137 lines
16 KiB
XML
137 lines
16 KiB
XML
<document id="E78058873803CE9676D9F9F65DFC3CCE" ID-CLB-Dataset="51238" ID-DOI="10.5281/zenodo.156768" ID-GBIF-Dataset="0ae8bbb8-3209-479e-bd4e-efa5e38555e4" ID-ISSN="11755326" ID-Zenodo-Dep="156768" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459639206332" checkinUser="plazi" docAuthor="Wappler, Torsten" docDate="2003" docId="D33887DEFF9A3B0CFEF104BCE481D71C" docLanguage="en" docName="zt00374.pdf" docOrigin="Zootaxa 374" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Chorotingioites Wappler, 2003, n. gen." docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="2F01FFA6FF9D3B04FFF9072DE524D40A" masterDocTitle="New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs" masterLastPageNumber="26" masterPageNumber="1" pageNumber="8" updateTime="1698205971885" updateUser="ExternalLinkService">
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<mods:mods id="CD64A0CFD941C58979F4ACA1ED14A0FA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D70ABF448E4CFD811D83C2383EC9CCAE">
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<mods:title id="80CF4FD08E069D9671B5A1B4D60E0335">New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs</mods:title>
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<mods:name id="2F9FC1DE08CC1CCD09BB8899E6EFFEDE" type="personal">
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<mods:roleTerm id="30540816612D40D34CAABC3A09AB7294">Author</mods:roleTerm>
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<mods:namePart id="A0F5176C0F6015D6DA1827A8BD3E36EF">Wappler, Torsten</mods:namePart>
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<mods:typeOfResource id="24A09041989E017ABDA94F3680AAD590">text</mods:typeOfResource>
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<mods:title id="2522EA7A594137C2FC657D391893E127">Zootaxa</mods:title>
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<mods:part id="6502332A7B9D46AC8BF893C3AC280F40">
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<mods:date id="460B924A02786B244A8CB608E5594E0F">2003</mods:date>
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<mods:detail id="8E08B94C5B9200849A16754EE7C7EC44" type="volume">
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<mods:number id="6687C55C2D42C571D4CB032EDD818E5B">374</mods:number>
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<mods:classification id="3A46186EFE0956ADF25145620B0AD5D3">journal article</mods:classification>
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<mods:identifier id="598740BEEB7FFA6EE59D5E4B8CA8E431" type="CLB-Dataset">51238</mods:identifier>
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<mods:identifier id="30430CEE4DEA182537BC2414ED1F5A55" type="DOI">10.5281/zenodo.156768</mods:identifier>
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<mods:identifier id="C616A95CFC022550FEE86805894B8853" type="Zenodo-Dep">156768</mods:identifier>
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<treatment id="D33887DEFF9A3B0CFEF104BCE481D71C" ID-DOI="http://doi.org/10.5281/zenodo.6273731" ID-GBIF-Taxon="119332621" ID-Zenodo-Dep="6273731" LSID="urn:lsid:plazi:treatment:D33887DEFF9A3B0CFEF104BCE481D71C" httpUri="http://treatment.plazi.org/id/D33887DEFF9A3B0CFEF104BCE481D71C" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<subSubSection id="138B6543FF9A3B03FEF104BCE738D7A1" box="[264,540,913,939]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="5B2E36C8FF9A3B03FEF104BCE738D7A1" blockId="7.[264,540,913,939]" box="[264,540,913,939]" pageId="7" pageNumber="8">
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<heading id="006681A4FF9A3B03FEF104BCE738D7A1" bold="true" box="[264,540,913,939]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<emphasis id="69E5EADAFF9A3B03FEF104BCE738D7A1" bold="true" box="[264,540,913,939]" pageId="7" pageNumber="8">
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<taxonomicName id="9C914D4BFF9A3B03FEF104BCE4E7D7A1" box="[264,451,913,939]" class="Insecta" family="Tingidae" genus="Chorotingioites" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="69E5EADAFF9A3B03FEF104BCE4E7D7A1" bold="true" box="[264,451,913,939]" italics="true" pageId="7" pageNumber="8">Chorotingioites</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="72D657A1FF9A3B03FE3204BCE738D7A1" box="[459,540,913,939]" pageId="7" pageNumber="8" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="138B6543FF9A3B03FEF104F1E687D7FC" box="[264,931,988,1014]" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="5B2E36C8FF9A3B03FEF104F1E687D7FC" blockId="7.[264,1323,988,1894]" box="[264,931,988,1014]" pageId="7" pageNumber="8">
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<emphasis id="69E5EADAFF9A3B03FEF104F1E488D7FC" bold="true" box="[264,428,988,1014]" pageId="7" pageNumber="8">
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<typeStatus id="842A886AFF9A3B03FEF104F1E461D7FC" box="[264,325,988,1014]" pageId="7" pageNumber="8">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="9C914D4BFF9A3B03FE4D04F1E7E3D7FF" box="[436,711,988,1013]" class="Insecta" family="Tingidae" genus="Chorotingioites" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="priscus" status="sp. nov.">
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<emphasis id="69E5EADAFF9A3B03FE4D04F1E7E3D7FF" box="[436,711,988,1013]" italics="true" pageId="7" pageNumber="8">Chorotingioites priscus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="72D657A1FF9A3B03FD3704F1E62BD7FC" box="[718,783,988,1014]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
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, monobasic.
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</paragraph>
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</subSubSection>
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<subSubSection id="138B6543FF9A3B03FEC10329E14DD04C" pageId="7" pageNumber="8" type="etymology">
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<paragraph id="5B2E36C8FF9A3B03FEC10329E14DD04C" blockId="7.[264,1323,988,1894]" pageId="7" pageNumber="8">
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<emphasis id="69E5EADAFF9A3B03FEC10329E707D014" bold="true" box="[312,547,1028,1054]" pageId="7" pageNumber="8">Derivatio nominis:</emphasis>
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<emphasis id="69E5EADAFF9A3B03FDCE0329E77BD017" box="[567,607,1028,1053]" italics="true" pageId="7" pageNumber="8">ites</emphasis>
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, from the Greek for “like”, in reference to the similarity of this genus to
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<taxonomicName id="9C914D4BFF9A3B03FE5D0301E70BD04F" box="[420,559,1068,1093]" class="Insecta" family="Tingidae" genus="Chorotingis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="69E5EADAFF9A3B03FE5D0301E70BD04F" box="[420,559,1068,1093]" italics="true" pageId="7" pageNumber="8">Chorotingis</emphasis>
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</taxonomicName>
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of the tribe Ypsotingini. The name is masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="138B6543FF9A3B03FEC10379E16CD1F4" pageId="7" pageNumber="8" type="diagnosis">
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<paragraph id="5B2E36C8FF9A3B03FEC10379E447D09C" blockId="7.[264,1323,988,1894]" pageId="7" pageNumber="8">
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<emphasis id="69E5EADAFF9A3B03FEC10379E49ED064" bold="true" box="[312,442,1108,1134]" pageId="7" pageNumber="8">Diagnosis:</emphasis>
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Large specimen, ca.
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<quantity id="9C699B2DFF9A3B03FD440379E63AD064" box="[701,798,1108,1134]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="7" pageNumber="8" unit="mm" value="4.8">4.8 mm</quantity>
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. Body moderately oval quite elongated and slender.
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</paragraph>
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<paragraph id="5B2E36C8FF9A3B03FEC10389E16CD1F4" blockId="7.[264,1323,988,1894]" pageId="7" pageNumber="8">Head reaching or almost reaching apex of antennal segment I, with 3 spines—1 dorsomedial and 2 occipital. Eyes globular, projecting laterally of the head. Antennae moderately long and thickened. Pronotum with three parallel carinae. Areolate posterior process well developed, elongated backwards along hemelytra covering whole clavi and approximately half of discoidal area. Macropterous form. Costal area moderately wide, along the entire length. Subcostal area nearly as wide as costal area, with three rows of cells in the widest par. Discoidal area in the widest place with 4 incompletely regular rows of cells. Sutural area (membrane) extremely widened forming the areolate membrane which consists in the broadest place of approximately 8–9 irregular rows of cells.</paragraph>
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</subSubSection>
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<subSubSection id="138B6543FF9A3B0CFEC10121E481D71C" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="discussion">
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<paragraph id="5B2E36C8FF9A3B03FEC10121E6E2D36C" blockId="7.[264,1323,988,1894]" pageId="7" pageNumber="8">
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<emphasis id="69E5EADAFF9A3B03FEC10121E4ECD22C" bold="true" box="[312,456,1548,1574]" pageId="7" pageNumber="8">Comments:</emphasis>
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The presence of a celllike hind outgrowth of the pronotum that completely covers the clavi and the absence of additional protruding transverse veins on the hemelytra indicates that
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<taxonomicName id="9C914D4BFF9A3B03FDD70171E7D4D27F" box="[558,752,1628,1653]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="69E5EADAFF9A3B03FDD70171E7D4D27F" box="[558,752,1628,1653]" italics="true" pageId="7" pageNumber="8">Oblongomorpha</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="72D657A1FF9A3B03FD000171E66ED27C" box="[761,842,1628,1654]" pageId="7" pageNumber="8" rank="genus">n. gen.</taxonomicNameLabel>
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belongs to the subfamily
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<taxonomicName id="9C914D4BFF9A3B03FB7D0171E1CBD27C" box="[1156,1263,1628,1654]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subFamily" subFamily="Tinginae">Tinginae</taxonomicName>
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. The elongated head, even reaching the apex of the first antennal segment, and the presence of a dorsomedial and two occipital spines exclude
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<taxonomicName id="9C914D4BFF9A3B03FCCF0181E6DCD2CF" box="[822,1016,1708,1733]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="69E5EADAFF9A3B03FCCF0181E6DCD2CF" box="[822,1016,1708,1733]" italics="true" pageId="7" pageNumber="8">Oblongomorpha</emphasis>
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</taxonomicName>
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from being a member of one of the tribes
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<taxonomicName id="9C914D4BFF9A3B03FE2C01F9E70ED2E4" box="[469,554,1748,1774]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="tribe" tribe="Tingini">Tingini</taxonomicName>
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and Litatdeini. Further apomorphies of the Ypsotingini, such as the reduced or depressed clavi, is difficult to observe in fossils and their presence can neither be confirmed nor excluded. Based on the available evidence, however, I am most inclined to classify the fossil genus in the tribe
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<emphasis id="69E5EADAFF9A3B03FCC80061E699D36C" bold="true" box="[817,957,1868,1894]" pageId="7" pageNumber="8">Ypsotingini</emphasis>
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.
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</paragraph>
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<paragraph id="5B2E36C8FF953B0CFEC10631E185D644" blockId="8.[264,1324,284,790]" pageId="8" pageNumber="9">
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The
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<typeStatus id="842A886AFF953B0CFE940631E484D53C" box="[365,416,284,310]" pageId="8" pageNumber="9">type</typeStatus>
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species
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<taxonomicName id="9C914D4BFF953B0CFDFA0631E75FD53F" box="[515,635,284,309]" class="Insecta" family="Tingidae" genus="Chorotingioites" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="priscus">
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<emphasis id="69E5EADAFF953B0CFDFA0631E75FD53F" box="[515,635,284,309]" italics="true" pageId="8" pageNumber="9">C. priscus</emphasis>
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</taxonomicName>
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, is relative large (estimated length about
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<quantity id="9C699B2DFF953B0CFB9B0631E183D53C" box="[1122,1191,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="mm" value="5.0">5 mm</quantity>
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) compared to most members of the Ypsotingini. On account of the presence of longitudinal carinae on the pronotum, the narrow paranota, the general shape of the body, and the separation of the hemelytra in costal, subcostal, discoidal, and sutural areas, with a 2seriated costal area along most of length of the discoidal area, the fossil species is seemingly more close related to the New World genera
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<emphasis id="69E5EADAFF953B0CFD6B06C9E62BD5F7" box="[658,783,484,509]" italics="true" pageId="8" pageNumber="9">Chortingis</emphasis>
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Drake, and
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<taxonomicName id="9C914D4BFF953B0CFC5806C9E184D5F4" authority="Drake. The" authorityName="Drake. The" box="[929,1184,484,510]" class="Insecta" family="Tingidae" genus="Euaulana" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="69E5EADAFF953B0CFC5806C9E137D5F7" box="[929,1043,484,509]" italics="true" pageId="8" pageNumber="9">Euaulana</emphasis>
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Drake. The
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</taxonomicName>
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incomplete state of the material, however, does not permit an evaluation of its closer relationships. Both genera at present restricted to
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<collectingCountry id="23867658FF953B0CFD5E0519E636D644" box="[679,786,564,590]" name="Australia" pageId="8" pageNumber="9">Australia</collectingCountry>
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and Tasmania (
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<bibRefCitation id="3F004B39FF953B0CFC330519E1B0D644" author="Froeschner" box="[970,1172,564,590]" pageId="8" pageNumber="18" refString="Froeschner, R. C. (2001) Lace bug genera of the world, II: Subfamily Tinginae: Tribes Litadeini and Ypsotingini (Heteroptera: Tingidae). Smithsonian Contributions to Zoology, 611, 1 - 28." type="journal article" year="2001">Froeschner 2001</bibRefCitation>
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).
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</paragraph>
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<paragraph id="5B2E36C8FF953B0CFEC10571E481D71C" blockId="8.[264,1324,284,790]" pageId="8" pageNumber="9">
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The current classification of this tribe contains only 7 modern extant genera with 76 species (
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<bibRefCitation id="3F004B39FF953B0CFE9405A9E71ED694" author="Froeschner" box="[365,570,644,670]" pageId="8" pageNumber="18" refString="Froeschner, R. C. (2001) Lace bug genera of the world, II: Subfamily Tinginae: Tribes Litadeini and Ypsotingini (Heteroptera: Tingidae). Smithsonian Contributions to Zoology, 611, 1 - 28." type="journal article" year="2001">Froeschner 2001</bibRefCitation>
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). So far no representative of the Ypsotingini has been reported from the fossil record. Lace bugs belonging to the Ypsotingini mainly distributed in the Paleartic and Oriental zoogeographic regions (
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<bibRefCitation id="3F004B39FF953B0CFCD205F9E6D2D6E4" author="Froeschner" box="[811,1014,724,750]" pageId="8" pageNumber="18" refString="Froeschner, R. C. (2001) Lace bug genera of the world, II: Subfamily Tinginae: Tribes Litadeini and Ypsotingini (Heteroptera: Tingidae). Smithsonian Contributions to Zoology, 611, 1 - 28." type="journal article" year="2001">Froeschner 2001</bibRefCitation>
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;
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<bibRefCitation id="3F004B39FF953B0CFBFA05F9E180D6E4" author="Pericart" box="[1027,1188,724,750]" pageId="8" pageNumber="19" refString="Pericart, J. (1983) Hemipteres Tingidae Euro-Mediterraneens. Faune de France, 69, Federation Francaise des Societe de Sciences Naturelles, Paris, 620 pp." type="book" year="1983">Péricart 1983</bibRefCitation>
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;
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<bibRefCitation id="3F004B39FF953B0CFB4805F9E4B3D71C" author="Pericart" pageId="8" pageNumber="19" refString="Pericart, J. & Golub, V. B. (1996) Tingidae. In: Aukema, B. & Rieger, Ch. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region, 2, Netherlands Entomological Society, Wageningen, 361 pp." type="book" year="1996">Péricart & Golub 1996</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |