Synopsis of the terrestrial Reptiles of Equatorial Guinea Author Sánchez-Vialas, Alberto Author Calvo-Revuelta, Marta 0000-0002-8628-1055 Author Riva, Ignacio De La 0000-0001-5064-4507 albertosv@mncn.csic.es text Zootaxa 2022 2022-11-03 5202 1 1 197 http://dx.doi.org/10.11646/zootaxa.5202.1.1 journal article 182444 10.11646/zootaxa.5202.1.1 45ba25c1-47a9-4001-93d9-294815bf37e7 1175-5326 7285600 BF4831D2-D98B-4265-9138-03DB8607B826 Atheris squamigera (Hallowell, 1854) Figure 32B Echis squamigera Hallowell, 1854a: 193 . Type locality: “ Near the river Gaboon , Guinea”, Gabon River, Gabon . Poecilostolus burtonii Günther, 1863b: 20 . Type locality: “ West Africa … Camaroon country”. Atheris lucani Rochebrune, 1885: 89 . Type locality: “Locum Landana dictum” (untraceable locality). Atheris squamigera robusta Laurent, 1956 . Type locality: “Nioka”, Congo Belge ( Democratic Republic of the Congo ). Distribution . This species extends from Nigeria to Kenya and southward to northern Angola . In Equatorial Guinea it has been reported from Bioko ( Broadley 1998b ; Drewes et al. 1999 ) at Moka ( Boulenger 1906 ; Capocaccia 1961a ; Mertens 1964a ) and between Moka and Malabo ( Mertens 1964a ; Ceríaco et al. 2020a ), and in Río Muni at Cabo San Juan ( Boulenger 1905 ) and Monte Alén National Park ( Lasso et al. 2002 ), including Monte Mitra ( Gonwouo & Nsang 2005 ). There is a record from iNaturalist close to Moka, Bioko ( 03°19′55.07″N , 08°38′39.04″E ) ( Map 38A ). Comments. Based on cytochrome b gene sequences, Atheris squamigera is closely related to A. subocularis Fischer, 1888 ( Lawson et al. 2001 ), and the two species form a clade sister to A. broadleyi Lawson, 1999 ( Lawson et al. 2001 ). Phylogenetic relationships between these species and A. hetfieldi remain unknow. Specimens examined. Three specimens. Bioko : red swamp in Caldera de Luba , 03°21′27.99″N , 08°30′52.11″E , 27 November 2003 ( MNCN 50275 ) . Río Muni : without specific locality, but likely Cabo San Juan as part of the material examined by Boulenger (1905) ( MNCN 22788 ) ; Ayamiken , July 1964 ( EBD 2875 ) .