500 lines
58 KiB
XML
500 lines
58 KiB
XML
<document id="0F130164282FF1746DFB1569E6DB3BE4" ID-DOI="10.3897/evolsyst.8.141847" ID-ZooBank="8E0DCC28-3448-4001-90B7-E675BC107381" ID-publisher-id="141847" URI-arpha="784085DF-996A-5FEC-89FE-F9FE7A70E5A1" URI-zoobank="http://zoobank.org/8E0DCC28-3448-4001-90B7-E675BC107381" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1732116009247" checkinUser="pensoft" docAuthor="Wermelinger-Moreira, Gabriel, de Souza Castanheira, Pedro & Baptista, Renner Luiz Cerqueira" docDate="2024" docId="CD5166CF00595E44A1C1A696A30D90E4" docLanguage="en" docName="Evolutionary Systematics 8(2): 261-271" docOrigin="Evolutionary Systematics 8 (2)" docSource="https://evolsyst.pensoft.net/article/141847/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Harmonicon rufescens F. O. Pickard-Cambridge 1896" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="271" masterDocId="784085DF996A5FEC89FEF9FE7A70E5A1" masterDocTitle="A new species of Harmonicon F. O. Pickard-Cambridge, 1896, from central-west Brazil and redescription of H. rufescens F. O. Pickard-Cambridge, 1896 (Araneae, Mygalomorphae, Dipluridae)" masterLastPageNumber="271" masterPageNumber="261" pageNumber="261" updateTime="1732123585974" updateUser="admin">
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<mods:titleInfo id="5D461432CF3799FB640A849E5215272F">
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<mods:title id="614544574D5979AB0BAF07D9B59095F4">A new species of Harmonicon F. O. Pickard-Cambridge, 1896, from central-west Brazil and redescription of H. rufescens F. O. Pickard-Cambridge, 1896 (Araneae, Mygalomorphae, Dipluridae)</mods:title>
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<mods:name id="01AF6A8B7710EFA0C9311F60E9E77ADF" type="personal">
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<mods:role id="1330470192A4F3312AEE80CACDE4C597">
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<mods:roleTerm id="4B48254F8D1F3433B986ACFDD489D813">Author</mods:roleTerm>
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<mods:namePart id="20AB82A391163D67E31D0BA9A5CC6B8D">Wermelinger-Moreira, Gabriel</mods:namePart>
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<mods:affiliation id="5E571D989EA546D95DF935BF5720BA8B">Laboratório de Diversidade de Aracnídeos, Universidade Federal do Rio de Janeiro, Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundão, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:name id="ECBAD33C8263EFBD0B06411373B2C726" type="personal">
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<mods:roleTerm id="E4518ADCBDC25830F25A7FAF3A30EC4F">Author</mods:roleTerm>
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<mods:namePart id="99870744D8AC4D85EBEB530E0EE409C3">de Souza Castanheira, Pedro</mods:namePart>
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<mods:nameIdentifier id="2B41DB96AE85A53E8B84DB5D1435A245" type="ORCID">0000-0002-0623-1622</mods:nameIdentifier>
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<mods:affiliation id="7D9C26A8745B4E8AE2A2B70C5533593D">Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia & Laboratório de Diversidade de Aracnídeos, Universidade Federal do Rio de Janeiro, Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundão, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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</mods:name>
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<mods:name id="921DF034E556BFBAEABE7254221D9B60" type="personal">
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<mods:roleTerm id="1E2FC89135197164F3698D38189C72F2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A824F883518691BB0C7B735A7CB3DA42">Baptista, Renner Luiz Cerqueira</mods:namePart>
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<mods:affiliation id="AC652EB20C75B5664CAA2F26B2981E37">Laboratório de Diversidade de Aracnídeos, Universidade Federal do Rio de Janeiro, Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundão, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:titleInfo id="9E632A0202A55AC513286E4339830A8E">
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<mods:title id="901E127CA0F1079B16D790C602E3F996">Evolutionary Systematics</mods:title>
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</mods:titleInfo>
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<mods:part id="B65C4DE5D003F35C12849139FF97A49D">
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<mods:date id="7E92728350D224D2DC17CEB08B015D55">2024</mods:date>
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<mods:number id="B40F8C1F5951081D86A78FD202265383">2024-11-20</mods:number>
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</mods:detail>
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<mods:detail id="60F52444EDBC5A907B79C321466A044C" type="volume">
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<mods:number id="77A196E8A320007AF9F7D88238F7BAD7">8</mods:number>
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</mods:detail>
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<mods:detail id="D10A8230D24F73CB57D7B35D867FFAC2" type="issue">
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<mods:number id="690271DBFD4BB76C69788ECA060E3A44">2</mods:number>
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<mods:start id="DCAF915E7B37D23FFC16261E0805C373">261</mods:start>
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<mods:end id="910F696D5A79536141FD1277B70F33DD">271</mods:end>
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<mods:classification id="9AF7ADED974DA51B7EF7E3025271F057">journal article</mods:classification>
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<mods:identifier id="A84449812E2E3D14CE6A3DF091E8131C" type="DOI">10.3897/evolsyst.8.141847</mods:identifier>
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<mods:identifier id="CB19C88B8B3F14EE80769BA986BF28D1" type="ZooBank">8E0DCC28-3448-4001-90B7-E675BC107381</mods:identifier>
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</mods:mods>
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<treatment id="CD5166CF00595E44A1C1A696A30D90E4" ID-arpha="CD5166CF-0059-5E44-A1C1-A696A30D90E4" LSID="urn:lsid:plazi:treatment:CD5166CF00595E44A1C1A696A30D90E4" httpUri="http://treatment.plazi.org/id/CD5166CF00595E44A1C1A696A30D90E4">
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<subSubSection id="646AED9FA76FBCA444410666FE578974" type="nomenclature">
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<paragraph id="717CF2EA41ADA6EBB823A0EA656F50F3">
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<taxonomicName id="E8BF43ADC9757ABC92F60DF179CB5D8C" ID-CoL="6LCCH" authority="F. O. Pickard-Cambridge, 1896" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
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<emphasis id="E36C899B4ADC3C1022CB9FB6DFE191F9" italics="true">Harmonicon rufescens</emphasis>
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F. O.
