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<document id="98EBDB6FC17DC5EAF7849A9639BBBE77" ID-DOI="10.11646/zootaxa.5375.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10170174" ID-ZooBank="30782E67-2E2C-4B63-9809-85C3945187AE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1700559471735" checkinUser="plazi" docAuthor="Inoue, Shota &amp; Nomura, Shûhei" docDate="2023" docId="03861C35FFD9FFF6FF698728FCC6F948" docLanguage="en" docName="zootaxa.5375.1.4.pdf" docOrigin="Zootaxa 5375 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.1.4/52314" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrodes Raffray 1908" docType="treatment" docVersion="4" lastPageNumber="84" masterDocId="FFBF644DFFD8FFF7FFFE846AFFACFFFB" masterDocTitle="New species and record of Tyrodes Raffray (Coleoptera: Staphylinidae: Pselaphinae) from Japan" masterLastPageNumber="92" masterPageNumber="83" pageNumber="84" updateTime="1700835184917" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="DE199D3BE55F8469F807935424BA42ED">New species and record of Tyrodes Raffray (Coleoptera: Staphylinidae: Pselaphinae) from Japan</mods:title>
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<mods:namePart id="C7D0F2FF9A8D98AD74FC7860596380E3">Inoue, Shota</mods:namePart>
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<mods:affiliation id="57E52E5DFAC49AA8659D00975151CBA9">University Education Center, Tokyo Metropolitan University, 1 - 1 Minami-Osawa, Hachioji, Tokyo, 192 - 0397 Japan shota 666. shigure @ gmail. com; https: // orcid. org / 0000 - 0002 - 7005 - 6531 &amp; The Kyushu University Museum, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<mods:namePart id="D388FDA0C3031F7FC00D03692F906603">Nomura, Shûhei</mods:namePart>
<mods:nameIdentifier id="C7FC7C0CAB071E1C6FEDCE7D43CA425A" type="ORCID">0000-0003-1086-5078</mods:nameIdentifier>
<mods:affiliation id="57EA91575AAC7777DCAE5AC26FA46DA4">Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1, Amakubo, Tsukuba-shi, Ibaraki, 305 - 0005 Japan nomura @ kahaku. go. jp; https: // orcid. org / 0000 - 0003 - 1086 - 5078</mods:affiliation>
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<subSubSection id="C335FEA8FFD9FFF6FF698728FCB9FC7A" pageId="1" pageNumber="84" type="nomenclature">
<paragraph id="8B90AD23FFD9FFF6FF698728FE53FCA7" blockId="1.[151,789,834,897]" box="[151,511,834,861]" pageId="1" pageNumber="84">
<heading id="D0D81A4FFFD9FFF6FF698728FE53FCA7" bold="true" box="[151,511,834,861]" fontSize="11" level="1" pageId="1" pageNumber="84" reason="1">
<emphasis id="B95B7131FFD9FFF6FF698728FE53FCA7" bold="true" box="[151,511,834,861]" pageId="1" pageNumber="84">
Genus
<taxonomicName id="4C2FD6A0FFD9FFF6FF108728FE53FCA7" ID-CoL="8XQM6" authority="Raffray, 1908" authorityName="Raffray" authorityYear="1908" box="[238,511,834,861]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FF108728FEE1FCA6" bold="true" box="[238,333,834,861]" italics="true" pageId="1" pageNumber="84">Tyrodes</emphasis>
<bibRefCitation id="EFBED0D2FFD9FFF6FEAB8728FE53FCA7" author="Raffray, A." box="[341,511,834,860]" pageId="1" pageNumber="84" pagination="1 - 487" refId="ref4869" refString="Raffray, A. (1908) Coleoptera. Fam. Pselaphidae. In: Wytsman, P. (Ed.), Genera insectorum. Fasc. 64. V. Verteneuil &amp; L. Desmet, Bruxelles, pp. 1 - 487, 9 pls." type="journal article" year="1908">Raffray, 1908</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B90AD23FFD9FFF6FF69870DFCB9FC7A" blockId="1.[151,789,834,897]" box="[151,789,871,897]" pageId="1" pageNumber="84">[Japanese common name: Nisekoke-arizukamushi Zoku]</paragraph>
</subSubSection>
<subSubSection id="C335FEA8FFD9FFF6FF6987C1FAE5FC19" pageId="1" pageNumber="84" type="reference_group">
<paragraph id="8B90AD23FFD9FFF6FF6987C1FAE5FC19" blockId="1.[151,1416,939,994]" pageId="1" pageNumber="84">
<treatmentCitationGroup id="AB3F8A0DFFD9FFF6FF6987C1FAE5FC19" pageId="1" pageNumber="84">
<treatmentCitation id="0A8E8B32FFD9FFF6FF6987C1FE01FC38" author="Raffray, A." box="[151,429,939,963]" page="379" pageId="1" pageNumber="84" year="1908">
<taxonomicName id="4C2FD6A0FFD9FFF6FF6987C1FE01FC38" ID-CoL="8XQM6" authority="Raffray, 1908: 379" authorityName="Raffray" authorityPageNumber="379" authorityYear="1908" box="[151,429,939,963]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FF6987C1FF4AFC38" box="[151,230,939,963]" italics="true" pageId="1" pageNumber="84">Tyrodes</emphasis>
<bibRefCitation id="EFBED0D2FFD9FFF6FF1287C1FE01FC38" author="Raffray, A." box="[236,429,939,963]" pageId="1" pageNumber="84" pagination="1 - 487" refId="ref4869" refString="Raffray, A. (1908) Coleoptera. Fam. Pselaphidae. In: Wytsman, P. (Ed.), Genera insectorum. Fasc. 64. V. Verteneuil &amp; L. Desmet, Bruxelles, pp. 1 - 487, 9 pls." type="journal article" year="1908">Raffray, 1908: 379</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54941381FFD9FFF6FE4987C6FE45FC3F" box="[439,489,940,964]" pageId="1" pageNumber="84">Type</typeStatus>
species:
<taxonomicName id="4C2FD6A0FFD9FFF6FDB887C6FC03FC38" ID-CoL="59ZTM" authority="L. W. Schaufass, 1887" authorityName="L. W. Schaufass" authorityYear="1887" box="[582,943,939,963]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="histrio">
<emphasis id="B95B7131FFD9FFF6FDB887C6FD6AFC38" box="[582,710,939,963]" italics="true" pageId="1" pageNumber="84">Tyrus histrio</emphasis>
L. W. Schaufass, 1887
</taxonomicName>
(subsequent designation by
<treatmentCitation id="0A8E8B32FFD9FFF6FB3187C6FAD0FC38" author="Lucas, R." box="[1231,1404,939,963]" page="663" pageId="1" pageNumber="84" year="1920">
<bibRefCitation id="EFBED0D2FFD9FFF6FB3187C6FAD0FC38" author="Lucas, R." box="[1231,1404,939,963]" pageId="1" pageNumber="84" pagination="1 - 696" refId="ref4522" refString="Lucas, R. (1920) Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Archiv f ¸ r Naturgeschichte, Series A, 84, i - xxxi + 1 - 696." type="journal article" year="1920">Lucas 1920: 663</bibRefCitation>
</treatmentCitation>
).
<taxonomicName id="4C2FD6A0FFD9FFF6FF6987A0FE10FC19" ID-CoL="8XQM7" authority="Jeannel, 1960: 455" authorityName="Jeannel" authorityPageNumber="455" authorityYear="1960" box="[151,444,970,994]" class="Insecta" family="Staphylinidae" genus="Tyrodites" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FF6987A0FF58FC19" box="[151,244,970,994]" italics="true" pageId="1" pageNumber="84">Tyrodites</emphasis>
Jeannel, 1960: 455
</taxonomicName>
.
<typeStatus id="54941381FFD9FFF6FE3B87A0FE5BFC19" box="[453,503,970,994]" pageId="1" pageNumber="84">Type</typeStatus>
species:
<taxonomicName id="4C2FD6A0FFD9FFF6FDAA87A0FC19FC19" ID-CoL="59Z7C" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" box="[596,949,970,994]" class="Insecta" family="Staphylinidae" genus="Tyrodites" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B95B7131FFD9FFF6FDAA87A0FC8FFC19" box="[596,803,970,994]" italics="true" pageId="1" pageNumber="84">Tyrodites championi</emphasis>
Jeannel, 1960
</taxonomicName>
(synonymized by
<treatmentCitation id="0A8E8B32FFD9FFF6FB8E87A0FA91FC19" author="Besuchet, C." box="[1136,1341,970,994]" page="316" pageId="1" pageNumber="84" year="1970">
<bibRefCitation id="EFBED0D2FFD9FFF6FB8E87A0FA91FC19" author="Besuchet, C." box="[1136,1341,970,994]" pageId="1" pageNumber="84" pagination="313 - 318" refId="ref4173" refString="Besuchet, C. (1970) Pselaphides recoltes au Nepal par le Prof. H. Janetschek (Coleoptera). Berichte des Naturwissenschafilichmedizinischen Vereins in Innsbruck, 58, 313 - 318." type="journal article" year="1970">Besuchet 1970: 316</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C335FEA8FFD9FFF6FF69807AFAADFB6D" pageId="1" pageNumber="84" type="diagnosis">
<paragraph id="8B90AD23FFD9FFF6FF69807AFAADFB6D" blockId="1.[151,1437,1040,1715]" pageId="1" pageNumber="84">
<emphasis id="B95B7131FFD9FFF6FF69807AFEBEFBD1" bold="true" box="[151,274,1040,1066]" pageId="1" pageNumber="84">Diagnosis.</emphasis>
Head with small, nude frontal median fovea and one pair of larger, setose vertexal foveae; maxillary palpi large, each with palpomeres 2–4 each pedunculate basally, widened apically; palpomeres 4 each with sensory appendage at apex; pronotum with antebasal sulcus connecting median and lateral foveae; abdomen with tergite IV longer than V; aedeagus stout, with ovoid median lobe (
<bibRefCitation id="EFBED0D2FFD9FFF6FCF98016FC0CFB6D" author="Raffray, A." box="[775,928,1148,1174]" pageId="1" pageNumber="84" pagination="1 - 487" refId="ref4869" refString="Raffray, A. (1908) Coleoptera. Fam. Pselaphidae. In: Wytsman, P. (Ed.), Genera insectorum. Fasc. 64. V. Verteneuil &amp; L. Desmet, Bruxelles, pp. 1 - 487, 9 pls." type="journal article" year="1908">Raffray 1908</bibRefCitation>
;
<bibRefCitation id="EFBED0D2FFD9FFF6FC528016FBEEFB6C" author="Jeannel, R." box="[940,1090,1148,1175]" pageId="1" pageNumber="84" pagination="5 - 32" refId="ref4418" refString="Jeannel, R. (1957) Sur quelques Pselaphides du Tonkin recueillis par le Pere A. de Cooman. Revue Francaise d'Entomologie, 24, 5 - 32." type="journal article" year="1957">Jeannel 1957</bibRefCitation>
;
<bibRefCitation id="EFBED0D2FFD9FFF6FBB28016FB5AFB6D" author="Yin, Z. - W. &amp; Li, L. - Z." box="[1100,1270,1148,1174]" pageId="1" pageNumber="84" pagination="101 - 105" refId="ref5050" refString="Yin, Z. - W. &amp; Li, L. - Z. (2013) First record of Tyrodes Raffray (Coleoptera, Staphylinidae, Pselaphinae) in China, with description of T. jenisi sp. n. from Yunnan Province. Zookeys, 301, 101 - 105. https: // doi. org / 10.3897 / zookeys. 301.4911" type="journal article" year="2013">Yin &amp; Li 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C335FEA8FFD9FFF6FF3980CAFCC6F948" pageId="1" pageNumber="84" type="discussion">
