD8DA40F" box="[558,611,1309,1328]" pageId="35" pageNumber="185" rank="species">n. sp.</taxonomicNameLabel>
, female: (A) – idiosoma, ventral view; (B) – genital field. Scale bars = 100 µm.
</paragraph>
</caption>
<paragraph id="C116101CFFA7CF0CDC35FA5EFC94A67D" blockId="35.[826,1429,1483,1858]" pageId="35" pageNumber="185">
Female — similar to male except for the shape of the genital field (
<figureCitation id="59920C99FFA7CF0CDB52FA7BFB9FA739" box="[1031,1137,1518,1542]" captionStart="FIGURE" captionStartId="35.[251,263,1309,1328]" captionTargetBox="[190,1406,326,1241]" captionTargetId="figure-4@35.[166,1421,302,1276]" captionTargetPageId="35" captionText="FIGURE 25: Axonopsis balneatoris n. sp., female: (A) – idiosoma, ventral view; (B) – genital field. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5404134" httpUri="https://zenodo.org/record/5404134/files/figure.png" pageId="35" pageNumber="185">Fig. 25B</figureCitation>
); dorsal shield L 428, W 234; ventral shield L 469, W 305, capitular bay L 99; width between most lateral pair of acetabula 123; gonopore relatively small, W 34; palp: total L 191, dL and %L (in parentheses): P-1, 28 (14.7); P-2, 42 (22.0); P-3, 32 (16.8); P-4, 65 (34.0); P-5, 24 (12.6); L P-2/P-4 ratio 0.65; legs: L of I-Leg-4-6: 42, 49, 63; I-Leg-6 L/H 1.8; L of IV-Leg-3-6: 65, 69, 82, 108; IV-Leg-6 L/H 2.7; leg and palp chaetotaxy as in the male.
</paragraph>
<paragraph id="C116101CFFA7CF0BDC35F8F4FEBDA04D" blockId="35.[826,1429,1889,1948]" lastBlockId="36.[159,762,311,779]" lastPageId="36" lastPageNumber="186" pageId="35" pageNumber="185">
Etymology —
<emphasis id="F3DDCC0EFFA7CF0CDB59F8F4FB81A646" box="[1036,1135,1889,1913]" italics="true" pageId="35" pageNumber="185">balneator</emphasis>
(lat.) = ’Bader’. Named after Dr Carl Bader who made the first contribution to the knowledge of the water mite fauna of the Arabian peninsula.
</paragraph>
<paragraph id="C116101CFFA0CF0BDF90FE1CFDE1A3E3" blockId="36.[159,762,311,779]" pageId="36" pageNumber="186">
Remarks — The new species resembles
<emphasis id="F3DDCC0EFFA0CF0BDDC8FE1CFEEEA0FC" italics="true" pageId="36" pageNumber="186">
<taxonomicName id="06A96B9FFFA0CF0BDDC8FE1CFF55A0FC" authorityName="Piersig" authorityYear="1893" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="36" pageNumber="186" phylum="Arthropoda" rank="genus">Axonopsis</taxonomicName>
vietsi
</emphasis>
Mota¸s and Tanasachi, 1947, a hyporheobiontic species known from the Central Europe and the Mediterranean area (Pe˘si´c and
<bibRefCitation id="A5386DEDFFA0CF0BDD10FE67FD01A335" author="Gerecke R." box="[581,751,498,522]" pageId="36" pageNumber="186" pagination="563 - 576" refId="ref18121" refString="Pe ˘ si'c, V., Gerecke R. 2003 - Water mites of the genera Albaxona, Axonopsis, Barbaxonella and Erebaxonopsis (Acari, Hydrachnidia: Aturidae: Axonopsinae) from Central Europe and Mediterranean area - Arch. Hydrobiol., 139: 563 - 576." type="journal article" year="2003">Gerecke, 2003</bibRefCitation>
).
<taxonomicName id="06A96B9FFFA0CF0BDFCAFD80FE61A312" authority="Smit &amp; Pesic, 2010" authorityName="Smit &amp; Pesic" authorityYear="2010" box="[159,399,533,557]" class="Arachnida" family="Aturidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="36" pageNumber="186" phylum="Arthropoda" rank="species" species="balneatoris" status="sp. nov.">
<emphasis id="F3DDCC0EFFA0CF0BDFCAFD80FE61A312" box="[159,399,533,557]" italics="true" pageId="36" pageNumber="186">Axonopsis balneatoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8EE7175FFA0CF0BDECEFD80FE05A313" box="[411,491,533,556]" pageId="36" pageNumber="186" rank="species">n. sp.</taxonomicNameLabel>
, can be distinguished from
<emphasis id="F3DDCC0EFFA0CF0BDFB7FDADFEAFA36F" box="[226,321,568,592]" italics="true" pageId="36" pageNumber="186">A. vietsi</emphasis>
by the more slender idiosoma with subparallel lateral margins. A further difference is found in the shape of IV-Leg-6 which is less elevated distally and with a straight dorsal margin in
<emphasis id="F3DDCC0EFFA0CF0BDFCAFD51FF17A3E3" box="[159,249,708,732]" italics="true" pageId="36" pageNumber="186">A. vietsi</emphasis>
(Gerecke pers. comm.).
</paragraph>
<paragraph id="C116101CFFA0CF0BDF90FD66FE22A234" blockId="36.[159,762,311,779]" box="[197,460,755,779]" pageId="36" pageNumber="186">
Distribution —
<collectingCountry id="B9BE508CFFA0CF0BDE29FD66FE26A234" box="[380,456,755,779]" name="Oman" pageId="36" pageNumber="186">Oman</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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<emphasis id="F3DDCC0EFFADCF06DC1DFE08FA87A08A" bold="true" box="[840,1385,413,437]" italics="true" pageId="41" pageNumber="191">Arrenurus (Brevicaudaturus) rectituberculatus</emphasis>
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(
<figureCitation id="59920C99FFADCF06DCB2FE69FB8EA32B" box="[999,1120,508,532]" captionStart="FIGURE" captionStartId="43.[159,171,1804,1823]" captionTargetBox="[189,1396,396,1729]" captionTargetId="figure-4@43.[189,1399,391,1771]" captionTargetPageId="43" captionText="FIGURE 31: Arrenurus rectituberculatus n. sp., male: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – idiosoma, lateral view; (D-E) – palp. Scale bars = 200 µm (A-C), 50 µm (D-E)." figureDoi="http://doi.org/10.5281/zenodo.5404146" httpUri="https://zenodo.org/record/5404146/files/figure.png" pageId="41" pageNumber="191">Figures 31</figureCitation>
A-E, 32A-C)
</paragraph>
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Type material —
<materialsCitation id="71C11A41FFADCF06DB66FDD3FA7FA3BE" collectingDate="2008-11-04" country="Oman" latitude="17.053934" location="Wadi Hanna" longLatPrecision="1" longitude="54.60815" pageId="41" pageNumber="191" specimenCount="1" typeStatus="holotype">
<typeStatus id="1E12AEBEFFADCF06DB66FDD3FB4CA361" box="[1075,1186,582,606]" pageId="41" pageNumber="191" type="holotype">Holotype</typeStatus>
male,
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,
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,
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,
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.
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: 1/2/0, same data as holotype.
</paragraph>
<paragraph id="C116101CFFADCF06DC35FD2FFBA5A203" blockId="41.[826,1429,582,1949]" pageId="41" pageNumber="191">Diagnosis — Male with a large rectangular hump near posterior margin, between pygal lobes; dorsal furrow very short. Female with laterally widened genital plates.</paragraph>
<paragraph id="C116101CFFADCF06DC35FCC7FC07A255" blockId="41.[826,1429,582,1949]" box="[864,1001,850,874]" pageId="41" pageNumber="191">Description</paragraph>
<paragraph id="C116101CFFADCF06DC35FC15FA9AA4D3" blockId="41.[826,1429,582,1949]" pageId="41" pageNumber="191">
Male — Idiosoma yellowish, L 1183 (1183), W (including humps) 1110 (1134). Anterior idiosoma margin straight to slightly convex. Eyes on small humps, D1 on very large humps. Cauda short, pygal lobes distinct. Between pygal lobes, dorsally a rectangular hump, which extends beyond posterior idiosoma margin. Dorsal shield incomplete, dorsal furrow very short, lying between D1 (
<figureCitation id="59920C99FFADCF06DBA7FBE3FAB7A5B1" box="[1266,1369,1142,1166]" captionStart="FIGURE" captionStartId="43.[159,171,1804,1823]" captionTargetBox="[189,1396,396,1729]" captionTargetId="figure-4@43.[189,1399,391,1771]" captionTargetPageId="43" captionText="FIGURE 31: Arrenurus rectituberculatus n. sp., male: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – idiosoma, lateral view; (D-E) – palp. Scale bars = 200 µm (A-C), 50 µm (D-E)." figureDoi="http://doi.org/10.5281/zenodo.5404146" httpUri="https://zenodo.org/record/5404146/files/figure.png" pageId="41" pageNumber="191">Fig. 31A</figureCitation>
). Cx- I extending beyond anterior idiosioma margin. Suture lines between coxal plates indistinct. Gonopore L 90. Genital dield wing-shaped, indistinct, posterior margin with a row of long setae (
<figureCitation id="59920C99FFADCF06DBBEFA97FAA0A425" box="[1259,1358,1282,1306]" captionStart="FIGURE" captionStartId="43.[159,171,1804,1823]" captionTargetBox="[189,1396,396,1729]" captionTargetId="figure-4@43.[189,1399,391,1771]" captionTargetPageId="43" captionText="FIGURE 31: Arrenurus rectituberculatus n. sp., male: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – idiosoma, lateral view; (D-E) – palp. Scale bars = 200 µm (A-C), 50 µm (D-E)." figureDoi="http://doi.org/10.5281/zenodo.5404146" httpUri="https://zenodo.org/record/5404146/files/figure.png" pageId="41" pageNumber="191">Fig. 31B</figureCitation>
). Palp (
<figureCitation id="59920C99FFADCF06DC17FAB0FC7FA402" box="[834,913,1317,1341]" captionStart="FIGURE" captionStartId="43.[159,171,1804,1823]" captionTargetBox="[189,1396,396,1729]" captionTargetId="figure-4@43.[189,1399,391,1771]" captionTargetPageId="43" captionText="FIGURE 31: Arrenurus rectituberculatus n. sp., male: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – idiosoma, lateral view; (D-E) – palp. Scale bars = 200 µm (A-C), 50 µm (D-E)." figureDoi="http://doi.org/10.5281/zenodo.5404146" httpUri="https://zenodo.org/record/5404146/files/figure.png" pageId="41" pageNumber="191">Fig. 31</figureCitation>
D-E): total L 364; L: P-1, 40; P-2, 102; P-3, 66; P-4, 106; P-5, 50; P-2 anteroventrally with three setae, more medially another seta; P-3 anterodorsally with one seta. L of I-Leg-4-6: 178, 194, 267. L of IV- Leg-4-6: 259, 227, 203; IV-Leg-4 without spur. Third and fourth legs with numerous swimming setae.
</paragraph>
<paragraph id="C116101CFFADCF05DC35F996FE0DA04E" blockId="41.[826,1429,582,1949]" lastBlockId="42.[159,762,311,451]" lastPageId="42" lastPageNumber="192" pageId="41" pageNumber="191">
Female — Idiosoma L 1539, W 1458. Anterior margin slightly concave. D1, D4 and L4 on large humps, L3, V2 and V3 on smaller humps. Posterior idiosoma margin indented. Dorsal shield (
<figureCitation id="59920C99FFADCF06DA3DF9F9FC85A798" captionStart="FIGURE" captionStartId="44.[159,171,1776,1795]" captionTargetBox="[189,1372,433,1704]" captionTargetId="figure-7@44.[189,1399,419,1744]" captionTargetPageId="44" captionText="FIGURE 32: Arrenurus rectituberculatus n. sp., female: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – idiosoma, lateral view. Scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.5404148" httpUri="https://zenodo.org/record/5404148/files/figure.png" pageId="41" pageNumber="191">Fig. 32A</figureCitation>
) complete, L 786, W 867. Cx-I extending just beyond anterior idiosoma margin (
<figureCitation id="59920C99FFADCF06DBB3F927FAB9A7F5" box="[1254,1367,1714,1738]" captionStart="FIGURE" captionStartId="44.[159,171,1776,1795]" captionTargetBox="[189,1372,433,1704]" captionTargetId="figure-7@44.[189,1399,419,1744]" captionTargetPageId="44" captionText="FIGURE 32: Arrenurus rectituberculatus n. sp., female: (A) – idiosoma, dorsal view; (B) – idiosoma, ventral view; (C) – idiosoma, lateral view. Scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.5404148" httpUri="https://zenodo.org/record/5404148/files/figure.png" pageId="41" pageNumber="191">Fig. 32B</figureCitation>
). Suture lines of coxal plates indistinct. Medial corner of Cx-IV rounded. Genital plates indistinct (and therefore not illustrated in lateral view), long and bowed, widened laterally. Gonopore L 156. Palp: total L 403; L: P-1, 48; P-2, 114; P-3, 60; P-4, 129; P-5, 52; palp as in male. L of I-Leg-4-6: 211, 211, 259. L of IV-Leg-4-6: 284, 255, 211. Third and fourth legs with numerous swimming setae.
</paragraph>
<paragraph id="C116101CFFAECF05DF90FE1DFE1AA0FC" blockId="42.[159,762,311,451]" pageId="42" pageNumber="192">Etymology — Named after the rectangular posterodorsal hump of the male.</paragraph>
<caption id="95D64094FFAECF05DFCAFABEFE97A465" ID-DOI="http://doi.org/10.5281/zenodo.5404144" ID-Zenodo-Dep="5404144" httpUri="https://zenodo.org/record/5404144/files/figure.png" pageId="42" pageNumber="192" startId="42.[159,171,1323,1342]" targetBox="[182,732,499,1286]" targetPageId="42">
<paragraph id="C116101CFFAECF05DFCAFABEFE97A465" blockId="42.[159,763,1323,1370]" pageId="42" pageNumber="192">
FIGURE 30:
<taxonomicName id="06A96B9FFFAECF05DE5AFABEFE20A401" authority="Smit" authorityName="Smit" authorityYear="2000" box="[271,462,1323,1342]" class="Arachnida" family="Arrenuridae" genus="Arrenurus" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="192" phylum="Arthropoda" rank="species" species="ortali">
<emphasis id="F3DDCC0EFFAECF05DE5AFABEFE73A401" box="[271,413,1323,1342]" italics="true" pageId="42" pageNumber="192">Arrenurus ortali</emphasis>
Smit
</taxonomicName>
, female: idiosoma, ventral view. Scale bar = 200 µm.
</paragraph>
</caption>
<paragraph id="C116101CFFAECF05DF90FA0FFB14A04D" blockId="42.[159,762,1434,1948]" lastBlockId="42.[826,1429,311,370]" pageId="42" pageNumber="192">
Remarks — The very short dorsal furrow of the male, resulting in the almost absence of a dorsal shield, is quite unusual. Complete absence of a dorsal shield is also found within the genus
<emphasis id="F3DDCC0EFFAECF05DFCAF9B3FEBBA701" box="[159,341,1574,1598]" italics="true" pageId="42" pageNumber="192">Thorophoracarus</emphasis>
Viets.
<bibRefCitation id="A5386DEDFFAECF05DEE6F9B3FDADA701" author="Cook, D. R." box="[435,579,1574,1598]" pageId="42" pageNumber="192" pagination="1 - 860" refId="ref17864" refString="Cook, D. R. 1974 - Water mite genera and subgenera - Mem. Amer. Ent. Inst., 21: 1 - 860." type="journal article" year="1974">Cook (1974)</bibRefCitation>
postulated the hypotheses that
<emphasis id="F3DDCC0EFFAECF05DE30F9DCFDF5A75E" box="[357,539,1609,1633]" italics="true" pageId="42" pageNumber="192">Thorophoracarus</emphasis>
has arisen several times from divergent
<taxonomicName id="06A96B9FFFAECF05DEF4F9F9FDFAA7BC" authorityName="DugLs" authorityYear="1834" box="[417,532,1644,1667]" class="Arachnida" family="Arrenuridae" genus="Arrenurus" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="F3DDCC0EFFAECF05DEF4F9F9FDFAA7BC" box="[417,532,1644,1667]" italics="true" pageId="42" pageNumber="192">Arrenurus</emphasis>
</taxonomicName>
stock. The male of the new species, with hardly a dorsal furrow, could support this hypotheses. The female shares some characters with other
<emphasis id="F3DDCC0EFFAECF05DEFEF940FDB0A7D2" box="[427,606,1749,1773]" italics="true" pageId="42" pageNumber="192">Brevicaudaturus</emphasis>
species, like the long bowed genital plates and the shape of the posterior idiosoma margin. It is most close to
<taxonomicName id="06A96B9FFFAECF05DDB5F889FE9AA669" authority="Green, 1974" authorityName="Green" authorityYear="1974" class="Arachnida" family="Arrenuridae" genus="Arrenurus" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="192" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="F3DDCC0EFFAECF05DDB5F889FF09A669" italics="true" pageId="42" pageNumber="192">A. moorei</emphasis>
<bibRefCitation id="A5386DEDFFAECF05DFB9F8ABFE9AA669" author="Green, J. &amp; Corbet S. A. &amp; Betney E." box="[236,372,1854,1878]" pageId="42" pageNumber="192" pagination="199 - 223" refId="ref17976" refString="Green, J., Corbet S. A., Betney E. 1974 - Ecological studies on crater lakes in West Cameroon. Debundsha Lake - J. Zool. Lond., 173: 199 - 223." type="journal article" year="1974">Green, 1974</bibRefCitation>
</taxonomicName>
from
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, an insufficiently described species, only known from the female sex. The female of the new species differs from
<taxonomicName id="06A96B9FFFAECF05DDC7F810FD14A6A3" authorityName="Green" authorityYear="1974" box="[658,762,1925,1948]" class="Arachnida" family="Arrenuridae" genus="Arrenurus" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="192" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="F3DDCC0EFFAECF05DDC7F810FD14A6A3" box="[658,762,1925,1948]" italics="true" pageId="42" pageNumber="192">A. moorei</emphasis>
</taxonomicName>
in the laterally widened and shorter genital plates (not widened and longer in
<taxonomicName id="06A96B9FFFAECF05DBD7FECFFB04A04D" authorityName="Green" authorityYear="1974" box="[1154,1258,346,370]" class="Arachnida" family="Arrenuridae" genus="Arrenurus" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="192" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="F3DDCC0EFFAECF05DBD7FECFFB04A04D" box="[1154,1258,346,370]" italics="true" pageId="42" pageNumber="192">A. moorei</emphasis>
</taxonomicName>
).
</paragraph>
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147
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<taxonomicName id="06A96B9FFFADCF06DFB1FD26FD5AA3F5" authority="Smit, 2000" authorityName="Smit" authorityYear="2000" box="[228,692,690,715]" class="Arachnida" family="Arrenuridae" genus="Arrenurus" kingdom="Animalia" order="Trombidiformes" pageId="41" pageNumber="191" phylum="Arthropoda" rank="species" species="ortali" subGenus="Arrenurus">
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<emphasis id="F3DDCC0EFFADCF06DFB1FD26FDDCA3F5" bold="true" box="[228,562,690,715]" italics="true" pageId="41" pageNumber="191">Arrenurus (Arrenurus) ortali</emphasis>
<bibRefCitation id="A5386DEDFFADCF06DD6CFD27FD5AA3F5" author="Smit, H. &amp; Gerecke R. &amp; Sabatino A. di" box="[569,692,690,714]" pageId="41" pageNumber="191" pagination="201 - 267" refId="ref18425" refString="Smit, H., Gerecke R., Sabatino A. di 2000 - A catalogue of water mites of the superfamily Arrenuroidea (Acari: Hydrachnidia) from the Mediterranean countries - Arch. Hydrobiol. Suppl., 121: 201 - 267." type="journal article" year="2000">Smit, 2000</bibRefCitation>
</emphasis>
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(
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)
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<paragraph id="C116101CFFADCF06DF90FCA0FD1FA24F" blockId="41.[159,762,821,1948]" pageId="41" pageNumber="191">
Type material —
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<typeStatus id="1E12AEBEFFADCF06DEFEFCA0FDF4A272" box="[427,538,821,845]" pageId="41" pageNumber="191" type="holotype">Holotype</typeStatus>
male,
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,
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,
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,
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</date>
, leg.
<collectorName id="6C5C75CAFFADCF06DEAEFCCDFD81A24F" box="[507,623,856,880]" pageId="41" pageNumber="191">Margalith</collectorName>
(
<collectionCode id="A7B888D9FFADCF06DD28FCCDFD3AA24F" box="[637,724,856,880]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" lsid="urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="41" pageNumber="191">ZMAN</collectionCode>
!).
</materialsCitation>
</paragraph>
<paragraph id="C116101CFFADCF06DF90FC10FDDAA2FF" blockId="41.[159,762,821,1948]" pageId="41" pageNumber="191">
Other material —
<materialsCitation id="71C11A41FFADCF06DEF0FC10FDDEA2FF" collectingDate="2008-11-04" country="Oman" latitude="17.053934" location="Wadi Hanna" longLatPrecision="1" longitude="54.60815" pageId="41" pageNumber="191" specimenCount="1">
1/4/0,
<location id="C47646C7FFADCF06DD55FC10FD74A2A2" LSID="urn:lsid:plazi:treatment:4900A10AFFADCF06DFB1FD26FB37A04D:C47646C7FFADCF06DD55FC10FD74A2A2" box="[512,666,901,925]" country="Oman" latitude="17.053934" longLatPrecision="1" longitude="54.60815" name="Wadi Hanna" pageId="41" pageNumber="191">Wadi Hanna</location>
,
<collectingCountry id="B9BE508CFFADCF06DDFFFC10FD1BA2A2" box="[682,757,901,925]" name="Oman" pageId="41" pageNumber="191">Oman</collectingCountry>
,
<geoCoordinate id="A49D76DBFFADCF06DFCAFC3DFEC4A2FF" box="[159,298,936,960]" degrees="17" direction="north" minutes="03.236" orientation="latitude" pageId="41" pageNumber="191" precision="1" value="17.053934">17°03.236 N</geoCoordinate>
<geoCoordinate id="A49D76DBFFADCF06DE65FC3DFE59A2FF" box="[304,439,936,960]" degrees="54" direction="east" minutes="36.489" orientation="longitude" pageId="41" pageNumber="191" precision="1" value="54.60815">54°36.489 E</geoCoordinate>
,
<date id="B51736DCFFADCF06DE96FC3DFDDEA2FF" box="[451,560,936,960]" pageId="41" pageNumber="191" value="2008-11-04">
<collectingDate id="A553CF34FFADCF06DE96FC3DFDDEA2FF" box="[451,560,936,960]" pageId="41" pageNumber="191" value="2008-11-04">4-xi-2008</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="C116101CFFADCF06DF90FC43FEA0A2D1" blockId="41.[159,762,821,1948]" box="[197,334,982,1006]" pageId="41" pageNumber="191">Description</paragraph>
<paragraph id="C116101CFFADCF06DF90FB96FDB8A5F5" blockId="41.[159,762,821,1948]" pageId="41" pageNumber="191">
<materialsCitation id="71C11A41FFADCF06DF90FB96FDBCA5F5" country="Oman" location="Male - Described" municipality="The" pageId="41" pageNumber="191" specimenCount="1">
<location id="C47646C7FFADCF06DF90FB96FE49A524" LSID="urn:lsid:plazi:treatment:4900A10AFFADCF06DFB1FD26FB37A04D:C47646C7FFADCF06DF90FB96FE49A524" box="[197,423,1027,1051]" country="Oman" municipality="The" name="Male - Described" pageId="41" pageNumber="191">Male — Described</location>
in
<bibRefCitation id="A5386DEDFFADCF06DE83FB96FD5CA524" author="Smit, H. &amp; Gerecke R. &amp; Sabatino A. di" box="[470,690,1027,1051]" pageId="41" pageNumber="191" pagination="201 - 267" refId="ref18425" refString="Smit, H., Gerecke R., Sabatino A. di 2000 - A catalogue of water mites of the superfamily Arrenuroidea (Acari: Hydrachnidia) from the Mediterranean countries - Arch. Hydrobiol. Suppl., 121: 201 - 267." type="journal article" year="2000">
Smit
<emphasis id="F3DDCC0EFFADCF06DD4DFB91FDBFA524" box="[536,593,1027,1051]" italics="true" pageId="41" pageNumber="191">et al.</emphasis>
(2000)
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.
<collectingMunicipality id="21728A66FFADCF06DD98FB96FD17A524" box="[717,761,1027,1051]" pageId="41" pageNumber="191">The</collectingMunicipality>
male from
<collectingCountry id="B9BE508CFFADCF06DE73FBB3FE81A501" box="[294,367,1062,1086]" name="Oman" pageId="41" pageNumber="191">Oman</collectingCountry>
is slightly larger than the
<typeStatus id="1E12AEBEFFADCF06DDEFFBB2FD17A500" box="[698,761,1063,1087]" pageId="41" pageNumber="191">types</typeStatus>
(given in parentheses): Idiosoma 1089 (1038-1057) long and 664 (660-669) wide. Following the original description the petiole should have grooves, but in fact these are short, stiff curved setae
</materialsCitation>
.
</paragraph>
<paragraph id="C116101CFFADCF06DF90FB75FEB6A673" blockId="41.[159,762,821,1948]" pageId="41" pageNumber="191">
Female — Idiosoma greenish-brownish, L 1085 (1049-1175), W 923 (915-1021). Anterior margin of idiosoma convex. Dorsal shield complete, L 842 (810-899), W 632 (624-688). Cx-I extending beyond anterior margin of idiosoma (
<figureCitation id="59920C99FFADCF06DD41FAF9FD93A4BB" box="[532,637,1388,1412]" captionStart="FIGURE" captionStartId="42.[159,171,1323,1342]" captionTargetBox="[182,732,499,1286]" captionTargetId="figure-31@42.[165,755,487,1291]" captionTargetPageId="42" captionText="FIGURE 30: Arrenurus ortali Smit, female: idiosoma, ventral view. Scale bar = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.5404144" httpUri="https://zenodo.org/record/5404144/files/figure.png" pageId="41" pageNumber="191">Fig. 30</figureCitation>
). Medial margin of Cx-IV longer than medial margin of Cx- III. Medial distance of Cx-IV longer than width of one genital valve. Gonopore L 136, W 154, genital valves with very small sclerotized patches. Genital plates tapering laterally, directed perpendicularly towards lateral idiosoma margin. Palp: total L 384; L: P-1, 46; P-2, 102; P-3, 66; P-4, 106; P-5, 64; P-2 anteroventrally with three setae (as in male), more anterodorsally one more seta and dorsal margin with three setae. L of I-Leg-4-6: 156, 166, 144. L of IV-Leg- 4-5: 208, 192. Third and fourth legs with numerous swimming setae. Claws of legs with large clawlet and claw blade.
</paragraph>
<paragraph id="C116101CFFADCF06DF90F8F4FB37A04D" blockId="41.[159,762,821,1948]" lastBlockId="41.[826,1429,311,370]" pageId="41" pageNumber="191">
Remarks — The female was not known before (but see note at the preceding species). Previously known from the Sinai (
<collectingCountry id="B9BE508CFFADCF06DB10FEA2FB67A070" box="[1093,1161,311,335]" name="Israel" pageId="41" pageNumber="191">Israel</collectingCountry>
, nowadays
<collectingCountry id="B9BE508CFFADCF06DA47FEA2FAB7A070" box="[1298,1369,311,335]" name="Egypt" pageId="41" pageNumber="191">Egypt</collectingCountry>
) and the Negev (
<collectingCountry id="B9BE508CFFADCF06DC94FECFFBEDA04D" box="[961,1027,346,370]" name="Israel" pageId="41" pageNumber="191">Israel</collectingCountry>
) (
<bibRefCitation id="A5386DEDFFADCF06DB4FFECFFB23A04D" author="Smit, H. &amp; Gerecke R. &amp; Sabatino A. di" box="[1050,1229,346,370]" pageId="41" pageNumber="191" pagination="201 - 267" refId="ref18425" refString="Smit, H., Gerecke R., Sabatino A. di 2000 - A catalogue of water mites of the superfamily Arrenuroidea (Acari: Hydrachnidia) from the Mediterranean countries - Arch. Hydrobiol. Suppl., 121: 201 - 267." type="journal article" year="2000">
Smit
<emphasis id="F3DDCC0EFFADCF06DB02FECEFB62A04D" box="[1111,1164,346,370]" italics="true" pageId="41" pageNumber="191">et al.</emphasis>
2000
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

69
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<taxonomicName LSID="4901932F-6B5A-95E9-730B-D2A144D8B289" authority="Banks, 1904" authorityName="Banks" authorityYear="1904" class="Arachnida" family="Philodromidae" genus="Apollophanes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apollophanes texanus" order="Araneae" pageId="263" pageNumber="264" phylum="Arthropoda" rank="species" species="texanus">Apollophanes texanus Banks, 1904</taxonomicName>
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<taxonomicName authorityName="Banks" authorityYear="1904" class="Arachnida" family="Philodromidae" genus="Apollophanes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apollophanes texanus" order="Araneae" pageId="263" pageNumber="264" phylum="Arthropoda" rank="species" species="texanus">Apollophanes texanus</taxonomicName>
<bibRefCitation author="Broussard, GH" pageId="437" pageNumber="438" publicationUrl="10.3157/0013-872X(2006)117[249:CSAAIT]2.0.CO;2" refId="B99" refString="Broussard, GH, Horner, NV, 2006. Cursorial spiders (Arachnida: Araneae) in the Chihuahuan desert of western Texas, U.S.A. Entomological News 117: 249-260. doi: 10.3157/0013-872X(2006)117[249:CSAAIT]2.0.CO;2" title="Cursorial spiders (Arachnida: Araneae) in the Chihuahuan desert of western Texas, U. S. A. Entomological News 117: 249 - 260. doi:" url="10.3157/0013-872X(2006)117[249:CSAAIT]2.0.CO;2" year="2006">Broussard and Horner 2006</bibRefCitation>
: 255;
<bibRefCitation author="Dondale, CD" pageId="443" pageNumber="444" publicationUrl="10.4039/Ent1071175-11" refId="B222" refString="Dondale, CD, Redner, JH, 1975c. Revision of the spider genus Apollophanes (Araneida: Thomisidae). Canadian Entomologist 107: 1175-1192. doi: 10.4039/Ent1071175-11" title="Revision of the spider genus Apollophanes (Araneida: Thomisidae). Canadian Entomologist 107: 1175 - 1192. doi:" url="10.4039/Ent1071175-11" year="1975 c">Dondale and Redner 1975c</bibRefCitation>
: 1181 [spelling], mf, desc. (figs 3, 6, 26-28);
<bibRefCitation author="Jackman, JA" pageId="451" pageNumber="452" refId="B378" refString="Jackman, JA, 1997. A field guide to spiders and scorpions of Texas. Texas Monthly Field Guide Series, Gulf Publishing Company, Houston, Texas, 201 pp." title="A field guide to spiders and scorpions of Texas. Texas Monthly Field Guide Series, Gulf Publishing Company, Houston, Texas, 201 pp." year="1997">Jackman 1997</bibRefCitation>
: 166;
<bibRefCitation author="Kaston, BJ" pageId="452" pageNumber="453" refId="B396" refString="Kaston, BJ, 1953. How to know the spiders, first edition, Dubuque, Iowa, 220 pp." title="How to know the spiders, first edition, Dubuque, Iowa, 220 pp." year="1953">Kaston 1953</bibRefCitation>
: 103 (fig. 253);
<bibRefCitation author="Kaston, BJ" pageId="452" pageNumber="453" refId="B398" refString="Kaston, BJ, 1972. How to know the spiders, second edition, Dubuque, Iowa, 289 pp." title="How to know the spiders, second edition, Dubuque, Iowa, 289 pp." year="1972">Kaston 1972</bibRefCitation>
: 247 (fig. 562);
<bibRefCitation author="Kaston, BJ" pageId="452" pageNumber="453" refId="B400" refString="Kaston, BJ, 1978. How to know the spiders, third edition, William C. Brown Company, Publishers, Dubuque, Iowa, 272 pp." title="How to know the spiders, third edition, William C. Brown Company, Publishers, Dubuque, Iowa, 272 pp." year="1978">Kaston 1978</bibRefCitation>
: 239 (fig. 609);
<bibRefCitation author="Petrunkevitch, A" pageId="461" pageNumber="462" publicationUrl="10.5962/bhl.title.23819" refId="B567" refString="Petrunkevitch, A, 1911. A synonymic index-catalogue of spiders of North, Central and South America with all adjacent islands, Greenland, Bermuda, West Indies, Terra del Fuego, Galapagos, etc. Bulletin of the American Museum of Natural History 29: 1-791. doi: 10.5962/bhl.title.23819" title="A synonymic index-catalogue of spiders of North, Central and South America with all adjacent islands, Greenland, Bermuda, West Indies, Terra del Fuego, Galapagos, etc. Bulletin of the American Museum of Natural History 29: 1 - 791. doi:" url="10.5962/bhl.title.23819" year="1911">Petrunkevitch 1911</bibRefCitation>
: 402;
<bibRefCitation author="Richman, DB" pageId="466" pageNumber="467" publicationUrl="10.1894/JC-31.1" refId="B665" refString="Richman, DB, Brantley, SL, Hu, DH-C, Whitehouse, MEA, 2011a. Spiders of the Chihuahuan Desert of Southern New Mexico and Western Texas. Southwestern Naturalist 56: 44-53. doi: 10.1894/JC-31.1" title="Spiders of the Chihuahuan Desert of Southern New Mexico and Western Texas. Southwestern Naturalist 56: 44 - 53. doi:" url="10.1894/JC-31.1" year="2011 a">Richman et al. 2011a</bibRefCitation>
: 49;
<bibRefCitation author="Roewer, CF" pageId="467" pageNumber="468" refId="B680" refString="Roewer, CF, 1955. Katalog der Araneae von 1758 bis 1940, bzw. 1954. Bruxelles, Belgium, volume 2." title="Katalog der Araneae von 1758 bis 1940, bzw. 1954. Bruxelles, Belgium, volume 2." year="1955">Roewer 1955</bibRefCitation>
: 767;
<bibRefCitation author="Vogel, BR" pageId="471" pageNumber="472" refId="B759" refString="Vogel, BR, 1970b. Bibliography of Texas Spiders. Armadillo Papers 2: 1-36." title="Bibliography of Texas Spiders. Armadillo Papers 2: 1 - 36." year="1970 b">Vogel 1970b</bibRefCitation>
: 27
</paragraph>
<paragraph pageId="263" pageNumber="264">
<taxonomicName class="Arachnida" family="Philodromidae" genus="Apollophanes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apollophanes texana" order="Araneae" pageId="263" pageNumber="264" phylum="Arthropoda" rank="species" species="texana">Apollophanes texana</taxonomicName>
Banks, 1904;
<bibRefCitation author="Banks, N" pageId="433" pageNumber="434" refId="B23" refString="Banks, N, 1904. New genera and species of nearctic spiders. Journal of the New York Entomological Society 12: 109-119." title="New genera and species of nearctic spiders. Journal of the New York Entomological Society 12: 109 - 119." year="1904">Banks 1904</bibRefCitation>
: 113, mf, desc. (figs 12, 20);
<bibRefCitation author="Banks, N" pageId="433" pageNumber="434" publicationUrl="10.5479/si.03629236.72.i" refId="B26" refString="Banks, N, 1910. Catalogue of nearctic spiders. Bulletin of the United States National Museum 72: 1-80. doi: 10.5479/si.03629236.72.i" title="Catalogue of nearctic spiders. Bulletin of the United States National Museum 72: 1 - 80. doi:" url="10.5479/si.03629236.72.i" year="1910">Banks 1910</bibRefCitation>
: 51
</paragraph>
</subSubSection>
<subSubSection pageId="263" pageNumber="264" type="distribution">
<paragraph pageId="263" pageNumber="264">Distribution.</paragraph>
<paragraph pageId="263" pageNumber="264">Central and west Texas; Bexar, Brewster, Culberson, El Paso, Hudspeth, Presidio, Terrell</paragraph>
</subSubSection>
<subSubSection pageId="263" pageNumber="264" type="locality">
<paragraph pageId="263" pageNumber="264">Locality.</paragraph>
<paragraph pageId="263" pageNumber="264">Big Bend National Park, Big Bend Ranch State Park, Chihuahuan desert, Dalquest Research Site, Davis Mountains, Guadalupe Mountains</paragraph>
</subSubSection>
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<paragraph pageId="263" pageNumber="264">Time of activity.</paragraph>
<paragraph pageId="263" pageNumber="264">Female (March)</paragraph>
</subSubSection>
<subSubSection pageId="263" pageNumber="264" type="habitat">
<paragraph pageId="263" pageNumber="264">Habitat.</paragraph>
<paragraph pageId="263" pageNumber="264">(landscape features: under rock)</paragraph>
</subSubSection>
<subSubSection pageId="263" pageNumber="264" type="method">
<paragraph pageId="263" pageNumber="264">Method.</paragraph>
<paragraph pageId="263" pageNumber="264">pitfall trap [m]</paragraph>
</subSubSection>
<subSubSection pageId="263" pageNumber="264" type="type">
<paragraph pageId="263" pageNumber="264">Type.</paragraph>
<paragraph pageId="263" pageNumber="264">Texas (male, female, Bexar Co., San Antonio, syntype, no date, no collector, MCZ)</paragraph>
</subSubSection>
<subSubSection pageId="263" pageNumber="264" type="etymology">
<paragraph pageId="263" pageNumber="264">Etymology.</paragraph>
<paragraph pageId="263" pageNumber="264">locality (state)</paragraph>
</subSubSection>
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<paragraph pageId="263" pageNumber="264">Collection.</paragraph>
<paragraph pageId="263" pageNumber="264">MCZ, MSU, NMSU</paragraph>
</subSubSection>
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1954
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M a t e r i a l: 1 (
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), 1 Corfu, in BM coll. S.S. Saunders nach
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. Der Nachweis von &quot;
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&quot; in
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ist falsch, es handelt sich um &quot;Corfu&quot; (Corcyra = Kérkyra), siehe Originalbeschreibung in
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.