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<bibRefCitation id="3121A5A5ED4589ABFA4FD3112AA4CE2E" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship " Faraday " up the river Amazons. Proceedings of the zoological Society of London 1896: 716 - 766. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03076. x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge, 1896</bibRefCitation>
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</taxonomicName>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C2497B9668D9AE37ACF8732E8A99FA1B" type="description">
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||
<paragraph id="D33D9FD8570CEE88BB20C5403704C761">
|
||
<figureCitation id="24407B43AC4ECC5A5952AE39664E7D34" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">Figs 1 A – G</figureCitation>
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,
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<figureCitation id="D0FBE8BA4747C1F86A321493AC4B010B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution records of H. rufescens (holotype = red and black circle, specimen here described = red circle) and H. candango sp. nov. (yellow square)." httpUri="https://binary.pensoft.net/fig/1178838">4</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="35BD7B2C43E834C57E29CFF9DAF36860" type="reference_group">
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||
<paragraph id="4B40EEB6020BEA06057BA867DC2F4EFF">
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||
<treatmentCitationGroup id="2B7D063F085C093BFCAF1FE4660248BA">
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||
<treatmentCitation id="9155088197E9CE933D47F8C3E44301BD" author="Pickard-Cambridge FO" page="756" year="1896">
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<taxonomicName id="FC953F1F97C8F2C0DAD54A9F46916E41" ID-CoL="6LCCH" authority="F. O. Pickard-Cambridge, 1896: 756" authorityName="F. O. Pickard-Cambridge" authorityPageNumber="756" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
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<emphasis id="DC9B844DD09176E8A83F847826857CAC" italics="true">Harmonicon rufescens</emphasis>
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<comment id="2158AC59DB2CB90A55EBA644FB881C37">
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||
F. O.
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<bibRefCitation id="8EE1372981B8410C19B0AE4FB88C74A3" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship " Faraday " up the river Amazons. Proceedings of the zoological Society of London 1896: 716 - 766. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03076. x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge, 1896: 756</bibRefCitation>
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, pl. 33, fig. 3, 6, pl. 35, fig. 2–3 (juvenile).
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||
</comment>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="55C12522A939CDA0DB41F17616C2DA3B">
|
||
<treatmentCitationGroup id="DE7EDE0F3A34F97416D2AEA5139A3B9C">
|
||
<taxonomicName id="864B29E6CAF52E4BD7178EBDDF8EAC62" authorityName="Masi" authorityYear="1917" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
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||
<emphasis id="2976E0A7CEAA6095773A3347F9D81123" italics="true">Diplura rufescens</emphasis>
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||
</taxonomicName>
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||
<comment id="BAD6E5AEB9B9AD41D0F84AB685E40602">
|
||
:
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||
<treatmentCitation id="1D8F0EE581DC1C9301DE7D5D59FE857B" author="Raven RJ" page="153" year="1985">
|
||
<bibRefCitation id="A57D244AC35AE96BB526CB592BABE460" author="Raven RJ" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="1-180" refId="B15" refString="Raven RJ (1985) The spider infraorder Mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History 182: 1 - 180." title="The spider infraorder Mygalomorphae (Araneae): cladistics and systematics." volume="182" year="1985">Raven, 1985: 153</bibRefCitation>
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||
</treatmentCitation>
|
||
(synonymy and transfer to
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<taxonomicName id="B3878E6F17024E191B977278BDF3B33A" ID-CoL="46FP" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
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||
<emphasis id="DA878982C9C5A0E05DF91ECC9B6D2266" italics="true">Diplura</emphasis>
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||
</taxonomicName>
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||
).
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||
</comment>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="984C2CEDC0297CEB6C3B02FA26FFCBC7">
|
||
<treatmentCitationGroup id="D071A8AF1E5E0A4F4C52C1446552F6C0">
|
||
<taxonomicName id="8369C5921118DF46072F4EEBCA907ED8" ID-CoL="6LCCH" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
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||
<emphasis id="216BA85F8862E0204654ED7F804E0126" italics="true">Harmonicon rufescens</emphasis>
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||
</taxonomicName>
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||
<comment id="ED85FBBC9F81747FEBF6C4A1A0192E40">
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||
:
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||
<treatmentCitation id="25A36FE157D3CDAA0C7CC3EA6D9AE31E" author="Maréchal P & Marty C" page="500" year="1998">
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||
<bibRefCitation id="6B987D9A1409F2A30306B7EC0ECE00EA" DOI="10.5962/p.268900" author="Maréchal P & Marty C" journalOrPublisher="Zoosystema" pagination="499-504" refId="B9" refString="Marechal P, Marty C (1998) Rehabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espece de Guyane francaise (Araneae, Mygalomorphae, Dipluridae). Zoosystema 20: 499 - 504. https: // doi. org / 10.5962 / p. 268900" title="Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae)." volume="20" year="1998">Maréchal & Marty, 1998: 500</bibRefCitation>
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||
</treatmentCitation>
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||
(rejected synonym with
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||
<taxonomicName id="077CCE6865368996EC5CDE2ED753AF9F" ID-CoL="46FP" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
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<emphasis id="0829A2E6FCF6DF70DBD5055DA10A245E" italics="true">Diplura</emphasis>
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</taxonomicName>
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||
).
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||
</comment>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0E56AE" type="material">
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||
<paragraph id="1E4FB6B79233F5A13A8033C53EA03687">
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||
<heading id="BA6AF0B713B17B338FD2E1EE121ED432" reason="title">Material examined.</heading>
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||
</paragraph>
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||
<paragraph id="F0D4B38EC4DB633BC8490DAAFFC67737">
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||
<materialsCitation id="FDF67CDED970F018E99995252D3E6B19" collectionCode="NHMUK" collectorName="F. O. Pickard-Cambridge & Austen" country="Brazil" county="lower Amazon" location="Santarem" specimenCount="1" specimenCount-male="1" stateProvince="Para" typeStatus="holotype">
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||
<emphasis id="E0E61B68E6C6FEBCE9A8F40B945E4D39" bold="true" italics="true">
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||
<typeStatus id="2A81585AD437CBE2B1367198AB519209" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
:
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||
<collectingCountry id="A43363A0366C849D305808BA63C40822" name="Brazil">Brazil</collectingCountry>
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||
•
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||
<specimenCount id="25F6324E8A1DB83A74674FAB78C6EC77" count="1" type="male">♂</specimenCount>
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||
imm.,
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||
<collectingCounty id="F79F50F53891D8B29C5ED693C23AB294">lower Amazon</collectingCounty>
|
||
,
|
||
<location id="75E5444D00E07C230848173114CC365C" LSID="urn:lsid:plazi:treatment:CD5166CF00595E44A1C1A696A30D90E4:75E5444D00E07C230848173114CC365C" country="Brazil" county="lower Amazon" name="Santarem" stateProvince="Para">Santarém</location>
|
||
,
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||
<collectingRegion id="E9B58214D9DE6E0DF5A73FEF58AC6162" country="Brazil" name="Para">Pará</collectingRegion>
|
||
; no date;
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||
<collectorName id="93721F41CA839D7E48051E919EBE8778">F. O. Pickard-Cambridge</collectorName>
|
||
and
|
||
<collectorName id="46A1EBA836FCAFF0ECF59454E96B8858">Austen</collectorName>
|
||
leg;
|
||
<named-content id="41489FBC9490A905CC663A1D84665E41" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum of the United Kingdom">
|
||
<collectionCode id="68A09A90DE2157845A0C842F37AE7730" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||
</named-content>
|
||
(no registration number). Not examined.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="BBDA7E3DB0EC9AAF7B4772A12F6B6D83">
|
||
<heading id="64D28AAA73CE70F34C9064FFACACAC4C" reason="title">Other material.</heading>
|
||
</paragraph>
|
||
<paragraph id="1DEBDB34B1C1E8AF51D4C016766D0243">
|
||
<materialsCitation id="16A195C7EC89E218D2004538EC293039" collectingDate="2009-04-14" collectionCode="UFRJ" collectorName="A. P. L. Giupponi & D. R. Pedroso" country="Brazil" latitude="-3.2023056" location="Altamira" longLatPrecision="1" longitude="-52.22678" specimenCode="UFRJ 2575" specimenCount="1" specimenCount-female="1" stateProvince="Para">
|
||
<collectingCountry id="9AD8C8DEEC121075C323CD12978C068F" name="Brazil">Brazil</collectingCountry>
|
||
•
|
||
<specimenCount id="4FC43E11EA59C26C1AC0B145B9D0D1E5" count="1" type="female">♀</specimenCount>
|
||
;
|
||
<collectingRegion id="8A28565F6CD1A0F4ECB498D9ECB02A52" country="Brazil" name="Para">Pará</collectingRegion>
|
||
,
|
||
<location id="5B24138D90A099738D9DD79BACB91727" LSID="urn:lsid:plazi:treatment:CD5166CF00595E44A1C1A696A30D90E4:5B24138D90A099738D9DD79BACB91727" country="Brazil" latitude="-3.2023056" longLatPrecision="1" longitude="-52.22678" name="Altamira" stateProvince="Para">Altamira</location>
|
||
[
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXAAG" specific-use="{"type":"Point","coordinates":[-52.226778,-3.202306]}">
|
||
<geoCoordinate id="E09C2A000C38A1F44AA0420FBDF02145" degrees="3" direction="south" minutes="12" orientation="latitude" precision="1" seconds="08.3" value="-3.2023056">3°12'08.3"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2AB38830FBFF433FB3459E756FBBE0F4" degrees="52" direction="west" minutes="13" orientation="longitude" precision="1" seconds="36.4" value="-52.22678">52°13'36.4"W</geoCoordinate>
|
||
</named-content>
|
||
];
|
||
<date id="789D83441CF7766DCC42976EA5D7BD60" value="2009-04-14">
|
||
<collectingDate id="FEEB5C5C4022CB4A3E6D167FE7714B8C" value="2009-04-14">14 April 2009</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="52F18FB007C3B5B8A263637FB9F135C9">A. P. L. Giupponi</collectorName>
|
||
and
|
||
<collectorName id="BC09FB56BAB4DDED8BF27161789BF089">D. R. Pedroso</collectorName>
|
||
leg.;
|
||
<specimenCode id="33285C9FB5F9D938F3BBA987A1790EF5" collectionCode="UFRJ">
|
||
<abbrev id="ABBRID0E3AAG" xlink_title="Laboratório de Diversidade de Aracnídeos (LABAR), Universidade Federal do Rio de Janeiro">UFRJ</abbrev>
|
||
2575
|
||
</specimenCode>
|
||
</materialsCitation>
|
||
) •
|
||
<materialsCitation id="8624681672DFD79405129B256C0C8226" collectingDate="2009-04-14" collectionCode="MNRJ" collectorName="A. P. L. Giupponi & D. R. Pedroso" country="Brazil" latitude="-3.2023056" location="Altamira" longLatPrecision="1" longitude="-52.22678" specimenCode="MNRJ 4371" specimenCount="2" specimenCount-female="2" stateProvince="Para">
|
||
<specimenCount id="BA0E3D1C7FE08D0AAC6CCDC542AE73D4" count="2" type="female">2 ♀♀</specimenCount>
|
||
, 5 j; same data as for previous;
|
||
<specimenCode id="3314D23DA1FBC2D56737AD37D72FCF1F" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" type="">
|
||
<named-content id="6CB8AD944DDD9505E75A3885BBEF02F9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">MNRJ</named-content>
|
||
4371
|
||
</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="F5A9E60AA741CB5582663538D5FF60B2" collectingDate="2011-04-02" collectingDateMax="2011-04-07" collectingDateMin="2011-04-02" collectionCode="MNRJ" collectorName="D. R. Pedroso" country="Brazil" latitude="-3.2023056" location="Altamira" longLatPrecision="1" longitude="-52.22678" specimenCode="MNRJ 4362" specimenCount="1" specimenCount-female="1" stateProvince="Para">
|
||
<specimenCount id="298D05EBD3724DC6370FB213FCD3D0EC" count="1" type="female">♀</specimenCount>
|
||
, 4 j; same locality as preceding;
|
||
<date id="042E6CCD79435BF62F6023BB540AD46A" value="2011-04-02" valueMax="2011-04-07" valueMin="2011-04-02">
|
||
<collectingDate id="EA0A7C722297691A1303827FB7157976" value="2011-04-02" valueMax="2011-04-07" valueMin="2011-04-02">2–7 April 2011</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="9F69B89FE04C41BFF8FE0E1713313A21">D. R. Pedroso</collectorName>
|
||
leg;
|
||
<specimenCode id="7FB4DB5E507AF7776E7C6B5F6808D1FD" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" type="">
|
||
<named-content id="E08279EA3EAA719DEBF20E64DEC681C0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">MNRJ</named-content>
|
||
4362
|
||
</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EIBAG" type="diagnosis">
|
||
<paragraph id="79FC623E5ED8DCB414D9657D8975E465">
|
||
<heading id="B3330D6363A54C43A3022C3CE2CF0D7F" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="D7831324D80D588EB5EDBCA64D49E234">
|
||
Females of
|
||
<taxonomicName id="D222D749D049427790DBF647E3A531CE" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
|
||
<emphasis id="53025C3CADDD651DDAEDAC2BFBF1E638" italics="true">H. rufescens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1605B172878A822D155B8DD69FB81B05" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
|
||
<emphasis id="E63984665C51BD4BF9993E85677B0F24" italics="true">H. oiapoqueae</emphasis>
|
||
</taxonomicName>
|
||
present spermathecae with median branch ending in a blunt rounded tip and without additional lobules (Fig.