<paragraph id="8B90AD23FFD9FFF6FF3980CAFD4DFA21" blockId="1.[151,1437,1040,1715]" pageId="1" pageNumber="84">
<emphasis id="B95B7131FFD9FFF6FF3980CAFEE4FB41" bold="true" box="[199,328,1184,1210]" pageId="1" pageNumber="84">Comments</emphasis>
. The genus
<taxonomicName id="4C2FD6A0FFD9FFF6FE3580CAFD8EFB41" authorityName="Raffray" authorityYear="1908" box="[459,546,1184,1210]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FE3580CAFD8EFB41" box="[459,546,1184,1210]" italics="true" pageId="1" pageNumber="84">Tyrodes</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C2FD6A0FFD9FFF6FD5280CBFCC2FB41" authority="Aube, 1833" authorityName="Aube" authorityYear="1833" box="[684,878,1184,1210]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FD5280CBFD46FB41" box="[684,746,1185,1210]" italics="true" pageId="1" pageNumber="84">Tyrus</emphasis>
Aubé,
<quantity id="4CD700C6FFD9FFF6FCC880CAFCC2FB41" box="[822,878,1184,1210]" metricMagnitude="1" metricUnit="m" metricValue="4.65582" pageId="1" pageNumber="84" unit="in" value="1833.0">1833</quantity>
</taxonomicName>
in their general appearances, but it is distinguished from the latter by the following characteristics: frontal median fovea small, nude (distinct, larger, setose in
<taxonomicName id="4C2FD6A0FFD9FFF6FAB380AFFA27FB25" authorityName="Aube" authorityYear="1833" box="[1357,1419,1221,1246]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FAB380AFFA27FB25" box="[1357,1419,1221,1246]" italics="true" pageId="1" pageNumber="84">Tyrus</emphasis>
</taxonomicName>
); abdominal tergite IV longer than V (IV and V subequal in
<taxonomicName id="4C2FD6A0FFD9FFF6FCE18083FCF1FAF9" authorityName="Aube" authorityYear="1833" box="[799,861,1257,1282]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FCE18083FCF1FAF9" box="[799,861,1257,1282]" italics="true" pageId="1" pageNumber="84">Tyrus</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFBED0D2FFD9FFF6FC8B8082FBB2FAF9" author="Yin, Z. - W. &amp; Li, L. - Z." box="[885,1054,1256,1282]" pageId="1" pageNumber="84" pagination="101 - 105" refId="ref5050" refString="Yin, Z. - W. &amp; Li, L. - Z. (2013) First record of Tyrodes Raffray (Coleoptera, Staphylinidae, Pselaphinae) in China, with description of T. jenisi sp. n. from Yunnan Province. Zookeys, 301, 101 - 105. https: // doi. org / 10.3897 / zookeys. 301.4911" type="journal article" year="2013">Yin &amp; Li 2013</bibRefCitation>
). In addition, after an examination of previous studies and the specimens studied in this paper, the following characteristics may also be useful for separation of both genera: male antennomeres 1 each with triangular projection on the external side of the basal part (
<figureCitation id="1314B1A6FFD9FFF6FF2C813EFE82FA95" box="[210,302,1364,1390]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[177,1411,188,1912]" captionTargetId="figure-27@2.[170,1417,181,1920]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes segrex Kurbatov (A, C–F. Male. B. Female). A, B. Dorsal habitus. C. Antennomere 1. D. Meso- and metaventrite and ventral side of abdomen. E. Metaventral projection. F. Lateral side of abdomen. Scale bars: 1.0 mm in A, B; 0.5 mm in D, F; 0.2 mm in E; 0.1 mm in C." figureDoi="http://doi.org/10.5281/zenodo.10170178" httpUri="https://zenodo.org/record/10170178/files/figure.png" pageId="1" pageNumber="84">Figs 1C</figureCitation>
,
<figureCitation id="1314B1A6FFD9FFF6FEC5813EFEF1FA94" box="[315,349,1364,1391]" captionStart="FIGURE 5" captionStartId="6.[151,250,1710,1734]" captionTargetBox="[158,1429,198,1681]" captionTargetId="figure-27@6.[151,1436,181,1686]" captionTargetPageId="6" captionText="FIGURE 5. Morphology of Tyrodes amamianus sp. nov., male. A. Dorsal habitus. B. Ventral habitus. C. Antennomere 1. D. Metaventral projection. E. Lateral side of abdomen. F. Tergite VIII. G. Sternite VIII. H. Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="1" pageNumber="84">5C</figureCitation>
) (usually lacking the triangular projection in
<taxonomicName id="4C2FD6A0FFD9FFF6FCA1813FFC31FA95" authorityName="Aube" authorityYear="1833" box="[863,925,1365,1390]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FCA1813FFC31FA95" box="[863,925,1365,1390]" italics="true" pageId="1" pageNumber="84">Tyrus</emphasis>
</taxonomicName>
, but
<taxonomicName id="4C2FD6A0FFD9FFF6FC29813FFAC3FA95" authority="Yin &amp; Nomura, 2013" authorityName="Yin &amp; Nomura" authorityYear="2013" box="[983,1391,1364,1391]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="sichuanicus">
<emphasis id="B95B7131FFD9FFF6FC29813FFBD4FA95" box="[983,1144,1364,1390]" italics="true" pageId="1" pageNumber="84">T. sichuanicus</emphasis>
<bibRefCitation id="EFBED0D2FFD9FFF6FB81813EFAC3FA95" author="Yin, Z. - W. &amp; Nomura, S. &amp; Li, L. - Z." box="[1151,1391,1364,1391]" pageId="1" pageNumber="84" pagination="47 - 53" refId="ref5117" refString="Yin, Z. - W., Nomura, S. &amp; Li, L. - Z. (2013) Redescription of Tyrus Aube and T. sinensis Raffray (Coleoptera: Staphylinidae: Pselaphinae), and two new species from Sichuan, Southwest China. Zootaxa, 3637 (1), 47 - 53. https: // doi. org / 10.11646 / zootaxa. 3637.1.5" type="journal article" year="2013">Yin &amp; Nomura, 2013</bibRefCitation>
</taxonomicName>
has laterally expanded antennomeres 1), and tergite IV lacking median longitudinal carina (
<figureCitation id="1314B1A6FFD9FFF6FB968112FB13FA69" box="[1128,1215,1400,1426]" captionStart="FIGURE 3" captionStartId="4.[151,250,1394,1418]" captionTargetBox="[151,1435,417,1370]" captionTargetId="figure-147@4.[151,1435,417,1370]" captionTargetPageId="4" captionText="FIGURE 3. Morphological details of Tyrodes segrex Kurbatov (A–B. Male. C–D. Female) in SEM images. A. Tergite IV. B, C. Abdomen in lateral view. D. Tergite VIII—enlarged image of the dashed white line square in Fig. 2C." figureDoi="http://doi.org/10.5281/zenodo.10170189" httpUri="https://zenodo.org/record/10170189/files/figure.png" pageId="1" pageNumber="84">Fig. 3A</figureCitation>
) (usually present in
<taxonomicName id="4C2FD6A0FFD9FFF6FF6981F7FF79FA4D" authorityName="Aube" authorityYear="1833" box="[151,213,1437,1462]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FF6981F7FF79FA4D" box="[151,213,1437,1462]" italics="true" pageId="1" pageNumber="84">Tyrus</emphasis>
</taxonomicName>
, but lacking in
<taxonomicName id="4C2FD6A0FFD9FFF6FE7781F7FD42FA4C" authority="Bryant, 1915" authorityName="Bryant" authorityYear="1915" box="[393,750,1436,1463]" class="Insecta" family="Staphylinidae" genus="Tyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="myrmecophilus">
<emphasis id="B95B7131FFD9FFF6FE7781F7FDFDFA4D" box="[393,593,1436,1462]" italics="true" pageId="1" pageNumber="84">T. myrmecophilus</emphasis>
<bibRefCitation id="EFBED0D2FFD9FFF6FDA781F7FD42FA4C" author="Bryant, G. E." box="[601,750,1436,1463]" pageId="1" pageNumber="84" pagination="297 - 302" refId="ref4277" refString="Bryant, G. E. (1915) New species of Pselaphidae (group Tyrini). Entomologist's Monthly Magazine, 51, 297 - 302, pl. 21. https: // doi. org / 10.5962 / bhl. part. 7791" type="journal article" year="1915">Bryant, 1915</bibRefCitation>
</taxonomicName>
) (
<bibRefCitation id="EFBED0D2FFD9FFF6FCFB81F6FC0CFA4D" author="Raffray, A." box="[773,928,1436,1462]" pageId="1" pageNumber="84" pagination="1 - 487" refId="ref4869" refString="Raffray, A. (1908) Coleoptera. Fam. Pselaphidae. In: Wytsman, P. (Ed.), Genera insectorum. Fasc. 64. V. Verteneuil &amp; L. Desmet, Bruxelles, pp. 1 - 487, 9 pls." type="journal article" year="1908">Raffray 1908</bibRefCitation>
;
<bibRefCitation id="EFBED0D2FFD9FFF6FC5381F7FB90FA4C" author="Bryant, G. E." box="[941,1084,1436,1463]" pageId="1" pageNumber="84" pagination="297 - 302" refId="ref4277" refString="Bryant, G. E. (1915) New species of Pselaphidae (group Tyrini). Entomologist's Monthly Magazine, 51, 297 - 302, pl. 21. https: // doi. org / 10.5962 / bhl. part. 7791" type="journal article" year="1915">Bryant 1915</bibRefCitation>
;
<bibRefCitation id="EFBED0D2FFD9FFF6FBB781F6FB4CFA4C" author="Jeannel, R." box="[1097,1248,1436,1463]" pageId="1" pageNumber="84" pagination="5 - 32" refId="ref4418" refString="Jeannel, R. (1957) Sur quelques Pselaphides du Tonkin recueillis par le Pere A. de Cooman. Revue Francaise d'Entomologie, 24, 5 - 32." type="journal article" year="1957">Jeannel 1957</bibRefCitation>
;
<bibRefCitation id="EFBED0D2FFD9FFF6FB1081F6FA3BFA4D" author="Besuchet, C." box="[1262,1431,1436,1462]" pageId="1" pageNumber="84" pagination="313 - 318" refId="ref4173" refString="Besuchet, C. (1970) Pselaphides recoltes au Nepal par le Prof. H. Janetschek (Coleoptera). Berichte des Naturwissenschafilichmedizinischen Vereins in Innsbruck, 58, 313 - 318." type="journal article" year="1970">Besuchet 1970</bibRefCitation>
;
<bibRefCitation id="EFBED0D2FFD9FFF6FF6981AAFE92FA21" author="Yin, Z. - W. &amp; Li, L. - Z." box="[151,318,1472,1498]" pageId="1" pageNumber="84" pagination="101 - 105" refId="ref5050" refString="Yin, Z. - W. &amp; Li, L. - Z. (2013) First record of Tyrodes Raffray (Coleoptera, Staphylinidae, Pselaphinae) in China, with description of T. jenisi sp. n. from Yunnan Province. Zookeys, 301, 101 - 105. https: // doi. org / 10.3897 / zookeys. 301.4911" type="journal article" year="2013">Yin &amp; Li 2013</bibRefCitation>
; Yin
<emphasis id="B95B7131FFD9FFF6FE8681ABFE1DFA21" box="[376,433,1472,1498]" italics="true" pageId="1" pageNumber="84">et al.</emphasis>
2013;
<bibRefCitation id="EFBED0D2FFD9FFF6FE0081AAFD79FA21" author="Zhang, Y. - Q. &amp; Yin, Z. - W." box="[510,725,1472,1498]" pageId="1" pageNumber="84" pagination="596 - 599" refId="ref5190" refString="Zhang, Y. - Q. &amp; Yin, Z. - W. (2023) A new species of Tyrodes Raffray (Coleoptera: Staphylinidae: Pselaphinae) from China. Zootaxa, 5293 (3), 596 - 599. https: // doi. org / 10.11646 / zootaxa. 5293.3.11" type="journal article" year="2023">Zhang &amp; Yin 2023</bibRefCitation>
).
</paragraph>
<paragraph id="8B90AD23FFD9FFF6FF39818EFA26F9BD" blockId="1.[151,1437,1040,1715]" pageId="1" pageNumber="84">
The genus
<taxonomicName id="4C2FD6A0FFD9FFF6FEBF818FFD7EFA05" authority="Hlavac &amp; Nomura, 2003" authorityName="Hlavac &amp; Nomura" authorityYear="2003" box="[321,722,1508,1535]" class="Insecta" family="Staphylinidae" genus="Megatyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FEBF818FFE15FA05" box="[321,441,1509,1534]" italics="true" pageId="1" pageNumber="84">Megatyrus</emphasis>
<bibRefCitation id="EFBED0D2FFD9FFF6FE41818EFD7EFA05" author="Hlavac, P. &amp; Nomura, S." box="[447,722,1508,1535]" pageId="1" pageNumber="84" pagination="165 - 174" refId="ref4323" refString="Hlavac, P. &amp; Nomura, S. (2003) A taxonomic revision of Tyrini of the Oriental region III. Megatyrus (Coleoptera, Staphylinidae, Pselaphinae), a new genus of the Tyrina from China and Vietnam. Elytra, 31, 165 - 174." type="journal article" year="2003">Hlaváč &amp; Nomura, 2003</bibRefCitation>
</taxonomicName>
share the characteristic of antennomeres 1 with
<taxonomicName id="4C2FD6A0FFD9FFF6FB15818EFAEEFA05" authorityName="Raffray" authorityYear="1908" box="[1259,1346,1508,1534]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FB15818EFAEEFA05" box="[1259,1346,1508,1534]" italics="true" pageId="1" pageNumber="84">Tyrodes</emphasis>
</taxonomicName>
, but the members of
<taxonomicName id="4C2FD6A0FFD9FFF6FEDF8263FE35F9D9" authorityName="Hlavac &amp; Nomura" authorityYear="2003" box="[289,409,1545,1570]" class="Insecta" family="Staphylinidae" genus="Megatyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FEDF8263FE35F9D9" box="[289,409,1545,1570]" italics="true" pageId="1" pageNumber="84">Megatyrus</emphasis>
</taxonomicName>
are easily separated from
<taxonomicName id="4C2FD6A0FFD9FFF6FD468262FCA3F9D9" authorityName="Raffray" authorityYear="1908" box="[696,783,1544,1570]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B95B7131FFD9FFF6FD468262FCA3F9D9" box="[696,783,1544,1570]" italics="true" pageId="1" pageNumber="84">Tyrodes</emphasis>
</taxonomicName>
by the larger body, maxillary palpi with palpomeres 4 each lacking apical appendage, and tergite IV with well-defined median longitudinal carina (
<bibRefCitation id="EFBED0D2FFD9FFF6FB948246FAD2F9BD" author="Hlavac, P. &amp; Nomura, S." box="[1130,1406,1580,1607]" pageId="1" pageNumber="84" pagination="165 - 174" refId="ref4323" refString="Hlavac, P. &amp; Nomura, S. (2003) A taxonomic revision of Tyrini of the Oriental region III. Megatyrus (Coleoptera, Staphylinidae, Pselaphinae), a new genus of the Tyrina from China and Vietnam. Elytra, 31, 165 - 174." type="journal article" year="2003">Hlaváč &amp; Nomura 2003</bibRefCitation>
).
</paragraph>
<paragraph id="8B90AD23FFD9FFF6FF39823AFCC6F948" blockId="1.[151,1437,1040,1715]" pageId="1" pageNumber="84">
Among the Japanese
<taxonomicName id="4C2FD6A0FFD9FFF6FE45823AFDC3F991" box="[443,623,1616,1642]" pageId="1" pageNumber="84">
<emphasis id="B95B7131FFD9FFF6FE45823AFDBEF991" box="[443,530,1616,1642]" italics="true" pageId="1" pageNumber="84">Tyrodes</emphasis>
species
</taxonomicName>
, the male shares following characteristics: posterolateral clypeal region strongly concave with demarcate edge (
<figureCitation id="1314B1A6FFD9FFF6FDA0821EFCADF975" box="[606,769,1652,1678]" captionStart="FIGURE 2" captionStartId="3.[151,250,1766,1790]" captionTargetBox="[151,1435,777,1743]" captionTargetId="figure-283@3.[151,1435,777,1743]" captionTargetPageId="3" captionText="FIGURE 2. Head of Tyrodes segrex Kurbatov (A, C–D. Male. B. Female) in SEM images. A, B. Head in lateral view; C. Posterolateral clypeal region in lateral view; D. Posterolateral clypeal region in dorsal view. Abbreviations: ant1—antennomere 1; mp—maxillary palpus." figureDoi="http://doi.org/10.5281/zenodo.10170184" httpUri="https://zenodo.org/record/10170184/files/figure.png" pageId="1" pageNumber="84">Fig. 2A, C, D</figureCitation>
); metaventrite with small setose projection (
<figureCitation id="1314B1A6FFD9FFF6FB01821EFAF6F975" box="[1279,1370,1652,1678]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[177,1411,188,1912]" captionTargetId="figure-27@2.[170,1417,181,1920]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes segrex Kurbatov (A, C–F. Male. B. Female). A, B. Dorsal habitus. C. Antennomere 1. D. Meso- and metaventrite and ventral side of abdomen. E. Metaventral projection. F. Lateral side of abdomen. Scale bars: 1.0 mm in A, B; 0.5 mm in D, F; 0.2 mm in E; 0.1 mm in C." figureDoi="http://doi.org/10.5281/zenodo.10170178" httpUri="https://zenodo.org/record/10170178/files/figure.png" pageId="1" pageNumber="84">Figs 1E</figureCitation>
,
<figureCitation id="1314B1A6FFD9FFF6FA97821EFA22F974" box="[1385,1422,1652,1679]" captionStart="FIGURE 5" captionStartId="6.[151,250,1710,1734]" captionTargetBox="[158,1429,198,1681]" captionTargetId="figure-27@6.[151,1436,181,1686]" captionTargetPageId="6" captionText="FIGURE 5. Morphology of Tyrodes amamianus sp. nov., male. A. Dorsal habitus. B. Ventral habitus. C. Antennomere 1. D. Metaventral projection. E. Lateral side of abdomen. F. Tergite VIII. G. Sternite VIII. H. Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="1" pageNumber="84">5D</figureCitation>
); sternite IV–V strongly concave at lateral parts (
<figureCitation id="1314B1A6FFD9FFF6FD5482F2FCACF949" box="[682,768,1688,1714]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[177,1411,188,1912]" captionTargetId="figure-27@2.[170,1417,181,1920]" captionTargetPageId="2" captionText="FIGURE 1. Morphology of Tyrodes segrex Kurbatov (A, C–F. Male. B. Female). A, B. Dorsal habitus. C. Antennomere 1. D. Meso- and metaventrite and ventral side of abdomen. E. Metaventral projection. F. Lateral side of abdomen. Scale bars: 1.0 mm in A, B; 0.5 mm in D, F; 0.2 mm in E; 0.1 mm in C." figureDoi="http://doi.org/10.5281/zenodo.10170178" httpUri="https://zenodo.org/record/10170178/files/figure.png" pageId="1" pageNumber="84">Figs 1F</figureCitation>
,
<figureCitation id="1314B1A6FFD9FFF6FCF282F2FC83F949" box="[780,815,1688,1714]" captionStart="FIGURE 3" captionStartId="4.[151,250,1394,1418]" captionTargetBox="[151,1435,417,1370]" captionTargetId="figure-147@4.[151,1435,417,1370]" captionTargetPageId="4" captionText="FIGURE 3. Morphological details of Tyrodes segrex Kurbatov (A–B. Male. C–D. Female) in SEM images. A. Tergite IV. B, C. Abdomen in lateral view. D. Tergite VIII—enlarged image of the dashed white line square in Fig. 2C." figureDoi="http://doi.org/10.5281/zenodo.10170189" httpUri="https://zenodo.org/record/10170189/files/figure.png" pageId="1" pageNumber="84">3B</figureCitation>
,
<figureCitation id="1314B1A6FFD9FFF6FCC582F2FCF1F948" box="[827,861,1688,1715]" captionStart="FIGURE 5" captionStartId="6.[151,250,1710,1734]" captionTargetBox="[158,1429,198,1681]" captionTargetId="figure-27@6.[151,1436,181,1686]" captionTargetPageId="6" captionText="FIGURE 5. Morphology of Tyrodes amamianus sp. nov., male. A. Dorsal habitus. B. Ventral habitus. C. Antennomere 1. D. Metaventral projection. E. Lateral side of abdomen. F. Tergite VIII. G. Sternite VIII. H. Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="1" pageNumber="84">5E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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