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<document id="7C80B40DF1F35B9AE6E792A9C3E1614E" ID-DOI="10.11646/zootaxa.5537.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14239439" ID-ZooBank="8A7AF43F-0E83-48A0-950E-0716CDC753A6" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1732800333346" checkinUser="plazi" docAuthor="Selis, Marco, Fateryga, Alexander V. &amp; Cilia, Giovanni" docDate="2024" docId="C866706D622A9A1E89FDF243D29BF954" docLanguage="en" docName="zootaxa.5537.2.1.pdf" docOrigin="Zootaxa 5537 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5537.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euodynerus Dalla Torre 1904" docType="key" docVersion="1" lastPageNumber="185" masterDocId="345F0815620A9A3C896AF525D707FFD1" masterDocTitle="The genus Euodynerus Dalla Torre in Europe and the Maghreb (Hymenoptera: Vespidae: Eumeninae)" masterLastPageNumber="194" masterPageNumber="151" pageNumber="183" updateTime="1733413254902" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="08D774C883F8D36CF2844194541871F4">The genus Euodynerus Dalla Torre in Europe and the Maghreb (Hymenoptera: Vespidae: Eumeninae)</mods:title>
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<mods:namePart id="6E51343C8B6835BDD0C4D411AC6570E0">Selis, Marco</mods:namePart>
<mods:affiliation id="6F3829CF2A8D186D27966E553D45BBF0">Via dei Tarquini, 22 - 01100 Viterbo, Italy</mods:affiliation>
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<mods:namePart id="5D255925309BBD63369AF02364037ED3">Fateryga, Alexander V.</mods:namePart>
<mods:affiliation id="527C1598E996C40A76AB592337B00FD8">T. I. Vyazemsky Karadag Scientific Station-Nature Reserve of RAS-Branch of A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, Nauki Str. 24, Kurortnoye, 298188 Feodosiya, Russia</mods:affiliation>
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<mods:namePart id="031ABE0C47B851426A53C54693C96DAD">Cilia, Giovanni</mods:namePart>
<mods:affiliation id="893BF6C7878039DA099FE87C1B5C6747">CREA Research Centre for Agriculture and Environment (CREA-AA), Via di Corticella 133, 40128, Bologna, Italy</mods:affiliation>
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<mods:date id="AD5DBE443E893465C4C20C326614B675">2024</mods:date>
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<treatment id="C866706D622A9A1E89FDF243D29BF954" LSID="urn:lsid:plazi:treatment:C866706D622A9A1E89FDF243D29BF954" httpUri="http://treatment.plazi.org/id/C866706D622A9A1E89FDF243D29BF954" lastPageId="34" lastPageNumber="185" pageId="32" pageNumber="183">
<subSubSection id="08D592F0622A9A1C89FDF243D313F851" box="[151,1044,1893,1920]" pageId="32" pageNumber="183" type="nomenclature">
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<heading id="1B387617622A9A1C89FDF243D313F851" bold="true" box="[151,1044,1893,1920]" fontSize="11" level="1" pageId="32" pageNumber="183" reason="1">
<emphasis id="72BB1D69622A9A1C89FDF243D313F851" bold="true" box="[151,1044,1893,1920]" pageId="32" pageNumber="183">
Key to the species of
<taxonomicName id="87CFBAF8622A9A1C88F7F240D537F851" ID-CoL="4G4Z" ID-ENA="187861" authority="Dalla Torre, 1904" authorityName="Dalla Torre" authorityYear="1904" box="[413,560,1893,1920]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="72BB1D69622A9A1C88F7F240D537F851" bold="true" box="[413,560,1893,1920]" italics="true" pageId="32" pageNumber="183">Euodynerus</emphasis>
</taxonomicName>
occurring in Europe and the Maghreb
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<keyStep id="FB3BD9DE622A9A1D89FDF28ED29BFE8C" lastPageId="33" lastPageNumber="184" pageId="32" pageNumber="183">
<paragraph id="4070C17B622A9A1D89FDF28ED29BFF1B" blockId="32.[151,1436,1962,2014]" lastBlockId="33.[151,1437,151,1992]" lastPageId="33" lastPageNumber="184" pageId="32" pageNumber="183">
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<footnote id="23D4DD75622A9A1C89FDF28ED29BF80F" pageId="32" pageNumber="183">1. S2 step-like in lateral view, with a basal vertical face sharply delimited by transverse ridge which is strongly projecting in the middle (Fig. 15A); surface of S2 shallowly but distinctly depressed just behind transverse ridge (Fig. 15A). Propodeum without</footnote>
dorsal carinae (
<figureCitation id="D8F4DDFE622B9A1D881CF5B2D6CBFF7C" box="[374,460,151,173]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15F</figureCitation>
). Vestiture of head, mesosoma and anterior part of metasoma made of long and wavy setae, reaching up to three ocellar diameters in length (
<figureCitation id="D8F4DDFE622B9A1D8B31F590D5AEFF1A" box="[603,681,181,203]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15J</figureCitation>
)............................................................. 2
</keyLead>
</paragraph>
<paragraph id="4070C17B622B9A1D89FDF5F6D29BFE8C" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF5F6D29BFE8C" pageId="33" pageNumber="184">
- Characters not combined as above. S2 usually evenly convex in lateral view and with a basal furrow in the middle (
<figureCitation id="D8F4DDFE622B9A1D8C49F5F7D7F3FED4" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Figs 15B, C, E</figureCitation>
); if step-like, then basal transverse ridge broadly interrupted in the middle by longitudinal furrow and not projecting as above (
<figureCitation id="D8F4DDFE622B9A1D8983F428D645FEF2" box="[233,322,269,291]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15D</figureCitation>
); surface of S2 usually not depressed in lateral view (depressed as above only in
<taxonomicName id="87CFBAF8622B9A1D8D2DF428D3B3FEF3" authorityName="RU Zeleny Sad" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[1095,1204,269,290]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="caspicus">
<emphasis id="72BB1D69622B9A1D8D2DF428D3B3FEF3" box="[1095,1204,269,290]" italics="true" pageId="33" pageNumber="184">E. caspicus</emphasis>
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). Propodeal carinae and vestiture variable, but never combined as above (e.g. if propodeum without dorsal carinae, then vestiture not made of long and wavy setae) (
<figureCitation id="D8F4DDFE622B9A1D880AF462D50DFE8C" box="[352,522,327,349]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Figs 15G–I, K–O</figureCitation>
).......................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FB3BD9DE622B9A1D89FDF440D29BFD6C" pageId="33" pageNumber="184">
<paragraph id="4070C17B622B9A1D89FDF440D29BFDDC" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF440D29BFDDC" pageId="33" pageNumber="184">
2. Disc of clypeus entirely flattened and not depressed above apex; apical margin wider than interantennal space, subtruncate in female and deeply emarginate in male (
<figureCitation id="D8F4DDFE622B9A1D8B34F4A7D5CEFE46" box="[606,713,386,408]" captionStart="FIGURE 4" captionStartId="13.[151,250,1804,1828]" captionTargetBox="[151,1435,181,1780]" captionTargetId="figure-17@13.[151,1435,181,1780]" captionTargetPageId="13" captionText="FIGURE 4. A, C–D, Euodynerus bidentatus bidentatus (Lepeletier). A, D, female from Morocco. C, male from Morocco. B, E, Euodynerus bidentatus puniceus Blüthgen, female from Libya. F–I, Euodynerus caspicus (Morawitz). F, H, male from Astrakhan Province, Russia. G, I, female from Kazakhstan." figureDoi="http://doi.org/10.5281/zenodo.14239447" httpUri="https://zenodo.org/record/14239447/files/figure.png" pageId="33" pageNumber="184">Figs 4C–E</figureCitation>
). Punctures on mesosoma finer and denser and with densely micropunctate interspaces, giving matte appearance. Scutellum with subtriangular lateral spots (absent in
<taxonomicName id="87CFBAF8622B9A1D8D26F485D3CBFE64" authorityName="Bluthgen" authorityYear="1956" box="[1100,1228,416,437]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="subSpecies" species="bidentatus" subSpecies="puniceus">
ssp.
<emphasis id="72BB1D69622B9A1D8D1DF485D3CBFE64" box="[1143,1228,416,437]" italics="true" pageId="33" pageNumber="184">puniceus</emphasis>
</taxonomicName>
), metanotum entirely black and metasoma with at least five tergites with apical yellow band (only two with ferruginous band in
<taxonomicName id="87CFBAF8622B9A1D8C61F498D28AFE03" authorityName="Bluthgen" authorityYear="1956" box="[1291,1421,445,466]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="subSpecies" species="bidentatus" subSpecies="puniceus">
ssp.
<emphasis id="72BB1D69622B9A1D8C52F498D28AFE03" box="[1336,1421,445,466]" italics="true" pageId="33" pageNumber="184">puniceus</emphasis>
</taxonomicName>
); wings with orange tinge in basal half and slightly infuscate apical half (
<figureCitation id="D8F4DDFE622B9A1D8AF9F4FFD307FE3E" box="[915,1024,474,496]" captionStart="FIGURE 4" captionStartId="13.[151,250,1804,1828]" captionTargetBox="[151,1435,181,1780]" captionTargetId="figure-17@13.[151,1435,181,1780]" captionTargetPageId="13" captionText="FIGURE 4. A, C–D, Euodynerus bidentatus bidentatus (Lepeletier). A, D, female from Morocco. C, male from Morocco. B, E, Euodynerus bidentatus puniceus Blüthgen, female from Libya. F–I, Euodynerus caspicus (Morawitz). F, H, male from Astrakhan Province, Russia. G, I, female from Kazakhstan." figureDoi="http://doi.org/10.5281/zenodo.14239447" httpUri="https://zenodo.org/record/14239447/files/figure.png" pageId="33" pageNumber="184">Figs 4A, B</figureCitation>
). Female: cephalic foveae placed in barely differentiated circular area, narrower than ocellar triangle (cf.
<figureCitation id="D8F4DDFE622B9A1D8A5FF4D2D48CFDDC" box="[821,907,503,525]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15X</figureCitation>
)..................
<taxonomicName id="87CFBAF8622B9A1D8D1AF4DDD29BFDDC" authority="(Lepeletier)" authorityName="MA Tafraoute" baseAuthorityName="Lepeletier" baseAuthorityYear="1841" box="[1136,1436,503,525]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="bidentatus" subGenus="Euodynerus">
<emphasis id="72BB1D69622B9A1D8D1AF4DDD29BFDDC" bold="true" box="[1136,1436,503,525]" pageId="33" pageNumber="184">
<emphasis id="72BB1D69622B9A1D8D1AF4DDD381FDDC" bold="true" box="[1136,1158,504,525]" italics="true" pageId="33" pageNumber="184">E.</emphasis>
(
<emphasis id="72BB1D69622B9A1D8DFEF4DDD3ADFDDC" bold="true" box="[1172,1194,504,525]" italics="true" pageId="33" pageNumber="184">E.</emphasis>
)
<emphasis id="72BB1D69622B9A1D8DD2F4D2D218FDDC" bold="true" box="[1208,1311,503,525]" italics="true" pageId="33" pageNumber="184">bidentatus</emphasis>
(Lepeletier)
</emphasis>
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</keyLead>
</paragraph>
<paragraph id="4070C17B622B9A1D89FDF730D29BFD6C" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF730D29BFD6C" pageId="33" pageNumber="184">
- Disc of clypeus convex, with small shallowly depressed area above apex; apical margin narrower than interantennal space, shallowly but distinctly emarginate (
<figureCitation id="D8F4DDFE622B9A1D8B29F717D59BFD96" box="[579,668,562,584]" captionStart="FIGURE 11" captionStartId="25.[151,250,1151,1175]" captionTargetBox="[151,1435,191,1127]" captionTargetId="figure-17@25.[151,1435,181,1127]" captionTargetPageId="25" captionText="FIGURE 11. A, D, Euodynerus bidentiformis (Giordani Soika), female from Sardinia. B–C, E–G, E. bidentoides (Giordani Soika), sp. resurr. B, E, male from Portugal. C, F, female from Morocco. G, female from Portugal." figureDoi="http://doi.org/10.5281/zenodo.14239461" httpUri="https://zenodo.org/record/14239461/files/figure.png" pageId="33" pageNumber="184">Fig. 11D</figureCitation>
). Punctures on mesosoma larger and sparser and with scattered micropunctures in interspaces, giving shiny appearance. Scutellum entirely black, metanotum with wide yellow band covering its upper half and metasoma with only three (
<specimenCount id="56C90AF2622B9A1D8B7BF748D526FD53" box="[529,545,621,642]" count="1" pageId="33" pageNumber="184" type="female">♀</specimenCount>
) or four (
<specimenCount id="56C90AF2622B9A1D8B14F748D596FD53" box="[638,657,621,642]" count="1" pageId="33" pageNumber="184" type="male">♂</specimenCount>
) apical bands; wings entirely hyaline with subtle grayish-brown tinge (
<figureCitation id="D8F4DDFE622B9A1D8C53F748D296FD53" box="[1337,1425,621,643]" captionStart="FIGURE 11" captionStartId="25.[151,250,1151,1175]" captionTargetBox="[151,1435,191,1127]" captionTargetId="figure-17@25.[151,1435,181,1127]" captionTargetPageId="25" captionText="FIGURE 11. A, D, Euodynerus bidentiformis (Giordani Soika), female from Sardinia. B–C, E–G, E. bidentoides (Giordani Soika), sp. resurr. B, E, male from Portugal. C, F, female from Morocco. G, female from Portugal." figureDoi="http://doi.org/10.5281/zenodo.14239461" httpUri="https://zenodo.org/record/14239461/files/figure.png" pageId="33" pageNumber="184">Fig. 11A</figureCitation>
). Female: cephalic foveae placed in clearly differentiated elliptical area, wider than ocellar triangle (cf.
<figureCitation id="D8F4DDFE622B9A1D8DDEF7AFD237FD71" box="[1204,1328,650,672]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Figs 15U–W</figureCitation>
).............................................................................
<taxonomicName id="87CFBAF8622B9A1D8D48F78DD29BFD6C" authority="(Giordani Soika)" authorityName="IT Tuili" baseAuthorityName="Giordani Soika" baseAuthorityYear="1942" box="[1058,1436,679,701]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="bidentiformis" subGenus="Pareuodynerus">
<emphasis id="72BB1D69622B9A1D8D48F78DD29BFD6C" bold="true" box="[1058,1436,679,701]" pageId="33" pageNumber="184">
<emphasis id="72BB1D69622B9A1D8D48F78DD33FFD6C" bold="true" box="[1058,1080,680,701]" italics="true" pageId="33" pageNumber="184">E.</emphasis>
(
<emphasis id="72BB1D69622B9A1D8D2CF78DD35FFD6C" bold="true" box="[1094,1112,680,701]" italics="true" pageId="33" pageNumber="184">P.</emphasis>
)
<emphasis id="72BB1D69622B9A1D8D0CF782D3EDFD6C" bold="true" box="[1126,1258,679,701]" italics="true" pageId="33" pageNumber="184">bidentiformis</emphasis>
(Giordani Soika)
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</paragraph>
</keyStep>
<keyStep id="FB3BD9DE622B9A1D89FDF7E0D29BFC33" pageId="33" pageNumber="184">
<paragraph id="4070C17B622B9A1D89FDF7E0D29BFC9E" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF7E0D29BFC9E" pageId="33" pageNumber="184">
3. Dorsal carinae of propodeum forming sharp lamellar tooth on medial end, adjacent to disc of metanotum (
<figureCitation id="D8F4DDFE622B9A1D8DACF7E0D219FD0A" box="[1222,1310,709,731]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15G</figureCitation>
). Metanotum with posterior margin of dorsal face finely denticulate and evenly rounded, margin complete or at most narrowly interrupted (
<figureCitation id="D8F4DDFE622B9A1D8983F7DAD646FCC4" box="[233,321,767,789]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15P</figureCitation>
). Male: apical segment of mid and hind tarsi variably darkened but always contrasting with yellowish preceding segments, and variably widened (
<figureCitation id="D8F4DDFE622B9A1D8B42F638D599FCE3" box="[552,670,797,819]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Figs 15R, S</figureCitation>
). Female: cephalic foveae placed in clearly differentiated elliptical area, wider than ocellar triangle (
<figureCitation id="D8F4DDFE622B9A1D88C4F61FD52BFC81" box="[430,556,826,848]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Figs 15U–W</figureCitation>
)....................................................................... 4
</keyLead>
</paragraph>
<paragraph id="4070C17B622B9A1D89FDF67DD29BFC33" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF67DD29BFC33" pageId="33" pageNumber="184">
- Dorsal carinae of propodeum variable, from absent (
<figureCitation id="D8F4DDFE622B9A1D8BB2F672D428FCBC" box="[728,815,855,877]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15H</figureCitation>
) to complete but not forming distinct tooth and widely separated from metanotum (
<figureCitation id="D8F4DDFE622B9A1D88FBF650D6E5FC5A" box="[401,482,885,907]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15I</figureCitation>
). Metanotum with posterior margin of dorsal face coarsely denticulate and laterally raised, margin usually depressed and interrupted in middle (
<figureCitation id="D8F4DDFE622B9A1D8BF6F6B7D5F0FC79" box="[668,759,914,936]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15Q</figureCitation>
). Male: tarsi of all legs entirely ferruginous-yellow and not widened, apical segment not contrasting with preceding ones (
<figureCitation id="D8F4DDFE622B9A1D8BB3F68AD428FC14" box="[729,815,943,965]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15T</figureCitation>
). Female: cephalic foveae placed in barely differentiated circular area, narrower than ocellar triangle (
<figureCitation id="D8F4DDFE622B9A1D8B55F6E8D59FFC32" box="[575,664,973,995]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15X</figureCitation>
), or mixed with surrounding punctures (
<figureCitation id="D8F4DDFE622B9A1D8D71F6E8D375FC32" box="[1051,1138,973,995]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15Y</figureCitation>
)....................... 8
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4. S
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lateral view abruptly convex basally, almost forming short vertical face separated from rest of sternite by shallow fold narrowly interrupted in middle (
<figureCitation id="D8F4DDFE622B9A1D8B7CF122D56BFBCC" box="[534,620,1031,1053]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15B</figureCitation>
).................................................................. 5
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- S
<quantity id="87376C9E622B9A1D8985F100D613FBEA" box="[239,276,1061,1083]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="33" pageNumber="184" unit="in" value="2.0">2 in</quantity>
lateral view evenly convex from base to apex, basal third of sternite somewhat depressed in middle (
<figureCitation id="D8F4DDFE622B9A1D8DB5F100D233FBEA" box="[1247,1332,1061,1083]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15C</figureCitation>
)...... 6
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<keyStep id="FB3BD9DE622B9A1D89FDF167D29BFA8E" pageId="33" pageNumber="184">
<paragraph id="4070C17B622B9A1D89FDF167D29BFB1C" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF167D29BFB1C" pageId="33" pageNumber="184">
5. Gena and vertex strongly developed, in dorsal view gena much longer than dorsal lobe of eye (
<figureCitation id="D8F4DDFE622B9A1D8D07F167D3C0FB89" box="[1133,1223,1090,1112]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15V</figureCitation>
). Apical lamella of T1 shorter, almost linear in middle and barely reaching one puncture diameter on sides; punctures of T2 becoming sparser apically (
<figureCitation id="D8F4DDFE622B9A1D8983F158D65AFB43" box="[233,349,1149,1171]" captionStart="FIGURE 11" captionStartId="25.[151,250,1151,1175]" captionTargetBox="[151,1435,191,1127]" captionTargetId="figure-17@25.[151,1435,181,1127]" captionTargetPageId="25" captionText="FIGURE 11. A, D, Euodynerus bidentiformis (Giordani Soika), female from Sardinia. B–C, E–G, E. bidentoides (Giordani Soika), sp. resurr. B, E, male from Portugal. C, F, female from Morocco. G, female from Portugal." figureDoi="http://doi.org/10.5281/zenodo.14239461" httpUri="https://zenodo.org/record/14239461/files/figure.png" pageId="33" pageNumber="184">Figs 11B, C</figureCitation>
). Vestiture of head and mesosoma, including base of legs, longer and wavier, reaching up to three ocellar diameters in length (cf.
<figureCitation id="D8F4DDFE622B9A1D8809F1BFD6B4FB61" box="[355,435,1178,1200]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15J</figureCitation>
). Light markings extensive and bright yellow, with at least four bands on tergites (
<figureCitation id="D8F4DDFE622B9A1D8DA3F1BFD239FB7E" box="[1225,1342,1178,1200]" captionStart="FIGURE 11" captionStartId="25.[151,250,1151,1175]" captionTargetBox="[151,1435,191,1127]" captionTargetId="figure-17@25.[151,1435,181,1127]" captionTargetPageId="25" captionText="FIGURE 11. A, D, Euodynerus bidentiformis (Giordani Soika), female from Sardinia. B–C, E–G, E. bidentoides (Giordani Soika), sp. resurr. B, E, male from Portugal. C, F, female from Morocco. G, female from Portugal." figureDoi="http://doi.org/10.5281/zenodo.14239461" httpUri="https://zenodo.org/record/14239461/files/figure.png" pageId="33" pageNumber="184">Figs 11B, C</figureCitation>
).............................................................................
<taxonomicName id="87CFBAF8622B9A1D8D5CF19DD29BFB1C" authority="(Giordani Soika)" baseAuthorityName="Giordani Soika" box="[1078,1436,1207,1229]" class="Insecta" family="Vespidae" genus="Odynerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="bidentoides" subGenus="Pareuodynerus">
<emphasis id="72BB1D69622B9A1D8D5CF19DD29BFB1C" bold="true" box="[1078,1436,1207,1229]" pageId="33" pageNumber="184">
<emphasis id="72BB1D69622B9A1D8D5CF19DD34BFB1C" bold="true" box="[1078,1100,1208,1229]" italics="true" pageId="33" pageNumber="184">E.</emphasis>
(
<emphasis id="72BB1D69622B9A1D8D30F19DD36BFB1C" bold="true" box="[1114,1132,1208,1229]" italics="true" pageId="33" pageNumber="184">P.</emphasis>
)
<emphasis id="72BB1D69622B9A1D8D10F192D3EDFB1C" bold="true" box="[1146,1258,1207,1229]" italics="true" pageId="33" pageNumber="184">bidentoides</emphasis>
(Giordani Soika)
</emphasis>
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</paragraph>
<paragraph id="4070C17B622B9A1D89FDF1F0D29BFA8E" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF1F0D29BFA8E" pageId="33" pageNumber="184">
- Gena and vertex normally developed, in dorsal view gena as long as dorsal lobe of eye (
<figureCitation id="D8F4DDFE622B9A1D8D49F1F0D385FB3A" box="[1059,1154,1237,1259]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15W</figureCitation>
). Apical lamella of T1 longer, well visible in middle and exceeding one puncture diameter on sides; punctures of T2 forming denser preapical band (
<figureCitation id="D8F4DDFE622B9A1D8C18F1D7D637FAF4" captionStart="FIGURE 13" captionStartId="29.[151,250,1873,1897]" captionTargetBox="[226,1361,188,1849]" captionTargetId="figure-17@29.[226,1361,181,1849]" captionTargetPageId="29" captionText="FIGURE 13. A–J, Euodynerus quadrifasciatus (Fabricius). A, female from Estonia. B, H, female from Italy. C, female from Crimea, Russia. D, F, male from China (E. quadrifasciatus atripes Giordani Soika, syn. nov.). E, male from Italy. G, female from Khabarovsk Territory, Russia. I–J, male from Turkey (paratype of E.quadrifasciatus rufipes Gusenleitner, syn. nov.). K–N, E. rubrosignatus Gusenleitner, stat. nov., female from Libya. K–L, paratype of E. quadrifasciatus rubrosignatus Gusenleitner. M–N, holotype of E. notatus cyrenaicus Giordani Soika, syn. nov." figureDoi="http://doi.org/10.5281/zenodo.14239465" httpUri="https://zenodo.org/record/14239465/files/figure.png" pageId="33" pageNumber="184">Figs 13K, M</figureCitation>
). Vestiture of head and mesosoma shorter and stiffer, not reaching two ocellar diameters in length (cf.
<figureCitation id="D8F4DDFE622B9A1D8C68F02AD250FAF4" box="[1282,1367,1295,1317]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15L</figureCitation>
). Light markings reduced and orange-red, with only three bands on tergites (third band laterally abbreviated) (
<figureCitation id="D8F4DDFE622B9A1D8DD2F008D232FA93" box="[1208,1333,1325,1347]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Figs 15K, M</figureCitation>
)..............................................................................
<taxonomicName id="87CFBAF8622B9A1D8D2EF06FD29BFA8E" authority="Gusenleitner" authorityName="Gusenleitner." authorityYear="1984" baseAuthorityName="Gusenleitner. M" baseAuthorityYear="1984" box="[1092,1436,1354,1376]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="rubrosignatus" subGenus="Pareuodynerus">
<emphasis id="72BB1D69622B9A1D8D2EF06FD29BFA8E" bold="true" box="[1092,1436,1354,1376]" pageId="33" pageNumber="184">
<emphasis id="72BB1D69622B9A1D8D2EF06FD35DFA8E" bold="true" box="[1092,1114,1354,1375]" italics="true" pageId="33" pageNumber="184">E.</emphasis>
(
<emphasis id="72BB1D69622B9A1D8D02F06FD37DFA8E" bold="true" box="[1128,1146,1354,1375]" italics="true" pageId="33" pageNumber="184">P.</emphasis>
)
<emphasis id="72BB1D69622B9A1D8DEDF06FD215FAB1" bold="true" box="[1159,1298,1354,1376]" italics="true" pageId="33" pageNumber="184">rubrosignatus</emphasis>
Gusenleitner
</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="FB3BD9DE622B9A1D89FDF042D29BFA69" pageId="33" pageNumber="184">
<paragraph id="4070C17B622B9A1D89FDF042D29BFA4B" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF042D29BFA4B" pageId="33" pageNumber="184">
6. Vestiture of mesosoma long and wavy, exceeding two ocellar diameters in length (
<figureCitation id="D8F4DDFE622B9A1D8A92F042D348FAAC" box="[1016,1103,1383,1405]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15K</figureCitation>
)...................................................................................................
<taxonomicName id="87CFBAF8622B9A1D8D22F0A0D29BFA4B" authority="(Fabricius)" authorityName="RU Tsey" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1096,1436,1412,1434]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="quadrifasciatus" subGenus="Pareuodynerus">
<emphasis id="72BB1D69622B9A1D8D22F0A0D29BFA4B" bold="true" box="[1096,1436,1412,1434]" pageId="33" pageNumber="184">
<emphasis id="72BB1D69622B9A1D8D22F0A0D359FA4B" bold="true" box="[1096,1118,1413,1434]" italics="true" pageId="33" pageNumber="184">E.</emphasis>
(
<emphasis id="72BB1D69622B9A1D8D06F0A0D379FA4B" bold="true" box="[1132,1150,1413,1434]" italics="true" pageId="33" pageNumber="184">P.</emphasis>
)
<emphasis id="72BB1D69622B9A1D8DE6F0A1D222FA4B" bold="true" box="[1164,1317,1412,1434]" italics="true" pageId="33" pageNumber="184">quadrifasciatus</emphasis>
(Fabricius)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4070C17B622B9A1D89FDF086D29BFA69" blockId="33.[151,1437,151,1992]" box="[151,1436,1442,1464]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF086D29BFA69" box="[151,1436,1442,1464]" pageId="33" pageNumber="184">
- Vestiture of mesosoma short and straight, brush-like, not exceeding one ocellar diameter in length (
<figureCitation id="D8F4DDFE622B9A1D8DF0F087D3E9FA69" box="[1178,1262,1442,1464]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15L</figureCitation>
)............ 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FB3BD9DE622B9A1D89FDF0E5D29BF92B" pageId="33" pageNumber="184">
<paragraph id="4070C17B622B9A1D89FDF0E5D29AF99B" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF0E5D29AF99B" pageId="33" pageNumber="184">
7. Clypeus more finely and sparsely punctate, interspaces on disc always exceeding puncture diameter in length (
<figureCitation id="D8F4DDFE622B9A1D8C7CF09AD294FA04" box="[1302,1427,1471,1493]" captionStart="FIGURE 12" captionStartId="27.[151,250,1887,1911]" captionTargetBox="[151,1435,189,1863]" captionTargetId="figure-17@27.[151,1435,181,1863]" captionTargetPageId="27" captionText="FIGURE 12. A–G, Euodynerus notatus (Jurine). A, E, male from Hungary. B, F, female from Altai, Russia. C, G, female from Krasnodar Territory, Russia. D, male from Nizhny Novgorod Province, Russia. H–K, E. posticus (Herrich-Schaeffer). H, J, male from Italy. I, female from Crimea, Russia. K, female from Italy." figureDoi="http://doi.org/10.5281/zenodo.14239463" httpUri="https://zenodo.org/record/14239463/files/figure.png" pageId="33" pageNumber="184">Figs 12D–G</figureCitation>
). Pronotal carina weakly lamellate. Propodeum always entirely black, tibiae usually marked with black at least on inner face. Male: clypeus more convex basally than apically, with long free apical part and clearly emarginate apical margin (
<figureCitation id="D8F4DDFE622B9A1D8C71F0DFD296F9DE" box="[1307,1425,1530,1552]" captionStart="FIGURE 12" captionStartId="27.[151,250,1887,1911]" captionTargetBox="[151,1435,189,1863]" captionTargetId="figure-17@27.[151,1435,181,1863]" captionTargetPageId="27" captionText="FIGURE 12. A–G, Euodynerus notatus (Jurine). A, E, male from Hungary. B, F, female from Altai, Russia. C, G, female from Krasnodar Territory, Russia. D, male from Nizhny Novgorod Province, Russia. H–K, E. posticus (Herrich-Schaeffer). H, J, male from Italy. I, female from Crimea, Russia. K, female from Italy." figureDoi="http://doi.org/10.5281/zenodo.14239463" httpUri="https://zenodo.org/record/14239463/files/figure.png" pageId="33" pageNumber="184">Figs 12D, E</figureCitation>
); mandible with basalmost tooth similar to other teeth, outer surface mostly black with yellow basal triangle (
<figureCitation id="D8F4DDFE622B9A1D8D9DF332D256F9FC" box="[1271,1361,1559,1581]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="33" pageNumber="184">Fig. 16A</figureCitation>
); apical segment of mid and hind tarsi black and distinctly widened (
<figureCitation id="D8F4DDFE622B9A1D8A4CF310D47BF99A" box="[806,892,1589,1611]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15R</figureCitation>
).........................
<taxonomicName id="87CFBAF8622B9A1D8DDDF310D29AF99B" authority="(Jurine)" authorityName="RU Sochi" baseAuthorityName="Jurine" baseAuthorityYear="1807" box="[1207,1437,1588,1610]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="notatus" subGenus="Pareuodynerus">
<emphasis id="72BB1D69622B9A1D8DDDF310D29AF99B" bold="true" box="[1207,1437,1588,1610]" pageId="33" pageNumber="184">
<emphasis id="72BB1D69622B9A1D8DDDF310D3CAF99B" bold="true" box="[1207,1229,1589,1610]" italics="true" pageId="33" pageNumber="184">E.</emphasis>
(
<emphasis id="72BB1D69622B9A1D8DB1F310D3EAF99B" bold="true" box="[1243,1261,1589,1610]" italics="true" pageId="33" pageNumber="184">P.</emphasis>
)
<emphasis id="72BB1D69622B9A1D8D90F310D244F99B" bold="true" box="[1274,1347,1589,1610]" italics="true" pageId="33" pageNumber="184">notatus</emphasis>
(Jurine)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4070C17B622B9A1D89FDF376D29BF92B" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF376D29BF92B" pageId="33" pageNumber="184">
- Clypeus more coarsely and densely punctate, interspaces on disc mostly not exceeding puncture diameter in length (
<figureCitation id="D8F4DDFE622B9A1D8C2EF377D7F1F954" captionStart="FIGURE 12" captionStartId="27.[151,250,1887,1911]" captionTargetBox="[151,1435,189,1863]" captionTargetId="figure-17@27.[151,1435,181,1863]" captionTargetPageId="27" captionText="FIGURE 12. A–G, Euodynerus notatus (Jurine). A, E, male from Hungary. B, F, female from Altai, Russia. C, G, female from Krasnodar Territory, Russia. D, male from Nizhny Novgorod Province, Russia. H–K, E. posticus (Herrich-Schaeffer). H, J, male from Italy. I, female from Crimea, Russia. K, female from Italy." figureDoi="http://doi.org/10.5281/zenodo.14239463" httpUri="https://zenodo.org/record/14239463/files/figure.png" pageId="33" pageNumber="184">Figs 12J, K</figureCitation>
). Pronotal carina strongly lamellate. Propodeum usually largely marked with yellow (yellow markings reduced in northeastern populations), tibiae entirely yellow with ferruginous inner face. Male: clypeus evenly convex from base to apex, with very short free apical part and subtruncate apical margin (
<figureCitation id="D8F4DDFE622B9A1D8BB6F38FD42AF96E" box="[732,813,1706,1728]" captionStart="FIGURE 12" captionStartId="27.[151,250,1887,1911]" captionTargetBox="[151,1435,189,1863]" captionTargetId="figure-17@27.[151,1435,181,1863]" captionTargetPageId="27" captionText="FIGURE 12. A–G, Euodynerus notatus (Jurine). A, E, male from Hungary. B, F, female from Altai, Russia. C, G, female from Krasnodar Territory, Russia. D, male from Nizhny Novgorod Province, Russia. H–K, E. posticus (Herrich-Schaeffer). H, J, male from Italy. I, female from Crimea, Russia. K, female from Italy." figureDoi="http://doi.org/10.5281/zenodo.14239463" httpUri="https://zenodo.org/record/14239463/files/figure.png" pageId="33" pageNumber="184">Fig. 12J</figureCitation>
); mandible with basalmost tooth larger and perpendicular to axis of other teeth, outer surface almost entirely yellow (
<figureCitation id="D8F4DDFE622B9A1D8B88F3E2D43BF90C" box="[738,828,1735,1757]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="33" pageNumber="184">Fig. 16B</figureCitation>
); apical segment of mid and hind tarsi reddish-brown and not markedly widened (
<figureCitation id="D8F4DDFE622B9A1D88CBF3C0D6F4F92A" box="[417,499,1765,1787]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15S</figureCitation>
)...............................................
<taxonomicName id="87CFBAF8622B9A1D8D52F3C0D29BF92B" authority="(Herrich-Schaeffer)" authorityName="RU Nikita" baseAuthorityName="Herrich-Schaeffer" baseAuthorityYear="1841" box="[1080,1436,1764,1786]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="posticus" subGenus="Pareuodynerus">
<emphasis id="72BB1D69622B9A1D8D52F3C0D29BF92B" bold="true" box="[1080,1436,1764,1786]" pageId="33" pageNumber="184">
<emphasis id="72BB1D69622B9A1D8D52F3C0D349F92B" bold="true" box="[1080,1102,1765,1786]" italics="true" pageId="33" pageNumber="184">E.</emphasis>
(
<emphasis id="72BB1D69622B9A1D8D36F3C0D369F92B" bold="true" box="[1116,1134,1765,1786]" italics="true" pageId="33" pageNumber="184">P.</emphasis>
)
<emphasis id="72BB1D69622B9A1D8D16F3C0D3CBF92B" bold="true" box="[1148,1228,1765,1786]" italics="true" pageId="33" pageNumber="184">posticus</emphasis>
(Herrich-Schaeffer)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FB3BD9DE622B9A1D89FDF227D29BF816" pageId="33" pageNumber="184">
<paragraph id="4070C17B622B9A1D89FDF227D29BF8A1" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF227D29BF8A1" pageId="33" pageNumber="184">
8. S2 step-like in lateral view, with basal vertical face sharply delimited by bulging transverse ridge widely interrupted in the middle; surface of S2 shallowly but distinctly depressed just behind transverse ridge (
<figureCitation id="D8F4DDFE622B9A1D8D40F23AD382F8E4" box="[1066,1157,1823,1845]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15D</figureCitation>
). Tegula with scattered deep punctures on whole surface and strongly pointed posterior lobe (
<figureCitation id="D8F4DDFE622B9A1D8A34F218D4BEF882" box="[862,953,1853,1875]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="33" pageNumber="184">Fig. 16C</figureCitation>
). Posterior horizontal face of T1 entirely orangeyellow, rarely with barely visible median dark line (
<figureCitation id="D8F4DDFE622B9A1D8BB8F27FD43FF8BE" box="[722,824,1882,1904]" captionStart="FIGURE 4" captionStartId="13.[151,250,1804,1828]" captionTargetBox="[151,1435,181,1780]" captionTargetId="figure-17@13.[151,1435,181,1780]" captionTargetPageId="13" captionText="FIGURE 4. A, C–D, Euodynerus bidentatus bidentatus (Lepeletier). A, D, female from Morocco. C, male from Morocco. B, E, Euodynerus bidentatus puniceus Blüthgen, female from Libya. F–I, Euodynerus caspicus (Morawitz). F, H, male from Astrakhan Province, Russia. G, I, female from Kazakhstan." figureDoi="http://doi.org/10.5281/zenodo.14239447" httpUri="https://zenodo.org/record/14239447/files/figure.png" pageId="33" pageNumber="184">Figs 4F, G</figureCitation>
)...........................
<taxonomicName id="87CFBAF8622B9A1D8DECF27FD29BF8A1" authority="(Morawitz)" authorityName="RU Zeleny Sad" baseAuthorityName="Morawitz" baseAuthorityYear="1873" box="[1158,1436,1882,1904]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="caspicus" subGenus="Euodynerus">
<emphasis id="72BB1D69622B9A1D8DECF27FD29BF8A1" bold="true" box="[1158,1436,1882,1904]" pageId="33" pageNumber="184">
<emphasis id="72BB1D69622B9A1D8DECF27FD39BF8BE" bold="true" box="[1158,1180,1882,1903]" italics="true" pageId="33" pageNumber="184">E.</emphasis>
(
<emphasis id="72BB1D69622B9A1D8DC0F27FD3C7F8BE" bold="true" box="[1194,1216,1882,1903]" italics="true" pageId="33" pageNumber="184">E.</emphasis>
)
<emphasis id="72BB1D69622B9A1D8DA4F27FD225F8BE" bold="true" box="[1230,1314,1882,1903]" italics="true" pageId="33" pageNumber="184">caspicus</emphasis>
(Morawitz)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4070C17B622B9A1D89FDF25DD29BF816" blockId="33.[151,1437,151,1992]" pageId="33" pageNumber="184">
<keyLead id="FB3E624E622B9A1D89FDF25DD29BF816" pageId="33" pageNumber="184">
- S2 more or less evenly convex in lateral view, without basal transverse ridge; surface of S2 not depressed (
<figureCitation id="D8F4DDFE622B9A1D8D98F252D24DF85C" box="[1266,1354,1911,1933]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="33" pageNumber="184">Fig. 15E</figureCitation>
). Tegula impunctate or with coarser punctures and less pointed posterior lobe (
<figureCitation id="D8F4DDFE622B9A1D8A1DF2B0D4F5F87A" box="[887,1010,1941,1963]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="33" pageNumber="184">Figs 16D–G</figureCitation>
). Posterior horizontal face of T1 at least with well-developed longitudinal black band, usually with semicircular or pentagonal large black area basally................ 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FB3BD9DE62289A1E89FDF5BDD29BFEAB" pageId="34" pageNumber="185">
<paragraph id="4070C17B62289A1E89FDF5BDD29BFF36" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF5BDD29BFF36" pageId="34" pageNumber="185">
9. Posterior lobe of tegula with dense coarse punctures, usually deep but still well-evident even when shallow (
<figureCitation id="D8F4DDFE62289A1E8C7FF5B2D295FF7C" box="[1301,1426,151,173]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Figs 16D, E</figureCitation>
). Dorsal side of mesosoma with very sparse and fine dust-like pubescence, almost bare (
<figureCitation id="D8F4DDFE62289A1E8D50F590D39EFF1A" box="[1082,1177,181,203]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Fig. 15M</figureCitation>
). Clypeus with coarse and dense longitudinal striae in female, with finer striae and elongate punctures in male (
<figureCitation id="D8F4DDFE62289A1E8D64F5F7D375FF39" box="[1038,1138,210,232]" captionStart="FIGURE 6" captionStartId="17.[151,250,1873,1897]" captionTargetBox="[221,1366,193,1849]" captionTargetId="figure-17@17.[221,1366,181,1849]" captionTargetPageId="17" captionText="FIGURE 6. Euodynerus dantici (Rossi). A, K, female from Italy (E. dantici s. str.). B, L, female from Montecristo, Italy (E. dantici poggii Giordani Soika, syn. nov.). C, M, female from Malta. D, J, male from Menorca, Spain (E. minoricensis Sanza, syn. nov.). E, female from Tadjikistan (E. dantici pamiricus Blüthgen). F, N, female from Buryatia, Russia (E. dantici brachytomus (Kostylev)). G, O, female from China (E. dantici violaceipennis Giordani Soika). H, female from Laos. I, male from Greece." figureDoi="http://doi.org/10.5281/zenodo.14239451" httpUri="https://zenodo.org/record/14239451/files/figure.png" pageId="34" pageNumber="185">Figs 6I–O</figureCitation>
;
<figureCitation id="D8F4DDFE62289A1E8D16F5F7D3B3FF36" box="[1148,1204,210,231]" captionStart="FIGURE 8" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[200,1386,192,1918]" captionTargetId="figure-20@20.[200,1386,181,1918]" captionTargetPageId="20" captionText="FIGURE 8. A–F, Euodynerus hellenicus Blüthgen. A, male from Bulgaria. B, E, female from Spain. C, F, female from Iran. D, male from Spain. G–M, Euodynerus semisaecularis (Dalla Torre). G, J, male from Greece. H, L, female from Crete, Greece. I, M, female from Afghanistan. K, male from Armenia." figureDoi="http://doi.org/10.5281/zenodo.14239455" httpUri="https://zenodo.org/record/14239455/files/figure.png" pageId="34" pageNumber="185">8D–F</figureCitation>
)................ 10
</keyLead>
</paragraph>
<paragraph id="4070C17B62289A1E89FDF5D5D29BFEAB" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF5D5D29BFEAB" pageId="34" pageNumber="185">
- Posterior lobe of tegula smooth or with few scattered fine punctures, visible only under incident light (
<figureCitation id="D8F4DDFE62289A1E8DAAF5CAD231FED4" box="[1216,1334,239,261]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Figs 16F, G</figureCitation>
). Vestiture of dorsal side of mesosoma variable, dust-like and sparse (cf.
<figureCitation id="D8F4DDFE62289A1E8A59F428D488FEF2" box="[819,911,269,291]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Fig. 15M</figureCitation>
), long and brush-like (
<figureCitation id="D8F4DDFE62289A1E8D03F428D3B8FEF2" box="[1129,1215,269,291]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Fig. 15N</figureCitation>
) or with scattered bent setae (
<figureCitation id="D8F4DDFE62289A1E8875F40FD67FFE91" box="[287,376,298,320]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Fig. 15O</figureCitation>
). Clypeus with fine and sparse longitudinal striae intermixed with punctures in female (in
<taxonomicName id="87CFBAF862289A1E8DB2F40ED26CFEEE" authorityName="UZ Uchkulach" baseAuthorityName="Lichtenstein" baseAuthorityYear="1884" box="[1240,1387,298,320]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="disconotatus">
<emphasis id="72BB1D6962289A1E8DB2F40ED26CFEEE" box="[1240,1387,298,320]" italics="true" pageId="34" pageNumber="185">E. disconotatus</emphasis>
</taxonomicName>
with some scattered coarser striae), without striae and with rounded punctures in male (
<figureCitation id="D8F4DDFE62289A1E8A91F462D36DFE8C" box="[1019,1130,327,349]" captionStart="FIGURE 5" captionStartId="15.[151,250,1495,1519]" captionTargetBox="[151,1435,639,1472]" captionTargetId="figure-221@15.[151,1435,633,1472]" captionTargetPageId="15" captionText="FIGURE 5. Euodynerus curictensis Blüthgen. A, E, male from Morocco. B, F, female from Greece. C, H, female from Sardinia (E. curictensis sardous Borsato, syn. nov.). D, male from Lebanon. G, female from Morocco." figureDoi="http://doi.org/10.5281/zenodo.14239449" httpUri="https://zenodo.org/record/14239449/files/figure.png" pageId="34" pageNumber="185">Figs 5D–H</figureCitation>
;
<figureCitation id="D8F4DDFE62289A1E8D19F46DD3F4FE8C" box="[1139,1267,328,349]" captionStart="FIGURE 7" captionStartId="19.[151,250,1908,1932]" captionTargetBox="[210,1376,183,1884]" captionTargetId="figure-17@19.[210,1376,181,1884]" captionTargetPageId="19" captionText="FIGURE 7. A–H, Euodynerus disconotatus (Lichtenstein). A, male from Azerbaijan. B, E, male from Italy. C, G, female from Italy. D, female from Greece. F, female from Sicily, Italy. H, female from Azerbaijan. I–O, Euodynerus fastidiosus (de Saussure). I, L, male from Italy. J, male from Greece. K, female from Crete, Greece. M, female from Italy. N, female from Crimea, Russia. O, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239453" httpUri="https://zenodo.org/record/14239453/files/figure.png" pageId="34" pageNumber="185">7E–H; 7L–O</figureCitation>
;
<figureCitation id="D8F4DDFE62289A1E8D97F46DD231FE8C" box="[1277,1334,328,349]" captionStart="FIGURE 8" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[200,1386,192,1918]" captionTargetId="figure-20@20.[200,1386,181,1918]" captionTargetPageId="20" captionText="FIGURE 8. A–F, Euodynerus hellenicus Blüthgen. A, male from Bulgaria. B, E, female from Spain. C, F, female from Iran. D, male from Spain. G–M, Euodynerus semisaecularis (Dalla Torre). G, J, male from Greece. H, L, female from Crete, Greece. I, M, female from Afghanistan. K, male from Armenia." figureDoi="http://doi.org/10.5281/zenodo.14239455" httpUri="https://zenodo.org/record/14239455/files/figure.png" pageId="34" pageNumber="185">8J–M</figureCitation>
;
<figureCitation id="D8F4DDFE62289A1E8C2BF46DD7FEFEAB" captionStart="FIGURE 9" captionStartId="22.[151,250,1908,1932]" captionTargetBox="[177,1411,192,1883]" captionTargetId="figure-20@22.[177,1411,181,1883]" captionTargetPageId="22" captionText="FIGURE 9. A–H, Euodynerus variegatus (Fabricius). A, E–F, female from Morocco. B, H, female from Egypt (E. variegatus kruegeri (von Schulthess), syn. nov.). C, G, female from Israel (E. variegatus kruegeri (von Schulthess), syn. nov.). D, male from Sardinia, Italy. I–M, Euodynerus velutinus Blüthgen. I, male from Azerbaijan, J, L, male from Greece. K, female from Greece. M, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239457" httpUri="https://zenodo.org/record/14239457/files/figure.png" pageId="34" pageNumber="185">9D–H, L, M</figureCitation>
)................................................................................................ 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FB3BD9DE62289A1E89FDF4A7D29AFDB4" pageId="34" pageNumber="185">
<paragraph id="4070C17B62289A1E89FDF4A7D29BFE3E" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF4A7D29BFE3E" pageId="34" pageNumber="185">
10. Apical margin of clypeus subtruncate (
<figureCitation id="D8F4DDFE62289A1E8B31F4A7D5C0FE46" box="[603,711,386,408]" captionStart="FIGURE 8" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[200,1386,192,1918]" captionTargetId="figure-20@20.[200,1386,181,1918]" captionTargetPageId="20" captionText="FIGURE 8. A–F, Euodynerus hellenicus Blüthgen. A, male from Bulgaria. B, E, female from Spain. C, F, female from Iran. D, male from Spain. G–M, Euodynerus semisaecularis (Dalla Torre). G, J, male from Greece. H, L, female from Crete, Greece. I, M, female from Afghanistan. K, male from Armenia." figureDoi="http://doi.org/10.5281/zenodo.14239455" httpUri="https://zenodo.org/record/14239455/files/figure.png" pageId="34" pageNumber="185">Figs 8D–F</figureCitation>
). Outer margin of tegula distinctly bent in posterior half, becoming almost perpendicular to longitudinal axis of body (
<figureCitation id="D8F4DDFE62289A1E8BFAF4BAD5EBFE64" box="[656,748,415,437]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Fig. 16D</figureCitation>
). Apical translucent margin of T2 longer and shallowly reflexed (
<figureCitation id="D8F4DDFE62289A1E8C1FF4BAD608FE02" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Fig. 16H</figureCitation>
). Female: clypeus as wide as long (
<figureCitation id="D8F4DDFE62289A1E8B34F498D5C2FE03" box="[606,709,445,467]" captionStart="FIGURE 8" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[200,1386,192,1918]" captionTargetId="figure-20@20.[200,1386,181,1918]" captionTargetPageId="20" captionText="FIGURE 8. A–F, Euodynerus hellenicus Blüthgen. A, male from Bulgaria. B, E, female from Spain. C, F, female from Iran. D, male from Spain. G–M, Euodynerus semisaecularis (Dalla Torre). G, J, male from Greece. H, L, female from Crete, Greece. I, M, female from Afghanistan. K, male from Armenia." figureDoi="http://doi.org/10.5281/zenodo.14239455" httpUri="https://zenodo.org/record/14239455/files/figure.png" pageId="34" pageNumber="185">Figs 8E, F</figureCitation>
); F1 and part of F2 usually reddish; ventral side of metasoma largely yellow with reduced black markings......................................................
<taxonomicName id="87CFBAF862289A1E8DE6F4FFD29BFE3E" authority="Bluthgen" authorityName="Bluthgen. A" authorityYear="1942" box="[1164,1436,474,496]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="hellenicus" subGenus="Euodynerus">
<emphasis id="72BB1D6962289A1E8DE6F4FFD29BFE3E" bold="true" box="[1164,1436,474,496]" pageId="34" pageNumber="185">
<emphasis id="72BB1D6962289A1E8DE6F4FFD3A5FE3E" bold="true" box="[1164,1186,474,495]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
(
<emphasis id="72BB1D6962289A1E8DDAF4FFD3C1FE3E" bold="true" box="[1200,1222,474,495]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
)
<emphasis id="72BB1D6962289A1E8DBEF4FFD23EFE21" bold="true" box="[1236,1337,474,496]" italics="true" pageId="34" pageNumber="185">hellenicus</emphasis>
Blüthgen
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4070C17B62289A1E89FDF4DDD29AFDB4" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF4DDD29AFDB4" pageId="34" pageNumber="185">
- Apical margin of clypeus shallowly but distinctly emarginate (
<figureCitation id="D8F4DDFE62289A1E8A51F4D2D4A7FDDC" box="[827,928,503,525]" captionStart="FIGURE 6" captionStartId="17.[151,250,1873,1897]" captionTargetBox="[221,1366,193,1849]" captionTargetId="figure-17@17.[221,1366,181,1849]" captionTargetPageId="17" captionText="FIGURE 6. Euodynerus dantici (Rossi). A, K, female from Italy (E. dantici s. str.). B, L, female from Montecristo, Italy (E. dantici poggii Giordani Soika, syn. nov.). C, M, female from Malta. D, J, male from Menorca, Spain (E. minoricensis Sanza, syn. nov.). E, female from Tadjikistan (E. dantici pamiricus Blüthgen). F, N, female from Buryatia, Russia (E. dantici brachytomus (Kostylev)). G, O, female from China (E. dantici violaceipennis Giordani Soika). H, female from Laos. I, male from Greece." figureDoi="http://doi.org/10.5281/zenodo.14239451" httpUri="https://zenodo.org/record/14239451/files/figure.png" pageId="34" pageNumber="185">Figs 6I–O</figureCitation>
). Outer margin of tegula evenly rounded in posterior half (
<figureCitation id="D8F4DDFE62289A1E8879F730D66EFDFA" box="[275,361,533,555]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Fig. 16E</figureCitation>
). Apical translucent margin of T2 very narrow and linear, barely developed (
<figureCitation id="D8F4DDFE62289A1E8D52F730D383FDFA" box="[1080,1156,533,555]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Fig. 16I</figureCitation>
) (rarely longer and shallowly reflexed in male specimens). Female: clypeus longer than wide (
<figureCitation id="D8F4DDFE62289A1E8A27F717D4BBFD99" box="[845,956,562,584]" captionStart="FIGURE 6" captionStartId="17.[151,250,1873,1897]" captionTargetBox="[221,1366,193,1849]" captionTargetId="figure-17@17.[221,1366,181,1849]" captionTargetPageId="17" captionText="FIGURE 6. Euodynerus dantici (Rossi). A, K, female from Italy (E. dantici s. str.). B, L, female from Montecristo, Italy (E. dantici poggii Giordani Soika, syn. nov.). C, M, female from Malta. D, J, male from Menorca, Spain (E. minoricensis Sanza, syn. nov.). E, female from Tadjikistan (E. dantici pamiricus Blüthgen). F, N, female from Buryatia, Russia (E. dantici brachytomus (Kostylev)). G, O, female from China (E. dantici violaceipennis Giordani Soika). H, female from Laos. I, male from Greece." figureDoi="http://doi.org/10.5281/zenodo.14239451" httpUri="https://zenodo.org/record/14239451/files/figure.png" pageId="34" pageNumber="185">Figs 6K–O</figureCitation>
); flagellum entirely black dorsally; ventral side of metasoma largely black with lateral yellow spots...........................................
<taxonomicName id="87CFBAF862289A1E8DAEF775D29AFDB4" authority="(Rossi)" authorityName="UZ Yangikishlak" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[1220,1437,591,613]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="dantici" subGenus="Euodynerus">
<emphasis id="72BB1D6962289A1E8DAEF775D29AFDB4" bold="true" box="[1220,1437,591,613]" pageId="34" pageNumber="185">
<emphasis id="72BB1D6962289A1E8DAEF775D3DDFDB4" bold="true" box="[1220,1242,592,613]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
(
<emphasis id="72BB1D6962289A1E8D82F775D3F9FDB4" bold="true" box="[1256,1278,592,613]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
)
<emphasis id="72BB1D6962289A1E8C66F76AD257FDB4" bold="true" box="[1292,1360,591,613]" italics="true" pageId="34" pageNumber="185">dantici</emphasis>
(Rossi)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FB3BD9DE62289A1E89FDF748D29BFD6C" pageId="34" pageNumber="185">
<paragraph id="4070C17B62289A1E89FDF748D29BFD53" blockId="34.[151,1437,151,1669]" box="[151,1436,621,643]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF748D29BFD53" box="[151,1436,621,643]" pageId="34" pageNumber="185">
11. Dorsal side of mesosoma with dense erect setae, brush-like (
<figureCitation id="D8F4DDFE62289A1E8A4CF748D47AFD52" box="[806,893,621,643]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Fig. 15N</figureCitation>
).......................................... 12
</keyLead>
</paragraph>
<paragraph id="4070C17B62289A1E89FDF7AED29BFD6C" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF7AED29BFD6C" pageId="34" pageNumber="185">
- Dorsal side of mesosoma with dust-like pubescence (cf.
<figureCitation id="D8F4DDFE62289A1E8A63F7AFD462FD71" box="[777,869,650,672]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Fig. 15M</figureCitation>
), only in
<taxonomicName id="87CFBAF862289A1E8AAEF7AED369FD4E" box="[964,1134,650,672]" class="Insecta" family="Vespidae" genus="Odynerus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="semisaecularis">
<emphasis id="72BB1D6962289A1E8AAEF7AED369FD4E" box="[964,1134,650,672]" italics="true" pageId="34" pageNumber="185">E. semisaecularis</emphasis>
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with sparse apically bent erect setae (
<figureCitation id="D8F4DDFE62289A1E8875F782D671FD6C" box="[287,374,679,701]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Fig. 15O</figureCitation>
)..................................................................................... 13
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<keyStep id="FB3BD9DE62289A1E89FDF7E0D29BFC76" pageId="34" pageNumber="185">
<paragraph id="4070C17B62289A1E89FDF7E0D29BFCE3" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF7E0D29BFCE3" pageId="34" pageNumber="185">
12. Punctures on mesepisternum arranged in longitudinal series, with interspaces forming irregular parallel striae. Male: apical margin of clypeus with deep semi-circular emargination, apical teeth acute and their distance equal to interantennal space (
<figureCitation id="D8F4DDFE62289A1E8C1FF7C7D7F8FCC4" captionStart="FIGURE 9" captionStartId="22.[151,250,1908,1932]" captionTargetBox="[177,1411,192,1883]" captionTargetId="figure-20@22.[177,1411,181,1883]" captionTargetPageId="22" captionText="FIGURE 9. A–H, Euodynerus variegatus (Fabricius). A, E–F, female from Morocco. B, H, female from Egypt (E. variegatus kruegeri (von Schulthess), syn. nov.). C, G, female from Israel (E. variegatus kruegeri (von Schulthess), syn. nov.). D, male from Sardinia, Italy. I–M, Euodynerus velutinus Blüthgen. I, male from Azerbaijan, J, L, male from Greece. K, female from Greece. M, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239457" httpUri="https://zenodo.org/record/14239457/files/figure.png" pageId="34" pageNumber="185">Fig. 9L</figureCitation>
); hind femur angularly expanded on base of hind margin (
<figureCitation id="D8F4DDFE62289A1E8A5DF7DAD48EFCC4" box="[823,905,767,789]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Fig. 16J</figureCitation>
). Female: apical margin of clypeus deeply emarginate; clypeus black or with small markings on basal corners (
<figureCitation id="D8F4DDFE62289A1E8B92F638D44FFCE3" box="[760,840,797,819]" captionStart="FIGURE 9" captionStartId="22.[151,250,1908,1932]" captionTargetBox="[177,1411,192,1883]" captionTargetId="figure-20@22.[177,1411,181,1883]" captionTargetPageId="22" captionText="FIGURE 9. A–H, Euodynerus variegatus (Fabricius). A, E–F, female from Morocco. B, H, female from Egypt (E. variegatus kruegeri (von Schulthess), syn. nov.). C, G, female from Israel (E. variegatus kruegeri (von Schulthess), syn. nov.). D, male from Sardinia, Italy. I–M, Euodynerus velutinus Blüthgen. I, male from Azerbaijan, J, L, male from Greece. K, female from Greece. M, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239457" httpUri="https://zenodo.org/record/14239457/files/figure.png" pageId="34" pageNumber="185">Fig. 9M</figureCitation>
)...........................
<taxonomicName id="87CFBAF862289A1E8DFCF638D29BFCE3" authority="Bluthgen" authorityName="Bluthgen. I" authorityYear="1951" box="[1174,1436,796,818]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="velutinus" subGenus="Euodynerus">
<emphasis id="72BB1D6962289A1E8DFCF638D29BFCE3" bold="true" box="[1174,1436,796,818]" pageId="34" pageNumber="185">
<emphasis id="72BB1D6962289A1E8DFCF638D3ABFCE3" bold="true" box="[1174,1196,797,818]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
(
<emphasis id="72BB1D6962289A1E8DD0F638D3D7FCE3" bold="true" box="[1210,1232,797,818]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
)
<emphasis id="72BB1D6962289A1E8DB4F639D23EFCE3" bold="true" box="[1246,1337,796,818]" italics="true" pageId="34" pageNumber="185">velutinus</emphasis>
Blüthgen
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="4070C17B62289A1E89FDF61ED29BFC76" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF61ED29BFC76" pageId="34" pageNumber="185">
- Punctures on mesepisternum not arranged in series, interspaces forming irregular reticulation. Male: apical margin of clypeus shallowly emarginate, apical teeth more or less right-angled and their distance wider than interantennal space (
<figureCitation id="D8F4DDFE62289A1E8D9DF672D266FCBC" box="[1271,1377,855,877]" captionStart="FIGURE 5" captionStartId="15.[151,250,1495,1519]" captionTargetBox="[151,1435,639,1472]" captionTargetId="figure-221@15.[151,1435,633,1472]" captionTargetPageId="15" captionText="FIGURE 5. Euodynerus curictensis Blüthgen. A, E, male from Morocco. B, F, female from Greece. C, H, female from Sardinia (E. curictensis sardous Borsato, syn. nov.). D, male from Lebanon. G, female from Morocco." figureDoi="http://doi.org/10.5281/zenodo.14239449" httpUri="https://zenodo.org/record/14239449/files/figure.png" pageId="34" pageNumber="185">Figs 5D, E</figureCitation>
); hind femur not expanded, hind margin entirely straight (
<figureCitation id="D8F4DDFE62289A1E8BAFF650D419FC5A" box="[709,798,885,907]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Fig. 16K</figureCitation>
). Female: apical margin of clypeus subtruncate; clypeus with broad basal yellow markings to almost entirely yellow (
<figureCitation id="D8F4DDFE62289A1E8BD6F6B7D437FC79" box="[700,816,914,936]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Figs 15F–H</figureCitation>
)............................
<taxonomicName id="87CFBAF862289A1E8DECF6B7D29BFC76" authority="Bluthgen" authorityName="Bluthgen. A, E" authorityYear="1940" box="[1158,1436,914,936]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="curictensis" subGenus="Euodynerus">
<emphasis id="72BB1D6962289A1E8DECF6B7D29BFC76" bold="true" box="[1158,1436,914,936]" pageId="34" pageNumber="185">
<emphasis id="72BB1D6962289A1E8DECF6B7D39BFC76" bold="true" box="[1158,1180,914,935]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
(
<emphasis id="72BB1D6962289A1E8DC0F6B7D3C7FC76" bold="true" box="[1194,1216,914,935]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
)
<emphasis id="72BB1D6962289A1E8DA4F6B7D23EFC76" bold="true" box="[1230,1337,914,935]" italics="true" pageId="34" pageNumber="185">curictensis</emphasis>
Blüthgen
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<keyStep id="FB3BD9DE62289A1E89FDF695D29BFB89" pageId="34" pageNumber="185">
<paragraph id="4070C17B62289A1E89FDF695D29BFC2E" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF695D29BFC2E" pageId="34" pageNumber="185">
13. Tegula narrower and with more or less evenly curved outer margin, with few scattered fine punctures (
<figureCitation id="D8F4DDFE62289A1E8DDDF68AD20AFC14" box="[1207,1293,943,965]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Fig. 16F</figureCitation>
). Male: S6 and S7 with erect setae, either long or short (
<figureCitation id="D8F4DDFE62289A1E8B1BF6E8D5F7FC33" box="[625,752,973,995]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Figs 16L, M</figureCitation>
). Female: cephalic foveae placed in differentiated area with shiny and smooth surface (
<figureCitation id="D8F4DDFE62289A1E88EBF6CFD6DFFBD1" box="[385,472,1002,1024]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Fig. 15X</figureCitation>
)............................................................................. 14
</keyLead>
</paragraph>
<paragraph id="4070C17B62289A1E89FDF12DD29BFB89" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF12DD29BFB89" pageId="34" pageNumber="185">
- Tegula wider and with outer margin more strongly rounded in posterior half, almost smooth (
<figureCitation id="D8F4DDFE62289A1E8DEFF122D3E5FBCC" box="[1157,1250,1031,1053]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Fig. 16G</figureCitation>
). Male: S6 and S7 with dust-like pubescence, some very short oblique setae at apex only (
<figureCitation id="D8F4DDFE62289A1E8ADAF100D30AFBEA" box="[944,1037,1061,1083]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Fig. 16N</figureCitation>
). Female: cephalic foveae not placed in differentiated area, mixed with surrounding punctures (
<figureCitation id="D8F4DDFE62289A1E8B9EF167D44CFB89" box="[756,843,1090,1112]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Fig. 15Y</figureCitation>
).............................................. 15
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<keyLead id="FB3E624E62289A1E89FDF145D29BFB61" pageId="34" pageNumber="185">
14. Dorsal side of mesosoma almost bare, with very sparse dust-like pubescence (cf.
<figureCitation id="D8F4DDFE62289A1E8A9CF17AD353FBA4" box="[1014,1108,1119,1141]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Fig. 15M</figureCitation>
). Male: clypeus longer than wide, with deep and dense punctures (
<figureCitation id="D8F4DDFE62289A1E8B7FF158D558FB43" box="[533,607,1149,1171]" captionStart="FIGURE 7" captionStartId="19.[151,250,1908,1932]" captionTargetBox="[210,1376,183,1884]" captionTargetId="figure-17@19.[210,1376,181,1884]" captionTargetPageId="19" captionText="FIGURE 7. A–H, Euodynerus disconotatus (Lichtenstein). A, male from Azerbaijan. B, E, male from Italy. C, G, female from Italy. D, female from Greece. F, female from Sicily, Italy. H, female from Azerbaijan. I–O, Euodynerus fastidiosus (de Saussure). I, L, male from Italy. J, male from Greece. K, female from Crete, Greece. M, female from Italy. N, female from Crimea, Russia. O, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239453" httpUri="https://zenodo.org/record/14239453/files/figure.png" pageId="34" pageNumber="185">Fig. 7L</figureCitation>
); S6 and S7 with long erect setae on whole surface (
<figureCitation id="D8F4DDFE62289A1E8D39F158D3ADFB42" box="[1107,1194,1149,1171]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Fig. 16L</figureCitation>
). Female: face with short pubescence around antennal insertions not reaching bottom of ocular sinus..............
<taxonomicName id="87CFBAF862289A1E8D31F1BFD29BFB61" authority="(de Saussure)" authorityName="UZ Uchkulach" baseAuthorityName="de Saussure" baseAuthorityYear="1853" box="[1115,1436,1178,1200]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="fastidiosus" subGenus="Euodynerus">
<emphasis id="72BB1D6962289A1E8D31F1BFD29BFB61" bold="true" box="[1115,1436,1178,1200]" pageId="34" pageNumber="185">
<emphasis id="72BB1D6962289A1E8D31F1BFD376FB7E" bold="true" box="[1115,1137,1178,1199]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
(
<emphasis id="72BB1D6962289A1E8D15F1BFD392FB7E" bold="true" box="[1151,1173,1178,1199]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
)
<emphasis id="72BB1D6962289A1E8DC9F1BFD20BFB61" bold="true" box="[1187,1292,1178,1200]" italics="true" pageId="34" pageNumber="185">fastidiosus</emphasis>
(de Saussure)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4070C17B62289A1E89FDF19DD29BFAF4" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF19DD29BFAF4" pageId="34" pageNumber="185">
- Dorsal side of mesosoma with sparse apically bent setae in addition to dust-like pubescence (
<figureCitation id="D8F4DDFE62289A1E8D32F192D3B6FB1C" box="[1112,1201,1207,1229]" captionStart="FIGURE 15" captionStartId="31.[151,250,1654,1678]" captionTargetBox="[151,1435,181,1630]" captionTargetId="figure-17@31.[151,1435,181,1630]" captionTargetPageId="31" captionText="FIGURE 15. A–E, S2 in lateral view. F–I, propodeum in posterodorsal view (arrowheads indicating dorsal carinae). J–O, vestiture of mesosoma. P–Q, metanotum in anterior view. R–T, male hind tarsus in dorsal view. U–Y, female head in dorsal view (arrowhead in Y indicating foveae not placed in depression). A, F, J, Euodynerus bidentiformis (Giordani Soika). B, V, E. bidentoides (Giordani Soika). C, K, E. quadrifasciatus (Fabricius). D, E. caspicus (Morawitz). E, H, M, T, E. dantici (Rossi). G, P, S, U, E. posticus (Herrich-Schaeffer). I, Y, E. disconotatus (Lichtenstein). L, R, E. notatus (Jurine). N, E. curictensis Blüthgen. O, E. semisaecularis (Dalla Torre). Q, X, E. fastidiosus (de Saussure). W, E. rubrosignatus Gusenleitner." figureDoi="http://doi.org/10.5281/zenodo.14239469" httpUri="https://zenodo.org/record/14239469/files/figure.png" pageId="34" pageNumber="185">Fig. 15O</figureCitation>
). Male: clypeus about as long as wide, with shallow sparse punctures (
<figureCitation id="D8F4DDFE62289A1E8BF0F1F0D406FB3B" box="[666,769,1237,1259]" captionStart="FIGURE 8" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[200,1386,192,1918]" captionTargetId="figure-20@20.[200,1386,181,1918]" captionTargetPageId="20" captionText="FIGURE 8. A–F, Euodynerus hellenicus Blüthgen. A, male from Bulgaria. B, E, female from Spain. C, F, female from Iran. D, male from Spain. G–M, Euodynerus semisaecularis (Dalla Torre). G, J, male from Greece. H, L, female from Crete, Greece. I, M, female from Afghanistan. K, male from Armenia." figureDoi="http://doi.org/10.5281/zenodo.14239455" httpUri="https://zenodo.org/record/14239455/files/figure.png" pageId="34" pageNumber="185">Figs 8J, K</figureCitation>
); S6 and S7 with short setae on apical half only (
<figureCitation id="D8F4DDFE62289A1E8DB5F1F0D23AFB3A" box="[1247,1341,1237,1259]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="34" pageNumber="185">Fig. 16M</figureCitation>
). Female: face with long dense pubescence in whole lower half and reaching bottom of ocular sinus...................................................................................................
<taxonomicName id="87CFBAF862289A1E8D5DF035D29BFAF4" authority="(Dalla Torre)" baseAuthorityName="Dalla Torre" box="[1079,1436,1295,1317]" class="Insecta" family="Vespidae" genus="Odynerus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="semisaecularis" subGenus="Euodynerus">
<emphasis id="72BB1D6962289A1E8D5DF035D29BFAF4" bold="true" box="[1079,1436,1295,1317]" pageId="34" pageNumber="185">
<emphasis id="72BB1D6962289A1E8D5DF035D34AFAF4" bold="true" box="[1079,1101,1296,1317]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
(
<emphasis id="72BB1D6962289A1E8D31F035D376FAF4" bold="true" box="[1115,1137,1296,1317]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
)
<emphasis id="72BB1D6962289A1E8D15F02AD217FAF4" bold="true" box="[1151,1296,1295,1317]" italics="true" pageId="34" pageNumber="185">semisaecularis</emphasis>
(Dalla Torre)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FB3BD9DE62289A1E89FDF008D29BF954" pageId="34" pageNumber="185">
<paragraph id="4070C17B62289A1E89FDF008D29BFA04" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF008D29BFA04" pageId="34" pageNumber="185">
15. Mesoscutum with subquadrate yellow marking in front of scutellum (
<figureCitation id="D8F4DDFE62289A1E8AE3F008D4D1FA93" box="[905,982,1325,1347]" captionStart="FIGURE 7" captionStartId="19.[151,250,1908,1932]" captionTargetBox="[210,1376,183,1884]" captionTargetId="figure-17@19.[210,1376,181,1884]" captionTargetPageId="19" captionText="FIGURE 7. A–H, Euodynerus disconotatus (Lichtenstein). A, male from Azerbaijan. B, E, male from Italy. C, G, female from Italy. D, female from Greece. F, female from Sicily, Italy. H, female from Azerbaijan. I–O, Euodynerus fastidiosus (de Saussure). I, L, male from Italy. J, male from Greece. K, female from Crete, Greece. M, female from Italy. N, female from Crimea, Russia. O, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239453" httpUri="https://zenodo.org/record/14239453/files/figure.png" pageId="34" pageNumber="185">Fig. 7C</figureCitation>
), usually absent in males (
<figureCitation id="D8F4DDFE62289A1E8DB1F008D221FA93" box="[1243,1318,1325,1347]" captionStart="FIGURE 7" captionStartId="19.[151,250,1908,1932]" captionTargetBox="[210,1376,183,1884]" captionTargetId="figure-17@19.[210,1376,181,1884]" captionTargetPageId="19" captionText="FIGURE 7. A–H, Euodynerus disconotatus (Lichtenstein). A, male from Azerbaijan. B, E, male from Italy. C, G, female from Italy. D, female from Greece. F, female from Sicily, Italy. H, female from Azerbaijan. I–O, Euodynerus fastidiosus (de Saussure). I, L, male from Italy. J, male from Greece. K, female from Crete, Greece. M, female from Italy. N, female from Crimea, Russia. O, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239453" httpUri="https://zenodo.org/record/14239453/files/figure.png" pageId="34" pageNumber="185">Fig. 7B</figureCitation>
) (present in Eastern populations,
<figureCitation id="D8F4DDFE62289A1E88C1F06FD6F1FA8E" box="[427,502,1354,1376]" captionStart="FIGURE 7" captionStartId="19.[151,250,1908,1932]" captionTargetBox="[210,1376,183,1884]" captionTargetId="figure-17@19.[210,1376,181,1884]" captionTargetPageId="19" captionText="FIGURE 7. A–H, Euodynerus disconotatus (Lichtenstein). A, male from Azerbaijan. B, E, male from Italy. C, G, female from Italy. D, female from Greece. F, female from Sicily, Italy. H, female from Azerbaijan. I–O, Euodynerus fastidiosus (de Saussure). I, L, male from Italy. J, male from Greece. K, female from Crete, Greece. M, female from Italy. N, female from Crimea, Russia. O, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239453" httpUri="https://zenodo.org/record/14239453/files/figure.png" pageId="34" pageNumber="185">Fig. 7A</figureCitation>
) and rarely reduced in females (
<figureCitation id="D8F4DDFE62289A1E8A41F06FD47FFA8E" box="[811,888,1354,1376]" captionStart="FIGURE 7" captionStartId="19.[151,250,1908,1932]" captionTargetBox="[210,1376,183,1884]" captionTargetId="figure-17@19.[210,1376,181,1884]" captionTargetPageId="19" captionText="FIGURE 7. A–H, Euodynerus disconotatus (Lichtenstein). A, male from Azerbaijan. B, E, male from Italy. C, G, female from Italy. D, female from Greece. F, female from Sicily, Italy. H, female from Azerbaijan. I–O, Euodynerus fastidiosus (de Saussure). I, L, male from Italy. J, male from Greece. K, female from Crete, Greece. M, female from Italy. N, female from Crimea, Russia. O, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239453" httpUri="https://zenodo.org/record/14239453/files/figure.png" pageId="34" pageNumber="185">Fig. 7D</figureCitation>
); scutellum usually with two separate subtriangular spots (
<figureCitation id="D8F4DDFE62289A1E8983F042D652FAAC" box="[233,341,1383,1405]" captionStart="FIGURE 7" captionStartId="19.[151,250,1908,1932]" captionTargetBox="[210,1376,183,1884]" captionTargetId="figure-17@19.[210,1376,181,1884]" captionTargetPageId="19" captionText="FIGURE 7. A–H, Euodynerus disconotatus (Lichtenstein). A, male from Azerbaijan. B, E, male from Italy. C, G, female from Italy. D, female from Greece. F, female from Sicily, Italy. H, female from Azerbaijan. I–O, Euodynerus fastidiosus (de Saussure). I, L, male from Italy. J, male from Greece. K, female from Crete, Greece. M, female from Italy. N, female from Crimea, Russia. O, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239453" httpUri="https://zenodo.org/record/14239453/files/figure.png" pageId="34" pageNumber="185">Figs 7B–D</figureCitation>
) (broad band in populations from Asia,
<figureCitation id="D8F4DDFE62289A1E8BB4F042D42BFAAC" box="[734,812,1383,1405]" captionStart="FIGURE 7" captionStartId="19.[151,250,1908,1932]" captionTargetBox="[210,1376,183,1884]" captionTargetId="figure-17@19.[210,1376,181,1884]" captionTargetPageId="19" captionText="FIGURE 7. A–H, Euodynerus disconotatus (Lichtenstein). A, male from Azerbaijan. B, E, male from Italy. C, G, female from Italy. D, female from Greece. F, female from Sicily, Italy. H, female from Azerbaijan. I–O, Euodynerus fastidiosus (de Saussure). I, L, male from Italy. J, male from Greece. K, female from Crete, Greece. M, female from Italy. N, female from Crimea, Russia. O, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239453" httpUri="https://zenodo.org/record/14239453/files/figure.png" pageId="34" pageNumber="185">Fig. 7A</figureCitation>
); metanotum with broad yellow band covering dorsal half (
<figureCitation id="D8F4DDFE62289A1E8C18F042D627FA4B" captionStart="FIGURE 7" captionStartId="19.[151,250,1908,1932]" captionTargetBox="[210,1376,183,1884]" captionTargetId="figure-17@19.[210,1376,181,1884]" captionTargetPageId="19" captionText="FIGURE 7. A–H, Euodynerus disconotatus (Lichtenstein). A, male from Azerbaijan. B, E, male from Italy. C, G, female from Italy. D, female from Greece. F, female from Sicily, Italy. H, female from Azerbaijan. I–O, Euodynerus fastidiosus (de Saussure). I, L, male from Italy. J, male from Greece. K, female from Crete, Greece. M, female from Italy. N, female from Crimea, Russia. O, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239453" httpUri="https://zenodo.org/record/14239453/files/figure.png" pageId="34" pageNumber="185">Figs 7A–D</figureCitation>
). Male: clypeus as long as wide with apical margin as wide as interantennal distance, surface with sparse but clearly visible punctures (
<figureCitation id="D8F4DDFE62289A1E88FCF087D6E5FA66" box="[406,482,1442,1464]" captionStart="FIGURE 7" captionStartId="19.[151,250,1908,1932]" captionTargetBox="[210,1376,183,1884]" captionTargetId="figure-17@19.[210,1376,181,1884]" captionTargetPageId="19" captionText="FIGURE 7. A–H, Euodynerus disconotatus (Lichtenstein). A, male from Azerbaijan. B, E, male from Italy. C, G, female from Italy. D, female from Greece. F, female from Sicily, Italy. H, female from Azerbaijan. I–O, Euodynerus fastidiosus (de Saussure). I, L, male from Italy. J, male from Greece. K, female from Crete, Greece. M, female from Italy. N, female from Crimea, Russia. O, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239453" httpUri="https://zenodo.org/record/14239453/files/figure.png" pageId="34" pageNumber="185">Fig. 7E</figureCitation>
). Female: clypeus coarsely punctate and with some longitudinal striae, apical carinae mixed with coarse sculpture and not well-evident (
<figureCitation id="D8F4DDFE62289A1E8B3FF09AD5B9FA04" box="[597,702,1471,1493]" captionStart="FIGURE 7" captionStartId="19.[151,250,1908,1932]" captionTargetBox="[210,1376,183,1884]" captionTargetId="figure-17@19.[210,1376,181,1884]" captionTargetPageId="19" captionText="FIGURE 7. A–H, Euodynerus disconotatus (Lichtenstein). A, male from Azerbaijan. B, E, male from Italy. C, G, female from Italy. D, female from Greece. F, female from Sicily, Italy. H, female from Azerbaijan. I–O, Euodynerus fastidiosus (de Saussure). I, L, male from Italy. J, male from Greece. K, female from Crete, Greece. M, female from Italy. N, female from Crimea, Russia. O, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239453" httpUri="https://zenodo.org/record/14239453/files/figure.png" pageId="34" pageNumber="185">Figs 7F–H</figureCitation>
)...............................
<taxonomicName id="87CFBAF862289A1E8D2EF0E5D29BFA04" authority="(Lichtenstein)" authorityName="UZ Uchkulach" baseAuthorityName="Lichtenstein" baseAuthorityYear="1884" box="[1092,1436,1471,1493]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="disconotatus" subGenus="Euodynerus">
<emphasis id="72BB1D6962289A1E8D2EF0E5D29BFA04" bold="true" box="[1092,1436,1471,1493]" pageId="34" pageNumber="185">
<emphasis id="72BB1D6962289A1E8D2EF0E5D35DFA04" bold="true" box="[1092,1114,1472,1493]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
(
<emphasis id="72BB1D6962289A1E8D02F0E5D379FA04" bold="true" box="[1128,1150,1472,1493]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
)
<emphasis id="72BB1D6962289A1E8DE6F09AD20FFA04" bold="true" box="[1164,1288,1471,1493]" italics="true" pageId="34" pageNumber="185">disconotatus</emphasis>
(Lichtenstein)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4070C17B62289A1E89FDF0F8D29BF954" blockId="34.[151,1437,151,1669]" pageId="34" pageNumber="185">
<keyLead id="FB3E624E62289A1E89FDF0F8D29BF954" pageId="34" pageNumber="185">
- Mesoscutum entirely black (
<figureCitation id="D8F4DDFE62289A1E8898F0F8D567FA23" box="[498,608,1501,1523]" captionStart="FIGURE 9" captionStartId="22.[151,250,1908,1932]" captionTargetBox="[177,1411,192,1883]" captionTargetId="figure-20@22.[177,1411,181,1883]" captionTargetPageId="22" captionText="FIGURE 9. A–H, Euodynerus variegatus (Fabricius). A, E–F, female from Morocco. B, H, female from Egypt (E. variegatus kruegeri (von Schulthess), syn. nov.). C, G, female from Israel (E. variegatus kruegeri (von Schulthess), syn. nov.). D, male from Sardinia, Italy. I–M, Euodynerus velutinus Blüthgen. I, male from Azerbaijan, J, L, male from Greece. K, female from Greece. M, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239457" httpUri="https://zenodo.org/record/14239457/files/figure.png" pageId="34" pageNumber="185">Figs 9A–C</figureCitation>
); scutellum usually with wide band covering most of its surface (
<figureCitation id="D8F4DDFE62289A1E8DBAF0F8D23CFA23" box="[1232,1339,1501,1523]" captionStart="FIGURE 9" captionStartId="22.[151,250,1908,1932]" captionTargetBox="[177,1411,192,1883]" captionTargetId="figure-20@22.[177,1411,181,1883]" captionTargetPageId="22" captionText="FIGURE 9. A–H, Euodynerus variegatus (Fabricius). A, E–F, female from Morocco. B, H, female from Egypt (E. variegatus kruegeri (von Schulthess), syn. nov.). C, G, female from Israel (E. variegatus kruegeri (von Schulthess), syn. nov.). D, male from Sardinia, Italy. I–M, Euodynerus velutinus Blüthgen. I, male from Azerbaijan, J, L, male from Greece. K, female from Greece. M, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239457" httpUri="https://zenodo.org/record/14239457/files/figure.png" pageId="34" pageNumber="185">Figs 9A, B</figureCitation>
) (reduced to spots or absent in populations from the Levant, which have red markings,
<figureCitation id="D8F4DDFE62289A1E8A81F0DFD33EF9DE" box="[1003,1081,1530,1552]" captionStart="FIGURE 9" captionStartId="22.[151,250,1908,1932]" captionTargetBox="[177,1411,192,1883]" captionTargetId="figure-20@22.[177,1411,181,1883]" captionTargetPageId="22" captionText="FIGURE 9. A–H, Euodynerus variegatus (Fabricius). A, E–F, female from Morocco. B, H, female from Egypt (E. variegatus kruegeri (von Schulthess), syn. nov.). C, G, female from Israel (E. variegatus kruegeri (von Schulthess), syn. nov.). D, male from Sardinia, Italy. I–M, Euodynerus velutinus Blüthgen. I, male from Azerbaijan, J, L, male from Greece. K, female from Greece. M, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239457" httpUri="https://zenodo.org/record/14239457/files/figure.png" pageId="34" pageNumber="185">Fig. 9C</figureCitation>
); metanotum at most with irregular band along transverse carina (
<figureCitation id="D8F4DDFE62289A1E8B60F332D57CF9FC" box="[522,635,1559,1581]" captionStart="FIGURE 9" captionStartId="22.[151,250,1908,1932]" captionTargetBox="[177,1411,192,1883]" captionTargetId="figure-20@22.[177,1411,181,1883]" captionTargetPageId="22" captionText="FIGURE 9. A–H, Euodynerus variegatus (Fabricius). A, E–F, female from Morocco. B, H, female from Egypt (E. variegatus kruegeri (von Schulthess), syn. nov.). C, G, female from Israel (E. variegatus kruegeri (von Schulthess), syn. nov.). D, male from Sardinia, Italy. I–M, Euodynerus velutinus Blüthgen. I, male from Azerbaijan, J, L, male from Greece. K, female from Greece. M, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239457" httpUri="https://zenodo.org/record/14239457/files/figure.png" pageId="34" pageNumber="185">Figs 9A–C</figureCitation>
). Male: clypeus longer than wide with apical margin narrower than interantennal distance, surface microsculpted with very fine punctures visible only at high magnification (
<figureCitation id="D8F4DDFE62289A1E8D0AF310D3A8F99B" box="[1120,1199,1589,1611]" captionStart="FIGURE 9" captionStartId="22.[151,250,1908,1932]" captionTargetBox="[177,1411,192,1883]" captionTargetId="figure-20@22.[177,1411,181,1883]" captionTargetPageId="22" captionText="FIGURE 9. A–H, Euodynerus variegatus (Fabricius). A, E–F, female from Morocco. B, H, female from Egypt (E. variegatus kruegeri (von Schulthess), syn. nov.). C, G, female from Israel (E. variegatus kruegeri (von Schulthess), syn. nov.). D, male from Sardinia, Italy. I–M, Euodynerus velutinus Blüthgen. I, male from Azerbaijan, J, L, male from Greece. K, female from Greece. M, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239457" httpUri="https://zenodo.org/record/14239457/files/figure.png" pageId="34" pageNumber="185">Fig. 9D</figureCitation>
). Female: clypeus finely and sparsely punctate, at most with scattered microstriation between punctures, apical carinae not mixed with sculpture and well visible (
<figureCitation id="D8F4DDFE62289A1E8834F34AD6CFF954" box="[350,456,1647,1669]" captionStart="FIGURE 9" captionStartId="22.[151,250,1908,1932]" captionTargetBox="[177,1411,192,1883]" captionTargetId="figure-20@22.[177,1411,181,1883]" captionTargetPageId="22" captionText="FIGURE 9. A–H, Euodynerus variegatus (Fabricius). A, E–F, female from Morocco. B, H, female from Egypt (E. variegatus kruegeri (von Schulthess), syn. nov.). C, G, female from Israel (E. variegatus kruegeri (von Schulthess), syn. nov.). D, male from Sardinia, Italy. I–M, Euodynerus velutinus Blüthgen. I, male from Azerbaijan, J, L, male from Greece. K, female from Greece. M, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239457" httpUri="https://zenodo.org/record/14239457/files/figure.png" pageId="34" pageNumber="185">Figs 9E–H</figureCitation>
).........................................................
<taxonomicName id="87CFBAF862289A1E8D1CF355D29BF954" authority="(Fabricius)" authorityName="IT Assemini" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1142,1436,1647,1669]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="variegatus" subGenus="Euodynerus">
<emphasis id="72BB1D6962289A1E8D1CF355D29BF954" bold="true" box="[1142,1436,1647,1669]" pageId="34" pageNumber="185">
<emphasis id="72BB1D6962289A1E8D1CF355D38BF954" bold="true" box="[1142,1164,1648,1669]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
(
<emphasis id="72BB1D6962289A1E8DF0F355D3B7F954" bold="true" box="[1178,1200,1648,1669]" italics="true" pageId="34" pageNumber="185">E.</emphasis>
)
<emphasis id="72BB1D6962289A1E8DD4F34AD222F955" bold="true" box="[1214,1317,1647,1668]" italics="true" pageId="34" pageNumber="185">variegatus</emphasis>
(Fabricius)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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