|
||
<figureCitation id="571A3A8AEE6F712BC18603EDDF5F8C4F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 D – F</figureCitation>
|
||
;
|
||
<bibRefCitation id="DB427A38D76CEA5214485ACB0C431BBF" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 89–96. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
|
||
, fig. 5). Females of
|
||
<taxonomicName id="BC0202742F5164B724E59F6D1BC45626" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
|
||
<emphasis id="04FAC87223D1DE47B217E695B531657B" italics="true">H. rufescens</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="48CC6A8DE0ED17A5FB057941AC100BA4" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
|
||
<emphasis id="2032B2D37EBD91E27263CA12378A6789" italics="true">H. oiapoqueae</emphasis>
|
||
</taxonomicName>
|
||
by the tarsal claw of the pedipalp with a double row of teeth (vs. unique row;
|
||
<bibRefCitation id="16A607A9491DFE390569D3F4A985D8A5" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 89–96. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
|
||
, p. 92); and spermathecae lateral branch with its tip folded over the stem and without lobules (vs. tip bearing several lobules) (Fig.
|
||
<figureCitation id="3C8D0179BE3DF994023D305F610AA466" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 D – F</figureCitation>
|
||
;
|
||
<bibRefCitation id="A422CD02690FECB0194BFC82E5A18A50" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 89–96. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
|
||
, fig. 5).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EODAG" type="Taxonomic notes">
|
||
<paragraph id="488545477EC089CB24E0754C42D3BBFB">
|
||
<heading id="26F0CC550291629FA3CAE0241658A55B" reason="title">Taxonomic notes.</heading>
|
||
</paragraph>
|
||
<paragraph id="10DC8E96CDDE442F3D055AB873414A89">
|
||
<taxonomicName id="33DD7F2AAE3A70581F509140E68E7808" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0BA8FC010C3CC4CAECE444677DB9D31" italics="true">Harmonicon</emphasis>
|
||
</taxonomicName>
|
||
was established by F. O.
|
||
<bibRefCitation id="2F173FCD41C39D6E91840E05CCC412CE" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" firstAuthor="Pickard-Cambridge" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons. Proceedings of the zoological Society of London 1896: 716–766. https://doi.org/10.1111/j.1096-3642.1896.tb03076.x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge (1896)</bibRefCitation>
|
||
to accommodate a single species,
|
||
<taxonomicName id="DA0B5D1E9721A0559CA4918B085F415A" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
|
||
<emphasis id="D02A4CC5FD746ADD355AC6FF7C625E8E" italics="true">H. rufescens</emphasis>
|
||
</taxonomicName>
|
||
, based on a presumed female specimen from Santarém,
|
||
<collectingCountry id="2A7DC6BF3589A4F974AA1EB2495E9873" name="Brazil">Brazil</collectingCountry>
|
||
.
|
||
<bibRefCitation id="B833D1164D9316F557FED8215AE2EBB4" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 89–96. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam (2011</bibRefCitation>
|
||
, p. 91) were able to examine the
|
||
<typeStatus id="21D28D2D8770E0F6E4E72897AA84D238" type="holotype">holotype</typeStatus>
|
||
and noticed that “ Cambridge (1896) was incorrect in stating that the
|
||
<typeStatus id="654912FDFE44C2BA5039AC2BEDEEB286" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="8B086A5B238696FB85DBB97C62024480" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
|
||
<emphasis id="E06E27131573A2CAF7AC6CBCBE2CCDDF" italics="true">Harmonicon rufescens</emphasis>
|
||
</taxonomicName>
|
||
is a female; it is in fact a juvenile male ”. The female described here is from Altamira, also in the
|
||
<collectingRegion id="FA076F934F2463491AA46812EDEF7CF7" country="Brazil" name="Para">Pará state</collectingRegion>
|
||
,
|
||
<collectingCountry id="512D30E8EC3F6CA7825C7BC9F8FE73E6" name="Brazil">Brazil</collectingCountry>
|
||
, and located ca.
|
||
<quantity id="073272E385553BA4AAB2AD7EA5E358E9" metricMagnitude="5" metricUnit="m" metricValue="3.0" unit="km" value="300.0">300 km</quantity>
|
||
from Santarém. Both localities are placed in the lower Amazons, have similar climate, low altitude, and are covered by dense or open Ombrophile Amazonian Forest and Savannas (
|
||
<bibRefCitation id="AFEC3F20B0387D2D9BC79CC7CD8F4FF2" DOI="10.1590/S0100-84042008000100015" author="Magnusson" etAl="et al." firstAuthor="Magnusson" issue="1" journalOrPublisher="Revista Brasileira de Botânica" pagination="165-177" refId="B8" refString="Magnusson WE, Lima AP, Albernaz ALKM, Sanaiotti TM, Guillaumet J-L (2008) Composição florística e cobertura vegetal das savanas na região de Alter do Chão, Santarém – PA. Revista Brasileira de Botânica 31 (1): 165–177. https://doi.org/10.1590/S0100-84042008000100015" title="Composição florística e cobertura vegetal das savanas na região de Alter do Chão, Santarém - PA." volume="31" year="2008">Magnusson et al. 2008</bibRefCitation>
|
||
). Other
|
||
<taxonomicName id="2DB8098E47A5B2A226B60661D06B541F" authorityName="Simon" authorityYear="1889" class="Arachnida" family="Dipluridae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="family">Dipluridae</taxonomicName>
|
||
species collected in Santarém at the same expedition to lower Amazon as
|
||
<taxonomicName id="8FB47447A1F92D37C9CB80F5B0C42955" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
|
||
<emphasis id="08BD7DA3964FC42FBAEFE21756DFB54F" italics="true">H. rufescens</emphasis>
|
||
</taxonomicName>
|
||
also have large distribution range.
|
||
<taxonomicName id="0A3549DA830057B77FC2E143C2E34AC3" authority="(F. O. Pickard-Cambridge, 1896)" baseAuthorityName="F. O. Pickard-Cambridge" baseAuthorityYear="1896" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="sanguinea">
|
||
<emphasis id="9183A2E7EEC5FE79815A9B51B28EF0A5" italics="true">Diplura sanguinea</emphasis>
|
||
(F. O.
|
||
<bibRefCitation id="0DDA3839AE32E3CF81F8A9D5F5EB34B8" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship " Faraday " up the river Amazons. Proceedings of the zoological Society of London 1896: 716 - 766. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03076. x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge, 1896</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
is widespread through North and Northeast regions of
|
||
<collectingCountry id="486BB66F2958048C9AE7781AB22A57A6" name="Brazil">Brazil</collectingCountry>
|
||
(
|
||
<bibRefCitation id="8BACF6D03E25DBA9C85DCB22D29D474D" DOI="10.13156/arac.2020.18.7.681" author="Brescovit" etAl="et al." firstAuthor="Brescovit" issue="7" journalOrPublisher="Arachnology" pagination="681-689" refId="B2" refString="Brescovit AD, Sherwood D, Lucas SM (2021) First description of the males of Diplura sanguinea (F. O. Pickard-Cambridge, 1896) and Diplura nigra (F. O. Pickard-Cambridge, 1896) from Brazilian Amazonia, with notes on biogeography and opisthosomal patterning (Araneae, Dipluridae). Arachnology 18 (7): 681–689. https://doi.org/10.13156/arac.2020.18.7.681" title="First description of the males of Diplura sanguinea (F. O. Pickard-Cambridge, 1896) and Diplura nigra (F. O. Pickard-Cambridge, 1896) from Brazilian Amazonia, with notes on biogeography and opisthosomal patterning (Araneae, Dipluridae)." volume="18" year="2021">Brescovit et al. 2021</bibRefCitation>
|
||
) and is also found in
|
||
<collectingCountry id="CC357469CBD4F5F1BDA30D03DD99A6E7" name="Colombia">Colombia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="2390F921A1FA7C6E74C81F9D34D55132" DOI="10.5281/zenodo.10201779" author="Sherwood" etAl="et al." firstAuthor="Sherwood" journalOrPublisher="ZooNova" pagination="1-22" refId="B16" refString="Sherwood D, Drolshagen B, Osorio LM, Benavides LR, Seiter M (2023) An inordinate fondness for spinnerets: on some spiders of the genera Diplura C. L. Koch, 1850 and Linothele Karsch, 1879 with new species, records, and notes on types (Araneae, Dipluridae). ZooNova 29: 1–22. https://doi.org/10.5281/zenodo.10201779" title="An inordinate fondness for spinnerets: on some spiders of the genera Diplura C. L. Koch, 1850 and Linothele Karsch, 1879 with new species, records, and notes on types (Araneae, Dipluridae)." volume="29" year="2023">Sherwood et al. 2023</bibRefCitation>
|
||
).
|
||
<taxonomicName id="85E7DC81C74BE3AE4F18FE6B45215F8A" authority="(F. O. Pickard-Cambridge, 1896)" baseAuthorityName="F. O. Pickard-Cambridge" baseAuthorityYear="1896" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nigra">
|
||
<emphasis id="710E6168927C29EF358623AB65FC056F" italics="true">Diplura nigra</emphasis>
|
||
(F. O.
|
||
<bibRefCitation id="CFC665F92B1249C84C6D173BEB37A3D2" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship " Faraday " up the river Amazons. Proceedings of the zoological Society of London 1896: 716 - 766. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03076. x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge, 1896</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
has been recorded from several localities around Santarém (
|
||
<bibRefCitation id="EE703EBF38205B9B22EB0E525035EF9E" DOI="10.13156/arac.2020.18.7.681" author="Brescovit" etAl="et al." firstAuthor="Brescovit" issue="7" journalOrPublisher="Arachnology" pagination="681-689" refId="B2" refString="Brescovit AD, Sherwood D, Lucas SM (2021) First description of the males of Diplura sanguinea (F. O. Pickard-Cambridge, 1896) and Diplura nigra (F. O. Pickard-Cambridge, 1896) from Brazilian Amazonia, with notes on biogeography and opisthosomal patterning (Araneae, Dipluridae). Arachnology 18 (7): 681–689. https://doi.org/10.13156/arac.2020.18.7.681" title="First description of the males of Diplura sanguinea (F. O. Pickard-Cambridge, 1896) and Diplura nigra (F. O. Pickard-Cambridge, 1896) from Brazilian Amazonia, with notes on biogeography and opisthosomal patterning (Araneae, Dipluridae)." volume="18" year="2021">Brescovit et al. 2021</bibRefCitation>
|
||
), and we examined specimens from other localities from
|
||
<collectingRegion id="15EF8AF7902B4A593F9EDC526745343A" country="Brazil" name="Para">Pará</collectingRegion>
|
||
, Amazonas,
|
||
<collectingRegion id="1CA6F6303AB0243912740AA7A4DCC4F9" country="Brazil" name="Roraima">Roraima</collectingRegion>
|
||
,
|
||
<collectingRegion id="E23DED24BBC3A9811A6DF16CFBA237E0" country="Brazil" name="Rondonia">Rondônia</collectingRegion>
|
||
, and
|
||
<collectingRegion id="EEA22AEDE22139A1D2FABE88BF4A960E" country="Brazil" name="Mato Grosso">Mato Grosso</collectingRegion>
|
||
states in
|
||
<collectingCountry id="06A965ADDD9CDFB4FAAD52DC2C74E186" name="Brazil">Brazil</collectingCountry>
|
||
. The only other species described for the same Brazilian state is
|
||
<taxonomicName id="3D90B4001B16ED85A5C5B6C4A4991171" authorityName="Pedroso & Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
|
||
<emphasis id="4F94638939AE1B5DC100A5B41F17E788" italics="true">H. cerberus</emphasis>
|
||
</taxonomicName>
|
||
, from Pequiá cave, near Carajás, in the municipality of Parauapebas, but its male
|
||
<typeStatus id="E153E4CFA82F0A95FEF9C5341C0B892D" type="holotype">holotype</typeStatus>
|
||
has several troglomorphic characteristics, and its type locality is placed ca.
|
||
<quantity id="D34B3C61666D5EB7731CEC2B5536ED29" metricMagnitude="5" metricUnit="m" metricValue="6.6" unit="km" value="660.0">660 km</quantity>
|
||
away from Santarém (
|
||
<bibRefCitation id="9C5434817ED2F5F612571C2F64DDEE4B" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 77–88. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
|
||
). Besides, the region around the cave where
|
||
<taxonomicName id="DE1242AA8DDF75D8A879DF83692991E9" authorityName="Pedroso & Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
|
||
<emphasis id="EAB616FCDE3D42DD5A77211F8A1E1120" italics="true">H. cerberus</emphasis>
|
||
</taxonomicName>
|
||
was collected is covered by the Amazon “ canga ” vegetation, which covers areas rich in iron ores and has unique vegetation (
|
||
<bibRefCitation id="224759803BB927558E9189CAD063BF23" DOI="10.1590/2175-7860201869336" author="Mota" etAl="et al." firstAuthor="Mota" issue="3" journalOrPublisher="Rodriguésia" pagination="1435-1488" refId="B10" refString="Mota NFO, Watanabe MTC, Zappi DC, Hiura AL, Pallos J, Viveros RS, Giulietti AM, Viana PL (2018) Amazon canga: the unique vegetation of Carajás revealed by the list of seed plants. Rodriguésia 69 (3): 1435–1488. https://doi.org/10.1590/2175-7860201869336" title="Amazon canga: the unique vegetation of Carajás revealed by the list of seed plants." volume="69" year="2018">Mota et al. 2018</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ENHAG" type="description">
|
||
<paragraph id="F5BE77306B89494E801C8154B13F02AF">
|
||
<heading id="D65F7C22B770350BF0D2936E3C826D4F" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="8693BDFBE6977ABF683D3E2D3263A9FB">
|
||
<emphasis id="08ADF6A3B66511E9878C3091785723E8" bold="true">
|
||
Female (
|
||
<abbrev id="ABBRID0EVHAG" xlink_title="Laboratório de Diversidade de Aracnídeos (LABAR), Universidade Federal do Rio de Janeiro">
|
||
<collectionCode id="94ADA5CE77F1B1B5F652A659301BA591">UFRJ</collectionCode>
|
||
</abbrev>
|
||
2575)
|
||
</emphasis>
|
||
. Carapace oblong, wider at middle portion, light orange-brown, covered with dark setae, caput slightly elevated, darker; cephalic area not shortened, darker furrows, margins of thoracic area with a row of long, thick, rigid, erect, and curved dark setae pointing ectally (Fig.
|
||
<figureCitation id="88B51B1CC0E403868D928F3ED26BCD90" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
|
||
). Clypeus long, with frontal margin carrying 19 median-to-long, thick setae facing forward (Fig.
|
||
<figureCitation id="158FF84A0509C2C5CC70D37F6805206D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
|
||
). Eye tubercle with 15 thick setae on its anterior margin; area between eyes with 11 short and thickened setae, and others abundant and dark but not thickened setae (Fig.
|
||
<figureCitation id="041920A4FCC929D6C2892DC7D59805FA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
|
||
). Eye formula:
|
||
<abbrev id="ABBRID0EGIAG" xlink_title="anterior lateral eyes">ALE</abbrev>
|
||
>
|
||
<abbrev id="ABBRID0EKIAG" xlink_title="posterior lateral eyes">PLE</abbrev>
|
||
>
|
||
<abbrev id="ABBRID0EOIAG" xlink_title="anterior median eyes">AME</abbrev>
|
||
>
|
||
<abbrev id="ABBRID0ESIAG" xlink_title="posterior median eyes">PME</abbrev>
|
||
. Ocular area much wider than long. Anterior eye row slightly recurved. Posterior eye row slightly recurved. Median ocular quadrangle wider than long, wider at its posterior margin.
|
||
<abbrev id="ABBRID0EWIAG" xlink_title="anterior median eyes">AME</abbrev>
|
||
rounded,
|
||
<abbrev id="ABBRID0E1IAG" xlink_title="anterior lateral eyes">ALE</abbrev>
|
||
and
|
||
<abbrev id="ABBRID0E5IAG" xlink_title="posterior lateral eyes">PLE</abbrev>
|
||
elliptical, and
|
||
<abbrev id="ABBRID0ECJAG" xlink_title="posterior median eyes">PME</abbrev>
|
||
reduced in size, oval.
|
||
<abbrev id="ABBRID0EGJAG" xlink_title="anterior median eyes">AME</abbrev>
|
||
-
|
||
<abbrev id="ABBRID0EKJAG" xlink_title="anterior median eyes">AME</abbrev>
|
||
separated by less than its diameter.
|
||
<abbrev id="ABBRID0EOJAG" xlink_title="anterior median eyes">AME</abbrev>
|
||
contiguous to
|
||
<abbrev id="ABBRID0ESJAG" xlink_title="posterior median eyes">PME</abbrev>
|
||
.
|
||
<abbrev id="ABBRID0EWJAG" xlink_title="anterior median eyes">AME</abbrev>
|
||
separated from
|
||
<abbrev id="ABBRID0E1JAG" xlink_title="posterior median eyes">PME</abbrev>
|
||
by around half of its diameter.
|
||
<abbrev id="ABBRID0E5JAG" xlink_title="anterior lateral eyes">ALE</abbrev>
|
||
and
|
||
<abbrev id="ABBRID0ECKAG" xlink_title="posterior median eyes">PME</abbrev>
|
||
contiguous to
|
||
<abbrev id="ABBRID0EGKAG" xlink_title="posterior lateral eyes">PLE</abbrev>
|
||
. Fovea transverse and slightly recurved (Fig.
|
||
<figureCitation id="3DFF5D8F221FC78A592703894DD8C67B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
|
||
). Labium wider than long with no cuspules (Fig.
|
||
<figureCitation id="13C17AF2159424739329682264213536" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 B</figureCitation>
|
||
). Sternum longer than wide, covered by black stiff setae and brownish thin recumbent setae, one pair of labial comma-shaped sigilla clearly separated from each other, three sternal sigilla near margin on each side, increasing in size from first to last, the third one much larger and elliptical (Fig.
|
||
<figureCitation id="0BF7593C41B54CFE2A64216E446777CB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 B</figureCitation>
|
||
). Chelicerae dark red with 10 promarginal teeth at left chelicera and 12 promarginal teeth at right chelicera, fangs furrow with denticles. Plectrum with 6–8 separated thick setae on left side and right side, respectively. Maxillae elongated with 25 cuspules at the right side and 31 at the left one (Fig.
|
||
<figureCitation id="08A839B1CA5B89202C64BB31F1EEEA97" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 B</figureCitation>
|
||
). Lyra at ventral side of maxilla, asymmetrical, formed by 6–7 modified thick and long setae, increasing in size in a basal to distal direction, with a distinct gap between the base of each seta (Fig.
|
||
<figureCitation id="8249D90DBF8BAC0BBD3422CCEE61EE73" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
|
||
). Left lyra with six setae (Fig.
|
||
<figureCitation id="44D9F133F9F199B095ED185488703FBA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
|
||
), right lyra with seven (two broken) setae. Left lyra thick, erect, regularly curved setae (Fig.
|
||
<figureCitation id="A6AC732A2A03DE820F9D3E640D62066D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
|
||
, arrows) placed distally and interspersed among the field of long and thin setae covering the retrolateral face of maxilla, five at the left side and four at the right side. Leg formula IV> I> II> III; length of segments (femur + patella + tibia + metatarsus + tarsus = total length): I – 8.4 + 2.8 + 8.3 + 6.9 + 5.1 = 31.5, II – 7.4 + 2.6 + 7.5 + 6.9 + 4.8 = 29.2, III – 6.7 + 2.3 + 7.0 + 8.2 + 4.6 = 28.8, IV – 8.6 + 2.7 + 9.6 + 10.8 + 4.9 = 36.6. Tarsi with few cracks covering almost all dorsal and lateral faces, except by basis and tip of article; all tarsi and metatarsi with thin scopula, metatarsi with inconspicuous scopula; pedipalp tarsi with middle to apical scopulae; pedipalp claw with seven teeth;
|
||
<abbrev id="ABBRID0EGLAG" xlink_title="inferior tarsal claw">ITC</abbrev>
|
||
without teeth;
|
||
<abbrev id="ABBRID0EKLAG" xlink_title="superior tarsal claw">STC</abbrev>
|
||
teeth: I: 9–5, II: 9–4, III – IV: 10–4. Macrosetae: Leg I: femur pld 0–0 – 2, d 1–1 – 1, rld 0–0 – 1, patella 0, tibia v 1–1 – 3 right, p 0–1 – 1, v 1–1 – 2 left, metatarsus v 2–1 – 2; Leg II: femur d 2–1 – 0, pld 0–0 – 1 left, pld 0–2 – 1 right, patella pld 0–0 – 1 left, tibia p 0–0 – 1 left, p 0–1 – 1 right, v 1–2 – 2 right, v 1–1 – 2 left, metatarsus v 1–2 – 2, p 0–1 – 0; Leg III: femur pld 0–0 – 2 left, d 2–0 – 0, rld 0–2 – 1 left, rld 2–2 – 1 right, patella 0, tibia p 0–1 – 1, rld 1–0 – 0, r 0–2 – 1 left, r 0–1 – 1 right, v 2–0 – 2 left, v 2–2 – 2 right, metatarsus d 0–0 – 2, r 0–2 – 0 left, rld 1–0 – 0 left, r 1–2 – 2 right, p 2–1 – 0, v 2–2 – 2 right, v 3–2 – 3 left; Leg IV: femur rld 2–2 – 1, d 1–1 – 0 right, d 2–1 – 0 left, patella r 0–1 – 0, tibia rld 1–0 – 0, p 1–0 – 1, r 1–1 – 1, v 2–2 – 2, metatarsus rld 2–1 – 1 left, rld 2–2 – 1 right, r 0–2 – 0 left, pld 2–1 – 1, v 2–1 – 3 left, v 3–1 – 3 right. Abdomen light brown with no stripes (Fig.
|
||
<figureCitation id="E341515C0B487F29AC0E8A241F3C543B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
|
||
), covered with dark setae, venter uniformly brown.
|
||
<abbrev id="ABBRID0ESLAG" xlink_title="posterior lateral spinnerets">PLS</abbrev>
|
||
longer than half of abdomen, with articulate fusules on all articles, and last article finger-like and with light patches of ventral cracks (Fig.
|
||
<figureCitation id="E556E51DD97B33061E33CFF1EA62F05A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
|
||
). Spermathecae separated by ca. 1.5 × its length, with two sclerotised thick stems (or branches), originating from a common basis poorly sclerotised and translucent, each branch with almost the same thickness and length; median branch slightly curved laterally and with large, rounded, and blunt tip without vesicles; lateral branch straight, with a wider rounded head, with its tip rounded and folded upon the stem (Fig.
|
||
<figureCitation id="89B889160A09F0407B3CF82AC65DD00F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D – F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D – E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 D – F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="3F9E43088F59A15FAC0BEE910D8F9639" ID-arpha="4EEF1C8A-5EA3-5D89-9CEB-2E845FB8C226" httpUri="https://binary.pensoft.net/fig/1178835" startId="F1">
|
||
<paragraph id="B3E5EEED8471BDAC8B2FEFDA27BB5AD7">
|
||
<label id="FAD2A957D21233AA572FAD3D8981E08F">Figure 1.</label>
|
||
</paragraph>
|
||
<paragraph id="B79DB207BCEBD2752C8771C53E42FC8F">
|
||
<taxonomicName id="FB7C67CCC4DAAD5239F170D3D8410B07" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
|
||
<emphasis id="90C3E09590BDC224ABB959D85D03858B" italics="true">Harmonicon rufescens</emphasis>
|
||
</taxonomicName>
|
||
, female (
|
||
<abbrev id="ABBRID0ERMAG" xlink_title="Laboratório de Diversidade de Aracnídeos (LABAR), Universidade Federal do Rio de Janeiro">UFRJ</abbrev>
|
||
2575).
|
||
<emphasis id="07FF4948EFD8584FB43688B0CC69C12C" bold="true">A.</emphasis>
|
||
Dorsal habitus;
|
||
<emphasis id="9FA99C8022C8691DADDD9A5E6D88C450" bold="true">B.</emphasis>
|
||
Sternum;
|
||
<emphasis id="AE2FCAC7825BA030C7214398C3F2B3D2" bold="true">C.</emphasis>
|
||
Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae);
|
||
<emphasis id="8D74EDCD25B23C4243435BA8503A2485" bold="true">D – F.</emphasis>
|
||
Spermathecae:
|
||
<emphasis id="87032CA1A9B7763F9EDA31B2DC02318A" bold="true">D.</emphasis>
|
||
Ventral view;
|
||
<emphasis id="852E183B5BC31819BAD8830F1189E1DF" bold="true">E.</emphasis>
|
||
Dorsal view;
|
||
<emphasis id="1711F92B07ED4D29079567C37D738E62" bold="true">F.</emphasis>
|
||
Ventral view, partially cleared;
|
||
<emphasis id="CAB600B54674C178942836665F23ACEA" bold="true">G.</emphasis>
|
||
Alive immature female specimen in nature (
|
||
<named-content id="2A0929D70E52092768AEF51B276A6043" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">MNRJ</named-content>
|
||
4362). Scale bars: 5 mm (
|
||
<emphasis id="2D9E3E5382F1AD1E600357D4BCF7CF18" bold="true">A</emphasis>
|
||
); 2 mm (
|
||
<emphasis id="FC6A919645B759CDBD8FF40B27B5777E" bold="true">B</emphasis>
|
||
); 1 mm (
|
||
<emphasis id="1511C35EEBE652B17B5637210B708D1B" bold="true">C</emphasis>
|
||
); 0.5 mm (
|
||
<emphasis id="E1BAFF4A97FCCBE1C97A1F3DC757793F" bold="true">D – E</emphasis>
|
||
). Living specimen photo by Denis Pedroso.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="927951B3D4E42C82E665B4FB605847FD">
|
||
<abbrev id="ABBRID0EYNAG" xlink_title="total length">TL</abbrev>
|
||
26.9. Carapace 10.2 long, 9.1 wide, carapace length / width 1.12.
|
||
<abbrev id="ABBRID0E3NAG" xlink_title="anterior median eyes">AME</abbrev>
|
||
<quantity id="ECF95AD1A60048451D43AA876D25521E" metricMagnitude="-4" metricUnit="m" metricValue="3.2" unit="mm" value="0.32">0.32 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0EAOAG" xlink_title="anterior median eyes">AME</abbrev>
|
||
-
|
||
<abbrev id="ABBRID0EEOAG" xlink_title="anterior median eyes">AME</abbrev>
|
||
<quantity id="68BEF96236DA348AC131F798B69106D6" metricMagnitude="-4" metricUnit="m" metricValue="2.4" unit="mm" value="0.24">0.24 mm</quantity>
|
||
. Abdomen 16.7 long, 8.5 wide. Spinnerets:
|
||
<abbrev id="ABBRID0EIOAG" xlink_title="posterior median spinnerets">PMS</abbrev>
|
||
2.6 long;
|
||
<abbrev id="ABBRID0EMOAG" xlink_title="posterior lateral spinnerets">PLS</abbrev>
|
||
16.2 long, basal article 4.7, middle 4.9, distal 6.6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EQOAG" type="variation">
|
||
<paragraph id="0A861BF8F0EDCAD3B22E6C2C0DFC33AA">
|
||
<heading id="392E00B5748BB201AF765FAC6690048D" reason="title">Variation.</heading>
|
||
</paragraph>
|
||
<paragraph id="27E604854406AFD6036DFBA53C3B94BE">
|
||
Females (n = 6).
|
||
<abbrev id="ABBRID0EWOAG" xlink_title="total length">TL</abbrev>
|
||
24.4–32.9. Chelicerae with 10 to 13 promarginal teeth, plectrum with five to eight setae, 25 to 40 maxillary cuspules, and lyra with five to seven bristles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E1OAG" type="distribution">
|
||
<paragraph id="FB4DA11230A85CB8DB6514A915E15973">
|
||
<heading id="9C84DA7CEA38FB557BB9F42BAE976EC2" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="5409048740D6B4D70A227E3A885BAC5D">
|
||
Only known from Altamira and Santarém cities,
|
||
<collectingRegion id="5B545C565CF7C06543107E1AB71FBA40" country="Brazil" name="Para">Pará State</collectingRegion>
|
||
,
|
||
<collectingCountry id="9243043F0F88CD4F799D831240A2D249" name="Brazil">Brazil</collectingCountry>
|
||
(Fig.
|
||
<figureCitation id="730C377838C8BF2AEDAB716B48DECDF3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution records of H. rufescens (holotype = red and black circle, specimen here described = red circle) and H. candango sp. nov. (yellow square)." httpUri="https://binary.pensoft.net/fig/1178838">4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |