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<document id="EC3E7D35B7E6F2341FB35509D42EC2A2" ID-DOI="10.37520/aemnp.2021.030" ID-ISSN="1804-6487" ID-Zenodo-Dep="5821367" ID-ZooBank="06FDFB43-0B61-4DA8-B260-D78ABD62756C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641391650078" checkinUser="plazi" docAuthor="Bezděk, Jan" docDate="2021" docId="492287F9DB65FF8DFEF3AA4CFB42F984" docLanguage="en" docName="ActaEntMusNatPra.61.2.529-614.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 61 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Liroetis medvedevi Bezděk, 2021, nom. nov." docType="treatment" docVersion="4" lastPageNumber="555" masterDocId="B51BFF81DB7FFF97FFD5AC4FFF9BFFF3" masterDocTitle="Redefinition of Liroetis, with descriptions of two new species and an annotated list of species (Coleoptera: Chrysomelidae: Galerucinae)" masterLastPageNumber="614" masterPageNumber="529" pageNumber="555" updateTime="1699285417030" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="02885013C60AAFC5B277D241834E0649">Redefinition of Liroetis, with descriptions of two new species and an annotated list of species (Coleoptera: Chrysomelidae: Galerucinae)</mods:title>
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<mods:namePart id="34FBA9C14714F7ECBC945FF26DD5E294">Bezděk, Jan</mods:namePart>
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<mods:title id="2A95F47AE3BA128A335A6AEA73642042">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="5AB61910ADC75C570E535F1248AF5A8D">2021</mods:date>
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<mods:number id="5C229C6D08B8B752A8EC1B7807C54C85">2021-12-31</mods:number>
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<treatment id="492287F9DB65FF8DFEF3AA4CFB42F984" ID-DOI="http://doi.org/10.5281/zenodo.5821682" ID-Zenodo-Dep="5821682" LSID="urn:lsid:plazi:treatment:492287F9DB65FF8DFEF3AA4CFB42F984" httpUri="http://treatment.plazi.org/id/492287F9DB65FF8DFEF3AA4CFB42F984" lastPageNumber="555" pageId="26" pageNumber="555">
<subSubSection id="89916564DB65FF8DFEF3AA4CFDF2F9E8" box="[294,617,1539,1563]" pageId="26" pageNumber="555" type="nomenclature">
<paragraph id="C13436EFDB65FF8DFEF3AA4CFDF2F9E8" blockId="26.[294,617,1539,1563]" box="[294,617,1539,1563]" pageId="26" pageNumber="555">
<heading id="9A7C8183DB65FF8DFEF3AA4CFDF2F9E8" box="[294,617,1539,1563]" centered="true" fontSize="10" level="2" pageId="26" pageNumber="555" reason="3">
<emphasis id="F3FFEAFDDB65FF8DFEF3AA4CFDF2F9E8" bold="true" box="[294,617,1539,1563]" pageId="26" pageNumber="555">
<taxonomicName id="068B4D6CDB65FF8DFEF3AA4CFE68F9E8" authorityName="Bezděk" authorityYear="2021" box="[294,499,1539,1563]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="medvedevi" status="nom. nov.">
<emphasis id="F3FFEAFDDB65FF8DFEF3AA4CFE68F9E8" bold="true" box="[294,499,1539,1563]" italics="true" pageId="26" pageNumber="555">Liroetis medvedevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8CC5786DB65FF8DFE2FAA4BFDF2F9E8" box="[506,617,1540,1563]" pageId="26" pageNumber="555">nom. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89916564DB65FF8DFE99AA6DFDD8F9C6" box="[332,579,1570,1589]" pageId="26" pageNumber="555" type="description">
<paragraph id="C13436EFDB65FF8DFE99AA6DFDD8F9C6" blockId="26.[151,761,1570,1898]" box="[332,579,1570,1589]" pageId="26" pageNumber="555">
(
<figureCitation id="59B02A6ADB65FF8DFE87AA6DFE5BF9C6" box="[338,448,1570,1589]" captionStart="Figs 95100" captionStartId="27.[151,188,1346,1365]" captionTargetBox="[151,1436,189,1313]" captionTargetId="figure-11@27.[151,1436,189,1988]" captionTargetPageId="27" captionText="Figs 95100. Liroetis medvedevi nom. nov. (type specimens of L. nigricollis Medvedev, 2009). 9597 Male, holotype (7.0 mm). 95 dorsal view; 96 labels; 97 head and pronotum. 98100 Female, paratype (9.3 mm). 98 dorsal view; 99 ventral view; 100 labels." figureDoi="http://doi.org/10.5281/zenodo.5821403" httpUri="https://zenodo.org/record/5821403/files/figure.png" pageId="26" pageNumber="555">Figs 95100</figureCitation>
,
<figureCitation id="59B02A6ADB65FF8DFE1DAA6DFE73F9C6" box="[456,488,1570,1589]" captionStart="Figs 108117" captionStartId="29.[151,188,1826,1845]" captionTargetBox="[157,1433,197,1783]" captionTargetId="figure-11@29.[151,1436,189,1988]" captionTargetPageId="29" captionText="Figs 108117.Aedeagus of Liroetis, dorsal and lateral views. 108 L. flavipennis Bryant, 1954; 109 L. humeralis Jiang, 1988; 110 L. lonicernis Jiang, 1988; 111 L. medvedevi nom. nov.; 112 L. nepalensis Chûjô, 1966; 113 L. prominensis Jiang, 1988; 114 L. sichuanensis Jiang, 1988; 115 L. tibetanus Jiang, 1988; 116 L. tibialis Jiang, 1988; 117 L. zhongdianicus Jiang, 1988. Figs 116117 reproduced from JൺඇǤ (1988). Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5821409" httpUri="https://zenodo.org/record/5821409/files/figure.png" pageId="26" pageNumber="555">111</figureCitation>
,
<figureCitation id="59B02A6ADB65FF8DFE24AA6DFDA6F9C6" box="[497,573,1570,1589]" captionStart="Figs 118131" captionStartId="30.[151,188,1809,1828]" captionTargetBox="[162,1428,212,1758]" captionTargetId="figure-13@30.[151,1436,189,1988]" captionTargetPageId="30" captionText="Figs 118131. Spermatheca (Figs 118124) and sternite VII (Figs 125131) of Liroetis. 118, 125 L. flavipennis Bryant, 1954; 119, 126 L. humeralis Jiang, 1988; 120, 127 L. medvedevi nom. nov.; 121, 128 L. nepalensis Chûjô, 1966; 122, 129 L. sichuanensis Jiang, 1988; 123, 130 L. tibetanus Jiang, 1988; 124, 131 L. zhongdianicus Jiang, 1988. Scales 0.25 mm for Figs 118124, 0.5 mm for Figs 125131." figureDoi="http://doi.org/10.5281/zenodo.5821413" httpUri="https://zenodo.org/record/5821413/files/figure.png" pageId="26" pageNumber="555">120, 127</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="89916564DB65FF8DFF42AA0BFD52F97E" pageId="26" pageNumber="555" type="reference_group">
<paragraph id="C13436EFDB65FF8DFF42AA0BFDE1F981" blockId="26.[151,761,1570,1898]" pageId="26" pageNumber="555">
<treatmentCitationGroup id="E19B11C1DB65FF8DFF42AA0BFDE1F981" pageId="26" pageNumber="555">
<treatmentCitation id="402A10FEDB65FF8DFF42AA0BFE6DF9A4" author="MEDVEDEV L. N." box="[151,502,1604,1623]" page="407" pageId="26" pageNumber="555" year="2009">
<taxonomicName id="068B4D6CDB65FF8DFF42AA0BFE6DF9A4" authority="Medvedev, 2009: 407" authorityName="Medvedev" authorityPageNumber="407" authorityYear="2009" box="[151,502,1604,1623]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="nigricollis">
<emphasis id="F3FFEAFDDB65FF8DFF42AA0BFEADF9A4" box="[151,310,1604,1623]" italics="true" pageId="26" pageNumber="555">Liroetis nigricollis</emphasis>
<bibRefCitation id="A51A4B1EDB65FF8DFEE9AA0BFE6DF9A4" author="MEDVEDEV L. N." box="[316,502,1604,1623]" pageId="26" pageNumber="555" pagination="407 - 410" refId="ref54748" refString="MEDVEDEV L. N. 2009: New species of leaf beetles from the collection of the Naturkundemuseum Erfurt (Insecta: Coleoptera: Chrysomelidae). Pp. 407 - 410. In: HARTMANN M. &amp; WEIPERT J. (eds): Biodiversitat und Naturausstattung im Himalaya III. Verein der Freun- de &amp; Forderer des Naturkundemuseums Erfurt e. V., Erfurt, 477 pp." type="book chapter" year="2009">Medvedev, 2009: 407</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<emphasis id="F3FFEAFDDB65FF8DFD69AA0BFEE9F981" bold="true" pageId="26" pageNumber="555">Junior secondary homonym</emphasis>
of
<taxonomicName id="068B4D6CDB65FF8DFE5AAA10FDEEF981" authority="(Jiang, 1990)" authorityName="Bezděk" authorityYear="2021" baseAuthorityName="Jiang" baseAuthorityYear="1990" box="[399,629,1631,1650]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="nigricollis">
<emphasis id="F3FFEAFDDB65FF8DFE5AAA10FE66F981" box="[399,509,1631,1650]" italics="true" pageId="26" pageNumber="555">L. nigricollis</emphasis>
(
<bibRefCitation id="A51A4B1EDB65FF8DFDDCAA10FDF4F981" author="JIANG S. - Q." box="[521,623,1631,1650]" pageId="26" pageNumber="555" pagination="141 - 144" refId="ref53495" refString="JIANG S. - Q. 1990: On the Chinese species of the Galerucinae genus Zangia (Coleoptera: Chrysomelidae). Sinozoologia 7: 141 - 144." type="journal article" year="1990">Jiang, 1990</bibRefCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C13436EFDB65FF8DFF42AA35FD52F97E" blockId="26.[151,761,1570,1898]" box="[151,713,1658,1677]" pageId="26" pageNumber="555">
<taxonomicName id="068B4D6CDB65FF8DFF42AA35FEADF97E" authorityName="Bezděk" authorityYear="2021" baseAuthorityName="Jiang" baseAuthorityYear="1990" box="[151,310,1658,1677]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="nigricollis">
<emphasis id="F3FFEAFDDB65FF8DFF42AA35FEADF97E" box="[151,310,1658,1677]" italics="true" pageId="26" pageNumber="555">Liroetis nigricollis</emphasis>
</taxonomicName>
: SඉඋൾർIJൾඋ-Uൾൻൾඋඌൺඑ (2011): 433 (catalogue).
</paragraph>
</subSubSection>
<subSubSection id="89916564DB65FF8DFF42AAD3FEE4F899" pageId="26" pageNumber="555" type="materials_examined">
<paragraph id="C13436EFDB65FF8DFF42AAD3FDECF95C" blockId="26.[151,761,1570,1898]" box="[151,631,1692,1711]" pageId="26" pageNumber="555">
<emphasis id="F3FFEAFDDB65FF8DFF42AAD3FE8BF95C" bold="true" box="[151,272,1692,1711]" pageId="26" pageNumber="555">
<typeStatus id="1E30884DDB65FF8DFF42AAD3FF5FF95C" box="[151,196,1692,1711]" pageId="26" pageNumber="555">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="B99C767FDB65FF8DFECDAAD3FED2F95C" box="[280,329,1692,1711]" name="India" pageId="26" pageNumber="555">India</collectingCountry>
,
<collectingRegion id="034FF80DDB65FF8DFE87AAD3FE5AF95C" box="[338,449,1692,1711]" country="India" name="West Bengal" pageId="26" pageNumber="555">West Bengal</collectingRegion>
, Darjeeling, Tonglu.
</paragraph>
<paragraph id="C13436EFDB65FF8DFF42AAF8FEE4F899" blockId="26.[151,761,1570,1898]" pageId="26" pageNumber="555">
<emphasis id="F3FFEAFDDB65FF8DFF42AAF8FEE6F939" bold="true" box="[151,381,1719,1738]" pageId="26" pageNumber="555">Type material examined.</emphasis>
Hඈඅඈඍඒඉൾ: J (
<figureCitation id="59B02A6ADB65FF8DFDDDAAF8FDF6F939" box="[520,621,1719,1738]" captionStart="Figs 95100" captionStartId="27.[151,188,1346,1365]" captionTargetBox="[151,1436,189,1313]" captionTargetId="figure-11@27.[151,1436,189,1988]" captionTargetPageId="27" captionText="Figs 95100. Liroetis medvedevi nom. nov. (type specimens of L. nigricollis Medvedev, 2009). 9597 Male, holotype (7.0 mm). 95 dorsal view; 96 labels; 97 head and pronotum. 98100 Female, paratype (9.3 mm). 98 dorsal view; 99 ventral view; 100 labels." figureDoi="http://doi.org/10.5281/zenodo.5821403" httpUri="https://zenodo.org/record/5821403/files/figure.png" pageId="26" pageNumber="555">Figs 9597</figureCitation>
),
<collectingCountry id="B99C767FDB65FF8DFD54AAF8FD58F939" box="[641,707,1719,1738]" name="India" pageId="26" pageNumber="555">INDIA</collectingCountry>
,
<collectingRegion id="034FF80DDB65FF8DFD1BAAF8FF48F917" country="India" name="West Bengal" pageId="26" pageNumber="555">West Bengal</collectingRegion>
/ Darjeeling, Tonglu /
<date id="B535102FDB65FF8DFE4AAA9EFD83F917" box="[415,536,1745,1764]" pageId="26" pageNumber="555" value="1998-05-19" valueMax="1998-05-23" valueMin="1998-05-19">19.23.v.1998</date>
/
<quantity id="06739B0ADB65FF8DFDF8AA9EFD05F917" box="[557,670,1745,1764]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="26" pageNumber="555" unit="m" value="2800.0" valueMax="3000.0" valueMin="2600.0">26003000m</quantity>
NN / leg. Fabrizi &amp;Ahrens [w, p] //
<typeStatus id="1E30884DDB65FF8DFEBAAAA3FE5CF90C" box="[367,455,1772,1791]" pageId="26" pageNumber="555" type="holotype">Holotypus</typeStatus>
/
<taxonomicName id="068B4D6CDB65FF8DFE00AAA3FD8CF90C" authorityName="Weise" authorityYear="1889" box="[469,535,1772,1791]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="genus">Liroetis</taxonomicName>
/ nigricollis / L. Medvedev [r, p] (
<collectionCode id="A79AAE2ADB65FF8DFF07AB48FE8DF8E9" box="[210,278,1799,1818]" country="Germany" httpUri="http://grbio.org/cool/3082-3vhh" name="Naturkundesmuseum" pageId="26" pageNumber="555">NMEG</collectionCode>
). Pൺඋൺඍඒඉൾඌ:
<specimenCount id="D78DFD66DB65FF8DFE52AB48FE2DF8EA" box="[391,438,1792,1818]" pageId="26" pageNumber="555" type="female">3 ♀♀</specimenCount>
(
<figureCitation id="59B02A6ADB65FF8DFE14AB48FDABF8E9" box="[449,560,1799,1818]" captionStart="Figs 95100" captionStartId="27.[151,188,1346,1365]" captionTargetBox="[151,1436,189,1313]" captionTargetId="figure-11@27.[151,1436,189,1988]" captionTargetPageId="27" captionText="Figs 95100. Liroetis medvedevi nom. nov. (type specimens of L. nigricollis Medvedev, 2009). 9597 Male, holotype (7.0 mm). 95 dorsal view; 96 labels; 97 head and pronotum. 98100 Female, paratype (9.3 mm). 98 dorsal view; 99 ventral view; 100 labels." figureDoi="http://doi.org/10.5281/zenodo.5821403" httpUri="https://zenodo.org/record/5821403/files/figure.png" pageId="26" pageNumber="555">Figs 98100</figureCitation>
),
<collectingCountry id="B99C767FDB65FF8DFD97AB48FD1FF8E9" box="[578,644,1799,1818]" name="India" pageId="26" pageNumber="555">INDIA</collectingCountry>
,
<collectingRegion id="034FF80DDB65FF8DFD59AB48FD6CF8E9" box="[652,759,1799,1818]" country="India" name="West Bengal" pageId="26" pageNumber="555">West Bengal</collectingRegion>
/ Darjeeling, Tonglu /
<date id="B535102FDB65FF8DFE8EAB6EFE4FF8C7" box="[347,468,1825,1844]" pageId="26" pageNumber="555" value="1998-05-19" valueMax="1998-05-23" valueMin="1998-05-19">19.23.v.1998</date>
/
<quantity id="06739B0ADB65FF8DFE3CAB6EFDC1F8C7" box="[489,602,1825,1844]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="26" pageNumber="555" unit="m" value="2800.0" valueMax="3000.0" valueMin="2600.0">26003000m</quantity>
NN / leg. Fabrizi &amp; Ahrens [w, p] //
<typeStatus id="1E30884DDB65FF8DFEE8AB73FE2EF8BC" box="[317,437,1852,1871]" pageId="26" pageNumber="555" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="068B4D6CDB65FF8DFE1DAB73FD91F8BC" authorityName="Weise" authorityYear="1889" box="[456,522,1852,1871]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="genus">Liroetis</taxonomicName>
/ nigricollis
<taxonomicNameLabel id="E8CC5786DB65FF8DFDAEAB73FD30F8BC" box="[635,683,1852,1871]" pageId="26" pageNumber="555" rank="species">n. sp.</taxonomicNameLabel>
/ des. L. Medvedev [r, p] (
<collectionCode id="A79AAE2ADB65FF8DFEE4AB18FEEEF899" box="[305,373,1879,1898]" country="Germany" httpUri="http://grbio.org/cool/3082-3vhh" name="Naturkundesmuseum" pageId="26" pageNumber="555">NMEG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="89916564DB65FF8DFF42ABCDFA06FA44" pageId="26" pageNumber="555" type="diagnosis">
<paragraph id="C13436EFDB65FF8DFF42ABCDFEB8F80A" blockId="26.[151,760,1922,2073]" pageId="26" pageNumber="555">
<emphasis id="F3FFEAFDDB65FF8DFF42ABCDFE92F86A" bold="true" box="[151,265,1922,1945]" pageId="26" pageNumber="555">Diagnosis.</emphasis>
<emphasis id="F3FFEAFDDB65FF8DFED8ABCDFE17F86A" box="[269,396,1922,1945]" italics="true" pageId="26" pageNumber="555">Colouration</emphasis>
. Head, scutellum and elytra brown, pronotum brownish black with paler basal part, antennae, legs and underside black, posterior part of last abdominal ventrite pale.
</paragraph>
<paragraph id="C13436EFDB65FF8DFF60A44DFC58FF04" blockId="26.[151,760,1922,2073]" lastBlockId="26.[828,1437,189,1655]" pageId="26" pageNumber="555">
<emphasis id="F3FFEAFDDB65FF8DFF60A44DFEA1F7EA" box="[181,314,2050,2073]" italics="true" pageId="26" pageNumber="555">Body length</emphasis>
. J (
<typeStatus id="1E30884DDB65FF8DFEB8A44DFE49F7EA" box="[365,466,2050,2073]" pageId="26" pageNumber="555" type="holotype">holotype</typeStatus>
): 7.0 mm,
<specimenCount id="D78DFD66DB65FF8DFD9BABB0FDF6F7EB" box="[590,621,2047,2072]" pageId="26" pageNumber="555" type="female">♀♀</specimenCount>
(
<typeStatus id="1E30884DDB65FF8DFDABA44DFD77F7EA" box="[638,748,2050,2073]" pageId="26" pageNumber="555" type="paratype">paratypes</typeStatus>
):
<quantity id="06739B0ADB65FF8DFCE9AC8FFC58FF24" box="[828,963,192,215]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="9.8" metricValueMin="8.7" pageId="26" pageNumber="555" unit="mm" value="9.25" valueMax="9.8" valueMin="8.7">8.79.8 mm</quantity>
(J
<specimenCount id="D78DFD66DB65FF8DFC36ACF2FC6CFF25" box="[995,1015,189,214]" pageId="26" pageNumber="555" type="female"></specimenCount>
: 7.210.0 mm based on the original description).
</paragraph>
<paragraph id="C13436EFDB65FF8DFC8FAD4FFA08FE04" blockId="26.[828,1437,189,1655]" pageId="26" pageNumber="555">
<emphasis id="F3FFEAFDDB65FF8DFC8FAD4FFC0BFEE4" box="[858,912,256,279]" italics="true" pageId="26" pageNumber="555">Male</emphasis>
(
<figureCitation id="59B02A6ADB65FF8DFC49AD4FFB88FEE4" box="[924,1043,256,279]" captionStart="Figs 95100" captionStartId="27.[151,188,1346,1365]" captionTargetBox="[151,1436,189,1313]" captionTargetId="figure-11@27.[151,1436,189,1988]" captionTargetPageId="27" captionText="Figs 95100. Liroetis medvedevi nom. nov. (type specimens of L. nigricollis Medvedev, 2009). 9597 Male, holotype (7.0 mm). 95 dorsal view; 96 labels; 97 head and pronotum. 98100 Female, paratype (9.3 mm). 98 dorsal view; 99 ventral view; 100 labels." figureDoi="http://doi.org/10.5281/zenodo.5821403" httpUri="https://zenodo.org/record/5821403/files/figure.png" pageId="26" pageNumber="555">Figs 95, 97</figureCitation>
).Antennae 0.96 times as long as body. Pronotum convex, 1.53 times as wide as long, lustrous, almost impunctate, anterior margin straight, bordered, anterior angles not projecting. Middle part of posterior margin of abdominal ventrite IV obliquely impressed, with two small subtriagular processes separated by shallow semicircular incision. Last abdominal ventrite with longitudinal impression narrowed subbasally. Metatibial spur absent.
</paragraph>
<paragraph id="C13436EFDB65FF8DFC8FAE4FFA11FC84" blockId="26.[828,1437,189,1655]" pageId="26" pageNumber="555">
<emphasis id="F3FFEAFDDB65FF8DFC8FAE4FFC58FDE4" box="[858,963,512,535]" italics="true" pageId="26" pageNumber="555">Aedeagus</emphasis>
(
<figureCitation id="59B02A6ADB65FF8DFC00AE4FFBAEFDE4" box="[981,1077,512,535]" captionStart="Figs 108117" captionStartId="29.[151,188,1826,1845]" captionTargetBox="[157,1433,197,1783]" captionTargetId="figure-11@29.[151,1436,189,1988]" captionTargetPageId="29" captionText="Figs 108117.Aedeagus of Liroetis, dorsal and lateral views. 108 L. flavipennis Bryant, 1954; 109 L. humeralis Jiang, 1988; 110 L. lonicernis Jiang, 1988; 111 L. medvedevi nom. nov.; 112 L. nepalensis Chûjô, 1966; 113 L. prominensis Jiang, 1988; 114 L. sichuanensis Jiang, 1988; 115 L. tibetanus Jiang, 1988; 116 L. tibialis Jiang, 1988; 117 L. zhongdianicus Jiang, 1988. Figs 116117 reproduced from JൺඇǤ (1988). Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5821409" httpUri="https://zenodo.org/record/5821409/files/figure.png" pageId="26" pageNumber="555">Fig. 111</figureCitation>
). Median lobe of aedeagus 3.16 times as long as wide; basal quarter wide, subparallel, middle part of median lobe gradually convergent anteriorly, distinctly constricted in anterior quarter, apical part narrow, subparallel with distinct narrow median furrow. Lateral view: median lobe widely rounded; lateral elevation rounded, placed in anterior 2/5 of aedeagus length. Dorsal process 5.00 times as long as wide, 0.65 times as long as median lobe; narrow, widest in middle part, apex narrow and sharp. Lateral view: dorsal process widely regularly rounded, apical part with large U-shaped incision, dorsal branch wide and short, ventral branch long and narrow.
</paragraph>
<paragraph id="C13436EFDB65FF8DFC8FAFCFFB28FB84" blockId="26.[828,1437,189,1655]" pageId="26" pageNumber="555">
<emphasis id="F3FFEAFDDB65FF8DFC8FAFCFFC31FC64" box="[858,938,896,919]" italics="true" pageId="26" pageNumber="555">Female</emphasis>
(
<figureCitation id="59B02A6ADB65FF8DFC62AFCFFBA8FC64" box="[951,1075,896,919]" captionStart="Figs 95100" captionStartId="27.[151,188,1346,1365]" captionTargetBox="[151,1436,189,1313]" captionTargetId="figure-11@27.[151,1436,189,1988]" captionTargetPageId="27" captionText="Figs 95100. Liroetis medvedevi nom. nov. (type specimens of L. nigricollis Medvedev, 2009). 9597 Male, holotype (7.0 mm). 95 dorsal view; 96 labels; 97 head and pronotum. 98100 Female, paratype (9.3 mm). 98 dorsal view; 99 ventral view; 100 labels." figureDoi="http://doi.org/10.5281/zenodo.5821403" httpUri="https://zenodo.org/record/5821403/files/figure.png" pageId="26" pageNumber="555">Figs 9899</figureCitation>
). Metatibial spur absent. Posterior margin of last abdominal ventrite entire. Sternite VIII umbrella-like with shallow emargination in middle of posterior margin, surface without setation; tignum 0.25 times as long as sternite VIII (
<figureCitation id="59B02A6ADB65FF8DFBA7A84FFB56FBE4" box="[1138,1229,1024,1047]" captionStart="Figs 118131" captionStartId="30.[151,188,1809,1828]" captionTargetBox="[162,1428,212,1758]" captionTargetId="figure-13@30.[151,1436,189,1988]" captionTargetPageId="30" captionText="Figs 118131. Spermatheca (Figs 118124) and sternite VII (Figs 125131) of Liroetis. 118, 125 L. flavipennis Bryant, 1954; 119, 126 L. humeralis Jiang, 1988; 120, 127 L. medvedevi nom. nov.; 121, 128 L. nepalensis Chûjô, 1966; 122, 129 L. sichuanensis Jiang, 1988; 123, 130 L. tibetanus Jiang, 1988; 124, 131 L. zhongdianicus Jiang, 1988. Scales 0.25 mm for Figs 118124, 0.5 mm for Figs 125131." figureDoi="http://doi.org/10.5281/zenodo.5821413" httpUri="https://zenodo.org/record/5821413/files/figure.png" pageId="26" pageNumber="555">Fig. 127</figureCitation>
). Spermatheca with well developed oval nodulus, cornu C-shaped, narrow, gradually narrowing towards sharp apex, spermathecal duct moderately sinuate (
<figureCitation id="59B02A6ADB65FF8DFB9CA82FFB3CFB84" box="[1097,1191,1120,1143]" captionStart="Figs 118131" captionStartId="30.[151,188,1809,1828]" captionTargetBox="[162,1428,212,1758]" captionTargetId="figure-13@30.[151,1436,189,1988]" captionTargetPageId="30" captionText="Figs 118131. Spermatheca (Figs 118124) and sternite VII (Figs 125131) of Liroetis. 118, 125 L. flavipennis Bryant, 1954; 119, 126 L. humeralis Jiang, 1988; 120, 127 L. medvedevi nom. nov.; 121, 128 L. nepalensis Chûjô, 1966; 122, 129 L. sichuanensis Jiang, 1988; 123, 130 L. tibetanus Jiang, 1988; 124, 131 L. zhongdianicus Jiang, 1988. Scales 0.25 mm for Figs 118124, 0.5 mm for Figs 125131." figureDoi="http://doi.org/10.5281/zenodo.5821413" httpUri="https://zenodo.org/record/5821413/files/figure.png" pageId="26" pageNumber="555">Fig. 120</figureCitation>
).
</paragraph>
<paragraph id="C13436EFDB65FF8DFCE9A8CFFA06FA44" blockId="26.[828,1437,189,1655]" pageId="26" pageNumber="555">
<emphasis id="F3FFEAFDDB65FF8DFCE9A8CFFBA8FB64" bold="true" box="[828,1075,1152,1175]" pageId="26" pageNumber="555">Differential diagnosis.</emphasis>
<taxonomicName id="068B4D6CDB65FF8DFBECA8CFFA99FB64" authorityName="Bezděk" authorityYear="2021" box="[1081,1282,1152,1175]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="medvedevi" status="nom. nov.">
<emphasis id="F3FFEAFDDB65FF8DFBECA8CFFA99FB64" box="[1081,1282,1152,1175]" italics="true" pageId="26" pageNumber="555">Liroetis medvedevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8CC5786DB65FF8DFADDA8CFFAE9FB64" box="[1288,1394,1152,1175]" pageId="26" pageNumber="555">nom. nov.</taxonomicNameLabel>
can be distinguished from all other representatives of
<taxonomicName id="068B4D6CDB65FF8DFA9FA8EFFC19FB24" authorityName="Bryant" authorityYear="1954" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="flavipennis">
<emphasis id="F3FFEAFDDB65FF8DFA9FA8EFFC19FB24" italics="true" pageId="26" pageNumber="555">L. flavipennis</emphasis>
</taxonomicName>
species-group by brownish black pronotum (pale brown in all other species). Dorsal process of aedeagus in lateral view are bifurcate, dorsal branch wide and short, ventral branch long and narrow, separated with large U-shaped incision (
<figureCitation id="59B02A6ADB65FF8DFBC7A90FFBEBFAA4" box="[1042,1136,1344,1367]" captionStart="Figs 108117" captionStartId="29.[151,188,1826,1845]" captionTargetBox="[157,1433,197,1783]" captionTargetId="figure-11@29.[151,1436,189,1988]" captionTargetPageId="29" captionText="Figs 108117.Aedeagus of Liroetis, dorsal and lateral views. 108 L. flavipennis Bryant, 1954; 109 L. humeralis Jiang, 1988; 110 L. lonicernis Jiang, 1988; 111 L. medvedevi nom. nov.; 112 L. nepalensis Chûjô, 1966; 113 L. prominensis Jiang, 1988; 114 L. sichuanensis Jiang, 1988; 115 L. tibetanus Jiang, 1988; 116 L. tibialis Jiang, 1988; 117 L. zhongdianicus Jiang, 1988. Figs 116117 reproduced from JൺඇǤ (1988). Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5821409" httpUri="https://zenodo.org/record/5821409/files/figure.png" pageId="26" pageNumber="555">Fig. 111</figureCitation>
). Similarly modified dorsal process of aedeagus is known also in
<taxonomicName id="068B4D6CDB65FF8DFB1AA92FFAD5FA84" authorityName="Jiang" authorityYear="1988" box="[1231,1358,1376,1399]" class="Insecta" family="Chrysomelidae" genus="Liroetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="tibetanus">
<emphasis id="F3FFEAFDDB65FF8DFB1AA92FFAD5FA84" box="[1231,1358,1376,1399]" italics="true" pageId="26" pageNumber="555">L. tibetanus</emphasis>
</taxonomicName>
which, however, has pale brown pronotum and dorsal process of aedeagus with ventral branch shorter than in
<taxonomicName id="068B4D6CDB65FF8DFAD9A9EFFA0DFA44" authorityName="Bezděk" authorityYear="2021" box="[1292,1430,1440,1463]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="medvedevi">
<emphasis id="F3FFEAFDDB65FF8DFAD9A9EFFA0DFA44" box="[1292,1430,1440,1463]" italics="true" pageId="26" pageNumber="555">L. medvedevi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="89916564DB65FF8DFCE9A98FFC1AFA04" pageId="26" pageNumber="555" type="distribution">
<paragraph id="C13436EFDB65FF8DFCE9A98FFC1AFA04" blockId="26.[828,1437,189,1655]" pageId="26" pageNumber="555">
<emphasis id="F3FFEAFDDB65FF8DFCE9A98FFC4FFA24" bold="true" box="[828,980,1472,1495]" pageId="26" pageNumber="555">Distribution.</emphasis>
<collectingCountry id="B99C767FDB65FF8DFC0AA98FFB86FA24" box="[991,1053,1472,1495]" name="India" pageId="26" pageNumber="555">India</collectingCountry>
:
<collectingRegion id="034FF80DDB65FF8DFBFEA98FFB26FA24" box="[1067,1213,1472,1495]" country="India" name="West Bengal" pageId="26" pageNumber="555">West Bengal</collectingRegion>
,
<collectingCountry id="B99C767FDB65FF8DFB18A98FFA94FA24" box="[1229,1295,1472,1495]" name="Nepal" pageId="26" pageNumber="555">Nepal</collectingCountry>
(Mൾൽඏൾൽൾඏ 2009).
</paragraph>
</subSubSection>
<subSubSection id="89916564DB65FF8DFCE9AA4FFB42F984" pageId="26" pageNumber="555" type="discussion">
<paragraph id="C13436EFDB65FF8DFCE9AA4FFB42F984" blockId="26.[828,1437,189,1655]" pageId="26" pageNumber="555">
<emphasis id="F3FFEAFDDB65FF8DFCE9AA4FFC27F9E4" bold="true" box="[828,956,1536,1559]" pageId="26" pageNumber="555">Comments.</emphasis>
<taxonomicName id="068B4D6CDB65FF8DFC15AA4FFAA8F9E4" authority="Medvedev, 2009" authorityName="Medvedev" authorityYear="2009" box="[960,1331,1536,1559]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="nigricollis">
<emphasis id="F3FFEAFDDB65FF8DFC15AA4FFB1AF9E4" box="[960,1153,1536,1559]" italics="true" pageId="26" pageNumber="555">Liroetis nigricollis</emphasis>
<bibRefCitation id="A51A4B1EDB65FF8DFB50AA4FFAA8F9E4" author="MEDVEDEV L. N." box="[1157,1331,1536,1559]" pageId="26" pageNumber="555" pagination="407 - 410" refId="ref54748" refString="MEDVEDEV L. N. 2009: New species of leaf beetles from the collection of the Naturkundemuseum Erfurt (Insecta: Coleoptera: Chrysomelidae). Pp. 407 - 410. In: HARTMANN M. &amp; WEIPERT J. (eds): Biodiversitat und Naturausstattung im Himalaya III. Verein der Freun- de &amp; Forderer des Naturkundemuseums Erfurt e. V., Erfurt, 477 pp." type="book chapter" year="2009">Medvedev, 2009</bibRefCitation>
</taxonomicName>
is a junior secondary homonym of
<taxonomicName id="068B4D6CDB65FF8DFBEAAA6FFAFBF9C4" authority="(Jiang, 1990)" authorityName="Bezděk" authorityYear="2021" baseAuthorityName="Jiang" baseAuthorityYear="1990" box="[1087,1376,1568,1591]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="nigricollis">
<emphasis id="F3FFEAFDDB65FF8DFBEAAA6FFB52F9C4" box="[1087,1225,1568,1591]" italics="true" pageId="26" pageNumber="555">L. nigricollis</emphasis>
(
<bibRefCitation id="A51A4B1EDB65FF8DFB02AA6FFAC3F9C4" author="JIANG S. - Q." box="[1239,1368,1568,1591]" pageId="26" pageNumber="555" pagination="141 - 144" refId="ref53495" refString="JIANG S. - Q. 1990: On the Chinese species of the Galerucinae genus Zangia (Coleoptera: Chrysomelidae). Sinozoologia 7: 141 - 144." type="journal article" year="1990">Jiang, 1990</bibRefCitation>
)
</taxonomicName>
. New substitute name
<taxonomicName id="068B4D6CDB65FF8DFC3AAA0FFB1BF9A4" authorityName="Bezděk" authorityYear="2021" box="[1007,1152,1600,1623]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="medvedevi" status="nom. nov.">
<emphasis id="F3FFEAFDDB65FF8DFC3AAA0FFB1BF9A4" box="[1007,1152,1600,1623]" italics="true" pageId="26" pageNumber="555">L. medvedevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8CC5786DB65FF8DFB5CAA0FFB6CF9A4" box="[1161,1271,1600,1623]" pageId="26" pageNumber="555">nom. nov.</taxonomicNameLabel>
is proposed to replace
<taxonomicName id="068B4D6CDB65FF8DFC5AAA2FFB4EF984" authority="Medvedev, 2009" authorityName="Medvedev" authorityYear="2009" box="[911,1237,1632,1655]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="nigricollis">
<emphasis id="F3FFEAFDDB65FF8DFC5AAA2FFB81F984" box="[911,1050,1632,1655]" italics="true" pageId="26" pageNumber="555">L. nigricollis</emphasis>
<bibRefCitation id="A51A4B1EDB65FF8DFBF5AA2FFB4EF984" author="MEDVEDEV L. N." box="[1056,1237,1632,1655]" pageId="26" pageNumber="555" pagination="407 - 410" refId="ref54748" refString="MEDVEDEV L. N. 2009: New species of leaf beetles from the collection of the Naturkundemuseum Erfurt (Insecta: Coleoptera: Chrysomelidae). Pp. 407 - 410. In: HARTMANN M. &amp; WEIPERT J. (eds): Biodiversitat und Naturausstattung im Himalaya III. Verein der Freun- de &amp; Forderer des Naturkundemuseums Erfurt e. V., Erfurt, 477 pp." type="book chapter" year="2009">Medvedev, 2009</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>