<documentid="A3F4E8AC84F0498E778248D857D6575E"ID-CLB-Dataset="298664"ID-DOI="10.3897/contrib.entomol.74.e107520"ID-GBIF-Dataset="a711c138-a6c3-46bd-b28e-a82cf000b0b8"ID-ZooBank="E314C311-AE79-4679-8EB6-99B63B4E8965"ID-publisher-id="107520"URI-arpha="691666F4-F906-58ED-B199-0AF6E963CEEE"URI-zoobank="http://zoobank.org/E314C311-AE79-4679-8EB6-99B63B4E8965"XM.bibliography_approvedBy="admin"XM.materialsCitations_approvedBy="admin"XM.treatmentCitations_approvedBy="admin"XM.treatments_approvedBy="admin"article-type="research-article"checkinTime="1718792832460"checkinUser="pensoft"docAuthor="Jusoh, Wan F. A. & Ballantyne, Lesley"docDate="2024"docId="AA9657FE72055D4485EF12F9718FB390"docLanguage="en"docName="Contributions to Entomology 74(1): 63-80"docOrigin="Contributions to Entomology 74 (1)"docSource="https://contributions-to-entomology.arphahub.com/article/107520/download/xml/"docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic"docStyleName="PensoftTaxPub.0000.journal_article.generic"docTitle="Luciola nicollieri Bugnion 1922"docType="treatment"docVersion="5"dtd-version="3.0"lastPageNumber="80"masterDocId="691666F4F90658EDB1990AF6E963CEEE"masterDocTitle="A catalogue and redescription of type specimens of fireflies (Coleoptera, Lampyridae, Luciolinae) deposited in Naturalis Biodiversity Center, Leiden"masterLastPageNumber="80"masterPageNumber="63"pageNumber="63"updateTime="1718822341776"updateUser="ExternalLinkService">
<mods:titleid="30C31B6EAA976696E8FDDCF87FD7E5C3">A catalogue and redescription of type specimens of fireflies (Coleoptera, Lampyridae, Luciolinae) deposited in Naturalis Biodiversity Center, Leiden</mods:title>
<mods:affiliationid="5B89D5C02384F511FAD9703794463A83">School of Science, Monash University Malaysia, Bandar Sunway 47500, Selangor, Malaysia</mods:affiliation>
<mods:affiliationid="C6FAE966825CEADC420945EDD9EA3024">School of Agricultural, Environmental and Veterinary Sciences, Charles Sturt University, Wagga Wagga 2678, Australia</mods:affiliation>
<figureCitationid="57EF77458039E7034552C12FD56DFE89"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">Fig. 4 A – Q</figureCitation>
<bibRefCitationid="289BB6A3DFAB6544AE4CFEE32367D529"author="Bugnion E"refId="B20"refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 1 - 14."year="1922">Bugnion, 1922: 1–14</bibRefCitation>
.
<treatmentCitationid="56CC82F22EBAF516E124298088344D2B"author="Ballantyne LA & Lambkin CL & Ho J-Z & Jusoh WFA & Nada B & Nak-Eiam S & Thancharoen A & Wattanachaiyingcharoen W & Yiu V"page="103"year="2019">
<bibRefCitationid="19D1E4A10060CCEB0AEE49F437AC3DF8"DOI="10.11646/zootaxa.4687.1.1"author="Ballantyne LA & Lambkin CL & Ho J-Z & Jusoh WFA & Nada B & Nak-Eiam S & Thancharoen A & Wattanachaiyingcharoen W & Yiu V"etAl="et al."firstAuthor="Ballantyne"journalOrPublisher="Zootaxa"page="103"pagination="1-174"refId="B14"refString="Ballantyne LA, Lambkin CL, Ho J-Z, Jusoh WFA, Nada B, Nak-Eiam S, Thancharoen A, Wattanachaiyingcharoen W, Yiu V (2019) The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species. Zootaxa 4687: 1–174. https://doi.org/10.11646/zootaxa.4687.1.1"title="The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species."volume="4687"year="2019">Ballantyne et al. 2019: 103</bibRefCitation>
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<treatmentCitationid="FBB949731869B9A0ED77ADFE88160E63"author="De Silva DR & Wijekoon CD & Wegiriya HE & Bandara SN & Madushanka TI"page="331"year="2023">
<bibRefCitationid="BFEFFAC4EDBB2C58F5D5FCD9EF7983A1"DOI="10.52547/jibs.9.2.331"author="De Silva DR & Wijekoon CD & Wegiriya HE & Bandara SN & Madushanka TI"etAl="et al."firstAuthor="De Silva"journalOrPublisher="Journal of Insect Biodiversity and Systematics"page="331"pagination="331-341"refId="B25"refString="De Silva DR, Wijekoon CD, Wegiriya HE, Bandara SN, Madushanka TI (2023) Re-description of Luciola nicollieri Bugnion, 1922 (Lampyridae, Luciolinae) in Sri Lanka. Journal of Insect Biodiversity and Systematics 9: 331–341. https://doi.org/10.52547/jibs.9.2.331"title="Re-description of Luciola nicollieri Bugnion, 1922 (Lampyridae, Luciolinae) in Sri Lanka."volume="9"year="2023">De Silva et al. 2023: 331</bibRefCitation>
<bibRefCitationid="E79524DCFC8A601A4EEAF68540DD3E1D"author="McDermott FA"editor="Steel WO"firstAuthor="McDermott"journalOrPublisher="Uitgeverij Dr. W. Junk, The Hague (‘ s-Gravenhage), the Netherlands"page="110"pagination="1-149"refId="B50"refString="McDermott FA (1966) Lampyridae. In: Steel WO (Ed.) Coleopterorum Catalogus Supplementa, pars 9. Uitgeverij Dr. W. Junk, The Hague (‘ s-Gravenhage), the Netherlands, 1–149. [Luciolini, 98–118]"title="Lampyridae."volumeTitle="Coleopterorum Catalogus Supplementa, pars 9."year="1966">McDermott 1966: 110</bibRefCitation>
(misspelling).
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</paragraph>
</subSubSection>
<subSubSectionid="SECID0EU6BG"type="Lectotype and paralectotype">
<bibRefCitationid="2047BFFC892F9CCB7F636FD5E62C5F79"author="Bugnion E"refId="B20"refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 1 - 14."year="1922">Bugnion, 1922</bibRefCitation>
/ ZMAN type COLE. 0929.1 ”; (5) “
<named-contentid="A343A343D09258172415E36F562E318F"content-type="dwc:institutional_code"xlink_href="http://grbio.org/institution/national-museum-natural-history-naturalis"xlink_title="Naturalis Biodiversity Center, Leiden">
<collectionCodeid="3B095862AA2793D1D6F446E1D8303B90"country="Netherlands"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992"lsid="urn:lsid:biocol.org:col:34992"name="National Museum of Natural History, Naturalis"type="Museum">RMNH</collectionCode>
<figureCitationid="283487C12EBB2626C95DF19850964476"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 A</figureCitation>
<bibRefCitationid="A5D42C33854E3FE2F571B12A4ADDBE59"author="Bugnion E"refId="B20"refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 1 - 14."year="1922">Bugnion, 1922</bibRefCitation>
/ ZMAN type / COLE. 0929.2 ”; (5) “
<named-contentid="B4AB4A33D5B52112574C34D163478D1E"content-type="dwc:institutional_code"xlink_href="http://grbio.org/institution/national-museum-natural-history-naturalis"xlink_title="Naturalis Biodiversity Center, Leiden">
<collectionCodeid="E78805A5A091770BBD527A9B784DED80"country="Netherlands"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992"lsid="urn:lsid:biocol.org:col:34992"name="National Museum of Natural History, Naturalis"type="Museum">RMNH</collectionCode>
<figureCitationid="71A859DE21E5BFAC9853ABE57FDB0FF0"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 B</figureCitation>
Male with orange pronotum, black elytra with narrow pale orange lateral and sutural margins, elytral apex appearing more widely pale due in part to an accumulation of fat body. Venter black except for yellowish creamy
<figureCitationid="E02CECF1B3AFCC26F455DF36D3CFB4A8"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 D</figureCitation>
<figureCitationid="6A28980766D22A6CE5501EBE29225AAB"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 C – G</figureCitation>
). Pronotum orange with faint thin black line visible from above along lateral margin and around anterolateral corners (not visible in figures);
very light brown; elytra very dark brown with epipleural ridge (from above) appearing narrowly paler brown; apical paler fat body extending narrowly anteriorly for 0.9 elytral length along lateral margin, scooped in median area, extending anteriorly 1 / 10 elytral length along suture; remainder of suture indistinctly slightly paler than rest of elytron; head between black eyes black; antennae and palpi dark brown; venter of thorax and basal abdominal ventrites black; legs 1, 2 with coxae, trochanters light brown, remainder very dark brown; legs 3 entirely very dark brown except for small light brown area where inner margins of coxae are contiguous;
in V 6, 7 orange with posterior margin of both yellowish; T 6–8 yellow semi – transparent with underlying fat bodies visible; T 3–5 dark brown; laterally reflexed margins of V 3–5 dark brown, of 6, 7 yellow.
<figureCitationid="921F0163EE17D15033AA2661A6FFE119"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 E</figureCitation>
<figureCitationid="E63E502C2B079BB283B7823B53242A92"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 D</figureCitation>
<figureCitationid="76B89B57EA90FD61D5C5BDE0A7C64DFD"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 F</figureCitation>
). Antennal sockets contiguous; head wider than width of prothoracic cavity; mouthparts well developed, and specimen could feed as adult. Antennae longer than, but less than twice
<abbrevid="ABBRID0EFIAI"xlink_title="greatest head width">GHW</abbrev>
<figureCitationid="ED033F59369592E01609FB24269D4A0C"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 C, G – J</figureCitation>
completely occupies V 6, and possibly also V 7 (Fig.
<figureCitationid="6F878A41DFDC7BC633C2471552E511E3"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 C</figureCitation>
); posterior margin of V 7 broadly rounded; posterior margin of T 7 entire (Fig.
<figureCitationid="188274E2014A7F210DAE105B37A7A760"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 G, H</figureCitation>
), not emarginated, with corners rounded; posterior margin of T 8 entire, not emarginated, lateral margins slightly divergent posteriorly, anterolateral prolongations narrow, apically acute and 0.4 as long as entire tergite; ventral surface smooth without ridges or flanges (Fig.
<figureCitationid="F5AA5D207981177E9ECFAFBF94FF2FE6"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 I, J</figureCitation>
<figureCitationid="55F80A66B302416B542BB932D8529F01"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 K – N</figureCitation>
). Slightly asymmetrical as right margin of sternite is narrowly emarginated (Fig.
<figureCitationid="34675601A4B544063C4930A96F9093BF"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 L</figureCitation>
); anterior margin sternite slightly produced on right side (Fig.
<figureCitationid="EE1214314BFACBA281E179D5D4AED026"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 L, M, N</figureCitation>
); posterior margin entire, rounded, hairy, extending only a little beyond the rounded hairy tergite apex (Fig.
<figureCitationid="816E22B180FF05C9BDBCDB6FFA1AF738"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 N</figureCitation>
); sheath tergite in two sections (Fig.
<figureCitationid="C0F07CC0E43E8ABD9CADD38BE1EC945E"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 K, L, N</figureCitation>
); anterior margin of tergite irregularly produced (Fig.
<figureCitationid="7F1D6C7A1BBE2DFDE3255F69A0E95483"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 L, M, N</figureCitation>
<figureCitationid="E418F5B288FB473ED389C2BAB79AF8A5"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 O – Q</figureCitation>
for slightly less than half aedeagal length, extent somewhat confused by underlying tissue (Fig.
<figureCitationid="D622D6554AD61B72A1DAB7B9D43B8576"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 O, P, Q</figureCitation>
<figureCitationid="707C420B425BCC635DFFD0B34F5251FE"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 O</figureCitation>
contiguous along basal 1 / 3 of their dorsal length, then with a slight separation before apices approach in median line (Fig.
<figureCitationid="81D1DEBCA692AD005AA4916FDC82017D"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 O</figureCitation>
<figureCitationid="351A9995D68326E47F3BA3E90BE29EA9"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 P</figureCitation>
); elongate slender, apically acute leaf like lobes present along outer ventral margins, extending behind
<figureCitationid="0CB7CA01204C6D89EFED93FF27892477"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 P</figureCitation>
subparallel-sided along basal 4 / 5 then abruptly narrowed (Fig.
<figureCitationid="9311990A1FCA1871C26F44A35D21D548"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 P</figureCitation>
just before narrowed area narrowly expanded laterally on ventral surface, incline slightly below; (these narrowed lateral expansions partly contribute to the narrowed appearance) (Fig.
<figureCitationid="F50118CE8C7FEE566365A458ED6A2DD3"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 P, Q</figureCitation>
; lower single arrow on left indicates the lateral expansion on the right side only);
when viewed from the side same width along most of length until the area above the lateral expansions of its ventral surface, when it expands to its apex (Fig.
<figureCitationid="AC16FE9FD8C6E4621474FAE14B4597FF"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 Q</figureCitation>
<figureCitationid="6934F1B4231C54CA27DC9D5DD78E7612"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 O, Q</figureCitation>
<figureCitationid="78077166F99A717BB342FC0B9599EA8D"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 O</figureCitation>
upper left arrow) to connect (upper arrow Fig.
<figureCitationid="6EBFE4D61444D440163D1B09EC461BD7"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 Q</figureCitation>
shows area of attachment between the two sets of lobes), with similar thickened, darkened paired lobes arising from inner basal margin of
<figureCitationid="B079BCA7BC39F4866CF13FEE223D6250"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 O</figureCitation>
lower left arrow), immediately behind anterior margin of
<figureCitationid="EE3436282E02D442FD59D36F2D396B11"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 O</figureCitation>
) (Fig.
<figureCitationid="9F5E0FD1A243E7B7A12B47300888D4A8"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 O, Q</figureCitation>
<figureCitationid="F030707BC6860D8E16449C0B69AF2E7E"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Luciola nicollieri Bugnion, 1922 lectotype male (A, C, E, F, H – Q) and paralectotype male (B, D, G. Without head and prothorax). A, B. Specimen labels; C. Ventral mesothorax – end of abdomen; D. Dorsal habitus; E. Dorsal head prothorax and anterior area of mesothorax; F. Anterior head; G. Ventral, V 5–7 and elytral apices; H. V 7 ventral; I, J. Tergite 8 – T 8 dorsal view (I) and ventral view (J); K – N. Aedeagal sheath: K. Dorsal with aedeagus ventral surface to right; L. Ventral view; M. Dorsal view; N. Left lateral; O – Q. Aedeagus: O. Dorsal view (arrow top left indicates thickened left margin of ML, lower arrow thickened lobe from inner margin of LL, lower arrow right side indicates area of attachment to inner surface of base of LL); P. Ventral view (upper oblique arrows left and right indicate leafy lobes from inner margins of LL, lower left arrow lateral expansion of ML margins); Q. Left lateral (upper arrow indicates junction between lobes from ML to left and lobes from LL to right; lower right arrow indicates area of attachment to inner base of LL). All images are to scale, except specimen labels."figureDoi="10.3897/contrib.entomol.74.e107520.figure4"httpUri="https://binary.pensoft.net/fig/1073386">4 Q</figureCitation>
lower left arrow); connection between the two areas probably muscle as attachment appears to permit some independent movement of the
<bibRefCitationid="ED863BA1447BFF9542BB2F2278DFB41D"author="Bugnion E"firstAuthor="Bugnion"refId="B20"refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 1–14."year="1922">Bugnion (1922)</bibRefCitation>
described two “ côtes ”, but interstitial lines were not clearly visible on this specimen. He considered
<bibRefCitationid="2AFEF7A094D5FBBC0B26D5681049F64D"author="Bugnion E"firstAuthor="Bugnion"page="2"refId="B20"refString="Bugnion E (1922) Luciola nicollieri sp. n. de Ceylan. Spolia Zeylandica XII: 1–14."year="1922">Bugnion 1922: 2</bibRefCitation>
).
<bibRefCitationid="2FC9BC3A35DF7BDD6FDBE9AFEFA69170"author="Yiu V"firstAuthor="Yiu"journalOrPublisher="Lampyrid"pagination="67-119"refId="B77"refString="Yiu V (2017) A study of Rhagophthalmidae and Lampyridae in Hong Kong with descriptions of new species (Coleoptera): Part 2. Lampyrid 4: 67–119."title="A study of Rhagophthalmidae and Lampyridae in Hong Kong with descriptions of new species (Coleoptera): Part 2."volume="4"year="2017">Yiu (2017)</bibRefCitation>
identified as near nicollieri, a population of males, brachelytral females, and larvae with laterally explanate tergal margins from
<bibRefCitationid="6AAA5ED6A1F4179F6984D212F7FD97C5"DOI="10.11646/zootaxa.4687.1.1"author="Ballantyne LA & Lambkin CL & Ho J-Z & Jusoh WFA & Nada B & Nak-Eiam S & Thancharoen A & Wattanachaiyingcharoen W & Yiu V"etAl="et al."firstAuthor="Ballantyne"journalOrPublisher="Zootaxa"pagination="1-174"refId="B14"refString="Ballantyne LA, Lambkin CL, Ho J-Z, Jusoh WFA, Nada B, Nak-Eiam S, Thancharoen A, Wattanachaiyingcharoen W, Yiu V (2019) The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species. Zootaxa 4687: 1–174. https://doi.org/10.11646/zootaxa.4687.1.1"title="The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species."volume="4687"year="2019">Ballantyne et al. (2019)</bibRefCitation>
s. lato and type not located, as they felt the distinctive colour pattern would allow subsequent association of specimens.
<bibRefCitationid="3E97948DB3FC4D3BAD79414117827359"DOI="10.52547/jibs.9.2.331"author="De Silva DR & Wijekoon CD & Wegiriya HE & Bandara SN & Madushanka TI"etAl="et al."firstAuthor="De Silva"journalOrPublisher="Journal of Insect Biodiversity and Systematics"pagination="331-341"refId="B25"refString="De Silva DR, Wijekoon CD, Wegiriya HE, Bandara SN, Madushanka TI (2023) Re-description of Luciola nicollieri Bugnion, 1922 (Lampyridae, Luciolinae) in Sri Lanka. Journal of Insect Biodiversity and Systematics 9: 331–341. https://doi.org/10.52547/jibs.9.2.331"title="Re-description of Luciola nicollieri Bugnion, 1922 (Lampyridae, Luciolinae) in Sri Lanka."volume="9"year="2023">De Silva et al. (2023)</bibRefCitation>
basing their identification on Ballantyne’s comparison with these dissections. It is difficult to reconcile their figure 3 d-f with what we illustrate here. They also associated a brachelytral and possibly flightless female with the male.
