357 lines
49 KiB
XML
357 lines
49 KiB
XML
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<mods:title id="5DE82C7C43B05A6E584DE2F66792D3B8">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="BC2286ED32ED8A5FEFFDE1864D78D9B5">Merle, Didier</mods:namePart>
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<mods:namePart id="86D0C4596DE727D2743DCF3CFE58C979">Ledon, Daniel</mods:namePart>
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<mods:affiliation id="29CF20C41134A952776DC59DB1A8712D">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:namePart id="57C0D714C2931D637800D57E09F0B33F">Goret, Bernard</mods:namePart>
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<mods:affiliation id="8A52A31DB8F1B89F496C3A4624A27048">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
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<mods:title id="4CC029F4822A02C116807911BC4831CA">Geodiversitas</mods:title>
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<mods:date id="59DEEA42438885F64036937093C1BC9D">2024</mods:date>
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<treatment id="038E87DAFFF3E411FC6CFAC3FD16FA1C" ID-DOI="http://doi.org/10.5281/zenodo.13989276" ID-GBIF-Taxon="244735063" ID-Zenodo-Dep="13989276" LSID="urn:lsid:zoobank.org:act:59D8D20D-41AD-43F9-8A3B-B247A150B335" httpUri="http://treatment.plazi.org/id/038E87DAFFF3E411FC6CFAC3FD16FA1C" lastPageId="23" lastPageNumber="516" pageId="21" pageNumber="514" scope_class="Gastropoda" scope_family="Muricidae" scope_phylum="Mollusca" scope_subFamily="Typhinae">
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<taxonomicName id="4C274D4FFFF3E413FC6CFAC3FB47FA9C" ID-CoL="CN5M3" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[980,1222,1301,1327]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="longicanalis" status="sp. nov.">
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<emphasis id="B953EADEFFF3E413FC6CFAC3FB47FA9C" bold="true" box="[980,1222,1301,1327]" italics="true" pageId="21" pageNumber="514">Timbellus longicanalis</emphasis>
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<subSubSection id="C33D6547FFF3E413FC42FAE2FA2EFA39" pageId="21" pageNumber="514" type="description">
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(
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A-D; 26E, F)
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urn:lsid:zoobank.org:act:
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<emphasis id="B953EADEFFF3E413FC95FA16FC56FA6B" box="[813,983,1472,1496]" italics="true" pageId="21" pageNumber="514">Timbellus fusoides</emphasis>
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–
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Merle
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<emphasis id="B953EADEFFF3E413FB89FA17FBE0FA6B" box="[1073,1121,1472,1496]" italics="true" pageId="21" pageNumber="514">et al.</emphasis>
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2011: 446
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, pl. 101 fig. 5a, b only [
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].
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. —
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.
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•
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,
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,
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;
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; middle Eocene (uppermost Lutetian);
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.F.A24101 (
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coll.), H:
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(
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;
|
||
