The Onthophagus fuscus (Coleoptera: Scarabaeidae) species complex: an update and the description of a new species Author Joaqui, Tatiana Author Moctezuma, Victor Author Sánchez-Huerta, José Luis Author Escobar, Federico text Zootaxa 2019 2019-02-13 4555 2 151 186 journal article 27553 10.11646/zootaxa.4555.2.1 95c8b580-1685-41d2-9853-1a0b5b83c327 1175-5326 2624219 E8F6B60F-3E7B-4BD1-938C-F46A840AA08D Onthophagus canescens Zunino & Halffter, 1988 , new rank Figs. 3–4 , 18–20 Onthophagus fuscus canescens Zunino & Halffter 1988 : 83 –85; Navarrete-Heredia & Galindo 1997 : 3 ; Anduaga 2000 : 130 ; Krajcik 2006 : 106 ; Pulido-Herrera & Zunino 2007 : 103 ; Moctezuma et al. 2016 : 33 , 35–36. Type material examined ( 1 male , 2 females ). Paratypes . 1 male , 1 female : “ México , Parque Nacional J. M. Morelos , Morelia , Michoacán , 25/VII/64 , P. Reyes C. col., en hongos” ( GHC ) ; 1 female : “ México , Valle de Bravo , Edo. de México , 25/VI/67 , col. G. y V. Halffter , cebo excremento humano” ( VMC ) . Non-type material examined. 1 female : “ México , Michoacán , Tejamaniles , 17/VIII/90 . A. Morales , col.” ( CEMT ) . Remarks. Length: 9.1 ± 0.1 mm . The following set of characters help us to support the new status of O. canescens ( Table 1 ): pronotal pubescence medially absent (densely covering the entire surface in O fuscus ); short elytral pubescence (long elytral pubescence in O. fuscus ); major male frontal carina completely impressed (not impressed laterally in O. fuscus ); major male pronotal prominence obtusely trapezoidal (obtusely triangular in O. fuscus ); female frontal carina complete, rounded (not impressed laterally, trapezoidal in O. fuscus ); apical-superior area of the parameres trapezoidal (obtuse in O. fuscus ); apex of the inferior right lobe concave (claw shaped in O. fuscus ). Onthophagus canescens occurs between 1700–2100 m in La Sierra de Mil Cumbres, central-western Trans-Mexican Volcanic Belt ( Fig. 17 ).