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<document id="B0E97409389D11AB6BB1D84960240FCE" ID-DOI="10.11646/zootaxa.5380.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10212761" ID-ZooBank="2E5011DC-FC41-46D7-A955-04C6694F291B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701163613994" checkinUser="plazi" docAuthor="Moradmand, Majid, Wesal, Mohammad Wasil &amp; Kulkarni, Siddharth" docDate="2023" docId="D33C87F9E322C655FF47F8826D9A8B8B" docLanguage="en" docName="zootaxa.5380.1.6.pdf" docOrigin="Zootaxa 5380 (1)" docSource="https://mapress.com/zt/article/download/zootaxa.5380.1.6/52360" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spariolenus omidvarbrothers Moradmand &amp; Wesal &amp; Kulkarni 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="81" masterDocId="2F05FF81E323C651FFD0FFAA680B8842" masterDocTitle="Taxonomic revision of the troglophile Spariolenus spiders (Araneae: Sparassidae) in South and West Asia" masterLastPageNumber="95" masterPageNumber="77" pageNumber="78" updateTime="1701164107850" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3CC634D3B3C359D5DE999DEDD36C8A46">Taxonomic revision of the troglophile Spariolenus spiders (Araneae: Sparassidae) in South and West Asia</mods:title>
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<mods:namePart id="E9C9F039D08C608CF5DDB3FF26D5DBF3">Moradmand, Majid</mods:namePart>
<mods:nameIdentifier id="E180A62768912745F6252F7ADAE0AD49" type="ORCID">0000-0001-8596-8448</mods:nameIdentifier>
<mods:affiliation id="872C9D74BF824F5403CA093D6E77B271">Department of Plant and Animal Biology, Faculty of Biological Science and Technology, University of Isfahan, Isfahan, Iran. m. moradmand @ sci. ui. ac. ir; moradmand. arachnids @ gmail. com; https: // orcid. org / 0000 - 0001 - 8596 - 8448 Department of Biology, Faculty of Education, University of Ghazni, Ghazni, Afghanistan. &amp; Arachnology, Senckenberg Research Institute, Frankfurt am Main, Germany. &amp; Department of Biogeography, Trier University, Trier, Germany.</mods:affiliation>
<mods:nameIdentifier id="5809600F1B8A856252D6B20EA72CD284" type="email">m.moradmand@sci.ui.ac.ir</mods:nameIdentifier>
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<mods:namePart id="4037911D1DDE63F5CC48C2BCD49ED5D5">Wesal, Mohammad Wasil</mods:namePart>
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<mods:namePart id="55427843C30F19DD4A2F3F6630609EA5">Kulkarni, Siddharth</mods:namePart>
<mods:affiliation id="6E227D4FCC128E97D0DF7D0AD5A640F6">Department of Integrative Biology, University of Wisconsin-Madison, United States.</mods:affiliation>
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<mods:title id="2CFF35448960CC6956353F06F020F003">Zootaxa</mods:title>
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<mods:date id="4B2AFD5B95564FF304AB02B57562B767">2023</mods:date>
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<mods:number id="E9F16ED358708D0BD3818B6D6AB03358">2023-11-28</mods:number>
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<mods:number id="3B487EBB874274E95CB2FE192AB1A1FA">5380</mods:number>
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<treatment id="D33C87F9E322C655FF47F8826D9A8B8B" LSID="urn:lsid:plazi:treatment:D33C87F9E322C655FF47F8826D9A8B8B" httpUri="http://treatment.plazi.org/id/D33C87F9E322C655FF47F8826D9A8B8B" lastPageId="4" lastPageNumber="81" pageId="1" pageNumber="78">
<subSubSection id="138F6564E322C650FF47F8826A498F01" box="[151,578,1832,1859]" pageId="1" pageNumber="78" type="nomenclature">
<paragraph id="5B2A36EFE322C650FF47F8826A498F01" blockId="1.[151,578,1832,1895]" box="[151,578,1832,1859]" pageId="1" pageNumber="78">
<heading id="00628183E322C650FF47F8826A498F01" bold="true" box="[151,578,1832,1859]" fontSize="11" level="1" pageId="1" pageNumber="78" reason="1">
<emphasis id="69E1EAFDE322C650FF47F8826A498F01" bold="true" box="[151,578,1832,1859]" pageId="1" pageNumber="78">
<taxonomicName id="9C954D6CE322C650FF47F88269F38F01" authority="Moradmand &amp; Wesal &amp; Kulkarni, 2023" authorityName="Moradmand &amp; Wesal &amp; Kulkarni" authorityYear="2023" box="[151,504,1832,1859]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="omidvarbrothers" status="sp. nov.">
<emphasis id="69E1EAFDE322C650FF47F88269F38F01" bold="true" box="[151,504,1832,1859]" italics="true" pageId="1" pageNumber="78">Spariolenus omidvarbrothers</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72D25786E322C650FE2FF8836A498F01" box="[511,578,1833,1859]" pageId="1" pageNumber="78" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="138F6564E322C650FF47F8E669F98F25" box="[151,498,1868,1895]" pageId="1" pageNumber="78" type="description">
<paragraph id="5B2A36EFE322C650FF47F8E669F98F25" blockId="1.[151,578,1832,1895]" box="[151,498,1868,1895]" pageId="1" pageNumber="78">
<figureCitation id="C3AE2A6AE322C650FF47F8E6691F8F25" box="[151,276,1868,1895]" captionStart="FIGURES 1" captionStartId="2.[151,264,1801,1825]" captionTargetBox="[166,1420,315,1761]" captionTargetId="figure-75@2.[151,1436,308,1777]" captionTargetPageId="2" captionText="FIGURES 1A–C. Spariolenus omidvarbrothers sp. n., male holotype, Iran (SMF). A–B left palp (A prolateral, B ventral); C Embolus tip, ventral. Abbreviations: BRB—Basal retrolateral bulge, C—Conductor, ET—embolus tip, PET—prolateral part of embolus, RET—retrolateral part of embolus, dRTA—dorsal part of retrolateral tibial apophysis, SP—spermophore, T— tegulum, vRTA—ventral part of retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.10212763" httpUri="https://zenodo.org/record/10212763/files/figure.png" pageId="1" pageNumber="78">Figs 1A–C</figureCitation>
,
<figureCitation id="C3AE2A6AE322C650FECFF8E7696C8F25" box="[287,359,1869,1895]" captionStart="FIGURES 2" captionStartId="3.[151,264,1513,1537]" captionTargetBox="[161,1420,185,1484]" captionTargetId="figure-19@3.[151,1436,181,1489]" captionTargetPageId="3" captionText="FIGURES 2A–D. Spariolenus omidvarbrothers sp. n., female paratype, Iran (SMF). A epigyne, ventral; B vulva, dorsal; C right vulva, lateral; D schematic course of internal duct system, lateral.Abbreviations: CO—Copulatory opening; EF—Epigynal field; EP—Epigynal pit; FC—First coil of vulva; FD—Fertilisation duct; LID—Lumen of internal duct system; MEP —Margin of epigynal pit; SC—Second coil of vulva; TC—Third coil of vulva." figureDoi="http://doi.org/10.5281/zenodo.10212765" httpUri="https://zenodo.org/record/10212765/files/figure.png" pageId="1" pageNumber="78">2A–D</figureCitation>
,
<figureCitation id="C3AE2A6AE322C650FEA2F8E7699C8F25" box="[370,407,1869,1895]" captionStart="FIGURES 9" captionStartId="13.[151,264,1908,1932]" captionTargetBox="[213,1375,181,1885]" captionTargetId="figure-19@13.[213,1375,181,1885]" captionTargetPageId="13" captionText="FIGURES 9A–G. Spariolenus spp. A–D, G frontal view of prosoma; E chelicerae and labium, ventral; F cheliceral dentition, ventral. A Spariolenus omidvarbrothers sp. n., male holotype; B Spariolenus lindbergi (Roewer, 1962) comb. nov., male; C Spariolenus baluchistanicus sp. n., female holotype; D–F Spariolenus bakasura sp. n., female paratype; G Spariolenus kabandha sp. n." figureDoi="http://doi.org/10.5281/zenodo.10212785" httpUri="https://zenodo.org/record/10212785/files/figure.png" pageId="1" pageNumber="78">9A</figureCitation>
,
<figureCitation id="C3AE2A6AE322C650FE72F8E769F98F25" box="[418,498,1869,1895]" captionStart="FIGURES 11" captionStartId="15.[151,264,1906,1930]" captionTargetBox="[159,1427,181,1882]" captionTargetId="figure-19@15.[159,1427,181,1882]" captionTargetPageId="15" captionText="FIGURES 11A–E. A–C Spariolenus omidvarbrothers sp. n. (A habitus of male, B female, C type locality and habitat, Bidan Oasis close to Barashk in Sistan &amp; Baluchistan Province, Iran); D Spariolenus mansourii Moradmand, 2017 preying on a cicada; E habitat of Spariolenus lindbergi (Roewer, 1962) comb. nov. in Afghanistan. Arrows shows the observation location of the specimens." figureDoi="http://doi.org/10.5281/zenodo.10212793" httpUri="https://zenodo.org/record/10212793/files/figure.png" pageId="1" pageNumber="78">11A–C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="138F6564E322C650FF47F83F69598FB5" pageId="1" pageNumber="78" type="materials_examined">
<paragraph id="5B2A36EFE322C650FF47F83F69598FB5" blockId="1.[151,1437,1940,2039]" pageId="1" pageNumber="78">
<emphasis id="69E1EAFDE322C650FF47F83F694D8FEC" bold="true" box="[151,326,1940,1967]" pageId="1" pageNumber="78">Type material.</emphasis>
<materialsCitation id="EBFD3CB2E322C650FE9EF83F6BA68F91" collectingDate="2017-06-05" collectionCode="SMF" collectorName="Sistan &amp; M. Moradmand &amp; H. Salehi" country="Iran" county="Bidan Oasis valley" elevation="1336" latitude="26.61" location="Baluchestan Province" longLatPrecision="744" longitude="60.35" municipality="Barashk" pageId="1" pageNumber="78" specimenCount="1" typeStatus="holotype">
<emphasis id="69E1EAFDE322C650FE9EF83F69CE8FED" bold="true" box="[334,453,1941,1967]" pageId="1" pageNumber="78">
<typeStatus id="842E884DE322C650FE9EF83F69B48FED" box="[334,447,1941,1967]" pageId="1" pageNumber="78" type="holotype">Holotype</typeStatus>
:
</emphasis>
male,
<emphasis id="69E1EAFDE322C650FDC3F83E6BE88FEC" bold="true" box="[531,995,1940,1967]" pageId="1" pageNumber="78">
<collectingCountry id="2382767FE322C650FDC3F83E6A548FEC" box="[531,607,1940,1966]" name="Iran" pageId="1" pageNumber="78">IRAN</collectingCountry>
:
<emphasis id="69E1EAFDE322C650FDBCF83E6BE88FEC" bold="true" box="[620,995,1940,1967]" italics="true" pageId="1" pageNumber="78">
<collectorName id="F6605339E322C650FDBCF83E6ABF8FEC" box="[620,692,1940,1966]" pageId="1" pageNumber="78">Sistan</collectorName>
&amp;
<location id="5E4A6034E322C650FD0AF83E6BD78FEC" LSID="urn:lsid:plazi:treatment:D33C87F9E322C655FF47F8826D9A8B8B:5E4A6034E322C650FD0AF83E6BD78FEC" box="[730,988,1940,1967]" country="Iran" county="Bidan Oasis valley" latitude="26.61" longLatPrecision="744" longitude="60.35" municipality="Barashk" name="Baluchestan Province" pageId="1" pageNumber="78">Baluchestan Province</location>
:
</emphasis>
</emphasis>
<collectingMunicipality id="BB4EAC95E322C650FC3BF83E6C418FEC" box="[1003,1098,1940,1966]" pageId="1" pageNumber="78">Barashk</collectingMunicipality>
,
<collectingCounty id="B24B4E63E322C650FB86F83E6D268FEC" box="[1110,1325,1940,1966]" pageId="1" pageNumber="78">Bidan Oasis valley</collectingCounty>
,
<geoCoordinate id="3EA15028E322C650FAEAF83E6D938FED" box="[1338,1432,1940,1967]" degrees="26.61" direction="north" orientation="latitude" pageId="1" pageNumber="78" precision="555" value="26.61">26.61 N</geoCoordinate>
,
<geoCoordinate id="3EA15028E322C650FF47F81268FB8F91" box="[151,240,1976,2003]" degrees="60.35" direction="east" orientation="longitude" pageId="1" pageNumber="78" precision="555" value="60.35">60.35 E</geoCoordinate>
,
<quantity id="9C6D9B0AE322C650FF2BF812695A8F91" box="[251,337,1976,2003]" metricMagnitude="3" metricUnit="m" metricValue="1.3359999999999999" pageId="1" pageNumber="78" unit="m" value="1336.0">
<elevation id="D0B8D1DCE322C650FF2BF812695A8F91" box="[251,337,1976,2003]" metricMagnitude="3" metricUnit="m" metricValue="1.3359999999999999" pageId="1" pageNumber="78" unit="m" value="1336.0">1336 m</elevation>
</quantity>
,
<date id="2F2B102FE322C650FE8DF81269ED8F91" box="[349,486,1976,2003]" pageId="1" pageNumber="78" value="2017-06-05">
<collectingDate id="3F6FE9C7E322C650FE8DF81269ED8F91" box="[349,486,1976,2003]" pageId="1" pageNumber="78" value="2017-06-05">5 June 2017</collectingDate>
</date>
,
<collectorName id="F6605339E322C650FE20F8136A948F90" box="[496,671,1976,2003]" pageId="1" pageNumber="78">M. Moradmand</collectorName>
&amp;
<collectorName id="F6605339E322C650FD12F8136B228F90" box="[706,809,1976,2003]" pageId="1" pageNumber="78">H. Salehi</collectorName>
leg. (
<collectionCode id="3D84AE2AE322C650FCB8F8136BAD8F91" box="[872,934,1977,2003]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="78">SMF</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="EBFD3CB2E322C650FC67F81369458FB5" collectingDate="2017-06-05" collectionCode="SMF" collectorName="Sistan &amp; M. Moradmand &amp; H. Salehi" country="Iran" county="Bidan Oasis valley" elevation="1336" latitude="26.61" location="Baluchestan Province" longLatPrecision="744" longitude="60.35" municipality="Barashk" pageId="1" pageNumber="78" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="69E1EAFDE322C650FC67F8136C328F91" bold="true" box="[951,1081,1977,2003]" pageId="1" pageNumber="78">
<typeStatus id="842E884DE322C650FC67F8136C388F91" box="[951,1075,1977,2003]" pageId="1" pageNumber="78" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="4D93FD66E322C650FBEFF8136CA18F90" box="[1087,1194,1976,2003]" count="2" pageId="1" pageNumber="78" type="female">2 females</specimenCount>
with same data as for holotype (
<collectionCode id="3D84AE2AE322C650FED8F877694D8FB5" box="[264,326,2013,2039]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="78">SMF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="138F6564E321C653FF17FF3C691F895E" pageId="2" pageNumber="79" type="etymology">
<paragraph id="5B2A36EFE321C653FF17FF3C691F895E" blockId="2.[151,1436,150,284]" pageId="2" pageNumber="79">
<emphasis id="69E1EAFDE321C653FF17FF3C694588F2" bold="true" box="[199,334,150,176]" pageId="2" pageNumber="79">Etymology.</emphasis>
The species is named in honour of the Omidvar brothers, two Iranian adventurers and explorers who travelled around the world in the 1950s and produced a diary and one of the first travel documentaries from extremely remote areas. Their biography inspired the life of many generations including the first author; noun in apposition.
</paragraph>
</subSubSection>
<caption id="0FEA6667E321C653FF47F8A36AF58FCF" ID-DOI="http://doi.org/10.5281/zenodo.10212763" ID-Zenodo-Dep="10212763" httpUri="https://zenodo.org/record/10212763/files/figure.png" pageId="2" pageNumber="79" startId="2.[151,264,1801,1825]" targetBox="[166,1420,315,1761]" targetPageId="2" targetType="figure">
<paragraph id="5B2A36EFE321C653FF47F8A36AF58FCF" blockId="2.[151,1436,1801,1933]" pageId="2" pageNumber="79">
<emphasis id="69E1EAFDE321C653FF47F8A3695F8F63" bold="true" box="[151,340,1801,1825]" pageId="2" pageNumber="79">FIGURES 1A–C.</emphasis>
<taxonomicName id="9C954D6CE321C653FE8BF8A36A8F8F63" authority="Moradmand &amp; Wesal &amp; Kulkarni, 2023" authorityName="Moradmand &amp; Wesal &amp; Kulkarni" authorityYear="2023" box="[347,644,1801,1825]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="omidvarbrothers" status="sp. nov.">
<emphasis id="69E1EAFDE321C653FE8BF8A36A8F8F63" box="[347,644,1801,1825]" italics="true" pageId="2" pageNumber="79">Spariolenus omidvarbrothers</emphasis>
</taxonomicName>
<emphasis id="69E1EAFDE321C653FD5CF8A06ACD8F60" bold="true" box="[652,710,1802,1826]" pageId="2" pageNumber="79">
<taxonomicNameLabel id="72D25786E321C653FD5CF8A06ACD8F60" box="[652,710,1802,1826]" pageId="2" pageNumber="79" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male holotype, Iran (SMF). A–B left palp (A prolateral, B ventral); C Embolus tip, ventral. Abbreviations: BRB—Basal retrolateral bulge, C—Conductor, ET—embolus tip, PET—prolateral part of embolus, RET—retrolateral part of embolus, dRTA—dorsal part of retrolateral tibial apophysis, SP—spermophore, T— tegulum, vRTA—ventral part of retrolateral tibial apophysis.
</paragraph>
</caption>
<caption id="0FEA6667E320C652FF47FA436B458E2F" ID-DOI="http://doi.org/10.5281/zenodo.10212765" ID-Zenodo-Dep="10212765" httpUri="https://zenodo.org/record/10212765/files/figure.png" pageId="3" pageNumber="80" startId="3.[151,264,1513,1537]" targetBox="[161,1420,185,1484]" targetPageId="3" targetType="figure">
<paragraph id="5B2A36EFE320C652FF47FA436B458E2F" blockId="3.[151,1437,1513,1645]" pageId="3" pageNumber="80">
<emphasis id="69E1EAFDE320C652FF47FA43695E8E43" bold="true" box="[151,341,1513,1537]" pageId="3" pageNumber="80">FIGURES 2A–D.</emphasis>
<taxonomicName id="9C954D6CE320C652FE8DFA436A8C8E43" authority="Moradmand &amp; Wesal &amp; Kulkarni, 2023" authorityName="Moradmand &amp; Wesal &amp; Kulkarni" authorityYear="2023" box="[349,647,1513,1537]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="80" phylum="Arthropoda" rank="species" species="omidvarbrothers" status="sp. nov.">
<emphasis id="69E1EAFDE320C652FE8DFA436A8C8E43" box="[349,647,1513,1537]" italics="true" pageId="3" pageNumber="80">Spariolenus omidvarbrothers</emphasis>
</taxonomicName>
<emphasis id="69E1EAFDE320C652FD40FA436AC08E43" bold="true" box="[656,715,1513,1537]" pageId="3" pageNumber="80">
<taxonomicNameLabel id="72D25786E320C652FD40FA436AC08E43" box="[656,715,1513,1537]" pageId="3" pageNumber="80" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, female paratype, Iran (SMF). A epigyne, ventral; B vulva, dorsal; C right vulva, lateral; D schematic course of internal duct system, lateral. Abbreviations: CO—Copulatory opening; EF—Epigynal field; EP—Epigynal pit; FC—First coil of vulva; FD—Fertilisation duct; LID—Lumen of internal duct system; MEP —Margin of epigynal pit; SC—Second coil of vulva; TC—Third coil of vulva.
</paragraph>
</caption>
<subSubSection id="138F6564E320C655FF17F9156C488B1F" lastPageId="4" lastPageNumber="81" pageId="3" pageNumber="80" type="diagnosis">
<paragraph id="5B2A36EFE320C652FF17F91569118FCF" blockId="3.[151,1436,1727,2042]" pageId="3" pageNumber="80">
<emphasis id="69E1EAFDE320C652FF17F91569498E9B" bold="true" box="[199,322,1727,1753]" pageId="3" pageNumber="80">Diagnosis.</emphasis>
Males of the new species share the bifurcated ET with
<taxonomicName id="9C954D6CE320C652FC1FF96A6D728E9B" authority="Moradmand &amp; Jager, 2011" authorityName="Moradmand &amp; Jager" authorityYear="2011" box="[975,1401,1727,1754]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="80" phylum="Arthropoda" rank="species" species="zagros">
<emphasis id="69E1EAFDE320C652FC1FF96A6C3C8E9B" box="[975,1079,1728,1754]" italics="true" pageId="3" pageNumber="80">S. zagros</emphasis>
<bibRefCitation id="3F044B1EE320C652FB91F9156D728E9B" author="Moradmand, M. &amp; Jager, P." box="[1089,1401,1727,1754]" pageId="3" pageNumber="80" pagination="46 - 62" refId="ref7525" refString="Moradmand, M. &amp; Jager, P. (2011) A review on the huntsman spider genus Spariolenus Simon, 1880 (Araneae: Sparassidae: Heteropodinae) in Iran with description of four new species. Zootaxa, 2910, 46 - 62. https: // doi. org / 10.11646 / zootaxa. 2910.1.2" type="journal article" year="2011">Moradmand &amp; Jäger, 2011</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="9C954D6CE320C652FA57F96A69D38EBF" authority="Moradmand, 2017" authorityName="Moradmand" authorityYear="2017" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="80" phylum="Arthropoda" rank="species" species="fathpouri">
<emphasis id="69E1EAFDE320C652FA57F96A68F48EBF" italics="true" pageId="3" pageNumber="80">S. fathpouri</emphasis>
<bibRefCitation id="3F044B1EE320C652FED6F94969D38EBF" author="Moradmand, M." box="[262,472,1763,1789]" pageId="3" pageNumber="80" pagination="11 - 25" refId="ref7477" refString="Moradmand, M. (2017) Three new species of the subterranean huntsman spider genus Spariolenus (Araneae, Sparassidae, Heteropodinae) in Iran. Subterranean Biology, 24, 11 - 25. https: // doi. org / 10.3897 / subtbiol. 24.20985" type="journal article" year="2017">Moradmand, 2017</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="9C954D6CE320C652FE34F94E6A798EBF" authorityName="Moradmand" authorityYear="2017" box="[484,626,1764,1790]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="80" phylum="Arthropoda" rank="species" species="mansourii">
<emphasis id="69E1EAFDE320C652FE34F94E6A798EBF" box="[484,626,1764,1790]" italics="true" pageId="3" pageNumber="80">S. mansourii</emphasis>
</taxonomicName>
, and
<taxonomicName id="9C954D6CE320C652FD7FF94E6B388EBF" authority="Moradmand &amp; Wesal &amp; Kulkarni, 2023" authorityName="Moradmand &amp; Wesal &amp; Kulkarni" authorityYear="2023" box="[687,819,1763,1790]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="80" phylum="Arthropoda" rank="species" species="bakasura" status="sp. n.">
<emphasis id="69E1EAFDE320C652FD7FF94E6B388EBF" box="[687,819,1763,1790]" italics="true" pageId="3" pageNumber="80">S. bakasura</emphasis>
</taxonomicName>
<emphasis id="69E1EAFDE320C652FCEBF9496B718EBF" bold="true" box="[827,890,1763,1789]" pageId="3" pageNumber="80">
<taxonomicNameLabel id="72D25786E320C652FCEBF9496B718EBF" box="[827,890,1763,1789]" pageId="3" pageNumber="80" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
). They can be distinguished from the latter three species by subequal RET and PET (vs. RET obviously longer than PET). They differ from
<taxonomicName id="9C954D6CE320C652FB7AF8A26D198F63" authorityName="Moradmand &amp; Jager" authorityYear="2011" box="[1194,1298,1800,1826]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="80" phylum="Arthropoda" rank="species" species="zagros">
<emphasis id="69E1EAFDE320C652FB7AF8A26D198F63" box="[1194,1298,1800,1826]" italics="true" pageId="3" pageNumber="80">S. zagros</emphasis>
</taxonomicName>
by PET 1/3 of RET in width (vs. similar width of PET and RET in
<taxonomicName id="9C954D6CE320C652FCC2F8866B728F07" authorityName="Moradmand &amp; Jager" authorityYear="2011" box="[786,889,1836,1862]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="80" phylum="Arthropoda" rank="species" species="zagros">
<emphasis id="69E1EAFDE320C652FCC2F8866B728F07" box="[786,889,1836,1862]" italics="true" pageId="3" pageNumber="80">S. zagros</emphasis>
</taxonomicName>
) (
<figureCitation id="C3AE2A6AE320C652FC43F8816BE78F07" box="[915,1004,1835,1861]" captionStart="FIGURES 1" captionStartId="2.[151,264,1801,1825]" captionTargetBox="[166,1420,315,1761]" captionTargetId="figure-75@2.[151,1436,308,1777]" captionTargetPageId="2" captionText="FIGURES 1A–C. Spariolenus omidvarbrothers sp. n., male holotype, Iran (SMF). A–B left palp (A prolateral, B ventral); C Embolus tip, ventral. Abbreviations: BRB—Basal retrolateral bulge, C—Conductor, ET—embolus tip, PET—prolateral part of embolus, RET—retrolateral part of embolus, dRTA—dorsal part of retrolateral tibial apophysis, SP—spermophore, T— tegulum, vRTA—ventral part of retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.10212763" httpUri="https://zenodo.org/record/10212763/files/figure.png" pageId="3" pageNumber="80">Fig. 1C</figureCitation>
). The female epigyne shows a unique MS (widened medially and extended longitudinally,
<figureCitation id="C3AE2A6AE320C652FD2BF8E56B5D8F2B" box="[763,854,1871,1897]" captionStart="FIGURES 2" captionStartId="3.[151,264,1513,1537]" captionTargetBox="[161,1420,185,1484]" captionTargetId="figure-19@3.[151,1436,181,1489]" captionTargetPageId="3" captionText="FIGURES 2A–D. Spariolenus omidvarbrothers sp. n., female paratype, Iran (SMF). A epigyne, ventral; B vulva, dorsal; C right vulva, lateral; D schematic course of internal duct system, lateral.Abbreviations: CO—Copulatory opening; EF—Epigynal field; EP—Epigynal pit; FC—First coil of vulva; FD—Fertilisation duct; LID—Lumen of internal duct system; MEP —Margin of epigynal pit; SC—Second coil of vulva; TC—Third coil of vulva." figureDoi="http://doi.org/10.5281/zenodo.10212765" httpUri="https://zenodo.org/record/10212765/files/figure.png" pageId="3" pageNumber="80">Fig. 2A</figureCitation>
) (vs. MS narrow and barely visible in the rest of congeners).
</paragraph>
<paragraph id="5B2A36EFE320C655FF17F83D6CE5895F" blockId="3.[151,1436,1727,2042]" lastBlockId="4.[151,1437,151,969]" lastPageId="4" lastPageNumber="81" pageId="3" pageNumber="80">
<emphasis id="69E1EAFDE320C652FF17F83D690F8FF3" bold="true" box="[199,260,1943,1969]" pageId="3" pageNumber="80">Male</emphasis>
(
<typeStatus id="842E884DE320C652FEC5F83D69758FF3" box="[277,382,1943,1969]" pageId="3" pageNumber="80" type="holotype">holotype</typeStatus>
):
<emphasis id="69E1EAFDE320C652FE42F8326A378FF3" box="[402,572,1944,1969]" italics="true" pageId="3" pageNumber="80">Measurements.</emphasis>
TL 14.1, PL 6.6, PW 5.7, AW 2.5, OL 7.5, OW 4.5.
<emphasis id="69E1EAFDE320C652FB72F8326CDC8FF3" box="[1186,1239,1944,1969]" italics="true" pageId="3" pageNumber="80">Eyes</emphasis>
(
<figureCitation id="C3AE2A6AE320C652FB37F83D6D418FF3" box="[1255,1354,1943,1969]" captionStart="FIGURES 11" captionStartId="15.[151,264,1906,1930]" captionTargetBox="[159,1427,181,1882]" captionTargetId="figure-19@15.[159,1427,181,1882]" captionTargetPageId="15" captionText="FIGURES 11A–E. A–C Spariolenus omidvarbrothers sp. n. (A habitus of male, B female, C type locality and habitat, Bidan Oasis close to Barashk in Sistan &amp; Baluchistan Province, Iran); D Spariolenus mansourii Moradmand, 2017 preying on a cicada; E habitat of Spariolenus lindbergi (Roewer, 1962) comb. nov. in Afghanistan. Arrows shows the observation location of the specimens." figureDoi="http://doi.org/10.5281/zenodo.10212793" httpUri="https://zenodo.org/record/10212793/files/figure.png" pageId="3" pageNumber="80">Fig 11A</figureCitation>
). AME 0.33, ALE 0.75, PME 0.48, PLE 0.83, eye interdistances: AME-AME 0.18, AME-ALE 0.03, PME-PME 0.17, PME-PLE 0.62, AME-PME 0.21, ALE-PLE 0.57, clypeus-AME 0.71, clypeus-ALE 0.21.
<emphasis id="69E1EAFDE320C652FB54F84A6CFD8FB8" box="[1156,1270,2016,2042]" italics="true" pageId="3" pageNumber="80">Spination.</emphasis>
Palp 131, 101, 2121; Legs: Femur I–III 323(2), IV 321; Patella I–IV 101; Tibia I 131 10, II 232 10,
<date id="2F2B102FE327C655FB9DFF3D6CA088F3" box="[1101,1195,151,177]" pageId="4" pageNumber="81" value="1318-03">III 1318</date>
,
<date id="2F2B102FE327C655FB66FF3D6D1D88F3" box="[1206,1302,151,177]" pageId="4" pageNumber="81" value="2226-04">IV 2226</date>
; Metatarsus
<date id="2F2B102FE327C655FF47FF1169078897" box="[151,268,187,213]" pageId="4" pageNumber="81" value="1014-01" valueMax="1014-03-31" valueMin="1014-01-00">I–III 1014</date>
, IV 3036.
<emphasis id="69E1EAFDE327C655FEAEFF116AD58897" box="[382,734,187,213]" italics="true" pageId="4" pageNumber="81">Measurements of palp and legs.</emphasis>
Palp 9.8 [3.2, 1.3, 2.2, 3.1], I 40.6 [10.8, 3.8, 11.5, 11.7, 2.8], II 46.0 [12.2, 4.1, 13.7, 13.2, 2.8], III 35.2 [10.1, 3.7, 10.2, 9.1, 2.1], IV 36.0 [10.2, 3.1, 10.2, 10.1, 2.4]. Leg formula: II I IV III. Chelicerae with 3 promarginal and 5 retromarginal teeth, and 10 intermarginal denticles.
</paragraph>
<paragraph id="5B2A36EFE327C655FF17FE8D692089EF" blockId="4.[151,1437,151,969]" pageId="4" pageNumber="81">
<emphasis id="69E1EAFDE327C655FF17FE8D690F8903" box="[199,260,295,321]" italics="true" pageId="4" pageNumber="81">Palp.</emphasis>
As in diagnosis, with cymbium slightly longer than tibia, BRB present, vRTA shorter than dRTA, dRTA pointed and vRTA rounded in retrolateral view, PET slightly shorter than RET, and both are long and slim, and not covering each other in ventral view. Conductor hyaline and extending beyond or roughly the same length as ET (
<figureCitation id="C3AE2A6AE327C655FF4FFE39691489EF" box="[159,287,403,429]" captionStart="FIGURES 1" captionStartId="2.[151,264,1801,1825]" captionTargetBox="[166,1420,315,1761]" captionTargetId="figure-75@2.[151,1436,308,1777]" captionTargetPageId="2" captionText="FIGURES 1A–C. Spariolenus omidvarbrothers sp. n., male holotype, Iran (SMF). A–B left palp (A prolateral, B ventral); C Embolus tip, ventral. Abbreviations: BRB—Basal retrolateral bulge, C—Conductor, ET—embolus tip, PET—prolateral part of embolus, RET—retrolateral part of embolus, dRTA—dorsal part of retrolateral tibial apophysis, SP—spermophore, T— tegulum, vRTA—ventral part of retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.10212763" httpUri="https://zenodo.org/record/10212763/files/figure.png" pageId="4" pageNumber="81">Figs 1A–C</figureCitation>
).
</paragraph>
<paragraph id="5B2A36EFE327C655FF17FE1D6A7A89B7" blockId="4.[151,1437,151,969]" pageId="4" pageNumber="81">
<emphasis id="69E1EAFDE327C655FF17FE1D69538993" box="[199,344,439,465]" italics="true" pageId="4" pageNumber="81">Colouration.</emphasis>
Light brown to beige in body with dim to dark grey bands on carapace and legs. Dorsal opisthosoma with chevron shaped markings (
<figureCitation id="C3AE2A6AE327C655FE2DFE716A6E89B7" box="[509,613,475,501]" captionStart="FIGURES 11" captionStartId="15.[151,264,1906,1930]" captionTargetBox="[159,1427,181,1882]" captionTargetId="figure-19@15.[159,1427,181,1882]" captionTargetPageId="15" captionText="FIGURES 11A–E. A–C Spariolenus omidvarbrothers sp. n. (A habitus of male, B female, C type locality and habitat, Bidan Oasis close to Barashk in Sistan &amp; Baluchistan Province, Iran); D Spariolenus mansourii Moradmand, 2017 preying on a cicada; E habitat of Spariolenus lindbergi (Roewer, 1962) comb. nov. in Afghanistan. Arrows shows the observation location of the specimens." figureDoi="http://doi.org/10.5281/zenodo.10212793" httpUri="https://zenodo.org/record/10212793/files/figure.png" pageId="4" pageNumber="81">Fig. 11A</figureCitation>
).
</paragraph>
<paragraph id="5B2A36EFE327C655FF17FE556C1B8AB3" blockId="4.[151,1437,151,969]" pageId="4" pageNumber="81">
<emphasis id="69E1EAFDE327C655FF17FE5569158A5B" bold="true" box="[199,286,511,537]" pageId="4" pageNumber="81">Female</emphasis>
(
<typeStatus id="842E884DE327C655FEFDFE55699E8A5B" box="[301,405,511,537]" pageId="4" pageNumber="81" type="paratype">paratype</typeStatus>
):
<emphasis id="69E1EAFDE327C655FE78FE556A598A5A" box="[424,594,511,536]" italics="true" pageId="4" pageNumber="81">Measurements.</emphasis>
TL 13.5, PL 7.2, PW 6.3, AW 3.6, OL 6.3, OW 4.5.
<emphasis id="69E1EAFDE327C655FB71FE556CD68A5A" box="[1185,1245,511,536]" italics="true" pageId="4" pageNumber="81">Eyes.</emphasis>
AME 0.31, ALE 0.80, PME 0.50, PLE 0.97, eye inter distances:AME-AME 0.18, AME-ALE 0.07, PME-PME 0.27, PME-PLE 0.63, AME-PME 0.32, ALE-PLE 0.71, clypeus-AME 0.87, clypeus-ALE 0.28.
<emphasis id="69E1EAFDE327C655FC3EFDED6D6B8A23" box="[1006,1376,583,609]" italics="true" pageId="4" pageNumber="81">Measurements of palp and legs.</emphasis>
Palp 10.4 [3.1, 1.7, 2.3, 3.3], I 32.8 [9.1, 3.8, 9.6, 8.1, 2.2], II [missing], III 29.6 [8.7, 3.5, 8.2, 7.1, 2.1], IV 31.1 [9.1, 3.6, 8.5, 7.8, 2.1].
<emphasis id="69E1EAFDE327C655FEB7FD2569D18AEB" box="[359,474,655,681]" italics="true" pageId="4" pageNumber="81">Spination.</emphasis>
Palp 131, 101, 2221, 1014; Legs (leg II missing): Femur I, III 323, IV 321; Patella I, III–IV 001; Tibia I 111(10),
<date id="2F2B102FE327C655FE05FD196A3A8A8F" box="[469,561,691,717]" pageId="4" pageNumber="81" value="1118-03">III 1118</date>
,
<date id="2F2B102FE327C655FDEDFD196A968A8F" box="[573,669,691,717]" pageId="4" pageNumber="81" value="2126-04">IV 2126</date>
; Metatarsus I,
<date id="2F2B102FE327C655FC92FD196BAB8A8F" box="[834,928,691,717]" pageId="4" pageNumber="81" value="1014-03">III 1014</date>
, IV 3036. Chelicerae with 3 promarginal and 5 retromarginal teeth, cheliceral furrow with around 20 intermarginal denticles.
</paragraph>
<paragraph id="5B2A36EFE327C655FF17FD516BD08B7B" blockId="4.[151,1437,151,969]" pageId="4" pageNumber="81">
<emphasis id="69E1EAFDE327C655FF17FD5169FA8B56" box="[199,497,763,789]" italics="true" pageId="4" pageNumber="81">Female copulatory organ.</emphasis>
As in diagnosis, with EF wider than long, AB absent, CO small, MEP diagonaly extended (
<figureCitation id="C3AE2A6AE327C655FEDCFCB5696D8B7B" box="[268,358,799,825]" captionStart="FIGURES 2" captionStartId="3.[151,264,1513,1537]" captionTargetBox="[161,1420,185,1484]" captionTargetId="figure-19@3.[151,1436,181,1489]" captionTargetPageId="3" captionText="FIGURES 2A–D. Spariolenus omidvarbrothers sp. n., female paratype, Iran (SMF). A epigyne, ventral; B vulva, dorsal; C right vulva, lateral; D schematic course of internal duct system, lateral.Abbreviations: CO—Copulatory opening; EF—Epigynal field; EP—Epigynal pit; FC—First coil of vulva; FD—Fertilisation duct; LID—Lumen of internal duct system; MEP —Margin of epigynal pit; SC—Second coil of vulva; TC—Third coil of vulva." figureDoi="http://doi.org/10.5281/zenodo.10212765" httpUri="https://zenodo.org/record/10212765/files/figure.png" pageId="4" pageNumber="81">Fig. 2A</figureCitation>
); TC extending laterad beyond FC and SC (
<figureCitation id="C3AE2A6AE327C655FC80FCB56BC48B7B" box="[848,975,799,825]" captionStart="FIGURES 2" captionStartId="3.[151,264,1513,1537]" captionTargetBox="[161,1420,185,1484]" captionTargetId="figure-19@3.[151,1436,181,1489]" captionTargetPageId="3" captionText="FIGURES 2A–D. Spariolenus omidvarbrothers sp. n., female paratype, Iran (SMF). A epigyne, ventral; B vulva, dorsal; C right vulva, lateral; D schematic course of internal duct system, lateral.Abbreviations: CO—Copulatory opening; EF—Epigynal field; EP—Epigynal pit; FC—First coil of vulva; FD—Fertilisation duct; LID—Lumen of internal duct system; MEP —Margin of epigynal pit; SC—Second coil of vulva; TC—Third coil of vulva." figureDoi="http://doi.org/10.5281/zenodo.10212765" httpUri="https://zenodo.org/record/10212765/files/figure.png" pageId="4" pageNumber="81">Figs 2B–C</figureCitation>
).
</paragraph>
<paragraph id="5B2A36EFE327C655FF17FCE96C488B1F" blockId="4.[151,1437,151,969]" box="[199,1091,835,861]" pageId="4" pageNumber="81">
<emphasis id="69E1EAFDE327C655FF17FCE969538B1F" box="[199,344,835,861]" italics="true" pageId="4" pageNumber="81">Colouration.</emphasis>
Same as for male but with lighter colour bands on legs (
<figureCitation id="C3AE2A6AE327C655FC01FCE96C3D8B1F" box="[977,1078,835,861]" captionStart="FIGURES 11" captionStartId="15.[151,264,1906,1930]" captionTargetBox="[159,1427,181,1882]" captionTargetId="figure-19@15.[159,1427,181,1882]" captionTargetPageId="15" captionText="FIGURES 11A–E. A–C Spariolenus omidvarbrothers sp. n. (A habitus of male, B female, C type locality and habitat, Bidan Oasis close to Barashk in Sistan &amp; Baluchistan Province, Iran); D Spariolenus mansourii Moradmand, 2017 preying on a cicada; E habitat of Spariolenus lindbergi (Roewer, 1962) comb. nov. in Afghanistan. Arrows shows the observation location of the specimens." figureDoi="http://doi.org/10.5281/zenodo.10212793" httpUri="https://zenodo.org/record/10212793/files/figure.png" pageId="4" pageNumber="81">Fig. 11B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="138F6564E327C655FF17FCCC6D9A8B8B" pageId="4" pageNumber="81" type="distribution">
<paragraph id="5B2A36EFE327C655FF17FCCC6D9A8B8B" blockId="4.[151,1437,151,969]" pageId="4" pageNumber="81">
<emphasis id="69E1EAFDE327C655FF17FCCC6A8B8BC2" bold="true" box="[199,640,870,897]" pageId="4" pageNumber="81">Distribution and habitat preferences.</emphasis>
Known only from the
<typeStatus id="842E884DE327C655FCACFCCD6BA78BC3" box="[892,940,871,897]" pageId="4" pageNumber="81">type</typeStatus>
locality (
<figureCitation id="C3AE2A6AE327C655FBC5FCCD6C6C8BC3" box="[1045,1127,871,897]" captionStart="FIGURE 13" captionStartId="17.[151,250,1099,1123]" captionTargetBox="[163,1424,181,1075]" captionTargetId="figure-19@17.[163,1424,181,1075]" captionTargetPageId="17" captionText="FIGURE 13. The distribution records map of all known Spariolenus species (studied here and revised to date) in South and West Asia. Localities for all new species described in this study are marked with a star." figureDoi="http://doi.org/10.5281/zenodo.10212798" httpUri="https://zenodo.org/record/10212798/files/figure.png" pageId="4" pageNumber="81">Fig. 13</figureCitation>
). Specimens were collected at night on boulders and rocks. The
<typeStatus id="842E884DE327C655FDF3FC216A588BE7" box="[547,595,907,933]" pageId="4" pageNumber="81">type</typeStatus>
locality was an oasis in a small canyon dominated by palm trees (
<figureCitation id="C3AE2A6AE327C655FAFBFC216D848BE7" box="[1323,1423,907,933]" captionStart="FIGURES 11" captionStartId="15.[151,264,1906,1930]" captionTargetBox="[159,1427,181,1882]" captionTargetId="figure-19@15.[159,1427,181,1882]" captionTargetPageId="15" captionText="FIGURES 11A–E. A–C Spariolenus omidvarbrothers sp. n. (A habitus of male, B female, C type locality and habitat, Bidan Oasis close to Barashk in Sistan &amp; Baluchistan Province, Iran); D Spariolenus mansourii Moradmand, 2017 preying on a cicada; E habitat of Spariolenus lindbergi (Roewer, 1962) comb. nov. in Afghanistan. Arrows shows the observation location of the specimens." figureDoi="http://doi.org/10.5281/zenodo.10212793" httpUri="https://zenodo.org/record/10212793/files/figure.png" pageId="4" pageNumber="81">Fig. 11C</figureCitation>
). An unknown species of the genus
<taxonomicName id="9C954D6CE327C655FDC7FC056B308B8B" authority="Simon, 1903" authorityName="Simon" authorityYear="1903" box="[535,827,943,969]" class="Arachnida" family="Sparassidae" genus="Eusparassus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="69E1EAFDE327C655FDC7FC056AAE8B8A" box="[535,677,943,968]" italics="true" pageId="4" pageNumber="81">Eusparassus</emphasis>
<bibRefCitation id="3F044B1EE327C655FD7CFC056B308B8B" author="Simon, E." box="[684,827,943,969]" pageId="4" pageNumber="81" refId="ref7869" refString="Simon, E. (1903) Histoire naturelle des Araignees. (2 nd Ed.). Roret, Paris, 1080 pp." type="book" year="1903">Simon, 1903</bibRefCitation>
</taxonomicName>
was sympatrically sampled on plants and vegetation.
</paragraph>
</subSubSection>
</treatment>
</document>
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