<figureCitation id="131C2A49FFF3E413FAC1F992FA25F9EE" box="[1401,1444,1604,1629]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="21" pageNumber="514">26E</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4F3C91FFF3E413FC95F989FB38F921" ID-GBIF-Occurrence="4958671362" collectionCode="MNHN" collectorName="Le Marchand" country="France" location="France" pageId="21" pageNumber="514" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="B953EADEFFF3E413FC95F989FCFEF9C4" bold="true" box="[813,895,1631,1655]" pageId="21" pageNumber="514">
|
||
<typeStatus id="549C886EFFF3E413FC95F989FCFEF9C4" box="[813,895,1631,1655]" pageId="21" pageNumber="514" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B953EADEFFF3E413FC31F989FC49F9C4" bold="true" box="[905,968,1631,1655]" pageId="21" pageNumber="514">
|
||
<collectingCountry id="F330765CFFF3E413FC31F989FC49F9C4" box="[905,968,1631,1655]" name="France" pageId="21" pageNumber="514">France</collectingCountry>
|
||
</emphasis>
|
||
• 1 spm; same as for the holotype;
|
||
<collectionCode id="ED36AE09FFF3E413FB44F9B6FACCF9C4" box="[1276,1357,1632,1655]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="514">MNHN</collectionCode>
|
||
.F.A24100 (
|
||
<collectorName id="26D2531AFFF3E413FC8BF9ACFC39F921" box="[819,952,1658,1682]" pageId="21" pageNumber="514">Le Marchand</collectorName>
|
||
coll.) (
|
||
<figureCitation id="131C2A49FFF3E413FC46F9ACFBF8F922" box="[1022,1145,1658,1682]" captionStart="FIG" captionStartId="22.[131,142,1581,1598]" captionTargetBox="[132,1455,215,1538]" captionTargetId="figure-331@22.[698,1267,781,1349]" captionTargetPageId="22" captionText="FIG. 11. — Spiral cords and internal denticles of the outer lip in Timbellus longicanalis n. sp. (A-D) and type material of Murex fusoides (Deshayes, 1865) (E-I): A, B, holotype,MNHN.F.A24101 (Le Marchand coll.),Fresville, Manche,Normandy, France,middle Eocene (upper Lutetian);C, D, paratype,MNHN.F.A24100, Fresville;E, type material of M. fusoides in the Deshayes’s collection corresponding to two eroded T. tripteroides; F, G, lectotype designated herein,UCBL-EM 33384, Valmondois, Val d’Oise, Paris Basin, France, middle Eocene (Bartonian); H, I, paralectotype, UCBL-EM 33383, Sainte-Aulde (Caumont), Seine-et-Marne, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E-I, E. Robert (UCBL)." figureDoi="http://doi.org/10.5281/zenodo.13985784" httpUri="https://zenodo.org/record/13985784/files/figure.png" pageId="21" pageNumber="514">Figs 11C, D</figureCitation>
|
||
;
|
||
<figureCitation id="131C2A49FFF3E413FB3BF9ACFB2FF921" box="[1155,1198,1658,1682]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="21" pageNumber="514">26F</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFF3E413FC95F966FC3AF94F" pageId="21" pageNumber="514" type="etymology">
|
||
<paragraph id="8B9836CCFFF3E413FC95F966FC3AF94F" blockId="21.[811,1455,1711,1788]" pageId="21" pageNumber="514">
|
||
<smallCapsWord id="8D7EA010FFF3E413FC95F966FC24F975" baselines="1729,1730" box="[813,933,1712,1735]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="21" pageNumber="514">ETYMOLOGY</smallCapsWord>
|
||
. — From the Latin adjective
|
||
<emphasis id="B953EADEFFF3E413FB01F979FB72F974" box="[1209,1267,1711,1735]" italics="true" pageId="21" pageNumber="514">longus</emphasis>
|
||
meaning elongated and
|
||
<emphasis id="B953EADEFFF3E413FCE0F91CFC19F951" box="[856,920,1738,1762]" italics="true" pageId="21" pageNumber="514">canalis</emphasis>
|
||
meaning canal, refecting the elongated siphonal canal of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFF3E413FC95F8CCFC41F82F" pageId="21" pageNumber="514" type="materials_examined">
|
||
<paragraph id="8B9836CCFFF3E413FC95F8CCFBFDF8F8" blockId="21.[811,1456,1817,1868]" pageId="21" pageNumber="514">
|
||
<smallCapsWord id="8D7EA010FFF3E413FC95F8CCFCDCF89C" baselines="1835,1835" box="[813,861,1818,1841]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="21" pageNumber="514">
|
||
<typeStatus id="549C886EFFF3E413FC95F8CCFCDCF89C" box="[813,861,1818,1841]" pageId="21" pageNumber="514">TYPE</typeStatus>
|
||
</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFF3E413FCDAF8C9FC3DF883" baselines="1836" box="[866,956,1823,1840]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="21" pageNumber="514">HORIZON</smallCapsWord>
|
||
. — Calcaire de Fresville, middle Eocene (uppermost Lutetian), see
|
||
<bibRefCitation id="EFB64B3DFFF3E413FC0EF8E2FBF7F8FF" author="BIGNOT G. & HOMMERIL P. & LARSONNEUR C." box="[950,1142,1844,1868]" pageId="21" pageNumber="514" pagination="405 - 416" refId="ref42941" refString="BIGNOT G., HOMMERIL P. & LARSONNEUR C. 1968. - Le Lutetien au large du Cotentin, in Colloque sur l'Eocene (Paris, mai 1968). Memoires BRGM 58: 405 - 416." type="journal article" year="1968">
|
||
Bignot
|
||
<emphasis id="B953EADEFFF3E413FC47F8E3FBAFF8FF" box="[1023,1070,1844,1868]" italics="true" pageId="21" pageNumber="514">et al.</emphasis>
|
||
(1968)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF3E413FC95F8BCFC41F82F" blockId="21.[813,1457,1897,1948]" pageId="21" pageNumber="514">
|
||
<smallCapsWord id="8D7EA010FFF3E413FC95F8BCFCDEF8CC" baselines="1915,1915" box="[813,863,1898,1921]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="21" pageNumber="514">
|
||
<typeStatus id="549C886EFFF3E413FC95F8BCFCDEF8CC" box="[813,863,1898,1921]" pageId="21" pageNumber="514">TYPE</typeStatus>
|
||
</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFF3E413FCD0F8B9FC47F833" baselines="1916" box="[872,966,1903,1920]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="21" pageNumber="514">LOCALITY</smallCapsWord>
|
||
. —
|
||
<collectingCountry id="F330765CFFF3E413FBBAF8BCFBC9F832" box="[1026,1096,1898,1921]" name="France" pageId="21" pageNumber="514">France</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E3F82EFFF3E413FBEDF8BFFB44F832" box="[1109,1221,1897,1921]" country="France" name="Haute-Normandie" pageId="21" pageNumber="514">Normandy</collectingRegion>
|
||
, Manche, Fresville, see
|
||
<bibRefCitation id="EFB64B3DFFF3E413FC96F852FC3DF82F" author="APGN & ASSOCIATION PATRIMOINE GEOLOGIQUE DE NORMANDIE" box="[814,956,1924,1948]" pageId="21" pageNumber="514" refId="ref42597" refString="APGN (ASSOCIATION PATRIMOINE GEOLOGIQUE DE NORMANDIE) 2024. - Faluns cretaces et tertiaires de Fresville, BNO 0182. Available at: http: // www. apgn. fr / apgn / carte _ site / fic _ s 50 _ 75. pdf (last access on 03 / 2024)." type="book" year="2024">APGN (2024)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFF3E413FC95F86FFB84F858" pageId="21" pageNumber="514" type="distribution">
|
||
<paragraph id="8B9836CCFFF3E413FC95F86FFB84F858" blockId="21.[811,1455,1977,2027]" pageId="21" pageNumber="514">
|
||
<smallCapsWord id="8D7EA010FFF3E413FC95F86FFC36F87C" baselines="1994,1995" box="[813,951,1977,2000]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="21" pageNumber="514">DISTRIBUTION</smallCapsWord>
|
||
. —
|
||
<collectingRegion id="49E3F82EFFF3E413FC67F86FFBC8F862" box="[991,1097,1977,2001]" country="France" name="Haute-Normandie" pageId="21" pageNumber="514">Normandy</collectingRegion>
|
||
(
|
||
<location id="8EF86017FFF3E413FBECF86FFA95F862" LSID="urn:lsid:plazi:treatment:038E87DAFFF3E411FC6CFAC3FD16FA1C:8EF86017FFF3E413FBECF86FFA95F862" box="[1108,1300,1977,2001]" name="Calcaire de Fresville" pageId="21" pageNumber="514" stateProvince="Haute-Normandie">
|
||
Calcaire
|
||
<collectorName id="26D2531AFFF3E413FB1FF86FFA95F862" box="[1191,1300,1977,2001]" pageId="21" pageNumber="514">de Fresville</collectorName>
|
||
</location>
|
||
); middle Eocene (uppermost Lutetian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF586644FFF0E410FF3BF9FBFB15F92A" ID-DOI="http://doi.org/10.5281/zenodo.13985784" ID-Zenodo-Dep="13985784" httpUri="https://zenodo.org/record/13985784/files/figure.png" pageId="22" pageNumber="515" startId="22.[131,142,1581,1598]" targetBox="[132,1455,215,1538]" targetPageId="22" targetType="figure">
|
||
<paragraph id="8B9836CCFFF0E410FF3BF9FBFB15F92A" blockId="22.[131,1457,1581,1689]" pageId="22" pageNumber="515">
|
||
<smallCapsWord id="8D7EA010FFF0E410FF3BF9FBFF1CF98E" baselines="1594,1594" box="[131,157,1581,1598]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="22" pageNumber="515">FIG</smallCapsWord>
|
||
. 11. — Spiral cords and internal denticles of the outer lip in
|
||
<taxonomicName id="4C274D4FFFF0E410FD37F9FBFCC4F98D" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[655,837,1581,1598]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="515" phylum="Mollusca" rank="species" species="longicanalis" status="sp. nov.">
|
||
<emphasis id="B953EADEFFF0E410FD37F9FBFCC4F98D" box="[655,837,1581,1598]" italics="true" pageId="22" pageNumber="515">Timbellus longicanalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF0E410FCF2F9FBFCFBF98D" box="[842,890,1581,1598]" pageId="22" pageNumber="515" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="B953EADEFFF0E410FC3AF9FBFC25F98D" bold="true" box="[898,932,1581,1598]" pageId="22" pageNumber="515">A -D</emphasis>
|
||
) and type material of
|
||
<taxonomicName id="4C274D4FFFF0E410FBE1F9FBFAEAF98D" authority="(Deshayes, 1865)" baseAuthorityName="Deshayes" baseAuthorityYear="1865" box="[1113,1387,1581,1598]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="515" phylum="Mollusca" rank="species" species="fusoides">
|
||
<emphasis id="B953EADEFFF0E410FBE1F9FBFB56F98D" box="[1113,1239,1581,1598]" italics="true" pageId="22" pageNumber="515">Murex fusoides</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFF0E410FB59F9FBFAE6F98D" author="DESHAYES G. - P." box="[1249,1383,1581,1598]" pageId="22" pageNumber="515" pagination="101 - 107" refId="ref44001" refString="DESHAYES G. - P. 1865. - Description des Animaux sans vertebres decouverts dans le bassin de Paris pour servir de supplement a la description des coquilles fossiles des environs de Paris, comprenant une revue generale de toutes les especes actuellement connues. Tome 3. Bailliere, Paris, Livraisons 45 - 46, p. 201 - 288; Atlas 2, pl. 86 - 93 [2 January]; Livraisons 47 - 48, p. 289 - 424, Atlas 2, pl. 94 - 100 [10 June]; Livraisons 49 - 50, p. 425 - 668; Atlas 2, pl. 101 - 107 [15 December]." type="journal article" year="1865">Deshayes, 1865</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B953EADEFFF0E410FACDF9FBFA0EF98D" bold="true" box="[1397,1423,1581,1598]" pageId="22" pageNumber="515">E -I</emphasis>
|
||
):
|
||
<emphasis id="B953EADEFFF0E410FA25F9FBFA2BF98D" bold="true" box="[1437,1450,1581,1598]" pageId="22" pageNumber="515">A</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFF0E410FF3CF992FF10F9E6" bold="true" box="[132,145,1604,1621]" pageId="22" pageNumber="515">B</emphasis>
|
||
, holotype,MNHN.F.A24101 (Le Marchand coll.),Fresville, Manche,Normandy, France,middle Eocene (upper Lutetian);
|
||
<emphasis id="B953EADEFFF0E410FBD3F992FBF8F9E6" bold="true" box="[1131,1145,1604,1621]" pageId="22" pageNumber="515">C</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFF0E410FB3AF992FB11F9E6" bold="true" box="[1154,1168,1604,1621]" pageId="22" pageNumber="515">D</emphasis>
|
||
, paratype,MNHN.F.A24100, Fresville;
|
||
<emphasis id="B953EADEFFF0E410FF15F98CFF38F9D8" bold="true" box="[173,185,1626,1643]" pageId="22" pageNumber="515">E</emphasis>
|
||
, type material of
|
||
<taxonomicName id="4C274D4FFFF0E410FEFEF98CFE27F9D8" baseAuthorityName="Deshayes" baseAuthorityYear="1865" box="[326,422,1626,1643]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="515" phylum="Mollusca" rank="species" species="fusoides">
|
||
<emphasis id="B953EADEFFF0E410FEFEF98CFE27F9D8" box="[326,422,1626,1643]" italics="true" pageId="22" pageNumber="515">M. fusoides</emphasis>
|
||
</taxonomicName>
|
||
in the Deshayes’s collection corresponding to two eroded
|
||
<taxonomicName id="4C274D4FFFF0E410FC3FF98CFC78F9D8" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[903,1017,1626,1643]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="515" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF0E410FC3FF98CFC78F9D8" box="[903,1017,1626,1643]" italics="true" pageId="22" pageNumber="515">T. tripteroides</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B953EADEFFF0E410FBBBF98CFB8FF9D8" bold="true" box="[1027,1038,1626,1643]" pageId="22" pageNumber="515">F</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFF0E410FBAFF98CFBA4F9D8" bold="true" box="[1047,1061,1626,1643]" pageId="22" pageNumber="515">G</emphasis>
|
||
, lectotype designated herein,UCBL-EM 33384, Valmondois, Val d’Oise, Paris Basin, France, middle Eocene (Bartonian);
|
||
<emphasis id="B953EADEFFF0E410FD5EF9A7FD75F931" bold="true" box="[742,756,1649,1666]" pageId="22" pageNumber="515">H</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFF0E410FD47F9A7FC84F931" bold="true" box="[767,773,1649,1666]" pageId="22" pageNumber="515">I</emphasis>
|
||
, paralectotype, UCBL-EM 33383, Sainte-Aulde (Caumont), Seine-et-Marne, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E-I, E. Robert (UCBL).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33D6547FFF0E411FF3CF904FD16FA1C" lastPageId="23" lastPageNumber="516" pageId="22" pageNumber="515" type="description">
|
||
<paragraph id="8B9836CCFFF0E411FF3CF904FD27FE22" blockId="22.[131,778,1746,2028]" lastBlockId="23.[131,775,215,401]" lastPageId="23" lastPageNumber="516" pageId="22" pageNumber="515">
|
||
<smallCapsWord id="8D7EA010FFF0E410FF3CF904FE94F959" baselines="1765,1765" box="[132,277,1746,1772]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="22" pageNumber="515">DESCRIPTION</smallCapsWord>
|
||
Protoconch not preserved. Teleoconch up to
|
||
<quantity id="4CDF9B29FFF0E410FD2AF924FD67F8BF" box="[658,742,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="22" pageNumber="515" unit="mm" value="35.0">35 mm</quantity>
|
||
in height, up to
|
||
<quantity id="4CDF9B29FFF0E410FEACF8C4FEE3F89F" box="[276,354,1810,1836]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="515" unit="mm" value="14.0">14 mm</quantity>
|
||
in width, fusiform in profile, composed of six whorls. Moderately high spire. Last whorl up to 77% of total length. Apical angle 41° excluding spine and 61° including spines. Spiral sculpture with moderately marked primary cords on varices. First and second whorl eroded. Third whorl: appearance of weak s1 and P2 on intervarices. Fourth whorl: appearance of weak P1 cord. Fifth whorl: no change. Sixth whorl: P1-P4 marked, P5-P6 weak, s1, s2, s6, P6 not atrophied, s1-s2, s6 weak; ADP, MP and ABP weak. Axial sculpture eroded on two early whorls. From third last whorl: three varices and one intervarix intercalated. Higher relief of intervarices on P2. Varices lamellose, slightly winged. Appearance of P1 spine on third whorl, no other spines adapically. On last whorl P1 spine short, straight, slightly bent adaxially. On last whorl growth lamellae erect giving scabrous aspect to shell surface. Ovate aperture up to 29% of diameter and up to 78% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, slightly erect anteriorly, forming very slightly curved inductura at base. Parietal lip adherent. Outer lip with simple, weak denticles including ID, D1 to D6. Siphonal canal narrow, open, straight, up to 59% of apertural length. Pseudoumbilicus narrow.