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B e m e r k u n g e n Frasspflanze unbekannt, bisher nur aus Kérkyra bekannt (
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M a t e r i a l: Exemplare Korfu, Tsavrou, 29.5., Gouvia, 23.5. und Korrision,
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, auf Wiesen nicht selten (
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B e m e r k u n g e n Frasspflanze der Larven unbekannt, die Art ist von Zentral- und SO-Europa und der
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M a t e r i a l: 1, 1 Korfu, siehe ENSLIN (1913) p. 117-118, 126-
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B e m e r k u n g e n Frasspflanze unbekannt, die Art ist in SO-Europa verbreitet (
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;
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;
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;
<bibRefCitation id="A5392AACFF98FFA619808F80FC25C5B6" author="TAEGER A &amp; BLANK S" box="[699,918,1180,1202]" pageId="6" pageNumber="841" pagination="399 - 504" refId="ref4962" refString="TAEGER A., BLANK S. M. &amp; A. D. LISTON (2006): European Sawflies (Hymenoptera: Symphyta) - A Species Checklist for the Countries. - In: Recent Sawfly Research: Synthesis and Prospects, Keltern p. 399 - 504." type="book chapter" year="2006">TAEGER et al. 2006</bibRefCitation>
); Farbfotos der Imagines in
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. Die Art ist also nicht auf Kérkyra beschränkt, z. B. in
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und in
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beim Mt. Ossa.
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M a t e r i a l: 1 Korfu, Gouvia,
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(
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B e m e r k u n g e n Frasspflanze unbekannt. Eine mediterrane Art, die sonst nur aus
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und den Pyrenäen bekannt ist (
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M a t e r i a l:
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B e m e r k u n g e n Frasspflanze der Larven sind Fiederblätter von
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(LORENZ- KRAUS 1957), andere Autoren nennen auch
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sp. Die Art ist neu für Korfu, ein nearktischer Einwanderer aus den
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M a t e r i a l: 1 Korfu, Pantokrator,
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, in
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B e m e r k u n g e n Larven bohren in Stängeln von
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, die Art kommt in Europa, im Norden bis
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, im Kaukasus und in
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vor (
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M a t e r i a l: 1 Korfu, Tsavrou,
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B e m e r k u n g e n:DieLarvenfressenanBlätternvon
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B e m e r k u n g e n:DieLarvenlebenanBrassicaceae wie
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spp.,
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<emphasis id="F3DC8B4FFF9AFFA418D58EEEFEBFC722" italics="true" pageId="4" pageNumber="839">Alliaria petiolata</emphasis>
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und
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sp., die Art ist weit verbreitet von Europa, der
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, Transkaukasien, dem
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, Sibirien bis Zentralasien (
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M a t e r i a l: 1
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, Korfu, Mesaria,
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,
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, leg. H. Malicky, in coll. et det. W. Sch. 2011.
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B e m e r k u n g e n Die Larven leben an den Blättern von
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(
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), vielleicht auch an mediterranen
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-Arten? Die Art ist von Zentral- und S-Europa sowie von der
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M a t e r i a l: Korfu, Tsavrou, 29.5., Gouvia, 23.5. und Temploni
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, auf Gebüsch nach
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.
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B e m e r k u n g e n:DieLarvensindanBlätternvon
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bekannt wie
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,
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spp. und
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spp. (
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).
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<emphasis id="F3DC8B4FFF9CFFA21B898863FE2AC290" bold="true" box="[178,409,894,916]" italics="true" pageId="2" pageNumber="837">Aprosthema fusicorne</emphasis>
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M a t e r i a l: 1 Korfu, Tsavrou, 29.5. u.
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; 1 Pantokrator,
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B e m e r k u n g e n:Larven an
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nach
<bibRefCitation id="A5392AACFF9CFFA219E188CCFC19C2E2" author="KONTUNIEMI T" box="[730,938,976,998]" pageId="2" pageNumber="837" pagination="1 - 104" refId="ref4572" refString="KONTUNIEMI T. (1960): Die Futterpflanzen der Sagewespenlarven (Hymenoptera, Symphyta) Finnlands. - Animalia fennica, Helsingfors 9: 1 - 104." type="journal article" year="1960">KONTUNIEMI (1960)</bibRefCitation>
, die Art ist in Europa weit verbreitet, auch von
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bekannt (
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).
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<paragraph id="C117575DFF9DFFA31B89889AFCC0C29F" blockId="3.[178,883,903,923]" box="[178,883,903,923]" pageId="3" pageNumber="838">M a t e r i a l:1 1 Corfu, leg. Erber, 1 leg. Pareys, nach STROBL (1895).</paragraph>
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B e m e r k u n g e n Die Larven fressen an Blättern von
<taxonomicName id="06A82CDEFF9DFFA3181E88B6FC26C2C4" box="[805,917,939,960]" class="Magnoliopsida" family="Rosaceae" genus="Prunus" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="838" phylum="Tracheophyta" rank="species" species="undetermined">
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(
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), der Verfasser hat sie an
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vorgefunden (SCHEDL &amp; KRAUS 1988). Die Art ist von
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, der
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, von
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, dem
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, von
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, dem Transkaukasus, dem
<collectingCountry id="B9BF17CDFF9DFFA31BDF8F1FFEBEC513" box="[228,269,1026,1047]" name="Iran" pageId="3" pageNumber="838">Iran</collectingCountry>
bis
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verbreitet (SCHEDL und KRAUS 1988).
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M a t e r i a l: 1, 1 Korfu, Pantokrator,
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B e m e r k u n g e n:DieLarvenfressenebenfallsan
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B e m e r k u n g e n Larven z.B. an
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(J.P.E.F. STEIN 1876)
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</paragraph>
<paragraph id="C117575DFF9DFFA31B898ECBFCDDC4EE" blockId="3.[178,878,1494,1514]" box="[178,878,1494,1514]" pageId="3" pageNumber="838">M a t e r i a l :: 1 Corfu, leg. Erber, in NHMW, nach STROBL (1895 p. 144).</paragraph>
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B e m e r k u n g e n Die Frasspflanze(n) der Larven ist unbekannt. Die Art ist am Balkan, der
<collectingCountry id="B9BF17CDFF9DFFA31A0F8D08FEC8C729" box="[308,379,1557,1581]" name="Turkey" pageId="3" pageNumber="838">Türkei</collectingCountry>
und in
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verbreitet (
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).
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<emphasis id="F3DC8B4FFF9DFFA31B898FB4FE81C5BB" bold="true" box="[178,306,1193,1215]" italics="true" pageId="3" pageNumber="838">Abia sericea</emphasis>
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<paragraph id="C117575DFF9DFFA31B898FC4FCCFC5E9" blockId="3.[178,892,1241,1261]" box="[178,892,1241,1261]" pageId="3" pageNumber="838">M a t e r i a l:1 Corfu, leg. Simony, in NHMW, nach STROBL (1895 p. 143).</paragraph>
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B e m e r k u n g e n Die Larven ernähren sich, soweit bekannt, von Blättern von
<emphasis id="F3DC8B4FFF9DFFA31B898E07FED0C42B" box="[178,355,1306,1327]" italics="true" pageId="3" pageNumber="838">Knautia arvensis</emphasis>
bzw.
<emphasis id="F3DC8B4FFF9DFFA31A918E07FDC0C42B" box="[426,627,1306,1327]" italics="true" pageId="3" pageNumber="838">Succisa pratensis (</emphasis>
TAEGER 1998), die Art ist in ganz Europa verbreitet (
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144
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K. Andersen 1910
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<emphasis id="2D5C61319B151C419A2ACD714EDB8AFC" inLineHeading="true" pageNumber="330">Distribution:</emphasis>
Arfak Mtns. (New
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). Specimens from Mansuar Isl. (Prov. of
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,
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) may also represent
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<emphasis id="E6FA84848D397C4AFA9C32D6C0773976" italics="true" pageNumber="330">cyclotis</emphasis>
(
<bibRefCitation id="FA60A08EFB0DD7AE5D23DB266C6D2AB8" author="Meinig, H." refId="ref236882" refString="Meinig, H. 2002. New records of bats (Chiroptera) from Indonesian Islands. Myotis, 40: 59 - 79." year="2002">Meinig, 2002</bibRefCitation>
)
</taxonomicName>
. Specimens from New Britain formerly assigned to this species apparently represent
<taxonomicName id="CAA19AD9659E14E3127AF924AB88D613" authority="(Bonaccorso, 1998)" baseAuthorityName="Bonaccorso" baseAuthorityYear="1998" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="species" species="vizcaccia">
<emphasis id="66A874DF1F23ECE87CDEC24129A66252" italics="true" pageNumber="330">vizcaccia</emphasis>
(
<bibRefCitation id="69E34AFC6453C86DBF2FFFBB65A7B465" author="Bonaccorso, F. J." refId="ref35461" refString="Bonaccorso, F. J. 1998. Bats of Papua New Guinea. Conservation International Tropical Field Guide Series, Conservation International, Washington, D. C., 489 pp." year="1998">Bonaccorso, 1998</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B0DDEDE285535C0E00DD048A0B9BF701" pageNumber="330" type="conservation">
<paragraph id="ADFC365A77FCD2F90B0ABFA2E6FAE2A0" pageNumber="330">
<emphasis id="D78F4D972E675028805C7326397DCCC8" inLineHeading="true" pageNumber="330">Conservation:</emphasis>
<collectionCode id="34492EB318743F3DD5F2E307642B9857">IUCN</collectionCode>
/
<collectionCode id="CA017719AC2FABE2D1C372417A23CBEE" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (1992) – No Data.
<collectionCode id="E42DDB0BF51BCFDAD8023F8BFA1F4A9C">IUCN</collectionCode>
2003 – Lower Risk (nt).
</paragraph>
</subSubSection>
<subSubSection id="BF816C70E452BDBBBF7BA3AE17FFF320" pageNumber="330" type="discussion">
<paragraph id="E8AF53DD0DBE293F717FE35BF1B65445" pageNumber="330">
<emphasis id="40657654902C470988F50D7D1F995FE9" inLineHeading="true" pageNumber="330">Discussion:</emphasis>
<taxonomicName id="27975708ED976BF1CB3A691F47321CE3" baseAuthorityName="Meinig" baseAuthorityYear="2002" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="species" species="cyclotis">
<emphasis id="288BC44F926498BC4A235A4B8138413B" italics="true" pageNumber="330">cyclotis</emphasis>
</taxonomicName>
species group. Apparently does not include
<taxonomicName id="9CBA73C523DA4C482B95CA36AC8B74FB" authorityName="K. Andersen" authorityYear="1912" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="species" species="certans">
<emphasis id="6AA3C7AFB620438EE3B1C7008E88A1FA" italics="true" pageNumber="330">certans</emphasis>
</taxonomicName>
; see
<bibRefCitation id="9828BB2C602D672127013009540058B1" author="Peterson, R. L." refId="ref282761" refString="Peterson, R. L. 1991. Systematic variation in the megachiropteran tube-nosed bats Nyctimene cyclotis and N. certans. Bulletin of the American Museum of Natural History, 206: 26 - 41." year="1991">Peterson (1991)</bibRefCitation>
and Flannery (1995
<emphasis id="9BE5C4D4D943C06B1339CDE3F45E75C6" italics="true" pageNumber="330">a</emphasis>
,
<emphasis id="7307CBA7CE7F15108D5D910807ABF692" italics="true" pageNumber="330">b)</emphasis>
, though also see
<bibRefCitation id="09D565B205D120F2999F39BF4722BF39" author="Bonaccorso, F. J." refId="ref35461" refString="Bonaccorso, F. J. 1998. Bats of Papua New Guinea. Conservation International Tropical Field Guide Series, Conservation International, Washington, D. C., 489 pp." year="1998">Bonaccorso (1998)</bibRefCitation>
and comments under
<taxonomicName id="77EC67E81B9116DCB8A7818680A484D9" authorityName="K. Andersen" authorityYear="1912" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="species" species="certans">
<emphasis id="5085FDF9D78C32B2656B69D519A47C41" italics="true" pageNumber="330">certans</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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80
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Pleijel, 1993
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Reported from Greece by
<bibRefCitation id="65A4AC03387ECC0BDEB5E989EE16F488" author="Chatzigeorgiou, G" journalOrPublisher="Hellenic Centre for Marine Research" pageId="0" pageNumber="20997" title="Polychaetes from two subtidal rocky shores of the North coast of Crete, collected for the NaGISA project 2007 - 2008." year="2016">Chatzigeorgiou et al. (2016)</bibRefCitation>
. Type locality: Mediterranean (Banyuls-sur-Mer, France).
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Art ISFS: 332900 Checklist: 1036970
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aggr.
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<paragraph id="E17D68A23F2ABA7B930F397643649228">
<heading id="261497FC4541D56E3A8D48FBAFBADE9E">Zusammenfassung</heading>
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Artbeschreibung
<emphasis id="6D65AA949DA67FD80A672B0694F8F5B1" italics="true">
(nach
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)
</emphasis>
:
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hoch.
<emphasis id="1A824FBFE675DE7CA2219D553F6526FA" bold="true">
Spreite der
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mit
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Grund
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,
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lang. Blattstiel meist
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als Spreite.
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meist deutlich weisslich gefleckt
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. Auf den
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neben den lockerstehenden, bis
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langen Borsten keine 0,1-0,5 mm langen Haare, nur weniger als 0,1 mm hohe
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.
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lang.
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<paragraph id="BBED2024F6C6C6C299C670A1D43810A9">
<normalizedToken id="CA84C2B2DBAE24014983146DBD0E2DE0" originalValue="Blütezeit">Bluetezeit</normalizedToken>
<emphasis id="5BDA3F24CEC9336426A1D72905C8579E" italics="true">
(nach
<bibRefCitation id="D2636D85C167079457020FCD4B9964C6" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
: 3-5
</paragraph>
<paragraph id="2DA8E583A39EE85BF31430666AD66B3B">
Standort und Verbreitung in der Schweiz
<emphasis id="8842E1E98529DFCF1BFE9389D5B71B1B" italics="true">
(nach
<bibRefCitation id="322F66FA0B2AF2F3BEF4155F795BC015" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
:
<normalizedToken id="A170F737C6556B87A54FC8E3AB31A899" originalValue="Wälder">Waelder</normalizedToken>
,
<normalizedToken id="F8305E261C57CF130A43DA22F43B0D29" originalValue="Gebüsche">Gebuesche</normalizedToken>
/ kollin-subalpin / M, ANW (VD, VS),
<normalizedToken id="CCE99D2A4678DD6A703F368912C7F410" originalValue="südliches">suedliches</normalizedToken>
TI u.a.
</paragraph>
</subSubSection>
<subSubSection id="3642D2875ECF3725D969E49C3698F834" type="distribution">
<paragraph id="D9A4070A94831ADCD7BE0CA34E9C4BD4">
Verbreitung global
<emphasis id="E0BF2B0F66F1FE1F22B394233A9CCA6E" italics="true">
(nach
<bibRefCitation id="476F2FD136B7A5708176D62AD69A6F09" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
:
<normalizedToken id="E8D0162D92F5C4B8E22772D220B20EB0" originalValue="Mitteleuropäisch">Mitteleuropaeisch</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="0F6400606847200E67A3687C2DC4C8D7" type="biology_ecology">
<paragraph id="1190D05C9E489AB710B3A9A13C3539F3">
<normalizedToken id="B0F1A0B1BFA8A0B95CCF2A3FF7831E81" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte
<emphasis id="8835723E0019DD70FBABF045EA95BF3F" italics="true">
(nach
<bibRefCitation id="D077BD1052A03FF726570A7C31E13160" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. 2010</bibRefCitation>
)
</emphasis>
3 + 43-24 + 4.h.2n=16
</paragraph>
</subSubSection>
<subSubSection id="2C5F2F334A1BE45BB99A71D0B9ED2FF3" type="multiple">
<paragraph id="F474C8532CFB96E08ACBC104B9B915C4">Status</paragraph>
</subSubSection>
<subSubSection id="B569AD74FC7F4CD8B96A5F1CA4114979" type="conservation">
<paragraph id="2E4B841C2F011CD0F11A311F7DE13FB2">
<heading id="21DEE0B98418F612A8DE27F26308115A">Status IUCN</heading>
: Potenziell
<normalizedToken id="BB99581DA89795BF6D9C076F063D8691" originalValue="gefährdet">gefaehrdet</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="1DF7105C57BFFE062F6393F8240560BA" type="multiple">
<paragraph id="ED24415F461A023659E7EC2733D3A0B1">
<heading id="4E32F20269E350B03A3134661DCB802D">
<normalizedToken id="E059628C23E9C1F4D47DE2805EC6D7BE" originalValue="Ökologie">Oekologie</normalizedToken>
</heading>
</paragraph>
<paragraph id="AAFB01CC634F9A5BD6C0E77ECF88B504">
Lebensform
<normalizedToken id="D85F7E411DA4DA2AC54FAE6B3750F70C" originalValue="Mehrjähriger">Mehrjaehriger</normalizedToken>
Hemikryptophyt
</paragraph>
<paragraph id="5A6A81045C24AE14D89966C0ED28E0E5">
<heading id="43C1E461AE269063440C59970EAAB4D8">Lebensraum Lebensraum</heading>
nach
<bibRefCitation id="B518C456080AB145C76D18F13E1494E1" author="Delarze, R. &amp; Gonseth, Y. &amp; Eggenberger, S. &amp; Vust, M." location="Bad Hersfeld, Germany" publisher="Ott Verlag" refString="Delarze, R., Gonseth, Y., Eggenberger, S., Vust, M. 2015. Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition). Ott Verlag, Bad Hersfeld, Germany, 456 pp." title="Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition)" year="2015">Delarze &amp; al. 2015</bibRefCitation>
</paragraph>
<paragraph id="BBA89B6AF527CB117D5330A86CE71758">
<table id="62FEBA878A8E9D6ED28EA0D5F36B38D6" class="table" inLine="true">
<tbody id="89AACED5B79DE8B773A5D1C318CA96C5">
<tr id="BD75D4B668BCE905C6B1E1BE3AAE2846">
<td id="8131CB716FC8FEBC393F2553F686FD36">
6.2.3 - Waldmeister-Buchenwald (
<emphasis id="E342B91A4CE9687D430E72D011DEE58C" italics="true">Galio-Fagenion</emphasis>
)
</td>
</tr>
<tr id="397212CBC1FBB0E37FBB5A17627A4569">
<td id="2BB7694891AFB86BAAC9BA13A211C474">
6.3.3 - Eichen- Hainbuchenwald (
<emphasis id="F4F44E91EB1112649A863D6FECE0F972" italics="true">Carpinion</emphasis>
)
</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="E5E7C500C920C1D48FBB2845A578AD35">
<tableNote id="C3B82D01FD9941A8513388F79B6CD243">
<emphasis id="90F69277B52E299C93834E6E9DD9D8D8" bold="true">fett</emphasis>
<emphasis id="ACE66F5902BBAFAF1E97FE5915E09927" italics="true">
Dominante Art, welche das Aussehen des Lebensraumes
<normalizedToken id="FA7075296645BC1BFBF5C63EE083A7FA" originalValue="mitprägt">mitpraegt</normalizedToken>
</emphasis>
<emphasis id="ED17B34553C54413B9BF6A6E62998C13" italics="true">Charakterart</emphasis>
<emphasis id="EFBE7C31751625D377D7262CBCADF4F3" italics="true">Weniger strikt an den Lebensraum gebundene Art</emphasis>
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="9632C05FC17ACEB305BAE89E180AC5E3" type="biology_ecology">
<paragraph id="6FAE311F8C9E8A4FD0F27A55414C8267">
<normalizedToken id="1DDB62A35E0A7800C534FF82282D6B7A" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="89065058B95ACA22887FFB2567666735" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="A7771AF3FEC420F2AEE96FCEAAE99EC4">
<table id="533A93F732B189089ADAFABFA5C798CB" class="table" inLine="true">
<tbody id="16C22E99CA7BE39DBFE8CE7E4C6E789D">
<tr id="AE283A3493934535E7974231333CF4E2">
<th id="37FEDECE699C2567E8AC4348447B8A9D" colspan="2">
<emphasis id="B5F03049B74A10B9FADF57607C396B9A" bold="true">Bodenfaktoren</emphasis>
</th>
<th id="8AAFD148ACFFEBEB720A044DFBA9A9C6" colspan="2">
<emphasis id="17DE0AE6AF84BE16352F64C490F12AF7" bold="true">Klimafaktoren</emphasis>
</th>
<th id="7BF5F2C6C05F657AB6F16814C3970318" colspan="2">
<emphasis id="B566351C888990F08E59BBDF90E9ECCE" bold="true">Salztoleranz</emphasis>
</th>
</tr>
<tr id="1D6A5CA83D98D6410FFEA6C6471BBB08">
<td id="4239FED8D0AAC3EA70EC2B7C58F8C807">Feuchtezahl F</td>
<td id="A5A90681B5895B100DE7A3AF975998D9" style="text-align:left;width:100px">feucht</td>
<td id="CA6973DA17E7749C6ADFA2B5F8C1264D">Lichtzahl L</td>
<td id="47564CB883B132CD5FB45801FAD97EDE" style="text-align:left;width:60px">schattig</td>
<td id="4DA12432DCC1C6B8B09DD9B5B49FBF1F">Salzzeichen</td>
<td id="2BC3952501827F8201D5B7DFE38AB09C" style="text-align:left;width:60px">--</td>
</tr>
<tr id="DC7547449B474F4BAF60F94398FFB945">
<td id="9E2B9AB26E63B01693042AA8B0BD27D0">Reaktionszahl R</td>
<td id="E4C6E249F35119272BDA7A15E43B53A9" style="text-align:left">neutral bis basisch (pH 5.5-8.5)</td>
<td id="B47C2909216803A5EBFCEF862210795C">Temperaturzahl T</td>
<td id="9A32DB601AF7D36FAA3FB4DFCBDAB694" style="text-align:left">warm-kollin</td>
</tr>
<tr id="02B5513BD9C1D77F57080747A04F0B14">
<td id="A7A222131235FB441C97DC286252CE4E">
<normalizedToken id="25DC606D42D8B0CA04657C71C14E9B72" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="96E856704CC3119F146C58E9A2315024" style="text-align:left">
<normalizedToken id="33872194012632C8987522A2ABAE5C2D" originalValue="mässig">maessig</normalizedToken>
<normalizedToken id="9B6E9798C7292E41F4C5C2E8E3C44AD3" originalValue="nährstoffarm">naehrstoffarm</normalizedToken>
bis
<normalizedToken id="5AF33324DCB972C2F0C7262F3F9A20B6" originalValue="mässig">maessig</normalizedToken>
<normalizedToken id="ACFF62DEBBFA80D8E17EEDDE422EE353" originalValue="nährstoffreich">naehrstoffreich</normalizedToken>
</td>
<td id="44B979D1DE775A927CF60506297A6193">
<normalizedToken id="1F9FAC636E51913768E9C457B4F41F05" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="FE10F94C75050ADBC858C9DD439B15A5" style="text-align:left">subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter)</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="25799A35BE784FC124CEE966E2F9DC50" type="reference_group">
<paragraph id="7CA39C06F6746DDC1185A298EBC1A24F">
<heading id="E7DF8AF87DC546EACFB55A0BF3F55A91">Nomenklatur</heading>
</paragraph>
<paragraph id="0800B2FB13936BE90CAD2AFD31BC637F">
<treatmentCitation id="466595A6F49BB2962D232948790F3A13">
<heading id="2465510CB6B17EEA8B8E876CE55AF281">
<normalizedToken id="1594367A11FF728B2DC704A903E48CA1" originalValue="Gültiger">Gueltiger</normalizedToken>
Name (
<bibRefCitation id="80FFBF66D4D273417893799D8646437C" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</heading>
:
<taxonomicName id="7B06CB405E19E48956A3A2C427B0DFEC" ID-CoL="4QGQF" authority="L." authorityName="L." class="Magnoliopsida" family="Boraginaceae" genus="Pulmonaria" kingdom="Plantae" order="Boraginales" phylum="Tracheophyta" rank="species" species="officinalis">
<emphasis id="644310F7E1E23F4217909546BE92636C" italics="true">Pulmonaria officinalis</emphasis>
L.
</taxonomicName>
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA6343FD22BC4414EBE7142889663DE1" type="vernacular_names">
<paragraph id="648CFCDB64671EEA0C62040AC38615A0">
Volksname Deutscher Name:
<vernacularName id="B71540014FE729738B9054FCB4139632" language="deu">
<normalizedToken id="3A49BE6277E312C47DDCCF75FE46AA9B" originalValue="Gewöhnliches">Gewoehnliches</normalizedToken>
Lungenkraut
</vernacularName>
Nom
<normalizedToken id="FB14A595F85A3669690FA73BC12CA567" originalValue="français">francais</normalizedToken>
:
<vernacularName id="B47D14D1FF3A2A5BCBBA05053B40665D" language="fra">Pulmonaire officinale</vernacularName>
Nome italiano: --
</paragraph>
</subSubSection>
<subSubSection id="A97395466FE151E16F74205971870706" type="reference_group">
<paragraph id="9097EFF15EBF870AA55FCCE17B862387">
<normalizedToken id="576C191D0AAF1B61DB8C1D8A8C54E778" originalValue="Übereinstimmung">Uebereinstimmung</normalizedToken>
mit anderen Referenzwerken
</paragraph>
<paragraph id="8419C75950F56F04B396B135E8E974E1">
<table id="B16A5A69F2498A9B99757740A2A23683" class="table table-condensed" inLine="true">
<tbody id="87A3FDF0FCFCE52A044F2B48FAF5FF09">
<tr id="B22025CE88ACBEEA03A21D6F41A27EE3">
<th id="D31A05E1882C5721B39811EC34C9750F" width="80px">Relation</th>
<th id="A3EE6B1285CB4148F49ACB09AD89A120">Nom</th>
<th id="05BB67E9D52EA3FC6B8C60C84F1A2722">Referenzwerke</th>
<th id="C33A9B1825533A77613CE1DE018CC84E" width="90px">No</th>
</tr>
<tr id="2A1E72AC6480879B90E5D9BD3C870D9C">
<td id="0753B3949A28D522F9E4040C3265F175" class="taxon-relation">=</td>
<td id="B73E3C66C7A25FA5FA4EE606BC584612">
<taxonomicName id="2C0DA2351DA491A86C888243BAFCD317" authority="L." authorityName="L." class="Magnoliopsida" family="Boraginaceae" genus="Pulmonaria" kingdom="Plantae" order="Boraginales" phylum="Tracheophyta" rank="species" species="officinalis">Pulmonaria officinalis L.</taxonomicName>
</td>
<td id="4E39CEE9D454767B646E8BF975CECC87">
<treatmentCitation id="A8A7F34A2DA0F64D5D87EBE28B2BD9BD">
<bibRefCitation id="60E4272E49023FF66122F7E8163FF099" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
</treatmentCitation>
</td>
<td id="36FAD6AEC2C3D5A179F776085F5F2C5D">332900</td>
</tr>
<tr id="6EADE796DC36EE79C634F7ED6DF00C58">
<td id="65E37DEA69A0D84FC2A9B4E7A23DD4FD" class="taxon-relation">=</td>
<td id="33C25123DDDAE62C4660D8CB4F73BC55">
<taxonomicName id="D423097B30C4AA7BB3616C09B3CD8FDC" authority="L." authorityName="L." class="Magnoliopsida" family="Boraginaceae" genus="Pulmonaria" kingdom="Plantae" order="Boraginales" phylum="Tracheophyta" rank="species" species="officinalis">Pulmonaria officinalis L.</taxonomicName>
</td>
<td id="8B743291043329EC118299AE1C525411">
<treatmentCitation id="75B93C46E8957E2F6FAB37011913F9A5">
<bibRefCitation id="94C1BD685C3BBE41E10EC3E0104566C5">Flora Helvetica 2001</bibRefCitation>
</treatmentCitation>
</td>
<td id="296A5814D7D6D99BAB6797D84640FA29">1607</td>
</tr>
<tr id="8B8E531865247920EF1235217162687C">
<td id="A0BAB1BA3E0317ECBB4AC9F0C8ED5075" class="taxon-relation">=</td>
<td id="89CAA4034AFD54856898F0A491EBBD30">
<taxonomicName id="AEE63F2A31E7E36493A4138A940C490A" authority="L." authorityName="L." class="Magnoliopsida" family="Boraginaceae" genus="Pulmonaria" kingdom="Plantae" order="Boraginales" phylum="Tracheophyta" rank="species" species="officinalis">Pulmonaria officinalis L.</taxonomicName>
</td>
<td id="6B9068E79E468E04015D82F689A3B3FE">
<treatmentCitation id="C160655F6321B41CBBB0B5DE93049043">
<bibRefCitation id="92360DB3114A6C9D01E4B25C951869EA">Flora Helvetica 2012</bibRefCitation>
</treatmentCitation>
</td>
<td id="C21C02B9558874E3783B7F487B038BDA">1481</td>
</tr>
<tr id="3907A33A57B8110512D5935737DA9D45">
<td id="186AE9702E25D6BD443879D4DF198B99" class="taxon-relation">=</td>
<td id="433A2B7426F65A42C3177C0FFE30A77B">
<taxonomicName id="F65CF4DBAD40D285B67E025BD1313260" authority="L." authorityName="L." class="Magnoliopsida" family="Boraginaceae" genus="Pulmonaria" kingdom="Plantae" order="Boraginales" phylum="Tracheophyta" rank="species" species="officinalis">Pulmonaria officinalis L.</taxonomicName>
</td>
<td id="28CA2A33F37C5A669A186A9FC21A8D5D">
<treatmentCitation id="3DDDB0FC8455051472ECBBC3AED61C52" httpUri="http://treatment.plazi.org/id/5644E4A6FBB417662A3075C4A279D443">
<bibRefCitation id="4A16030291BD0C7C36104386E38C0E68">Flora Helvetica 2018</bibRefCitation>
</treatmentCitation>
</td>
<td id="BB81DC298AE2872C5BEB8BDB089FC14D">1481</td>
</tr>
<tr id="AD6BCDC4A3B7EAAB6AF0BB4334C33C06">
<td id="11C05207D32C95BAF3DB7320776B3A66" class="taxon-relation">=</td>
<td id="3C3CD79C97354B6A3681E0666D6B980C">
<taxonomicName id="93FF4EF10D32C8B1EDDA676A7C6F707B" authority="L." authorityName="L." class="Magnoliopsida" family="Boraginaceae" genus="Pulmonaria" kingdom="Plantae" order="Boraginales" phylum="Tracheophyta" rank="species" species="officinalis">Pulmonaria officinalis L.</taxonomicName>
</td>
<td id="C722BB715B3AA0DA760914879E0142D7">
<treatmentCitation id="F3D852B5A794581C5AF80DD60EBD9A76">
<bibRefCitation id="50D9D0B5C1E407252595BA639D9D47AF">Index synonymique 1996</bibRefCitation>
</treatmentCitation>
</td>
<td id="97232231DBEC623FC20C76D3D250AD82">332900</td>
</tr>
<tr id="222C37DDC7345F234BD965AB88AD5C5E">
<td id="69AE079C05DE4D1870E8B0412526C00A" class="taxon-relation">=</td>
<td id="299590E37B627CF59C588ED691AF6360">
<taxonomicName id="152830268C13A5573EA25CC307417778" authority="L." authorityName="L." class="Magnoliopsida" family="Boraginaceae" genus="Pulmonaria" kingdom="Plantae" order="Boraginales" phylum="Tracheophyta" rank="species" species="officinalis">Pulmonaria officinalis L.</taxonomicName>
</td>
<td id="271507F061898336416ED64234D23AD9">
<treatmentCitation id="4C71ACF25B2D66A716EC74E1AB5BFC7F">
<bibRefCitation id="6943DE0DDAD5344B4E8428ED94BAF5B1">SISF/ISFS 2</bibRefCitation>
</treatmentCitation>
</td>
<td id="E8B21117F3210BE833D1E48C2EB5E0FD">332900</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="5FF1179D8304064738757768409094DF">
<tableNote id="1F51389E1746DE4BE39FEEC92AEF003E">
= Taxon stimmt mit akzeptiertem Taxon
<normalizedToken id="A760B144672687FE376E3CD682C54E1C" originalValue="überein">ueberein</normalizedToken>
(
<bibRefCitation id="AF233570A9D86D058B618F252F32DF0D" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) &lt;Taxon ist im akzeptierten Taxon (
<bibRefCitation id="FA6D618D6F9E305783C9AB991474149E" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) enthalten&gt; Taxon
<normalizedToken id="BF5948106B701846CB169514D3828740" originalValue="enthält">enthaelt</normalizedToken>
(neben anderen) auch das akzeptierte Taxon (
<bibRefCitation id="6DF97F2700AA104A9B9D8ECCCE8DDD17" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="8E8A61F536DEF09A80F3891E71063115" type="conservation">
<paragraph id="64E90A0177A45BB3CE5E6F63B3DCA4DB">
<heading id="6C500098F919A67EAC5E46041D505C9F">Status Indigenat</heading>
: Indigen
</paragraph>
<paragraph id="711FD36293DA26D05197B41B2C59FD7B">
<emphasis id="4E323375B9EB8FA3C70004021F827928" bold="true">
<heading id="4D7A3B4F339768D2424DF16C7B2C3AE1">
Liste der
<normalizedToken id="FB961AAEB1287EA3F218EE62EEE9F60E" originalValue="gefährdeten">gefaehrdeten</normalizedToken>
Pflanzen IUCN
</heading>
(nach
<bibRefCitation id="8248AA658CBAA39D912F12711FC75751" author="Walter, K.S. &amp; Gillett, H.J." location="Gland, Switzerland and Cambridge, UK" publicationUrl="https://doi.org/10.5962/bhl.title.44833" publisher="IUCN" refString="Walter, K.S. and Gillett, H.J. (editors). 1998. 1997 IUCN Red List of Threatened Plants. IUCN, Gland, Switzerland and Cambridge, UK. DOI: https://doi.org/10.5962/bhl.title.44833" title="IUCN Red List of Threatened Plants" year="1997">Walter &amp; Gillett 1997</bibRefCitation>
):
</emphasis>
Nein
</paragraph>
<paragraph id="9AA1BEAD76AB2D62A7973FA2F0BE8ADA">
<heading id="8D30AA9BCE822BA2D925290D1DB0451E">Status Rote Liste national 2016</heading>
</paragraph>
<paragraph id="83AB4BAB194F765802C3CF156094BD5F">
<heading id="A4C720A25DC256DD30BAB59BE3CDD399">Status IUCN</heading>
: Potenziell
<normalizedToken id="103F63242DA3BD89A9A490CD77BF6DBE" originalValue="gefährdet">gefaehrdet</normalizedToken>
</paragraph>
<paragraph id="D640D7C3CB4BABF862D1654DDD22F267">
<heading id="72B50C65294EBA17344920C978ADA94C">
<normalizedToken id="2A2F0949798962970BB68064F008B43B" originalValue="Zusätzliche">Zusaetzliche</normalizedToken>
Informationen
</heading>
Kriterien IUCN: A2c
</paragraph>
<paragraph id="E35E0D0CA026117C2EE1B2658336455C">
<heading id="BD5EC867265BC5C25785811F70D1B1F3">Status Rote Liste regional 2019</heading>
</paragraph>
<paragraph id="B044B29837FB756C13AAFD0B56D802F3">
<table id="FAB57BA5CE1249725ED88C714CE2AF17" class="table table-condensed" inLine="true">
<tbody id="FC7955F4681D3553D9AFFF3067F13C53">
<tr id="633CCCD69CB0644C5EDBC57B7347A0C2">
<th id="3C0822125988747CF55DB091ECE75D7B">Biogeografische Regionen</th>
<th id="9127B889E2567E7049A4B4CFBFB277E7">Status</th>
<th id="B968E67A7CD475DCE7EFFCADC843A87B">Kriterien IUCN</th>
</tr>
<tr id="6301A6043FE486806BEC3C6CECF64FFE">
<td id="B190FEECAACCAE80BAA59E9374C606B5" class="regional_status">Jura (JU)</td>
<td id="7BE4B0C480D1FDB4250FD18A53EEE31A">
potenziell
<normalizedToken id="D39B98FCE4C28F9F02DCB6E2586DE804" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Near Threatened)
</td>
<td id="85C58C79D7368C29EE6CF5217BE26459">A2c</td>
</tr>
<tr id="30FFE0B7098F394E4BEC351C236770BC">
<td id="E2D8AC936F096E422F10243BC15321C0" class="regional_status">Mittelland (MP)</td>
<td id="EB59DD4B145286DD03E76D312008779D">
potenziell
<normalizedToken id="BB0798458D944C3798BEB90298498E76" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Near Threatened)
</td>
<td id="2CCB06B1B9D525D3481F725A98A7149B">A2c</td>
</tr>
<tr id="92779416B2184CC30B40759C2F0B9665">
<td id="37F2DD5D79B1A8932E8E102AB872C194" class="regional_status">Alpennordflanke (NA)</td>
<td id="D62D09F16CA873FB3B16B1C0764F5C43">
potenziell
<normalizedToken id="9C700CF57823C59D69057AF08C3C4DEA" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Near Threatened)
</td>
<td id="0BB658BDE0389F4598726419D3DEC2F6">A2c</td>
</tr>
<tr id="DD02E300B593347E5F381059083425F7">
<td id="90F8F47124DEAF71338E806604A86363" class="regional_status">
<normalizedToken id="143322AD78D1D9581C9BC14C379CAF74" originalValue="Alpensüdflanke">Alpensuedflanke</normalizedToken>
(SA)
</td>
<td id="1904FFBBFF4CB1FE9B4D6363F355CFA1">
potenziell
<normalizedToken id="64A27E890102301AAF10FE1DFB6DBBB5" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Near Threatened)
</td>
<td id="5304C817A2731A7BF33F4570883F448A">A2c</td>
</tr>
<tr id="41D54147D44B2851D36F3DF3FCEC50D7">
<td id="6843CD7D706A5D0FBDE6ED03B7E92AB9" class="regional_status">
<normalizedToken id="DEA9EE884023E2A60CDC7A7AB93E4AC5" originalValue="Östliche">Oestliche</normalizedToken>
Zentralalpen (EA)
</td>
<td id="3E3D3E84EF797B26185F19D754E407E1">
<normalizedToken id="4E4FE4A3E06200F983F638D8CAE49529" originalValue="ungenügende">ungenuegende</normalizedToken>
Datengrundlage (Data Deficient)
</td>
</tr>
<tr id="A9C9121568645343645961E76A2AC196">
<td id="24E3487942037A264AD9F8E5D632FF42" class="regional_status">Westliche Zentralalpen (WA)</td>
<td id="005A2799E2E4C4EF9D5A792522F2526A">nicht beurteilt (Not Evaluated)</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="A48EC447F6942267E2EC23AABDBCEA95">
<heading id="B44F16B07554F9D4D6651CFA84ED527A">
Status nationale
<normalizedToken id="C449E27AAD6259632194D21D03AA558F" originalValue="Priorität">Prioritaet</normalizedToken>
/Verantwortung
</heading>
</paragraph>
<paragraph id="E959976C1C9564FD9CE91C8981FFF45D">
<table id="9D290D9127538ED8D0A805297D160300" class="table" inLine="true">
<tr id="2DF4AC099125492D3E0736B4266F5771">
<td id="0B427DF61A7A8EEFA7C627F8BC45BF7D">
Keine nationale
<normalizedToken id="C6F7F537CE64D76C2B2EC6E7E87F5091" originalValue="Priorität">Prioritaet</normalizedToken>
oder internationale Verantwortung
</td>
</tr>
</table>
</paragraph>
<paragraph id="8F209EE85A54C7327B79F579D6E905AB">
<heading id="3A34D9D8411B36A9FB0E79A95E860A30">Schutzstatus</heading>
</paragraph>
<paragraph id="BCD3B35ED9CAF527BF2C1A386D498267">
<table id="052D6B4DD2270589C3D181EBE0731CD6" class="table" inLine="true">
<tr id="C591C3F15BD5EF8AB6FF2B6F176739D6">
<td id="ACE12F4442632757CBBC0821D9C900FF">Kein internationaler, nationaler oder kantonaler Schutz</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

135
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<treatment id="490322FC03B327EC13AAA0134E9FCCAA" ID-DOI="http://doi.org/10.5281/zenodo.4373177" ID-GBIF-Taxon="164015576" ID-Zenodo-Dep="4373177" LSID="urn:lsid:plazi:treatment:490322FC03B327EC13AAA0134E9FCCAA" httpUri="http://treatment.plazi.org/id/490322FC03B327EC13AAA0134E9FCCAA" lastPageNumber="827" pageNumber="827">
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<paragraph id="056410C3E96B3283781A4E9AF6EF7B95" pageNumber="827">
<treatmentCitation id="1CB69FAB8DB5BE12654FC35D93693CBE" author="Linnaeus" pageNumber="63" year="1762">
<bibCitation id="57F3351DEE2529F4421C12E3FDB70A88" author="Linnaeus" pagination="63" title="Schoenus spathaceus" volumeTitle="Species Plantarum, ed. 2, 1" year="1762">
<taxonomicName id="1D9CA91C3CB539BC347B4DA00476DEE5" ID-CoL="6F69K" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1762" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="spathaceus">
<emphasis id="A3DD900E3A0A1AFD4D8608829B72FE26" bold="true" pageNumber="827">Schoenus spathaceus</emphasis>
Linnaeus
</taxonomicName>
,
<volumeTitle id="E45F85F64525377B3BA231EC5A9F0101">
<emphasis id="8E17DA65959CAA16406190C77EB15BB5" italics="true" pageNumber="827">Species Plantarum</emphasis>
, ed. 2, 1
</volumeTitle>
: 63. 1762
</bibCitation>
</treatmentCitation>
,
<emphasis id="5E757D595D570BA86FC5FB4F03711293" italics="true" pageNumber="827">nom. illeg.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="2189395042B5A0FEB759D4314D361382" pageNumber="827" type="distribution">
<paragraph id="6ED960B8EE70F938F6945F2C99C35539" pageNumber="827">&quot;Habitat in Virginia, Cap. b. spei.&quot; RCN: 400.</paragraph>
</subSubSection>
<subSubSection id="3A3AC7CD0FF9BD368D9AEDE9F91CCC6C" pageNumber="827" type="reference_group">
<paragraph id="77ED583788065C5F5B160AFC7CA8D15C" pageNumber="827">
Replaced synonym:
<taxonomicName id="70A1D7716CEDFF37984145E83A0F929E" authority="L. (1753)" authorityName="Linnaeus" authorityYear="1753" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="arundinaceus">
<emphasis id="01F2BB0C6EE5D8E7DE974ABBA8556123" italics="true" pageNumber="827">Cyperus arundinaceus</emphasis>
L. (1753)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C94793034DA48B5F4346CE6AAAF17638" pageNumber="827" type="reference_group">
<paragraph id="41FB6FDEFF4216C0FF07E95BA4FCE8EA" pageNumber="827">
Replaced synonym of:
<taxonomicName id="5DE08679B83021D8F369EE45DCA2242E" authority="L. (1767)" authorityName="Linnaeus" authorityYear="1767" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="spathaceus">
<emphasis id="2553FB5A55780F30625AE2EC0FB2E5C2" italics="true" pageNumber="827">Cyperus spathaceus</emphasis>
L. (1767)
</taxonomicName>
,
<emphasis id="1966DF26CAD4072EED89838F281D0D0D" italics="true" pageNumber="827">nom. illeg.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="89F18AB8422ECFFC7FF3F6481C57BC06" pageNumber="827" type="materials_examined">
<paragraph id="02494EFD11B1BB597E948B479B5ED55D" pageNumber="827">
<materialsCitation id="01FB70C6E21CD34E58D3F437BEB2F0F6" ID-GBIF-Occurrence="2612020359" collectionCode="LINN BM" specimenCode="70.39" typeStatus="lectotype">
<typeStatus id="EADC75929B62015792EE16D50B8DD9F7">Lectotype</typeStatus>
(Kukkonen in
<emphasis id="A49D3D355E0095EDDF061387521C9C0E" italics="true" pageNumber="827">Ann. Bot. Fenn.</emphasis>
27: 65. 1990): Herb. Linn. No. 70.39 (
<collectionCode id="7BB0C0DCB168882C2122F1AB5D468347">LINN</collectionCode>
;
<typeStatus id="7CAA0D67A93F8C92981A993C30AD05D5">iso-</typeStatus>
<collectionCode id="77434940B8639B12C9B8D6DEDF8636D5">BM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EF08D589183CE3019CA6E0A8D284D8C8" pageNumber="827" type="nomenclature">
<paragraph id="DB14D917C7DF9B537D94EF595BE88F61" pageNumber="827">
Current name:
<taxonomicName id="036B094799E86C5EEDD39A7DECAC615D" authority="(L.) Britton" authorityName="Britton" authorityYear="1894" baseAuthorityName="Linnaeus" baseAuthorityYear="1762" class="Magnoliopsida" family="Cyperaceae" genus="Dulichium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="arundinaceum">
<emphasis id="88A4828EA7C2522D35BA6610FDE7FF04" italics="true" pageNumber="827">
<emphasis id="1187FF9883C02D14E61A5CF19995851E" bold="true" italics="true" pageNumber="827">Dulichium arundinaceum</emphasis>
</emphasis>
(L.) Britton
</taxonomicName>
(
<taxonomicName id="47B04D871C5C997543F1D38A473BAD59" authorityName="de Jussieu" authorityYear="1789" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="D70744A832BD8BA6AEFDD70A56C54048" pageNumber="827" type="discussion">
<paragraph id="974A3BE27F16967F8BD1F0593D8EFD99" pageNumber="827">
<emphasis id="DC58DF44CDB92D4FF7CF8738C6741024" italics="true" pageNumber="827">Note:</emphasis>
Linnaeus erroneously cites &quot;
<taxonomicName id="E9857CDC2C6C69E8A505F82E98C2D903" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="ferrugineus">Cyperus ferrugineus</taxonomicName>
.
<emphasis id="E30A053ADA66D563EAAD132E5DD92FA0" italics="true" pageNumber="827">Sp. pl.</emphasis>
44.&quot; in synonymy in the protologue. However, there is no
<taxonomicName id="81D0BACA58310AE97E7DA040FDC24B5A" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="ferrugineus">
<emphasis id="EC5286466DDE5B4F9501EE3009856065" italics="true" pageNumber="827">C. ferrugineus</emphasis>
</taxonomicName>
on p. 44 of
<emphasis id="0785E9670902D389E41910DD99883448" italics="true" pageNumber="827">
<volumeTitle id="7B1E8A6158FE1726955342BDD8E84E40">Species Plantarum</volumeTitle>
</emphasis>
, or indeed anywhere else in that work. The similarities of the phrase names, inclusion of the same three pre-1753 synonyms, and the same habitat statement show that
<taxonomicName id="A3D42608CFC96C9A54280BCA2DE120C5" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="arundinaceus">
<emphasis id="C8A2F4671E1F870D3D5A254B0FBB6909" italics="true" pageNumber="827">C. arundinaceus</emphasis>
</taxonomicName>
is the name Linnaeus was intending.
</paragraph>
</subSubSection>
</treatment>
</document>

114
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<mods:title>Updating the knowledge of sand flies (Diptera, Psychodidae) in Rondonia State, Brazil</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pereira Junior, Antonio Marques</mods:namePart>
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<mods:affiliation>Fundacao Oswaldo Cruz, Fiocruz Rondonia, Porto Velho, Brazil &amp; Instituto Nacional de Ciencia e Tecnologia de Epidemiologia da Amazonia Ocidental, Porto Velho, Brazil</mods:affiliation>
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</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rodrigues, Moreno Magalhaes de Souza</mods:namePart>
<mods:affiliation>Fundacao Oswaldo Cruz, Fiocruz Rondonia, Porto Velho, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Medeiros, Jansen Fernandes</mods:namePart>
<mods:affiliation>Fundacao Oswaldo Cruz, Fiocruz Rondonia, Porto Velho, Brazil &amp; Instituto Nacional de Ciencia e Tecnologia de Epidemiologia da Amazonia Ocidental, Porto Velho, Brazil</mods:affiliation>
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<taxonomicName LSID="49033A29-A793-5BDB-92B4-5E170F0D1F0B" authority="(Franca, 1920)" baseAuthorityName="Franca" baseAuthorityYear="1920" class="Insecta" family="Psychodidae" genus="Migonemyia" kingdom="Animalia" lsidName="Migonemyia migonei" order="Diptera" pageId="0" pageNumber="90015" phylum="Arthropoda" rank="species" species="migonei">
Migonemyia migonei (
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, 1920)
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<paragraph pageId="0" pageNumber="90015">Distribution</paragraph>
<paragraph pageId="0" pageNumber="90015">
Buritis,
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, Cacoal, Campo Novo,
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,
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do Oeste,
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, Monte Negro, Nova
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, Porto Velho
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<paragraph pageId="0" pageNumber="90015">Notes</paragraph>
<paragraph pageId="0" pageNumber="90015">
<bibRefCitation author="Galardo, Allan Kardec Ribeiro" journalOrPublisher="Revista da Sociedade Brasileira de Medicina Tropical" pageId="0" pageNumber="90015" pagination="265 - 271" refId="B7910410" refString="Galardo, Allan Kardec Ribeiro, Galardo, Clicia Denis, Silveira, Guilherme Abbad, Ribeiro, Kaio Augusto Nabas, Hijjar, Andrea Valadao, Oliveira, Liliane Leite, Dos Santos, Thiago Vasconcelos, 2015. Phlebotominae sand flies (Diptera: Psychodidae): potential vectors of American cutaneous leishmaniasis agents in the area associated with the Santo Antonio Hydroelectric System in Western Amazonian Brazil. Revista da Sociedade Brasileira de Medicina Tropical 48 (3): 265 - 271" title="Phlebotominae sand flies (Diptera: Psychodidae): potential vectors of American cutaneous leishmaniasis agents in the area associated with the Santo Antonio Hydroelectric System in Western Amazonian Brazil." volume="48" year="2015">Galardo et al. 2015</bibRefCitation>
,
<bibRefCitation author="Gil, Luis H S" journalOrPublisher="Memorias do Instituto Oswaldo Cruz" pageId="0" pageNumber="90015" pagination="751 - 5" refId="B7910455" refString="Gil, Luis H S, Basano, Sergio A, Souza, Adelson A, Silva, Maria G S, Barata, Iorlando, Ishikawa, Edna A, Camargo, Luis M A, Shaw, Jeffrey J, 2003. Recent observations on the sand fly (Diptera: Psychodidae) fauna of the State of Rondonia, Western Amazonia, Brazil: the importance of Psychdopygus davisi as a vector of zoonotic cutaneous leishmaniasis. Memorias do Instituto Oswaldo Cruz 98 (6): 751 - 5" title="Recent observations on the sand fly (Diptera: Psychodidae) fauna of the State of Rondonia, Western Amazonia, Brazil: the importance of Psychdopygus davisi as a vector of zoonotic cutaneous leishmaniasis." volume="98" year="2003">Gil et al. 2003</bibRefCitation>
,
<bibRefCitation author="Leao, Paula de Oliveira" journalOrPublisher="Parasites &amp; Vectors" pageId="0" pageNumber="90015" pagination=": 422" refId="B7910525" refString="Leao, Paula de Oliveira, Pereira Junior, Antonio Marques, de Paulo, Paula Frassinetti Medeiros, Carvalho, Luis Paulo Costa, Souza, Ana Beatriz Nascimento, da Silva, Michelli Santos, Castro, Thais Santos, Freitas, Moises Thiago de Souza, Rodrigues, Moreno Magalhaes de Souza, Ferreira, Gabriel Eduardo Melim, Medeiros, Jansen Fernandes, 2020. Vertical stratification of sand fly diversity in relation to natural infections of Leishmania sp. and blood-meal sources in Jamari National Forest, Rondonia State, Brazil. Parasites &amp; Vectors 13 (1): 422" title="Vertical stratification of sand fly diversity in relation to natural infections of Leishmania sp. and blood-meal sources in Jamari National Forest, Rondonia State, Brazil." volume="13" year="2020">
<normalizedToken originalValue="Leão">Leao</normalizedToken>
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,
<bibRefCitation author="Pereira Junior, Antonio Marques" journalOrPublisher="Zootaxa" pageId="0" pageNumber="90015" pagination="270 - 278" refId="B7910585" refString="Pereira Junior, Antonio Marques, Marialva, Eric Fabricio, Juliao, Genimar Reboucas, Pessoa, Felipe Arley Costa, Medeiros, Jansen Fernandes, 2019. Survey of sand flies (Diptera: Psychodidae: Phlebotominae) in Guajara-Mirim State Park forest reserve, near the Brazil-Bolivian border, with a description of Pintomyia fiocruzi, a new sand fly species. Zootaxa 4691 (3): 270 - 278" title="Survey of sand flies (Diptera: Psychodidae: Phlebotominae) in Guajara-Mirim State Park forest reserve, near the Brazil-Bolivian border, with a description of Pintomyia fiocruzi, a new sand fly species" volume="4691" year="2019">
Pereira
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et al. 2019a
</bibRefCitation>
,
<bibRefCitation author="Pereira Junior, Antonio Marques" journalOrPublisher="Memorias do Instituto Oswaldo Cruz" pageId="0" pageNumber="90015" pagination=": e 190170" refId="B7910573" refString="Pereira Junior, Antonio Marques, Souza, Ana Beatriz Nascimento, Castro, Thais Santos, da Silva, Michelli Santos, de Paulo, Paula Frassinetti Medeiros, Ferreira, Gabriel Eduardo Melim, de Medeiros, Jansen Fernandes, 2019. Diversity, natural infection and blood meal sources of phlebotomine sandflies (Diptera, Psychodidae) in the western Brazilian Amazon. Memorias do Instituto Oswaldo Cruz 114: e190170" title="Diversity, natural infection and blood meal sources of phlebotomine sandflies (Diptera, Psychodidae) in the western Brazilian Amazon." volume="114" year="2019">
Pereira
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et al. 2019b
</bibRefCitation>
,
<bibRefCitation author="Resadore, Fabio" journalOrPublisher="Vector Borne and Zoonotic Diseases" pageId="0" pageNumber="90015" pagination="347 - 357" refId="B7910606" refString="Resadore, Fabio, Junior, Antonio Marques Pereira, de Paulo, Paula Frassinetti Medeiros, Gil, Luiz Herman Soares, Rodrigues, Moreno Magalhaes de Souza, Araujo, Maisa da Silva, Juliao, Genimar Reboucas, Medeiros, Jansen Fernandes, 2018. Composition and vertical stratification of phlebotomine sand fly fauna and the molecular detection of Leishmania in forested areas in Rondonia State municipalities, Western Amazon, Brazil. Vector Borne and Zoonotic Diseases 19 (5): 347 - 357" title="Composition and vertical stratification of phlebotomine sand fly fauna and the molecular detection of Leishmania in forested areas in Rondonia State municipalities, Western Amazon, Brazil" volume="19" year="2018">Resadore et al. 2018</bibRefCitation>
,
<bibRefCitation author="Silva, Adriele Nunes Rodrigues" journalOrPublisher="Acta Tropica" pageId="0" pageNumber="90015" pagination=": 105757" refId="B7910637" refString="Silva, Adriele Nunes Rodrigues, Junior, Antonio Marques Pereira, de Paulo, Paula Frassinetti Medeiros, da Silva, Michelli Santos, Castro, Thais Santos, Costa, Glaucilene da Silva, Freitas, Moises Thiago de Souza, Rodrigues, Moreno Magalhaes de Souza, Medeiros, Jansen Fernandes, 2021. Detection of Leishmania species (Kinetoplastida, Trypanosomatidae) in phlebotomine sand flies (Diptera, Psychodidae) from Porto Velho, Northern Brazil. Acta Tropica 213: 105757" title="Detection of Leishmania species (Kinetoplastida, Trypanosomatidae) in phlebotomine sand flies (Diptera, Psychodidae) from Porto Velho, Northern Brazil." volume="213" year="2021">Silva et al. 2021</bibRefCitation>
,
<bibRefCitation author="Torchitte, A P A" journalOrPublisher="Brazilian Journal of Biology" pageId="0" pageNumber="90015" pagination="225 - 227" refId="B7910691" refString="Torchitte, A P A, Pereira Junior, A M, Paulo, P F M, Costa, G S, Castro, T S, Ferreira, R G M, Medeiros, J F, 2020. Identification of sand flies (Diptera: Psychodidae) and blood meal sources in periurban areas of Ji-Parana municipality, Western Brazilian Amazon. Brazilian Journal of Biology 81 (1): 225 - 227" title="Identification of sand flies (Diptera: Psychodidae) and blood meal sources in periurban areas of Ji-Parana municipality, Western Brazilian Amazon." volume="81" year="2020">Torchitte et al. 2020</bibRefCitation>
</paragraph>
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</treatment>
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157
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<taxonomicName LSID="49039DDC-219F-958A-FA8C-A222DB2E50B1" authority="Dahlbom, 1845" authorityName="Dahlbom" authorityYear="1845" class="Insecta" family="Chrysididae" genus="Chrysura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chrysura foveata" order="Hymenoptera" pageId="26" pageNumber="105" phylum="Arthropoda" rank="species" species="foveata">Chrysura foveata Dahlbom, 1845</taxonomicName>
<figureCitation captionStart="Plate 24" captionStartId="F24" captionText="Plate 24. Chrysura foveata Dahlbom, 1845, syntype male. A Habitus, dorso-lateral view B head, frontal view C head and mesosoma, dorsal view D metasoma, dorsal view E third metasomal tergite, dorsal view F metasomal sternites, ventral view." pageId="26" pageNumber="105">Plate 24</figureCitation>
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<paragraph pageId="27" pageNumber="106">
<taxonomicName authorityName="Dahlbom" authorityYear="1845" class="Insecta" family="Chrysididae" genus="Chrysura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chrysura foveata" order="Hymenoptera" pageId="27" pageNumber="106" phylum="Arthropoda" rank="species" species="foveata">
<pageBreakToken pageId="27" pageNumber="106" start="start">Chrysura</pageBreakToken>
foveata
</taxonomicName>
:
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: 6.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="106" type="type locality">
<paragraph pageId="27" pageNumber="106">Type locality.</paragraph>
<paragraph pageId="27" pageNumber="106">Egypt.</paragraph>
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<subSubSection pageId="27" pageNumber="106" type="syntype ♀">
<paragraph pageId="27" pageNumber="106">Syntype ♀.</paragraph>
<paragraph pageId="27" pageNumber="106">[Egypt] [Hedb.] [NHRS-HEVA000001083].</paragraph>
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<subSubSection pageId="27" pageNumber="106" type="syntype ♂">
<paragraph pageId="27" pageNumber="106">Syntype ♂.</paragraph>
<paragraph pageId="27" pageNumber="106">[Egypt] [Hedb.] [NHRS-HEVA000001084].</paragraph>
<caption pageId="27" pageNumber="106" start="Plate 24" startId="F24">
<paragraph pageId="27" pageNumber="106">
<emphasis bold="true" pageId="27" pageNumber="106">Plate 24.</emphasis>
<taxonomicName authorityName="Dahlbom" authorityYear="1845" class="Insecta" family="Chrysididae" genus="Chrysura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chrysura foveata" order="Hymenoptera" pageId="27" pageNumber="106" phylum="Arthropoda" rank="species" species="foveata">
<emphasis italics="true" pageId="27" pageNumber="106">Chrysura foveata</emphasis>
</taxonomicName>
Dahlbom, 1845, syntype male.
<emphasis bold="true" pageId="27" pageNumber="106">A</emphasis>
Habitus, dorso-lateral view
<emphasis bold="true" pageId="27" pageNumber="106">B</emphasis>
head, frontal view
<emphasis bold="true" pageId="27" pageNumber="106">C</emphasis>
head and mesosoma, dorsal view
<emphasis bold="true" pageId="27" pageNumber="106">D</emphasis>
metasoma, dorsal view
<emphasis bold="true" pageId="27" pageNumber="106">E</emphasis>
third metasomal tergite, dorsal view
<emphasis bold="true" pageId="27" pageNumber="106">F</emphasis>
metasomal sternites, ventral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="27" pageNumber="106" type="remarks">
<paragraph pageId="27" pageNumber="106">Remarks.</paragraph>
<paragraph pageId="27" pageNumber="106">
<taxonomicName authorityName="Dahlbom" authorityYear="1845" class="Insecta" family="Chrysididae" genus="Chrysura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chrysura foveata" order="Hymenoptera" pageId="27" pageNumber="106" phylum="Arthropoda" rank="species" species="foveata">
<emphasis italics="true" pageId="27" pageNumber="106">Chrysura foveata</emphasis>
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was described based on few specimens (
<emphasis italics="true" pageId="27" pageNumber="106">rar.</emphasis>
) considered as females, but in the collection one female and one male are found.
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: 172) gave a subsequent description of the species, which is not very precise, especially with respect to the colour.
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: 497) placed
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<emphasis italics="true" pageId="27" pageNumber="106">Chrysura foveata</emphasis>
</taxonomicName>
in synonym with
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1853" class="Insecta" family="Chrysididae" genus="Chrysura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chrysura trimaculata" order="Hymenoptera" pageId="27" pageNumber="106" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis italics="true" pageId="27" pageNumber="106">Chrysura trimaculata</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
, 1853), without type examination. This synonym is in error;
<taxonomicName authorityName="Dahlbom" authorityYear="1845" class="Insecta" family="Chrysididae" genus="Chrysura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chrysura foveata" order="Hymenoptera" pageId="27" pageNumber="106" phylum="Arthropoda" rank="species" species="foveata">
<emphasis italics="true" pageId="27" pageNumber="106">Chrysura foveata</emphasis>
</taxonomicName>
was described from Egypt, whereas
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1853" class="Insecta" family="Chrysididae" genus="Chrysura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chrysura trimaculata" order="Hymenoptera" pageId="27" pageNumber="106" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis italics="true" pageId="27" pageNumber="106">Chrysura trimaculata</emphasis>
</taxonomicName>
is a Euro-Sibiric species, not distributed in northern Africa and belonging to a different genus.
<taxonomicName authorityName="Dahlbom" authorityYear="1845" class="Insecta" family="Chrysididae" genus="Chrysura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chrysura foveata" order="Hymenoptera" pageId="27" pageNumber="106" phylum="Arthropoda" rank="species" species="foveata">
<emphasis italics="true" pageId="27" pageNumber="106">Chrysura foveata</emphasis>
</taxonomicName>
belongs to the
<taxonomicName authorityName="Abeille de Perrin" authorityYear="1878" class="Insecta" family="Chrysididae" genus="Chrysis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysis hydropica" order="Hymenoptera" pageId="27" pageNumber="106" phylum="Arthropoda" rank="species" species="hydropica">
<emphasis italics="true" pageId="27" pageNumber="106">Chrysis hydropica</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="106" type="current status">
<paragraph pageId="27" pageNumber="106">Current status.</paragraph>
<paragraph pageId="27" pageNumber="106">
<taxonomicName class="Insecta" family="Chrysididae" genus="Chrysis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysis foveata" order="Hymenoptera" pageId="27" pageNumber="106" phylum="Arthropoda" rank="species" species="foveata">
<emphasis italics="true" pageId="27" pageNumber="106">Chrysis foveata</emphasis>
</taxonomicName>
(Dahlbom, 1845) (transferred by
<bibRefCitation author="Mocsary, A" journalOrPublisher="Hungarian Academy of Sciences, Budapest" pageId="46" pageNumber="125" refId="B62" refString="Mocsary, A, 1889. Monographia Chrysididarum Orbis Terrestris Universi. Hungarian Academy of Sciences, Budapest" title="Monographia Chrysididarum Orbis Terrestris Universi." year="1889">
<normalizedToken originalValue="Mocsáry">Mocsary</normalizedToken>
1889
</bibRefCitation>
: 292).
</paragraph>
</subSubSection>
</treatment>
</document>

179
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<mods:title id="D73ED7209B17AFA47EEFEF4245C770C5">A new species of the genus Bryodemella (s. str.) Yin, 1982 from China (Orthoptera: Acridoidea: Oedipodidae)</mods:title>
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<treatment id="490487B1A23DFFEDFF0EFB1BFB3433DE" ID-DOI="http://doi.org/10.5281/zenodo.5954742" ID-GBIF-Taxon="143637689" ID-Zenodo-Dep="5954742" LSID="urn:lsid:plazi:treatment:490487B1A23DFFEDFF0EFB1BFB3433DE" httpUri="http://treatment.plazi.org/id/490487B1A23DFFEDFF0EFB1BFB3433DE" lastPageId="2" lastPageNumber="388" pageId="0" pageNumber="386">
<subSubSection id="89B7652CA23DFFEFFF0EFB1BFCCA36AC" box="[151,768,1180,1206]" pageId="0" pageNumber="386" type="nomenclature">
<paragraph id="C11236A7A23DFFEFFF0EFB1BFCCA36AC" blockId="0.[151,1436,1180,2001]" box="[151,768,1180,1206]" pageId="0" pageNumber="386">
<heading id="9A5A81CBA23DFFEFFF0EFB1BFCCA36AC" bold="true" box="[151,768,1180,1206]" fontSize="11" level="1" pageId="0" pageNumber="386" reason="1">
<emphasis id="F3D9EAB5A23DFFEFFF0EFB1BFCCA36AC" bold="true" box="[151,768,1180,1206]" pageId="0" pageNumber="386">
<taxonomicName id="06AD4D24A23DFFEFFF0EFB1BFEFA36AC" ID-CoL="8NNMT" baseAuthorityName="Zacher" baseAuthorityYear="1919" box="[151,304,1180,1206]" class="Insecta" family="Acrididae" genus="Bryodemella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="0" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="F3D9EAB5A23DFFEFFF0EFB1BFEFA36AC" bold="true" box="[151,304,1180,1206]" italics="true" pageId="0" pageNumber="386">Bryodemella</emphasis>
</taxonomicName>
(
<emphasis id="F3D9EAB5A23DFFEFFED8FB1BFEB536AC" bold="true" box="[321,383,1180,1206]" italics="true" pageId="0" pageNumber="386">s. str.</emphasis>
)
<emphasis id="F3D9EAB5A23DFFEFFE08FB1BFDDD36AC" bold="true" box="[401,535,1180,1206]" italics="true" pageId="0" pageNumber="386">rufifemura</emphasis>
sp. nov. (Figs. a–f)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89B7652CA23DFFEFFF0EFB38FE0436E3" pageId="0" pageNumber="386" type="materials_examined">
<paragraph id="C11236A7A23DFFEFFF0EFB38FE0436E3" blockId="0.[151,1436,1180,2001]" pageId="0" pageNumber="386">
Holotype: male.
<materialsCitation id="71C53CFAA23DFFEFFED5FB38FE0436E3" ID-GBIF-Occurrence="1838302512" collectingDate="2013-09-05" collectorName="Zhang Dao-Chuan &amp; Zhi &amp; Chao &amp; Wang Peng-Xiang." location="Langkazi" pageId="0" pageNumber="386" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="paratype">
<typeStatus id="1E168805A23DFFEFFED5FB38FE7236CC" box="[332,440,1215,1238]" pageId="0" pageNumber="386" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="D7ABFD2EA23DFFEFFE5AFB38FDD936CC" box="[451,531,1215,1238]" pageId="0" pageNumber="386" type="male">1 male</specimenCount>
,
<specimenCount id="D7ABFD2EA23DFFEFFD8EFB38FDB936CC" box="[535,627,1215,1238]" pageId="0" pageNumber="386" type="female">1 female</specimenCount>
,
<location id="C472607CA23DFFEFFDE6FB38FD2F36CC" LSID="urn:lsid:plazi:treatment:490487B1A23DFFEDFF0EFB1BFB3433DE:C472607CA23DFFEFFDE6FB38FD2F36CC" box="[639,741,1215,1238]" name="Langkazi" pageId="0" pageNumber="386" stateProvince="Xizang">Langkazi</location>
,
<collectingRegion id="0369F845A23DFFEFFD68FB38FC8B36CC" box="[753,833,1215,1238]" country="China" name="Xizang" pageId="0" pageNumber="386">Xizang</collectingRegion>
,
<collectingDate id="A557E98FA23DFFEFFCD4FB38FC0A36CC" box="[845,960,1215,1238]" pageId="0" pageNumber="386" value="2013-09-05">
<date id="B5131067A23DFFEFFCD4FB38FC0A36CC" box="[845,960,1215,1238]" pageId="0" pageNumber="386" value="2013-09-05">2013-IX-5</date>
</collectingDate>
, collected by
<collectorName id="6C585371A23DFFEFFBCEFB38FAEA36CC" box="[1111,1312,1215,1238]" pageId="0" pageNumber="386">Zhang Dao-Chuan</collectorName>
,
<collectorName id="6C585371A23DFFEFFAB4FB38FA9836CC" box="[1325,1362,1215,1238]" pageId="0" pageNumber="386">Zhi</collectorName>
Yong-
<collectorName id="6C585371A23DFFEFFF0EFB65FF0536E3" box="[151,207,1250,1273]" pageId="0" pageNumber="386">Chao</collectorName>
and
<collectorName id="6C585371A23DFFEFFE9AFB65FE0436E3" box="[259,462,1250,1273]" pageId="0" pageNumber="386">Wang Peng-Xiang.</collectorName>
</materialsCitation>
</paragraph>
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<subSubSection id="89B7652CA23DFFEDFF5EFA83FB393323" lastPageId="2" lastPageNumber="388" pageId="0" pageNumber="386" type="description">
<paragraph id="C11236A7A23DFFEFFF5EFA83FE8E353E" blockId="0.[151,1436,1180,2001]" pageId="0" pageNumber="386">
<emphasis id="F3D9EAB5A23DFFEFFF5EFA83FECB3706" bold="true" box="[199,257,1284,1308]" pageId="0" pageNumber="386">Male</emphasis>
. Body medium in size. Length of head obviously shorter than that of pronotum. Vertex short and broad, lateral margins distinct. Frons upright from lateral view, frontal ridge distinctly broad between antennae, constricted just below median ocellus, and not reaching to clypeus downward. Antennae filiform, reaching posterior margin of pronotum, length of a middle segment 2.0 times width. Foveolae slightly distinct, almost equilateral triangle. Eyes oval?vertical diameter 1.2 times horizontal diameter, and slightly longer than that of subocular groove. Median carina of pronotum thin, distinct in prozona yet indistinct in metazona, obviously incised by hind sulcus, metazona about 2 times longer than prozona, lateral keels of pronotum slightly distinct on metazona. Pronotum with dense strumae and short carinae, anterior margin straight, posterior margin almost right-angular. Prosternum appreciably swelled. Both elytra and hindwings developed, almost extending to the end of hind tibia, intercalary vein in medial area of elytra irregular and very weak. Main longitudinal veins of hindwings obviously thickened in basal half. Minimum width of interspace between mesosternal lobes 1.5 times its length. Hind femur dumpy, length 3.0 times maximal width, upper basal lobi longer than lower basal lobi, dorsal carina wholly smooth, lower knee lobes of inner side tilted inward. Outer side of hind tibia with 8 spines, and the inner side with 10 spines, outer apical spine absent. Arolium between claws not reaching the middle of claws. Tympanal organ developed, tympanic flap small, only covering minor part of tympanic cavity. Cerci long cylindrical. Supra-anal plate triangular, dorsal side flat. Subgenital plate brevi-conic, apex blunt. Epiphallus as shown in Fig. b.
</paragraph>
<paragraph id="C11236A7A23DFFEFFF5EF8A8FBD035CB" blockId="0.[151,1436,1180,2001]" pageId="0" pageNumber="386">
<emphasis id="F3D9EAB5A23DFFEFFF5EF8A8FEEB355D" bold="true" box="[199,289,1839,1863]" pageId="0" pageNumber="386">Female.</emphasis>
Body larger and more robust than male. Antennae shorter, not reaching hind margin of pronotum, length of a middle segment 2 times width. Vertical diameter of eyes shorter than that of subocular groove. Metazona of pronotum about 1.9 times prozona in length. Minimum width of interspace between mesosternal lobes 2.0 times its length. Elytra shorter, extending over the end of hind femur, intercalary vein of medial area absent. Ovipositor short and thick, apical part hook-like, upper external margin of dorsal valves smooth. Others same as male.
</paragraph>
<paragraph id="C11236A7A23CFFEEFF0EF833FC9A35FD" blockId="1.[151,1436,1972,2023]" pageId="1" pageNumber="387">
<emphasis id="F3D9EAB5A23CFFEEFF0EF833FD1D35D3" bold="true" box="[151,727,1972,1994]" pageId="1" pageNumber="387">
FIGURES a–f.
<taxonomicName id="06AD4D24A23CFFEEFEAAF833FE7A35D0" baseAuthorityName="Zacher" baseAuthorityYear="1919" box="[307,432,1972,1994]" class="Insecta" family="Acrididae" genus="Bryodemella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="F3D9EAB5A23CFFEEFEAAF833FE7A35D0" bold="true" box="[307,432,1972,1994]" italics="true" pageId="1" pageNumber="387">Bryodemella</emphasis>
</taxonomicName>
(s. str.)
<emphasis id="F3D9EAB5A23CFFEEFD99F833FDA735D0" bold="true" box="[512,621,1972,1994]" italics="true" pageId="1" pageNumber="387">rufifemura</emphasis>
sp. nov. a.
</emphasis>
Body dorsal view
<emphasis id="F3D9EAB5A23CFFEEFC12F833FC5735D3" bold="true" box="[907,925,1972,1993]" pageId="1" pageNumber="387">Ƌ</emphasis>
;
<emphasis id="F3D9EAB5A23CFFEEFC30F833FC7635D3" bold="true" box="[937,956,1972,1993]" pageId="1" pageNumber="387">b.</emphasis>
Epiphallus;
<emphasis id="F3D9EAB5A23CFFEEFBACF833FB8C35D3" bold="true" box="[1077,1094,1972,1993]" pageId="1" pageNumber="387">c.</emphasis>
Head and pronotum dorsal view
<emphasis id="F3D9EAB5A23CFFEEFA1AF833FA5F35D3" bold="true" box="[1411,1429,1972,1993]" pageId="1" pageNumber="387">Ƌ</emphasis>
;
<emphasis id="F3D9EAB5A23CFFEEFF0EF855FF6135FD" bold="true" box="[151,171,2002,2023]" pageId="1" pageNumber="387">d.</emphasis>
Head lateral view
<specimenCount id="D7ABFD2EA23CFFEEFEC6F855FEB835FD" box="[351,370,2002,2023]" pageId="1" pageNumber="387" type="female">♀</specimenCount>
;
<emphasis id="F3D9EAB5A23CFFEEFEE7F855FE4435FD" bold="true" box="[382,398,2002,2023]" pageId="1" pageNumber="387">e.</emphasis>
Hind femur inner side;
<emphasis id="F3D9EAB5A23CFFEEFDEDF855FD4835FD" bold="true" box="[628,642,2002,2023]" pageId="1" pageNumber="387">f.</emphasis>
Body lateral view
<specimenCount id="D7ABFD2EA23CFFEEFCA1F855FC8135FD" box="[824,843,2002,2023]" pageId="1" pageNumber="387" type="female">♀</specimenCount>
.
</paragraph>
<paragraph id="C11236A7A23FFFEDFF5EFF10FB0532EE" blockId="2.[151,1436,151,452]" pageId="2" pageNumber="388">
<emphasis id="F3D9EAB5A23FFFEDFF5EFF10FE8832B5" bold="true" box="[199,322,151,175]" pageId="2" pageNumber="388">Coloration</emphasis>
: Body brown or grayish brown. The base of a quarter of elytra darkish. Base of hindwings red except darker pre-anal area, fascia darker from anterior to posterior margins. Hind femur with red on full inner side, with a darker fascia in the middle, and with two dark bands on upper side. Hind tibia orange red. Tarsus red.
</paragraph>
<paragraph id="C11236A7A23FFFEDFF5EFF78FB393323" blockId="2.[151,1436,151,452]" pageId="2" pageNumber="388">
<emphasis id="F3D9EAB5A23FFFEDFF5EFF78FE7E330D" bold="true" box="[199,436,255,279]" pageId="2" pageNumber="388">Measurement (mm):</emphasis>
Length of body: male 26.9–27.5, female 34.6. Length of pronotum: male 7.5–8.7, female 10.4. Length of elytra: male 27.4–28.2, female 24.1. Length of hind femur: male 11.8–12.6, female 17.5.
</paragraph>
</subSubSection>
<subSubSection id="89B7652CA23FFFEDFF5EFEC3FB1C33BB" pageId="2" pageNumber="388" type="diagnosis">
<paragraph id="C11236A7A23FFFEDFF5EFEC3FB1C33BB" blockId="2.[151,1436,151,452]" pageId="2" pageNumber="388">
<emphasis id="F3D9EAB5A23FFFEDFF5EFEC3FEF63346" bold="true" box="[199,316,324,348]" pageId="2" pageNumber="388">Diagnosis.</emphasis>
The new species
<taxonomicName id="06AD4D24A23FFFEDFD98FEC3FD433346" baseAuthorityName="Zacher" baseAuthorityYear="1919" box="[513,649,324,348]" class="Insecta" family="Acrididae" genus="Bryodemella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="genus">
<emphasis id="F3D9EAB5A23FFFEDFD98FEC3FD433346" box="[513,649,324,348]" italics="true" pageId="2" pageNumber="388">Bryodemella</emphasis>
</taxonomicName>
(
<emphasis id="F3D9EAB5A23FFFEDFD03FEC2FD1F3346" box="[666,725,325,348]" italics="true" pageId="2" pageNumber="388">s. str.</emphasis>
)
<emphasis id="F3D9EAB5A23FFFEDFD7CFEC3FC9D3346" box="[741,855,324,348]" italics="true" pageId="2" pageNumber="388">rufifemura</emphasis>
<emphasis id="F3D9EAB5A23FFFEDFCF9FEC3FC723346" bold="true" box="[864,952,324,348]" pageId="2" pageNumber="388">sp. nov.</emphasis>
is similar to
<taxonomicName id="06AD4D24A23FFFEDFBD2FEC3FB193346" baseAuthorityName="Zacher" baseAuthorityYear="1919" box="[1099,1235,324,348]" class="Insecta" family="Acrididae" genus="Bryodemella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="genus">
<emphasis id="F3D9EAB5A23FFFEDFBD2FEC3FB193346" box="[1099,1235,324,348]" italics="true" pageId="2" pageNumber="388">Bryodemella</emphasis>
</taxonomicName>
(
<emphasis id="F3D9EAB5A23FFFEDFB7DFEC2FAD53346" box="[1252,1311,325,348]" italics="true" pageId="2" pageNumber="388">s. str.</emphasis>
)
<emphasis id="F3D9EAB5A23FFFEDFAA9FEC3FA563346" box="[1328,1436,324,348]" italics="true" pageId="2" pageNumber="388">diamesum</emphasis>
(Bei-Bienko, 1930), but differs from the latter by red color of almost full inner side of hind femur, median keel of pronotum indistinct in metazoan and vertical diameter of eye shorter than subocular groove in female.
</paragraph>
</subSubSection>
<subSubSection id="89B7652CA23FFFEDFF5EFE2BFB3433DE" box="[199,1278,428,452]" pageId="2" pageNumber="388" type="etymology">
<paragraph id="C11236A7A23FFFEDFF5EFE2BFB3433DE" blockId="2.[151,1436,151,452]" box="[199,1278,428,452]" pageId="2" pageNumber="388">
<emphasis id="F3D9EAB5A23FFFEDFF5EFE2BFE8833DE" bold="true" box="[199,322,428,452]" pageId="2" pageNumber="388">Etymology</emphasis>
: The specific epithet is named for coloration of hind femur by
<emphasis id="F3D9EAB5A23FFFEDFC7BFE2BFBDB33DE" box="[994,1041,428,452]" italics="true" pageId="2" pageNumber="388">rufi-</emphasis>
meaning red in Latin.
</paragraph>
</subSubSection>
</treatment>
</document>

153
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@ -0,0 +1,153 @@
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<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
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<mods:namePart>Means, Jackson C.</mods:namePart>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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<mods:namePart>Hennen, Derek A.</mods:namePart>
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<mods:namePart>Marek, Paul E.</mods:namePart>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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<treatment ID-GBIF-Taxon="181004724" LSID="urn:lsid:plazi:treatment:4904BD244F185008A2DBCFB047DF8648" httpUri="http://treatment.plazi.org/id/4904BD244F185008A2DBCFB047DF8648" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="7PLGS" ID-ENA="2675563" LSID="4904BD24-4F18-5008-A2DB-CFB047DF8648" authority="“ Hanging Rock ”" authorityName="&quot; Hanging Rock &quot;" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus" status="incertae sedis">Nannaria &quot;Hanging Rock&quot;</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">incertae sedis</taxonomicNameLabel>
<figureCitation captionStart="Figure 118" captionStartId="F118" captionText="Figure 118. Nannaria &quot; Hanging Rock &quot; incertae sedis ♂ (VMNH, NAN 0360) left gonopod A anterior view; red arrow indicates pointed lateral flange B medial view C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure118" httpUri="https://binary.pensoft.net/fig/531221" pageId="0" pageNumber="1">Fig. 118</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355691" collectingDate="2007-11-14" collectorName="R. Hoffman, For, Suppl." country="United States" county="Scott County" elevation="152" latitude="36.8589" location="Rec Area" longLatPrecision="6" longitude="-82.434" municipality="Hanging Rock" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<collectingCountry name="United States of America">United States</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
</emphasis>
•
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Scott County</collectingCounty>
, woods off Rt. 72, ca
<locationDeviation location="Dungannon">
<quantity metricMagnitude="3" metricUnit="m" metricValue="4.828032" unit="mi" value="3.0">3 mi</quantity>
NE of Dungannon
</locationDeviation>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.524" unit="ft" value="500.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.524" unit="ft" value="500.0">500 ft.</elevation>
</quantity>
beyond entrance to
<collectingMunicipality>Hanging Rock</collectingMunicipality>
USFS
<location LSID="urn:lsid:plazi:treatment:4904BD244F185008A2DBCFB047DF8648:CBD79087B9F2F7ADD9F3BF110987CD66" country="United States" county="Scott County" latitude="36.8589" longLatPrecision="6" longitude="-82.434" municipality="Hanging Rock" name="Rec Area" stateProvince="Virginia">Rec Area</location>
;
<geoCoordinate degrees="36.8589" direction="north" orientation="latitude" precision="5" value="36.8589">36.8589°N</geoCoordinate>
, -
<geoCoordinate degrees="82.4340" direction="west" orientation="longitude" precision="5" value="-82.434">82.4340°W</geoCoordinate>
;
<collectingDate value="2007-11-14">14 Nov. 2007</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0360.
