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<document id="90DA69CA6AA351D93D321C2C66D709CB" ID-CLB-Dataset="27553" ID-DOI="10.11646/zootaxa.4555.2.1" ID-GBIF-Dataset="95c8b580-1685-41d2-9853-1a0b5b83c327" ID-ISSN="1175-5326" ID-Zenodo-Dep="2624219" ID-ZooBank="E8F6B60F-3E7B-4BD1-938C-F46A840AA08D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554272112745" checkinUser="plazi" docAuthor="Joaqui, Tatiana, Moctezuma, Victor, Sánchez-Huerta, José Luis &amp; Escobar, Federico" docDate="2019" docId="BA2787A68069FFF87AEAFE4BFDC4195F" docLanguage="en" docName="zootaxa.4555.2.1.pdf" docOrigin="Zootaxa 4555 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus canescens Zunino &amp; Halffter 1988" docType="treatment" docVersion="8" lastPageNumber="160" masterDocId="461EFFDE8060FFF17A7DFFFCFF8B1D13" masterDocTitle="The Onthophagus fuscus (Coleoptera: Scarabaeidae) species complex: an update and the description of a new species" masterLastPageNumber="186" masterPageNumber="151" pageNumber="160" updateTime="1698716389512" updateUser="ExternalLinkService">
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<mods:title id="EEEDBC070375B571A1212798B63CA9EA">The Onthophagus fuscus (Coleoptera: Scarabaeidae) species complex: an update and the description of a new species</mods:title>
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<mods:namePart id="202807BF33A01BBE7D92C84AE44DC014">Joaqui, Tatiana</mods:namePart>
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<mods:namePart id="8253F3627BA2647DAED1A9A23E335E5D">Sánchez-Huerta, José Luis</mods:namePart>
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<taxonomicName id="F58E4D338069FFF87AEAFE4BFD781CC1" ID-CoL="7XMJZ" authority="Zunino &amp; Halffter, 1988" authorityName="Zunino &amp; Halffter" authorityYear="1988" box="[151,755,439,466]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="canescens">
<emphasis id="00FAEAA28069FFF87AEAFE4BFE361CC2" bold="true" box="[151,445,439,465]" italics="true" pageId="9" pageNumber="160">Onthophagus canescens</emphasis>
<bibRefCitation id="561F4B418069FFF87BB9FE44FD781CC1" author="Zunino, M. &amp; Halffter, G." box="[452,755,440,466]" pageId="9" pageNumber="160" pagination="1 - 211" refId="ref14729" refString="Zunino, M. &amp; Halffter, G. (1988) Analisis taxonomico, ecologico y biogeografico de un grupo americano de Onthophagus (Coleoptera: Scarabaeidae). Museo Regionale di Scienze Naturali Torino, 9, 1 - 211." type="journal article" year="1988">Zunino &amp; Halffter, 1988</bibRefCitation>
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, new rank
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</paragraph>
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<figureCitation id="AAB52A358069FFF87AEAFE26FE8A1CE0" box="[151,257,474,499]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="10.[151,250,2003,2025]" captionStartId-1="11.[151,250,1990,2012]" captionTargetBox-0="[180,1421,322,1833]" captionTargetBox-1="[175,1417,288,1891]" captionTargetId-0="figure@10.[151,1437,288,1876]" captionTargetId-1="figure@11.[150,1437,238,1924]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 3. Onthophagus canescens male dorsal habitus, paratype." captionText-1="FIGURE 4. Onthophagus canescens female dorsal habitus, paratype." httpUri-0="https://zenodo.org/record/2624223/files/figure.png" httpUri-1="https://zenodo.org/record/2624225/files/figure.png" pageId="9" pageNumber="160">Figs. 3–4</figureCitation>
