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<mods:title id="F60D1A30057C069D7128AAE7E0884378">A new species of Harmonicon F. O. Pickard-Cambridge, 1896, from central-west Brazil and redescription of H. rufescens F. O. Pickard-Cambridge, 1896 (Araneae, Mygalomorphae, Dipluridae)</mods:title>
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<mods:namePart id="83BF1BE2ED00BD3506135D665AB4A2F0">Wermelinger-Moreira, Gabriel</mods:namePart>
<mods:affiliation id="B5AD3BB7F19692DA347901B342F54D02">Laboratório de Diversidade de Aracnídeos, Universidade Federal do Rio de Janeiro, Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundão, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:namePart id="99674540CBEEE44A7F1904EC42461026">de Souza Castanheira, Pedro</mods:namePart>
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<mods:affiliation id="0DA0231C69D0F4B1B3975615F5205481">Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Laboratório de Diversidade de Aracnídeos, Universidade Federal do Rio de Janeiro, Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundão, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:namePart id="72BC1CD9EAEC05C9EAE6269466EFAF44">Baptista, Renner Luiz Cerqueira</mods:namePart>
<mods:affiliation id="020B326F24EB3A0A6B440B4C7C42C884">Laboratório de Diversidade de Aracnídeos, Universidade Federal do Rio de Janeiro, Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundão, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<subSubSection id="B2E9EBE5CCCABDE15DEA3A003FFAB3B3" type="nomenclature">
<paragraph id="8BE63168850246DFB07D8D609AD5284A">
<taxonomicName id="72DC8FEC699CE7DA17F8C158C602BB79" ID-CoL="4TR4" authority="F. O. Pickard-Cambridge, 1896" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="F35E445B32EE72A1C562024E1591E37E" italics="true">Harmonicon</emphasis>
F. O.
<bibRefCitation id="4C538407ED605B2685A0218CAEE9CE46" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship &quot; Faraday &quot; up the river Amazons. Proceedings of the zoological Society of London 1896: 716 - 766. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03076. x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge, 1896</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5KAC" type="type_taxon">
<paragraph id="6E432BB1406FC7B4536D8058E3CA0D4C">
<heading id="4128D0A7FA50751E4DDA9B1B776C8898" reason="title">Type species.</heading>
</paragraph>
<paragraph id="B5FD654538F6AF95A058D028F14CFCE0">
<taxonomicName id="458C59130BB1CD2CE6303493E9B6600E" authority="F. O. Pickard-Cambridge, 1896" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="DA03D09BE8A2CCC94F863F8BBE87CD2D" italics="true">Harmonicon rufescens</emphasis>
F. O.
<bibRefCitation id="5330A120BD12D1E23ED51920D6B2F293" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship &quot; Faraday &quot; up the river Amazons. Proceedings of the zoological Society of London 1896: 716 - 766. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03076. x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge, 1896</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPLAC" type="taxon_list">
<paragraph id="73F8B0AABAC556F51A16FD5B6D25F612">
<heading id="2F848FF2BD7BF8F4E075ED26A31A4AF5" reason="title">Composition.</heading>
</paragraph>
<paragraph id="F35523DE11416CB87E5862DF0702ECFE">
Five species:
<taxonomicName id="695462DAB659CEF92BFFCC9B75D72352" authority="Marechal &amp; Marty, 1998" authorityName="Marechal &amp; Marty" authorityYear="1998" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="audeae">
<emphasis id="6DFFBC55F2099A7F0B55CA2BA4F31A91" italics="true">Harmonicon audeae</emphasis>
<bibRefCitation id="C6F286F7004CBFF4718CCA0FD07E7608" DOI="10.5962/p.268900" author="Maréchal P &amp; Marty C" journalOrPublisher="Zoosystema" pagination="499-504" refId="B9" refString="Marechal P, Marty C (1998) Rehabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espece de Guyane francaise (Araneae, Mygalomorphae, Dipluridae). Zoosystema 20: 499 - 504. https: // doi. org / 10.5962 / p. 268900" title="Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae)." volume="20" year="1998">Maréchal &amp; Marty, 1998</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="F247EE259A7FF9DCD72C2D1C26D78D29" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="2D1285BBD267E07E468FAB23F67A58BA" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AED65D0FCC41789D18763DC05558264E" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="99AA2A360B86E55F3F1061F4858186C0" authority="Pedroso &amp; Baptista, 2014" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="194A00CE44AEC8904B741A19CDC77E5A" italics="true">H. cerberus</emphasis>
<bibRefCitation id="D87EF88AE3D19C21C913AA3E47FDEA24" DOI="10.3897/zookeys.389.6693" author="Pedroso DR &amp; Baptista RLC" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Para, Brazil, with notes on the genus. ZooKeys 389: 77 - 88. https: // doi. org / 10.3897 / zookeys. 389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso &amp; Baptista, 2014</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="A0CCAF6A45C22851A0CC263213979C13" authority="F. O. Pickard-Cambridge, 1896" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="6D0112C233B6CB88B63DE4292477AE23" italics="true">H. rufescens</emphasis>
F. O.
<bibRefCitation id="45130E18A53FD2EC0CF7324DF3025F31" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship &quot; Faraday &quot; up the river Amazons. Proceedings of the zoological Society of London 1896: 716 - 766. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03076. x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge, 1896</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="C23D2518E04F9C639F14746AA29255C3" authority="Drolshagen &amp; Backstam, 2011" authorityName="Drolshagen &amp; Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="098859B56E35E47C2ABFB5DC49A6A6CC" italics="true">H. oiapoqueae</emphasis>
<bibRefCitation id="4A40DA76A9A3F170D9CA8C1FC6984642" DOI="10.3897/zookeys.112.1205" author="Drolshagen B &amp; Bäckstam CM" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Backstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 89 - 96. https: // doi. org / 10.3897 / zookeys. 112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen &amp; Bäckstam, 2011</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMNAC" type="diagnosis">
<paragraph id="386E7BF69F1BC881B4E570675C0E070F">
<heading id="B233F4C05A27C12EA3C0E4F1C4928236" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="4EEAEB4D12720DBEC400A102384AA355">
<taxonomicName id="E7FC3DD5D6C213FC31956FF76A014318" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="138521EFC95B195A480189032CD566A2" italics="true">Harmonicon</emphasis>
</taxonomicName>
and the other three lyrate genera of
<taxonomicName id="5B728075CF985791439CBB3491B5B47C" authorityName="Simon" authorityYear="1889" class="Arachnida" family="Dipluridae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="subFamily" subFamily="Diplurinae">Diplurinae</taxonomicName>
(i. e.,
<taxonomicName id="75BB06053C5721EAD1D35EBEA98567E8" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="EA51035385F90F18EE0A6CB162DF4285" italics="true">Diplura</emphasis>
</taxonomicName>
,
<taxonomicName id="933C517AB3B74A4B97571566CB00BFFA" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Dipluridae" genus="Trechona" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="82EF92620F83F1ECD737918639A186BB" italics="true">Trechona</emphasis>
</taxonomicName>
and
<emphasis id="447992CCFC4EB0779F41C9F8FE8915FC" italics="true">Harpathele</emphasis>
) can be separated from
<taxonomicName id="BD4181FAE35FCB47D013FA64111B2013" authorityName="Karsch" authorityYear="1879" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="C02B7B9D0E57ACDD44DAEAC8F1A4A297" italics="true">Linothele</emphasis>
</taxonomicName>
, by the presence of lyra on the ventral side of each maxilla (Fig.
<figureCitation id="92CDBD1EA213AC977E626CA0A6F15E81" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
;
<bibRefCitation id="0B3EAB119DDCA1FD654DB5223A982F2F" DOI="10.3897/zookeys.771.24921" author="Pedroso" etAl="et al." firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="57-71" refId="B11" refString="Pedroso DR, Giupponi APdL, Baptista RLC (2018) Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae). ZooKeys 771: 5771. https://doi.org/10.3897/zookeys.771.24921" title="Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae)." volume="771" year="2018">Pedroso et al. 2018</bibRefCitation>
, fig. 9;
<bibRefCitation id="FF0F465B068D8EA0F8D9CA967D1B9EEC" DOI="10.1111/aen.12364" author="Pedroso" etAl="et al." firstAuthor="Pedroso" issue="1" journalOrPublisher="Austral Entomology" pagination="204-219" refId="B13" refString="Pedroso DR, De Miranda GS, Baptista RLC (2019) Further advancements in the taxonomy of Trechona spiders (Araneae, Dipluridae): two new species, redescription of Trechona uniformis Mello-Leitão, 1935, additions to descriptions of Trechona rufa Vellard, 1924 and Trechona venosa (Latreille, 1832) and key to the species. Austral Entomology 58 (1): 204219. https://doi.org/10.1111/aen.12364" title="Further advancements in the taxonomy of Trechona spiders (Araneae, Dipluridae): two new species, redescription of Trechona uniformis Mello-Leitão, 1935, additions to descriptions of Trechona rufa Vellard, 1924 and Trechona venosa (Latreille, 1832) and key to the species." volume="58" year="2019">Pedroso et al. 2019</bibRefCitation>
, fig. 3 A H;
<bibRefCitation id="5FFC0CDCA48A0BC7291FDD70A1639608" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, figs 1 C, 3 G).
<taxonomicName id="8FAD804A96B3C5F591D8D35FE5E37BE8" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="74E2F1E3288C00A05040267CC54C72BE" italics="true">Harmonicon</emphasis>
</taxonomicName>
,
<taxonomicName id="92324CDF8C71E0F16F9D6006EAFB9ECA" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="E3685E36CED534860B4A6CFD8A3C7813" italics="true">Diplura</emphasis>
</taxonomicName>
and
<emphasis id="7376B7629F26B9C2C74E2BBA0A6825DD" italics="true">Harpathele</emphasis>
differ from
<taxonomicName id="2C5B75A5AE2F18A274AC9099033B3F86" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Dipluridae" genus="Trechona" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="F175CEAC1830B29279AE635690C452A6" italics="true">Trechona</emphasis>
</taxonomicName>
by having lyra with less than 35 macrosetae in a unique row (Fig.
<figureCitation id="BDCF9E25E537484397F039EDDBD00138" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
;
<bibRefCitation id="1F8306D0F4D566935E214291D30F6C32" DOI="10.3897/zookeys.771.24921" author="Pedroso" etAl="et al." firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="57-71" refId="B11" refString="Pedroso DR, Giupponi APdL, Baptista RLC (2018) Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae). ZooKeys 771: 5771. https://doi.org/10.3897/zookeys.771.24921" title="Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae)." volume="771" year="2018">Pedroso et al. 2018</bibRefCitation>
, fig. 9;
<bibRefCitation id="5669644A98E74D6EFA7C2E12B9A5789E" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, figs 1 C, 3 G) (vs. more than 35 setae arranged in a plate with several rows;
<bibRefCitation id="0871C84B0659932C9FB5180321D318A1" DOI="10.1111/aen.12364" author="Pedroso" etAl="et al." firstAuthor="Pedroso" issue="1" journalOrPublisher="Austral Entomology" pagination="204-219" refId="B13" refString="Pedroso DR, De Miranda GS, Baptista RLC (2019) Further advancements in the taxonomy of Trechona spiders (Araneae, Dipluridae): two new species, redescription of Trechona uniformis Mello-Leitão, 1935, additions to descriptions of Trechona rufa Vellard, 1924 and Trechona venosa (Latreille, 1832) and key to the species. Austral Entomology 58 (1): 204219. https://doi.org/10.1111/aen.12364" title="Further advancements in the taxonomy of Trechona spiders (Araneae, Dipluridae): two new species, redescription of Trechona uniformis Mello-Leitão, 1935, additions to descriptions of Trechona rufa Vellard, 1924 and Trechona venosa (Latreille, 1832) and key to the species." volume="58" year="2019">Pedroso et al. 2019</bibRefCitation>
, fig. 3 A H), spinnerets usually longer than abdomen (Fig.
<figureCitation id="841657836034244A778E6564574F31A6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
;
<bibRefCitation id="BD9DCEFA0A9601B96EF4567A4075BB96" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, figs 5 A) (vs. reaching up to ½ abdomen length on females and ⅔ on males;
<bibRefCitation id="B6A6C4665D5FCD4DF9C19F7EE4798DE7" DOI="10.1111/aen.12364" author="Pedroso" etAl="et al." firstAuthor="Pedroso" issue="1" journalOrPublisher="Austral Entomology" pagination="204-219" refId="B13" refString="Pedroso DR, De Miranda GS, Baptista RLC (2019) Further advancements in the taxonomy of Trechona spiders (Araneae, Dipluridae): two new species, redescription of Trechona uniformis Mello-Leitão, 1935, additions to descriptions of Trechona rufa Vellard, 1924 and Trechona venosa (Latreille, 1832) and key to the species. Austral Entomology 58 (1): 204219. https://doi.org/10.1111/aen.12364" title="Further advancements in the taxonomy of Trechona spiders (Araneae, Dipluridae): two new species, redescription of Trechona uniformis Mello-Leitão, 1935, additions to descriptions of Trechona rufa Vellard, 1924 and Trechona venosa (Latreille, 1832) and key to the species." volume="58" year="2019">Pedroso et al. 2019</bibRefCitation>
, fig. 4 A, B), fovea slightly transverse (Fig.
<figureCitation id="F7CC9C1AB1F4FD28AA0725138BD33FF8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
;
<bibRefCitation id="1007CE05E375E8E7CB60A8A47B8448AC" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, fig. 1 A) (vs. fovea strongly recurved;
<bibRefCitation id="2A5EEB2945F61747EAFEDE612D6A619C" DOI="10.1111/aen.12364" author="Pedroso" etAl="et al." firstAuthor="Pedroso" issue="1" journalOrPublisher="Austral Entomology" pagination="204-219" refId="B13" refString="Pedroso DR, De Miranda GS, Baptista RLC (2019) Further advancements in the taxonomy of Trechona spiders (Araneae, Dipluridae): two new species, redescription of Trechona uniformis Mello-Leitão, 1935, additions to descriptions of Trechona rufa Vellard, 1924 and Trechona venosa (Latreille, 1832) and key to the species. Austral Entomology 58 (1): 204219. https://doi.org/10.1111/aen.12364" title="Further advancements in the taxonomy of Trechona spiders (Araneae, Dipluridae): two new species, redescription of Trechona uniformis Mello-Leitão, 1935, additions to descriptions of Trechona rufa Vellard, 1924 and Trechona venosa (Latreille, 1832) and key to the species." volume="58" year="2019">Pedroso et al. 2019</bibRefCitation>
, fig. 4 A C) and medium-sized to large adult body size (females at most
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, males
<quantity id="FF775AECF6B90CF21B49648A61A67EE7" metricMagnitude="-2" metricUnit="m" metricValue="2.0" unit="mm" value="20.0">20 mm</quantity>
) (Figs
<figureCitation id="393B5F6EB870F7D12F6FF39106E80900" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
,
<figureCitation id="C629347F5528EC53F1A8AD185D66B164" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 A</figureCitation>
;
<bibRefCitation id="F12D0FE018F42F101EAF77A4B665C4BA" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, figs 1 B, 3 B) (vs. large adults, females at least
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, males
<quantity id="C500602A8F24A988D0FC36ADCFBFC538" metricMagnitude="-2" metricUnit="m" metricValue="2.3" unit="mm" value="23.0">23 mm</quantity>
;
<bibRefCitation id="B1DED951E73C52FF8DE80AC11ED5094D" DOI="10.1111/aen.12364" author="Pedroso" etAl="et al." firstAuthor="Pedroso" issue="1" journalOrPublisher="Austral Entomology" pagination="204-219" refId="B13" refString="Pedroso DR, De Miranda GS, Baptista RLC (2019) Further advancements in the taxonomy of Trechona spiders (Araneae, Dipluridae): two new species, redescription of Trechona uniformis Mello-Leitão, 1935, additions to descriptions of Trechona rufa Vellard, 1924 and Trechona venosa (Latreille, 1832) and key to the species. Austral Entomology 58 (1): 204219. https://doi.org/10.1111/aen.12364" title="Further advancements in the taxonomy of Trechona spiders (Araneae, Dipluridae): two new species, redescription of Trechona uniformis Mello-Leitão, 1935, additions to descriptions of Trechona rufa Vellard, 1924 and Trechona venosa (Latreille, 1832) and key to the species." volume="58" year="2019">Pedroso et al. 2019</bibRefCitation>
, fig. 4 A C).
<taxonomicName id="AC950FF6E5DDB3E3D2764BC46B746190" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="04EEF44B015C41E27874FBBB5A4B2F38" italics="true">Harmonicon</emphasis>
</taxonomicName>
and
<emphasis id="48B09C8F59F10284F16057313FFCD8A4" italics="true">Harpathele</emphasis>
differ from
<taxonomicName id="EB71A116D6EAB18FF598EF4F2FEAD980" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="A9C497A9C00713E685C43AE4F3654BB6" italics="true">Diplura</emphasis>
</taxonomicName>
by the maxilla of pedipalp with a dorsal transverse suture bearing a strong curve at its basal third (Fig.
<figureCitation id="CD27C505A86CA160AA4EE2AA67430F40" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
;
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, fig. 1 C) (vs. straight or slightly curved at its basal third;
<bibRefCitation id="F5239579B2F5A30C246F18330DBC4F63" DOI="10.3897/zookeys.771.24921" author="Pedroso" etAl="et al." firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="57-71" refId="B11" refString="Pedroso DR, Giupponi APdL, Baptista RLC (2018) Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae). ZooKeys 771: 5771. https://doi.org/10.3897/zookeys.771.24921" title="Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae)." volume="771" year="2018">Pedroso et al. 2018</bibRefCitation>
, fig. 9); a field of numerous long and thin setae between the lyra and the transverse suture (Fig.
<figureCitation id="9BF206D11454D3F4DB5A8C04B2F7D379" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
;
<bibRefCitation id="55E1CC070CB05BA05433D0CF9EAC6344" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, fig. 1 C) (vs. no or just few interspersed short thin setae;
<bibRefCitation id="233C135FB112F22D200C5FDFD08B0D75" DOI="10.3897/zookeys.771.24921" author="Pedroso" etAl="et al." firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="57-71" refId="B11" refString="Pedroso DR, Giupponi APdL, Baptista RLC (2018) Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae). ZooKeys 771: 5771. https://doi.org/10.3897/zookeys.771.24921" title="Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae)." volume="771" year="2018">Pedroso et al. 2018</bibRefCitation>
, fig. 9); and a field of small rigid bristles (spiniform setae) below the same suture (Fig.
<figureCitation id="A291C7DD5B2ACEE3F78FD9AE8802FE84" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
;
<bibRefCitation id="BF8C47ED8ACF5A5CFA26B761436110E5" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, fig. 1 C) (vs. no rigid bristles;
<bibRefCitation id="B27FBB923079FF113C18A68379B295D1" DOI="10.3897/zookeys.771.24921" author="Pedroso" etAl="et al." firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="57-71" refId="B11" refString="Pedroso DR, Giupponi APdL, Baptista RLC (2018) Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae). ZooKeys 771: 5771. https://doi.org/10.3897/zookeys.771.24921" title="Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae)." volume="771" year="2018">Pedroso et al. 2018</bibRefCitation>
, fig. 9). Furthermore,
<taxonomicName id="09C4C01A01E897E7E2B17C8EDA9EA368" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="A1EA5669C59C9B6B52242F662E4BBF9F" italics="true">Harmonicon</emphasis>
</taxonomicName>
and
<emphasis id="B19E4A51BB4DCEBBAAD4C88F178CB581" italics="true">Harpathele</emphasis>
males do not bear a field of rigid macrosetae on the prolateral face of metatarsus I and have the tibia of pedipalp long and thin (Fig.
<figureCitation id="E3AA3A7E132AA64C752C1AEE56DB97C8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 B</figureCitation>
;
<bibRefCitation id="751F50B068A672E00AF92F76150E2970" DOI="10.3897/zookeys.771.24921" author="Pedroso" etAl="et al." firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="57-71" refId="B11" refString="Pedroso DR, Giupponi APdL, Baptista RLC (2018) Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae). ZooKeys 771: 5771. https://doi.org/10.3897/zookeys.771.24921" title="Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae)." volume="771" year="2018">Pedroso et al. 2018</bibRefCitation>
, fig. 22) (vs. with such field or clasper and tibia of pedipalp clearly inflated and relatively short;
<bibRefCitation id="BA37E99DF65D9420FD741491BB3328F0" DOI="10.3897/zookeys.771.24921" author="Pedroso" etAl="et al." firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="57-71" refId="B11" refString="Pedroso DR, Giupponi APdL, Baptista RLC (2018) Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae). ZooKeys 771: 5771. https://doi.org/10.3897/zookeys.771.24921" title="Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae)." volume="771" year="2018">Pedroso et al. 2018</bibRefCitation>
, figs 14, 21).
</paragraph>
<paragraph id="5BC537BBD7DE5DF84798EC383D650B0A">
<taxonomicName id="A5E0FAFF2141B02AEA534CF8C777B690" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="937664A2CD4EB452F0C3B4EFE07E4EDA" italics="true">Harmonicon</emphasis>
</taxonomicName>
can be distinguished from
<emphasis id="1591ACA27E2CE7D08188BE16A5DE71DC" italics="true">Harpathele</emphasis>
by carapace brownish red, strongly contrasting with uniform dark grey abdomen in living specimens (Figs
<figureCitation id="FA245150F84CE9EC4302CACDB425B9FE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 G</figureCitation>
,
<figureCitation id="5B6315AAC10A869B61F99BCF7123B483" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 G</figureCitation>
) (vs. carapace brownish not strongly contrasting with abdomen colour;
<bibRefCitation id="CA0994C6551070F89B04939DB6F403A7" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, figs 1 A, 3 A), dorsum of abdomen without transverse stripes (Figs
<figureCitation id="B04BD207396CA52810486010D285396B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
,
<figureCitation id="991CC1B8A9F0C2995A44F256E0B6D1C0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 A</figureCitation>
,
<figureCitation id="91344511402A9DD0C8D3F3298AB71F1F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 A</figureCitation>
) (vs. with transverse light stripes, sometimes difficult to see in males;
<bibRefCitation id="D5B8C3A924C86DFD54CE1F8C784EB7A8" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, figs 1 B, 3 B, 4 A, 5 A), and lyra formed by 58 macrosetae relatively thick and short, each one with basis clearly separated by a gap from the other setae, and apex usually strongly curved (Fig.
<figureCitation id="F67AF5D7161FFC6C144EBBCDF755768A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
;
<bibRefCitation id="649A095973A11DB5EEDC3B3BFF23E8B6" DOI="10.5962/p.268900" author="Marechal and Marty" firstAuthor="Marechal" journalOrPublisher="Zoosystema" pagination="499-504" refId="B9" refString="Maréchal P, Marty C (1998) Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae). Zoosystema 20: 499504. https://doi.org/10.5962/p.268900" title="Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae)." volume="20" year="1998">Maréchal and Marty 1998</bibRefCitation>
, fig. 2 A B;
<bibRefCitation id="4201105B0AC0F7BE76068BBC62A41266" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 6;
<bibRefCitation id="05A8ED15B1DA486A70BF07C975176E68" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
, fig. 6) (except
<taxonomicName id="C871B24B756203E2BE9DCA2AC341291F" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="A3E107AFE059CC92F0A325A07C0A0407" italics="true">Harmonicon candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA91A36F9D91583F60133712A1966636" rank="species">sp. nov.</taxonomicNameLabel>
, figs 2 H, 3 C) (vs. 2531 macrosetae, usually long, with bases closely placed and apex almost straight or weakly curved, but with some clavate tips in females;
<bibRefCitation id="F4181E17362EA883571E59B46C9A570A" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, figs 1 C, 3 G, 4 B, 5 G, 6 A D). Furthermore,
<taxonomicName id="4950EAC697DCD05600B2466744B7CA0B" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="8A303448E1BDD416BD8A73124E99AE99" italics="true">Harmonicon</emphasis>
</taxonomicName>
females have spermathecae formed by two branches rising from one base (Figs
<figureCitation id="2A215567AB3D671CF092F5E8192C4474" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 D F</figureCitation>
,
<figureCitation id="DBE9C3C0E65B91419B36A7797312B358" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 D F</figureCitation>
;
<bibRefCitation id="BB323777742AAA53874457F70CAF8CDF" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 5) (vs. one long, curved, and flattened branch, sometimes with a small round lateral vesicle or lobule;
<bibRefCitation id="6C215CA4E94AD13B006756E43764B440" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, figs 2 A D, 4 D), and males have embolus reaching between 1.1 and 1.5 as long as the palpal bulb in retrolateral view (Fig.
<figureCitation id="3DD07A01C699A7DFBB84E4BC6C83500C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 B, D</figureCitation>
;
<bibRefCitation id="BDA294F39296392A524A6D705A713012" DOI="10.5962/p.268900" author="Marechal and Marty" firstAuthor="Marechal" journalOrPublisher="Zoosystema" pagination="499-504" refId="B9" refString="Maréchal P, Marty C (1998) Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae). Zoosystema 20: 499504. https://doi.org/10.5962/p.268900" title="Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae)." volume="20" year="1998">Maréchal and Marty 1998</bibRefCitation>
, fig. 4;
<bibRefCitation id="73F08B1B7372DA0053CB643236891CC1" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 1;
<bibRefCitation id="DBB4BC12B48C1BDB33EB3CC47D59AA57" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
, fig. 7) (vs. 2 or more times longer than bulb;
<bibRefCitation id="072A540239A4E8244829EB13B7B5BB38" DOI="10.13156/arac.2024.19.8.1060" author="Wermelinger-Moreira" etAl="et al." firstAuthor="Wermelinger-Moreira" issue="8" journalOrPublisher="Arachnology" pagination="1060-1073" refId="B18" refString="Wermelinger-Moreira G, Pedroso DR, Castanheira PdS, Baptista RLC (2024) A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species. Arachnology 19 (8): 10601073. https://doi.org/10.13156/arac.2024.19.8.1060" title="A new genus of lyrate curtain-web spiders (Araneae, Mygalomorphae, Dipluridae) from southeastern Brazil, with two new species and revalidation of a formerly described species." volume="19" year="2024">Wermelinger-Moreira et al. 2024</bibRefCitation>
, figs 3 C, F, 5 E).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIIAE" type="Taxonomic notes">
<paragraph id="03EFDF7106903EA08FBF07714516B93F">
<heading id="FCC318F097459908BC049E0C572A5451" reason="title">Taxonomic notes.</heading>
</paragraph>
<paragraph id="FDDB8882046D3AE937FCED32091F0613">
A comparative diagnosis of
<taxonomicName id="7ABFF4012C67F53E3EF5EF4BE7202E25" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="6E79BB6D14D4C7CBEC4AF5932DC82F84" italics="true">Harmonicon</emphasis>
</taxonomicName>
and other genera of
<taxonomicName id="04D671AED3843D556CDC256CEB5A23EA" authorityName="Simon" authorityYear="1889" class="Arachnida" family="Dipluridae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="family">Dipluridae</taxonomicName>
was published by
<bibRefCitation id="127292D1ACE5FB8889BE485FA48D8453" DOI="10.3897/zookeys.771.24921" author="Pedroso DR &amp; Giupponi APdL &amp; Baptista RLC" etAl="et al." firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="57-71" refId="B11" refString="Pedroso DR, Giupponi APdL, Baptista RLC (2018) Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae). ZooKeys 771: 5771. https://doi.org/10.3897/zookeys.771.24921" title="Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae)." volume="771" year="2018">Pedroso et al. (2018)</bibRefCitation>
. Since then,
<emphasis id="C78FEA7D30ACA4C55A5110C4D114A79F" italics="true">Harpathele</emphasis>
was added to the subfamily, and some updates were made to the diagnosis presented above.
</paragraph>
<paragraph id="9BE4D37B0E2513FC68599FF3A756BFC5">
All described
<taxonomicName id="B6222647CFCB3B030545043D50AFED37">
<emphasis id="E33FC511BC1B4CFF402C237E7AD5DDC1" italics="true">Harmonicon</emphasis>
species
</taxonomicName>
have colour pattern and somatic traits generally similar and less variable, with the exception of
<taxonomicName id="364C24195F73BF4DBE5484E22E490223" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="809179E8E0AC592E1B69F47E8B05A796" italics="true">H. cerberus</emphasis>
</taxonomicName>
, which exhibits some troglomorphic characters, such as the fused and reduced lateral eyes and the pale yellow carapace (
<bibRefCitation id="AA6F74AA6CA62CED652A296743919580" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
, fig. 13). Most diagnostic characters are related to the genitalia or to sexual dimorphic traits.
</paragraph>
<paragraph id="938F5BF725166E2D93DBDDA1D9E5C039">
An interesting diagnostic somatic character is the shape of the sternal sigilla III.
<taxonomicName id="492715E49C6A95AE1184B593C866B003" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="E03C41352FDB0221D5BBA000371A2066" italics="true">Harmonicon rufescens</emphasis>
</taxonomicName>
,
<taxonomicName id="CE8CB2E4E07953002F910A45FB1F255F" authorityName="Drolshagen &amp; Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="9924AD1D71810E90ADA0A18F36525DC0" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
, and
<taxonomicName id="B523FA9240BAF411F7A0E1BDAA0A9953" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="7CFAB2077EBCB68D86D0A1EF5C34AAAE" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BC1040D37F70A5E4C5FB2C1611C30F5B" rank="species">sp. nov.</taxonomicNameLabel>
share sternal sigilla III with similar longer than wide ellipsoid shape (Figs
<figureCitation id="86B16C312032EC5B1FA01641BC5DEF68" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 B</figureCitation>
,
<figureCitation id="40F97AC2ECD30C28906B30C84FA58F42" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 G</figureCitation>
,
<figureCitation id="E4EEDF6BC7DA0A500DD4B48BB38289F9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 B</figureCitation>
;
<bibRefCitation id="9E08664343732EB718F9D57B9D9D4BA0" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 3). Those sigilla strongly contrast with the rounded (slightly longer than wide) sigilla of
<taxonomicName id="E7BEDA77CF38537848369B681A53B415" authorityName="Marechal &amp; Marty" authorityYear="1998" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="audeae">
<emphasis id="D3F1294E38CB8E69D3BA0A7E3AD1319F" italics="true">H. audeae</emphasis>
</taxonomicName>
(
<bibRefCitation id="6B4B5B2E637317B170C780EE5A9B6B14" DOI="10.5962/p.268900" author="Marechal and Marty" firstAuthor="Marechal" journalOrPublisher="Zoosystema" pagination="499-504" refId="B9" refString="Maréchal P, Marty C (1998) Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae). Zoosystema 20: 499504. https://doi.org/10.5962/p.268900" title="Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae)." volume="20" year="1998">Maréchal and Marty 1998</bibRefCitation>
, fig. 1 C). As noted by
<bibRefCitation id="34BC9A0EF925B9C29E4D8C6710ED6813" DOI="10.3897/zookeys.771.24921" author="Pedroso DR &amp; Giupponi APdL &amp; Baptista RLC" etAl="et al." firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="57-71" refId="B11" refString="Pedroso DR, Giupponi APdL, Baptista RLC (2018) Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae). ZooKeys 771: 5771. https://doi.org/10.3897/zookeys.771.24921" title="Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae)." volume="771" year="2018">Pedroso et al. (2018)</bibRefCitation>
, some species of
<taxonomicName id="CF8A5A5765B8522AC9BFC804601A1271" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="DFF8960D46E158E2951CB2A87532EC33" italics="true">Harmonicon</emphasis>
</taxonomicName>
do not bear a lyra with hook-shaped macrosetae. Instead, they are clavate with a long and weaker bent beginning at the median third of the seta, as in
<taxonomicName id="D222C0B4439085A7C5D460FB57ECE2CD" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="BC421F463AC8E1FAD72DEA8C3F8B3C10" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F750AB4EC38F6E78067CAFD9EDBDFEA" rank="species">sp. nov.</taxonomicNameLabel>
(Fig.
<figureCitation id="C9686FBF54CAC399FCE36C52FC9D0866" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 H</figureCitation>
,
<figureCitation id="BE010F9D923E888908AD8BE395109BED" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 C</figureCitation>
). Additional thickened and more sclerotised setae on the maxillae may represent a diagnostic character for at least some species of
<taxonomicName id="B3E21B32AF447F798507A21924090B9C" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="EA24736A8D87F54855247F303BDCC985" italics="true">Harmonicon</emphasis>
</taxonomicName>
. They are represented by three to five stiff setae placed at the distal half of the maxilla, away from the most basally placed lyra, and at the beginning of the large field of elongated and thin setae that cover its prolateral face, as in
<taxonomicName id="04D2698345BD09697B2882463F54FA21" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="CBBCCD37E450D9CE6C1589C7BE9104B9" italics="true">H. cerberus</emphasis>
</taxonomicName>
(
<bibRefCitation id="91608C4F62F6C76332636589A1B5214B" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
, fig. 6),
<taxonomicName id="79D53B2613DA2C012E6C837161C94C34" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="7E1F3B6DF7EA87098B30314C73CEB98D" italics="true">H. rufescens</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="F55CF648302DE23E7A7F43F81B6D886D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
), and also in
<taxonomicName id="6875AEECA6C5E04D4249D3CCA7FF0621" authority="(Pedroso et al. 2018)" baseAuthorityName="Pedroso" baseAuthorityYear="2018" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="6E8CDE7B78F5FEECBF15A757A446F723" italics="true">H. oiapoqueae</emphasis>
(
<bibRefCitation id="3C506E4570D34B15226AFC8C32365081" DOI="10.3897/zookeys.771.24921" author="Pedroso" etAl="et al." firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="57-71" refId="B11" refString="Pedroso DR, Giupponi APdL, Baptista RLC (2018) Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae). ZooKeys 771: 5771. https://doi.org/10.3897/zookeys.771.24921" title="Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae)." volume="771" year="2018">Pedroso et al. 2018</bibRefCitation>
)
</taxonomicName>
. In females, the tarsal claw of the pedipalp has double row of teeth in
<taxonomicName id="701280136953E5F60D1531C3130B6360" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="EC1518E258A9FEB08E3948DA92DA7744" italics="true">H. rufescens</emphasis>
</taxonomicName>
,
<taxonomicName id="C2D62AB681C40571BD4598D0053348B1" authorityName="Marechal &amp; Marty" authorityYear="1998" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="audeae">
<emphasis id="7F3DB9A032D25C1D39E948F1DE2713DA" italics="true">H. audeae</emphasis>
</taxonomicName>
, and
<taxonomicName id="C0C6C97F5A4707E474ED71FC00BFEACA" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="E0BAE6FA29283BC33DC865246B83F3BD" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4ABC0D24EE4FDEAC1BB0B8F6B67E679" rank="species">sp. nov.</taxonomicNameLabel>
, but only one row of teeth in
<taxonomicName id="DA54584026D2CD8D49427103C7FA8D4E" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="7C4CA3B420025D9FFD9F722351940EC7" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
(see
<bibRefCitation id="791F257E359E47726EDEC7B06851054D" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, p. 92).
</paragraph>
<paragraph id="6E9610EDCD318914927F37827A98B186">
The most useful sexual characters in males are those from the palpal bulb and modified tibia and metatarsus of leg I. Their tibia I bears one distal prolateral macroseta besides the retrolateral spur. In
<taxonomicName id="6AE989F30B69335FD65B73A6B5199BB7" authorityName="Marechal &amp; Marty" authorityYear="1998" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="audeae">
<emphasis id="BA930579CAB4FD220BBAFBD7BBE9CCA8" italics="true">H. audeae</emphasis>
</taxonomicName>
, the prolateral macroseta is thin and elongated (
<bibRefCitation id="083BC574C4B4ADA1EFB277CBAEC4442F" DOI="10.5962/p.268900" author="Marechal and Marty" firstAuthor="Marechal" journalOrPublisher="Zoosystema" pagination="499-504" refId="B9" refString="Maréchal P, Marty C (1998) Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae). Zoosystema 20: 499504. https://doi.org/10.5962/p.268900" title="Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae)." volume="20" year="1998">Maréchal and Marty 1998</bibRefCitation>
, fig. 3 C, D). In
<taxonomicName id="006958B4A04B67F86664C3F17D30707D" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="6FF48C213A02E2AD0621D7AA039B080B" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="63A1A2DD1846B71EB8A6FD5FCC21CC79" rank="species">sp. nov.</taxonomicNameLabel>
, that macroseta is lost, but the dark insertion plate is clearly visible (Fig.
<figureCitation id="D1641EE1A575076365A1F8C7844B0913" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 J</figureCitation>
), and in
<taxonomicName id="DACBB3A2872F0C0561260A90E90632D1" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="BC32BE57903025FB49EE941FF2DEA7B1" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
, a similar plate seems to be present in an Internet photograph taken by Droslhagen, but it is not mentioned or illustrated in the description (
<bibRefCitation id="C2F19DE1803DB32A75D12D0CC9892A2C" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 7). The only species apparently without such macroseta is
<taxonomicName id="D931341B46C0155CCEBFB0285727946B" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="3F886D1E723BCFED56A0611DE52E4DC8" italics="true">H. cerberus</emphasis>
</taxonomicName>
(
<bibRefCitation id="19557924863997C6928B4F1B99F25E19" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
, fig. 11), but its
<typeStatus id="C8888B758E3EFD5F891AB2EC4C8DA407" type="holotype">holotype</typeStatus>
has been destroyed during the
<named-content id="DC05DC337BD50F392306DA1372D22CB8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">
<collectionCode id="D506C6C57CFADC36DFF965ACC85D1393" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" type="">MNRJ</collectionCode>
</named-content>
s fire. The distal retrolateral spur of tibia I is similar in all described species, projected like a small, weakly curved rod in retrolateral view and slightly projected laterally in ventral view. The distal megaspine over the spur is a little more variable in both thickness and curvature. It is more elongate and curved and has a basis slightly thicker in
<taxonomicName id="5F19C7CBC386C5C2069A2A6574BB1199" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="128FB9FCEB11997A739CD89E85647E4D" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
(see
<bibRefCitation id="5FFA43BC4164DF03BB80705478732469" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 7) and
<taxonomicName id="C3367DD26BD462777AAB1452112ECE6B" authorityName="Marechal &amp; Marty" authorityYear="1998" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="audeae">
<emphasis id="5AE7246EAAF37977F7C5E0745CD0AA94" italics="true">H. audeae</emphasis>
</taxonomicName>
(see
<bibRefCitation id="E4B12CE9B4B1E53B9984D56261AAADD1" DOI="10.5962/p.268900" author="Marechal and Marty" firstAuthor="Marechal" journalOrPublisher="Zoosystema" pagination="499-504" refId="B9" refString="Maréchal P, Marty C (1998) Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae). Zoosystema 20: 499504. https://doi.org/10.5962/p.268900" title="Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae)." volume="20" year="1998">Maréchal and Marty 1998</bibRefCitation>
, fig. 3 C, D) than in
<taxonomicName id="79E1C1740A95C5CADF334EE4F434A624" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="831EA6F417987DF386C785F4C11A4D60" italics="true">H. cerberus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="AAFB2CC5343B31EA6AA7FD8290C6DA13" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
, fig. 11) and
<taxonomicName id="61E118FEF14F09B6D23AA73B48937BDD" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="C84BB8C15000A82246536C973C0FC5BD" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A059AED88744EA0C19FC4819E7636728" rank="species">sp. nov.</taxonomicNameLabel>
(Fig.
<figureCitation id="A2BFA8B583DB73D36296B0A2570C5A4D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 I, J</figureCitation>
).
</paragraph>
<paragraph id="12BC7DF36583CAEFBE28CD315DCE7E54">
In relation to metatarsus I, most species have a retrolateral tubercle at its basal third, but it is absent in
<taxonomicName id="B1163BAABB5C78AA6788CAB87B85AE59" authorityName="Marechal &amp; Marty" authorityYear="1998" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="audeae">
<emphasis id="4F0BF02A4DDD48252471D9F94AFBC6C9" italics="true">H. audeae</emphasis>
</taxonomicName>
(see
<bibRefCitation id="9111D7D0F6BB1D2C0FDAEDCF5485CF39" DOI="10.5962/p.268900" author="Marechal and Marty" firstAuthor="Marechal" journalOrPublisher="Zoosystema" pagination="499-504" refId="B9" refString="Maréchal P, Marty C (1998) Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae). Zoosystema 20: 499504. https://doi.org/10.5962/p.268900" title="Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae)." volume="20" year="1998">Maréchal and Marty 1998</bibRefCitation>
, fig. 3 D). The shape and size of the tubercle are similar in the three species where it has been recorded, where it appears as a small mound with a rounded point (Fig.
<figureCitation id="D889461C082B349C0C0524D68FBDDD04" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 J</figureCitation>
;
<bibRefCitation id="CA0087C82096F1AF228324941C448645" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 7;
<bibRefCitation id="AF93F1C3313D277CF9E4221269F397FA" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
, fig. 11). In
<taxonomicName id="F1FDED18F131A25F2B8AC8B85541E832" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="CDB6890E4365642825757F56D07E9AB9" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2821A4F16C6B5BBF254EB0FA436DFF9" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="EB04F827C52B3672FFBD84DA6C31C9C0" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="C40A216D172270700ABF448BC8CF354F" italics="true">H. cerberus</emphasis>
</taxonomicName>
, there is a retrolateral macroseta at the ventral side near the retrolateral tubercle of metatarsus I (Fig.
<figureCitation id="7FF1EEEB2D2AC5AC57AF1D4F85FB8CA7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 J</figureCitation>
;
<bibRefCitation id="475C7A6CD6B4CC1A1940D2F1782C9C5B" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
, fig. 11), which is not found in
<taxonomicName id="B935D834C2CD5F9FED9B05DEDAA8E2ED" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="76741C6FADF52EF5F923AD403199B1C5" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
(see
<bibRefCitation id="D292D735F788D999734D45BCD5089053" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 7). In
<taxonomicName id="DA1D30CC78EB117758657657584ABD61" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="E738A2F610C74923AD71190A3A6A802A" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90CE39DD34B9D351231826CC9302421F" rank="species">sp. nov.</taxonomicNameLabel>
, that macrosetae is placed at the same level of the tubercle (Fig.
<figureCitation id="5A3731F64CE1EDF21A06C7998EF42876" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 J</figureCitation>
), but it is placed basally to the tubercle in
<taxonomicName id="458AB86AA7FD0A1F400E1B801B874309" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="C0758F85F2C2EAAA7406E76F21730172" italics="true">H. cerberus</emphasis>
</taxonomicName>
(
<bibRefCitation id="FAA6F4BC8C2800660656A7F6A955C9F6" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
, fig. 11).
</paragraph>
<paragraph id="1B7B19585530AB17801566B6F58E8643">
The palpal bulb differs among the species. In
<taxonomicName id="EB4C3A3BD8DEAFDF96AA780E36C28014" authorityName="Marechal &amp; Marty" authorityYear="1998" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="audeae">
<emphasis id="13E122658CF00B45F536012200053FAD" italics="true">H. audeae</emphasis>
</taxonomicName>
, it is thin and piriform (
<bibRefCitation id="1813CF980C4B2DF97A0FE5492C710C72" DOI="10.5962/p.268900" author="Marechal and Marty" firstAuthor="Marechal" journalOrPublisher="Zoosystema" pagination="499-504" refId="B9" refString="Maréchal P, Marty C (1998) Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae). Zoosystema 20: 499504. https://doi.org/10.5962/p.268900" title="Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae)." volume="20" year="1998">Maréchal and Marty 1998</bibRefCitation>
, fig. 4 A C), while it is wider and more rounded in
<taxonomicName id="4FA0027D34AFB42624183022DCA95327" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="D144B2AB0EBB17B0475D890346C2DEEA" italics="true">H. cerberus</emphasis>
</taxonomicName>
,
<taxonomicName id="FAC26BBEB3D4B02B717A67E346F6421A" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="E31EE296A2DB0233C66F1A040E62A988" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
, and
<taxonomicName id="C2D14DBFAF50FCAF9201873220EC10B9" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="C476506911456428A6631DCB1FF3D073" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1276BB81BBB9C589BC4A925C9D0DA0E" rank="species">sp. nov.</taxonomicNameLabel>
(Fig.
<figureCitation id="1A0846B5E79137C69580CD443588D7A2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 C F</figureCitation>
;
<bibRefCitation id="440F4CF933DF23A40F607C52A67FACFB" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
, fig. 79;
<bibRefCitation id="3D5D2A6315FCFDE01592B46A15D8BED5" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 1, 2). In retrolateral view, the embolus may be about as long as (
<taxonomicName id="0DECFC2C6EB69F32188234BAF083A0F5" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="1E48C7B111296631069F5435C1788064" italics="true">H. cerberus</emphasis>
</taxonomicName>
, see
<bibRefCitation id="2EA6695DAE488F1304DEE191EFB53B84" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
, fig. 7) or a little longer (
<taxonomicName id="5E1EF63C57B17F6D643286181437397B" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="55D5B05DC278C45DD60DC72C82BBC03A" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
, ca. 1.2 ×,
<bibRefCitation id="B5F9ADD39E5370FC0E6D640A7E5D4B5D" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 2) than the bulb proper, but may also be elongated (
<taxonomicName id="5930D9C95F263C3B0B55096118211AA0" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="502093576FABD96BB2378043F27D8EAC" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4F61F5969CF4D3429E90408D33861B6A" rank="species">sp. nov.</taxonomicNameLabel>
, ca. 1, 5 × longer, Fig.
<figureCitation id="9B95FB35572D5633BFBECC6CF1C25313" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 B, D</figureCitation>
).
</paragraph>
<paragraph id="7FB3C2142EE95041D9E8A0E25BAAA496">
In females, the spermathecae and its accessory parts are very useful for diagnosis. The median branch of the spermathecae varies from about the same length as the lateral branch and ending in a rounded, blunt tip (
<taxonomicName id="C23750AC2DF78A8AD8EEA60211EE67D6" authority=", Drolshagen and Backstam 2011" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="2D57E152E84704CB223FA9F521A3B5C8" italics="true">H. oiapoqueae</emphasis>
,
<bibRefCitation id="A99DB46D6B90FF7FFD9A1E94DB8C2D71" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
</taxonomicName>
, fig. 5) to a little longer than the lateral branch and ending in a tip with 34 lobules (ex.
<taxonomicName id="9337C12500960C0F89410C19D5F8E11C" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="3B56A7A5197666687E77A15103BD08DA" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54BB776E7358D554A88C929125A3C4CD" rank="species">sp. nov.</taxonomicNameLabel>
, Fig.
