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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.990.55542" ID-GBIF-Dataset="1cede2a7-3880-480c-8401-1015e5440145" ID-PMC="PMC7674391" ID-Pensoft-Pub="1313-2970-990-1" ID-Pensoft-UUID="9F51F78CD53F5005A305DE65494002C4" ID-PubMed="33269011" ID-ZooBank="7B73642C278D40F89091B26213C9A704" ModsDocID="1313-2970-990-1" checkinTime="1605112396458" checkinUser="pensoft" docAuthor="Shimizu, So, Broad, Gavin R. &amp; Maeto, Kaoru" docDate="2020" docId="D38784CF1CA451CBBC205286DEDB654E" docLanguage="en" docName="ZooKeys 990: 1-144" docOrigin="ZooKeys 990" docSource="http://dx.doi.org/10.3897/zookeys.990.55542" docTitle="Enicospilus takakuwai Shimizu 2020, sp. nov." docType="treatment" docUuid="761D3C15-1EDF-4D58-AE38-FA778F7E9087" docUuidSource="ZooBank" docVersion="5" id="9F51F78CD53F5005A305DE65494002C4" lastPageNumber="1" masterDocId="9F51F78CD53F5005A305DE65494002C4" masterDocTitle="Integrative taxonomy and analysis of species richness patterns of nocturnal Darwin wasps of the genus Enicospilus Stephens (Hymenoptera, Ichneumonidae, Ophioninae) in Japan" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1668169845364" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Integrative taxonomy and analysis of species richness patterns of nocturnal Darwin wasps of the genus Enicospilus Stephens (Hymenoptera, Ichneumonidae, Ophioninae) in Japan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shimizu, So</mods:namePart>
<mods:affiliation>Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan &amp; DC and Overseas Challenge Program for Young Researchers, Japan Society for the Promotion of Science, Tokyo, Japan &amp; Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5202-4552</mods:nameIdentifier>
<mods:nameIdentifier type="email">parasitoidwasp.sou@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Broad, Gavin R.</mods:namePart>
<mods:affiliation>Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-7223-5333</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maeto, Kaoru</mods:namePart>
<mods:affiliation>Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>990</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>144</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.990.55542</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.990.55542</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-990-1</mods:identifier>
<mods:identifier type="ZooBank">7B73642C278D40F89091B26213C9A704</mods:identifier>
<mods:identifier type="Pensoft-UUID">9F51F78CD53F5005A305DE65494002C4</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="170296363" LSID="urn:lsid:zoobank.org:act:761D3C15-1EDF-4D58-AE38-FA778F7E9087" httpUri="http://treatment.plazi.org/id/D38784CF1CA451CBBC205286DEDB654E" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="D38784CF-1CA4-51CB-BC20-5286DEDB654E" authority="Shimizu" authorityName="Shimizu" authorityYear="2020" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus takakuwai" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="takakuwai" status="sp. nov.">Enicospilus takakuwai Shimizu</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">Figure 48</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The specific name is derived from the name of the collector of the holotype specimen, a famous Japanese insect taxonomist, Masatoshi Takakuwa.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type series">
<paragraph pageId="0" pageNumber="1">Type series.</paragraph>
<paragraph pageId="0" pageNumber="1">A holotype male only.</paragraph>
<paragraph pageId="0" pageNumber="1">
HT: ♂, Tairoike, Miyake-jima, Izu Iss.,
<normalizedToken originalValue="Tôkyô">Tokyo</normalizedToken>
,
<normalizedToken originalValue="Kantô-Kôshin">Kanto-Koshin</normalizedToken>
, JAPAN, 10-11.VI.2010, M. Takakuwa leg. (KPMNH).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Eastern Palaearctic region.</paragraph>
<paragraph pageId="0" pageNumber="1">
JAPAN: [
<normalizedToken originalValue="Kantô-Kôshin">Kanto-Koshin</normalizedToken>
]
<normalizedToken originalValue="Tôkyô">Tokyo</normalizedToken>
(Miyake-jima Island).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="bionomics">
<paragraph pageId="0" pageNumber="1">Bionomics.</paragraph>
<paragraph pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is more or less similar to
<taxonomicName lsidName="E. laqueatus" pageId="0" pageNumber="1" rank="species" species="laqueatus">
<emphasis italics="true" pageId="0" pageNumber="1">E. laqueatus</emphasis>
</taxonomicName>
, but easily distinguished by the position and shape of the central sclerite: central sclerite positioned in the anterodistal part of the fenestra, smaller and moderately sclerotised in
<taxonomicName lsidName="E. takakuwai" pageId="0" pageNumber="1" rank="species" species="takakuwai">
<emphasis italics="true" pageId="0" pageNumber="1">E. takakuwai</emphasis>
</taxonomicName>
sp. nov., as in Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48F</figureCitation>
, but positioned in the centrodistal part of the fenestra, larger and strongly sclerotised in
<taxonomicName lsidName="E. laqueatus" pageId="0" pageNumber="1" rank="species" species="laqueatus">
<emphasis italics="true" pageId="0" pageNumber="1">E. laqueatus</emphasis>
</taxonomicName>
, as in Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Enicospilus laqueatus (Enderlein, 1921) ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure24" httpUri="https://binary.pensoft.net/fig/473790" pageId="0" pageNumber="1">24F</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Male (HT). Body length ca. 16.5 mm.</paragraph>
<paragraph pageId="0" pageNumber="1">
Head with GOI = 3.2 (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48D</figureCitation>
). Lower face 0.8
<normalizedToken originalValue="×">x</normalizedToken>
as wide as high, strongly shiny, finely punctate with setae (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48B, D</figureCitation>
). Clypeus 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as wide as high, finely punctate with setae, convex in profile, and ventral margin acute (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48B, D</figureCitation>
). Malar space 0.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as basal mandibular width (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48B, D</figureCitation>
). Mandible moderately twisted by 15°, moderately long, evenly narrowed, outer surface with diagonal setose groove (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48B, D</figureCitation>
). Upper tooth of mandible 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as lower (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48B</figureCitation>
). Frons, vertex and gena strongly shiny with fine setae (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48B-D</figureCitation>
). Posterior ocellus touching eye (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48B-D</figureCitation>
). Ventral end of occipital carina joining oral carina. Antennae with 59 flagellomeres; first flagellomere 1.8
<normalizedToken originalValue="×">x</normalizedToken>
as long as second; 20th flagellomere 2.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Mesosoma entirely rather weakly shiny with fine setae (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48E</figureCitation>
). Pronotum fairly extensively diagonally striate (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48E</figureCitation>
). Mesoscutum 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum width, finely punctate, and evenly rounded in profile (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48E</figureCitation>
). Notauli absent (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48E</figureCitation>
). Scutellum with lateral longitudinal carinae along entire length of scutellum. Epicnemium punctate with setae. Epicnemial carina strong, almost straight, inclined to anterior, dorsal end not reaching anterior margin of mesopleuron (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48E</figureCitation>
). Meso- and metapleuron evenly moderately longitudinally striate (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48E</figureCitation>
). Submetapleural carina broadened anteriorly (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48E</figureCitation>
). Propodeum declivous in profile; anterior transverse carina complete; anterior area longitudinally striate; spiracular area smooth; posterior area entirely moderately reticulate; propodeal spiracle elliptical and joining pleural carina by ridge (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Wings (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48F</figureCitation>
). Fore wing length ca. 11.0 mm with AI = 0.4, CI = 0.4, DI = 0.3, ICI = 0.5, SDI = 1.3, SI = 0.1, SRI = 0.3; vein 1m-cu&amp;M evenly weakly curved; vein 2r&amp;RS almost straight; vein RS evenly curved; fenestra and sclerites of discosubmarginal cell as in Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48F</figureCitation>
; proximal sclerite triangular, not confluent with distal sclerite, strongly pigmented; central sclerite D-shaped, moderately sclerotised, and positioned in anterodistal part of fenestra; distal sclerite weakly pigmented; proximal corner of marginal cell rather sparsely setose, but without glabrous area; posterodistal corner of second discal cell ca. 100° and of subbasal cell ca. 90°; vein 1cu-a antefurcal to M&amp;RS by 0.2
<normalizedToken originalValue="×">x</normalizedToken>
length of 1cu-a. Hind wing with NI = 2.3; vein RS straight; vein RA with 7 uniform hamuli.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Legs. Hind leg with coxa in profile 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as deep; basitarsus 1.9
<normalizedToken originalValue="×">x</normalizedToken>
as long as second tarsomere; fourth tarsomere 3.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide; tarsal claw simply pectinate.
</paragraph>
<paragraph pageId="0" pageNumber="1">Metasoma with DMI = 1.4, PI = 3.3, THI = 4.5; thyridium oval.</paragraph>
<paragraph pageId="0" pageNumber="1">
Colour (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Enicospilus takakuwai Shimizu, sp. nov. ♂ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1">48</figureCitation>
). Head and mesosoma entirely yellow-brown except for apex of mandible black. Metasoma dark brown. Wings hyaline. Fore wing sclerites pigmented and amber. Wing veins red-brown to amber.
</paragraph>
<paragraph pageId="0" pageNumber="1">Female. Unknown.</paragraph>
<caption doi="10.3897/zookeys.990.55542.figure48" httpUri="https://binary.pensoft.net/fig/473814" pageId="0" pageNumber="1" start="Figure 48" startId="F48">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 48.</emphasis>
<taxonomicName authorityName="Shimizu" authorityYear="2020" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus takakuwai" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="takakuwai">
<emphasis italics="true" pageId="0" pageNumber="1">Enicospilus takakuwai</emphasis>
</taxonomicName>
Shimizu, sp. nov. ♂ (HT) from Japan
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
head, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
head, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
head, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
mesosoma, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
central part of fore wing.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>