,
<figureCitation id="AAB52A358069FFF87B70FE27FED81CE0" box="[269,339,475,499]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStart-2="FIGURE 20" captionStartId-0="26.[151,250,1082,1104]" captionStartId-1="27.[151,250,1965,1987]" captionStartId-2="28.[151,250,1594,1616]" captionTargetBox-0="[190,1399,210,1069]" captionTargetBox-1="[251,1337,210,1936]" captionTargetBox-2="[158,1410,200,1557]" captionTargetId-0="figure@26.[186,1407,195,1069]" captionTargetId-1="figure@27.[251,1346,195,1950]" captionTargetId-2="figure@28.[150,1436,195,1572]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionText-0="FIGURE 18. Aedeagus: A, Onthophagus fuscus; B, O. canescens; C, O. mycetorum; D, O. parafuscus; E, O. pseudofuscus. Redrawn from Zunino &amp; Halffter (1988)." captionText-1="FIGURE 19. Lamella copulatrix: A, O. fuscus; B, O. canescens; C, O. mycetorum; D, O. parafuscus; E, O. pseudofuscus. Redrawn from Zunino &amp; Halffter (1988)." captionText-2="FIGURE 20. Female genitalia: A, O. fuscus; B, O. canescens; C, O. mycetorum; D, O. parafuscus. Redrawn from Zunino &amp; Halffter (1988)." httpUri-0="https://zenodo.org/record/2624253/files/figure.png" httpUri-1="https://zenodo.org/record/2624257/files/figure.png" httpUri-2="https://zenodo.org/record/2624259/files/figure.png" pageId="9" pageNumber="160">18–20</figureCitation>
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<taxonomicName id="F58E4D338069FFF87AEAFDDEFD751F2B" ID-CoL="7XMJZ" authority="Zunino &amp; Halffter 1988: 83" authorityName="Zunino &amp; Halffter" authorityPageNumber="83" authorityYear="1988" box="[151,766,546,568]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="subSpecies" species="fuscus" subSpecies="canescens">
<emphasis id="00FAEAA28069FFF87AEAFDDEFE5D1F2B" box="[151,470,546,568]" italics="true" pageId="9" pageNumber="160">Onthophagus fuscus canescens</emphasis>
<treatmentCitation id="B32F10A18069FFF87BA0FDDEFD751F2B" author="Zunino, M. &amp; Halffter, G." box="[477,766,546,568]" page="83" pageId="9" pageNumber="160" year="1988">
<bibRefCitation id="561F4B418069FFF87BA0FDDEFD531F2B" author="Zunino, M. &amp; Halffter, G." box="[477,728,546,568]" pageId="9" pageNumber="160" pagination="1 - 211" refId="ref14729" refString="Zunino, M. &amp; Halffter, G. (1988) Analisis taxonomico, ecologico y biogeografico de un grupo americano de Onthophagus (Coleoptera: Scarabaeidae). Museo Regionale di Scienze Naturali Torino, 9, 1 - 211." type="journal article" year="1988">Zunino &amp; Halffter 1988</bibRefCitation>
: 83
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–85;
<treatmentCitation id="B32F10A18069FFF8794FFDDEFB361F2B" author="Navarrete-Heredia, J. L. &amp; Galindo, N. E." box="[818,1213,546,568]" page="3" pageId="9" pageNumber="160" year="1997">
<bibRefCitation id="561F4B418069FFF8794FFDDEFB291F2B" author="Navarrete-Heredia, J. L. &amp; Galindo, N. E." box="[818,1186,546,568]" pageId="9" pageNumber="160" pagination="1 - 16" refId="ref13974" refString="Navarrete-Heredia, J. L. &amp; Galindo, N. E. (1997) Escarabajos asociados a Basidiomycetes en San Jose de Los Laureles, Morelos, Mexico (Coleoptera: Scarabaeidae). Folia Entomologica Mexicana, 99, 1 - 16." type="journal article" year="1997">Navarrete-Heredia &amp; Galindo 1997</bibRefCitation>
: 3
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;
<treatmentCitation id="B32F10A18069FFF87EB4FDDEFA131F2B" author="Anduaga, S." box="[1225,1432,546,568]" page="130" pageId="9" pageNumber="160" year="2000">
