227 lines
32 KiB
XML
227 lines
32 KiB
XML
<document id="54C19FF714455EB112A680F8D6BBE994" ID-CLB-Dataset="32039" ID-DOI="10.11646/zootaxa.4323.1.8" ID-GBIF-Dataset="16e495f6-1e96-4e0a-af86-df00e3a1fdb5" ID-ISSN="1175-5326" ID-Zenodo-Dep="897413" ID-ZooBank="081A06F5-76A4-438A-85C8-959E1E183C73" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505882101095" checkinUser="plazi" docAuthor="Novelo-Gutiérrez, Rodolfo" docDate="2017" docId="BE3A6F5B8141EF4216AE0AD2FE030A5B" docLanguage="en" docName="zootaxa.4323.1.8.pdf" docOrigin="Zootaxa 4323 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pspehenops triangularis Novelo-Gutiérrez 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="115" masterDocId="420317238144EF4416390A45FFFB0B3F" masterDocTitle="Two new species of Psephenops Grouvelle (Coleoptera: Byrrhoidea: Psephenidae) from Panama" masterLastPageNumber="118" masterPageNumber="109" pageNumber="114" updateTime="1698468758633" updateUser="ExternalLinkService">
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<mods:titleInfo id="140D1CAACE76DAE0D3B0FAA3408A2CAF">
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<mods:title id="41885E015A4219782E2CBA058DD49462">Two new species of Psephenops Grouvelle (Coleoptera: Byrrhoidea: Psephenidae) from Panama</mods:title>
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</mods:titleInfo>
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<mods:name id="A2C8D6FB94B2143A3A1934D44D047FEB" type="personal">
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<mods:namePart id="3D4F88DD1EDFC0643984B844237C4D89">Novelo-Gutiérrez, Rodolfo</mods:namePart>
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<mods:title id="1D906ABD68EC87C3C87A691D966D3E94">Zootaxa</mods:title>
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<mods:part id="2084A702B2AF4F081ADED3696756C092">
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<mods:date id="3618236BFD0822DF2202ED85A49B4361">2017</mods:date>
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<mods:number id="D875667CC001C46950E4AE74236437E6">2017-09-20</mods:number>
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<mods:number id="A318A5F9E7995D0A0169DA7172ABBF8B">4323</mods:number>
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<treatment id="BE3A6F5B8141EF4216AE0AD2FE030A5B" ID-DOI="http://doi.org/10.5281/zenodo.6025170" ID-GBIF-Taxon="134313899" ID-Zenodo-Dep="6025170" LSID="urn:lsid:plazi:treatment:BE3A6F5B8141EF4216AE0AD2FE030A5B" httpUri="http://treatment.plazi.org/id/BE3A6F5B8141EF4216AE0AD2FE030A5B" lastPageId="6" lastPageNumber="115" pageId="5" pageNumber="114">
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<subSubSection id="7E898DC68141EF4116AE0AD2FEE30BEC" pageId="5" pageNumber="114" type="nomenclature">
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<paragraph id="362CDE4D8141EF4116AE0AD2FDDA0B8D" blockId="5.[151,545,151,211]" box="[151,545,151,178]" pageId="5" pageNumber="114">
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<heading id="6D6469218141EF4116AE0AD2FDDA0B8D" bold="true" box="[151,545,151,178]" fontSize="11" level="1" pageId="5" pageNumber="114" reason="1">
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<emphasis id="04E7025F8141EF4116AE0AD2FDDA0B8D" bold="true" box="[151,545,151,178]" pageId="5" pageNumber="114">
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<taxonomicName id="F193A5CE8141EF4116AE0AD2FE460B8E" authority="Novelo-Gutiérrez, 2017" authorityName="Novelo-Gutiérrez" authorityYear="2017" box="[151,445,151,177]" class="Insecta" family="Psephenidae" genus="Pspehenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="114" phylum="Arthropoda" rank="species" species="triangularis" status="sp. nov.">
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<emphasis id="04E7025F8141EF4116AE0AD2FE460B8E" bold="true" box="[151,445,151,177]" italics="true" pageId="5" pageNumber="114">Pspehenops triangularis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1FD4BF248141EF4117FD0ADDFDDA0B8D" box="[452,545,152,178]" pageId="5" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="362CDE4D8141EF4116AE0AFFFEE30BEC" blockId="5.[151,545,151,211]" box="[151,280,186,211]" pageId="5" pageNumber="114">
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(
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<figureCitation id="AEA8C2C88141EF4116A60AFFFEEB0BEC" box="[159,272,186,211]" captionStart-0="FIGURES 9 – 12" captionStart-1="FIGURES 13 – 16" captionStartId-0="6.[151,264,1893,1916]" captionStartId-1="7.[151,264,1970,1993]" captionTargetBox-0="[295,1278,455,1846]" captionTargetBox-1="[271,1339,203,1949]" captionTargetId-0="figure@6.[246,1334,427,1873]" captionTargetId-1="figure@7.[233,1359,193,1949]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 9 – 12. Psephenops triangularis sp. nov., male: 9) habitus, dorsal view; 10) habitus, ventral view; 11) A variant male, prosternum and prosternal process; 12) Ídem, dorsal view, with antennae 10 - segmented (in alcohol). Figs 9 – 10, dry specimens. Scale bar for Figs 9, 10 and 12 = 0.5 mm." captionText-1="FIGURES 13 – 16. Psephenops triangularis sp. nov., male genitalia: 13) dorsal; 14) ventral; 15) lateral; 16) ventrolateral. Scale bar = 0.12 mm." httpUri-0="https://zenodo.org/record/897419/files/figure.png" httpUri-1="https://zenodo.org/record/897421/files/figure.png" pageId="5" pageNumber="114">Figs 9–16</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="7E898DC68141EF4116AE0AB9FF0F0A64" pageId="5" pageNumber="114" type="materials_examined">
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<paragraph id="362CDE4D8141EF4116AE0AB9FF0F0A64" blockId="5.[151,1437,252,2015]" pageId="5" pageNumber="114">
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<materialsCitation id="86FBD4108141EF4116AE0AB9FC360A07" ID-GBIF-Occurrence="1639434059" collectingDate="2015-04-26" collectionCode="IEXA" collectorName="R. Novelo" country="Panama" county="Rio Marias" elevation="325" latitude="8.633667" location="Chame District Rio Marias" longLatPrecision="1" longitude="-80.02517" municipality="Chame District" pageId="5" pageNumber="114" specimenCount="2" stateProvince="Panama" typeStatus="holotype">
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<emphasis id="04E7025F8141EF4116AE0AB9FEC50A2A" bold="true" box="[151,318,252,277]" pageId="5" pageNumber="114">
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<typeStatus id="E92860EF8141EF4116AE0AB9FF290A2A" box="[151,210,252,277]" pageId="5" pageNumber="114">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="E92860EF8141EF4117740AB9FE4E0A2A" box="[333,437,252,277]" pageId="5" pageNumber="114" type="holotype">Holotype</typeStatus>
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male labeled:
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<collectingCountry id="4E849EDD8141EF4114620AB9FD2C0A2B" box="[603,727,252,276]" name="Panama" pageId="5" pageNumber="114">PANAMA</collectingCountry>
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: Province of
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<collectingRegion id="F45710AF8141EF4115520AB8FC3C0A2B" box="[875,967,253,276]" country="Panama" name="Panama" pageId="5" pageNumber="114">Panama</collectingRegion>
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;
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<location id="334C88968141EF4115EA0AB9FAFC0A2B" LSID="urn:lsid:plazi:treatment:BE3A6F5B8141EF4216AE0AD2FE030A5B:334C88968141EF4115EA0AB9FAFC0A2B" box="[979,1287,252,277]" country="Panama" county="Rio Marias" latitude="8.633667" longLatPrecision="1" longitude="-80.02517" municipality="Chame District" name="Chame District Rio Marias" pageId="5" pageNumber="114" stateProvince="Panama">
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<collectingMunicipality id="D64844378141EF4115EA0AB9FB850A2A" box="[979,1150,252,277]" pageId="5" pageNumber="114">Chame District</collectingMunicipality>
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<collectingCounty id="DF4DA6C18141EF4112BF0AB9FAFC0A2B" box="[1158,1287,252,276]" pageId="5" pageNumber="114">Río Marías</collectingCounty>
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</location>
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,
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<elevation id="BDBE397E8141EF41132B0AB9FA6C0A2A" box="[1298,1431,252,277]" metricMagnitude="2" metricUnit="m" metricValue="3.25" pageId="5" pageNumber="114" unit="m" value="325.0">
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<quantity id="F16B73A88141EF41132B0AB9FAA10A2B" box="[1298,1370,252,277]" metricMagnitude="2" metricUnit="m" metricValue="3.25" pageId="5" pageNumber="114" unit="m" value="325.0">325 m</quantity>
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a.s.l.
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</elevation>
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, (
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<geoCoordinate id="53A7B88A8141EF4116A60B65FEC40A08" box="[159,319,288,312]" direction="north" orientation="latitude" pageId="5" pageNumber="114" precision="1" value="8.633667">08°38.020’ N</geoCoordinate>
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,
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<geoCoordinate id="53A7B88A8141EF4117740B5AFE080A07" box="[333,499,287,312]" direction="west" orientation="longitude" pageId="5" pageNumber="114" precision="1" value="-80.02517">80°01.510’ W</geoCoordinate>
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),
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<date id="422DF88D8141EF4114320B5AFD4A0A07" box="[523,689,287,312]" pageId="5" pageNumber="114" value="2015-04-26">
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<collectingDate id="526901658141EF4114320B5AFD4A0A07" box="[523,689,287,312]" pageId="5" pageNumber="114" value="2015-04-26">26 April 2015</collectingDate>
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</date>
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,
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<collectorName id="9B66BB9B8141EF4114860B65FCCC0A07" box="[703,823,287,312]" pageId="5" pageNumber="114">R. Novelo</collectorName>
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col. (
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<collectionCode id="508246888141EF4115450B65FC3E0A07" box="[892,965,288,312]" pageId="5" pageNumber="114">IEXA</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="86FBD4108141EF4115E20B65FF0F0A64" ID-GBIF-Occurrence="1639434061" collectingDate="2015-04-26" collectionCode="IEXA" collectorName="R. Novelo" country="Panama" county="Rio Marias" elevation="1" latitude="8.633667" location="Chame District Rio Marias" longLatPrecision="1" longitude="-80.02517" municipality="Chame District" pageId="5" pageNumber="114" specimenCount="1" stateProvince="Panama" typeStatus="paratype">
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||
<typeStatus id="E92860EF8141EF4115E20B65FBBB0A07" box="[987,1088,288,312]" pageId="5" pageNumber="114" type="paratype">Paratype</typeStatus>
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,
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||
<quantity id="F16B73A88141EF4112770B65FB850A08" box="[1102,1150,288,312]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="114" unit="m" value="1.0">
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||
<elevation id="BDBE397E8141EF4112770B65FB850A08" box="[1102,1150,288,312]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="114" unit="m" value="1.0">1 m</elevation>
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||
</quantity>
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||
#, same data as holotype (
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||
<collectionCode id="508246888141EF4116A60B06FF130A64" box="[159,232,323,347]" pageId="5" pageNumber="114">IEXA</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="7E898DC68141EF4116FE0B23FBBB0CE1" pageId="5" pageNumber="114" type="description">
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<paragraph id="362CDE4D8141EF4116FE0B23FDBC0932" blockId="5.[151,1437,252,2015]" pageId="5" pageNumber="114">
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||
<emphasis id="04E7025F8141EF4116FE0B23FEA20A40" bold="true" box="[199,345,358,383]" pageId="5" pageNumber="114">Description.</emphasis>
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<typeStatus id="E92860EF8141EF4117510B23FE2B0A40" box="[360,464,358,383]" pageId="5" pageNumber="114" type="holotype">Holotype</typeStatus>
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Male: Total length,
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<quantity id="F16B73A88141EF4114EE0B23FCBD0A41" box="[727,838,358,383]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="114" unit="mm" value="2.5">2.50 mm</quantity>
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; humeral width, 1.0 mm. Body oval, depressed; integument dark red, head black; pronotum yellowish-red, darker on disk; elytra reddish-yellow, completely covered with short yellow setae with golden reflections, intermingled with dark red long setae (
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<figureCitation id="AEA8C2C88141EF4112CB0BE8FAC10AF9" box="[1266,1338,429,454]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1893,1916]" captionTargetBox="[295,1278,455,1846]" captionTargetId="figure@6.[246,1334,427,1873]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Psephenops triangularis sp. nov., male: 9) habitus, dorsal view; 10) habitus, ventral view; 11) A variant male, prosternum and prosternal process; 12) Ídem, dorsal view, with antennae 10 - segmented (in alcohol). Figs 9 – 10, dry specimens. Scale bar for Figs 9, 10 and 12 = 0.5 mm." httpUri="https://zenodo.org/record/897419/files/figure.png" pageId="5" pageNumber="114">Fig. 9</figureCitation>
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); coxae, trochanters and femora reddish-yellow; tibiae, tarsi and venter dark red, vestiture short, fine and dense yellowishgray, with golden reflections (
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<figureCitation id="AEA8C2C88141EF4117DF0BB1FDC10932" box="[486,570,500,525]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1893,1916]" captionTargetBox="[295,1278,455,1846]" captionTargetId="figure@6.[246,1334,427,1873]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Psephenops triangularis sp. nov., male: 9) habitus, dorsal view; 10) habitus, ventral view; 11) A variant male, prosternum and prosternal process; 12) Ídem, dorsal view, with antennae 10 - segmented (in alcohol). Figs 9 – 10, dry specimens. Scale bar for Figs 9, 10 and 12 = 0.5 mm." httpUri="https://zenodo.org/record/897419/files/figure.png" pageId="5" pageNumber="114">Fig. 10</figureCitation>
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).
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</paragraph>
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||
<paragraph id="362CDE4D8141EF4116FE0852FEBC08AD" blockId="5.[151,1437,252,2015]" pageId="5" pageNumber="114">
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||
<emphasis id="04E7025F8141EF4116FE0852FEF6090F" box="[199,269,535,560]" italics="true" pageId="5" pageNumber="114">Head:</emphasis>
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||
Short, transverse; clypeus subrectangular, declivous at an angle of less than 90 degrees from the plane of the frons, distal margin widely emarginated; fronto-clypeal surface and vertex with short yellow setae and long, dark red setae, vertex minutely punctate. Antennae short (e.g. when fully extended backwards, not reaching posterior margin of pronotum), moniliform, 11-segmented; scape longest, pedicel 0.50 length of scape and subglobose, third antennomere subtriangular and shorter than pedicel, antennomeres 4–10 small, moniliform, apical antennomere acuminate; scape and pedicel yellow (
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<figureCitation id="AEA8C2C88141EF41151D088CFC9409DD" box="[804,879,713,738]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1893,1916]" captionTargetBox="[295,1278,455,1846]" captionTargetId="figure@6.[246,1334,427,1873]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Psephenops triangularis sp. nov., male: 9) habitus, dorsal view; 10) habitus, ventral view; 11) A variant male, prosternum and prosternal process; 12) Ídem, dorsal view, with antennae 10 - segmented (in alcohol). Figs 9 – 10, dry specimens. Scale bar for Figs 9, 10 and 12 = 0.5 mm." httpUri="https://zenodo.org/record/897419/files/figure.png" pageId="5" pageNumber="114">Fig.11</figureCitation>
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). Eyes lateral, rounded, very prominent, dark red, with a yellow post-ocular area. Labrum subrectangular, similar to clypeus but shorter. Maxillary palpi 4- segmented, covered with short setae; basal palpomere smallest, apical palpomere largest, relative length of palpomeres: 0.04, 0.6, 0.05, 0.11; palpomeres 1–3 subconical, palpomere 4 acuminate. Labial palpi very short, 3- segmented, palpomeres 1–2 rounded and wide, palpomere 3 subconical, nipple-shaped, slightly longer than preceding ones.
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</paragraph>
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<paragraph id="362CDE4D8141EF4116FE09D8FD600EF5" blockId="5.[151,1437,252,2015]" pageId="5" pageNumber="114">
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<emphasis id="04E7025F8141EF4116FE09D8FEDA0889" box="[199,289,925,950]" italics="true" pageId="5" pageNumber="114">Thorax:</emphasis>
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Pronotum subtrapezoidal, almost as wide as elytral base; anterior margin arcuate, posterior margin bisinuate and
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<quantity id="F16B73A88141EF4117050987FE5F08E6" box="[316,420,962,986]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="114" unit="mm" value="0.9">0.90 mm</quantity>
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wide; lateral margin
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||
<quantity id="F16B73A88141EF41149C0984FCF508E6" box="[677,782,961,986]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="114" unit="mm" value="0.52">0.52 mm</quantity>
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wide, slightly curved along anterior half and expanded laterally on posterior half; posterolateral angles subacute. Pronotal disc with a well-developed, dorsomedial, longitudinal carina on the distal half, and a wide, oval, shallow depression to each side of carina; pronotal surface punctate, completely covered with short yellow setae, and long, dark red setae, more dense on disc (
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<figureCitation id="AEA8C2C88141EF41137D0E6EFA750F7B" box="[1348,1422,1067,1092]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1893,1916]" captionTargetBox="[295,1278,455,1846]" captionTargetId="figure@6.[246,1334,427,1873]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Psephenops triangularis sp. nov., male: 9) habitus, dorsal view; 10) habitus, ventral view; 11) A variant male, prosternum and prosternal process; 12) Ídem, dorsal view, with antennae 10 - segmented (in alcohol). Figs 9 – 10, dry specimens. Scale bar for Figs 9, 10 and 12 = 0.5 mm." httpUri="https://zenodo.org/record/897419/files/figure.png" pageId="5" pageNumber="114">Fig. 9</figureCitation>
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). Scutellum short, wider basally than long, apex rounded. Elytra reddish-yellow, covered with short, yellow setae with golden reflections, and long, dark red setae (
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<figureCitation id="AEA8C2C88141EF4114CE0E37FCB90FB4" box="[759,834,1138,1163]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1893,1916]" captionTargetBox="[295,1278,455,1846]" captionTargetId="figure@6.[246,1334,427,1873]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Psephenops triangularis sp. nov., male: 9) habitus, dorsal view; 10) habitus, ventral view; 11) A variant male, prosternum and prosternal process; 12) Ídem, dorsal view, with antennae 10 - segmented (in alcohol). Figs 9 – 10, dry specimens. Scale bar for Figs 9, 10 and 12 = 0.5 mm." httpUri="https://zenodo.org/record/897419/files/figure.png" pageId="5" pageNumber="114">Fig. 9</figureCitation>
|
||
); lateral margins parallel on basal 0.75, gradually converging posteriorly; humeral width 1.0 mm. Prosternum short, with anterolateral corners subtrapezoidal; prosternal process long and wide, flat, slightly carinate, with lateral margins not raised, ending in a lanceolate tip (
|
||
<figureCitation id="AEA8C2C88141EF4116A60E98FED90FCA" box="[159,290,1245,1270]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1893,1916]" captionTargetBox="[295,1278,455,1846]" captionTargetId="figure@6.[246,1334,427,1873]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Psephenops triangularis sp. nov., male: 9) habitus, dorsal view; 10) habitus, ventral view; 11) A variant male, prosternum and prosternal process; 12) Ídem, dorsal view, with antennae 10 - segmented (in alcohol). Figs 9 – 10, dry specimens. Scale bar for Figs 9, 10 and 12 = 0.5 mm." httpUri="https://zenodo.org/record/897419/files/figure.png" pageId="5" pageNumber="114">Figs 10–11</figureCitation>
|
||
) which reaches anterior half of mesoventrite. Mesoventrite shorter than metaventrite, with a wide groove for reception of prosternal process. Metaventrite large and bulky. Legs: Femora robust; tibiae slender, metatibia longer than metafemur, protibia with a tiny apicolateral denticle. Tarsomeres 1 and 2 ventrally lobed with a thick sponge-like vestiture; ventral lobe of tarsomere 1 extending distally over basal 0.30 of tarsomere 2; tarsomere 2 the longest, twice as long as tarsomere 1, extending distally and completely covering tarsomeres 3–4, and basal 0.80 of tarsomere 5; proportional length of tarsomeres: 0.50, 1.1, 0.18, 0.15, 0.55; apical claws long, curved, slender, each with a small basal tooth.
|
||
</paragraph>
|
||
<paragraph id="362CDE4D8141EF4116FE0F90FE410C4C" blockId="5.[151,1437,252,2015]" pageId="5" pageNumber="114">
|
||
<emphasis id="04E7025F8141EF4116FE0F90FEC10ED1" box="[199,314,1493,1518]" italics="true" pageId="5" pageNumber="114">Abdomen:</emphasis>
|
||
Vestiture short, fine and dense, yellowish-gray with golden reflections. Ventrite VI usually hidden under ventrite V; ventrites I and II wide, with posterior margin curved anteriorly in the middle; ventrite III with posterior margin curved posteriorly in the middle, ventrite IV narrow, with posterior margin straight; ventrite V with posterior margin slightly emarginate; ventrite VI very short, subtriangular, with posterior margin widely curved; ventrite VII or anal plate, oval, subtriangular, narrow, covered with long yellowish-gray setae (
|
||
<figureCitation id="AEA8C2C88141EF4113010C26FA750D44" box="[1336,1422,1635,1660]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1893,1916]" captionTargetBox="[295,1278,455,1846]" captionTargetId="figure@6.[246,1334,427,1873]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Psephenops triangularis sp. nov., male: 9) habitus, dorsal view; 10) habitus, ventral view; 11) A variant male, prosternum and prosternal process; 12) Ídem, dorsal view, with antennae 10 - segmented (in alcohol). Figs 9 – 10, dry specimens. Scale bar for Figs 9, 10 and 12 = 0.5 mm." httpUri="https://zenodo.org/record/897419/files/figure.png" pageId="5" pageNumber="114">Fig. 10</figureCitation>
|
||
); pygidium subtriangular covered with long dark red setae. Genitalia: Short, wide, subrectagular, trilobate (
|
||
<figureCitation id="AEA8C2C88141EF41130A0CC3FF4D0DFD" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1970,1993]" captionTargetBox="[271,1339,203,1949]" captionTargetId="figure@7.[233,1359,193,1949]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Psephenops triangularis sp. nov., male genitalia: 13) dorsal; 14) ventral; 15) lateral; 16) ventrolateral. Scale bar = 0.12 mm." httpUri="https://zenodo.org/record/897421/files/figure.png" pageId="5" pageNumber="114">Figs. 13– 14</figureCitation>
|
||
); total length,
|
||
<quantity id="F16B73A88141EF4117590CEFFE310DFD" box="[352,458,1706,1731]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="114" unit="mm" value="0.46">0.46 mm</quantity>
|
||
; total width,
|
||
<quantity id="F16B73A88141EF41145C0CEFFD340DFD" box="[613,719,1706,1730]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="114" unit="mm" value="0.2">0.20 mm</quantity>
|
||
. Parameres robust,
|
||
<quantity id="F16B73A88141EF4115880CEFFBE30DFD" box="[945,1048,1706,1731]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="114" unit="mm" value="0.25">0.25 mm</quantity>
|
||
long, longer than penis; internal margins fused in dorsal view (
|
||
<figureCitation id="AEA8C2C88141EF4117D50C88FDC40DD9" box="[492,575,1741,1766]" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1970,1993]" captionTargetBox="[271,1339,203,1949]" captionTargetId="figure@7.[233,1359,193,1949]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Psephenops triangularis sp. nov., male genitalia: 13) dorsal; 14) ventral; 15) lateral; 16) ventrolateral. Scale bar = 0.12 mm." httpUri="https://zenodo.org/record/897421/files/figure.png" pageId="5" pageNumber="114">Fig. 13</figureCitation>
|
||
), in ventrolateral view with an evident tooth on internal margin near to apex (
|
||
<figureCitation id="AEA8C2C88141EF4116A60CB4FF000C35" box="[159,251,1777,1802]" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1970,1993]" captionTargetBox="[271,1339,203,1949]" captionTargetId="figure@7.[233,1359,193,1949]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Psephenops triangularis sp. nov., male genitalia: 13) dorsal; 14) ventral; 15) lateral; 16) ventrolateral. Scale bar = 0.12 mm." httpUri="https://zenodo.org/record/897421/files/figure.png" pageId="5" pageNumber="114">Fig. 16</figureCitation>
|
||
). Penis
|
||
<quantity id="F16B73A88141EF41175D0CB4FE2A0C36" box="[356,465,1777,1801]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="114" unit="mm" value="0.21">0.21 mm</quantity>
|
||
long, shorter than parameres (
|
||
<figureCitation id="AEA8C2C88141EF4115680CB4FC270C35" box="[849,988,1777,1802]" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1970,1993]" captionTargetBox="[271,1339,203,1949]" captionTargetId="figure@7.[233,1359,193,1949]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Psephenops triangularis sp. nov., male genitalia: 13) dorsal; 14) ventral; 15) lateral; 16) ventrolateral. Scale bar = 0.12 mm." httpUri="https://zenodo.org/record/897421/files/figure.png" pageId="5" pageNumber="114">Figs 14–15</figureCitation>
|
||
); in ventral view finger-like, with a subtriangular lateral projection to each side of basal 0.50 of penis, bearing a short, wide, hourglass-shaped sclerite, which is 0.20 as long as the penis. Phallobase concave in dorsal view; bulky, subcylindrical in ventral view; length
|
||
<quantity id="F16B73A88141EF4116AE0D19FF060C4C" box="[151,253,1884,1908]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="114" unit="mm" value="0.23">0.23 mm</quantity>
|
||
, width
|
||
<quantity id="F16B73A88141EF4117760D19FE4D0C4C" box="[335,438,1884,1908]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="114" unit="mm" value="0.2">0.20 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="362CDE4D8141EF4116FE0D3BFBBB0CE1" blockId="5.[151,1437,252,2015]" pageId="5" pageNumber="114">
|
||
<emphasis id="04E7025F8141EF4116FE0D3BFEC70CA8" bold="true" box="[199,316,1918,1943]" pageId="5" pageNumber="114">Variation.</emphasis>
|
||
A specimen with a similar genital structure was collected, but with a yellow pronotum, and 10- segmented, yellowish-red antennae (
|
||
<figureCitation id="AEA8C2C88141EF4114780DE7FD330C84" box="[577,712,1954,1979]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1893,1916]" captionTargetBox="[295,1278,455,1846]" captionTargetId="figure@6.[246,1334,427,1873]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Psephenops triangularis sp. nov., male: 9) habitus, dorsal view; 10) habitus, ventral view; 11) A variant male, prosternum and prosternal process; 12) Ídem, dorsal view, with antennae 10 - segmented (in alcohol). Figs 9 – 10, dry specimens. Scale bar for Figs 9, 10 and 12 = 0.5 mm." httpUri="https://zenodo.org/record/897419/files/figure.png" pageId="5" pageNumber="114">Figs 11–12</figureCitation>
|
||
).
|
||
<typeStatus id="E92860EF8141EF4114DB0DE6FCBF0C84" box="[738,836,1955,1979]" pageId="5" pageNumber="114" type="paratype">Paratype</typeStatus>
|
||
measurements (in mm) [N=1]: Total length, 2.45. Pronotum: posterior margin, 0.85; lateral margin, 0.50. Elytra: humeral width, 0.90.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7E898DC68142EF4216FE0AD2FD7E0BEA" pageId="6" pageNumber="115" type="etymology">
|
||
<paragraph id="362CDE4D8142EF4216FE0AD2FD7E0BEA" blockId="6.[151,1436,151,357]" pageId="6" pageNumber="115">
|
||
<emphasis id="04E7025F8142EF4216FE0AD2FEB50B8F" bold="true" box="[199,334,151,176]" pageId="6" pageNumber="115">Etymology.</emphasis>
|
||
From the latin adjective “
|
||
<taxonomicName id="F193A5CE8142EF42144D0AD2FD030B8F" authorityName="Novelo-Gutiérrez" authorityYear="2017" box="[628,760,151,176]" class="Insecta" family="Psephenidae" genus="Pspehenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="115" phylum="Arthropoda" rank="species" species="triangularis">triangularis</taxonomicName>
|
||
” (with shape of triangle), this species is named
|
||
<taxonomicName id="F193A5CE8142EF42132F0AD2FA670B8F" authorityName="Novelo-Gutiérrez" authorityYear="2017" box="[1302,1436,151,176]" class="Insecta" family="Psephenidae" genus="Pspehenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="115" phylum="Arthropoda" rank="species" species="triangularis">
|
||
<emphasis id="04E7025F8142EF42132F0AD2FA670B8F" box="[1302,1436,151,176]" italics="true" pageId="6" pageNumber="115">triangularis</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the penis lateral projections.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7E898DC68142EF4216FE0A9AFC8C0A22" pageId="6" pageNumber="115" type="biology_ecology">
|
||
<paragraph id="362CDE4D8142EF4216FE0A9AFC8C0A22" blockId="6.[151,1436,151,357]" pageId="6" pageNumber="115">
|
||
<emphasis id="04E7025F8142EF4216FE0A9AFEDF0BC7" bold="true" box="[199,292,223,248]" pageId="6" pageNumber="115">Ecology</emphasis>
|
||
. The two specimens of
|
||
<taxonomicName id="F193A5CE8142EF4214150AA4FD360BC7" authority="Novelo-Gutiérrez, 2017" authorityName="Novelo-Gutiérrez" authorityYear="2017" box="[556,717,223,248]" class="Insecta" family="Psephenidae" genus="Pspehenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="115" phylum="Arthropoda" rank="species" species="triangularis" status="sp. nov.">
|
||
<emphasis id="04E7025F8142EF4214150AA4FD360BC7" box="[556,717,223,248]" italics="true" pageId="6" pageNumber="115">P. triangularis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="04E7025F8142EF4214EA0A9AFCD70BC7" bold="true" box="[723,812,223,248]" pageId="6" pageNumber="115">
|
||
<taxonomicNameLabel id="1FD4BF248142EF4214EA0A9AFCD70BC7" box="[723,812,223,248]" pageId="6" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were collected flying over turbulent rapids in the center of the stream, together with some specimens of
|
||
<taxonomicName id="F193A5CE8142EF4214970B40FCED0A22" authority="Novelo-Gutiérrez, 2017" authorityName="Novelo-Gutiérrez" authorityYear="2017" box="[686,790,260,285]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="115" phylum="Arthropoda" rank="species" species="bifidus" status="sp. nov.">
|
||
<emphasis id="04E7025F8142EF4214970B40FCED0A22" box="[686,790,260,285]" italics="true" pageId="6" pageNumber="115">P. bifidus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="04E7025F8142EF4215240B41FC8C0A22" bold="true" box="[797,887,260,285]" pageId="6" pageNumber="115">
|
||
<taxonomicNameLabel id="1FD4BF248142EF4215240B41FC8C0A22" box="[797,887,260,285]" pageId="6" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7E898DC68142EF4216FE0B62FE030A5B" pageId="6" pageNumber="115" type="discussion">
|
||
<paragraph id="362CDE4D8142EF4216FE0B62FE030A5B" blockId="6.[151,1436,151,357]" pageId="6" pageNumber="115">
|
||
<emphasis id="04E7025F8142EF4216FE0B62FEC00A7F" bold="true" box="[199,315,295,320]" pageId="6" pageNumber="115">Remarks.</emphasis>
|
||
<taxonomicName id="F193A5CE8142EF42177D0B62FDA10A7F" authority="Novelo-Gutiérrez, 2017" authorityName="Novelo-Gutiérrez" authorityYear="2017" box="[324,602,295,320]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="115" phylum="Arthropoda" rank="species" species="triangularis" status="sp. nov.">
|
||
<emphasis id="04E7025F8142EF42177D0B62FDA10A7F" box="[324,602,295,320]" italics="true" pageId="6" pageNumber="115">Psephenops triangularis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="04E7025F8142EF42145A0B62FD440A7F" bold="true" box="[611,703,295,320]" pageId="6" pageNumber="115">
|
||
<taxonomicNameLabel id="1FD4BF248142EF42145A0B62FD440A7F" box="[611,703,295,320]" pageId="6" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be separated from other known
|
||
<taxonomicName id="F193A5CE8142EF4212530B62FB140A7F" authorityName="Grouvelle" authorityYear="1898" box="[1130,1263,295,320]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="115" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="04E7025F8142EF4212530B62FB140A7F" box="[1130,1263,295,320]" italics="true" pageId="6" pageNumber="115">Psephenops</emphasis>
|
||
</taxonomicName>
|
||
species by the characteristics listed in
|
||
<tableCitation id="7B11EBF68142EF4217A70B09FE090A5B" box="[414,498,332,357]" captionStart="TABLE 1" captionStartId="4.[120,193,558,578]" captionText="TABLE 1. Comparative features of the species of Psephenops Grouvelle from Mesoamerica." pageId="6" pageNumber="115">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |