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<mods:title id="6BFE6554F29E96D34914D7C37D0FC791">Synopsis of the terrestrial Reptiles of Equatorial Guinea</mods:title>
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<treatment id="03C287E8FFDDFFECFF4BFC76FB10E4AF" ID-DOI="http://doi.org/10.5281/zenodo.7293531" ID-GBIF-Taxon="204305681" ID-Zenodo-Dep="7293531" LSID="urn:lsid:plazi:treatment:03C287E8FFDDFFECFF4BFC76FB10E4AF" httpUri="http://treatment.plazi.org/id/03C287E8FFDDFFECFF4BFC76FB10E4AF" lastPageNumber="128" pageId="127" pageNumber="128" scope_family="Viperinae" scope_order="Squamata">
<subSubSection id="C3716575FFDDFFECFF4BFC76FDEAE634" box="[151,607,1003,1030]" pageId="127" pageNumber="128" type="nomenclature">
<paragraph id="8BD436FEFFDDFFECFF4BFC76FDEAE634" blockId="127.[151,607,1003,1066]" box="[151,607,1003,1030]" pageId="127" pageNumber="128">
<heading id="D09C8192FFDDFFECFF4BFC76FDEAE634" bold="true" box="[151,607,1003,1030]" fontSize="11" level="2" pageId="127" pageNumber="128" reason="4">
<taxonomicName id="4C6B4D7DFFDDFFECFF4BFC76FDEAE634" ID-CoL="87QVQ" authority="(Hallowell, 1854)" baseAuthorityName="Hallowell" baseAuthorityYear="1854" box="[151,607,1003,1030]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="127" pageNumber="128" phylum="Chordata" rank="species" species="squamigera">
<emphasis id="B91FEAECFFDDFFECFF4BFC76FDEAE634" bold="true" box="[151,607,1003,1030]" pageId="127" pageNumber="128">
<emphasis id="B91FEAECFFDDFFECFF4BFC76FE30E634" bold="true" box="[151,389,1003,1030]" italics="true" pageId="127" pageNumber="128">Atheris squamigera</emphasis>
(Hallowell, 1854)
</emphasis>
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</heading>
</paragraph>
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</paragraph>
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<paragraph id="8BD436FEFFDDFFECFF4BFBC9FABCE65E" blockId="127.[151,1355,1108,1225]" box="[151,1289,1108,1133]" pageId="127" pageNumber="128">
<treatmentCitationGroup id="AB7B11D0FFDDFFECFF4BFBC9FABCE65E" box="[151,1289,1108,1133]" pageId="127" pageNumber="128">
<treatmentCitation id="0ACA10EFFFDDFFECFF4BFBC9FD8CE65F" author="Hallowell, E." box="[151,569,1108,1133]" page="193" pageId="127" pageNumber="128" year="1854">
<taxonomicName id="4C6B4D7DFFDDFFECFF4BFBC9FD8CE65F" authority="Hallowell, 1854 a: 193" authorityName="Hallowell" authorityPageNumber="193" authorityYear="1854" box="[151,569,1108,1133]" class="Reptilia" family="Viperidae" genus="Echis" kingdom="Animalia" order="Squamata" pageId="127" pageNumber="128" phylum="Chordata" rank="species" species="squamigera">
<emphasis id="B91FEAECFFDDFFECFF4BFBC9FEFBE65E" box="[151,334,1108,1132]" italics="true" pageId="127" pageNumber="128">Echis squamigera</emphasis>
<bibRefCitation id="EFFA4B0FFFDDFFECFE88FBC9FD8CE65F" author="Hallowell, E." box="[340,569,1108,1133]" pageId="127" pageNumber="128" pagination="193 - 194" refId="ref79657" refString="Hallowell, E. (1854 a) Descriptions of new reptiles from Guinea. Proceedings of the Academy of Natural Sciences of Philadelphia, 1854, 193 - 194." type="journal article" year="1854">Hallowell, 1854a: 193</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54D0885CFFDDFFECFD9EFBC8FDC1E65F" box="[578,628,1109,1133]" pageId="127" pageNumber="128">Type</typeStatus>
locality: “
<location id="8EB46025FFDDFFECFD01FBC8FCA4E65F" LSID="urn:lsid:plazi:treatment:03C287E8FFDDFFECFF4BFC76FB10E4AF:8EB46025FFDDFFECFD01FBC8FCA4E65F" box="[733,785,1109,1133]" country="Gabon" municipality="river Gaboon" name="Near" pageId="127" pageNumber="128">Near</location>
the
<collectingMunicipality id="6BB0AC84FFDDFFECFCE0FBC9FC70E65E" box="[828,965,1108,1132]" pageId="127" pageNumber="128">river Gaboon</collectingMunicipality>
, Guinea”,
<collectingCountry id="F37C766EFFDDFFECFBECFBC9FBC1E65E" box="[1072,1140,1108,1132]" name="Gabon" pageId="127" pageNumber="128">Gabon</collectingCountry>
River,
<collectingCountry id="F37C766EFFDDFFECFB62FBC9FAB0E65E" box="[1214,1285,1108,1132]" name="Gabon" pageId="127" pageNumber="128">Gabon</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD436FEFFDDFFECFF4BFBEEFBF8E6B9" blockId="127.[151,1355,1108,1225]" box="[151,1101,1139,1164]" pageId="127" pageNumber="128">
<treatmentCitationGroup id="AB7B11D0FFDDFFECFF4BFBEEFBF8E6B9" box="[151,1101,1139,1164]" pageId="127" pageNumber="128">
<treatmentCitation id="0ACA10EFFFDDFFECFF4BFBEEFDF0E6B9" author="Gunther, A." box="[151,581,1139,1163]" page="20" pageId="127" pageNumber="128" year="1863">
<taxonomicName id="4C6B4D7DFFDDFFECFF4BFBEEFDF0E6B9" ID-CoL="C8284" authority="Gunther, 1863 b: 20" authorityName="Gunther" authorityPageNumber="20" authorityYear="1863" box="[151,581,1139,1163]" class="Reptilia" family="Viperidae" genus="Poecilostolus" kingdom="Animalia" order="Squamata" pageId="127" pageNumber="128" phylum="Chordata" rank="species" species="burtonii">
<emphasis id="B91FEAECFFDDFFECFF4BFBEEFECDE6B9" box="[151,376,1139,1163]" italics="true" pageId="127" pageNumber="128">Poecilostolus burtonii</emphasis>
<bibRefCitation id="EFFA4B0FFFDDFFECFEA2FBEEFDF0E6B9" author="Gunther, A." box="[382,581,1139,1163]" pageId="127" pageNumber="128" pagination="20 - 25" refId="ref78945" refString="Gunther, A. (1863 b) On new species of snakes in the collection of the British Museum. Annals &amp; Magazine of Natural History, Series 3, 11 (61), 20 - 25. https: // doi. org / 10.1080 / 00222936308681373" type="journal article" year="1863">Günther, 1863b: 20</bibRefCitation>
</taxonomicName>
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.
<typeStatus id="54D0885CFFDDFFECFD93FBE9FD34E6BE" box="[591,641,1140,1164]" pageId="127" pageNumber="128">Type</typeStatus>
locality: “
<location id="8EB46025FFDDFFECFD36FBEEFCD2E6B9" LSID="urn:lsid:plazi:treatment:03C287E8FFDDFFECFF4BFC76FB10E4AF:8EB46025FFDDFFECFD36FBEEFCD2E6B9" box="[746,871,1139,1163]" municipality="Camaroon" name="West Africa" pageId="127" pageNumber="128">West Africa</location>
<collectingMunicipality id="6BB0AC84FFDDFFECFCA7FBEEFC52E6B9" box="[891,999,1139,1163]" pageId="127" pageNumber="128">Camaroon</collectingMunicipality>
country”.
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</paragraph>
<paragraph id="8BD436FEFFDDFFECFF4BFB0FFB29E698" blockId="127.[151,1355,1108,1225]" box="[151,1180,1170,1194]" pageId="127" pageNumber="128">
<treatmentCitationGroup id="AB7B11D0FFDDFFECFF4BFB0FFB29E698" box="[151,1180,1170,1194]" pageId="127" pageNumber="128">
<treatmentCitation id="0ACA10EFFFDDFFECFF4BFB0FFDBAE698" author="Rochebrune, A. T." box="[151,527,1170,1194]" page="89" pageId="127" pageNumber="128" year="1885">
<taxonomicName id="4C6B4D7DFFDDFFECFF4BFB0FFDBAE698" ID-CoL="688YY" authority="Rochebrune, 1885: 89" authorityName="Rochebrune" authorityPageNumber="89" authorityYear="1885" box="[151,527,1170,1194]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="127" pageNumber="128" phylum="Chordata" rank="species" species="lucani">
<emphasis id="B91FEAECFFDDFFECFF4BFB0FFE93E698" box="[151,294,1170,1194]" italics="true" pageId="127" pageNumber="128">Atheris lucani</emphasis>
<bibRefCitation id="EFFA4B0FFFDDFFECFEF0FB0FFDBAE698" author="Rochebrune, A. T." box="[300,527,1170,1194]" pageId="127" pageNumber="128" pagination="86 - 99" refId="ref88071" refString="Rochebrune, A. T. (1885) Vertebratonum novorum vel minus cognitorum orae Africae occidentalis in colarum. Bulletin de la Societe Philomathique de Paris, 9, 86 - 99." type="journal article" year="1885">Rochebrune, 1885: 89</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54D0885CFFDDFFECFDC5FB0FFDFEE698" box="[537,587,1170,1194]" pageId="127" pageNumber="128">Type</typeStatus>
locality: “Locum
<emphasis id="B91FEAECFFDDFFECFCD8FB0FFCEBE698" box="[772,862,1170,1194]" italics="true" pageId="127" pageNumber="128">Landana</emphasis>
dictum” (untraceable locality).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD436FEFFDDFFECFF4BFB2DFAFEE6FB" blockId="127.[151,1355,1108,1225]" box="[151,1355,1200,1225]" pageId="127" pageNumber="128">
<taxonomicName id="4C6B4D7DFFDDFFECFF4BFB2DFDFCE6FA" authority="Laurent, 1956" authorityName="Laurent" authorityYear="1956" box="[151,585,1200,1224]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="127" pageNumber="128" phylum="Chordata" rank="subSpecies" species="squamigera" subSpecies="robusta">
<emphasis id="B91FEAECFFDDFFECFF4BFB2DFE04E6FA" box="[151,433,1200,1224]" italics="true" pageId="127" pageNumber="128">Atheris squamigera robusta</emphasis>
Laurent, 1956
</taxonomicName>
.
<typeStatus id="54D0885CFFDDFFECFD8EFB2CFD31E6FB" box="[594,644,1201,1225]" pageId="127" pageNumber="128">Type</typeStatus>
locality: “Nioka”,
<collectingCountry id="F37C766EFFDDFFECFC99FB2CFC7FE6FA" box="[837,970,1200,1225]" name="Democratic Republic of the Congo" pageId="127" pageNumber="128">Congo Belge</collectingCountry>
(
<collectingCountry id="F37C766EFFDDFFECFC0BFB2DFAF5E6FB" box="[983,1344,1200,1225]" name="Democratic Republic of the Congo" pageId="127" pageNumber="128">Democratic Republic of the Congo</collectingCountry>
).
</paragraph>
</subSubSection>
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<paragraph id="8BD436FEFFDDFFECFF4BFB6BFF6DE7F7" blockId="127.[151,1437,1270,1693]" pageId="127" pageNumber="128">
<emphasis id="B91FEAECFFDDFFECFF4BFB6BFE9CE722" bold="true" box="[151,297,1270,1296]" pageId="127" pageNumber="128">Distribution</emphasis>
. This species extends from
<collectingCountry id="F37C766EFFDDFFECFDBCFB6AFD01E723" box="[608,692,1271,1297]" name="Nigeria" pageId="127" pageNumber="128">Nigeria</collectingCountry>
to
<collectingCountry id="F37C766EFFDDFFECFD0BFB6AFC95E723" box="[727,800,1271,1297]" name="Kenya" pageId="127" pageNumber="128">Kenya</collectingCountry>
and southward to northern
<collectingCountry id="F37C766EFFDDFFECFB8EFB6AFB1DE723" box="[1106,1192,1271,1297]" name="Angola" pageId="127" pageNumber="128">Angola</collectingCountry>
. In
<collectingCountry id="F37C766EFFDDFFECFB0CFB6AFA29E723" box="[1232,1436,1271,1297]" name="Equatorial Guinea" pageId="127" pageNumber="128">Equatorial Guinea</collectingCountry>
it has been reported from Bioko (
<bibRefCitation id="EFFA4B0FFFDDFFECFDDDFA86FD0CE707" author="Broadley, D. G." box="[513,697,1307,1333]" pageId="127" pageNumber="128" pagination="117 - 135" refId="ref73029" refString="Broadley, D. G. (1998 b) A review of the genus Atheris Cope (Serpentes: Viperidae), with the description of a new species from Uganda. Herpetological Journal, 8 (3), 117 - 135." type="journal article" year="1998">Broadley 1998b</bibRefCitation>
;
<bibRefCitation id="EFFA4B0FFFDDFFECFD1FFA86FC26E707" author="Drewes, R. C. &amp; Vindum, J. V. &amp; Henwood, L." box="[707,915,1307,1333]" pageId="127" pageNumber="128" pagination="58 - 61" refId="ref75376" refString="Drewes, R. C., Vindum, J. V. &amp; Henwood, L. (1999) Preliminary report on a survey of the herpetofauna of Bioko; Equatorial Guinea. In: Larison, B., Smith, T. B., Girman, D., Stauffer, D., Mila, B., Drewes, R. C., Griswold, C. E., Vindum, J. V., Ubick, D., O'Keefe, K., Nguema, J. &amp; Henwood, L. (Eds.), Biotic surveys of Bioko and Rio Muni, Equatorial Guinea. Technical Report submitted to Biodiversity Support Program, 1999, pp. 58 - 61." type="journal article" year="1999">
Drewes
<emphasis id="B91FEAECFFDDFFECFCC2FA86FCE1E707" box="[798,852,1307,1333]" italics="true" pageId="127" pageNumber="128">et al.</emphasis>
1999
</bibRefCitation>
) at Moka (
<bibRefCitation id="EFFA4B0FFFDDFFECFBD4FA86FB0AE707" author="Boulenger, G. A." box="[1032,1215,1307,1333]" pageId="127" pageNumber="128" pagination="196 - 216" refId="ref72267" refString="Boulenger, G. A. (1906) Report on the reptiles collected by the late L. Fea in West Africa. Annali del Museo Civico di Storia Naturale di Genova, 3 (2), 196 - 216." type="journal article" year="1906">Boulenger 1906</bibRefCitation>
;
<bibRefCitation id="EFFA4B0FFFDDFFECFB15FA86FA22E707" author="Capocaccia, L." box="[1225,1431,1307,1333]" pageId="127" pageNumber="128" pagination="285 - 309" refId="ref73329" refString="Capocaccia, L. (1961 a) Contributo allo studio dei serpenti delle isole del Golfo di Guinea. Annali del Museo Civico di Storia Naturale Giacomo Doria, 72, 285 - 309." type="journal article" year="1961">Capocaccia 1961a</bibRefCitation>
;
<bibRefCitation id="EFFA4B0FFFDDFFECFF4BFAA2FEF4E76B" author="Mertens, R." box="[151,321,1343,1369]" pageId="127" pageNumber="128" pagination="211 - 238" refId="ref84765" refString="Mertens, R. (1964 a) Die Reptilien von Fernando Poo. Bonner zoologische Beitrage, 15, 211 - 238." type="journal article" year="1964">Mertens 1964a</bibRefCitation>
) and between Moka and Malabo (
<bibRefCitation id="EFFA4B0FFFDDFFECFD1BFAA2FCC0E76B" author="Mertens, R." box="[711,885,1343,1369]" pageId="127" pageNumber="128" pagination="211 - 238" refId="ref84765" refString="Mertens, R. (1964 a) Die Reptilien von Fernando Poo. Bonner zoologische Beitrage, 15, 211 - 238." type="journal article" year="1964">Mertens 1964a</bibRefCitation>
;
<bibRefCitation id="EFFA4B0FFFDDFFECFC5EFAA2FBDEE76B" author="Ceriaco, L. M. &amp; Marques, M. P. &amp; Bauer, A. M." box="[898,1131,1343,1369]" pageId="127" pageNumber="128" pagination="581 - 593" refId="ref73831" refString="Ceriaco, L. M., Marques, M. P. &amp; Bauer, A. M. (2020 a) The Bush Vipers, genus Atheris Cope, 1862 (Squamata: Viperidae) of Bioko Island, Gulf of Guinea, with the description of a new species. Zootaxa, 4838 (4), 581 - 593. https: // doi. org / 10.11646 / zootaxa. 4838.4.9" type="journal article" year="2020">
Ceríaco
<emphasis id="B91FEAECFFDDFFECFC3DFAA2FBAEE76B" box="[993,1051,1343,1369]" italics="true" pageId="127" pageNumber="128">et al.</emphasis>
2020a
</bibRefCitation>
), and in
<collectingCountry id="F37C766EFFDDFFECFB13FAA2FA88E76B" box="[1231,1341,1343,1369]" name="Equatorial Guinea" pageId="127" pageNumber="128">Río Muni</collectingCountry>
at Cabo San Juan (
<bibRefCitation id="EFFA4B0FFFDDFFECFED0FAFEFE71E74F" author="Boulenger, G. A." box="[268,452,1378,1405]" pageId="127" pageNumber="128" pagination="183 - 186" refId="ref72234" refString="Boulenger, G. A. (1905) Reptiles de la Guinee Espagnole. Memorias de la Sociedad Espanola de Historia Natural, 1 (8), 183 - 186." type="journal article" year="1905">Boulenger 1905</bibRefCitation>
) and Monte Alén National Park (
<bibRefCitation id="EFFA4B0FFFDDFFECFCE2FAFEFBB2E74F" author="Lasso, C. A. &amp; Rial, A. I. &amp; Castroviejo, J. &amp; De la Riva, I." box="[830,1031,1379,1405]" pageId="127" pageNumber="128" pagination="21 - 34" refId="ref81678" refString="Lasso, C. A., Rial, A. I., Castroviejo, J. &amp; De la Riva, I. (2002) Herpetofauna del Parque Nacional de Monte Alen (Rio Muni, Guinea Ecuatorial). Graellsia, 58, 21 - 34. https: // doi. org / 10.3989 / graellsia. 2002. v 58. i 2.276" type="journal article" year="2002">
Lasso
<emphasis id="B91FEAECFFDDFFECFC56FAFEFC76E74F" box="[906,963,1379,1405]" italics="true" pageId="127" pageNumber="128">et al.</emphasis>
2002
</bibRefCitation>
), including Monte Mitra (
<bibRefCitation id="EFFA4B0FFFDDFFECFAF0FAFEFEF1E793" author="Gonwouo, N. L. &amp; Nsang, A." pageId="127" pageNumber="128" pagination="32 - 49" refId="ref78060" refString="Gonwouo, N. L. &amp; Nsang, A. (2005) An assessment of the reptiles and amphibians of the Monte Mitra forest, Monte Alen National Park. In: Sunderland, T. C. H. (Ed.), A Biodiversity Assessment of the Monte Mitra Forest, Monte Alen National Park, Equatorial Guinea. Smithsonian Institution, Washington, D. C., pp. 32 - 49." type="book chapter" year="2005">Gonwouo &amp; Nsang 2005</bibRefCitation>
). There is a record from iNaturalist close to Moka, Bioko (
<geoCoordinate id="EE5F5039FFDDFFECFC32FA15FB2FE791" box="[1006,1178,1416,1443]" degrees="03" direction="north" minutes="19" orientation="latitude" pageId="127" pageNumber="128" precision="1" seconds="55.07" value="3.3319638">03°1955.07″N</geoCoordinate>
,
<geoCoordinate id="EE5F5039FFDDFFECFB74FA15FAFAE791" box="[1192,1359,1416,1443]" degrees="08" direction="east" minutes="38" orientation="longitude" pageId="127" pageNumber="128" precision="1" seconds="39.04" value="8.644177">08°3839.04″E</geoCoordinate>
) (
<figureCitation id="13502A7BFFDDFFECFABBFA1AFF79E7F7" captionStart="MAPS 38" captionStartId="131.[151,223,1943,1967]" captionTargetBox="[202,1385,189,1906]" captionTargetId="figure-18@131.[159,1429,181,1919]" captionTargetPageId="131" captionText="MAPS 38 AB. Distribution maps for Equatorial Guinean records of (A) Atheris squamigera; (B) Bitis gabonica; (C) Bitis nasicornis." figureDoi="http://doi.org/10.5281/zenodo.7305205" httpUri="https://zenodo.org/record/7305205/files/figure.png" pageId="127" pageNumber="128">Map 38A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFDDFFECFF1BFA53FB10E4AF" pageId="127" pageNumber="128" type="discussion">
<paragraph id="8BD436FEFFDDFFECFF1BFA53FB0DE403" blockId="127.[151,1437,1270,1693]" pageId="127" pageNumber="128">
<emphasis id="B91FEAECFFDDFFECFF1BFA53FEFAE7DA" bold="true" box="[199,335,1486,1512]" pageId="127" pageNumber="128">Comments.</emphasis>
Based on cytochrome b gene sequences,
<taxonomicName id="4C6B4D7DFFDDFFECFCF0FA52FBBCE7DA" baseAuthorityName="Hallowell" baseAuthorityYear="1854" box="[812,1033,1487,1513]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="127" pageNumber="128" phylum="Chordata" rank="species" species="squamigera">
<emphasis id="B91FEAECFFDDFFECFCF0FA52FBBCE7DA" box="[812,1033,1487,1513]" italics="true" pageId="127" pageNumber="128">Atheris squamigera</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C6B4D7DFFDDFFECFB25FA52FE9AE43F" authority="Fischer, 1888" authorityName="Fischer" authorityYear="1888" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="127" pageNumber="128" phylum="Chordata" rank="species" species="subocularis">
<emphasis id="B91FEAECFFDDFFECFB25FA52FA28E7DB" box="[1273,1437,1487,1513]" italics="true" pageId="127" pageNumber="128">A. subocularis</emphasis>
<bibRefCitation id="EFFA4B0FFFDDFFECFF4BFA6EFE9AE43F" author="Fischer, J. G." box="[151,303,1523,1549]" pageId="127" pageNumber="128" pagination="1 - 10" refId="ref77090" refString="Fischer, J. G. (1888) Uber zwei neue Schlangen und einen neuen Laubfrosch von Kamerun. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 5, 1 - 10." type="journal article" year="1888">Fischer, 1888</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFFA4B0FFFDDFFECFEE2FA6EFDAEE43F" author="Lawson, D. P. &amp; Noonan, B. P. &amp; Ustach, P. C." box="[318,539,1523,1549]" pageId="127" pageNumber="128" pagination="737 - 744" refId="ref82090" refString="Lawson, D. P., Noonan, B. P. &amp; Ustach, P. C. (2001) Atheris subocularis (Serpentes: Viperidae) revisited: Molecular and morphological evidence for the resurrection of an enigmatic taxon. Copeia, 2001 (3), 737 - 744. https: // doi. org / 10.1643 / 0045 - 8511 (2001) 001 [0737: ASSVRM] 2.0. CO; 2" type="journal article" year="2001">
Lawson
<emphasis id="B91FEAECFFDDFFECFE43FA6EFE6DE43F" box="[415,472,1523,1549]" italics="true" pageId="127" pageNumber="128">et al.</emphasis>
2001
</bibRefCitation>
), and the two species form a clade sister to
<taxonomicName id="4C6B4D7DFFDDFFECFBDFFA6EFA86E43F" authority="Lawson, 1999" authorityName="Lawson" authorityYear="1999" box="[1027,1331,1523,1549]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="127" pageNumber="128" phylum="Chordata" rank="species" species="broadleyi">
<emphasis id="B91FEAECFFDDFFECFBDFFA6EFB38E43F" box="[1027,1165,1523,1549]" italics="true" pageId="127" pageNumber="128">A. broadleyi</emphasis>
Lawson, 1999
</taxonomicName>
(
<bibRefCitation id="EFFA4B0FFFDDFFECFA9EFA6EFEA7E403" author="Lawson, D. P. &amp; Noonan, B. P. &amp; Ustach, P. C." pageId="127" pageNumber="128" pagination="737 - 744" refId="ref82090" refString="Lawson, D. P., Noonan, B. P. &amp; Ustach, P. C. (2001) Atheris subocularis (Serpentes: Viperidae) revisited: Molecular and morphological evidence for the resurrection of an enigmatic taxon. Copeia, 2001 (3), 737 - 744. https: // doi. org / 10.1643 / 0045 - 8511 (2001) 001 [0737: ASSVRM] 2.0. CO; 2" type="journal article" year="2001">
Lawson
<emphasis id="B91FEAECFFDDFFECFF4BF98AFF7AE403" box="[151,207,1559,1585]" italics="true" pageId="127" pageNumber="128">et al.</emphasis>
2001
</bibRefCitation>
). Phylogenetic relationships between these species and
<taxonomicName id="4C6B4D7DFFDDFFECFC5DF98AFC49E403" box="[897,1020,1559,1585]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="127" pageNumber="128" phylum="Chordata" rank="species" species="hetfieldi">
<emphasis id="B91FEAECFFDDFFECFC5DF98AFC49E403" box="[897,1020,1559,1585]" italics="true" pageId="127" pageNumber="128">A. hetfieldi</emphasis>
</taxonomicName>
remain unknow.
</paragraph>
<paragraph id="8BD436FEFFDDFFECFF1BF9A7FB10E4AF" blockId="127.[151,1437,1270,1693]" pageId="127" pageNumber="128">
<emphasis id="B91FEAECFFDDFFECFF1BF9A7FE70E466" bold="true" box="[199,453,1594,1620]" pageId="127" pageNumber="128">Specimens examined.</emphasis>
<materialsCitation id="3B033CA3FFDDFFECFE16F9A6FD81E44B" ID-GBIF-Occurrence="3962324375" collectingDate="2003-11-27" collectionCode="MNCN" latitude="3.357775" location="Caldera de Luba" longLatPrecision="1" longitude="8.514475" pageId="127" pageNumber="128" specimenCode="MNCN 50275" specimenCount="1">
<collectorName id="269E5328FFDDFFECFE16F9A6FDBEE467" box="[458,523,1595,1621]" pageId="127" pageNumber="128">Three</collectorName>
specimens.
<collectorName id="269E5328FFDDFFECFD4DF9A6FD6FE467" box="[657,730,1595,1621]" pageId="127" pageNumber="128">Bioko</collectorName>
: red swamp in
<location id="8EB46025FFDDFFECFCA7F9A6FB86E467" LSID="urn:lsid:plazi:treatment:03C287E8FFDDFFECFF4BFC76FB10E4AF:8EB46025FFDDFFECFCA7F9A6FB86E467" box="[891,1075,1595,1621]" latitude="3.357775" longLatPrecision="1" longitude="8.514475" name="Caldera de Luba" pageId="127" pageNumber="128">
Caldera
<collectorName id="269E5328FFDDFFECFC0BF9A6FB86E467" box="[983,1075,1595,1621]" pageId="127" pageNumber="128">de Luba</collectorName>
</location>
,
<geoCoordinate id="EE5F5039FFDDFFECFBE1F9A1FB52E465" box="[1085,1255,1596,1623]" degrees="03" direction="north" minutes="21" orientation="latitude" pageId="127" pageNumber="128" precision="1" seconds="27.99" value="3.357775">03°2127.99″N</geoCoordinate>
,
<geoCoordinate id="EE5F5039FFDDFFECFB2DF9A1FA2DE465" box="[1265,1432,1596,1623]" degrees="08" direction="east" minutes="30" orientation="longitude" pageId="127" pageNumber="128" precision="1" seconds="52.11" value="8.514475">08°3052.11″E</geoCoordinate>
,
<date id="FFD5103EFFDDFFECFF4BF9C2FEC6E44B" box="[151,371,1631,1657]" pageId="127" pageNumber="128" value="2003-11-27">
<collectingDate id="EF91E9D6FFDDFFECFF4BF9C2FEC6E44B" box="[151,371,1631,1657]" pageId="127" pageNumber="128" value="2003-11-27">27 November 2003</collectingDate>
</date>
(
<specimenCode id="DBCD9E85FFDDFFECFE5FF9C2FD99E44B" box="[387,556,1630,1657]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="127" pageNumber="128" type="Museum">MNCN 50275</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B033CA3FFDDFFECFD9CF9C2FCA0E4AF" ID-GBIF-Occurrence="3962324689" collectionCode="MNCN" country="Equatorial Guinea" location="Rio Muni" pageId="127" pageNumber="128" specimenCode="MNCN 22788" specimenCount="1">
<collectingCountry id="F37C766EFFDDFFECFD9CF9C2FD06E44B" box="[576,691,1631,1657]" name="Equatorial Guinea" pageId="127" pageNumber="128">Río Muni</collectingCountry>
: without specific locality, but likely Cabo San Juan as part of the material examined by
<bibRefCitation id="EFFA4B0FFFDDFFECFE4DF91EFDE2E4AF" author="Boulenger, G. A." box="[401,599,1666,1693]" pageId="127" pageNumber="128" pagination="183 - 186" refId="ref72234" refString="Boulenger, G. A. (1905) Reptiles de la Guinee Espagnole. Memorias de la Sociedad Espanola de Historia Natural, 1 (8), 183 - 186." type="journal article" year="1905">Boulenger (1905)</bibRefCitation>
(
<specimenCode id="DBCD9E85FFDDFFECFDB9F91EFCB8E4AF" box="[613,781,1667,1693]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="127" pageNumber="128" type="Museum">MNCN 22788</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFDDFFECFCC3F91EFB14E4AF" ID-GBIF-Occurrence="3962324686" box="[799,1185,1666,1693]" collectingDate="1964-07" collectionCode="EBD" collectorName="Ayamiken" country="Equatorial Guinea" pageId="127" pageNumber="128" specimenCode="EBD 2875" specimenCount="1">
<collectorName id="269E5328FFDDFFECFCC3F91EFC21E4AF" box="[799,916,1667,1693]" pageId="127" pageNumber="128">Ayamiken</collectorName>
,
<date id="FFD5103EFFDDFFECFC43F91EFBB8E4AF" box="[927,1037,1667,1693]" pageId="127" pageNumber="128" value="1964-07">
<collectingDate id="EF91E9D6FFDDFFECFC43F91EFBB8E4AF" box="[927,1037,1667,1693]" pageId="127" pageNumber="128" value="1964-07">July 1964</collectingDate>
</date>
(
<specimenCode id="DBCD9E85FFDDFFECFBC7F91EFB2CE4AF" box="[1051,1177,1666,1693]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="127" pageNumber="128">EBD 2875</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>