<taxonomicNameid="4C274D4FFFF3E413FC6CFAC3FB47FA9C"ID-CoL="CN5M3"authorityName="Merle & Pacaud & Ledon & Goret"authorityYear="2024"box="[980,1222,1301,1327]"class="Gastropoda"family="Muricidae"genus="Timbellus"kingdom="Animalia"order="Neogastropoda"pageId="21"pageNumber="514"phylum="Mollusca"rank="species"species="longicanalis"status="sp. nov.">
<figureCitationid="131C2A49FFF3E413FBBBFAE2FBD0FAFD"box="[1027,1105,1332,1358]"captionStart="FIG"captionStartId="22.[131,142,1581,1598]"captionTargetBox="[132,1455,215,1538]"captionTargetId="figure-331@22.[698,1267,781,1349]"captionTargetPageId="22"captionText="FIG. 11. — Spiral cords and internal denticles of the outer lip in Timbellus longicanalis n. sp. (A-D) and type material of Murex fusoides (Deshayes, 1865) (E-I): A, B, holotype,MNHN.F.A24101 (Le Marchand coll.),Fresville, Manche,Normandy, France,middle Eocene (upper Lutetian);C, D, paratype,MNHN.F.A24100, Fresville;E, type material of M. fusoides in the Deshayes’s collection corresponding to two eroded T. tripteroides; F, G, lectotype designated herein,UCBL-EM 33384, Valmondois, Val d’Oise, Paris Basin, France, middle Eocene (Bartonian); H, I, paralectotype, UCBL-EM 33383, Sainte-Aulde (Caumont), Seine-et-Marne, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E-I, E. Robert (UCBL)."figureDoi="http://doi.org/10.5281/zenodo.13985784"httpUri="https://zenodo.org/record/13985784/files/figure.png"pageId="21"pageNumber="514">Figs 11</figureCitation>
<treatmentCitationid="0A8610DDFFF3E413FC4AFA16FB4EFA6B"author="MERLE D. & GARRIGUES B. & POINTIER J. - P."box="[1010,1231,1472,1496]"page="446"pageId="21"pageNumber="514"year="2011">
<bibRefCitationid="EFB64B3DFFF3E413FC4AFA16FB4EFA6B"author="MERLE D. & GARRIGUES B. & POINTIER J. - P."box="[1010,1231,1472,1496]"pageId="21"pageNumber="514"refId="ref47234"refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p."type="book"year="2011">
<bibRefCitationid="EFB64B3DFFF3E413FCE7FA0CFC78FA41"author="DESHAYES G. - P."box="[863,1017,1498,1522]"pageId="21"pageNumber="514"pagination="101 - 107"refId="ref44001"refString="DESHAYES G. - P. 1865. - Description des Animaux sans vertebres decouverts dans le bassin de Paris pour servir de supplement a la description des coquilles fossiles des environs de Paris, comprenant une revue generale de toutes les especes actuellement connues. Tome 3. Bailliere, Paris, Livraisons 45 - 46, p. 201 - 288; Atlas 2, pl. 86 - 93 [2 January]; Livraisons 47 - 48, p. 289 - 424, Atlas 2, pl. 94 - 100 [10 June]; Livraisons 49 - 50, p. 425 - 668; Atlas 2, pl. 101 - 107 [15 December]."type="journal article"year="1865">Deshayes, 1865</bibRefCitation>
<locationid="8EF86017FFF3E413FCDBF9FCFBACF9F1"LSID="urn:lsid:plazi:treatment:038E87DAFFF3E411FC6CFAC3FD16FA1C:8EF86017FFF3E413FCDBF9FCFBACF9F1"box="[867,1069,1578,1602]"country="France"name="Calcaire de Fresville"pageId="21"pageNumber="514"stateProvince="Haute-Normandie">Calcaire de Fresville</location>
<collectionCodeid="ED36AE09FFF3E413FC96F993FCFCF9EF"box="[814,893,1605,1628]"country="France"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988"lsid="urn:lsid:biocol.org:col:34988"name="Museum National d'Histoire Naturelle"pageId="21"pageNumber="514">MNHN</collectionCode>
<figureCitationid="131C2A49FFF3E413FAB2F992FAF5F9EF"box="[1290,1396,1604,1629]"captionStart="FIG"captionStartId="22.[131,142,1581,1598]"captionTargetBox="[132,1455,215,1538]"captionTargetId="figure-331@22.[698,1267,781,1349]"captionTargetPageId="22"captionText="FIG. 11. — Spiral cords and internal denticles of the outer lip in Timbellus longicanalis n. sp. (A-D) and type material of Murex fusoides (Deshayes, 1865) (E-I): A, B, holotype,MNHN.F.A24101 (Le Marchand coll.),Fresville, Manche,Normandy, France,middle Eocene (upper Lutetian);C, D, paratype,MNHN.F.A24100, Fresville;E, type material of M. fusoides in the Deshayes’s collection corresponding to two eroded T. tripteroides; F, G, lectotype designated herein,UCBL-EM 33384, Valmondois, Val d’Oise, Paris Basin, France, middle Eocene (Bartonian); H, I, paralectotype, UCBL-EM 33383, Sainte-Aulde (Caumont), Seine-et-Marne, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E-I, E. Robert (UCBL)."figureDoi="http://doi.org/10.5281/zenodo.13985784"httpUri="https://zenodo.org/record/13985784/files/figure.png"pageId="21"pageNumber="514">Figs 11A, B</figureCitation>
;
<figureCitationid="131C2A49FFF3E413FAC1F992FA25F9EE"box="[1401,1444,1604,1629]"captionStart="FIG"captionStartId="45.[132,143,1622,1639]"captionTargetBox="[132,1455,215,1562]"captionTargetId="figure-370@45.[700,1259,799,1371]"captionTargetPageId="45"captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm."figureDoi="http://doi.org/10.5281/zenodo.13985825"httpUri="https://zenodo.org/record/13985825/files/figure.png"pageId="21"pageNumber="514">26E</figureCitation>
<collectionCodeid="ED36AE09FFF3E413FB44F9B6FACCF9C4"box="[1276,1357,1632,1655]"country="France"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988"lsid="urn:lsid:biocol.org:col:34988"name="Museum National d'Histoire Naturelle"pageId="21"pageNumber="514">MNHN</collectionCode>
<figureCitationid="131C2A49FFF3E413FC46F9ACFBF8F922"box="[1022,1145,1658,1682]"captionStart="FIG"captionStartId="22.[131,142,1581,1598]"captionTargetBox="[132,1455,215,1538]"captionTargetId="figure-331@22.[698,1267,781,1349]"captionTargetPageId="22"captionText="FIG. 11. — Spiral cords and internal denticles of the outer lip in Timbellus longicanalis n. sp. (A-D) and type material of Murex fusoides (Deshayes, 1865) (E-I): A, B, holotype,MNHN.F.A24101 (Le Marchand coll.),Fresville, Manche,Normandy, France,middle Eocene (upper Lutetian);C, D, paratype,MNHN.F.A24100, Fresville;E, type material of M. fusoides in the Deshayes’s collection corresponding to two eroded T. tripteroides; F, G, lectotype designated herein,UCBL-EM 33384, Valmondois, Val d’Oise, Paris Basin, France, middle Eocene (Bartonian); H, I, paralectotype, UCBL-EM 33383, Sainte-Aulde (Caumont), Seine-et-Marne, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E-I, E. Robert (UCBL)."figureDoi="http://doi.org/10.5281/zenodo.13985784"httpUri="https://zenodo.org/record/13985784/files/figure.png"pageId="21"pageNumber="514">Figs 11C, D</figureCitation>
;
<figureCitationid="131C2A49FFF3E413FB3BF9ACFB2FF921"box="[1155,1198,1658,1682]"captionStart="FIG"captionStartId="45.[132,143,1622,1639]"captionTargetBox="[132,1455,215,1562]"captionTargetId="figure-370@45.[700,1259,799,1371]"captionTargetPageId="45"captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm."figureDoi="http://doi.org/10.5281/zenodo.13985825"httpUri="https://zenodo.org/record/13985825/files/figure.png"pageId="21"pageNumber="514">26F</figureCitation>
. — Calcaire de Fresville, middle Eocene (uppermost Lutetian), see
<bibRefCitationid="EFB64B3DFFF3E413FC0EF8E2FBF7F8FF"author="BIGNOT G. & HOMMERIL P. & LARSONNEUR C."box="[950,1142,1844,1868]"pageId="21"pageNumber="514"pagination="405 - 416"refId="ref42941"refString="BIGNOT G., HOMMERIL P. & LARSONNEUR C. 1968. - Le Lutetien au large du Cotentin, in Colloque sur l'Eocene (Paris, mai 1968). Memoires BRGM 58: 405 - 416."type="journal article"year="1968">
<bibRefCitationid="EFB64B3DFFF3E413FC96F852FC3DF82F"author="APGN & ASSOCIATION PATRIMOINE GEOLOGIQUE DE NORMANDIE"box="[814,956,1924,1948]"pageId="21"pageNumber="514"refId="ref42597"refString="APGN (ASSOCIATION PATRIMOINE GEOLOGIQUE DE NORMANDIE) 2024. - Faluns cretaces et tertiaires de Fresville, BNO 0182. Available at: http: // www. apgn. fr / apgn / carte _ site / fic _ s 50 _ 75. pdf (last access on 03 / 2024)."type="book"year="2024">APGN (2024)</bibRefCitation>
<locationid="8EF86017FFF3E413FBECF86FFA95F862"LSID="urn:lsid:plazi:treatment:038E87DAFFF3E411FC6CFAC3FD16FA1C:8EF86017FFF3E413FBECF86FFA95F862"box="[1108,1300,1977,2001]"name="Calcaire de Fresville"pageId="21"pageNumber="514"stateProvince="Haute-Normandie">
. 11. — Spiral cords and internal denticles of the outer lip in
<taxonomicNameid="4C274D4FFFF0E410FD37F9FBFCC4F98D"authorityName="Merle & Pacaud & Ledon & Goret"authorityYear="2024"box="[655,837,1581,1598]"class="Gastropoda"family="Muricidae"genus="Timbellus"kingdom="Animalia"order="Neogastropoda"pageId="22"pageNumber="515"phylum="Mollusca"rank="species"species="longicanalis"status="sp. nov.">
<bibRefCitationid="EFB64B3DFFF0E410FB59F9FBFAE6F98D"author="DESHAYES G. - P."box="[1249,1383,1581,1598]"pageId="22"pageNumber="515"pagination="101 - 107"refId="ref44001"refString="DESHAYES G. - P. 1865. - Description des Animaux sans vertebres decouverts dans le bassin de Paris pour servir de supplement a la description des coquilles fossiles des environs de Paris, comprenant une revue generale de toutes les especes actuellement connues. Tome 3. Bailliere, Paris, Livraisons 45 - 46, p. 201 - 288; Atlas 2, pl. 86 - 93 [2 January]; Livraisons 47 - 48, p. 289 - 424, Atlas 2, pl. 94 - 100 [10 June]; Livraisons 49 - 50, p. 425 - 668; Atlas 2, pl. 101 - 107 [15 December]."type="journal article"year="1865">Deshayes, 1865</bibRefCitation>
, paralectotype, UCBL-EM 33383, Sainte-Aulde (Caumont), Seine-et-Marne, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E-I, E. Robert (UCBL).
in width, fusiform in profile, composed of six whorls. Moderately high spire. Last whorl up to 77% of total length. Apical angle 41° excluding spine and 61° including spines. Spiral sculpture with moderately marked primary cords on varices. First and second whorl eroded. Third whorl: appearance of weak s1 and P2 on intervarices. Fourth whorl: appearance of weak P1 cord. Fifth whorl: no change. Sixth whorl: P1-P4 marked, P5-P6 weak, s1, s2, s6, P6 not atrophied, s1-s2, s6 weak; ADP, MP and ABP weak. Axial sculpture eroded on two early whorls. From third last whorl: three varices and one intervarix intercalated. Higher relief of intervarices on P2. Varices lamellose, slightly winged. Appearance of P1 spine on third whorl, no other spines adapically. On last whorl P1 spine short, straight, slightly bent adaxially. On last whorl growth lamellae erect giving scabrous aspect to shell surface. Ovate aperture up to 29% of diameter and up to 78% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, slightly erect anteriorly, forming very slightly curved inductura at base. Parietal lip adherent. Outer lip with simple, weak denticles including ID, D1 to D6. Siphonal canal narrow, open, straight, up to 59% of apertural length. Pseudoumbilicus narrow.
<bibRefCitationid="EFB64B3DFFF1E411FF34FE2EFEB8FDA0"author="DESHAYES G. - P."box="[140,313,504,531]"pageId="23"pageNumber="516"pagination="101 - 107"refId="ref44001"refString="DESHAYES G. - P. 1865. - Description des Animaux sans vertebres decouverts dans le bassin de Paris pour servir de supplement a la description des coquilles fossiles des environs de Paris, comprenant une revue generale de toutes les especes actuellement connues. Tome 3. Bailliere, Paris, Livraisons 45 - 46, p. 201 - 288; Atlas 2, pl. 86 - 93 [2 January]; Livraisons 47 - 48, p. 289 - 424, Atlas 2, pl. 94 - 100 [10 June]; Livraisons 49 - 50, p. 425 - 668; Atlas 2, pl. 101 - 107 [15 December]."type="journal article"year="1865">Deshayes, 1865</bibRefCitation>
)
</taxonomicName>
by
<bibRefCitationid="EFB64B3DFFF1E411FED2FE2EFDADFDA1"author="MERLE D. & GARRIGUES B. & POINTIER J. - P."box="[362,556,504,530]"pageId="23"pageNumber="516"refId="ref47234"refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p."type="book"year="2011">
<taxonomicNameid="4C274D4FFFF1E411FED2FE2EFE28FDA1"authorityName="Merle & Pacaud & Ledon & Goret"authorityYear="2024"box="[362,425,504,530]"class="Gastropoda"family="Muricidae"kingdom="Animalia"order="Neogastropoda"pageId="23"pageNumber="543"phylum="Mollusca"rank="subFamily"status="subfam. nov."subFamily="Merle">Merle</taxonomicName>
housed in the collection of the University Lyon (no. UCBL-EM 33383-84;
<figureCitationid="131C2A49FFF1E411FF3AFD41FF60FD01"box="[130,225,663,690]"captionStart="FIG"captionStartId="22.[131,142,1581,1598]"captionTargetBox="[132,1455,215,1538]"captionTargetId="figure-331@22.[698,1267,781,1349]"captionTargetPageId="22"captionText="FIG. 11. — Spiral cords and internal denticles of the outer lip in Timbellus longicanalis n. sp. (A-D) and type material of Murex fusoides (Deshayes, 1865) (E-I): A, B, holotype,MNHN.F.A24101 (Le Marchand coll.),Fresville, Manche,Normandy, France,middle Eocene (upper Lutetian);C, D, paratype,MNHN.F.A24100, Fresville;E, type material of M. fusoides in the Deshayes’s collection corresponding to two eroded T. tripteroides; F, G, lectotype designated herein,UCBL-EM 33384, Valmondois, Val d’Oise, Paris Basin, France, middle Eocene (Bartonian); H, I, paralectotype, UCBL-EM 33383, Sainte-Aulde (Caumont), Seine-et-Marne, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E-I, E. Robert (UCBL)."figureDoi="http://doi.org/10.5281/zenodo.13985784"httpUri="https://zenodo.org/record/13985784/files/figure.png"pageId="23"pageNumber="516">Fig. 11E</figureCitation>
