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<document id="7E724FF3C71C8F21B0BA87979B974103" ID-CLB-Dataset="22067" ID-DOI="10.5852/ejt.2017.337" ID-GBIF-Dataset="5a7f73c5-5646-446f-a72c-3ea18d8f634f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832483" ID-ZooBank="2669CB69-5064-4B5D-AC25-856680E3B9AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589830819450" checkinUser="carolina" docAuthor="Budha, Prem B., Naggs, Fred &amp; Backeljau, Thierry" docDate="2017" docId="03C6CF41FFF4FFFBFD94451FD6E4F868" docLanguage="en" docName="ejt-2017-337.pdf.imf" docOrigin="European Journal of Taxonomy 337" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Diplommatina regularis Fulton 1901" docType="treatment" docVersion="6" lastPageNumber="17" masterDocId="FFFFB739FFE4FFEBFFAE4762D552FF83" masterDocTitle="The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="17" updateTime="1698840088040" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="83372707A2330C652D571F682F54253F">The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species</mods:title>
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<mods:namePart id="7D7C84C2096C70040CDA618D50053B96">Budha, Prem B.</mods:namePart>
<mods:affiliation id="CE8C22B015B8C4296D7D0EBFC5901C2D">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Central Department of Zoology, Tribhuvan University, Kirtipur, Kathmandu, Nepal. &amp; Corresponding author: prembudha @ yahoo. com</mods:affiliation>
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<mods:namePart id="2BA8C06350245EF684739A7840125964">Naggs, Fred</mods:namePart>
<mods:affiliation id="C4A7D6114B7C69BDD7BC1BC690C4FE25">Natural History Museum, Cromwell Road, London, SW 7 5 BD, UK. &amp; Email: f. naggs @ nhm. ac. uk</mods:affiliation>
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<mods:namePart id="FC34B8D6980EEE1156BDDBFC91D5DB51">Backeljau, Thierry</mods:namePart>
<mods:affiliation id="32B4A2D2D3B2EE4AC55153366F3ED517">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B- 1000, Brussels, Belgium. &amp; Email: thierry. backeljau @ naturalsciences. be</mods:affiliation>
<mods:nameIdentifier id="F91A70F5A36762378942D4A8FA9D4111" type="email">thierry.backeljau@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="CAAD4966DC1572A0BA17116E58477A99">European Journal of Taxonomy</mods:title>
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<mods:date id="EED77E9C5E56E6F1C026FDD18F418A64">2017</mods:date>
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<mods:number id="5CF01AB22235D3AA001978344018DB72">2017-07-21</mods:number>
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<treatment id="03C6CF41FFF4FFFBFD94451FD6E4F868" ID-DOI="http://doi.org/10.5281/zenodo.3850694" ID-GBIF-Taxon="164167124" ID-Zenodo-Dep="3850694" LSID="urn:lsid:plazi:treatment:03C6CF41FFF4FFFBFD94451FD6E4F868" httpUri="http://treatment.plazi.org/id/03C6CF41FFF4FFFBFD94451FD6E4F868" lastPageNumber="17" pageId="16" pageNumber="17">
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<taxonomicName id="4C6F05D4FFF4FFFBFD94451FD6A8FD14" ID-CoL="36M3R" authority="Fulton, 1901" authorityName="Fulton" authorityYear="1901" box="[570,1018,637,664]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="regularis">
<emphasis id="B91BA245FFF4FFFBFD94451FD60EFD1B" bold="true" box="[570,860,637,664]" italics="true" pageId="16" pageNumber="17">Diplommatina regularis</emphasis>
<bibRefCitation id="EFFE03A6FFF4FFFBFCCA451FD6A8FD14" author="Fulton H." box="[868,1018,637,663]" pageId="16" pageNumber="17" pagination="242 - 245" refId="ref13161" refString="Fulton H. 1901. Descriptions of some supposed new species of Diplommatina, Opisthostoma, and a new variety of Alycaeus from N. Borneo, Banguey Island and Darjeeling. Annals and Magazine of Natural History, Series 7 8: 242 - 245. https: // doi. org / 10.1080 / 03745480109442915" type="journal article" year="1901">Fulton, 1901</bibRefCitation>
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<subSubSection id="C3752DDCFFF4FFFBFD7845C1D60FFD3D" box="[726,861,675,702]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BD07E57FFF4FFFBFD7845C1D60FFD3D" blockId="16.[570,1018,637,702]" box="[726,861,675,702]" pageId="16" pageNumber="17">
<figureCitation id="135462D2FFF4FFFBFD7845C1D665FD3D" box="[726,823,675,702]" captionStart="Fig" captionStartId="17.[189,232,1930,1956]" captionTargetBox="[189,1397,264,1875]" captionTargetId="figure@17.[189,1397,261,1877]" captionTargetPageId="17" captionText="Fig. 7. Diplommatina Benson, 1849 of Nepal. A. D. regularis Fulton, 1901 (CDZMTU045). B. D. salgharica sp. nov., holotype (CDZMTU046). C. D. shivapuriensis sp. nov., holotype (CDZMTU048). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832498" httpUri="https://zenodo.org/record/3832498/files/figure.png" pageId="16" pageNumber="17">Figs 7A</figureCitation>
,
<figureCitation id="135462D2FFF4FFFBFCED45C6D60FFD3D" box="[835,861,676,702]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="16" pageNumber="17">9J</figureCitation>
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<treatmentCitation id="0ACE5846FFF4FFFBFF13458AD7E7FC80" author="Fulton H." box="[189,693,744,771]" page="245" pageId="16" pageNumber="17" year="1901">
<taxonomicName id="4C6F05D4FFF4FFFBFF13458AD7E7FC80" ID-CoL="36M3R" authority="Fulton, 1901: 245" authorityName="Fulton" authorityPageNumber="245" authorityYear="1901" box="[189,693,744,771]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="regularis">
<emphasis id="B91BA245FFF4FFFBFF13458AD48AFC81" box="[189,472,744,770]" italics="true" pageId="16" pageNumber="17">Diplommatina regularis</emphasis>
<bibRefCitation id="EFFE03A6FFF4FFFBFE4E458AD7E7FC80" author="Fulton H." box="[480,693,744,771]" pageId="16" pageNumber="17" pagination="242 - 245" refId="ref13161" refString="Fulton H. 1901. Descriptions of some supposed new species of Diplommatina, Opisthostoma, and a new variety of Alycaeus from N. Borneo, Banguey Island and Darjeeling. Annals and Magazine of Natural History, Series 7 8: 242 - 245. https: // doi. org / 10.1080 / 03745480109442915" type="journal article" year="1901">Fulton, 1901: 245</bibRefCitation>
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.
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</paragraph>
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<treatmentCitationGroup id="AB7F5979FFF4FFFBFF13444ED6E9FCC4" box="[189,955,812,839]" pageId="16" pageNumber="17">
<taxonomicName id="4C6F05D4FFF4FFFBFF13444ED48AFCC5" ID-CoL="36M3R" authorityName="Fulton" authorityYear="1901" box="[189,472,812,838]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="regularis">
<emphasis id="B91BA245FFF4FFFBFF13444ED48AFCC5" box="[189,472,812,838]" italics="true" pageId="16" pageNumber="17">Diplommatina regularis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ACE5846FFF4FFFBFE58444ED795FCC4" author="Kobelt W." box="[502,711,812,839]" page="442" pageId="16" pageNumber="17" year="1902">
<bibRefCitation id="EFFE03A6FFF4FFFBFE58444ED795FCC4" author="Kobelt W." box="[502,711,812,839]" pageId="16" pageNumber="17" refId="ref13702" refString="Kobelt W. 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. Mollusca: Cyclophoridae, Lief. 16. R. Friedlander und Sohn Verlag, Berlin." type="book" year="1902">Kobelt 1902: 442</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ACE5846FFF4FFFBFD58444ED6E5FCC4" author="Gude G. K." box="[758,951,812,839]" page="330" pageId="16" pageNumber="17" year="1921">
<bibRefCitation id="EFFE03A6FFF4FFFBFD58444ED6E5FCC4" author="Gude G. K." box="[758,951,812,839]" pageId="16" pageNumber="17" refId="ref13576" refString="Gude G. K. 1921. The fauna of British India. Mollusca-III (Land operculates: Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Today &amp; Tomorrow's Printers &amp; Publishers, New Delhi." type="book" year="1921">Gude 1921: 330</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BD07E57FFF4FFFBFF134431D753FC17" blockId="16.[189,1398,812,916]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB7F5979FFF4FFFBFF134431D753FC17" pageId="16" pageNumber="17">
<taxonomicName id="4C6F05D4FFF4FFFBFF134431D7CCFCEE" ID-CoL="36M3R" authorityName="Fulton" authorityYear="1901" box="[189,670,851,877]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="regularis" subGenus="Diplommatina">
<emphasis id="B91BA245FFF4FFFBFF134431D7CCFCEE" box="[189,670,851,877]" italics="true" pageId="16" pageNumber="17">Diplommatina (Diplommatina) regularis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ACE5846FFF4FFFBFD104431D622FCEE" author="Zilch V. A." box="[702,880,851,878]" page="21" pageId="16" pageNumber="17" year="1953">
<bibRefCitation id="EFFE03A6FFF4FFFBFD104431D622FCEE" author="Zilch V. A." box="[702,880,851,878]" pageId="16" pageNumber="17" pagination="1 - 47" refId="ref14451" refString="Zilch V. A. 1953. Die Typen und Typoide des Natur-Museums Senckenberg, 9: Mollusca, Cyclophoridae, Diplommatininae. Archiv fur Molluskenkunde 82: 1 - 47." type="journal article" year="1953">Zilch 1953: 21</bibRefCitation>
</treatmentCitation>
, figs 125–126. —
<treatmentCitation id="0ACE5846FFF4FFFBFBE24431D58FFC10" author="Ramakrishna &amp; Mitra S. C. &amp; Dey A." page="91" pageId="16" pageNumber="17" year="2010">
<bibRefCitation id="EFFE03A6FFF4FFFBFBE24431D58FFC10" author="Ramakrishna &amp; Mitra S. C. &amp; Dey A." pageId="16" pageNumber="17" pagination="1 - 359" refId="ref14255" refString="Ramakrishna, Mitra S. C. &amp; Dey A. 2010. Annotated checklist of Indian land molluscs. Records of Zoological Survey of India, Occasional Paper 306: 1 - 359." type="journal article" year="2010">
Ramakrishna
<emphasis id="B91BA245FFF4FFFBFB414436D079FCEE" box="[1263,1323,851,877]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2010: 91
</bibRefCitation>
</treatmentCitation>
.
<emphasis id="B91BA245FFF4FFFBFF474418D451FC10" box="[233,259,890,915]" italics="true" pageId="16" pageNumber="17">—</emphasis>
<treatmentCitation id="0ACE5846FFF4FFFBFEA4441BD4A9FC17" author="Budha P. B. &amp; Naggs F. &amp; Backeljau T." box="[266,507,889,916]" page="7" pageId="16" pageNumber="17" year="2015">
<bibRefCitation id="EFFE03A6FFF4FFFBFEA4441BD4A9FC17" author="Budha P. B. &amp; Naggs F. &amp; Backeljau T." box="[266,507,889,916]" pageId="16" pageNumber="17" pagination="1 - 48" refId="ref13118" refString="Budha P. B., Naggs F. &amp; Backeljau T. 2015. Annotated checklist of the terrestrial gastropods of Nepal. ZooKeys 492: 1 - 48. https: // doi. org / 10.3897 / zookeys. 492.9175" type="journal article" year="2015">
Budha
<emphasis id="B91BA245FFF4FFFBFEF04418D4CBFC10" box="[350,409,889,915]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2015: 7
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFF4FFFBFF1344DFD03AFA0B" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BD07E57FFF4FFFBFF1344DFD464FC54" blockId="16.[189,310,957,983]" box="[189,310,957,983]" pageId="16" pageNumber="17">
<heading id="D098C93BFFF4FFFBFF1344DFD464FC54" bold="true" box="[189,310,957,983]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B91BA245FFF4FFFBFF1344DFD464FC54" bold="true" box="[189,310,957,983]" pageId="16" pageNumber="17">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFF4FFFBFF134492D03AFA0B" blockId="16.[189,1399,1007,1416]" pageId="16" pageNumber="17">
Measurements (n = 8): SH 2.5–3.0 mm, SW
<quantity id="4C97D3B2FFF4FFFBFD42448DD6D3FB89" box="[748,897,1007,1034]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="16" pageNumber="17" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
, Wh 7.0–7.5. Dextral, penultimate whorl slightly larger than the body whorl. Constriction at columellar margin slightly towards the inner lip of the parietal wall (
<figureCitation id="135462D2FFF4FFFBFEF2435ED4FDFBD5" box="[348,431,1084,1110]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="16" pageNumber="17">Fig. 9J</figureCitation>
). One parietalis, short (
<figureCitation id="135462D2FFF4FFFBFD12435ED648FBD5" box="[700,794,1084,1110]" captionStart="Fig" captionStartId="17.[189,232,1930,1956]" captionTargetBox="[189,1397,264,1875]" captionTargetId="figure@17.[189,1397,261,1877]" captionTargetPageId="17" captionText="Fig. 7. Diplommatina Benson, 1849 of Nepal. A. D. regularis Fulton, 1901 (CDZMTU045). B. D. salgharica sp. nov., holotype (CDZMTU046). C. D. shivapuriensis sp. nov., holotype (CDZMTU048). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832498" httpUri="https://zenodo.org/record/3832498/files/figure.png" pageId="16" pageNumber="17">Fig. 7A</figureCitation>
<figureCitation id="135462D2FFF4FFFBFCB4435ED67AFBD5" box="[794,808,1084,1110]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="16" pageNumber="17">3</figureCitation>
). One vertical palatalis, parallel to the constriction. One short horizontal palatalis, situated just above the columella (
<figureCitation id="135462D2FFF4FFFBFC734300D167FBFF" box="[989,1077,1122,1148]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="16" pageNumber="17">Fig. 9J</figureCitation>
). Columellaris moderately thin. Columellar tooth weak, only visible in mature shells (
<figureCitation id="135462D2FFF4FFFBFCD143EAD6B2FB21" box="[895,992,1160,1186]" captionStart="Fig" captionStartId="17.[189,232,1930,1956]" captionTargetBox="[189,1397,264,1875]" captionTargetId="figure@17.[189,1397,261,1877]" captionTargetPageId="17" captionText="Fig. 7. Diplommatina Benson, 1849 of Nepal. A. D. regularis Fulton, 1901 (CDZMTU045). B. D. salgharica sp. nov., holotype (CDZMTU046). C. D. shivapuriensis sp. nov., holotype (CDZMTU048). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832498" httpUri="https://zenodo.org/record/3832498/files/figure.png" pageId="16" pageNumber="17">Fig. 7A</figureCitation>
<figureCitation id="135462D2FFF4FFFBFC4E43EAD6BCFB21" box="[992,1006,1160,1186]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="16" pageNumber="17">3</figureCitation>
). Protoconch smooth, with small pits distributed all over (
<figureCitation id="135462D2FFF4FFFBFE7043CCD712FB4A" box="[478,576,1198,1225]" captionStart="Fig" captionStartId="17.[189,232,1930,1956]" captionTargetBox="[189,1397,264,1875]" captionTargetId="figure@17.[189,1397,261,1877]" captionTargetPageId="17" captionText="Fig. 7. Diplommatina Benson, 1849 of Nepal. A. D. regularis Fulton, 1901 (CDZMTU045). B. D. salgharica sp. nov., holotype (CDZMTU046). C. D. shivapuriensis sp. nov., holotype (CDZMTU048). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832498" httpUri="https://zenodo.org/record/3832498/files/figure.png" pageId="16" pageNumber="17">Fig. 7A</figureCitation>
<figureCitation id="135462D2FFF4FFFBFDEF43CDD71EFB4A" box="[577,588,1199,1225]" captionStart="Fig" captionStartId="2.[189,235,1999,2025]" captionTargetBox="[189,1399,1129,1958]" captionTargetId="figure@2.[187,1401,1126,1960]" captionTargetPageId="2" captionText="Fig.1. Map of the localities and areas surveyed for Diplommatina spp. in Nepal." figureDoi="http://doi.org/10.5281/zenodo.3832486" httpUri="https://zenodo.org/record/3832486/files/figure.png" pageId="16" pageNumber="17">1</figureCitation>
). Teleoconch with regular radial ribs; about 8–11 ribs/
<quantity id="4C97D3B2FFF4FFFBFB6343CCD074FB4A" box="[1229,1318,1198,1225]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="mm" value="0.5">0.5 mm</quantity>
on the penultimate whorl; 5–8 ribs/
<quantity id="4C97D3B2FFF4FFFBFDA143B7D73BFB6C" box="[527,617,1237,1264]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="mm" value="0.5">0.5 mm</quantity>
on the body whorl towards the aperture (
<figureCitation id="135462D2FFF4FFFBFBFB43B7D1E1FB6C" box="[1109,1203,1237,1263]" captionStart="Fig" captionStartId="17.[189,232,1930,1956]" captionTargetBox="[189,1397,264,1875]" captionTargetId="figure@17.[189,1397,261,1877]" captionTargetPageId="17" captionText="Fig. 7. Diplommatina Benson, 1849 of Nepal. A. D. regularis Fulton, 1901 (CDZMTU045). B. D. salgharica sp. nov., holotype (CDZMTU046). C. D. shivapuriensis sp. nov., holotype (CDZMTU048). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832498" httpUri="https://zenodo.org/record/3832498/files/figure.png" pageId="16" pageNumber="17">Fig. 7A</figureCitation>
<figureCitation id="135462D2FFF4FFFBFB1D43B7D190FB6C" box="[1203,1218,1237,1263]" captionStart="Fig" captionStartId="4.[189,232,1895,1921]" captionTargetBox="[189,1398,285,1853]" captionTargetId="figure@4.[189,1398,285,1853]" captionTargetPageId="4" captionText="Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two)." figureDoi="http://doi.org/10.5281/zenodo.3832488" httpUri="https://zenodo.org/record/3832488/files/figure.png" pageId="16" pageNumber="17">2</figureCitation>
, A
<figureCitation id="135462D2FFF4FFFBFB4843B7D1A4FB6C" box="[1254,1270,1237,1263]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="16" pageNumber="17">4</figureCitation>
). Very fine spiral striation present. Umbilicus closed. Outer lip at BeP abruptly curved on the columella, closing the umbilicus. Peristome thin, with two close lips (
<figureCitation id="135462D2FFF4FFFBFCB64243D624FAB8" box="[792,886,1313,1339]" captionStart="Fig" captionStartId="17.[189,232,1930,1956]" captionTargetBox="[189,1397,264,1875]" captionTargetId="figure@17.[189,1397,261,1877]" captionTargetPageId="17" captionText="Fig. 7. Diplommatina Benson, 1849 of Nepal. A. D. regularis Fulton, 1901 (CDZMTU045). B. D. salgharica sp. nov., holotype (CDZMTU046). C. D. shivapuriensis sp. nov., holotype (CDZMTU048). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832498" httpUri="https://zenodo.org/record/3832498/files/figure.png" pageId="16" pageNumber="17">Fig. 7A</figureCitation>
<figureCitation id="135462D2FFF4FFFBFCD84243D6D4FAB8" box="[886,902,1313,1339]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="16" pageNumber="17">4</figureCitation>
); AeP slightly curved, a single lip in most immature shells. Very thin parietal callus, extending at the base. Operculum paucispiral, corneous, thin, transparent, slightly concave, outer surface smooth, inner surface with raised peripheral circular edge.
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFF4FFFBFF1342D0D012F8AA" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BD07E57FFF4FFFBFF1342D0D4F8FA4F" blockId="16.[189,426,1458,1484]" box="[189,426,1458,1484]" pageId="16" pageNumber="17">
<heading id="D098C93BFFF4FFFBFF1342D0D4F8FA4F" bold="true" box="[189,426,1458,1484]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B91BA245FFF4FFFBFF1342D0D4F8FA4F" bold="true" box="[189,426,1458,1484]" pageId="16" pageNumber="17">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFF4FFFBFF444286D4E5FA7D" blockId="16.[189,1398,1508,1611]" box="[234,439,1508,1534]" pageId="16" pageNumber="17">
<emphasis id="B91BA245FFF4FFFBFF444286D4E5FA7D" bold="true" box="[234,439,1508,1534]" pageId="16" pageNumber="17">
Possible
<typeStatus id="54D4C0F5FFF4FFFBFEF94286D4E5FA7D" box="[343,439,1508,1534]" pageId="16" pageNumber="17" type="syntype">syntype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD07E57FFF4FFFBFF134168D467F9C8" blockId="16.[189,1398,1508,1611]" pageId="16" pageNumber="17">
<collectingCountry id="F3783EC7FFF4FFFBFF134168D447F9A7" box="[189,277,1546,1572]" name="India" pageId="16" pageNumber="17">INDIA</collectingCountry>
: 1 shell [
<taxonomicName id="4C6F05D4FFF4FFFBFE254169D6C3F9A7" authority="Fulton, 1901 Darjeeling Sow." authorityName="Fulton" authorityYear="1901" box="[395,913,1546,1572]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="regularis">
<emphasis id="B91BA245FFF4FFFBFE254169D74CF9A7" box="[395,542,1546,1572]" italics="true" pageId="16" pageNumber="17">D. regularis</emphasis>
<bibRefCitation id="EFFE03A6FFF4FFFBFD844168D797F9A7" author="Fulton H." box="[554,709,1546,1572]" pageId="16" pageNumber="17" pagination="242 - 245" refId="ref13161" refString="Fulton H. 1901. Descriptions of some supposed new species of Diplommatina, Opisthostoma, and a new variety of Alycaeus from N. Borneo, Banguey Island and Darjeeling. Annals and Magazine of Natural History, Series 7 8: 242 - 245. https: // doi. org / 10.1080 / 03745480109442915" type="journal article" year="1901">Fulton, 1901</bibRefCitation>
Darjeeling Sow.
</taxonomicName>
&amp; F.
<date id="FFD15897FFF4FFFBFC4E4168D136F9A7" box="[992,1124,1546,1572]" pageId="16" pageNumber="17" value="1901-01-10">01.10.1901</date>
1/6] (
<collectionCode id="ED7EE692FFF4FFFBFB1D4168D059F9A7" box="[1203,1291,1546,1572]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="16" pageNumber="17" type="Museum">RBINS</collectionCode>
525556, MT1099).
</paragraph>
<paragraph id="8BD07E57FFF4FFFBFF444117D774F90C" blockId="16.[189,1399,1653,1833]" box="[234,550,1653,1679]" pageId="16" pageNumber="17">
<emphasis id="B91BA245FFF4FFFBFF444117D774F90C" bold="true" box="[234,550,1653,1679]" pageId="16" pageNumber="17">Other material examined</emphasis>
</paragraph>
<paragraph id="8BD07E57FFF4FFFBFF1341F9D012F8AA" blockId="16.[189,1399,1653,1833]" pageId="16" pageNumber="17">
<collectingCountry id="F3783EC7FFF4FFFBFF1341F9D44CF936" box="[189,286,1691,1717]" name="Nepal" pageId="16" pageNumber="17">NEPAL</collectingCountry>
: 20 shells, Baghdwar, Shivapuri-Nagarjun National Park,
<quantity id="4C97D3B2FFF4FFFBFC6641F9D171F935" box="[968,1059,1691,1718]" metricMagnitude="3" metricUnit="m" metricValue="2.314" pageId="16" pageNumber="17" unit="m" value="2314.0">2314 m</quantity>
,
<geoCoordinate id="EE5B1890FFF4FFFBFB9E41F9D19FF935" box="[1072,1229,1691,1718]" degrees="27.805203" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="1" value="27.805202">27.805203 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFF4FFFBFB7741F9D023F935" box="[1241,1393,1691,1718]" degrees="85.390029" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="85.39003">85.390029 E</geoCoordinate>
,
<date id="FFD15897FFF4FFFBFF1341A0D40DF95F" box="[189,351,1730,1756]" pageId="16" pageNumber="17" value="2007-06-24">24 Jun. 2007</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFF4FFFBFDEE41A3D791F95F" box="[576,707,1729,1756]" pageId="16" pageNumber="17">CDZMTU</collectionCode>
045); 9 shells, Ghopte, Langtang National Park,
<quantity id="4C97D3B2FFF4FFFBFABC41A0D023F95F" box="[1298,1393,1730,1756]" metricMagnitude="3" metricUnit="m" metricValue="3.4189999999999996" pageId="16" pageNumber="17" unit="m" value="3419.0">3419 m</quantity>
,
<geoCoordinate id="EE5B1890FFF4FFFBFF13418AD412F881" box="[189,320,1768,1794]" degrees="28.0193" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="5" value="28.0193">28.0193 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFF4FFFBFEFE418AD49DF881" box="[336,463,1768,1795]" degrees="85.4619" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="5" value="85.4619">85.4619 E</geoCoordinate>
,
<date id="FFD15897FFF4FFFBFE4E418AD727F881" box="[480,629,1768,1794]" pageId="16" pageNumber="17" value="2007-06-08">8 Jun. 2007</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFF4FFFBFCF8418AD68AF880" box="[854,984,1768,1795]" pageId="16" pageNumber="17">CDZMTU</collectionCode>
060); 16 shells, Deurali, Langtang National Park,
<quantity id="4C97D3B2FFF4FFFBFEC0406CD740F8AA" box="[366,530,1806,1833]" metricMagnitude="3" metricUnit="m" metricValue="3.2619999999999996" metricValueMax="3.2769999999999997" metricValueMin="3.247" pageId="16" pageNumber="17" unit="m" value="3262.0" valueMax="3277.0" valueMin="3247.0">3247–3277 m</quantity>
,
<geoCoordinate id="EE5B1890FFF4FFFBFDB1406CD7C4F8AA" box="[543,662,1806,1833]" degrees="28.1059" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="5" value="28.1059">28.1059N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFF4FFFBFD0C406CD64EF8AA" box="[674,796,1806,1833]" degrees="85.3266" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="5" value="85.3266">85.3266 E</geoCoordinate>
,
<date id="FFD15897FFF4FFFBFC87406CD6E6F8AB" box="[809,948,1806,1833]" pageId="16" pageNumber="17" value="2007-06-06">6 Jun. 2007</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFF4FFFBFB2B406CD05BF8AA" box="[1157,1289,1806,1833]" pageId="16" pageNumber="17">CDZMTU</collectionCode>
061).
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFF4FFFBFF134031D6E4F868" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BD07E57FFF4FFFBFF134031D47DF8EE" blockId="16.[189,303,1875,1901]" box="[189,303,1875,1901]" pageId="16" pageNumber="17">
<heading id="D098C93BFFF4FFFBFF134031D47DF8EE" bold="true" box="[189,303,1875,1901]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B91BA245FFF4FFFBFF134031D47DF8EE" bold="true" box="[189,303,1875,1901]" pageId="16" pageNumber="17">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFF4FFFBFF1340E7D6E4F868" blockId="16.[189,1399,1924,2027]" pageId="16" pageNumber="17">
<taxonomicName id="4C6F05D4FFF4FFFBFF1340E7D419F81D" authorityName="Fulton" authorityYear="1901" box="[189,331,1924,1950]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="regularis">
<emphasis id="B91BA245FFF4FFFBFF1340E7D419F81D" box="[189,331,1924,1950]" italics="true" pageId="16" pageNumber="17">D. regularis</emphasis>
</taxonomicName>
is a very distinct species due to its regularly increasing whorls. Many shells with adult size show a single, very thin peristomal lip typical of juvenile shells. Juvenile shells show a more distinct constriction at the columellar margin than the fully grown shells.
</paragraph>
</subSubSection>
</treatment>
</document>
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