treatments-xml/data/49/22/87/492287F9DB35FFDBFC63AD58FD09FCA6.xml
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<document id="DFFC5B281A2C4855DFCE38309669C8E5" ID-DOI="10.37520/aemnp.2021.030" ID-ISSN="1804-6487" ID-Zenodo-Dep="5821367" ID-ZooBank="06FDFB43-0B61-4DA8-B260-D78ABD62756C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641391650078" checkinUser="plazi" docAuthor="Bezděk, Jan" docDate="2021" docId="492287F9DB35FFDBFC63AD58FD09FCA6" docLanguage="en" docName="ActaEntMusNatPra.61.2.529-614.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 61 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Liroetis apicicornis Jacoby 1896" docType="treatment" docVersion="3" lastPageNumber="605" masterDocId="B51BFF81DB7FFF97FFD5AC4FFF9BFFF3" masterDocTitle="Redefinition of Liroetis, with descriptions of two new species and an annotated list of species (Coleoptera: Chrysomelidae: Galerucinae)" masterLastPageNumber="614" masterPageNumber="529" pageNumber="603" updateTime="1699285417030" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="3E84A02E89C1122BB898F0B31E1F3C2E">Redefinition of Liroetis, with descriptions of two new species and an annotated list of species (Coleoptera: Chrysomelidae: Galerucinae)</mods:title>
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<mods:namePart id="BD8B8DF832065F40D50137D90D49AB52">Bezděk, Jan</mods:namePart>
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<mods:title id="96D3910D0EF432DB21391C0A438020DC">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="8B8D808577801ACA73F9306D94B0F3C2">2021</mods:date>
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<mods:number id="BFB3D7AD6831600D8BB98E44C33035D1">2021-12-31</mods:number>
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<subSubSection id="89916564DB35FFDDFC63AD58FAB9FEDC" box="[950,1314,279,303]" pageId="74" pageNumber="603" type="nomenclature">
<paragraph id="C13436EFDB35FFDDFC63AD58FAB9FEDC" blockId="74.[950,1314,279,303]" box="[950,1314,279,303]" pageId="74" pageNumber="603">
<heading id="9A7C8183DB35FFDDFC63AD58FAB9FEDC" box="[950,1314,279,303]" centered="true" fontSize="10" level="2" pageId="74" pageNumber="603" reason="3">
<taxonomicName id="068B4D6CDB35FFDDFC63AD58FAB9FEDC" authority="Jacoby, 1896" authorityName="Jacoby" authorityYear="1896" box="[950,1314,279,303]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="603" phylum="Arthropoda" rank="species" species="apicicornis">
<emphasis id="F3FFEAFDDB35FFDDFC63AD58FAB9FEDC" bold="true" box="[950,1314,279,303]" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFC63AD58FB11FEDC" bold="true" box="[950,1162,279,303]" italics="true" pageId="74" pageNumber="603">Liroetis apicicornis</emphasis>
<bibRefCitation id="A51A4B1EDB35FFDDFB44AD57FAB9FEDC" author="JACOBY M." box="[1169,1314,280,303]" pageId="74" pageNumber="603" pagination="250 - 304" refId="ref53423" refString="JACOBY M. 1896: Descriptions of the new genera and species of phytophagous Coleoptera obtained by Mr Andrewes in India. Part II. - Chrysomelinae, Halticinae and Galerucinae. Annales de la Societe Entomologique de Belgique 40: 250 - 304." type="journal article" year="1896">Jacoby, 1896</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89916564DB35FFDDFC3FAD79FB76FEBA" box="[1002,1261,310,329]" pageId="74" pageNumber="603" type="description">
<paragraph id="C13436EFDB35FFDDFC3FAD79FB76FEBA" blockId="74.[828,1438,310,665]" box="[1002,1261,310,329]" pageId="74" pageNumber="603">
(
<figureCitation id="59B02A6ADB35FFDDFC25AD79FBF2FEBA" box="[1008,1129,310,329]" captionStart="Figs 333338" captionStartId="75.[151,188,1908,1927]" captionTargetBox="[159,1436,189,1878]" captionTargetId="figure-54@75.[151,1436,189,1897]" captionTargetPageId="75" captionText="Figs 333338. Liroetis apicicornis Jacoby, 1896. 333335 Male, syntype (9.7 mm). 333 dorsal view; 334 ventral view; 335 labels. 336337 Male (9.5 mm): 336 dorsal view; 337 head and pronotum. 338 Female (10.3 mm)." figureDoi="http://doi.org/10.5281/zenodo.5821500" httpUri="https://zenodo.org/record/5821500/files/figure.png" pageId="74" pageNumber="603">Figs 333338</figureCitation>
,
<figureCitation id="59B02A6ADB35FFDDFBA4AD79FB08FEBA" box="[1137,1171,310,329]" captionStart="Figs 347351" captionStartId="78.[151,188,1354,1373]" captionTargetBox="[179,1399,212,1327]" captionTargetId="figure-13@78.[151,1436,189,1350]" captionTargetPageId="78" captionText="Figs 347351. Aedeagus of Liroetis, dorsal and lateral views, apex of dorsal process in frontal view. 347 L. apicicornis Jacoby, 1896; 348 L. latispinus (Chen, 1976); 349 L. pallidulus (Jiang, 1990); 350 L. reitteri (Pic, 1934); 351 L. violaceipennis Zhang, Li &amp; Yang, 2008. Fig. 348 reproduced from JൺඇǤ (1990). Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5821506" httpUri="https://zenodo.org/record/5821506/files/figure.png" pageId="74" pageNumber="603">347</figureCitation>
,
<figureCitation id="59B02A6ADB35FFDDFB49AD79FB7CFEBA" box="[1180,1255,310,329]" captionStart="Figs 352358" captionStartId="79.[151,188,1246,1265]" captionTargetBox="[157,1424,200,1227]" captionTargetId="figure-11@79.[151,1436,189,1243]" captionTargetPageId="79" captionText="Figs 352358. Spermatheca (Figs 352354) and sternite VII (Figs 355358) of Liroetis. 352, 355 L. apicicornis Jacoby, 1896; 353, 356 L. pallidulus (Jiang, 1990); 354, 358 L. violaceipennis Zhang, Li &amp; Yang, 2008; 357 L. reitteri (Pic, 1934). Scales 0.25 mm for Figs 352354, 0.5 mm for Figs 355358." figureDoi="http://doi.org/10.5281/zenodo.5821508" httpUri="https://zenodo.org/record/5821508/files/figure.png" pageId="74" pageNumber="603">352, 355</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="89916564DB35FFDDFCE9AD16FB63FE75" pageId="74" pageNumber="603" type="reference_group">
<paragraph id="C13436EFDB35FFDDFCE9AD16FADCFE9F" blockId="74.[828,1438,310,665]" box="[828,1351,345,364]" pageId="74" pageNumber="603">
<treatmentCitationGroup id="E19B11C1DB35FFDDFCE9AD16FADCFE9F" box="[828,1351,345,364]" pageId="74" pageNumber="603">
<treatmentCitation id="402A10FEDB35FFDDFCE9AD16FB1DFE9F" author="JACOBY M." box="[828,1158,345,364]" page="282" pageId="74" pageNumber="603" year="1896">
<taxonomicName id="068B4D6CDB35FFDDFCE9AD16FB1DFE9F" authority="Jacoby, 1896: 282" authorityName="Jacoby" authorityPageNumber="282" authorityYear="1896" box="[828,1158,345,364]" class="Insecta" family="Chrysomelidae" genus="Liroetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="603" phylum="Arthropoda" rank="species" species="apicicornis">
<emphasis id="F3FFEAFDDB35FFDDFCE9AD16FC7DFE9F" box="[828,998,345,364]" italics="true" pageId="74" pageNumber="603">Liroëtes apicicornis</emphasis>
<bibRefCitation id="A51A4B1EDB35FFDDFC39AD16FB1DFE9F" author="JACOBY M." box="[1004,1158,345,364]" pageId="74" pageNumber="603" pagination="250 - 304" refId="ref53423" refString="JACOBY M. 1896: Descriptions of the new genera and species of phytophagous Coleoptera obtained by Mr Andrewes in India. Part II. - Chrysomelinae, Halticinae and Galerucinae. Annales de la Societe Entomologique de Belgique 40: 250 - 304." type="journal article" year="1896">Jacoby, 1896: 282</bibRefCitation>
</taxonomicName>
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(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="C13436EFDB35FFDDFCE9AD3CFB63FE75" blockId="74.[828,1438,310,665]" box="[828,1272,371,390]" pageId="74" pageNumber="603">
<taxonomicName id="068B4D6CDB35FFDDFCE9AD3CFC78FE75" authorityName="Jacoby" authorityYear="1896" box="[828,995,371,390]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="603" phylum="Arthropoda" rank="species" species="apicicornis">
<emphasis id="F3FFEAFDDB35FFDDFCE9AD3CFC78FE75" box="[828,995,371,390]" italics="true" pageId="74" pageNumber="603">Liroëtis apicicornis</emphasis>
</taxonomicName>
: Wൾඌൾ (1924): 128 (catalogue).
</paragraph>
</subSubSection>
<subSubSection id="89916564DB35FFDDFCE9ADC1FA12FE4F" pageId="74" pageNumber="603" type="description">
<paragraph id="C13436EFDB35FFDDFCE9ADC1FA12FE4F" blockId="74.[828,1438,310,665]" pageId="74" pageNumber="603">
<taxonomicName id="068B4D6CDB35FFDDFCE9ADC1FC7EFE52" authorityName="Jacoby" authorityYear="1896" box="[828,997,398,417]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="603" phylum="Arthropoda" rank="species" species="apicicornis">
<emphasis id="F3FFEAFDDB35FFDDFCE9ADC1FC7EFE52" box="[828,997,398,417]" italics="true" pageId="74" pageNumber="603">Liroetis apicicornis</emphasis>
</taxonomicName>
: Mൺඎඅĸ (1936): 311; Wඅർඈඑ (1973): 476 (catalogue); Kආඈඍඈ (1989a): 82 (misidentification, =
<taxonomicName id="068B4D6CDB35FFDDFAD5ADE6FAE6FE4F" authorityName="Bezděk" authorityYear="2021" box="[1280,1405,425,444]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="603" phylum="Arthropoda" rank="species" species="aurantiacus">
<emphasis id="F3FFEAFDDB35FFDDFAD5ADE6FAE6FE4F" box="[1280,1405,425,444]" italics="true" pageId="74" pageNumber="603">L. aurantiacus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="89916564DB35FFDDFCE9AD84FADFFD6B" pageId="74" pageNumber="603" type="materials_examined">
<paragraph id="C13436EFDB35FFDDFCE9AD84FC73FE0B" blockId="74.[828,1438,310,665]" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFCE9AD84FC29FE2D" bold="true" box="[828,946,459,478]" pageId="74" pageNumber="603">
<typeStatus id="1E30884DDB35FFDDFCE9AD84FCF3FE2D" box="[828,872,459,478]" pageId="74" pageNumber="603">Type</typeStatus>
locality.
</emphasis>
Kanara, S.
<collectingCountry id="B99C767FDB35FFDDFBC2AD84FBC4FE2D" box="[1047,1119,459,478]" name="India" pageId="74" pageNumber="603">Bombay</collectingCountry>
[=
<collectingCountry id="B99C767FDB35FFDDFBAFAD84FB32FE2D" box="[1146,1193,459,478]" name="India" pageId="74" pageNumber="603">India</collectingCountry>
:
<collectingRegion id="034FF80DDB35FFDDFB7AAD84FA92FE2D" box="[1199,1289,459,478]" country="India" name="Karnataka" pageId="74" pageNumber="603">Karnataka</collectingRegion>
:Karavali (former Northern Kanara)].
</paragraph>
<paragraph id="C13436EFDB35FFDDFCE9AE4FFB44FD90" blockId="74.[828,1438,310,665]" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFCE9AE4FFB85FDE0" bold="true" box="[828,1054,512,531]" pageId="74" pageNumber="603">
<typeStatus id="1E30884DDB35FFDDFCE9AE4FFCF2FDE0" box="[828,873,512,531]" pageId="74" pageNumber="603">Type</typeStatus>
material examined.
</emphasis>
Sඒඇඍඒඉൾ: 1 J (
<figureCitation id="59B02A6ADB35FFDDFB73AE4FFABBFDE0" box="[1190,1312,512,531]" captionStart="Figs 333338" captionStartId="75.[151,188,1908,1927]" captionTargetBox="[159,1436,189,1878]" captionTargetId="figure-54@75.[151,1436,189,1897]" captionTargetPageId="75" captionText="Figs 333338. Liroetis apicicornis Jacoby, 1896. 333335 Male, syntype (9.7 mm). 333 dorsal view; 334 ventral view; 335 labels. 336337 Male (9.5 mm): 336 dorsal view; 337 head and pronotum. 338 Female (10.3 mm)." figureDoi="http://doi.org/10.5281/zenodo.5821500" httpUri="https://zenodo.org/record/5821500/files/figure.png" pageId="74" pageNumber="603">Figs 333335</figureCitation>
),
<typeStatus id="1E30884DDB35FFDDFAE7AE4FFAFBFDE0" box="[1330,1376,512,531]" pageId="74" pageNumber="603">Type</typeStatus>
/ H. T. [white round label with red collar, p] // Kanara [w, p] // Jacoby Coll. / 190928a [w, p] //
<taxonomicName id="068B4D6CDB35FFDDFC31AE7AFBB1FDBB" authorityName="Jacoby" authorityYear="1890" box="[996,1066,565,584]" class="Insecta" family="Chrysomelidae" genus="Liroetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="603" phylum="Arthropoda" rank="genus">Liroëtes</taxonomicName>
/ apicicornis / Jac. [b, h] // SYN- /
<typeStatus id="1E30884DDB35FFDDFAB2AE7AFA07FDBB" box="[1383,1436,565,584]" pageId="74" pageNumber="603">TYPE</typeStatus>
[white round label with blue collar, p] (
<collectionCode id="A79AAE2ADB35FFDDFB45AE1FFB4EFD90" box="[1168,1237,592,611]" pageId="74" pageNumber="603">BMNH</collectionCode>
).
</paragraph>
<paragraph id="C13436EFDB35FFDDFCE9AE24FADFFD6B" blockId="74.[828,1438,310,665]" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFCE9AE24FB00FD8D" bold="true" box="[828,1179,619,638]" pageId="74" pageNumber="603">
Material examined.
<collectingCountry id="B99C767FDB35FFDDFC2FAE24FBA4FD8D" box="[1018,1087,619,638]" name="India" pageId="74" pageNumber="603">INDIA</collectingCountry>
: Kൾඋൺඅൺ:
</emphasis>
Kallar valley,
<quantity id="06739B0ADB35FFDDFAF4AE24FAC1FD8D" box="[1313,1370,619,638]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="74" pageNumber="603" unit="km" value="15.0">15 km</quantity>
SW of Munnar,
<quantity id="06739B0ADB35FFDDFC5CAECAFC57FD6B" box="[905,972,645,664]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="74" pageNumber="603" unit="m" value="1250.0">1250 m</quantity>
,
<date id="B535102FDB35FFDDFC01AECAFBA1FD6B" box="[980,1082,645,664]" pageId="74" pageNumber="603" value="1997-05-01" valueMax="1997-05-09" valueMin="1997-05-01">1.9.v.1997</date>
, 1 J
<specimenCount id="D78DFD66DB35FFDDFBBDAECAFB13FD6B" box="[1128,1160,639,664]" pageId="74" pageNumber="603" type="female">1 ♀</specimenCount>
, R. Peša leg. (
<collectionCode id="A79AAE2ADB35FFDDFAD6AECAFAA1FD6B" box="[1283,1338,645,664]" pageId="74" pageNumber="603">JBCB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="89916564DB35FFDDFCE9AEFEFB17F87B" pageId="74" pageNumber="603" type="diagnosis">
<paragraph id="C13436EFDB35FFDDFCE9AEFEFA17FD1B" blockId="74.[828,1437,689,2056]" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFCE9AEFEFC2BFD3B" bold="true" box="[828,944,689,712]" pageId="74" pageNumber="603">Diagnosis.</emphasis>
<emphasis id="F3FFEAFDDB35FFDDFC60AEFEFBACFD3B" box="[949,1079,689,712]" italics="true" pageId="74" pageNumber="603">Colouration</emphasis>
. Body, legs and antennae completely pale brown, except two apical antennomeres black.
</paragraph>
<paragraph id="C13436EFDB35FFDDFC8FAEBEFABDFCFB" blockId="74.[828,1437,689,2056]" box="[858,1318,750,776]" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFC8FAEBEFC41FCFB" box="[858,986,753,776]" italics="true" pageId="74" pageNumber="603">Body length</emphasis>
. JJ:
<quantity id="06739B0ADB35FFDDFBC0AEBEFB0CFCFB" box="[1045,1175,753,776]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="9.7" metricValueMin="9.5" pageId="74" pageNumber="603" unit="mm" value="9.6" valueMax="9.7" valueMin="9.5">9.59.7 mm</quantity>
,
<specimenCount id="D78DFD66DB35FFDDFB77AEA1FB29FCF4" box="[1186,1202,750,775]" pageId="74" pageNumber="603" type="female"></specimenCount>
:
<quantity id="06739B0ADB35FFDDFB15AEBEFAB9FCFB" box="[1216,1314,753,776]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="74" pageNumber="603" unit="mm" value="10.3">10.3 mm</quantity>
.
</paragraph>
<paragraph id="C13436EFDB35FFDDFC8FAF5EFB79FBFB" blockId="74.[828,1437,689,2056]" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFC8FAF5EFC09FCDB" box="[858,914,785,808]" italics="true" pageId="74" pageNumber="603">Male</emphasis>
(
<figureCitation id="59B02A6ADB35FFDDFC71AF5EFBD8FCDB" box="[932,1091,785,808]" captionStart="Figs 333338" captionStartId="75.[151,188,1908,1927]" captionTargetBox="[159,1436,189,1878]" captionTargetId="figure-54@75.[151,1436,189,1897]" captionTargetPageId="75" captionText="Figs 333338. Liroetis apicicornis Jacoby, 1896. 333335 Male, syntype (9.7 mm). 333 dorsal view; 334 ventral view; 335 labels. 336337 Male (9.5 mm): 336 dorsal view; 337 head and pronotum. 338 Female (10.3 mm)." figureDoi="http://doi.org/10.5281/zenodo.5821500" httpUri="https://zenodo.org/record/5821500/files/figure.png" pageId="74" pageNumber="603">Figs 336337</figureCitation>
). Antennae 0.62 times as long as body. Pronotum 1.95 times as wide as long, lustrous, almost impunctate, anterior margin bordered. Abdominal ventrite IV with small subtriangular process in middle of posterior margin, surrounding surface slightly impressed. Last abdominal ventrite with longitudinal impression narrower in posterior part, wider basally. Protarsomere I narrow, metatibial spur short, very thin.
</paragraph>
<paragraph id="C13436EFDB35FFDDFC8FA85EFA06FB1B" blockId="74.[828,1437,689,2056]" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFC8FA85EFC58FBDB" box="[858,963,1041,1064]" italics="true" pageId="74" pageNumber="603">Aedeagus</emphasis>
(
<figureCitation id="59B02A6ADB35FFDDFC03A85EFBA2FBDB" box="[982,1081,1041,1064]" captionStart="Figs 347351" captionStartId="78.[151,188,1354,1373]" captionTargetBox="[179,1399,212,1327]" captionTargetId="figure-13@78.[151,1436,189,1350]" captionTargetPageId="78" captionText="Figs 347351. Aedeagus of Liroetis, dorsal and lateral views, apex of dorsal process in frontal view. 347 L. apicicornis Jacoby, 1896; 348 L. latispinus (Chen, 1976); 349 L. pallidulus (Jiang, 1990); 350 L. reitteri (Pic, 1934); 351 L. violaceipennis Zhang, Li &amp; Yang, 2008. Fig. 348 reproduced from JൺඇǤ (1990). Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5821506" httpUri="https://zenodo.org/record/5821506/files/figure.png" pageId="74" pageNumber="603">Fig. 347</figureCitation>
). Median lobe of aedeagus flat, lanceolate, 2.47 times as long as wide, widest in middle part; apical part subtriangular with incised tip. Lateral view: slightly bent, lateral elevation absent. Dorsal process short, 1.66 times as long as wide, 0.36 times as long as median lobe; apical part wide, covered with very long setae, in middle with wing-like structure; in lateral view Y-shaped.
</paragraph>
<paragraph id="C13436EFDB35FFDDFC8FA8BEFB82FA1B" blockId="74.[828,1437,689,2056]" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFC8FA8BEFC32FAFB" box="[858,937,1265,1288]" italics="true" pageId="74" pageNumber="603">Female</emphasis>
. Metatibial spur absent. Last abdominal ventrite with posterior margin entire. Sternite VIII transversely suboval with triangular median process, posterior half of sternite VIII and median process covered with very long setae; tignum as long as sternite VIII, with asymmetrical oblique apex (
<figureCitation id="59B02A6ADB35FFDDFC19A9DEFBBDFA5B" box="[972,1062,1425,1448]" captionStart="Figs 352358" captionStartId="79.[151,188,1246,1265]" captionTargetBox="[157,1424,200,1227]" captionTargetId="figure-11@79.[151,1436,189,1243]" captionTargetPageId="79" captionText="Figs 352358. Spermatheca (Figs 352354) and sternite VII (Figs 355358) of Liroetis. 352, 355 L. apicicornis Jacoby, 1896; 353, 356 L. pallidulus (Jiang, 1990); 354, 358 L. violaceipennis Zhang, Li &amp; Yang, 2008; 357 L. reitteri (Pic, 1934). Scales 0.25 mm for Figs 352354, 0.5 mm for Figs 355358." figureDoi="http://doi.org/10.5281/zenodo.5821508" httpUri="https://zenodo.org/record/5821508/files/figure.png" pageId="74" pageNumber="603">Fig. 355</figureCitation>
). Spermatheca with suboval nodulus covered with traces of transverse wrinkles, cornu short C-shaped (
<figureCitation id="59B02A6ADB35FFDDFC7AA99EFB96FA1B" box="[943,1037,1489,1512]" captionStart="Figs 352358" captionStartId="79.[151,188,1246,1265]" captionTargetBox="[157,1424,200,1227]" captionTargetId="figure-11@79.[151,1436,189,1243]" captionTargetPageId="79" captionText="Figs 352358. Spermatheca (Figs 352354) and sternite VII (Figs 355358) of Liroetis. 352, 355 L. apicicornis Jacoby, 1896; 353, 356 L. pallidulus (Jiang, 1990); 354, 358 L. violaceipennis Zhang, Li &amp; Yang, 2008; 357 L. reitteri (Pic, 1934). Scales 0.25 mm for Figs 352354, 0.5 mm for Figs 355358." figureDoi="http://doi.org/10.5281/zenodo.5821508" httpUri="https://zenodo.org/record/5821508/files/figure.png" pageId="74" pageNumber="603">Fig. 352</figureCitation>
).
</paragraph>
<paragraph id="C13436EFDB35FFDDFCE9A9BEFB17F87B" blockId="74.[828,1437,689,2056]" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFCE9A9BEFBAFF9FB" bold="true" box="[828,1076,1521,1544]" pageId="74" pageNumber="603">Differential diagnosis.</emphasis>
<taxonomicName id="068B4D6CDB35FFDDFBEFA9BEFA90F9FB" authorityName="Jacoby" authorityYear="1896" box="[1082,1291,1521,1544]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="603" phylum="Arthropoda" rank="species" species="apicicornis">
<emphasis id="F3FFEAFDDB35FFDDFBEFA9BEFA90F9FB" box="[1082,1291,1521,1544]" italics="true" pageId="74" pageNumber="603">Liroetis apicicornis</emphasis>
</taxonomicName>
is characterised by body, legs and antennae completely orange brown, with exception of two apical antennomeres that are black, median lobe of aedeagus flat and wide, with dorsal process short, and by pronotum nearly twice as wide as long. It is the only known species of
<taxonomicName id="068B4D6CDB35FFDDFB99AADEFB06F95B" authorityName="Weise" authorityYear="1889" box="[1100,1181,1681,1704]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="603" phylum="Arthropoda" rank="genus">
<emphasis id="F3FFEAFDDB35FFDDFB99AADEFB06F95B" box="[1100,1181,1681,1704]" italics="true" pageId="74" pageNumber="603">Liroetis</emphasis>
</taxonomicName>
inhabiting south-eastern
<collectingCountry id="B99C767FDB35FFDDFCE9AAFEFCE8F93B" box="[828,883,1713,1736]" name="India" pageId="74" pageNumber="603">India</collectingCountry>
and its distribution is disjunct from other species of the genus (the nearest species occur in Himalayas and easternmost
<collectingCountry id="B99C767FDB35FFDDFC10AABEFB9BF8FB" box="[965,1024,1777,1800]" name="India" pageId="74" pageNumber="603">India</collectingCountry>
).
<taxonomicName id="068B4D6CDB35FFDDFBC1AABEFB74F8FB" authority="Bezděk, 2021" authorityName="Bezděk" authorityYear="2021" box="[1044,1263,1777,1800]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="603" phylum="Arthropoda" rank="species" species="aurantiacus" status="sp. nov.">
<emphasis id="F3FFEAFDDB35FFDDFBC1AABEFB74F8FB" box="[1044,1263,1777,1800]" italics="true" pageId="74" pageNumber="603">Liroetis aurantiacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8CC5786DB35FFDDFB2CAABEFAD5F8FB" box="[1273,1358,1777,1800]" pageId="74" pageNumber="603" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="068B4D6CDB35FFDDFA52AABEFC2CF8DB" authority="Bezděk, 2021" authorityName="Bezděk" authorityYear="2021" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="603" phylum="Arthropoda" rank="species" species="baolocanus" status="sp. nov.">
<emphasis id="F3FFEAFDDB35FFDDFA52AABEFC2CF8DB" italics="true" pageId="74" pageNumber="603">L. baolocanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8CC5786DB35FFDDFC15AB5EFB8EF8DB" box="[960,1045,1809,1832]" pageId="74" pageNumber="603" rank="species">sp. nov.</taxonomicNameLabel>
from SE Asia have similarly transverse pronotum and similar colouration but they differ in completely brown antennae and very different structure of aedeagus (cf.
<figureCitation id="59B02A6ADB35FFDDFC1AAB3EFBD2F87B" box="[975,1097,1905,1928]" captionStart="Figs 6571" captionStartId="20.[151,188,1622,1641]" captionTargetBox="[161,1414,192,1585]" captionTargetId="figure-13@20.[151,1436,176,1975]" captionTargetPageId="20" captionText="Figs 6571. Details of new species of Liroetis. 6570 L. aurantiacus sp. nov. 65 Aedegus in dorsal, lateral and ventral views; 66 female pygidium; 67 spermatheca; 68 sternite VIII; 69 male abdomen in ventral and lateral views. 7071 L. baolocanus sp. nov. 70 Aedegus in dorsal, lateral and ventral views; 71 male abdomen in ventral and lateral views. Scales: 0.5 mm for Figs 6566, 68 and 70, 0.25 mm for Fig. 67; 1 mm for Figs 69, 71." figureDoi="http://doi.org/10.5281/zenodo.5821391" httpUri="https://zenodo.org/record/5821391/files/figure.png" pageId="74" pageNumber="603">Figs 65, 70</figureCitation>
,
<figureCitation id="59B02A6ADB35FFDDFB81AB3EFB1BF87B" box="[1108,1152,1905,1928]" captionStart="Figs 347351" captionStartId="78.[151,188,1354,1373]" captionTargetBox="[179,1399,212,1327]" captionTargetId="figure-13@78.[151,1436,189,1350]" captionTargetPageId="78" captionText="Figs 347351. Aedeagus of Liroetis, dorsal and lateral views, apex of dorsal process in frontal view. 347 L. apicicornis Jacoby, 1896; 348 L. latispinus (Chen, 1976); 349 L. pallidulus (Jiang, 1990); 350 L. reitteri (Pic, 1934); 351 L. violaceipennis Zhang, Li &amp; Yang, 2008. Fig. 348 reproduced from JൺඇǤ (1990). Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5821506" httpUri="https://zenodo.org/record/5821506/files/figure.png" pageId="74" pageNumber="603">347</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89916564DB35FFDDFCE9ABDEFC7FF83B" pageId="74" pageNumber="603" type="distribution">
<paragraph id="C13436EFDB35FFDDFCE9ABDEFC7FF83B" blockId="74.[828,1437,689,2056]" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFCE9ABDEFC4AF85B" bold="true" box="[828,977,1937,1960]" pageId="74" pageNumber="603">Distribution.</emphasis>
<collectingCountry id="B99C767FDB35FFDDFC0EABDEFB8CF85B" box="[987,1047,1937,1960]" name="India" pageId="74" pageNumber="603">India</collectingCountry>
:
<collectingRegion id="034FF80DDB35FFDDFBF0ABDEFB0EF85B" box="[1061,1173,1937,1960]" country="India" name="Karnataka" pageId="74" pageNumber="603">Karnataka</collectingRegion>
(Jൺർඈൻඒ 1896),
<collectingRegion id="034FF80DDB35FFDDFA81ABDEFA07F85B" box="[1364,1436,1937,1960]" country="India" name="Kerala" pageId="74" pageNumber="603">Kerala</collectingRegion>
(present paper).
</paragraph>
</subSubSection>
<subSubSection id="89916564DB35FFDBFCE9AB9EFD09FCA6" lastPageId="76" lastPageNumber="605" pageId="74" pageNumber="603" type="discussion">
<paragraph id="C13436EFDB35FFDBFCE9AB9EFE26FE66" blockId="74.[828,1437,689,2056]" lastBlockId="76.[151,761,190,853]" lastPageId="76" lastPageNumber="605" pageId="74" pageNumber="603">
<emphasis id="F3FFEAFDDB35FFDDFCE9AB9EFC25F81B" bold="true" box="[828,958,2001,2024]" pageId="74" pageNumber="603">Comments.</emphasis>
In the original description, Jൺർඈൻඒ (1896) indicated that only the ultimate antennomere is black. The two additional specimens examined are identical with the type specimens including the aedeagus but have the two last antennomeres black. Unfortunately, the antennae of the
<typeStatus id="1E30884DDB33FFDBFF6BAD51FE8AFEC6" box="[190,273,286,309]" pageId="76" pageNumber="605" type="syntype">syntype</typeStatus>
are largely missing. I am unable to determine whether Jൺർඈൻඒ (1896) made a mistake in the original description or whether there is variability in colouration of the apical antennomeres.
</paragraph>
<caption id="95F46667DB34FFDCFF42AB3BFC1EF852" ID-DOI="http://doi.org/10.5281/zenodo.5821500" ID-Zenodo-Dep="5821500" httpUri="https://zenodo.org/record/5821500/files/figure.png" pageId="75" pageNumber="604" startId="75.[151,188,1908,1927]" targetBox="[159,1436,189,1878]" targetPageId="75">
<paragraph id="C13436EFDB34FFDCFF42AB3BFC1EF852" blockId="75.[151,1437,1908,1953]" pageId="75" pageNumber="604">
Figs 333338.
<taxonomicName id="068B4D6CDB34FFDCFEC2AB3BFDACF874" authority="Jacoby, 1896" authorityName="Jacoby" authorityYear="1896" box="[279,567,1908,1927]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="604" phylum="Arthropoda" rank="species" species="apicicornis">
<emphasis id="F3FFEAFDDB34FFDCFEC2AB3BFE24F874" box="[279,447,1908,1927]" italics="true" pageId="75" pageNumber="604">Liroetis apicicornis</emphasis>
<bibRefCitation id="A51A4B1EDB34FFDCFE10AB3BFDACF874" author="JACOBY M." box="[453,567,1908,1927]" pageId="75" pageNumber="604" pagination="250 - 304" refId="ref53423" refString="JACOBY M. 1896: Descriptions of the new genera and species of phytophagous Coleoptera obtained by Mr Andrewes in India. Part II. - Chrysomelinae, Halticinae and Galerucinae. Annales de la Societe Entomologique de Belgique 40: 250 - 304." type="journal article" year="1896">Jacoby, 1896</bibRefCitation>
</taxonomicName>
. 333335 Male, syntype (9.7 mm). 333 dorsal view; 334 ventral view; 335 labels. 336337 Male (9.5 mm): 336 dorsal view; 337 head and pronotum. 338 Female (10.3 mm).
</paragraph>
</caption>
<paragraph id="C13436EFDB33FFDBFF60ADD1FDC5FD86" blockId="76.[151,761,190,853]" pageId="76" pageNumber="605">
<taxonomicName id="068B4D6CDB33FFDBFF60ADD1FE1DFE46" authorityName="Jacoby" authorityYear="1896" box="[181,390,414,437]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="605" phylum="Arthropoda" rank="species" species="apicicornis">
<emphasis id="F3FFEAFDDB33FFDBFF60ADD1FE1DFE46" box="[181,390,414,437]" italics="true" pageId="76" pageNumber="605">Liroetis apicicornis</emphasis>
</taxonomicName>
has metatibial spur present in males and absent in females, pronotum ca. twice as wide as long with anterior margin narrowly bordered and median lobe of aedeagus flat and wide, without lateral elevation. Such combination of characters is unique within the genus
<taxonomicName id="068B4D6CDB33FFDBFF42AE71FF72FDA6" authorityName="Weise" authorityYear="1889" box="[151,233,574,597]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="605" phylum="Arthropoda" rank="genus">
<emphasis id="F3FFEAFDDB33FFDBFF42AE71FF72FDA6" box="[151,233,574,597]" italics="true" pageId="76" pageNumber="605">Liroetis</emphasis>
</taxonomicName>
and thus
<taxonomicName id="068B4D6CDB33FFDBFE9BAE71FE79FDA6" authorityName="Jacoby" authorityYear="1896" box="[334,482,574,597]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="605" phylum="Arthropoda" rank="species" species="apicicornis">
<emphasis id="F3FFEAFDDB33FFDBFE9BAE71FE79FDA6" box="[334,482,574,597]" italics="true" pageId="76" pageNumber="605">L. apicicornis</emphasis>
</taxonomicName>
is for now not assigned to any of the species groups herein proposed.
</paragraph>
<paragraph id="C13436EFDB33FFDBFF60AE31FD09FCA6" blockId="76.[151,761,190,853]" pageId="76" pageNumber="605">
The records from
<collectingCountry id="B99C767FDB33FFDBFEBBAE31FE54FD66" box="[366,463,638,661]" name="Thailand" pageId="76" pageNumber="605">Thailand</collectingCountry>
,
<collectingCountry id="B99C767FDB33FFDBFE0DAE31FD97FD66" box="[472,524,638,661]" name="Laos" pageId="76" pageNumber="605">Laos</collectingCountry>
and
<collectingCountry id="B99C767FDB33FFDBFDEEAE31FD0EFD66" box="[571,661,638,661]" name="Vietnam" pageId="76" pageNumber="605">Vietnam</collectingCountry>
(Kආඈඍඈ 1989a) evidently refer to another species (at least part of them to
<taxonomicName id="068B4D6CDB33FFDBFF22AEF1FE4CFD26" authority="Bezděk, 2021" authorityName="Bezděk" authorityYear="2021" box="[247,471,702,725]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="605" phylum="Arthropoda" rank="species" species="aurantiacus" status="sp. nov.">
<emphasis id="F3FFEAFDDB33FFDBFF22AEF1FE4CFD26" box="[247,471,702,725]" italics="true" pageId="76" pageNumber="605">Liroetis aurantiacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8CC5786DB33FFDBFE34AEF1FDA6FD26" box="[481,573,702,725]" pageId="76" pageNumber="605" rank="species">sp. nov.</taxonomicNameLabel>
). The specimens published from
<collectingCountry id="B99C767FDB33FFDBFE93AE91FE1DFD06" box="[326,390,734,757]" name="Nepal" pageId="76" pageNumber="605">Nepal</collectingCountry>
(Mൾൽඏൾൽൾඏ &amp; SඉඋൾർIJൾඋ-Uൾൻൾඋ- ඌൺඑ 1998, 1999) were later described as
<taxonomicName id="068B4D6CDB33FFDBFDB1AEB1FECBFCC6" authority="Medvedev, 2007" authorityName="Medvedev" authorityYear="2007" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="605" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="F3FFEAFDDB33FFDBFDB1AEB1FD63FCE6" box="[612,760,766,789]" italics="true" pageId="76" pageNumber="605">L. brancuccii</emphasis>
<bibRefCitation id="A51A4B1EDB33FFDBFF42AF51FECBFCC6" author="MEDVEDEV L. N." box="[151,336,798,821]" pageId="76" pageNumber="605" pagination="289 - 305" refId="ref54721" refString="MEDVEDEV L. N. 2007: New species of Oriental Chrysomelidae (Coleoptera). Entomologica Basiliensia et Collectionis Frey 29: 289 - 305." type="journal article" year="2007">Medvedev, 2007</bibRefCitation>
</taxonomicName>
(Mൾൽඏൾൽൾඏ 2007), now classified as
<taxonomicName id="068B4D6CDB33FFDBFF42AF71FE31FCA6" box="[151,426,830,853]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="605" phylum="Arthropoda" rank="species" species="brancuccii">
<emphasis id="F3FFEAFDDB33FFDBFF42AF71FE31FCA6" box="[151,426,830,853]" italics="true" pageId="76" pageNumber="605">Hesperopenna brancuccii</emphasis>
</taxonomicName>
(see Bൾඓൽෂĸ 2016b).
</paragraph>
</subSubSection>
</treatment>
</document>