). The largest is a strongly eroded specimen with most of its axial sculpture, particularly its winged varices lost. This loss of sculpture gives the shell an artificially fusiform shape. The second specimen is smaller and also strongly eroded, but P1 spine is rather turned adapically as in some
<bibRefCitationid="EFB64B3DFFF1E411FEBDFCE1FE2CFCE2"author="LAMARCK J. - B."box="[261,429,823,849]"pageId="23"pageNumber="516"refId="ref45696"refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p."type="book"year="1822">Lamarck, 1822</bibRefCitation>
. We designate herein this smaller specimen (UCBL-EM 33384;
<figureCitationid="131C2A49FFF1E411FEE7FCA1FE58FC22"box="[351,473,887,913]"captionStart="FIG"captionStartId="22.[131,142,1581,1598]"captionTargetBox="[132,1455,215,1538]"captionTargetId="figure-331@22.[698,1267,781,1349]"captionTargetPageId="22"captionText="FIG. 11. — Spiral cords and internal denticles of the outer lip in Timbellus longicanalis n. sp. (A-D) and type material of Murex fusoides (Deshayes, 1865) (E-I): A, B, holotype,MNHN.F.A24101 (Le Marchand coll.),Fresville, Manche,Normandy, France,middle Eocene (upper Lutetian);C, D, paratype,MNHN.F.A24100, Fresville;E, type material of M. fusoides in the Deshayes’s collection corresponding to two eroded T. tripteroides; F, G, lectotype designated herein,UCBL-EM 33384, Valmondois, Val d’Oise, Paris Basin, France, middle Eocene (Bartonian); H, I, paralectotype, UCBL-EM 33383, Sainte-Aulde (Caumont), Seine-et-Marne, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E-I, E. Robert (UCBL)."figureDoi="http://doi.org/10.5281/zenodo.13985784"httpUri="https://zenodo.org/record/13985784/files/figure.png"pageId="23"pageNumber="516">Fig. 11F, G</figureCitation>
are present in several collections housed in the MNHN (e.g., Faullummel, Pons and Schtrock colls). Examination of these specimens shows them to represent eroded
<bibRefCitationid="EFB64B3DFFF1E411FDE8FB40FD7AFB03"author="LAMARCK J. - B."box="[592,763,1174,1200]"pageId="23"pageNumber="516"pagination="217 - 227"refId="ref45566"refString="LAMARCK J. - B. 1803. - Memoires sur les fossiles des environs de Paris (suite 1). Annales du Museum d'Histoire naturelle 2 (9): 217 - 227."type="journal article"year="1803">Lamarck, 1803</bibRefCitation>
)
</taxonomicName>
. The type material of
<taxonomicNameid="4C274D4FFFF1E411FED1FB60FD81FB63"authorityName="Merle & Pacaud & Ledon & Goret"authorityYear="2024"box="[361,512,1206,1232]"class="Gastropoda"family="Muricidae"genus="Timbellus"kingdom="Animalia"order="Neogastropoda"pageId="23"pageNumber="516"phylum="Mollusca"rank="species"species="longicanalis"status="n. sp.">
. It is very well preserved and its delicate and winged varices are not eroded. Thus, its fusoid shape is not due to a post-mortem alteration. The species differs from
by having more staged and more quadrate whorls, more adaxially oriented P1 spines, a straighter siphonal canal, a less rounded aperture, and by having weaker internal denticles and spiral cords.