<bibRefCitation id="561F4B418069FFF87EB4FDDEFAEF1F2B" author="Anduaga, S." box="[1225,1380,546,568]" pageId="9" pageNumber="160" pagination="119 - 130" refId="ref11922" refString="Anduaga, S. (2000) Escarabajos coprofagos (Coleoptera: Scarabaeoidea) asociados a hongos en la Sierra Madre Occidental, Durango, Mexico: con una compilacion de las especies micetofagas. Acta Zoologica Mexicana, New Series, 80, 119 - 130." type="journal article" year="2000">Anduaga 2000</bibRefCitation>
: 130
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;
<treatmentCitation id="B32F10A18069FFF87ABAFDBDFE091F44" author="Krajcik, M." box="[199,386,577,599]" page="106" pageId="9" pageNumber="160" year="2006">
<bibRefCitation id="561F4B418069FFF87ABAFDBDFEDB1F44" author="Krajcik, M." box="[199,336,577,599]" pageId="9" pageNumber="160" pagination="1 - 190" refId="ref13559" refString="Krajcik, M. (2006) Checklist of Scarabaeoidea of the world. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma X, 3, 1 - 190." type="journal article" year="2006">Krajcik 2006</bibRefCitation>
: 106
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;
<treatmentCitation id="B32F10A18069FFF87BF1FDBDFD761F44" author="Pulido-Herrera, L. A. &amp; Zunino, M." box="[396,765,577,599]" page="103" pageId="9" pageNumber="160" year="2007">
<bibRefCitation id="561F4B418069FFF87BF1FDBDFD411F44" author="Pulido-Herrera, L. A. &amp; Zunino, M." box="[396,714,577,599]" pageId="9" pageNumber="160" pagination="93 - 129" refId="ref14096" refString="Pulido-Herrera, L. A. &amp; Zunino, M. (2007) Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeinae). Sociedad Entomologica Aragonesa, Monografias Tercer Milenio, 7, 93 - 129. Available from: http: // sea-entomologia. org / PDF / PDFSM 3 MVOL 7 / Pdf 9093129 PulidoYZunino. pdf (accessed 28 October 2018)" type="journal article" year="2007">Pulido-Herrera &amp; Zunino 2007</bibRefCitation>
: 103
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;
<treatmentCitation id="B32F10A18069FFF8797AFDBEFB921F44" author="Moctezuma, V. &amp; Rossini, M. &amp; Zunino, M. &amp; Halffter, G." box="[775,1049,577,599]" page="33" pageId="9" pageNumber="160" year="2016">
<bibRefCitation id="561F4B418069FFF8797AFDBEFC7F1F44" author="Moctezuma, V. &amp; Rossini, M. &amp; Zunino, M. &amp; Halffter, G." box="[775,1012,577,599]" pageId="9" pageNumber="160" pagination="23 - 50" refId="ref13844" refString="Moctezuma, V., Rossini, M., Zunino, M. &amp; Halffter, G. (2016) A contribution to the knowledge of the mountain entomofauna of Mexico with a description of two new species of Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae). Zookeys, 572, 23 - 50. https: // doi. org / 10.3897 / zookeys. 572.6763" type="journal article" year="2016">
Moctezuma
<emphasis id="00FAEAA28069FFF879F8FDBEFC331F44" box="[901,952,577,599]" italics="true" pageId="9" pageNumber="160">et al.</emphasis>
2016
</bibRefCitation>
: 33
</treatmentCitation>
, 35–36.
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<emphasis id="00FAEAA28069FFF87AEAFD78FE581F8E" bold="true" box="[151,467,644,669]" pageId="9" pageNumber="160">Type material examined (</emphasis>
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,
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<emphasis id="00FAEAA28069FFF878D8FD78FCB41F8E" bold="true" box="[677,831,644,669]" pageId="9" pageNumber="160">
).
<typeStatus id="ED3588128069FFF878BDFD78FCB11F8E" box="[704,826,644,669]" pageId="9" pageNumber="160" type="paratype">Paratypes</typeStatus>
.
</emphasis>
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<specimenCount id="2488FD398069FFF87937FD78FC171F8E" box="[842,924,644,669]" pageId="9" pageNumber="160" type="male">1 male</specimenCount>
,
<specimenCount id="2488FD398069FFF879D6FD78FB9E1F8E" box="[939,1045,644,669]" pageId="9" pageNumber="160" type="female">1 female</specimenCount>
: “
<collectingCountry id="4A9976208069FFF87E50FD78FB0C1F8E" box="[1069,1159,644,669]" name="Mexico" pageId="9" pageNumber="160">México</collectingCountry>
, Parque Nacional J. M.
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,
<location id="3751606B8069FFF87B7FFD5BFED51FD3" LSID="urn:lsid:plazi:treatment:BA2787A68069FFF87AEAFE4BFDC4195F:3751606B8069FFF87B7FFD5BFED51FD3" box="[258,350,679,704]" country="Mexico" name="Morelia" pageId="9" pageNumber="160" stateProvince="Morelos">Morelia</location>
,
<collectingRegion id="F04AF8528069FFF87B14FD5BFE631FD3" box="[361,488,679,704]" country="Mexico" name="Michoacan de Ocampo" pageId="9" pageNumber="160">Michoacán</collectingRegion>
,
<date id="463010708069FFF87B8EFD5BFDEF1FD3" box="[499,612,679,704]" pageId="9" pageNumber="160" value="1964-07-25">
<collectingDate id="5674E9988069FFF87B8EFD5BFDEF1FD3" box="[499,612,679,704]" pageId="9" pageNumber="160" value="1964-07-25">25/VII/64</collectingDate>
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,
<collectorName id="9F7B53668069FFF87812FD54FD451FD3" box="[623,718,680,704]" pageId="9" pageNumber="160">P. Reyes</collectorName>
C. col., en hongos” (
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)
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;
<materialsCitation id="82E63CED8069FFF87E72FD54FBB01FF7" ID-GBIF-Occurrence="2234218860" collectingDate="1967-06-25" collectionCode="VMC" collectorName="G. y V. Halffter" country="Mexico" location="Valle de Bravo" pageId="9" pageNumber="160" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="2488FD398069FFF87E72FD54FBFE1FD3" box="[1039,1141,679,704]" pageId="9" pageNumber="160" type="female">1 female</specimenCount>
: “
<collectingCountry id="4A9976208069FFF87EF4FD5BFB681FD3" box="[1161,1251,679,704]" name="Mexico" pageId="9" pageNumber="160">México</collectingCountry>
,
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, Edo. de
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,
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<collectingDate id="5674E9988069FFF87B2AFD30FE341FF6" box="[343,447,716,741]" pageId="9" pageNumber="160" value="1967-06-25">25/VI/67</collectingDate>
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, col.
<collectorName id="9F7B53668069FFF87B87FD30FD2C1FF6" box="[506,679,716,741]" pageId="9" pageNumber="160">G. y V. Halffter</collectorName>
, cebo excremento humano” (
<collectionCode id="549FAE758069FFF87990FD30FBB81FF7" box="[1005,1075,716,740]" pageId="9" pageNumber="160">VMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="323136B08069FFF87ABAFD13FF741E3F" blockId="9.[151,1436,644,1101]" pageId="9" pageNumber="160">
<emphasis id="00FAEAA28069FFF87ABAFD13FDA01E1B" bold="true" box="[199,555,751,776]" pageId="9" pageNumber="160">Non-type material examined.</emphasis>
<materialsCitation id="82E63CED8069FFF87848FD0CFF701E3F" ID-GBIF-Occurrence="2234218970" collectingDate="1990-08-17" collectionCode="CEMT" collectorName="A. Morales" country="Mexico" location="Tejamaniles" pageId="9" pageNumber="160" specimenCount="1" specimenCount-female="1" stateProvince="Michoacan de Ocampo">
<specimenCount id="2488FD398069FFF87848FD0CFD141E1B" box="[565,671,751,776]" pageId="9" pageNumber="160" type="female">1 female</specimenCount>
: “
<collectingCountry id="4A9976208069FFF878C5FD13FC991E1B" box="[696,786,751,776]" name="Mexico" pageId="9" pageNumber="160">México</collectingCountry>
,
<collectingRegion id="F04AF8528069FFF8795CFD13FC2A1E1B" box="[801,929,751,776]" country="Mexico" name="Michoacan de Ocampo" pageId="9" pageNumber="160">Michoacán</collectingRegion>
,
<location id="3751606B8069FFF879CDFD13FBB11E1B" LSID="urn:lsid:plazi:treatment:BA2787A68069FFF87AEAFE4BFDC4195F:3751606B8069FFF879CDFD13FBB11E1B" box="[944,1082,751,776]" country="Mexico" name="Tejamaniles" pageId="9" pageNumber="160" stateProvince="Michoacan de Ocampo">Tejamaniles</location>
,
<date id="463010708069FFF87E34FD0CFB4F1E1B" box="[1097,1220,752,776]" pageId="9" pageNumber="160" value="1990-08-17">
<collectingDate id="5674E9988069FFF87E34FD0CFB4F1E1B" box="[1097,1220,752,776]" pageId="9" pageNumber="160" value="1990-08-17">17/VIII/90</collectingDate>
</date>
.
<collectorName id="9F7B53668069FFF87EAFFD0CFADC1E1B" box="[1234,1367,751,776]" pageId="9" pageNumber="160">A. Morales</collectorName>
, col.” (
<collectionCode id="549FAE758069FFF87AE2FCE8FF781E3F" box="[159,243,788,812]" pageId="9" pageNumber="160">CEMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7A94653B8069FFF87ABAFCCBFDC4195F" pageId="9" pageNumber="160" type="discussion">
<paragraph id="323136B08069FFF87ABAFCCBFDC4195F" blockId="9.[151,1436,644,1101]" pageId="9" pageNumber="160">
<emphasis id="00FAEAA28069FFF87ABAFCCBFEB01E43" bold="true" box="[199,315,823,848]" pageId="9" pageNumber="160">Remarks.</emphasis>
Length: 9.1 ±
<quantity id="F5769B558069FFF87B92FCC4FDC71E5C" box="[495,588,824,848]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="160" unit="mm" value="0.1">0.1 mm</quantity>
. The following set of characters help us to support the new status of
<taxonomicName id="F58E4D338069FFF87FFCFCC4FE821E67" authorityName="Zunino &amp; Halffter" authorityYear="1988" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="canescens">
<emphasis id="00FAEAA28069FFF87FFCFCC4FE821E67" italics="true" pageId="9" pageNumber="160">O. canescens</emphasis>
</taxonomicName>
(
<tableCitation id="7F0C030B8069FFF87B65FCA0FEE61E67" box="[280,365,860,885]" captionStart="TABLE 1" captionStartId="7.[147,233,637,661]" captionText="TABLE 1. List of diagnostic characters for the former subspecies of Onthophagus fuscus." pageId="9" pageNumber="160">Table 1</tableCitation>
): pronotal pubescence medially absent (densely covering the entire surface in
<taxonomicName id="F58E4D338069FFF87E95FCA0FAC01E67" authorityName="Boucomont" authorityYear="1932" box="[1256,1355,860,884]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="00FAEAA28069FFF87E95FCA0FAC01E67" box="[1256,1355,860,884]" italics="true" pageId="9" pageNumber="160">O fuscus</emphasis>
</taxonomicName>
); short elytral pubescence (long elytral pubescence in
<taxonomicName id="F58E4D338069FFF878CBFC7CFCA91E8B" authorityName="Boucomont" authorityYear="1932" box="[694,802,896,920]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="00FAEAA28069FFF878CBFC7CFCA91E8B" box="[694,802,896,920]" italics="true" pageId="9" pageNumber="160">O. fuscus</emphasis>
</taxonomicName>
); major male frontal carina completely impressed (not impressed laterally in
<taxonomicName id="F58E4D338069FFF87BEEFC58FE771EAF" authorityName="Boucomont" authorityYear="1932" box="[403,508,932,956]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="00FAEAA28069FFF87BEEFC58FE771EAF" box="[403,508,932,956]" italics="true" pageId="9" pageNumber="160">O. fuscus</emphasis>
</taxonomicName>
); major male pronotal prominence obtusely trapezoidal (obtusely triangular in
<taxonomicName id="F58E4D338069FFF87FFCFC58FF561EF3" authorityName="Boucomont" authorityYear="1932" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="00FAEAA28069FFF87FFCFC58FF561EF3" italics="true" pageId="9" pageNumber="160">O. fuscus</emphasis>
</taxonomicName>
); female frontal carina complete, rounded (not impressed laterally, trapezoidal in
<taxonomicName id="F58E4D338069FFF87E0EFC34FB571EF3" authorityName="Boucomont" authorityYear="1932" box="[1139,1244,968,992]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="00FAEAA28069FFF87E0EFC34FB571EF3" box="[1139,1244,968,992]" italics="true" pageId="9" pageNumber="160">O. fuscus</emphasis>
</taxonomicName>
); apical-superior area of the parameres trapezoidal (obtuse in
<taxonomicName id="F58E4D338069FFF878F1FC10FD7E1917" authorityName="Boucomont" authorityYear="1932" box="[652,757,1004,1028]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="00FAEAA28069FFF878F1FC10FD7E1917" box="[652,757,1004,1028]" italics="true" pageId="9" pageNumber="160">O. fuscus</emphasis>
</taxonomicName>
); apex of the inferior right lobe concave (claw shaped in
<taxonomicName id="F58E4D338069FFF87FFCFC10FF56193B" authorityName="Boucomont" authorityYear="1932" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="00FAEAA28069FFF87FFCFC10FF56193B" italics="true" pageId="9" pageNumber="160">O. fuscus</emphasis>
</taxonomicName>
).
<taxonomicName id="F58E4D338069FFF87A84FBECFD86193B" authorityName="Zunino &amp; Halffter" authorityYear="1988" box="[249,525,1040,1064]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="canescens">
<emphasis id="00FAEAA28069FFF87A84FBECFD86193B" box="[249,525,1040,1064]" italics="true" pageId="9" pageNumber="160">Onthophagus canescens</emphasis>
</taxonomicName>
occurs between
<quantity id="F5769B558069FFF878AAFBECFCFC1934" box="[727,887,1040,1064]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="9" pageNumber="160" unit="m" value="1900.0" valueMax="2100.0" valueMin="1700.0">1700–2100 m</quantity>
in La Sierra de Mil Cumbres, central-western Trans-Mexican Volcanic Belt (
<figureCitation id="AAB52A358069FFF87B93FBC8FDC9195F" box="[494,578,1076,1101]" captionStart="FIGURE 17" captionStartId="25.[151,250,1249,1271]" captionTargetBox="[207,1386,404,1237]" captionTargetId="figure@25.[207,1386,404,1237]" captionTargetPageId="25" captionText="FIGURE 17. Onthophagus fuscus species group distribution. Species distributions were determined according to specimen labels and specialized literature (Bates 1886–1890; Zunino &amp; Halffter 1988; Delgado &amp; Capistan 1996; Navarrete-Heredia et al. 2001; Zunino 2003; Moctezuma et al. 2016)." httpUri="https://zenodo.org/record/2624251/files/figure.png" pageId="9" pageNumber="160">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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