<figureCitation id="057C80FD60954CB22AC21153B36F1C61" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 D F</figureCitation>
). The lateral branch may have its tip folded over its stem (ex.
<taxonomicName id="0545D78D65D2879774F10A1E9B59AD47" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="C031D5696C5BB4C0545D15C2B8251F8B" italics="true">H. rufescens</emphasis>
</taxonomicName>
, Fig.
<figureCitation id="92531C8A0CDE00842BC99A8E96DBA760" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 D F</figureCitation>
) or with several lobules. The lobulated tip may have up to 45 lobules (
<taxonomicName id="11447595399A657DB2ABBEDD9EF12DB7" authority=", Drolshagen and Backstam 2011" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="D99DF8A6023199B1694680CBDD84C45A" italics="true">H. oiapoqueae</emphasis>
,
<bibRefCitation id="25B36F2383BC451588B760A2A684CF09" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
</taxonomicName>
, fig. 5) or be very wide and multilobulate (Fig.
<figureCitation id="1087F51D379670A29BEA551969A14188" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 D F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EU3AE" type="distribution">
<paragraph id="6095D4F641C400FD5BA29197537F52D9">
<heading id="9E3FF086BC5ED03549F6963F9E9C1739" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="92E1F7A4BC00EC27203AAC7BBFAE2CF3">
The genus can be found in several localities in the Amazon region, in
<collectingCountry id="3D64CDCFB8F7A51F9CAB101FA99AF3BC" name="French Guiana">French Guiana</collectingCountry>
and
<collectingRegion id="283710A6A280B6426E54147CBF8CD161" country="Brazil" name="Para">Pará State</collectingRegion>
, North
<collectingCountry id="684804E6ECC452A1EA8B4DA0CE13B850" name="Brazil">Brazil</collectingCountry>
. Its geographical range is here extended to Brasília,
<collectingRegion id="5556BCFAB3C687507037EB4C64E9B87B" country="Brazil" name="Distrito Federal">Distrito Federal</collectingRegion>
, Central-West region of
<collectingCountry id="8347E723FB7833C1C5B3ADE01E570433" name="Brazil">Brazil</collectingCountry>
, in the Cerrado biome. There are also records of undescribed species from Amazonian
<collectingCountry id="33E56BCD522FA7BE8AE67177DEB7ED3D" name="Peru">Peru</collectingCountry>
, the Northeast region, and additional states from the Central-West region of
<collectingCountry id="D9CAD25A682C106B754553C1D0039B5B" name="Brazil">Brazil</collectingCountry>
, according to
<bibRefCitation id="A0993207DBEC78E6DFD3C77FD6519686" DOI="10.3897/zookeys.771.24921" author="Pedroso DR &amp; Giupponi APdL &amp; Baptista RLC" etAl="et al." firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="57-71" refId="B11" refString="Pedroso DR, Giupponi APdL, Baptista RLC (2018) Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae). ZooKeys 771: 5771. https://doi.org/10.3897/zookeys.771.24921" title="Comments on the genus Diplura C. L. Koch, 1850, with description of two new species (Araneae, Mygalomorphae, Dipluridae)." volume="771" year="2018">Pedroso et al. (2018)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,500 @@
<document id="0F130164282FF1746DFB1569E6DB3BE4" ID-DOI="10.3897/evolsyst.8.141847" ID-ZooBank="8E0DCC28-3448-4001-90B7-E675BC107381" ID-publisher-id="141847" URI-arpha="784085DF-996A-5FEC-89FE-F9FE7A70E5A1" URI-zoobank="http://zoobank.org/8E0DCC28-3448-4001-90B7-E675BC107381" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1732116009247" checkinUser="pensoft" docAuthor="Wermelinger-Moreira, Gabriel, de Souza Castanheira, Pedro &amp; Baptista, Renner Luiz Cerqueira" docDate="2024" docId="CD5166CF00595E44A1C1A696A30D90E4" docLanguage="en" docName="Evolutionary Systematics 8(2): 261-271" docOrigin="Evolutionary Systematics 8 (2)" docSource="https://evolsyst.pensoft.net/article/141847/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Harmonicon rufescens F. O. Pickard-Cambridge 1896" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="271" masterDocId="784085DF996A5FEC89FEF9FE7A70E5A1" masterDocTitle="A new species of Harmonicon F. O. Pickard-Cambridge, 1896, from central-west Brazil and redescription of H. rufescens F. O. Pickard-Cambridge, 1896 (Araneae, Mygalomorphae, Dipluridae)" masterLastPageNumber="271" masterPageNumber="261" pageNumber="261" updateTime="1732123585974" updateUser="admin">
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<mods:title id="614544574D5979AB0BAF07D9B59095F4">A new species of Harmonicon F. O. Pickard-Cambridge, 1896, from central-west Brazil and redescription of H. rufescens F. O. Pickard-Cambridge, 1896 (Araneae, Mygalomorphae, Dipluridae)</mods:title>
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</mods:role>
<mods:namePart id="20AB82A391163D67E31D0BA9A5CC6B8D">Wermelinger-Moreira, Gabriel</mods:namePart>
<mods:affiliation id="5E571D989EA546D95DF935BF5720BA8B">Laboratório de Diversidade de Aracnídeos, Universidade Federal do Rio de Janeiro, Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundão, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:namePart id="99870744D8AC4D85EBEB530E0EE409C3">de Souza Castanheira, Pedro</mods:namePart>
<mods:nameIdentifier id="2B41DB96AE85A53E8B84DB5D1435A245" type="ORCID">0000-0002-0623-1622</mods:nameIdentifier>
<mods:affiliation id="7D9C26A8745B4E8AE2A2B70C5533593D">Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Laboratório de Diversidade de Aracnídeos, Universidade Federal do Rio de Janeiro, Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundão, Rio de Janeiro, RJ, Brazil</mods:affiliation>
</mods:name>
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<mods:roleTerm id="1E2FC89135197164F3698D38189C72F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A824F883518691BB0C7B735A7CB3DA42">Baptista, Renner Luiz Cerqueira</mods:namePart>
<mods:affiliation id="AC652EB20C75B5664CAA2F26B2981E37">Laboratório de Diversidade de Aracnídeos, Universidade Federal do Rio de Janeiro, Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundão, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:title id="901E127CA0F1079B16D790C602E3F996">Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part id="B65C4DE5D003F35C12849139FF97A49D">
<mods:date id="7E92728350D224D2DC17CEB08B015D55">2024</mods:date>
<mods:detail id="79639DADF38D1407611CD817345D1CD1" type="pubDate">
<mods:number id="B40F8C1F5951081D86A78FD202265383">2024-11-20</mods:number>
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<mods:number id="77A196E8A320007AF9F7D88238F7BAD7">8</mods:number>
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<treatment id="CD5166CF00595E44A1C1A696A30D90E4" ID-arpha="CD5166CF-0059-5E44-A1C1-A696A30D90E4" LSID="urn:lsid:plazi:treatment:CD5166CF00595E44A1C1A696A30D90E4" httpUri="http://treatment.plazi.org/id/CD5166CF00595E44A1C1A696A30D90E4">
<subSubSection id="646AED9FA76FBCA444410666FE578974" type="nomenclature">
<paragraph id="717CF2EA41ADA6EBB823A0EA656F50F3">
<taxonomicName id="E8BF43ADC9757ABC92F60DF179CB5D8C" ID-CoL="6LCCH" authority="F. O. Pickard-Cambridge, 1896" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="E36C899B4ADC3C1022CB9FB6DFE191F9" italics="true">Harmonicon rufescens</emphasis>
F. O.
<bibRefCitation id="3121A5A5ED4589ABFA4FD3112AA4CE2E" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship &quot; Faraday &quot; up the river Amazons. Proceedings of the zoological Society of London 1896: 716 - 766. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03076. x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge, 1896</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C2497B9668D9AE37ACF8732E8A99FA1B" type="description">
<paragraph id="D33D9FD8570CEE88BB20C5403704C761">
<figureCitation id="24407B43AC4ECC5A5952AE39664E7D34" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">Figs 1 A G</figureCitation>
,
<figureCitation id="D0FBE8BA4747C1F86A321493AC4B010B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution records of H. rufescens (holotype = red and black circle, specimen here described = red circle) and H. candango sp. nov. (yellow square)." httpUri="https://binary.pensoft.net/fig/1178838">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="35BD7B2C43E834C57E29CFF9DAF36860" type="reference_group">
<paragraph id="4B40EEB6020BEA06057BA867DC2F4EFF">
<treatmentCitationGroup id="2B7D063F085C093BFCAF1FE4660248BA">
<treatmentCitation id="9155088197E9CE933D47F8C3E44301BD" author="Pickard-Cambridge FO" page="756" year="1896">
<taxonomicName id="FC953F1F97C8F2C0DAD54A9F46916E41" ID-CoL="6LCCH" authority="F. O. Pickard-Cambridge, 1896: 756" authorityName="F. O. Pickard-Cambridge" authorityPageNumber="756" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="DC9B844DD09176E8A83F847826857CAC" italics="true">Harmonicon rufescens</emphasis>
<comment id="2158AC59DB2CB90A55EBA644FB881C37">
F. O.
<bibRefCitation id="8EE1372981B8410C19B0AE4FB88C74A3" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship &quot; Faraday &quot; up the river Amazons. Proceedings of the zoological Society of London 1896: 716 - 766. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03076. x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge, 1896: 756</bibRefCitation>
, pl. 33, fig. 3, 6, pl. 35, fig. 23 (juvenile).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="55C12522A939CDA0DB41F17616C2DA3B">
<treatmentCitationGroup id="DE7EDE0F3A34F97416D2AEA5139A3B9C">
<taxonomicName id="864B29E6CAF52E4BD7178EBDDF8EAC62" authorityName="Masi" authorityYear="1917" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="2976E0A7CEAA6095773A3347F9D81123" italics="true">Diplura rufescens</emphasis>
</taxonomicName>
<comment id="BAD6E5AEB9B9AD41D0F84AB685E40602">
:
<treatmentCitation id="1D8F0EE581DC1C9301DE7D5D59FE857B" author="Raven RJ" page="153" year="1985">
<bibRefCitation id="A57D244AC35AE96BB526CB592BABE460" author="Raven RJ" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="1-180" refId="B15" refString="Raven RJ (1985) The spider infraorder Mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History 182: 1 - 180." title="The spider infraorder Mygalomorphae (Araneae): cladistics and systematics." volume="182" year="1985">Raven, 1985: 153</bibRefCitation>
</treatmentCitation>
(synonymy and transfer to
<taxonomicName id="B3878E6F17024E191B977278BDF3B33A" ID-CoL="46FP" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="DA878982C9C5A0E05DF91ECC9B6D2266" italics="true">Diplura</emphasis>
</taxonomicName>
).
</comment>
</treatmentCitationGroup>
</paragraph>
<paragraph id="984C2CEDC0297CEB6C3B02FA26FFCBC7">
<treatmentCitationGroup id="D071A8AF1E5E0A4F4C52C1446552F6C0">
<taxonomicName id="8369C5921118DF46072F4EEBCA907ED8" ID-CoL="6LCCH" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="216BA85F8862E0204654ED7F804E0126" italics="true">Harmonicon rufescens</emphasis>
</taxonomicName>
<comment id="ED85FBBC9F81747FEBF6C4A1A0192E40">
:
<treatmentCitation id="25A36FE157D3CDAA0C7CC3EA6D9AE31E" author="Maréchal P &amp; Marty C" page="500" year="1998">
<bibRefCitation id="6B987D9A1409F2A30306B7EC0ECE00EA" DOI="10.5962/p.268900" author="Maréchal P &amp; Marty C" journalOrPublisher="Zoosystema" pagination="499-504" refId="B9" refString="Marechal P, Marty C (1998) Rehabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espece de Guyane francaise (Araneae, Mygalomorphae, Dipluridae). Zoosystema 20: 499 - 504. https: // doi. org / 10.5962 / p. 268900" title="Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae)." volume="20" year="1998">Maréchal &amp; Marty, 1998: 500</bibRefCitation>
</treatmentCitation>
(rejected synonym with
<taxonomicName id="077CCE6865368996EC5CDE2ED753AF9F" ID-CoL="46FP" authorityName="C. L. Koch" authorityYear="1850" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="0829A2E6FCF6DF70DBD5055DA10A245E" italics="true">Diplura</emphasis>
</taxonomicName>
).
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E56AE" type="material">
<paragraph id="1E4FB6B79233F5A13A8033C53EA03687">
<heading id="BA6AF0B713B17B338FD2E1EE121ED432" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="F0D4B38EC4DB633BC8490DAAFFC67737">
<materialsCitation id="FDF67CDED970F018E99995252D3E6B19" collectionCode="NHMUK" collectorName="F. O. Pickard-Cambridge &amp; Austen" country="Brazil" county="lower Amazon" location="Santarem" specimenCount="1" specimenCount-male="1" stateProvince="Para" typeStatus="holotype">
<emphasis id="E0E61B68E6C6FEBCE9A8F40B945E4D39" bold="true" italics="true">
<typeStatus id="2A81585AD437CBE2B1367198AB519209" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="A43363A0366C849D305808BA63C40822" name="Brazil">Brazil</collectingCountry>
<specimenCount id="25F6324E8A1DB83A74674FAB78C6EC77" count="1" type="male"></specimenCount>
imm.,
<collectingCounty id="F79F50F53891D8B29C5ED693C23AB294">lower Amazon</collectingCounty>
,
<location id="75E5444D00E07C230848173114CC365C" LSID="urn:lsid:plazi:treatment:CD5166CF00595E44A1C1A696A30D90E4:75E5444D00E07C230848173114CC365C" country="Brazil" county="lower Amazon" name="Santarem" stateProvince="Para">Santarém</location>
,
<collectingRegion id="E9B58214D9DE6E0DF5A73FEF58AC6162" country="Brazil" name="Para">Pará</collectingRegion>
; no date;
<collectorName id="93721F41CA839D7E48051E919EBE8778">F. O. Pickard-Cambridge</collectorName>
and
<collectorName id="46A1EBA836FCAFF0ECF59454E96B8858">Austen</collectorName>
leg;
<named-content id="41489FBC9490A905CC663A1D84665E41" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum of the United Kingdom">
<collectionCode id="68A09A90DE2157845A0C842F37AE7730" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
(no registration number). Not examined.
</materialsCitation>
</paragraph>
<paragraph id="BBDA7E3DB0EC9AAF7B4772A12F6B6D83">
<heading id="64D28AAA73CE70F34C9064FFACACAC4C" reason="title">Other material.</heading>
</paragraph>
<paragraph id="1DEBDB34B1C1E8AF51D4C016766D0243">
<materialsCitation id="16A195C7EC89E218D2004538EC293039" collectingDate="2009-04-14" collectionCode="UFRJ" collectorName="A. P. L. Giupponi &amp; D. R. Pedroso" country="Brazil" latitude="-3.2023056" location="Altamira" longLatPrecision="1" longitude="-52.22678" specimenCode="UFRJ 2575" specimenCount="1" specimenCount-female="1" stateProvince="Para">
<collectingCountry id="9AD8C8DEEC121075C323CD12978C068F" name="Brazil">Brazil</collectingCountry>
<specimenCount id="4FC43E11EA59C26C1AC0B145B9D0D1E5" count="1" type="female"></specimenCount>
;
<collectingRegion id="8A28565F6CD1A0F4ECB498D9ECB02A52" country="Brazil" name="Para">Pará</collectingRegion>
,
<location id="5B24138D90A099738D9DD79BACB91727" LSID="urn:lsid:plazi:treatment:CD5166CF00595E44A1C1A696A30D90E4:5B24138D90A099738D9DD79BACB91727" country="Brazil" latitude="-3.2023056" longLatPrecision="1" longitude="-52.22678" name="Altamira" stateProvince="Para">Altamira</location>
[
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXAAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-52.226778,-3.202306]}">
<geoCoordinate id="E09C2A000C38A1F44AA0420FBDF02145" degrees="3" direction="south" minutes="12" orientation="latitude" precision="1" seconds="08.3" value="-3.2023056">3°12'08.3&quot;S</geoCoordinate>
,
<geoCoordinate id="2AB38830FBFF433FB3459E756FBBE0F4" degrees="52" direction="west" minutes="13" orientation="longitude" precision="1" seconds="36.4" value="-52.22678">52°13'36.4&quot;W</geoCoordinate>
</named-content>
];
<date id="789D83441CF7766DCC42976EA5D7BD60" value="2009-04-14">
<collectingDate id="FEEB5C5C4022CB4A3E6D167FE7714B8C" value="2009-04-14">14 April 2009</collectingDate>
</date>
;
<collectorName id="52F18FB007C3B5B8A263637FB9F135C9">A. P. L. Giupponi</collectorName>
and
<collectorName id="BC09FB56BAB4DDED8BF27161789BF089">D. R. Pedroso</collectorName>
leg.;
<specimenCode id="33285C9FB5F9D938F3BBA987A1790EF5" collectionCode="UFRJ">
<abbrev id="ABBRID0E3AAG" xlink_title="Laboratório de Diversidade de Aracnídeos (LABAR), Universidade Federal do Rio de Janeiro">UFRJ</abbrev>
2575
</specimenCode>
</materialsCitation>
) •
<materialsCitation id="8624681672DFD79405129B256C0C8226" collectingDate="2009-04-14" collectionCode="MNRJ" collectorName="A. P. L. Giupponi &amp; D. R. Pedroso" country="Brazil" latitude="-3.2023056" location="Altamira" longLatPrecision="1" longitude="-52.22678" specimenCode="MNRJ 4371" specimenCount="2" specimenCount-female="2" stateProvince="Para">
<specimenCount id="BA0E3D1C7FE08D0AAC6CCDC542AE73D4" count="2" type="female">2 ♀♀</specimenCount>
, 5 j; same data as for previous;
<specimenCode id="3314D23DA1FBC2D56737AD37D72FCF1F" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" type="">
<named-content id="6CB8AD944DDD9505E75A3885BBEF02F9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">MNRJ</named-content>
4371
</specimenCode>
</materialsCitation>
<materialsCitation id="F5A9E60AA741CB5582663538D5FF60B2" collectingDate="2011-04-02" collectingDateMax="2011-04-07" collectingDateMin="2011-04-02" collectionCode="MNRJ" collectorName="D. R. Pedroso" country="Brazil" latitude="-3.2023056" location="Altamira" longLatPrecision="1" longitude="-52.22678" specimenCode="MNRJ 4362" specimenCount="1" specimenCount-female="1" stateProvince="Para">
<specimenCount id="298D05EBD3724DC6370FB213FCD3D0EC" count="1" type="female"></specimenCount>
, 4 j; same locality as preceding;
<date id="042E6CCD79435BF62F6023BB540AD46A" value="2011-04-02" valueMax="2011-04-07" valueMin="2011-04-02">
<collectingDate id="EA0A7C722297691A1303827FB7157976" value="2011-04-02" valueMax="2011-04-07" valueMin="2011-04-02">27 April 2011</collectingDate>
</date>
;
<collectorName id="9F69B89FE04C41BFF8FE0E1713313A21">D. R. Pedroso</collectorName>
leg;
<specimenCode id="7FB4DB5E507AF7776E7C6B5F6808D1FD" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" type="">
<named-content id="E08279EA3EAA719DEBF20E64DEC681C0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">MNRJ</named-content>
4362
</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIBAG" type="diagnosis">
<paragraph id="79FC623E5ED8DCB414D9657D8975E465">
<heading id="B3330D6363A54C43A3022C3CE2CF0D7F" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="D7831324D80D588EB5EDBCA64D49E234">
Females of
<taxonomicName id="D222D749D049427790DBF647E3A531CE" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="53025C3CADDD651DDAEDAC2BFBF1E638" italics="true">H. rufescens</emphasis>
</taxonomicName>
and
<taxonomicName id="1605B172878A822D155B8DD69FB81B05" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="E63984665C51BD4BF9993E85677B0F24" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
present spermathecae with median branch ending in a blunt rounded tip and without additional lobules (Fig.
<figureCitation id="571A3A8AEE6F712BC18603EDDF5F8C4F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 D F</figureCitation>
;
<bibRefCitation id="DB427A38D76CEA5214485ACB0C431BBF" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 5). Females of
<taxonomicName id="BC0202742F5164B724E59F6D1BC45626" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="04FAC87223D1DE47B217E695B531657B" italics="true">H. rufescens</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="48CC6A8DE0ED17A5FB057941AC100BA4" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="2032B2D37EBD91E27263CA12378A6789" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
by the tarsal claw of the pedipalp with a double row of teeth (vs. unique row;
<bibRefCitation id="16A607A9491DFE390569D3F4A985D8A5" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, p. 92); and spermathecae lateral branch with its tip folded over the stem and without lobules (vs. tip bearing several lobules) (Fig.
<figureCitation id="3C8D0179BE3DF994023D305F610AA466" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 D F</figureCitation>
;
<bibRefCitation id="A422CD02690FECB0194BFC82E5A18A50" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 5).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EODAG" type="Taxonomic notes">
<paragraph id="488545477EC089CB24E0754C42D3BBFB">
<heading id="26F0CC550291629FA3CAE0241658A55B" reason="title">Taxonomic notes.</heading>
</paragraph>
<paragraph id="10DC8E96CDDE442F3D055AB873414A89">
<taxonomicName id="33DD7F2AAE3A70581F509140E68E7808" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="A0BA8FC010C3CC4CAECE444677DB9D31" italics="true">Harmonicon</emphasis>
</taxonomicName>
was established by F. O.
<bibRefCitation id="2F173FCD41C39D6E91840E05CCC412CE" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" firstAuthor="Pickard-Cambridge" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons. Proceedings of the zoological Society of London 1896: 716766. https://doi.org/10.1111/j.1096-3642.1896.tb03076.x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge (1896)</bibRefCitation>
to accommodate a single species,
<taxonomicName id="DA0B5D1E9721A0559CA4918B085F415A" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="D02A4CC5FD746ADD355AC6FF7C625E8E" italics="true">H. rufescens</emphasis>
</taxonomicName>
, based on a presumed female specimen from Santarém,
<collectingCountry id="2A7DC6BF3589A4F974AA1EB2495E9873" name="Brazil">Brazil</collectingCountry>
.
<bibRefCitation id="B833D1164D9316F557FED8215AE2EBB4" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam (2011</bibRefCitation>
, p. 91) were able to examine the
<typeStatus id="21D28D2D8770E0F6E4E72897AA84D238" type="holotype">holotype</typeStatus>
and noticed that “ Cambridge (1896) was incorrect in stating that the
<typeStatus id="654912FDFE44C2BA5039AC2BEDEEB286" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8B086A5B238696FB85DBB97C62024480" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="E06E27131573A2CAF7AC6CBCBE2CCDDF" italics="true">Harmonicon rufescens</emphasis>
</taxonomicName>
is a female; it is in fact a juvenile male ”. The female described here is from Altamira, also in the
<collectingRegion id="FA076F934F2463491AA46812EDEF7CF7" country="Brazil" name="Para">Pará state</collectingRegion>
,
<collectingCountry id="512D30E8EC3F6CA7825C7BC9F8FE73E6" name="Brazil">Brazil</collectingCountry>
, and located ca.
<quantity id="073272E385553BA4AAB2AD7EA5E358E9" metricMagnitude="5" metricUnit="m" metricValue="3.0" unit="km" value="300.0">300 km</quantity>
from Santarém. Both localities are placed in the lower Amazons, have similar climate, low altitude, and are covered by dense or open Ombrophile Amazonian Forest and Savannas (
<bibRefCitation id="AFEC3F20B0387D2D9BC79CC7CD8F4FF2" DOI="10.1590/S0100-84042008000100015" author="Magnusson" etAl="et al." firstAuthor="Magnusson" issue="1" journalOrPublisher="Revista Brasileira de Botânica" pagination="165-177" refId="B8" refString="Magnusson WE, Lima AP, Albernaz ALKM, Sanaiotti TM, Guillaumet J-L (2008) Composição florística e cobertura vegetal das savanas na região de Alter do Chão, Santarém PA. Revista Brasileira de Botânica 31 (1): 165177. https://doi.org/10.1590/S0100-84042008000100015" title="Composição florística e cobertura vegetal das savanas na região de Alter do Chão, Santarém - PA." volume="31" year="2008">Magnusson et al. 2008</bibRefCitation>
). Other
<taxonomicName id="2DB8098E47A5B2A226B60661D06B541F" authorityName="Simon" authorityYear="1889" class="Arachnida" family="Dipluridae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="family">Dipluridae</taxonomicName>
species collected in Santarém at the same expedition to lower Amazon as
<taxonomicName id="8FB47447A1F92D37C9CB80F5B0C42955" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="08BD7DA3964FC42FBAEFE21756DFB54F" italics="true">H. rufescens</emphasis>
</taxonomicName>
also have large distribution range.
<taxonomicName id="0A3549DA830057B77FC2E143C2E34AC3" authority="(F. O. Pickard-Cambridge, 1896)" baseAuthorityName="F. O. Pickard-Cambridge" baseAuthorityYear="1896" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="sanguinea">
<emphasis id="9183A2E7EEC5FE79815A9B51B28EF0A5" italics="true">Diplura sanguinea</emphasis>
(F. O.
<bibRefCitation id="0DDA3839AE32E3CF81F8A9D5F5EB34B8" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship &quot; Faraday &quot; up the river Amazons. Proceedings of the zoological Society of London 1896: 716 - 766. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03076. x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge, 1896</bibRefCitation>
)
</taxonomicName>
is widespread through North and Northeast regions of
<collectingCountry id="486BB66F2958048C9AE7781AB22A57A6" name="Brazil">Brazil</collectingCountry>
(
<bibRefCitation id="8BACF6D03E25DBA9C85DCB22D29D474D" DOI="10.13156/arac.2020.18.7.681" author="Brescovit" etAl="et al." firstAuthor="Brescovit" issue="7" journalOrPublisher="Arachnology" pagination="681-689" refId="B2" refString="Brescovit AD, Sherwood D, Lucas SM (2021) First description of the males of Diplura sanguinea (F. O. Pickard-Cambridge, 1896) and Diplura nigra (F. O. Pickard-Cambridge, 1896) from Brazilian Amazonia, with notes on biogeography and opisthosomal patterning (Araneae, Dipluridae). Arachnology 18 (7): 681689. https://doi.org/10.13156/arac.2020.18.7.681" title="First description of the males of Diplura sanguinea (F. O. Pickard-Cambridge, 1896) and Diplura nigra (F. O. Pickard-Cambridge, 1896) from Brazilian Amazonia, with notes on biogeography and opisthosomal patterning (Araneae, Dipluridae)." volume="18" year="2021">Brescovit et al. 2021</bibRefCitation>
) and is also found in
<collectingCountry id="CC357469CBD4F5F1BDA30D03DD99A6E7" name="Colombia">Colombia</collectingCountry>
(
<bibRefCitation id="2390F921A1FA7C6E74C81F9D34D55132" DOI="10.5281/zenodo.10201779" author="Sherwood" etAl="et al." firstAuthor="Sherwood" journalOrPublisher="ZooNova" pagination="1-22" refId="B16" refString="Sherwood D, Drolshagen B, Osorio LM, Benavides LR, Seiter M (2023) An inordinate fondness for spinnerets: on some spiders of the genera Diplura C. L. Koch, 1850 and Linothele Karsch, 1879 with new species, records, and notes on types (Araneae, Dipluridae). ZooNova 29: 122. https://doi.org/10.5281/zenodo.10201779" title="An inordinate fondness for spinnerets: on some spiders of the genera Diplura C. L. Koch, 1850 and Linothele Karsch, 1879 with new species, records, and notes on types (Araneae, Dipluridae)." volume="29" year="2023">Sherwood et al. 2023</bibRefCitation>
).
<taxonomicName id="85E7DC81C74BE3AE4F18FE6B45215F8A" authority="(F. O. Pickard-Cambridge, 1896)" baseAuthorityName="F. O. Pickard-Cambridge" baseAuthorityYear="1896" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="710E6168927C29EF358623AB65FC056F" italics="true">Diplura nigra</emphasis>
(F. O.
<bibRefCitation id="CFC665F92B1249C84C6D173BEB37A3D2" DOI="10.1111/j.1096-3642.1896.tb03076.x" author="Pickard-Cambridge FO" journalOrPublisher="Proceedings of the zoological Society of London" pagination="716-766" refId="B14" refString="Pickard-Cambridge FO (1896) On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship &quot; Faraday &quot; up the river Amazons. Proceedings of the zoological Society of London 1896: 716 - 766. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03076. x" title="On the Theraphosidae of the lower Amazons: being an account of the new genera and species of this group of spiders discovered during the expedition of the steamship “ Faraday ” up the river Amazons." volume="1896" year="1896">Pickard-Cambridge, 1896</bibRefCitation>
)
</taxonomicName>
has been recorded from several localities around Santarém (
<bibRefCitation id="EE703EBF38205B9B22EB0E525035EF9E" DOI="10.13156/arac.2020.18.7.681" author="Brescovit" etAl="et al." firstAuthor="Brescovit" issue="7" journalOrPublisher="Arachnology" pagination="681-689" refId="B2" refString="Brescovit AD, Sherwood D, Lucas SM (2021) First description of the males of Diplura sanguinea (F. O. Pickard-Cambridge, 1896) and Diplura nigra (F. O. Pickard-Cambridge, 1896) from Brazilian Amazonia, with notes on biogeography and opisthosomal patterning (Araneae, Dipluridae). Arachnology 18 (7): 681689. https://doi.org/10.13156/arac.2020.18.7.681" title="First description of the males of Diplura sanguinea (F. O. Pickard-Cambridge, 1896) and Diplura nigra (F. O. Pickard-Cambridge, 1896) from Brazilian Amazonia, with notes on biogeography and opisthosomal patterning (Araneae, Dipluridae)." volume="18" year="2021">Brescovit et al. 2021</bibRefCitation>
), and we examined specimens from other localities from
<collectingRegion id="15EF8AF7902B4A593F9EDC526745343A" country="Brazil" name="Para">Pará</collectingRegion>
, Amazonas,
<collectingRegion id="1CA6F6303AB0243912740AA7A4DCC4F9" country="Brazil" name="Roraima">Roraima</collectingRegion>
,
<collectingRegion id="E23DED24BBC3A9811A6DF16CFBA237E0" country="Brazil" name="Rondonia">Rondônia</collectingRegion>
, and
<collectingRegion id="EEA22AEDE22139A1D2FABE88BF4A960E" country="Brazil" name="Mato Grosso">Mato Grosso</collectingRegion>
states in
<collectingCountry id="06A965ADDD9CDFB4FAAD52DC2C74E186" name="Brazil">Brazil</collectingCountry>
. The only other species described for the same Brazilian state is
<taxonomicName id="3D90B4001B16ED85A5C5B6C4A4991171" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="4F94638939AE1B5DC100A5B41F17E788" italics="true">H. cerberus</emphasis>
</taxonomicName>
, from Pequiá cave, near Carajás, in the municipality of Parauapebas, but its male
<typeStatus id="E153E4CFA82F0A95FEF9C5341C0B892D" type="holotype">holotype</typeStatus>
has several troglomorphic characteristics, and its type locality is placed ca.
<quantity id="D34B3C61666D5EB7731CEC2B5536ED29" metricMagnitude="5" metricUnit="m" metricValue="6.6" unit="km" value="660.0">660 km</quantity>
away from Santarém (
<bibRefCitation id="9C5434817ED2F5F612571C2F64DDEE4B" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista 2014</bibRefCitation>
). Besides, the region around the cave where
<taxonomicName id="DE1242AA8DDF75D8A879DF83692991E9" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="EAB616FCDE3D42DD5A77211F8A1E1120" italics="true">H. cerberus</emphasis>
</taxonomicName>
was collected is covered by the Amazon “ canga ” vegetation, which covers areas rich in iron ores and has unique vegetation (
<bibRefCitation id="224759803BB927558E9189CAD063BF23" DOI="10.1590/2175-7860201869336" author="Mota" etAl="et al." firstAuthor="Mota" issue="3" journalOrPublisher="Rodriguésia" pagination="1435-1488" refId="B10" refString="Mota NFO, Watanabe MTC, Zappi DC, Hiura AL, Pallos J, Viveros RS, Giulietti AM, Viana PL (2018) Amazon canga: the unique vegetation of Carajás revealed by the list of seed plants. Rodriguésia 69 (3): 14351488. https://doi.org/10.1590/2175-7860201869336" title="Amazon canga: the unique vegetation of Carajás revealed by the list of seed plants." volume="69" year="2018">Mota et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENHAG" type="description">
<paragraph id="F5BE77306B89494E801C8154B13F02AF">
<heading id="D65F7C22B770350BF0D2936E3C826D4F" reason="title">Description.</heading>
</paragraph>
<paragraph id="8693BDFBE6977ABF683D3E2D3263A9FB">
<emphasis id="08ADF6A3B66511E9878C3091785723E8" bold="true">
Female (
<abbrev id="ABBRID0EVHAG" xlink_title="Laboratório de Diversidade de Aracnídeos (LABAR), Universidade Federal do Rio de Janeiro">
<collectionCode id="94ADA5CE77F1B1B5F652A659301BA591">UFRJ</collectionCode>
</abbrev>
2575)
</emphasis>
. Carapace oblong, wider at middle portion, light orange-brown, covered with dark setae, caput slightly elevated, darker; cephalic area not shortened, darker furrows, margins of thoracic area with a row of long, thick, rigid, erect, and curved dark setae pointing ectally (Fig.
<figureCitation id="88B51B1CC0E403868D928F3ED26BCD90" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
). Clypeus long, with frontal margin carrying 19 median-to-long, thick setae facing forward (Fig.
<figureCitation id="158FF84A0509C2C5CC70D37F6805206D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
). Eye tubercle with 15 thick setae on its anterior margin; area between eyes with 11 short and thickened setae, and others abundant and dark but not thickened setae (Fig.
<figureCitation id="041920A4FCC929D6C2892DC7D59805FA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
). Eye formula:
<abbrev id="ABBRID0EGIAG" xlink_title="anterior lateral eyes">ALE</abbrev>
&gt;
<abbrev id="ABBRID0EKIAG" xlink_title="posterior lateral eyes">PLE</abbrev>
&gt;
<abbrev id="ABBRID0EOIAG" xlink_title="anterior median eyes">AME</abbrev>
&gt;
<abbrev id="ABBRID0ESIAG" xlink_title="posterior median eyes">PME</abbrev>
. Ocular area much wider than long. Anterior eye row slightly recurved. Posterior eye row slightly recurved. Median ocular quadrangle wider than long, wider at its posterior margin.
<abbrev id="ABBRID0EWIAG" xlink_title="anterior median eyes">AME</abbrev>
rounded,
<abbrev id="ABBRID0E1IAG" xlink_title="anterior lateral eyes">ALE</abbrev>
and
<abbrev id="ABBRID0E5IAG" xlink_title="posterior lateral eyes">PLE</abbrev>
elliptical, and
<abbrev id="ABBRID0ECJAG" xlink_title="posterior median eyes">PME</abbrev>
reduced in size, oval.
<abbrev id="ABBRID0EGJAG" xlink_title="anterior median eyes">AME</abbrev>
-
<abbrev id="ABBRID0EKJAG" xlink_title="anterior median eyes">AME</abbrev>
separated by less than its diameter.
<abbrev id="ABBRID0EOJAG" xlink_title="anterior median eyes">AME</abbrev>
contiguous to
<abbrev id="ABBRID0ESJAG" xlink_title="posterior median eyes">PME</abbrev>
.
<abbrev id="ABBRID0EWJAG" xlink_title="anterior median eyes">AME</abbrev>
separated from
<abbrev id="ABBRID0E1JAG" xlink_title="posterior median eyes">PME</abbrev>
by around half of its diameter.
<abbrev id="ABBRID0E5JAG" xlink_title="anterior lateral eyes">ALE</abbrev>
and
<abbrev id="ABBRID0ECKAG" xlink_title="posterior median eyes">PME</abbrev>
contiguous to
<abbrev id="ABBRID0EGKAG" xlink_title="posterior lateral eyes">PLE</abbrev>
. Fovea transverse and slightly recurved (Fig.
<figureCitation id="3DFF5D8F221FC78A592703894DD8C67B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
). Labium wider than long with no cuspules (Fig.
<figureCitation id="13C17AF2159424739329682264213536" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 B</figureCitation>
). Sternum longer than wide, covered by black stiff setae and brownish thin recumbent setae, one pair of labial comma-shaped sigilla clearly separated from each other, three sternal sigilla near margin on each side, increasing in size from first to last, the third one much larger and elliptical (Fig.
<figureCitation id="0BF7593C41B54CFE2A64216E446777CB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 B</figureCitation>
). Chelicerae dark red with 10 promarginal teeth at left chelicera and 12 promarginal teeth at right chelicera, fangs furrow with denticles. Plectrum with 68 separated thick setae on left side and right side, respectively. Maxillae elongated with 25 cuspules at the right side and 31 at the left one (Fig.
<figureCitation id="08A839B1CA5B89202C64BB31F1EEEA97" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 B</figureCitation>
). Lyra at ventral side of maxilla, asymmetrical, formed by 67 modified thick and long setae, increasing in size in a basal to distal direction, with a distinct gap between the base of each seta (Fig.
<figureCitation id="8249D90DBF8BAC0BBD3422CCEE61EE73" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
). Left lyra with six setae (Fig.
<figureCitation id="44D9F133F9F199B095ED185488703FBA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
), right lyra with seven (two broken) setae. Left lyra thick, erect, regularly curved setae (Fig.
<figureCitation id="A6AC732A2A03DE820F9D3E640D62066D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
, arrows) placed distally and interspersed among the field of long and thin setae covering the retrolateral face of maxilla, five at the left side and four at the right side. Leg formula IV&gt; I&gt; II&gt; III; length of segments (femur + patella + tibia + metatarsus + tarsus = total length): I 8.4 + 2.8 + 8.3 + 6.9 + 5.1 = 31.5, II 7.4 + 2.6 + 7.5 + 6.9 + 4.8 = 29.2, III 6.7 + 2.3 + 7.0 + 8.2 + 4.6 = 28.8, IV 8.6 + 2.7 + 9.6 + 10.8 + 4.9 = 36.6. Tarsi with few cracks covering almost all dorsal and lateral faces, except by basis and tip of article; all tarsi and metatarsi with thin scopula, metatarsi with inconspicuous scopula; pedipalp tarsi with middle to apical scopulae; pedipalp claw with seven teeth;
<abbrev id="ABBRID0EGLAG" xlink_title="inferior tarsal claw">ITC</abbrev>
without teeth;
<abbrev id="ABBRID0EKLAG" xlink_title="superior tarsal claw">STC</abbrev>
teeth: I: 95, II: 94, III IV: 104. Macrosetae: Leg I: femur pld 00 2, d 11 1, rld 00 1, patella 0, tibia v 11 3 right, p 01 1, v 11 2 left, metatarsus v 21 2; Leg II: femur d 21 0, pld 00 1 left, pld 02 1 right, patella pld 00 1 left, tibia p 00 1 left, p 01 1 right, v 12 2 right, v 11 2 left, metatarsus v 12 2, p 01 0; Leg III: femur pld 00 2 left, d 20 0, rld 02 1 left, rld 22 1 right, patella 0, tibia p 01 1, rld 10 0, r 02 1 left, r 01 1 right, v 20 2 left, v 22 2 right, metatarsus d 00 2, r 02 0 left, rld 10 0 left, r 12 2 right, p 21 0, v 22 2 right, v 32 3 left; Leg IV: femur rld 22 1, d 11 0 right, d 21 0 left, patella r 01 0, tibia rld 10 0, p 10 1, r 11 1, v 22 2, metatarsus rld 21 1 left, rld 22 1 right, r 02 0 left, pld 21 1, v 21 3 left, v 31 3 right. Abdomen light brown with no stripes (Fig.
<figureCitation id="E341515C0B487F29AC0E8A241F3C543B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
), covered with dark setae, venter uniformly brown.
<abbrev id="ABBRID0ESLAG" xlink_title="posterior lateral spinnerets">PLS</abbrev>
longer than half of abdomen, with articulate fusules on all articles, and last article finger-like and with light patches of ventral cracks (Fig.
<figureCitation id="E556E51DD97B33061E33CFF1EA62F05A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 A</figureCitation>
). Spermathecae separated by ca. 1.5 × its length, with two sclerotised thick stems (or branches), originating from a common basis poorly sclerotised and translucent, each branch with almost the same thickness and length; median branch slightly curved laterally and with large, rounded, and blunt tip without vesicles; lateral branch straight, with a wider rounded head, with its tip rounded and folded upon the stem (Fig.
<figureCitation id="89B889160A09F0407B3CF82AC65DD00F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 D F</figureCitation>
).
</paragraph>
<caption id="3F9E43088F59A15FAC0BEE910D8F9639" ID-arpha="4EEF1C8A-5EA3-5D89-9CEB-2E845FB8C226" httpUri="https://binary.pensoft.net/fig/1178835" startId="F1">
<paragraph id="B3E5EEED8471BDAC8B2FEFDA27BB5AD7">
<label id="FAD2A957D21233AA572FAD3D8981E08F">Figure 1.</label>
</paragraph>
<paragraph id="B79DB207BCEBD2752C8771C53E42FC8F">
<taxonomicName id="FB7C67CCC4DAAD5239F170D3D8410B07" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="90C3E09590BDC224ABB959D85D03858B" italics="true">Harmonicon rufescens</emphasis>
</taxonomicName>
, female (
<abbrev id="ABBRID0ERMAG" xlink_title="Laboratório de Diversidade de Aracnídeos (LABAR), Universidade Federal do Rio de Janeiro">UFRJ</abbrev>
2575).
<emphasis id="07FF4948EFD8584FB43688B0CC69C12C" bold="true">A.</emphasis>
Dorsal habitus;
<emphasis id="9FA99C8022C8691DADDD9A5E6D88C450" bold="true">B.</emphasis>
Sternum;
<emphasis id="AE2FCAC7825BA030C7214398C3F2B3D2" bold="true">C.</emphasis>
Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae);
<emphasis id="8D74EDCD25B23C4243435BA8503A2485" bold="true">D F.</emphasis>
Spermathecae:
<emphasis id="87032CA1A9B7763F9EDA31B2DC02318A" bold="true">D.</emphasis>
Ventral view;
<emphasis id="852E183B5BC31819BAD8830F1189E1DF" bold="true">E.</emphasis>
Dorsal view;
<emphasis id="1711F92B07ED4D29079567C37D738E62" bold="true">F.</emphasis>
Ventral view, partially cleared;
<emphasis id="CAB600B54674C178942836665F23ACEA" bold="true">G.</emphasis>
Alive immature female specimen in nature (
<named-content id="2A0929D70E52092768AEF51B276A6043" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">MNRJ</named-content>
4362). Scale bars: 5 mm (
<emphasis id="2D9E3E5382F1AD1E600357D4BCF7CF18" bold="true">A</emphasis>
); 2 mm (
<emphasis id="FC6A919645B759CDBD8FF40B27B5777E" bold="true">B</emphasis>
); 1 mm (
<emphasis id="1511C35EEBE652B17B5637210B708D1B" bold="true">C</emphasis>
); 0.5 mm (
<emphasis id="E1BAFF4A97FCCBE1C97A1F3DC757793F" bold="true">D E</emphasis>
). Living specimen photo by Denis Pedroso.
</paragraph>
</caption>
<paragraph id="927951B3D4E42C82E665B4FB605847FD">
<abbrev id="ABBRID0EYNAG" xlink_title="total length">TL</abbrev>
26.9. Carapace 10.2 long, 9.1 wide, carapace length / width 1.12.
<abbrev id="ABBRID0E3NAG" xlink_title="anterior median eyes">AME</abbrev>
<quantity id="ECF95AD1A60048451D43AA876D25521E" metricMagnitude="-4" metricUnit="m" metricValue="3.2" unit="mm" value="0.32">0.32 mm</quantity>
,
<abbrev id="ABBRID0EAOAG" xlink_title="anterior median eyes">AME</abbrev>
-
<abbrev id="ABBRID0EEOAG" xlink_title="anterior median eyes">AME</abbrev>
<quantity id="68BEF96236DA348AC131F798B69106D6" metricMagnitude="-4" metricUnit="m" metricValue="2.4" unit="mm" value="0.24">0.24 mm</quantity>
. Abdomen 16.7 long, 8.5 wide. Spinnerets:
<abbrev id="ABBRID0EIOAG" xlink_title="posterior median spinnerets">PMS</abbrev>
2.6 long;
<abbrev id="ABBRID0EMOAG" xlink_title="posterior lateral spinnerets">PLS</abbrev>
16.2 long, basal article 4.7, middle 4.9, distal 6.6.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQOAG" type="variation">
<paragraph id="0A861BF8F0EDCAD3B22E6C2C0DFC33AA">
<heading id="392E00B5748BB201AF765FAC6690048D" reason="title">Variation.</heading>
</paragraph>
<paragraph id="27E604854406AFD6036DFBA53C3B94BE">
Females (n = 6).
<abbrev id="ABBRID0EWOAG" xlink_title="total length">TL</abbrev>
24.432.9. Chelicerae with 10 to 13 promarginal teeth, plectrum with five to eight setae, 25 to 40 maxillary cuspules, and lyra with five to seven bristles.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1OAG" type="distribution">
<paragraph id="FB4DA11230A85CB8DB6514A915E15973">
<heading id="9C84DA7CEA38FB557BB9F42BAE976EC2" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="5409048740D6B4D70A227E3A885BAC5D">
Only known from Altamira and Santarém cities,
<collectingRegion id="5B545C565CF7C06543107E1AB71FBA40" country="Brazil" name="Para">Pará State</collectingRegion>
,
<collectingCountry id="9243043F0F88CD4F799D831240A2D249" name="Brazil">Brazil</collectingCountry>
(Fig.
<figureCitation id="730C377838C8BF2AEDAB716B48DECDF3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution records of H. rufescens (holotype = red and black circle, specimen here described = red circle) and H. candango sp. nov. (yellow square)." httpUri="https://binary.pensoft.net/fig/1178838">4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,55 +1,55 @@
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<mods:namePart id="75DC8C710FB37C3B631E8A2AC15704F8">Wermelinger-Moreira, Gabriel</mods:namePart>
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<mods:namePart id="CE7E8BF3237F403CC0A1F30DB6ACE4DF">Wermelinger-Moreira, Gabriel</mods:namePart>
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<mods:namePart id="CC578AC1628D9FBA4DD1CF2590552EDC">de Souza Castanheira, Pedro</mods:namePart>
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; same data as for preceding;
; same data as for preceding
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