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<document id="A3F4E8AC84F0498E778248D857D6575E" ID-CLB-Dataset="298664" ID-DOI="10.3897/contrib.entomol.74.e107520" ID-GBIF-Dataset="a711c138-a6c3-46bd-b28e-a82cf000b0b8" ID-ZooBank="E314C311-AE79-4679-8EB6-99B63B4E8965" ID-publisher-id="107520" URI-arpha="691666F4-F906-58ED-B199-0AF6E963CEEE" URI-zoobank="http://zoobank.org/E314C311-AE79-4679-8EB6-99B63B4E8965" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1718792832460" checkinUser="pensoft" docAuthor="Jusoh, Wan F. A. &amp; Ballantyne, Lesley" docDate="2024" docId="AA9657FE72055D4485EF12F9718FB390" docLanguage="en" docName="Contributions to Entomology 74(1): 63-80" docOrigin="Contributions to Entomology 74 (1)" docSource="https://contributions-to-entomology.arphahub.com/article/107520/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Luciola nicollieri Bugnion 1922" docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="80" masterDocId="691666F4F90658EDB1990AF6E963CEEE" masterDocTitle="A catalogue and redescription of type specimens of fireflies (Coleoptera, Lampyridae, Luciolinae) deposited in Naturalis Biodiversity Center, Leiden" masterLastPageNumber="80" masterPageNumber="63" pageNumber="63" updateTime="1718822341776" updateUser="ExternalLinkService">
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<mods:title id="30C31B6EAA976696E8FDDCF87FD7E5C3">A catalogue and redescription of type specimens of fireflies (Coleoptera, Lampyridae, Luciolinae) deposited in Naturalis Biodiversity Center, Leiden</mods:title>
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<mods:name id="231A568267A1E19DDD261C1F48EC09D5" type="personal">
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<mods:roleTerm id="A86038E482371D590CD84BD29FCC7BD4">Author</mods:roleTerm>
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<mods:namePart id="A3CF21206283E490EE9A21EE3BAE4E54">Jusoh, Wan F. A.</mods:namePart>
<mods:nameIdentifier id="B1F0E0279D9FBF17E486F9C0CFB9A6B4" type="ORCID">0000-0002-2995-8429</mods:nameIdentifier>
<mods:affiliation id="5B89D5C02384F511FAD9703794463A83">School of Science, Monash University Malaysia, Bandar Sunway 47500, Selangor, Malaysia</mods:affiliation>
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<mods:namePart id="BAF815C042EED3C04C05191D2A7F3FCF">Ballantyne, Lesley</mods:namePart>
<mods:nameIdentifier id="2819398CFCC19A30A8C37121723930A1" type="ORCID">0000-0002-2029-3918</mods:nameIdentifier>
<mods:affiliation id="C6FAE966825CEADC420945EDD9EA3024">School of Agricultural, Environmental and Veterinary Sciences, Charles Sturt University, Wagga Wagga 2678, Australia</mods:affiliation>
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<mods:title id="E87FA406CF87E74C6BFD56662FFF601B">Contributions to Entomology</mods:title>
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<mods:part id="8475A8F034646E882F704C5B6898D41B">
<mods:date id="275CE0BA96A67875AE8470B4315C7650">2024</mods:date>
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<mods:number id="78A4CEB5081CA19F4B8A18B93307DDEA">74</mods:number>
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<treatment id="AA9657FE72055D4485EF12F9718FB390" ID-DOI="http://doi.org/10.5281/zenodo.12162256" ID-GBIF-Taxon="233233893" ID-Zenodo-Dep="12162256" ID-arpha="AA9657FE-7205-5D44-85EF-12F9718FB390" LSID="urn:lsid:plazi:treatment:AA9657FE72055D4485EF12F9718FB390" httpUri="http://treatment.plazi.org/id/AA9657FE72055D4485EF12F9718FB390">
<subSubSection id="3F23498D97BABEA384569BF03A3FEFEA" type="nomenclature">
<paragraph id="25895216D22763037730EE3EF257F301">
<taxonomicName id="FDD866C6385F205887A10960D53B37E1" authority="Bugnion, 1922" authorityName="Bugnion" authorityYear="1922" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nicollieri">
<emphasis id="419255E998AC54BD50956F0E70EECF66" italics="true">Luciola nicollieri</emphasis>
<bibRefCitation id="987B98469165F572BAEA9980394A9789" author="Bugnion E" refId="B20" refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 1 - 14." year="1922">Bugnion, 1922</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="95B25DB63884909DF784AE02EC5FCBD5" type="description">
<paragraph id="7C4DD9CD6790929D84819FF8E0495206">
<figureCitation id="57EF77458039E7034552C12FD56DFE89" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">Fig. 4 A Q</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="71490223A2C7A473EA2870947EF2B0E2" type="reference_group">
<paragraph id="59A455ED4C01A0D15FDC438E26759DC4">
<treatmentCitationGroup id="8F98AC89C6297FCB1C0F9D4EBD2ACC3A">
<treatmentCitation id="814B7A075799B7D097E783170A1AF13D" author="Bugnion E" page="14" year="1922">
<taxonomicName id="86B391EAA3DFD73D884142ACA8D3119C" authority="Bugnion, 1922: 1 - 14" authorityName="Bugnion" authorityPageNumber="1 - 14" authorityYear="1922" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nicollieri">
<emphasis id="56589330E166F859AEA7324FCBAE3C23" italics="true">Luciola nicollieri</emphasis>
<comment id="B20C2708D982097843E0AF4F28DD159D">
<bibRefCitation id="289BB6A3DFAB6544AE4CFEE32367D529" author="Bugnion E" refId="B20" refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 1 - 14." year="1922">Bugnion, 1922: 114</bibRefCitation>
.
<treatmentCitation id="56CC82F22EBAF516E124298088344D2B" author="Ballantyne LA &amp; Lambkin CL &amp; Ho J-Z &amp; Jusoh WFA &amp; Nada B &amp; Nak-Eiam S &amp; Thancharoen A &amp; Wattanachaiyingcharoen W &amp; Yiu V" page="103" year="2019">
<bibRefCitation id="19D1E4A10060CCEB0AEE49F437AC3DF8" DOI="10.11646/zootaxa.4687.1.1" author="Ballantyne LA &amp; Lambkin CL &amp; Ho J-Z &amp; Jusoh WFA &amp; Nada B &amp; Nak-Eiam S &amp; Thancharoen A &amp; Wattanachaiyingcharoen W &amp; Yiu V" etAl="et al." firstAuthor="Ballantyne" journalOrPublisher="Zootaxa" page="103" pagination="1-174" refId="B14" refString="Ballantyne LA, Lambkin CL, Ho J-Z, Jusoh WFA, Nada B, Nak-Eiam S, Thancharoen A, Wattanachaiyingcharoen W, Yiu V (2019) The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species. Zootaxa 4687: 1174. https://doi.org/10.11646/zootaxa.4687.1.1" title="The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species." volume="4687" year="2019">Ballantyne et al. 2019: 103</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="FBB949731869B9A0ED77ADFE88160E63" author="De Silva DR &amp; Wijekoon CD &amp; Wegiriya HE &amp; Bandara SN &amp; Madushanka TI" page="331" year="2023">
<bibRefCitation id="BFEFFAC4EDBB2C58F5D5FCD9EF7983A1" DOI="10.52547/jibs.9.2.331" author="De Silva DR &amp; Wijekoon CD &amp; Wegiriya HE &amp; Bandara SN &amp; Madushanka TI" etAl="et al." firstAuthor="De Silva" journalOrPublisher="Journal of Insect Biodiversity and Systematics" page="331" pagination="331-341" refId="B25" refString="De Silva DR, Wijekoon CD, Wegiriya HE, Bandara SN, Madushanka TI (2023) Re-description of Luciola nicollieri Bugnion, 1922 (Lampyridae, Luciolinae) in Sri Lanka. Journal of Insect Biodiversity and Systematics 9: 331341. https://doi.org/10.52547/jibs.9.2.331" title="Re-description of Luciola nicollieri Bugnion, 1922 (Lampyridae, Luciolinae) in Sri Lanka." volume="9" year="2023">De Silva et al. 2023: 331</bibRefCitation>
</treatmentCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="B88BC5EC06C2059A65E77F0EA0198665">
<treatmentCitationGroup id="C1AC0FB76D98875317258369A5E1F17D">
<treatmentCitation id="F66DE4016F1087DFCDF47BF4C4D57C02" author="McDermott FA" page="110" year="1966">
<taxonomicName id="9D6FB01C0096EDB55E0215D82A1A6460" ID-CoL="3WCTX" authorityName="Bugnion" authorityYear="1922" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nicolleri">
<emphasis id="7698285E1FE034E0E6E10EAABB1ED09E" italics="true">Luciola nicolleri</emphasis>
</taxonomicName>
<comment id="F534950AB0AD3A84E5C122F042274C48">
.
<bibRefCitation id="E79524DCFC8A601A4EEAF68540DD3E1D" author="McDermott FA" editor="Steel WO" firstAuthor="McDermott" journalOrPublisher="Uitgeverij Dr. W. Junk, The Hague ( s-Gravenhage), the Netherlands" page="110" pagination="1-149" refId="B50" refString="McDermott FA (1966) Lampyridae. In: Steel WO (Ed.) Coleopterorum Catalogus Supplementa, pars 9. Uitgeverij Dr. W. Junk, The Hague ( s-Gravenhage), the Netherlands, 1149. [Luciolini, 98118]" title="Lampyridae." volumeTitle="Coleopterorum Catalogus Supplementa, pars 9." year="1966">McDermott 1966: 110</bibRefCitation>
(misspelling).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EU6BG" type="Lectotype and paralectotype">
<paragraph id="13F7AFAECC7FB976411D86A7F7E5FAA0">
<heading id="0DEA8BFC52040CE0A154E6E013686DC1" reason="title">
<typeStatus id="5EC31DA3FC902D4E73F0B08341C02063" type="lectotype">Lectotype</typeStatus>
and
<typeStatus id="CDA336DFF01B3F0ECD9B599F8BE9D06B" type="paralectotype">paralectotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="D2B3823E72F2978A1F6042A813FE5AA7">
<specimenCount id="AB90AE5DF676E974E329600AFC3010D5" count="2" type="male">2 ♂</specimenCount>
(herein designated).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZ6BG" type="type locality">
<paragraph id="6F71BEE9BDD37790ACFF12A2894E7274">
<heading id="FFA0D1CC970569CA84D95D579BF2B43A" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="62B55A89D30C72BA9E4F70AAB4A28B34">
<collectingCountry id="8895DDB533219D2E3D382C817C00E91F" name="Sri Lanka">Ceylan</collectingCountry>
, Talgaswella, district dElpitiya ”.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E56BG" type="material">
<paragraph id="D5E1CB20116CB2319C4DCC2B1982D774">
<heading id="594D19828CAEAE1DEB91F0BFEC624C8A" reason="title">Material examined</heading>
</paragraph>
<paragraph id="00FE423477EAF2646BAF298076C4F4DD">
<emphasis id="3C8D80EC5957B97145C16585E900213E" bold="true">
(
<specimenCount id="1BD16D34046AFC191DFB36D5533378E9" count="2" type="male">2 ♂</specimenCount>
specimens).
<materialsCitation id="303F7D8EC3D258366F40C3D4C080E701" ID-GBIF-Occurrence="4895467313" collectionCode="RMNH" collectorName="E. BUGNION" country="Sri Lanka" location="Talgaswella" municipality="Hiver" specimenCode="INS / 968348" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="0C2C2B0A476BF52429D58CCD4140DFF0" bold="true" italics="true">
<typeStatus id="1092AF025A7554FB97D2BE270C54DCFE" type="lectotype">Lectotype</typeStatus>
</emphasis>
(herein designated):
<collectingCountry id="DBADB92CB0915AC683816D21FAE40CB5" name="Sri Lanka">SRI LANKA</collectingCountry>
<specimenCount id="766C9487A971E13212849337C151F0AD" count="1" type="male"></specimenCount>
: (1) “
<collectorName id="91985EE296C4BBC4400C304BDB22C6EB">E. BUGNION</collectorName>
/
<collectingMunicipality id="17DE8C90338538D8FD8466A1A03BEFA2">Hiver</collectingMunicipality>
19067 / Ceylan /
<location id="0D47AADAFC8B53C8495609F1F5E27DE7" LSID="urn:lsid:plazi:treatment:AA9657FE72055D4485EF12F9718FB390:0D47AADAFC8B53C8495609F1F5E27DE7" country="Sri Lanka" municipality="Hiver" name="Talgaswella">Talgaswella</location>
”; (2) “ don. E. BUGNION 22 ”; (3) “ COTYPE: Luciola / Nicollieri / Bugnion / ♂ ”; (4) “ Luciola. / nicollieri /
<bibRefCitation id="2047BFFC892F9CCB7F636FD5E62C5F79" author="Bugnion E" refId="B20" refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 1 - 14." year="1922">Bugnion, 1922</bibRefCitation>
/ ZMAN type COLE. 0929.1 ”; (5) “
<named-content id="A343A343D09258172415E36F562E318F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-natural-history-naturalis" xlink_title="Naturalis Biodiversity Center, Leiden">
<collectionCode id="3B095862AA2793D1D6F446E1D8303B90" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</named-content>
.
<specimenCode id="EBD715357FD9F467068749951092F8A0">INS / 968348</specimenCode>
” (Fig.
<figureCitation id="283487C12EBB2626C95DF19850964476" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 A</figureCitation>
)
</materialsCitation>
</emphasis>
;
<materialsCitation id="BF49B43C7ADCB500D89A0E26F31225E6" ID-GBIF-Occurrence="4895467310" collectionCode="RMNH" collectorName="E. BUGNION" country="Sri Lanka" location="Talgaswella" municipality="Hiver" specimenCode="INS / 968347" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<typeStatus id="C7DD0E6A69060DC773C78E703F0DD6DB" type="paralectotype">Paralectotype</typeStatus>
:
<collectingCountry id="7856CDEA5012B240940C7A5E9A393DE2" name="Sri Lanka">SRI LANKA</collectingCountry>
<specimenCount id="F61D27FC1768DE3461A874D72FD8F10D" count="1" type="male"></specimenCount>
(missing head and prothorax): (1) “
<collectorName id="5D6ACEA78DF9931021411D10483C7917">E. BUGNION</collectorName>
/
<collectingMunicipality id="7F21ADCFD6809E28767DE4A37C1A48EA">Hiver</collectingMunicipality>
19067 / Ceylan /
<location id="12F54F20B8F5FCA0083FEA3BBF4A802E" LSID="urn:lsid:plazi:treatment:AA9657FE72055D4485EF12F9718FB390:12F54F20B8F5FCA0083FEA3BBF4A802E" country="Sri Lanka" municipality="Hiver" name="Talgaswella">Talgaswella</location>
”; (2) “ don. E. BUGNION 22 ”; (3) “ COTYPE: / Luciola / Nicollieri / Bugnion / ♂ ”; (4) “ Luciola / nicollieri /
<bibRefCitation id="A5D42C33854E3FE2F571B12A4ADDBE59" author="Bugnion E" refId="B20" refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 1 - 14." year="1922">Bugnion, 1922</bibRefCitation>
/ ZMAN type / COLE. 0929.2 ”; (5) “
<named-content id="B4AB4A33D5B52112574C34D163478D1E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-natural-history-naturalis" xlink_title="Naturalis Biodiversity Center, Leiden">
<collectionCode id="E78805A5A091770BBD527A9B784DED80" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</named-content>
.
<specimenCode id="7A8DA0AE7C02C7B73C49C81EC6F6BDC9">INS / 968347</specimenCode>
” (Fig.
<figureCitation id="71A859DE21E5BFAC9853ABE57FDB0FF0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 B</figureCitation>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDCAI" type="diagnosis">
<paragraph id="0DB9D49387AF5A53F95942A8B0C45C4C">
<heading id="ABC797BB5224BBC56F6149F861D13F5B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="65A411CC5980FB38B353874B4B2198DA">
Male with orange pronotum, black elytra with narrow pale orange lateral and sutural margins, elytral apex appearing more widely pale due in part to an accumulation of fat body. Venter black except for yellowish creamy
<abbrev id="ABBRID0EJCAI" xlink_title="light organ">LO</abbrev>
in V 6, 7. The only
<taxonomicName id="385ED02F2E422D472F9E5B3823C4DA7E" authorityName="Laporte" authorityYear="1833" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="7E23BADDB5CC26B3CB87A1CB818D6F64" italics="true">Luciola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2D65D4A335D8A1B4F96B0EAA976D3DEF" sensu="stricto">s. str.</taxonomicNameLabel>
so far recorded with pale coloured pronotum without darker markings, and dark brown to black elytra with all margins pale except at the base.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUCAI" type="redescription">
<paragraph id="D41304F75AB81D632279384E809F207F">
<heading id="72489EED4438333237F177D4FCD78536" reason="title">
Redescription of
<typeStatus id="3685CB86A14883CF8D2F05A85EFFD26B" type="lectotype">lectotype</typeStatus>
male.
</heading>
</paragraph>
<paragraph id="A1BF128A3457A7F29CE581A2DFA702AA">
<emphasis id="75F244FD64F4346B469D383A2B9DDA8D" bold="true" italics="true">Body length</emphasis>
(Fig.
<figureCitation id="E02CECF1B3AFCC26F455DF36D3CFB4A8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 D</figureCitation>
).
<quantity id="633B217288834F4C937CF44674606C8E" metricMagnitude="-3" metricUnit="m" metricValue="6.5" unit="mm" value="6.5">6.5 mm</quantity>
long (intact specimen only).
</paragraph>
<caption id="84C267B1D21BD8188C32676EE20DD1DA" ID-DOI="10.3897/contrib.entomol.74.e107520.figure4" ID-arpha="7FA26537-03AF-56F0-A150-4AB4A88AF075" httpUri="https://binary.pensoft.net/fig/1073386" startId="F4">
<paragraph id="12F1884B35FA1115D668BAE1EFDFF49F">
<label id="2276B3B82A7E8F982D0370F644E2DDD0">Figure 4.</label>
</paragraph>
<paragraph id="6320C2DC1903BA5826B420D12F03202A">
<taxonomicName id="D2227B6D22DB8ABEF638DBEBDD420A0C" authority="Bugnion, 1922" authorityName="Bugnion" authorityYear="1922" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nicollieri">
<emphasis id="79461AAE7B8A113B19B0F654D877D44A" italics="true">Luciola nicollieri</emphasis>
<bibRefCitation id="E259A44629A1D4713C47D606BD2722D0" author="Bugnion E" refId="B20" refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 1 - 14." year="1922">Bugnion, 1922</bibRefCitation>
</taxonomicName>
lectotype male (
<emphasis id="9B41D5A67E091867602D7E4F1B6A8E11" bold="true">A</emphasis>
,
<emphasis id="B2E7EFAB2CE3023B431DC967506BD19A" bold="true">C, E, F, H Q</emphasis>
) and paralectotype male (
<emphasis id="783A08DA14FBEF6573DF7B44B8F6ED6F" bold="true">B, D, G.</emphasis>
Without head and prothorax).
<emphasis id="C0EC266094C1D7AE5A747B348BE6E268" bold="true">A, B.</emphasis>
Specimen labels;
<emphasis id="406E5570F12747C0DC5C69D3BCA57EA1" bold="true">C.</emphasis>
Ventral mesothorax end of abdomen;
<emphasis id="91ED3A53E170011AFC709367E73E0B28" bold="true">D.</emphasis>
Dorsal habitus;
<emphasis id="918FE0DAE3C3A7BE104B93FE2EBD2F33" bold="true">E.</emphasis>
Dorsal head prothorax and anterior area of mesothorax;
<emphasis id="D6E2D75ABC02F526B29980FEECDB788D" bold="true">F.</emphasis>
Anterior head;
<emphasis id="C02047BBA3C0F32435BB0948A509DA00" bold="true">G.</emphasis>
Ventral, V 57 and elytral apices;
<emphasis id="503DBBE973E1A11292179047DC806811" bold="true">H.</emphasis>
V 7 ventral;
<emphasis id="0141CCEFB34A4B16456D6CCB703D63DE" bold="true">I, J.</emphasis>
Tergite 8 T 8 dorsal view (I) and ventral view (J);
<emphasis id="5432149150B1DAA61D180E6720E89243" bold="true">K N.</emphasis>
Aedeagal sheath:
<emphasis id="E61B442D3CBDA9E5FEBE16E7AF772270" bold="true">K.</emphasis>
Dorsal with aedeagus ventral surface to right;
<emphasis id="FBCA7458BFE50E8BAD8E9469D039941B" bold="true">L.</emphasis>
Ventral view;
<emphasis id="95CB0C135A079F63F8DF9A2054581447" bold="true">M.</emphasis>
Dorsal view;
<emphasis id="14444A88A6FDC79F506B9DA54FA710BA" bold="true">N.</emphasis>
Left lateral;
<emphasis id="391623818C731E2E3870D980B0BC8326" bold="true">O Q.</emphasis>
Aedeagus:
<emphasis id="1B7BE3A2EDA459D51046FDC7971DF68C" bold="true">O.</emphasis>
Dorsal view (arrow top left indicates thickened left margin of
<abbrev id="ABBRID0EEFAI" xlink_title="median lobe">ML</abbrev>
, lower arrow thickened lobe from inner margin of
<abbrev id="ABBRID0EIFAI" xlink_title="lateral lobes">LL</abbrev>
, lower arrow right side indicates area of attachment to inner surface of base of
<abbrev id="ABBRID0EMFAI" xlink_title="lateral lobes">LL</abbrev>
);
<emphasis id="4FB939E146EBE396BE33565A66E642C8" bold="true">P.</emphasis>
Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of
<abbrev id="ABBRID0ESFAI" xlink_title="lateral lobes">LL</abbrev>
, lower left arrow lateral expansion of
<abbrev id="ABBRID0EWFAI" xlink_title="median lobe">ML</abbrev>
margins);
<emphasis id="62EDED232D3F608EBB61CC0E8CFC7DD7" bold="true">Q.</emphasis>
Left lateral (upper arrow indicates junction between lobes from
<abbrev id="ABBRID0E3FAI" xlink_title="median lobe">ML</abbrev>
to left and lobes from
<abbrev id="ABBRID0EAGAI" xlink_title="lateral lobes">LL</abbrev>
to right; lower right arrow indicates area of attachment to inner base of
<abbrev id="ABBRID0EEGAI" xlink_title="lateral lobes">LL</abbrev>
). All images are to scale, except specimen labels.
</paragraph>
</caption>
<paragraph id="CC8EE3A7AD9FB7D67CFB775FCC570790">
<emphasis id="D2AB67B9208A3FFFF01C616D3598A4B9" bold="true" italics="true">Colour</emphasis>
(Fig.
<figureCitation id="6A28980766D22A6CE5501EBE29225AAB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 C G</figureCitation>
). Pronotum orange with faint thin black line visible from above along lateral margin and around anterolateral corners (not visible in figures);
<abbrev id="ABBRID0EYGAI" xlink_title="mesoscutellum">MS</abbrev>
,
<abbrev id="ABBRID0E3GAI" xlink_title="mesonotal plates">MN</abbrev>
very light brown; elytra very dark brown with epipleural ridge (from above) appearing narrowly paler brown; apical paler fat body extending narrowly anteriorly for 0.9 elytral length along lateral margin, scooped in median area, extending anteriorly 1 / 10 elytral length along suture; remainder of suture indistinctly slightly paler than rest of elytron; head between black eyes black; antennae and palpi dark brown; venter of thorax and basal abdominal ventrites black; legs 1, 2 with coxae, trochanters light brown, remainder very dark brown; legs 3 entirely very dark brown except for small light brown area where inner margins of coxae are contiguous;
<abbrev id="ABBRID0EAHAI" xlink_title="light organ">LO</abbrev>
in V 6, 7 orange with posterior margin of both yellowish; T 68 yellow semi transparent with underlying fat bodies visible; T 35 dark brown; laterally reflexed margins of V 35 dark brown, of 6, 7 yellow.
</paragraph>
<paragraph id="DFEEC88B8648289CE1F9FF5CE432746C">
<emphasis id="BEDF5A9A4D6AA8270C48685D531C618B" bold="true" italics="true">Pronotum</emphasis>
(Fig.
<figureCitation id="921F0163EE17D15033AA2661A6FFE119" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 E</figureCitation>
). Width slightly exceeds humeral width.
</paragraph>
<paragraph id="68E06BC75F650F735B71CC5322E41F7E">
<emphasis id="1C58A0C4E819D0A4CEE290A95279732D" bold="true" italics="true">Elytra</emphasis>
(Fig.
<figureCitation id="E63E502C2B079BB283B7823B53242A92" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 D</figureCitation>
). Interstitial lines not obvious.
</paragraph>
<paragraph id="DD7057DC950522145D5AD6FACEEFD874">
<emphasis id="81EA2D91DF0AF0F3FAD86AF309992843" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="76B89B57EA90FD61D5C5BDE0A7C64DFD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 F</figureCitation>
). Antennal sockets contiguous; head wider than width of prothoracic cavity; mouthparts well developed, and specimen could feed as adult. Antennae longer than, but less than twice
<abbrev id="ABBRID0EFIAI" xlink_title="greatest head width">GHW</abbrev>
, all flagellar segments elongate slender.
</paragraph>
<paragraph id="7A21007C45AA2F0A71A2A7EC642976A5">
<emphasis id="27322975776A16D9CA8CEC9CA3B202B9" bold="true" italics="true">Abdomen</emphasis>
(Fig.
<figureCitation id="ED033F59369592E01609FB24269D4A0C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 C, G J</figureCitation>
).
<abbrev id="ABBRID0EUIAI" xlink_title="light organ">LO</abbrev>
completely occupies V 6, and possibly also V 7 (Fig.
<figureCitation id="6F878A41DFDC7BC633C2471552E511E3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 C</figureCitation>
); posterior margin of V 7 broadly rounded; posterior margin of T 7 entire (Fig.
<figureCitation id="188274E2014A7F210DAE105B37A7A760" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 G, H</figureCitation>
), not emarginated, with corners rounded; posterior margin of T 8 entire, not emarginated, lateral margins slightly divergent posteriorly, anterolateral prolongations narrow, apically acute and 0.4 as long as entire tergite; ventral surface smooth without ridges or flanges (Fig.
<figureCitation id="F5AA5D207981177E9ECFAFBF94FF2FE6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 I, J</figureCitation>
).
</paragraph>
<paragraph id="96A38ED8CD77A8B0F313D8E2C056278A">
<emphasis id="3242237CC3F9646809AB0C40CAC9B3EB" bold="true" italics="true">Aedeagal sheath</emphasis>
(Fig.
<figureCitation id="55F80A66B302416B542BB932D8529F01" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 K N</figureCitation>
). Slightly asymmetrical as right margin of sternite is narrowly emarginated (Fig.
<figureCitation id="34675601A4B544063C4930A96F9093BF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 L</figureCitation>
); anterior margin sternite slightly produced on right side (Fig.
<figureCitation id="EE1214314BFACBA281E179D5D4AED026" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 L, M, N</figureCitation>
); posterior margin entire, rounded, hairy, extending only a little beyond the rounded hairy tergite apex (Fig.
<figureCitation id="816E22B180FF05C9BDBCDB6FFA1AF738" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 N</figureCitation>
); sheath tergite in two sections (Fig.
<figureCitation id="C0F07CC0E43E8ABD9CADD38BE1EC945E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 K, L, N</figureCitation>
); anterior margin of tergite irregularly produced (Fig.
<figureCitation id="7F1D6C7A1BBE2DFDE3255F69A0E95483" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 L, M, N</figureCitation>
).
</paragraph>
<paragraph id="247F9A78C2ED08B81428347670CFC5F4">
<emphasis id="68EA7F0C39EFF54F8296AF5759CC1F08" bold="true" italics="true">Aedeagus</emphasis>
(Fig.
<figureCitation id="E418F5B288FB473ED389C2BAB79AF8A5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 O Q</figureCitation>
). L /
<geoCoordinate id="A05A6E672D7AB0D122A9619D0705DC46" degrees="2.5" direction="west" orientation="longitude" precision="5555" value="-2.5">W 2.5</geoCoordinate>
;
<abbrev id="ABBRID0EALAI" xlink_title="basal piece">BP</abbrev>
narrow extending along sides of
<abbrev id="ABBRID0EELAI" xlink_title="lateral lobes">LL</abbrev>
for slightly less than half aedeagal length, extent somewhat confused by underlying tissue (Fig.
<figureCitation id="D622D6554AD61B72A1DAB7B9D43B8576" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 O, P, Q</figureCitation>
); anterior dorsal margin of
<abbrev id="ABBRID0EMLAI" xlink_title="lateral lobes">LL</abbrev>
neither emarginated nor produced (Fig.
<figureCitation id="707C420B425BCC635DFFD0B34F5251FE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 O</figureCitation>
);
<abbrev id="ABBRID0EULAI" xlink_title="lateral lobes">LL</abbrev>
contiguous along basal 1 / 3 of their dorsal length, then with a slight separation before apices approach in median line (Fig.
<figureCitation id="81D1DEBCA692AD005AA4916FDC82017D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 O</figureCitation>
); inner preapical area of right
<abbrev id="ABBRID0E3LAI" xlink_title="lateral lobes">LL</abbrev>
hooked (unclear if this is also on the other lobe), apices
<abbrev id="ABBRID0EAMAI" xlink_title="lateral lobes">LL</abbrev>
expanded, membranous, wrap around on ventral surface beside
<abbrev id="ABBRID0EEMAI" xlink_title="median lobe">ML</abbrev>
apex, not covering apex (Fig.
<figureCitation id="351A9995D68326E47F3BA3E90BE29EA9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 P</figureCitation>
); elongate slender, apically acute leaf like lobes present along outer ventral margins, extending behind
<abbrev id="ABBRID0EMMAI" xlink_title="median lobe">ML</abbrev>
(Fig.
<figureCitation id="0CB7CA01204C6D89EFED93FF27892477" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 P</figureCitation>
upper arrows); when viewed from beneath
<abbrev id="ABBRID0EUMAI" xlink_title="median lobe">ML</abbrev>
subparallel-sided along basal 4 / 5 then abruptly narrowed (Fig.
<figureCitation id="9311990A1FCA1871C26F44A35D21D548" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 P</figureCitation>
); sides of
<abbrev id="ABBRID0E3MAI" xlink_title="median lobe">ML</abbrev>
just before narrowed area narrowly expanded laterally on ventral surface, incline slightly below; (these narrowed lateral expansions partly contribute to the narrowed appearance) (Fig.
<figureCitation id="F50118CE8C7FEE566365A458ED6A2DD3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 P, Q</figureCitation>
; lower single arrow on left indicates the lateral expansion on the right side only);
<abbrev id="ABBRID0EENAI" xlink_title="median lobe">ML</abbrev>
only slightly expanded at its rounded tip;
<abbrev id="ABBRID0EINAI" xlink_title="median lobe">ML</abbrev>
when viewed from the side same width along most of length until the area above the lateral expansions of its ventral surface, when it expands to its apex (Fig.
<figureCitation id="AC16FE9FD8C6E4621474FAE14B4597FF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 Q</figureCitation>
). Attachment of
<abbrev id="ABBRID0EQNAI" xlink_title="median lobe">ML</abbrev>
to
<abbrev id="ABBRID0EUNAI" xlink_title="lateral lobes">LL</abbrev>
(Fig.
<figureCitation id="6934F1B4231C54CA27DC9D5DD78E7612" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 O, Q</figureCitation>
): lateral margins of anterior dorsal
<abbrev id="ABBRID0E3NAI" xlink_title="median lobe">ML</abbrev>
thickened, darkened, extend obliquely dorsally (Fig.
<figureCitation id="78077166F99A717BB342FC0B9599EA8D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 O</figureCitation>
upper left arrow) to connect (upper arrow Fig.
<figureCitation id="6EBFE4D61444D440163D1B09EC461BD7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 Q</figureCitation>
shows area of attachment between the two sets of lobes), with similar thickened, darkened paired lobes arising from inner basal margin of
<abbrev id="ABBRID0EIOAI" xlink_title="lateral lobes">LL</abbrev>
(Fig.
<figureCitation id="B079BCA7BC39F4866CF13FEE223D6250" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 O</figureCitation>
lower left arrow), immediately behind anterior margin of
<abbrev id="ABBRID0EQOAI" xlink_title="lateral lobes">LL</abbrev>
(right arrow Fig.
<figureCitation id="EE3436282E02D442FD59D36F2D396B11" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 O</figureCitation>
) (Fig.
<figureCitation id="9F5E0FD1A243E7B7A12B47300888D4A8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 O, Q</figureCitation>
); mid anterior margin of
<abbrev id="ABBRID0E3OAI" xlink_title="lateral lobes">LL</abbrev>
heavily sclerotised and darkened, inclining dorsally so in lateral view the dorsal anterior
<abbrev id="ABBRID0EAPAI" xlink_title="lateral lobes">LL</abbrev>
margin appears concave (Fig.
<figureCitation id="F030707BC6860D8E16449C0B69AF2E7E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 57 and elytral apices; H. V 7 ventral; I, J. Tergite 8 T 8 dorsal view (I) and ventral view (J); K N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels." figureDoi="10.3897/contrib.entomol.74.e107520.figure4" httpUri="https://binary.pensoft.net/fig/1073386">4 Q</figureCitation>
lower left arrow); connection between the two areas probably muscle as attachment appears to permit some independent movement of the
<abbrev id="ABBRID0EIPAI" xlink_title="median lobe">ML</abbrev>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMPAI" type="notes">
<paragraph id="A567F8964B7CDBCB5451A4EEC1C16335">
<heading id="8A53F7930C01E6A3270D56B7C4949993" reason="title">Notes.</heading>
</paragraph>
<paragraph id="18B1C6FBF99E78F3DE3D880A2BF10575">
<bibRefCitation id="ED863BA1447BFF9542BB2F2278DFB41D" author="Bugnion E" firstAuthor="Bugnion" refId="B20" refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 114." year="1922">Bugnion (1922)</bibRefCitation>
described two “ côtes ”, but interstitial lines were not clearly visible on this specimen. He considered
<taxonomicName id="317DBE13F56BE0026DBDC1C3F196FA45" authorityName="Bugnion" authorityYear="1922" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nicollieri">
<emphasis id="9036E7E2E7114FDB75218A9D82F39DF8" italics="true">nicollieri</emphasis>
</taxonomicName>
most closely resembled
<taxonomicName id="F0BFA8DF6A93CD16F628FF039B2FBC55" authorityName="Bourgeois" authorityYear="1905" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="horni">
<emphasis id="F62F6444AF3D16EDF1CB30CCBBEAF6A8" italics="true">Luciola horni</emphasis>
</taxonomicName>
and attempted to distinguish the two species (see
<bibRefCitation id="2AFEF7A094D5FBBC0B26D5681049F64D" author="Bugnion E" firstAuthor="Bugnion" page="2" refId="B20" refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 114." year="1922">Bugnion 1922: 2</bibRefCitation>
).
<bibRefCitation id="2FC9BC3A35DF7BDD6FDBE9AFEFA69170" author="Yiu V" firstAuthor="Yiu" journalOrPublisher="Lampyrid" pagination="67-119" refId="B77" refString="Yiu V (2017) A study of Rhagophthalmidae and Lampyridae in Hong Kong with descriptions of new species (Coleoptera): Part 2. Lampyrid 4: 67119." title="A study of Rhagophthalmidae and Lampyridae in Hong Kong with descriptions of new species (Coleoptera): Part 2." volume="4" year="2017">Yiu (2017)</bibRefCitation>
identified as near nicollieri, a population of males, brachelytral females, and larvae with laterally explanate tergal margins from
<collectingCountry id="EE3F8277EC1ECD53E06798C8919F0ECD" name="Hong Kong">Hong Kong</collectingCountry>
. The specimens Yiu illustrated are inconsistent with what we describe here, as the
<abbrev id="ABBRID0E6QAI" xlink_title="median lobe">ML</abbrev>
of the aedeagus narrows to its apex.
<bibRefCitation id="6AAA5ED6A1F4179F6984D212F7FD97C5" DOI="10.11646/zootaxa.4687.1.1" author="Ballantyne LA &amp; Lambkin CL &amp; Ho J-Z &amp; Jusoh WFA &amp; Nada B &amp; Nak-Eiam S &amp; Thancharoen A &amp; Wattanachaiyingcharoen W &amp; Yiu V" etAl="et al." firstAuthor="Ballantyne" journalOrPublisher="Zootaxa" pagination="1-174" refId="B14" refString="Ballantyne LA, Lambkin CL, Ho J-Z, Jusoh WFA, Nada B, Nak-Eiam S, Thancharoen A, Wattanachaiyingcharoen W, Yiu V (2019) The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species. Zootaxa 4687: 1174. https://doi.org/10.11646/zootaxa.4687.1.1" title="The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species." volume="4687" year="2019">Ballantyne et al. (2019)</bibRefCitation>
listed
<taxonomicName id="196D9D0714C502DCFE87DA7377666B03" authorityName="Bugnion" authorityYear="1922" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nicollieri">
<emphasis id="F34DC8F0FE46E65BB13F29FB758A3C88" italics="true">L. nicollieri</emphasis>
</taxonomicName>
under
<taxonomicName id="05B96CBCB78632E4F2B82A2FC676B4F1" authorityName="Laporte" authorityYear="1833" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="55676ECE147F10082D794931AE9848BA" italics="true">Luciola</emphasis>
</taxonomicName>
s. lato and type not located, as they felt the distinctive colour pattern would allow subsequent association of specimens.
<bibRefCitation id="3E97948DB3FC4D3BAD79414117827359" DOI="10.52547/jibs.9.2.331" author="De Silva DR &amp; Wijekoon CD &amp; Wegiriya HE &amp; Bandara SN &amp; Madushanka TI" etAl="et al." firstAuthor="De Silva" journalOrPublisher="Journal of Insect Biodiversity and Systematics" pagination="331-341" refId="B25" refString="De Silva DR, Wijekoon CD, Wegiriya HE, Bandara SN, Madushanka TI (2023) Re-description of Luciola nicollieri Bugnion, 1922 (Lampyridae, Luciolinae) in Sri Lanka. Journal of Insect Biodiversity and Systematics 9: 331341. https://doi.org/10.52547/jibs.9.2.331" title="Re-description of Luciola nicollieri Bugnion, 1922 (Lampyridae, Luciolinae) in Sri Lanka." volume="9" year="2023">De Silva et al. (2023)</bibRefCitation>
described a specimen from
<collectingCountry id="9D9AD5CA8955621B110032FD4796BCAE" name="Sri Lanka">Sri Lanka</collectingCountry>
basing their identification on Ballantynes comparison with these dissections. It is difficult to reconcile their figure 3 d-f with what we illustrate here. They also associated a brachelytral and possibly flightless female with the male.
</paragraph>
</subSubSection>
</treatment>
</document>