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<document id="F0923A1C26223A8B0F6733C46E28CD0E" ID-CLB-Dataset="51238" ID-DOI="10.5281/zenodo.156768" ID-GBIF-Dataset="0ae8bbb8-3209-479e-bd4e-efa5e38555e4" ID-ISSN="1175­5326" ID-Zenodo-Dep="156768" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459639206332" checkinUser="plazi" docAuthor="Wappler, Torsten" docDate="2003" docId="D33887DEFF983B03FEF10519E02ED6CC" docLanguage="en" docName="zt00374.pdf" docOrigin="Zootaxa 374" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Exmesselensis disspinosus Wappler, 2003, n. sp." docType="treatment" docVersion="10" lastPageNumber="8" masterDocId="2F01FFA6FF9D3B04FFF9072DE524D40A" masterDocTitle="New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs" masterLastPageNumber="26" masterPageNumber="1" pageNumber="6" updateTime="1698205971885" updateUser="ExternalLinkService">
<mods:mods id="C77C527E08D5F0912E7623F5DB416B23" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A1B750A37FB2320BF9710601FAAD7053">
<mods:title id="89D23BC4D0D6B6C4DA2D89930BA7F12A">New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs</mods:title>
</mods:titleInfo>
<mods:name id="B9C8BBF2E195A72D9FD6437B088CE897" type="personal">
<mods:role id="190969AB8349976AC62C1E1CBA417C32">
<mods:roleTerm id="0E2366F75A5D44A4E72983532EB0B00B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D8DA9EE9911D5ACFBF8657B6305FD058">Wappler, Torsten</mods:namePart>
</mods:name>
<mods:typeOfResource id="AAEB10968AD26B4203D82B73547F9D81">text</mods:typeOfResource>
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<mods:title id="F6CDEB278FFB548DBB23266369F116EC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AE887AF812B2C18A1E636D33BD3568AA">
<mods:date id="C87B20175D9E48B787EA9314E5A33A4B">2003</mods:date>
<mods:detail id="6579C6A43F38CB149550376C53ACE4B8" type="volume">
<mods:number id="949F9A6BD1826CAE604B3437170CD652">374</mods:number>
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<mods:start id="CD51EDF2564FAC26D7695DADD03AE2CE">1</mods:start>
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<mods:classification id="02861075A16C16CCADDE74D854C92DCE">journal article</mods:classification>
<mods:identifier id="FCC583AB2B60FC4007ACDA091DCF658A" type="CLB-Dataset">51238</mods:identifier>
<mods:identifier id="E113CE7E90E2874C0866213E293988B8" type="DOI">10.5281/zenodo.156768</mods:identifier>
<mods:identifier id="559073275CD45FAC2126D8355DC73E24" type="GBIF-Dataset">0ae8bbb8-3209-479e-bd4e-efa5e38555e4</mods:identifier>
<mods:identifier id="410E3EE99AC370CED714242696EE9DFE" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="26EA69C9863BAD9CC8FDB26824D3349B" type="Zenodo-Dep">156768</mods:identifier>
</mods:mods>
<treatment id="D33887DEFF983B03FEF10519E02ED6CC" ID-DOI="http://doi.org/10.5281/zenodo.6273729" ID-GBIF-Taxon="119332623" ID-Zenodo-Dep="6273729" LSID="urn:lsid:zoobank.org:act:D88F0F11-7754-4222-A058-E744B8EABBCA" httpUri="http://treatment.plazi.org/id/D33887DEFF983B03FEF10519E02ED6CC" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<subSubSection id="138B6543FF983B01FEF10519E62BD647" box="[264,783,563,590]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="5B2E36C8FF983B01FEF10519E62BD647" blockId="5.[264,783,563,590]" box="[264,783,563,590]" pageId="5" pageNumber="6">
<heading id="006681A4FF983B01FEF10519E62BD647" bold="true" box="[264,783,563,590]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="69E5EADAFF983B01FEF10519E62BD647" bold="true" box="[264,783,563,590]" pageId="5" pageNumber="6">
<taxonomicName id="9C914D4BFF983B01FEF10519E76DD644" box="[264,585,564,590]" class="Insecta" family="Tingidae" genus="Exmesselensis" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="disspinosus" status="sp. nov.">
<emphasis id="69E5EADAFF983B01FEF10519E76DD644" bold="true" box="[264,585,564,590]" italics="true" pageId="5" pageNumber="6">Exmesselensis disspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72D657A1FF983B01FDB6051EE7B6D647" box="[591,658,563,589]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="C3AA2A4DFF983B01FD58051EE623D647" box="[673,775,563,589]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3 – 6. 3" captionStartId-0="4.[264,368,1541,1565]" captionStartId-1="6.[264,368,1650,1674]" captionTargetBox-0="[461,1161,454,1498]" captionTargetBox-1="[264,1323,490,1624]" captionTargetId-0="figure@4.[435,1161,449,1512]" captionTargetId-1="figure@6.[264,1323,490,1624]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="FIGURE 2. Exmesselensis disspinosus gen. et sp. n., reconstruction of the holotype, (macropterous form), compression fossil; coll. Forschungsinstitut Senckenberg (SMF), Frankfurt am Main (Germany), specimen SMF MeI 6301; scale bar represents 0.5 mm." captionText-1="FIGURE 3 – 6. 3, Dorsal habitus of holotype of Exmesselensis disspinosus (SMF MeI 6301); 4, Detail of areolate hemelytra of E. disspinosus (SMF MeI 6301); 5, Lateral habitus of paratype (SMF MeI 6958); 6, Detail of head and antennae structures (SMF MeI 6958)." httpUri-0="https://zenodo.org/record/156770/files/figure.png" httpUri-1="https://zenodo.org/record/168855/files/figure.png" pageId="5" pageNumber="6">Fig. 2–6</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="138B6543FF983B01FEF10553E4E3D6E3" pageId="5" pageNumber="6" type="etymology">
<paragraph id="5B2E36C8FF983B01FEF10553E4E3D6E3" blockId="5.[264,1325,638,1905]" pageId="5" pageNumber="6">
<emphasis id="69E5EADAFF983B01FEF10553E4D1D692" bold="true" box="[264,501,638,664]" pageId="5" pageNumber="6">Derivatio nominis:</emphasis>
The specific epithet is derived from the Latin words
<emphasis id="69E5EADAFF983B01FB770552E195D692" box="[1166,1201,639,664]" italics="true" pageId="5" pageNumber="6">dis</emphasis>
(meaning “without”) and
<emphasis id="69E5EADAFF983B01FE47058AE707D6CA" box="[446,547,679,704]" italics="true" pageId="5" pageNumber="6">spinosus</emphasis>
(meaning “thorny”) and is a reference to the absence of conspicuous head spines.
</paragraph>
</subSubSection>
<subSubSection id="138B6543FF983B01FEC105DBE126D7BB" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="5B2E36C8FF983B01FEC105DBE002D733" blockId="5.[264,1325,638,1905]" pageId="5" pageNumber="6">
<emphasis id="69E5EADAFF983B01FEC105DBE497D71A" bold="true" box="[312,435,758,784]" pageId="5" pageNumber="6">
<typeStatus id="842A886AFF983B01FEC105DBE48AD71A" box="[312,430,758,784]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
:
</emphasis>
A complete individual, dorsoventrally compressed, coll. Forschungsinstitut Senckenberg (
<collectionCode id="3D80AE0DFF983B01FE560432E4CBD733" LSID="urn:lsid:biocol.org:col:34838" box="[431,495,799,825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="6">SMF</collectionCode>
), Frankfurt am Main (
<collectingCountry id="23867658FF983B01FD080432E641D733" box="[753,869,799,825]" name="Germany" pageId="5" pageNumber="6">Germany</collectingCountry>
), specimen
<collectionCode id="3D80AE0DFF983B01FC170432E10DD733" LSID="urn:lsid:biocol.org:col:34838" box="[1006,1065,799,825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="6">SMF</collectionCode>
MeI 6301 (
<figureCitation id="C3AA2A4DFF983B01FB4B0432E03DD733" box="[1202,1305,799,825]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3 – 6. 3" captionStartId-0="4.[264,368,1541,1565]" captionStartId-1="6.[264,368,1650,1674]" captionTargetBox-0="[461,1161,454,1498]" captionTargetBox-1="[264,1323,490,1624]" captionTargetId-0="figure@4.[435,1161,449,1512]" captionTargetId-1="figure@6.[264,1323,490,1624]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="FIGURE 2. Exmesselensis disspinosus gen. et sp. n., reconstruction of the holotype, (macropterous form), compression fossil; coll. Forschungsinstitut Senckenberg (SMF), Frankfurt am Main (Germany), specimen SMF MeI 6301; scale bar represents 0.5 mm." captionText-1="FIGURE 3 – 6. 3, Dorsal habitus of holotype of Exmesselensis disspinosus (SMF MeI 6301); 4, Detail of areolate hemelytra of E. disspinosus (SMF MeI 6301); 5, Lateral habitus of paratype (SMF MeI 6958); 6, Detail of head and antennae structures (SMF MeI 6958)." httpUri-0="https://zenodo.org/record/156770/files/figure.png" httpUri-1="https://zenodo.org/record/168855/files/figure.png" pageId="5" pageNumber="6">Fig. 2–4</figureCitation>
).
</paragraph>
<paragraph id="5B2E36C8FF983B01FEC1046BE126D7BB" blockId="5.[264,1325,638,1905]" pageId="5" pageNumber="6">
<emphasis id="69E5EADAFF983B01FEC1046BE49BD76A" bold="true" box="[312,447,838,864]" pageId="5" pageNumber="6">
<typeStatus id="842A886AFF983B01FEC1046BE49ED76A" box="[312,442,838,864]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="3D80AE0DFF983B01FE33046AE721D76B" LSID="urn:lsid:biocol.org:col:34838" box="[458,517,839,865]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="6">SMF</collectionCode>
MeI 6958 (laterally compressed),
<collectionCode id="3D80AE0DFF983B01FC49046AE6CFD76B" LSID="urn:lsid:biocol.org:col:34838" box="[944,1003,839,865]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="6">SMF</collectionCode>
MeI 7798 (dorsoventrally compressed),
<collectionCode id="3D80AE0DFF983B01FE560442E4CED783" LSID="urn:lsid:biocol.org:col:34838" box="[431,490,879,905]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="6">SMF</collectionCode>
MeI 7877 (dorsoventrally compressed),
<collectionCode id="3D80AE0DFF983B01FC180442E138D783" LSID="urn:lsid:biocol.org:col:34838" box="[993,1052,879,905]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="6">SMF</collectionCode>
MeI 7920 (dorsoventrally compressed),
<collectionCode id="3D80AE0DFF983B01FE0904BAE70FD7BB" LSID="urn:lsid:biocol.org:col:34838" box="[496,555,919,945]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="6">SMF</collectionCode>
MeI 8082 (dorsoventrally compressed).
</paragraph>
</subSubSection>
<subSubSection id="138B6543FF983B01FEC10493E1BBD073" pageId="5" pageNumber="6" type="description">
<paragraph id="5B2E36C8FF983B01FEC10493E1BBD073" blockId="5.[264,1325,638,1905]" pageId="5" pageNumber="6">
<emphasis id="69E5EADAFF983B01FEC10493E761D7D2" bold="true" box="[312,581,958,984]" pageId="5" pageNumber="6">Dimensions (in mm):</emphasis>
Body length 2.56, width 1.08; length of head (from apex of clypeus to posterior margin of pronotum) 0.93, width 0.63; interocular width 0.26; antennomeres 2­4: 0.16, 0.99, 0.46; length of pronotum 0.83, width 1.36; length of fore leg (SMF MeI 6958): femur 0.69, tibia 0.99 (incomplete); length of middle leg (SMF MeI 6958): femur 0.79; length of hind leg (SMF MeI 6958): femur 0.96, tibia 1.19.
</paragraph>
</subSubSection>
<subSubSection id="138B6543FF983B01FEC103ABE4AED113" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="5B2E36C8FF983B01FEC103ABE132D0AB" blockId="5.[264,1325,638,1905]" box="[312,1046,1158,1185]" pageId="5" pageNumber="6">
<emphasis id="69E5EADAFF983B01FEC103ABE4C5D0AA" bold="true" box="[312,481,1158,1184]" pageId="5" pageNumber="6">
<typeStatus id="842A886AFF983B01FEC103ABE451D0AA" box="[312,373,1158,1184]" pageId="5" pageNumber="6">Type</typeStatus>
locality:
</emphasis>
Grube Messel near Darmstadt, grid square E89.
</paragraph>
<paragraph id="5B2E36C8FF983B01FEC10383E4AED113" blockId="5.[264,1325,638,1905]" pageId="5" pageNumber="6">
<emphasis id="69E5EADAFF983B01FEC10383E65ED0C2" bold="true" box="[312,890,1198,1224]" pageId="5" pageNumber="6">Stratigraphical and geographical distribution:</emphasis>
Known only from the lower Middle Eocene (Lower Lutetian,
<collectionCode id="3D80AE0DFF983B01FDC703FAE7B5D0FB" box="[574,657,1239,1265]" pageId="5" pageNumber="6">ELMA</collectionCode>
Geiseltalian, MP11), Messel Formation;
<quantity id="9C699B2DFF983B01FB7E03FAE00DD0FB" box="[1159,1321,1239,1265]" metricMagnitude="0" metricUnit="m" metricValue="2.83" metricValueMax="3.09" metricValueMin="2.57" pageId="5" pageNumber="6" unit="m" value="2.83" valueMax="3.09" valueMin="2.57">2.57 –3.09 m</quantity>
over alpha.
</paragraph>
</subSubSection>
<subSubSection id="138B6543FF983B01FEC1020BE632D14B" box="[312,790,1318,1345]" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="5B2E36C8FF983B01FEC1020BE632D14B" blockId="5.[264,1325,638,1905]" box="[312,790,1318,1345]" pageId="5" pageNumber="6">
<emphasis id="69E5EADAFF983B01FEC1020BE49ED14A" bold="true" box="[312,442,1318,1344]" pageId="5" pageNumber="6">Diagnosis:</emphasis>
<collectingCountry id="23867658FF983B01FE38020AE4C5D14B" box="[449,481,1319,1345]" name="American Samoa" pageId="5" pageNumber="6">As</collectingCountry>
for the genus (see above).
</paragraph>
</subSubSection>
<subSubSection id="138B6543FF983B03FEC10263E02ED6CC" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="5B2E36C8FF983B01FEC10263E4EDD1B3" blockId="5.[264,1325,638,1905]" pageId="5" pageNumber="6">
<emphasis id="69E5EADAFF983B01FEC10263E4F6D162" bold="true" box="[312,466,1358,1384]" pageId="5" pageNumber="6">Description:</emphasis>
Small specimen, ca.
<quantity id="9C699B2DFF983B01FD2F0262E612D163" box="[726,822,1359,1385]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="6" unit="mm" value="2.5">2.5 mm</quantity>
. Body oblong­oval, 2.3 times longer than wide, dorsal and ventral side of the body black to dark­brown, legs and antennae lightbrown posterior.
</paragraph>
<paragraph id="5B2E36C8FF983B01FEC102EAE1DDD2DB" blockId="5.[264,1325,638,1905]" pageId="5" pageNumber="6">Head elongated, 1.47 times as long as wide (from clypeus to anterior margin of pronotum). Vertex between eyes slightly elevated. Eyes globular, strongly projecting laterally of the head. Head with one significant interocular (dorsomedial) tubercle. Antenniferous tubercles short (half as long as the first antennal segment) and oblique diverge, with their apex thin and slightly curved. Antennae long and thin, especially the 3rd is 2.66 times longer than the 4th. First two segments very short. Bucullae distinctly protruding beyond the apex of the clypeus, directed to each other posteriorly, and not touching anteriorly.</paragraph>
<paragraph id="5B2E36C8FF983B02FEC101F2E6E5D58C" blockId="5.[264,1325,638,1905]" lastBlockId="6.[264,1322,284,390]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">Pronotum 1.63 times wide as long, covered with rough punctuation. Pronotal disc perceptible convex, with 3 longitudinal carinae. Median carina stretching from a posterior margin to the middle of the pronotum. Lateral carinae nearly as long as the median carinae, and weakly curved. Anterior margin of pronotum weakly waved, anterior margin medially strongly concave, anterolateral angles rounded. Paranota with one row of medium size cells of irregular and round form, getting gradually smaller to the anterior margin. Scutellum completely exposed, triangular in shape.</paragraph>
<caption id="0FEE6640FF9B3B02FEF1015FE166D2C0" httpUri="https://zenodo.org/record/168855/files/figure.png" pageId="6" pageNumber="7" targetBox="[264,1323,490,1624]" targetPageId="6">
<paragraph id="5B2E36C8FF9B3B02FEF1015FE166D2C0" blockId="6.[264,1323,1650,1738]" pageId="6" pageNumber="7">
<emphasis id="69E5EADAFF9B3B02FEF1015FE48CD280" bold="true" box="[264,424,1650,1674]" pageId="6" pageNumber="7">FIGURE 3–6.</emphasis>
3, Dorsal habitus of holotype of
<taxonomicName id="9C914D4BFF9B3B02FCD8015EE11AD280" box="[801,1086,1651,1674]" class="Insecta" family="Tingidae" genus="Exmesselensis" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="disspinosus">
<emphasis id="69E5EADAFF9B3B02FCD8015EE11AD280" box="[801,1086,1651,1674]" italics="true" pageId="6" pageNumber="7">Exmesselensis disspinosus</emphasis>
</taxonomicName>
(SMF MeI 6301); 4, Detail of areolate hemelytra of
<taxonomicName id="9C914D4BFF9B3B02FD9301BEE623D2A0" box="[618,775,1683,1706]" class="Insecta" family="Tingidae" genus="Exmesselensis" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="disspinosus">
<emphasis id="69E5EADAFF9B3B02FD9301BEE623D2A0" box="[618,775,1683,1706]" italics="true" pageId="6" pageNumber="7">E. disspinosus</emphasis>
</taxonomicName>
(SMF MeI 6301); 5, Lateral habitus of paratype (SMF MeI 6958); 6, Detail of head and antennae structures (SMF MeI 6958).
</paragraph>
</caption>
<paragraph id="5B2E36C8FF9B3B02FEC10036E00FD357" blockId="6.[264,1323,1819,1885]" pageId="6" pageNumber="7">Hemelytra (macropterous form) surpassing apex of abdomen, flattened, with distinctly carinated veins, separating the hemelytra in costal, subcostal, discoidal, and sutural areas.</paragraph>
<paragraph id="5B2E36C8FF9A3B03FEF10631E02ED6CC" blockId="7.[264,1323,284,710]" pageId="7" pageNumber="8">Lateral margins right in the midst imperceptible broadened, and slightly narrowed at base. Costal and subcostal area not wide, each with two rows of cells, and in the last third only with one row. Discoidal area triangular in shape, and in the widest place with 6­7 incompletely regular rows of cells of mainly oval shape. Sutural area (membrane) along clavus with one row of cells, and apical the clavus the sutural area is extremely widened forming the areolate membrane which consists in the broadest place of approximately 12–14 irregular rows of cells of round to oval shape. Tips of sutural areas of both hemelytra fully overlapping at rest. Clavus triangular in shape, and clearly separated by a claval commissure from the corium. In the broadest place with 4(?) rows of cells of mainly oval shape. In regard to the bad preservation the hypocosta is probably 1­serated (SMF MeI 7877, 7798). Femora anterior slightly swollen. Tibiae straight and cylindrical. Tarsi 2­segmented (?).</paragraph>
</subSubSection>
</treatment>
</document>
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