<collectorName>For</collectorName>
detailed collection data see
<collectorName>Suppl.</collectorName>
material 7
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Hypothesized placement.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="&quot; Hanging Rock &quot;" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
&quot;Hanging Rock&quot; specimens are hypothesized to be closely related to
<taxonomicName authorityName="&quot; Hanging Rock &quot;" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
<normalizedToken originalValue="“Bina”">&quot;Bina&quot;</normalizedToken>
based on the following combination of gonopod characters: With predominantly straight acropodite and short distal zone. Prefemoral process arising medially from pronounced, tooth-like prefemoral spine.
<taxonomicName authorityName="&quot; Hanging Rock &quot;" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
&quot;Hanging Rock&quot; can be separated from
<normalizedToken originalValue="“Bina”">&quot;Bina&quot;</normalizedToken>
based on the following combination of characters: Acropodite tip pointed, not rounded as in
<normalizedToken originalValue="“Bina”">&quot;Bina&quot;</normalizedToken>
. Acropodite with pointed lateral flange near apex (Fig.
<figureCitation captionStart="Figure 118" captionStartId="F118" captionText="Figure 118. Nannaria &quot; Hanging Rock &quot; incertae sedis ♂ (VMNH, NAN 0360) left gonopod A anterior view; red arrow indicates pointed lateral flange B medial view C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure118" httpUri="https://binary.pensoft.net/fig/531221" pageId="0" pageNumber="1">118A</figureCitation>
, red arrow), not rounded as in
<normalizedToken originalValue="“Bina”">&quot;Bina&quot;</normalizedToken>
. Prefemoral process curving towards but not crossing acropodite in anterior view.
</paragraph>
<caption doi="10.3897/zookeys.1030.62544.figure118" httpUri="https://binary.pensoft.net/fig/531221" pageId="0" pageNumber="1" start="Figure 118" startId="F118">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 118.</emphasis>
<taxonomicName authorityName="&quot; Hanging Rock &quot;" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
&quot;Hanging Rock&quot;
<emphasis italics="true" pageId="0" pageNumber="1">incertae sedis</emphasis>
♂ (VMNH, NAN0360) left gonopod
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
anterior view; red arrow indicates pointed lateral flange
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
medial view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
posterior view. Scale bar: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

183
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<mods:title>A synoptic review of the aloes (Asphodelaceae, Alooideae) of KwaZulu-Natal, an ecologically diverse province in eastern South Africa</mods:title>
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<taxonomicName LSID="4904DE90-0D29-541B-84DC-353662B4BF57" authority="(Dyer) Klopper &amp; Gideon F. Sm." class="Liliopsida" family="Asphodelaceae" genus="Aloidendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aloidendron barberae" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="barberae">Aloidendron barberae (Dyer) Klopper &amp; Gideon F.Sm.</taxonomicName>
</paragraph>
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<paragraph pageId="0" pageNumber="1">Syn.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Dyer" authorityYear="1874" class="Liliopsida" family="Asphodelaceae" genus="Aloe" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aloe barberae" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="barberae">
<emphasis italics="true" pageId="0" pageNumber="1">Aloe barberae</emphasis>
</taxonomicName>
Dyer.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="common names">
<paragraph pageId="0" pageNumber="1">Common names.</paragraph>
<paragraph pageId="0" pageNumber="1">Tree aloe (English); boomaalwyn, mikaalwyn (Afrikaans); impondondo, indlabendlazi, inkalane unkulu, umgxwala (Zulu).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Arborescent plant, up to 18 m high.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Stem</emphasis>
10-18 m high, profusely branched dichotomously and rebranched from about middle, erect, without persistent dried leaves.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Leaves</emphasis>
densely rosulate, recurved, dull green, without spots, ensiform, deeply channelled, 60-90 cm long, 7-9 cm wide at base; sheath with greenish-white marginal border; margin narrow, white, cartilaginous, with firm, horny, brownish tipped, dull white, deltoid teeth, 2-3 mm long, 10-25 mm apart.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Inflorescences</emphasis>
0.4-0.6 m high, erect, dichotomously 3-branched.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Racemes</emphasis>
cylindrical, slightly acuminate, 20-30 cm long, dense.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Floral bracts</emphasis>
8-10 mm long,
<normalizedToken originalValue="±">+/-</normalizedToken>
1 mm wide.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pedicels</emphasis>
7-10 mm long.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Flowers</emphasis>
:
<emphasis italics="true" pageId="0" pageNumber="1">perianth</emphasis>
rose to rose-pink, greenish tipped, 33-37 mm long,
<normalizedToken originalValue="±">+/-</normalizedToken>
9 mm across ovary, not narrowed above ovary, widening towards middle, narrowing somewhat towards upturned mouth, cylindrical-ventricose; outer segments free almost to base;
<emphasis italics="true" pageId="0" pageNumber="1">stamens</emphasis>
exserted to 15 mm;
<emphasis italics="true" pageId="0" pageNumber="1">style</emphasis>
exserted 15-20 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="flowering time">
<paragraph pageId="0" pageNumber="1">Flowering time.</paragraph>
<paragraph pageId="0" pageNumber="1">May-August.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">Dense, tall bush and low forest, rocky slopes of wooded valleys.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Diagnostic characters.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Liliopsida" family="Asphodelaceae" genus="Aloidendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aloidendron barberae" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="barberae">
<emphasis italics="true" pageId="0" pageNumber="1">Aloidendron barberae</emphasis>
</taxonomicName>
is one of only two large-growing tree aloes indigenous to KwaZulu-Natal. These two aloes both have dichotomously branched stems and branches that lack persistent dried leaves.
<taxonomicName class="Liliopsida" family="Asphodelaceae" genus="Aloidendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aloidendron barberae" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="barberae">
<emphasis italics="true" pageId="0" pageNumber="1">Aloidendron barberae</emphasis>
</taxonomicName>
differs from
<taxonomicName class="Liliopsida" family="Asphodelaceae" genus="Aloidendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aloidendron tongaense" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tongaense">
<emphasis italics="true" pageId="0" pageNumber="1">Aloidendron tongaense</emphasis>
</taxonomicName>
in being much taller (up to 18 m) with more branches and having larger bright green leaves of 60-90 cm long (not dull green and 40-59 cm); their distribution ranges are also mutually exclusive. The inflorescence is also slightly taller at 0.4-0.6 m (not
<normalizedToken originalValue="±">+/-</normalizedToken>
0.35 m) and 3-branched from a single point (not up to 6-branched), with longer cylindrical racemes of 20-30 cm long (not capitate and 4-6 cm), bearing straight rose-pink flowers that are 33-37 mm long (not curved yellow flowers of 47-50 mm) with stamens exserted to 15 mm at anthesis (not 3-5 mm).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="conservation status">
<paragraph pageId="0" pageNumber="1">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="1">
Least Concern (
<bibRefCitation author="Raimondo, D" journalOrPublisher="South African National Biodiversity Institute, Pretoria" pageId="0" pageNumber="1" refId="B37" refString="Raimondo, D, Von Staden, L, Foden, W, Victor, JE, Helme, NA, Turner, RC, Kamundi, DA, Manyama, PA, 2009. Red List of South African Plants. Strelitzia 25. South African National Biodiversity Institute, Pretoria." title="Red List of South African Plants. Strelitzia 25. South African National Biodiversity Institute, Pretoria." year="2009">Raimondo et al. 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Occurs in scattered localities, often in inaccessible sites (with steep gradients), in a broad coastal zone from East London in the Eastern Cape, through KwaZulu-Natal and Mpumalanga, South Africa, also in Eswatini (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aloidendron barberae. Photo: G. W. Reynolds." figureDoi="10.3897/phytokeys.142.48365.figure3" httpUri="https://binary.pensoft.net/fig/388229" pageId="0" pageNumber="1">3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="notes">
<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Liliopsida" family="Asphodelaceae" genus="Aloidendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aloidendron barberae" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="barberae">
<emphasis italics="true" pageId="0" pageNumber="1">Aloidendron barberae</emphasis>
</taxonomicName>
is often cited as occurring in Mozambique, the latest of these being
<bibRefCitation author="Van Jaarsveld, E" journalOrPublisher="Aloe" pageId="0" pageNumber="1" refId="B52" refString="Van Jaarsveld, E, Judd, E, 2015. Tree Aloes of Africa. Penrock Publications, Cape Town." title="Tree Aloes of Africa. Penrock Publications, Cape Town." year="2015">Van Jaarsveld and Judd (2015)</bibRefCitation>
. However, an examination of available herbarium specimens at several South African and European herbaria has shown that specimens from Mozambique all represent
<taxonomicName lsidName="A. tongaense" pageId="0" pageNumber="1" rank="species" species="tongaense">
<emphasis italics="true" pageId="0" pageNumber="1">A. tongaense</emphasis>
</taxonomicName>
(
<bibRefCitation author="Walker, CC" journalOrPublisher="Bradleya" pageId="0" pageNumber="1" refId="B61" refString="Walker, CC, Smith, GF, Figueiredo, E, Klopper, RR, Crouch, NR, Condy, G, 2019b. Aloidendronbarberae. Flowering Plants of Africa 66: 8-21, t.2342." title="Aloidendronbarberae. Flowering Plants of Africa 66: 8 - 21, t. 2342." year="2019 b">Walker et al. 2019b</bibRefCitation>
). This is supported by
<bibRefCitation author="Burrows, JE" pageId="0" pageNumber="1" refId="B2" refString="Burrows, JE, Burrows, SM, Loetter, MC, Schmidt, E, 2018. Trees and Shrubs Mozambique. Publishing Print Matters (Pty) Ltd., Cape Town." title="Trees and Shrubs Mozambique. Publishing Print Matters (Pty) Ltd., Cape Town." year="2018">Burrows et al. (2018)</bibRefCitation>
who only treat the latter species. However, considering that
<taxonomicName lsidName="A. barberae" pageId="0" pageNumber="1" rank="species" species="barberae">
<emphasis italics="true" pageId="0" pageNumber="1">A. barberae</emphasis>
</taxonomicName>
is common on the South African side of the Lebombo range, it may well be present in nearby southern Mozambique, which borders on the foothills of the range. Further investigation is needed to confirm whether or not
<taxonomicName lsidName="A. barberae" pageId="0" pageNumber="1" rank="species" species="barberae">
<emphasis italics="true" pageId="0" pageNumber="1">A. barberae</emphasis>
</taxonomicName>
is present in this botanically under-explored part of Mozambique.
</paragraph>
<caption doi="10.3897/phytokeys.142.48365.figure3" httpUri="https://binary.pensoft.net/fig/388229" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName class="Liliopsida" family="Asphodelaceae" genus="Aloidendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aloidendron barberae" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="barberae">
<emphasis italics="true" pageId="0" pageNumber="1">Aloidendron barberae</emphasis>
</taxonomicName>
. Photo: G.W. Reynolds.
</paragraph>
</caption>
</subSubSection>
</treatment>
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185
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(L.) R. Br.
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<paragraph id="91F3297380BF66D3A3F4A260090612ED" pageNumber="1342">
Artbeschreibung:
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<quantity id="1DAB932A0DDD086C68FBD431B4B792AC" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageNumber="1334" unit="cm" value="12.5" valueMax="20.0" valueMin="5.0">5-20 cm</quantity>
hoch,
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dem Grund mit 2 fast
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, 3
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.
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in kurzer, 5-10
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Traube.
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bis rot
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, etwas abstehend, ca.
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lang.
<emphasis id="B4098EB64A59FCDDB265B5C721338730" bold="true" pageNumber="1342">Lippe braunrot</emphasis>
,
<quantity id="ED5E92F1DA1DAF45218C2922A5FCB617" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageNumber="1334" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3-4 mm</quantity>
lang, am Grund jederseits mit einem
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Zahn, bis
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die Mitte eingeschnitten, mit zugespitzten, spreizenden Abschnitten. Kein Sporn.
</paragraph>
</subSubSection>
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<paragraph id="2BB76E6F6E814BE40B86F09DC9EC2A7D" pageNumber="1342">
<normalizedToken id="7E6E4A52F23112DA1E617A10D288113E" originalValue="Blütezeit" pageNumber="1334">Bluetezeit</normalizedToken>
: 5-7
</paragraph>
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Standort und Verbreitung in der Schweiz: Moosige
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/ montan-subalpin / A, M in
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, J (
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bis SO)
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<paragraph id="9B42D482A6C2BA0390292A57457FC810" pageNumber="1342">Verbreitung global: Eurosibirisch-nordamerikanisch</paragraph>
</subSubSection>
<subSubSection id="78D3881ED56D19DED41A0956A23790A5" pageNumber="1342" type="biology_ecology">
<paragraph id="E22FF6CB92F200F77341B5C53CBECF8C" pageNumber="1342">
<normalizedToken id="23D9E84A31619C1E38E9B78344492273" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="95D0E51EAFA390C4C97943F7673A16EC" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." pageNumber="1342" publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="A8810B58673414610A543DC463C9179F" pageNumber="1342">
<table id="E31276AA2D3FA48FD348AC7528E18915" class="table" inLine="true" pageNumber="1342">
<tr id="A7F51F64B613E51E77740FF660EBE222" pageNumber="1342">
<th id="DCD233890A02A169543DF377218041F5" colspan="2" pageNumber="1342">
<emphasis id="FCC106C503810494AF17AFD9F492D11C" bold="true" pageNumber="1342">Bodenfaktoren</emphasis>
</th>
<th id="DC4225A307B72DBFEC2672A47C846381" colspan="2" pageNumber="1342">
<emphasis id="A20A503C1CF2E4DCFEED47B1630EA107" bold="true" pageNumber="1342">Klimafaktoren</emphasis>
</th>
<th id="FAC3B665933D73BD7D16A646F2210E80" colspan="2" pageNumber="1342">
<emphasis id="2EF7D093122B8A03FA3A8E253E27C464" bold="true" pageNumber="1342">Salztoleranz</emphasis>
</th>
</tr>
<tr id="688C0788F0720FBB26A026D253BAC452" pageNumber="1342">
<td id="F87FBF360EF19701A24C199944272EC2" pageNumber="1342">Feuchtezahl F</td>
<td id="18C9C3722E3B30FD94E2AD0B3D5CB265" pageNumber="1342" style="text-align:left;width:100px">
sehr feucht; Feuchtigkeit
<normalizedToken id="C0B60B423E7CA62A296174C24230EB4A" originalValue="mässig">maessig</normalizedToken>
wechselnd (
<normalizedToken id="B600BCDB635FD1154D528F59473FB52E" originalValue="±">+/-</normalizedToken>
1-2 Stufen)
</td>
<td id="7993875EDAC0C700E22AFEFC131B6F11" pageNumber="1342">Lichtzahl L</td>
<td id="E42522AE9928BCA242339842C0872E4A" pageNumber="1342" style="text-align:left;width:60px">sehr schattig</td>
<td id="61407A20F1908C299B7CFBB4726B43B6" pageNumber="1342">Salzzeichen</td>
<td id="9BC28AD4559F4290BC81C0359BD1A00D" pageNumber="1342" style="text-align:left;width:60px">--</td>
</tr>
<tr id="ABC7625FA2B73CF9F9A63BB76FC5A165" pageNumber="1342">
<td id="11ACE7C52EAF4D694364C399DC5BF3D1" pageNumber="1342">Reaktionszahl R</td>
<td id="6BA23BA6E2DC791100160CBFCC5A65C3" pageNumber="1342" style="text-align:left">stark sauer (pH 2.5-5.5)</td>
<td id="5BB7205EE661D68353EB44410619EBB0" pageNumber="1342">Temperaturzahl T</td>
<td id="27838CE0BDF8AE9688C50FA777742EC9" pageNumber="1342" style="text-align:left">unter-subalpin und ober-montan</td>
</tr>
<tr id="D5DD477D94B1E6E2F713C1EE1FEEEAE4" pageNumber="1342">
<td id="F24927F79671E90927EEB8EC13590FE3" pageNumber="1342">
<normalizedToken id="177C62C15ED1CAC476E877A61FBBB6B4" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
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sehr
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<td id="A1DE4F5B9E4E52AEA9F93153C1891DE6" pageNumber="1342">
<normalizedToken id="5F78BAC907F0A3066A42F6F2F82F1420" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
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<td id="DB401DEB2A6798F4E6CB48487996CD6C" pageNumber="1342" style="text-align:left">subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter)</td>
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</table>
</paragraph>
</subSubSection>
<subSubSection id="CA178265376E116A2099E7ABB3490AB4" pageNumber="1342" type="vernacular_names">
<paragraph id="2E8429FBD5F32E2BCD41984D3C7B16C1" pageNumber="1342">
Volksname Deutscher Name:
<vernacularName id="607E71A327E96836B41D64BF3B8D49B5" language="deu" pageNumber="1342">Kleines Zweiblatt</vernacularName>
Nom
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:
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<normalizedToken id="87002C14465F3B131EAE2D6A7095BC14" originalValue="Listère">Listere</normalizedToken>
en coeur
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Nome italiano:
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<taxonomicName id="7630B2B3A561394516F09A7F25CB524A" class="Liliopsida" family="Orchidaceae" genus="Listera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="genus">Listera</taxonomicName>
minore
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</paragraph>
</subSubSection>
</treatment>
</document>

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