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF1E411FF3CFE61FE9EFE63" blockId="23.[130,778,439,1455]" box="[132,287,439,465]" pageId="23" pageNumber="516">
|
||
<smallCapsWord id="8D7EA010FFF1E411FF3CFE61FE9EFE63" baselines="458,459" box="[132,287,439,465]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comparisons" pageId="23" pageNumber="516">COMPARISONS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF1E411FF3CFE0EFD16FA1C" blockId="23.[130,778,439,1455]" pageId="23" pageNumber="516">
|
||
This species was previously identified as
|
||
<taxonomicName id="4C274D4FFFF1E411FDF0FE0EFEC0FDA0" authority="(Deshayes, 1865)" baseAuthorityName="Deshayes" baseAuthorityYear="1865" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="fusoides">
|
||
<emphasis id="B953EADEFFF1E411FDF0FE0EFC89FE41" box="[584,776,472,498]" italics="true" pageId="23" pageNumber="516">Timbellus fusoides</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFF1E411FF34FE2EFEB8FDA0" author="DESHAYES G. - P." box="[140,313,504,531]" pageId="23" pageNumber="516" pagination="101 - 107" refId="ref44001" refString="DESHAYES G. - P. 1865. - Description des Animaux sans vertebres decouverts dans le bassin de Paris pour servir de supplement a la description des coquilles fossiles des environs de Paris, comprenant une revue generale de toutes les especes actuellement connues. Tome 3. Bailliere, Paris, Livraisons 45 - 46, p. 201 - 288; Atlas 2, pl. 86 - 93 [2 January]; Livraisons 47 - 48, p. 289 - 424, Atlas 2, pl. 94 - 100 [10 June]; Livraisons 49 - 50, p. 425 - 668; Atlas 2, pl. 101 - 107 [15 December]." type="journal article" year="1865">Deshayes, 1865</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFB64B3DFFF1E411FED2FE2EFDADFDA1" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[362,556,504,530]" pageId="23" pageNumber="516" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFF1E411FED2FE2EFE28FDA1" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[362,425,504,530]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFF1E411FE09FE2FFE64FDA1" box="[433,485,504,530]" italics="true" pageId="23" pageNumber="516">et al.</emphasis>
|
||
(2011
|
||
</bibRefCitation>
|
||
: pl. 101, fig. 5a, b). Nevertheless, attribution to
|
||
<taxonomicName id="4C274D4FFFF1E411FE17FDCFFDC6FD81" baseAuthorityName="Deshayes" baseAuthorityYear="1865" box="[431,583,536,562]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="fusoides">
|
||
<emphasis id="B953EADEFFF1E411FE17FDCFFDC6FD81" box="[431,583,536,562]" italics="true" pageId="23" pageNumber="516">Murex fusoides</emphasis>
|
||
</taxonomicName>
|
||
is problematic, as the type material on which that Bartonian species is based is poorly preserved. It corresponds to
|
||
<specimenCount id="9D21FD45FFF1E411FE57FD8EFDF4FDC1" box="[495,629,600,626]" count="2" pageId="23" pageNumber="516" type="generic" typeStatus="syntype">two syntypes</specimenCount>
|
||
housed in the collection of the University Lyon (no. UCBL-EM 33383-84;
|
||
<figureCitation id="131C2A49FFF1E411FF3AFD41FF60FD01" box="[130,225,663,690]" captionStart="FIG" captionStartId="22.[131,142,1581,1598]" captionTargetBox="[132,1455,215,1538]" captionTargetId="figure-331@22.[698,1267,781,1349]" captionTargetPageId="22" captionText="FIG. 11. — Spiral cords and internal denticles of the outer lip in Timbellus longicanalis n. sp. (A-D) and type material of Murex fusoides (Deshayes, 1865) (E-I): A, B, holotype,MNHN.F.A24101 (Le Marchand coll.),Fresville, Manche,Normandy, France,middle Eocene (upper Lutetian);C, D, paratype,MNHN.F.A24100, Fresville;E, type material of M. fusoides in the Deshayes’s collection corresponding to two eroded T. tripteroides; F, G, lectotype designated herein,UCBL-EM 33384, Valmondois, Val d’Oise, Paris Basin, France, middle Eocene (Bartonian); H, I, paralectotype, UCBL-EM 33383, Sainte-Aulde (Caumont), Seine-et-Marne, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E-I, E. Robert (UCBL)." figureDoi="http://doi.org/10.5281/zenodo.13985784" httpUri="https://zenodo.org/record/13985784/files/figure.png" pageId="23" pageNumber="516">Fig. 11E</figureCitation>
|
||
). The largest is a strongly eroded specimen with most of its axial sculpture, particularly its winged varices lost. This loss of sculpture gives the shell an artificially fusiform shape. The second specimen is smaller and also strongly eroded, but P1 spine is rather turned adapically as in some
|
||
<taxonomicName id="4C274D4FFFF1E411FD27FCC1FE34FCE2" authority="(Lamarck, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF1E411FD27FCC1FF77FCE2" italics="true" pageId="23" pageNumber="516">Timbellus tripteroides</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFF1E411FEBDFCE1FE2CFCE2" author="LAMARCK J. - B." box="[261,429,823,849]" pageId="23" pageNumber="516" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and its aperture resembles that of
|
||
<taxonomicName id="4C274D4FFFF1E411FF19FC8EFEB1FCC2" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[161,304,855,881]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF1E411FF19FC8EFEB1FCC2" box="[161,304,855,881]" italics="true" pageId="23" pageNumber="516">T. tripteroides</emphasis>
|
||
</taxonomicName>
|
||
. We designate herein this smaller specimen (UCBL-EM 33384;
|
||
<figureCitation id="131C2A49FFF1E411FEE7FCA1FE58FC22" box="[351,473,887,913]" captionStart="FIG" captionStartId="22.[131,142,1581,1598]" captionTargetBox="[132,1455,215,1538]" captionTargetId="figure-331@22.[698,1267,781,1349]" captionTargetPageId="22" captionText="FIG. 11. — Spiral cords and internal denticles of the outer lip in Timbellus longicanalis n. sp. (A-D) and type material of Murex fusoides (Deshayes, 1865) (E-I): A, B, holotype,MNHN.F.A24101 (Le Marchand coll.),Fresville, Manche,Normandy, France,middle Eocene (upper Lutetian);C, D, paratype,MNHN.F.A24100, Fresville;E, type material of M. fusoides in the Deshayes’s collection corresponding to two eroded T. tripteroides; F, G, lectotype designated herein,UCBL-EM 33384, Valmondois, Val d’Oise, Paris Basin, France, middle Eocene (Bartonian); H, I, paralectotype, UCBL-EM 33383, Sainte-Aulde (Caumont), Seine-et-Marne, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E-I, E. Robert (UCBL)." figureDoi="http://doi.org/10.5281/zenodo.13985784" httpUri="https://zenodo.org/record/13985784/files/figure.png" pageId="23" pageNumber="516">Fig. 11F, G</figureCitation>
|
||
) as the
|
||
<typeStatus id="549C886EFFF1E411FD9FFCA1FD08FC22" box="[551,649,887,913]" pageId="23" pageNumber="516" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C274D4FFFF1E411FD10FCAEFF23FC02" baseAuthorityName="Deshayes" baseAuthorityYear="1865" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="fusoides">
|
||
<emphasis id="B953EADEFFF1E411FD10FCAEFF23FC02" italics="true" pageId="23" pageNumber="516">M. fusoides</emphasis>
|
||
</taxonomicName>
|
||
. Therefore,
|
||
<taxonomicName id="4C274D4FFFF1E411FEA7FC41FE10FC02" baseAuthorityName="Deshayes" baseAuthorityYear="1865" box="[287,401,919,945]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="fusoides">
|
||
<emphasis id="B953EADEFFF1E411FEA7FC41FE10FC02" box="[287,401,919,945]" italics="true" pageId="23" pageNumber="516">M. fusoides</emphasis>
|
||
</taxonomicName>
|
||
can be regarded as a possible junior synonym of
|
||
<taxonomicName id="4C274D4FFFF1E411FEB2FC61FE19FC63" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[266,408,950,976]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF1E411FEB2FC61FE19FC63" box="[266,408,950,976]" italics="true" pageId="23" pageNumber="516">T. tripteroides</emphasis>
|
||
</taxonomicName>
|
||
or a
|
||
<emphasis id="B953EADEFFF1E411FE76FC61FDEDFC63" box="[462,620,950,976]" italics="true" pageId="23" pageNumber="516">nomen dubium</emphasis>
|
||
as it is impossible to clearly identify a species based on the type material. Moreover, Bartonian specimens from the Paris Basin identified as
|
||
<taxonomicName id="4C274D4FFFF1E411FF18FBC1FE8AFB83" baseAuthorityName="Deshayes" baseAuthorityYear="1865" box="[160,267,1046,1072]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="fusoides">
|
||
<emphasis id="B953EADEFFF1E411FF18FBC1FE8AFB83" box="[160,267,1046,1072]" italics="true" pageId="23" pageNumber="516">T. fusoides</emphasis>
|
||
</taxonomicName>
|
||
are present in several collections housed in the MNHN (e.g., Faullummel, Pons and Schtrock colls). Examination of these specimens shows them to represent eroded
|
||
<taxonomicName id="4C274D4FFFF1E411FF3EFBA1FE97FB23" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[134,278,1142,1168]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF1E411FF3EFBA1FE97FB23" box="[134,278,1142,1168]" italics="true" pageId="23" pageNumber="516">T. tripteroides</emphasis>
|
||
</taxonomicName>
|
||
(probably reworked from the Lutetian) or to large and eroded
|
||
<taxonomicName id="4C274D4FFFF1E411FEFCFB40FC82FB03" authority="(Lamarck, 1803)" baseAuthorityName="Lamarck" baseAuthorityYear="1803" box="[324,771,1174,1200]" class="Gastropoda" family="Muricidae" genus="Pterochelus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="contabulatus">
|
||
<emphasis id="B953EADEFFF1E411FEFCFB40FDC3FB03" box="[324,578,1174,1200]" italics="true" pageId="23" pageNumber="516">Pterochelus contabulatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFF1E411FDE8FB40FD7AFB03" author="LAMARCK J. - B." box="[592,763,1174,1200]" pageId="23" pageNumber="516" pagination="217 - 227" refId="ref45566" refString="LAMARCK J. - B. 1803. - Memoires sur les fossiles des environs de Paris (suite 1). Annales du Museum d'Histoire naturelle 2 (9): 217 - 227." type="journal article" year="1803">Lamarck, 1803</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The type material of
|
||
<taxonomicName id="4C274D4FFFF1E411FED1FB60FD81FB63" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[361,512,1206,1232]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="longicanalis" status="n. sp.">
|
||
<emphasis id="B953EADEFFF1E411FED1FB60FD81FB63" box="[361,512,1206,1232]" italics="true" pageId="23" pageNumber="516">T. longicanalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF1E411FDBFFB60FDC2FB63" box="[519,579,1206,1232]" pageId="23" pageNumber="516" rank="species">n. sp.</taxonomicNameLabel>
|
||
comes exclusively from the upper Lutetian of
|
||
<collectingRegion id="49E3F82EFFF1E411FE02FB03FDB2FB5C" box="[442,563,1237,1263]" country="France" name="Haute-Normandie" pageId="23" pageNumber="516">Normandy</collectingRegion>
|
||
. It is very well preserved and its delicate and winged varices are not eroded. Thus, its fusoid shape is not due to a post-mortem alteration. The species differs from
|
||
<taxonomicName id="4C274D4FFFF1E411FE3CFAE0FD90FAFC" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[388,529,1333,1359]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF1E411FE3CFAE0FD90FAFC" box="[388,529,1333,1359]" italics="true" pageId="23" pageNumber="516">T. tripteroides</emphasis>
|
||
</taxonomicName>
|
||
by having more staged and more quadrate whorls, more adaxially oriented P1 spines, a straighter siphonal canal, a less rounded aperture, and by having weaker internal denticles and spiral cords.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |