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<document id="B817970DA6E2D520DC9B00EA7D2EAABA" ID-DOI="10.5281/zenodo.173802" ID-GBIF-Dataset="0a6c9168-e24b-4d02-955f-29d133c8890a" ID-ISSN="1175­5326" ID-Zenodo-Dep="173802" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459784372853" checkinUser="plazi" docAuthor="Bousquet, Yves" docDate="2006" docId="B74587BDFF88FFC8D36C3DD79DADE0DF" docLanguage="en" docName="zt01308p029.pdf" docOrigin="Zootaxa 1308" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Aspidoglossa subangulata Chaudoir 1843" docType="treatment" docVersion="8" lastPageNumber="10" masterDocId="4B7CFFC5FF8FFFC1D264384E9E5CE171" masterDocTitle="Review of the species of Ardistomina (Coleoptera: Carabidae: Clivinini) in America north of Mexico" masterLastPageNumber="29" masterPageNumber="1" pageNumber="8" updateTime="1698218318931" updateUser="plazi">
<mods:mods id="99CA3EFD803FFD882846869C077C98FF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D73538EBFA07C35A9FC4BC7C0BC53C61">
<mods:title id="FB7BD30EB17D52187E506D63E01940D0">Review of the species of Ardistomina (Coleoptera: Carabidae: Clivinini) in America north of Mexico</mods:title>
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<mods:namePart id="ADCB216280DC1C29DEB8F6EE7C1CA9BD">Bousquet, Yves</mods:namePart>
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<mods:date id="FBFE2B613E82965E2BA8C693F5EA8BE2">2006</mods:date>
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<mods:number id="A35147883C49AA2B8FC3EA3918E0D940">1308</mods:number>
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<mods:classification id="6BDB684582F8601873E0286AADFC5DBD">journal article</mods:classification>
<mods:identifier id="83F59929745B77350C6FEACB64CF7A03" type="DOI">10.5281/zenodo.173802</mods:identifier>
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<mods:identifier id="E22049C624B6545E892899D16D6AF92F" type="ISSN">1175­5326</mods:identifier>
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<treatment id="B74587BDFF88FFC8D36C3DD79DADE0DF" ID-DOI="http://doi.org/10.5281/zenodo.6262681" ID-GBIF-Taxon="119348754" ID-Zenodo-Dep="6262681" LSID="urn:lsid:plazi:treatment:B74587BDFF88FFC8D36C3DD79DADE0DF" httpUri="http://treatment.plazi.org/id/B74587BDFF88FFC8D36C3DD79DADE0DF" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
<subSubSection id="77F66520FF88FFC6D36C3DD79CCAE4A7" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="3F5336ABFF88FFC6D36C3DD79D40E4C2" blockId="7.[264,796,1433,1494]" box="[264,796,1433,1459]" pageId="7" pageNumber="8">
<heading id="641B81C7FF88FFC6D36C3DD79D40E4C2" bold="true" box="[264,796,1433,1459]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<taxonomicName id="F8EC4D28FF88FFC6D36C3DD79D40E4C2" ID-CoL="HH4B" authority="Chaudoir, 1843" authorityName="Chaudoir" authorityYear="1843" box="[264,796,1433,1459]" class="Insecta" family="Carabidae" genus="Aspidoglossa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="subangulata">
<emphasis id="0D98EAB9FF88FFC6D36C3DD79D40E4C2" bold="true" box="[264,796,1433,1459]" pageId="7" pageNumber="8">
<emphasis id="0D98EAB9FF88FFC6D36C3DD79C1EE4C2" bold="true" box="[264,578,1433,1459]" italics="true" pageId="7" pageNumber="8">Aspidoglossa subangulata</emphasis>
(
<bibRefCitation id="5B7D4B5AFF88FFC6D0363DD79D48E4C2" author="Chaudoir" box="[594,788,1433,1459]" pageId="7" pageNumber="27" refString="Chaudoir, M. de. (1843) Carabiques nouveaux. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16, 671 - 791." type="journal article" year="1843">Chaudoir, 1843</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="3F5336ABFF88FFC6D36C3DF29CCAE4A7" blockId="7.[264,796,1433,1494]" box="[264,662,1468,1494]" pageId="7" pageNumber="8">
(
<figureCitation id="A7D72A2EFF88FFC6D3743DF29F00E4A7" box="[272,348,1468,1494]" captionStart="FIGURES 4 – 8" captionStartId="12.[264,382,1796,1820]" captionTargetBox="[361,1232,331,1758]" captionTargetId="figure@12.[335,1252,306,1791]" captionTargetPageId="12" captionText="FIGURES 4 – 8. Median lobe (a) and paremeres (b). 4, Aspidoglossa subangulata; 5, Semiardistomis viridis; 6, S. puncticollis; 7, Ardistomis schaumii; 8, A. obliquatus." httpUri="https://zenodo.org/record/173805/files/figure.png" pageId="7" pageNumber="8">Figs 4</figureCitation>
,
<figureCitation id="A7D72A2EFF88FFC6D30D3DF29FEFE4A7" box="[361,435,1468,1494]" captionStart="FIGURES 11 – 18. 11 – 14" captionStartId="14.[264,382,1651,1675]" captionTargetBox="[372,1224,323,1650]" captionTargetId="figure@14.[356,1238,316,1655]" captionTargetPageId="14" captionText="FIGURES 11 – 18. 11 – 14. Apical stylomere. 11, Aspidoglossa subangulata; 12, Semiardistomis viridis; 13, Ardistomis schaumii; 14, A. obliquatus. 15 – 18. Spermatheca. 15, Aspidoglossa subangulata; 16, Semiardistomis viridis; 17, S. puncticollis; 18, Ardistomis schaumii." httpUri="https://zenodo.org/record/173807/files/figure.png" pageId="7" pageNumber="8">11, 15</figureCitation>
,
<figureCitation id="A7D72A2EFF88FFC6D3A43DF29F83E4A7" box="[448,479,1468,1494]" captionStart="FIGURES 19 – 22. 19 – 21" captionStartId="15.[264,382,1139,1163]" captionTargetBox="[269,1319,329,1113]" captionTargetId="figure@15.[264,1323,326,1115]" captionTargetPageId="15" captionText="FIGURES 19 – 22. 19 – 21. Head (dorsal view). 19, Aspidoglossa subangulata; 20, Semiardistomis viridis; 21, Ardistomis schaumii. 22, Extremity of body (lateral view) of Ardistomis schaumii. Scale bar = 0.2 mm (Figs 20 – 22), 0.5 mm (Fig. 19)." httpUri="https://zenodo.org/record/173808/files/figure.png" pageId="7" pageNumber="8">19</figureCitation>
,
<figureCitation id="A7D72A2EFF88FFC6D3883DF29C64E4A7" box="[492,568,1468,1494]" captionStart="FIGURES 23 – 26. 23 – 24" captionStartId="16.[264,382,1139,1163]" captionTargetBox="[267,1323,326,1113]" captionTargetId="figure@16.[264,1323,326,1113]" captionTargetPageId="16" captionText="FIGURES 23 – 26. 23 – 24. Protarsomeres (ventral view). 23, Aspidoglossa subangulata ɗ; 24, Aspidoglossa subangulata Ψ. 25 – 26. Humeral region (dorsal view). 25, Aspidoglossa subangulata; 26, Semiardistomis viridis. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/173809/files/figure.png" pageId="7" pageNumber="8">23–25</figureCitation>
,
<figureCitation id="A7D72A2EFF88FFC6D0273DF29C3FE4A7" box="[579,611,1468,1494]" captionStart="FIGURES 27 – 30. 27" captionStartId="17.[264,383,1120,1144]" captionTargetBox="[264,1318,314,1105]" captionTargetId="figure@17.[264,1323,314,1106]" captionTargetPageId="17" captionText="FIGURES 27 – 30. 27, Prosternum of Ardistomis schaumii. 28 – 30. Protibia (ventral view). 28, Aspidoglossa subangulata; 29, Semiardistomis viridis; 30, Ardistomis schaumii. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/173810/files/figure.png" pageId="7" pageNumber="8">28</figureCitation>
,
<figureCitation id="A7D72A2EFF88FFC6D00B3DF29CD1E4A7" box="[623,653,1468,1494]" captionStart="FIGURE 31" captionStartId="19.[264,367,1030,1054]" captionTargetBox="[338,1248,327,1006]" captionTargetId="figure@19.[338,1248,326,1006]" captionTargetPageId="19" captionText="FIGURE 31. Collection localities for Aspidoglossa subangulata (Chaudoir) in United States of America." httpUri="https://zenodo.org/record/173811/files/figure.png" pageId="7" pageNumber="8">31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="77F66520FF88FFC6D36C3E429B77E632" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="3F5336ABFF88FFC6D36C3E429FE8E732" blockId="7.[264,1323,1548,1859]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="1FFC1185FF88FFC6D36C3E429FE8E732" pageId="7" pageNumber="8">
<taxonomicName id="F8EC4D28FF88FFC6D36C3E429CAEE752" ID-CoL="3882P" authority="Chaudoir 1843: 738" authorityName="Chaudoir" authorityPageNumber="738" authorityYear="1843" box="[264,754,1548,1571]" class="Insecta" family="Carabidae" genus="Dyschirius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="subangulatus">
<emphasis id="0D98EAB9FF88FFC6D36C3E429C4DE752" box="[264,529,1548,1571]" italics="true" pageId="7" pageNumber="8">Dyschirius subangulatus</emphasis>
<treatmentCitation id="BE4D10BAFF88FFC6D07D3E429CAEE752" author="Chaudoir" box="[537,754,1548,1571]" page="738" pageId="7" pageNumber="8" year="1843">
<bibRefCitation id="5B7D4B5AFF88FFC6D07D3E429CE0E752" author="Chaudoir" box="[537,700,1548,1571]" pageId="7" pageNumber="27" refString="Chaudoir, M. de. (1843) Carabiques nouveaux. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16, 671 - 791." type="journal article" year="1843">Chaudoir 1843</bibRefCitation>
: 738
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="E0578809FF88FFC6D0993E429D6EE752" box="[765,818,1548,1571]" pageId="7" pageNumber="8">Type</typeStatus>
locality: New Orleans, Orleans Parish, Louisiana,
<collectingCountry id="47FB763BFF88FFC6D30E3E629FE8E732" box="[362,436,1580,1603]" name="United States of America" pageId="7" pageNumber="8">U.S.A.</collectingCountry>
</treatmentCitationGroup>
</paragraph>
<paragraph id="3F5336ABFF88FFC6D36C3E029D26E7F2" blockId="7.[264,1323,1548,1859]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="1FFC1185FF88FFC6D36C3E029D26E7F2" pageId="7" pageNumber="8">
<taxonomicName id="F8EC4D28FF88FFC6D36C3E029C93E712" ID-CoL="387VB" authority="Chaudoir 1843: 737" authorityName="Chaudoir" authorityPageNumber="737" authorityYear="1843" box="[264,719,1612,1635]" class="Insecta" family="Carabidae" genus="Dyschirius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="0D98EAB9FF88FFC6D36C3E029FB1E712" box="[264,493,1612,1635]" italics="true" pageId="7" pageNumber="8">Dyschirius humeralis</emphasis>
<treatmentCitation id="BE4D10BAFF88FFC6D3913E029C93E712" author="Chaudoir" box="[501,719,1612,1635]" page="737" pageId="7" pageNumber="8" year="1843">
<bibRefCitation id="5B7D4B5AFF88FFC6D3913E029CC5E712" author="Chaudoir" box="[501,665,1612,1635]" pageId="7" pageNumber="27" refString="Chaudoir, M. de. (1843) Carabiques nouveaux. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16, 671 - 791." type="journal article" year="1843">Chaudoir 1843</bibRefCitation>
: 737
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="E0578809FF88FFC6D0BD3E029D52E712" box="[729,782,1612,1635]" pageId="7" pageNumber="8">Type</typeStatus>
locality: New Orleans, Orleans Parish, Louisiana,
<collectingCountry id="47FB763BFF88FFC6D35C3E229FDEE7F2" box="[312,386,1644,1667]" name="United States of America" pageId="7" pageNumber="8">U.S.A.</collectingCountry>
Synonymy established by
<treatmentCitation id="BE4D10BAFF88FFC6D0C43E229D29E7F2" author="LeConte" box="[672,885,1644,1667]" page="80" pageId="7" pageNumber="8" year="1857">
<bibRefCitation id="5B7D4B5AFF88FFC6D0C43E229D29E7F2" author="LeConte" box="[672,885,1644,1667]" pageId="7" pageNumber="28" refString="LeConte, J. L. (1857) Synopsis of the species Clivina and allied genera inhabiting the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, [9], 75 - 83." type="journal article" year="1857">LeConte (1857: 80)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3F5336ABFF88FFC6D36C3EC29C4DE7B2" blockId="7.[264,1323,1548,1859]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="1FFC1185FF88FFC6D36C3EC29C4DE7B2" pageId="7" pageNumber="8">
<taxonomicName id="F8EC4D28FF88FFC6D36C3EC29CF6E7D2" ID-CoL="HH4F" authority="Putzeys 1846: 632" authorityName="Putzeys" authorityPageNumber="632" authorityYear="1846" box="[264,682,1676,1699]" class="Insecta" family="Carabidae" genus="Aspidoglossa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="0D98EAB9FF88FFC6D36C3EC29F81E7D2" box="[264,477,1676,1699]" italics="true" pageId="7" pageNumber="8">Aspidoglossa vicina</emphasis>
<treatmentCitation id="BE4D10BAFF88FFC6D3803EC29CF6E7D2" author="Putzeys" box="[484,682,1676,1699]" page="632" pageId="7" pageNumber="8" year="1846">
<bibRefCitation id="5B7D4B5AFF88FFC6D3803EC29C2AE7D2" author="Putzeys" box="[484,630,1676,1699]" pageId="7" pageNumber="29" refString="Putzeys, J. A. A. H. (1846) Monographie des Clivina et genres voisins, precedee d'un tableau synoptique des genres de la tribu des scaritides. Memoires de la Societe Royale des Sciences de Liege, 2, 521 - 663." type="journal article" year="1846">Putzeys 1846</bibRefCitation>
: 632
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="E0578809FF88FFC6D0D03EC29CB5E7D2" box="[692,745,1676,1699]" pageId="7" pageNumber="8">Type</typeStatus>
locality: &quot;Caroline&quot;,
<collectingCountry id="47FB763BFF88FFC6D1AF3EC29A49E7D2" box="[971,1045,1676,1699]" name="United States of America" pageId="7" pageNumber="8">U.S.A.</collectingCountry>
Synonymy established by
<treatmentCitation id="BE4D10BAFF88FFC6D35C3EE29C51E7B2" author="LeConte" box="[312,525,1708,1731]" page="80" pageId="7" pageNumber="8" year="1857">
<bibRefCitation id="5B7D4B5AFF88FFC6D35C3EE29C51E7B2" author="LeConte" box="[312,525,1708,1731]" pageId="7" pageNumber="28" refString="LeConte, J. L. (1857) Synopsis of the species Clivina and allied genera inhabiting the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, [9], 75 - 83." type="journal article" year="1857">LeConte (1857: 80)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3F5336ABFF88FFC6D36C3E829C25E672" blockId="7.[264,1323,1548,1859]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="1FFC1185FF88FFC6D36C3E829C25E672" pageId="7" pageNumber="8">
<taxonomicName id="F8EC4D28FF88FFC6D36C3E829C8FE792" ID-CoL="HH3S" authority="Putzeys 1846: 632" authorityName="Putzeys" authorityPageNumber="632" authorityYear="1846" box="[264,723,1740,1763]" class="Insecta" family="Carabidae" genus="Aspidoglossa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="0D98EAB9FF88FFC6D36C3E829FABE792" box="[264,503,1740,1763]" italics="true" pageId="7" pageNumber="8">Aspidoglossa fraterna</emphasis>
<treatmentCitation id="BE4D10BAFF88FFC6D0673E829C8FE792" author="Putzeys" box="[515,723,1740,1763]" page="632" pageId="7" pageNumber="8" year="1846">
<bibRefCitation id="5B7D4B5AFF88FFC6D0673E829CC5E792" author="Putzeys" box="[515,665,1740,1763]" pageId="7" pageNumber="29" refString="Putzeys, J. A. A. H. (1846) Monographie des Clivina et genres voisins, precedee d'un tableau synoptique des genres de la tribu des scaritides. Memoires de la Societe Royale des Sciences de Liege, 2, 521 - 663." type="journal article" year="1846">Putzeys 1846</bibRefCitation>
: 632
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="E0578809FF88FFC6D0853E829D4AE792" box="[737,790,1740,1763]" pageId="7" pageNumber="8">Type</typeStatus>
locality: &quot;Amérique boréale&quot;. Synonymy established by
<treatmentCitation id="BE4D10BAFF88FFC6D3FB3EA29C28E672" author="LeConte" box="[415,628,1772,1795]" page="80" pageId="7" pageNumber="8" year="1857">
<bibRefCitation id="5B7D4B5AFF88FFC6D3FB3EA29C28E672" author="LeConte" box="[415,628,1772,1795]" pageId="7" pageNumber="28" refString="LeConte, J. L. (1857) Synopsis of the species Clivina and allied genera inhabiting the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, [9], 75 - 83." type="journal article" year="1857">LeConte (1857: 80)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3F5336ABFF88FFC6D36C3F429B77E632" blockId="7.[264,1323,1548,1859]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="1FFC1185FF88FFC6D36C3F429B77E632" pageId="7" pageNumber="8">
<taxonomicName id="F8EC4D28FF88FFC6D36C3F429D5AE652" ID-CoL="HH4B" authority="LeConte (1857: 80)" authorityName="LeConte" authorityPageNumber="80" authorityYear="1857" box="[264,774,1804,1827]" class="Insecta" family="Carabidae" genus="Aspidoglossa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="subangulata">
<emphasis id="0D98EAB9FF88FFC6D36C3F429C7DE652" box="[264,545,1804,1827]" italics="true" pageId="7" pageNumber="8">Aspidoglossa subangulata</emphasis>
:
<treatmentCitation id="BE4D10BAFF88FFC6D0543F429D5AE652" author="LeConte" box="[560,774,1804,1827]" page="80" pageId="7" pageNumber="8" year="1857">
<bibRefCitation id="5B7D4B5AFF88FFC6D0543F429D5AE652" author="LeConte" box="[560,774,1804,1827]" pageId="7" pageNumber="28" refString="LeConte, J. L. (1857) Synopsis of the species Clivina and allied genera inhabiting the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, [9], 75 - 83." type="journal article" year="1857">LeConte (1857: 80)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="BE4D10BAFF88FFC6D1773F429DB2E652" author="Putzeys" box="[787,1006,1804,1827]" page="216" pageId="7" pageNumber="8" year="1866">
<bibRefCitation id="5B7D4B5AFF88FFC6D1773F429DB2E652" author="Putzeys" box="[787,1006,1804,1827]" pageId="7" pageNumber="29" refString="Putzeys, J. A. A. H. (1866) Revision generale des clivinides. Annales de la Societe Entomologique de Belgique, 10, 3 - 242." type="journal article" year="1866">Putzeys (1866: 216)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="BE4D10BAFF88FFC6D19D3F429A8CE652" author="LeConte" box="[1017,1232,1804,1827]" page="32" pageId="7" pageNumber="8" year="1879">
<bibRefCitation id="5B7D4B5AFF88FFC6D19D3F429A8CE652" author="LeConte" box="[1017,1232,1804,1827]" pageId="7" pageNumber="28" refString="LeConte, J. L. (1879) Synoptic tables: Ardistomis, Putz.; Aspidoglossa, Putz.; Clivina, Latr.; Schizogenius, Putz. Bulletin of the Brooklyn Entomological Society, 2, 32 - 34." type="journal article" year="1879">LeConte (1879: 32)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="BE4D10BAFF88FFC6D6B83F429F8DE632" author="Blatchley" page="61" pageId="7" pageNumber="8" year="1910">
<bibRefCitation id="5B7D4B5AFF88FFC6D6B83F429F8DE632" author="Blatchley" pageId="7" pageNumber="26" refString="Blatchley, W. S. (1910) An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana - with bibliography and descriptions of new species. The Nature Publishing Co., Indianapolis, 1386 pp." type="book" year="1910">Blatchley (1910: 61)</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="5B7D4B5AFF88FFC6D3B93F629CAAE632" author="Nichols" box="[477,758,1836,1859]" lastPageId="7" lastPageNumber="29" pageId="7" pageNumber="28" refString="Nichols, S. W. (1988 a) Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae). Ph. D. thesis, Cornell University, 393 pp." type="book" year="1988" yearSuffix="a">Nichols (1988a: 105, 115)</bibRefCitation>
;
<treatmentCitation id="BE4D10BAFF88FFC6D1653F629A08E632" author="Downie" box="[769,1108,1836,1859]" page="112" pageId="7" pageNumber="8" year="1996">
<bibRefCitation id="5B7D4B5AFF88FFC6D1653F629A08E632" author="Downie" box="[769,1108,1836,1859]" pageId="7" pageNumber="27" refString="Downie, N. M, &amp; Arnett, RH. Jr. (1996) The beetles of northeastern North America. Volume I: Introduction; suborders Archostemata, Adephaga, and Polyphaga, thru superfamily Cantharoidea. The Sandhill Crane Press, Gainesville, 880 pp." type="book" year="1996">Downie and Arnett (1996: 112)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="BE4D10BAFF88FFC6D63B3F629B7BE632" author="Ciegler" box="[1119,1319,1836,1859]" page="44" pageId="7" pageNumber="8" year="2000">
<bibRefCitation id="5B7D4B5AFF88FFC6D63B3F629B7BE632" author="Ciegler" box="[1119,1319,1836,1859]" pageId="7" pageNumber="27" refString="Ciegler, J. C. (2000). Ground beetles and wrinkled bark beetles of South Carolina (Coleoptera: Geadephaga: Carabidae and Rhysodidae). Biota of South Carolina. Vol. 1. South Carolina: Clemson University, Clemson, 149 pp." type="book" year="2000">Ciegler (2000: 44)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="77F66520FF87FFC9D36C39559CC2E39F" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="3F5336ABFF87FFC9D36C39559C8BE0A7" blockId="8.[264,1325,283,1870]" pageId="8" pageNumber="9">
<emphasis id="0D98EAB9FF87FFC9D36C39559F9EE044" bold="true" box="[264,450,283,309]" pageId="8" pageNumber="9">
<typeStatus id="E0578809FF87FFC9D36C39559F19E044" box="[264,325,283,309]" pageId="8" pageNumber="9">Type</typeStatus>
Material
</emphasis>
. Chaudoir originally described both
<taxonomicName id="F8EC4D28FF87FFC9D1FE39529A99E044" box="[922,1221,284,309]" class="Insecta" family="Carabidae" genus="Dyschirius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="subangulatus">
<emphasis id="0D98EAB9FF87FFC9D1FE39529A99E044" box="[922,1221,284,309]" italics="true" pageId="8" pageNumber="9">Dyschirius subangulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="F8EC4D28FF87FFC9D76A39529F21E02C" class="Insecta" family="Carabidae" genus="Dyschirius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="0D98EAB9FF87FFC9D76A39529F21E02C" italics="true" pageId="8" pageNumber="9">D. humeralis</emphasis>
</taxonomicName>
from an unspecified number of specimens. One specimen in
<collectionCode id="59FDAE6EFF87FFC9D632390A9AF7E02F" box="[1110,1195,324,350]" httpUri="http://grbio.org/cool/a5xg-x4h4" name="Museum d'Histoire Naturelle Perpignan" pageId="8" pageNumber="9">MHNP</collectionCode>
is labelled &quot;
<taxonomicName id="F8EC4D28FF87FFC9D37439229C4AE0F7" authority="Chaud." authorityName="Chaud." box="[272,534,364,390]" class="Insecta" family="Carabidae" genus="Dyschirius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="subangulatus">subangulatus Chaud.</taxonomicName>
[handwritten]/ Louisiane Chevrolat [handwritten]/
<typeStatus id="E0578809FF87FFC9D6D139229B76E0F7" box="[1205,1322,364,390]" pageId="8" pageNumber="9" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="F8EC4D28FF87FFC9D36C39DA9DFEE0DF" authority="Chd. Des. S.W. Nichols 1984" authorityName="Chd. Des. S.W. Nichols" authorityYear="1984" box="[264,930,404,430]" class="Insecta" family="Carabidae" genus="Dyschirius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="subangulatus">Dyschirius subangulatus Chd. Des. S.W. Nichols 1984</taxonomicName>
&quot; (
<emphasis id="0D98EAB9FF87FFC9D1DA39DA9DB6E0DC" box="[958,1002,404,429]" italics="true" pageId="8" pageNumber="9">fide</emphasis>
<bibRefCitation id="5B7D4B5AFF87FFC9D19139DA9AFBE0DF" author="Nichols" box="[1013,1191,404,430]" lastPageId="8" lastPageNumber="29" pageId="8" pageNumber="28" refString="Nichols, S. W. (1988 a) Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae). Ph. D. thesis, Cornell University, 393 pp." type="book" year="1988" yearSuffix="a">Nichols 1988a</bibRefCitation>
: 116). The
<typeStatus id="E0578809FF87FFC9D36C39F29F2FE0A7" box="[264,371,444,470]" pageId="8" pageNumber="9" type="lectotype">lectotype</typeStatus>
designation is here published.
</paragraph>
<paragraph id="3F5336ABFF87FFC9D35C39AA9CC2E39F" blockId="8.[264,1325,283,1870]" pageId="8" pageNumber="9">
Putzeys originally described
<taxonomicName id="F8EC4D28FF87FFC9D0FE39AA9DD6E08C" box="[666,906,484,509]" class="Insecta" family="Carabidae" genus="Aspidoglossa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="0D98EAB9FF87FFC9D0FE39AA9DD6E08C" box="[666,906,484,509]" italics="true" pageId="8" pageNumber="9">Aspidoglossa vicina</emphasis>
</taxonomicName>
based on two male specimens in Chevrolat's collection (which is now housed in the Hope Entomological Collections, The University Museum, at Oxford,
<collectingCountry id="47FB763BFF87FFC9D0EA3A7A9CA9E33F" box="[654,757,564,590]" name="United Kingdom" pageId="8" pageNumber="9">England</collectingCountry>
). He originally described
<taxonomicName id="F8EC4D28FF87FFC9D6553A7A9AE9E33C" box="[1073,1205,564,589]" class="Insecta" family="Carabidae" genus="Aspidoglossa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="0D98EAB9FF87FFC9D6553A7A9AE9E33C" box="[1073,1205,564,589]" italics="true" pageId="8" pageNumber="9">A. fraterna</emphasis>
</taxonomicName>
based on seven specimens in Dejean (housed in
<collectionCode id="59FDAE6EFF87FFC9D0B53A129D76E307" box="[721,810,604,630]" httpUri="http://grbio.org/cool/a5xg-x4h4" name="Museum d'Histoire Naturelle Perpignan" pageId="8" pageNumber="9">MHNP</collectionCode>
), Hope (housed in the Hope Entomological Collections), Buquet (location unknown to me) and Schaum (housed in the Zoologisches Museum, Berlin,
<collectingCountry id="47FB763BFF87FFC9D3B13AE29C18E3B7" box="[469,580,684,710]" name="Germany" pageId="8" pageNumber="9">Germany</collectingCountry>
) collections. I have not seen any
<typeStatus id="E0578809FF87FFC9D1AF3AE29A6EE3B7" box="[971,1074,684,710]" pageId="8" pageNumber="9" type="syntype">syntypes</typeStatus>
of these two taxa, or of
<taxonomicName id="F8EC4D28FF87FFC9D34C3A9A9CC2E39F" authority="Chaudoir." authorityName="Chaudoir." box="[296,670,724,750]" class="Insecta" family="Carabidae" genus="Dyschirius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="0D98EAB9FF87FFC9D34C3A9A9C7FE39C" box="[296,547,724,749]" italics="true" pageId="8" pageNumber="9">Dyschirius humeralis</emphasis>
Chaudoir.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="77F66520FF87FFC9D35C3AB29C16E217" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="3F5336ABFF87FFC9D35C3AB29C16E217" blockId="8.[264,1325,283,1870]" pageId="8" pageNumber="9">
<emphasis id="0D98EAB9FF87FFC9D35C3AB29FECE267" bold="true" box="[312,432,764,790]" pageId="8" pageNumber="9">Diagnosis</emphasis>
. Distinguished from all other Nearctic Ardistomina by the presence of a row of punctures along the metepisternal suture on the metasternum and the proeminent lateral lobes of the clypeus.
</paragraph>
</subSubSection>
<subSubSection id="77F66520FF87FFC8D35C3B3A9DADE0DF" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="3F5336ABFF87FFC9D35C3B3A9D47E437" blockId="8.[264,1325,283,1870]" pageId="8" pageNumber="9">
<emphasis id="0D98EAB9FF87FFC9D35C3B3A9F95E2FF" bold="true" box="[312,457,884,910]" pageId="8" pageNumber="9">Description</emphasis>
. Color: Upper surface black except head capsule with sides of frons and clypeus and labrum reddish­brown; elytron with preapical reddish spot on intervals 4–7 extended narrowly to apex; antennae and palpi pale reddish­brown; legs reddish­brown. Microsculpture: Pronotum and elytra devoid of microlines, smooth. Head: Eyes large and hemispherical. Frons with more or less V­shaped groove varied from distinct to suggested only. Antennomeres 5–10 longer than broad. Prothorax: Pronotum with midline not margined on each side. Elytra: Strial punctures coarse but progressively finer toward apex. Interval 3 with 6–10 setae, interval 5 with 5–8 setae. Abdomen: Last visible sternite with 4 setae, distance between 2 setae on each side shorter than that between medial setae. Male Genitalia: Median lobe elongate (
<figureCitation id="A7D72A2EFF87FFC9D0F73C929C86E587" box="[659,730,1244,1270]" captionStart="FIGURES 4 – 8" captionStartId="12.[264,382,1796,1820]" captionTargetBox="[361,1232,331,1758]" captionTargetId="figure@12.[335,1252,306,1791]" captionTargetPageId="12" captionText="FIGURES 4 – 8. Median lobe (a) and paremeres (b). 4, Aspidoglossa subangulata; 5, Semiardistomis viridis; 6, S. puncticollis; 7, Ardistomis schaumii; 8, A. obliquatus." httpUri="https://zenodo.org/record/173805/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
a), parameres markedly long and narrrow, more or less similar (
<figureCitation id="A7D72A2EFF87FFC9D3FB3D4A9FB4E46F" box="[415,488,1284,1310]" captionStart="FIGURES 4 – 8" captionStartId="12.[264,382,1796,1820]" captionTargetBox="[361,1232,331,1758]" captionTargetId="figure@12.[335,1252,306,1791]" captionTargetPageId="12" captionText="FIGURES 4 – 8. Median lobe (a) and paremeres (b). 4, Aspidoglossa subangulata; 5, Semiardistomis viridis; 6, S. puncticollis; 7, Ardistomis schaumii; 8, A. obliquatus." httpUri="https://zenodo.org/record/173805/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
b). Female Genitalia: Apical stylomere as illustrated (
<figureCitation id="A7D72A2EFF87FFC9D6053D4A9AE4E46F" box="[1121,1208,1284,1310]" captionStart="FIGURES 11 – 18. 11 – 14" captionStartId="14.[264,382,1651,1675]" captionTargetBox="[372,1224,323,1650]" captionTargetId="figure@14.[356,1238,316,1655]" captionTargetPageId="14" captionText="FIGURES 11 – 18. 11 – 14. Apical stylomere. 11, Aspidoglossa subangulata; 12, Semiardistomis viridis; 13, Ardistomis schaumii; 14, A. obliquatus. 15 – 18. Spermatheca. 15, Aspidoglossa subangulata; 16, Semiardistomis viridis; 17, S. puncticollis; 18, Ardistomis schaumii." httpUri="https://zenodo.org/record/173807/files/figure.png" pageId="8" pageNumber="9">Fig. 11</figureCitation>
); seminal tube of spermatheca markedly long (
<figureCitation id="A7D72A2EFF87FFC9D0D13D629D50E437" box="[693,780,1324,1350]" captionStart="FIGURES 11 – 18. 11 – 14" captionStartId="14.[264,382,1651,1675]" captionTargetBox="[372,1224,323,1650]" captionTargetId="figure@14.[356,1238,316,1655]" captionTargetPageId="14" captionText="FIGURES 11 – 18. 11 – 14. Apical stylomere. 11, Aspidoglossa subangulata; 12, Semiardistomis viridis; 13, Ardistomis schaumii; 14, A. obliquatus. 15 – 18. Spermatheca. 15, Aspidoglossa subangulata; 16, Semiardistomis viridis; 17, S. puncticollis; 18, Ardistomis schaumii." httpUri="https://zenodo.org/record/173807/files/figure.png" pageId="8" pageNumber="9">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="3F5336ABFF87FFC9D35C3D1A9C37E41F" blockId="8.[264,1325,283,1870]" box="[312,619,1364,1390]" pageId="8" pageNumber="9">Body length: 6.5–8.0 mm.</paragraph>
<paragraph id="3F5336ABFF87FFC9D35C3D329A17E7DF" blockId="8.[264,1325,283,1870]" pageId="8" pageNumber="9">
<emphasis id="0D98EAB9FF87FFC9D35C3D329CDFE4E7" bold="true" box="[312,643,1404,1430]" pageId="8" pageNumber="9">Geographical Distribution</emphasis>
. This species ranges over eastern
<collectingCountry id="47FB763BFF87FFC9D67F3D329AE5E4E7" box="[1051,1209,1404,1430]" name="United States of America" pageId="8" pageNumber="9">United States</collectingCountry>
(
<figureCitation id="A7D72A2EFF87FFC9D6AF3D329B7EE4E7" box="[1227,1314,1404,1430]" captionStart="FIGURE 31" captionStartId="19.[264,367,1030,1054]" captionTargetBox="[338,1248,327,1006]" captionTargetId="figure@19.[338,1248,326,1006]" captionTargetPageId="19" captionText="FIGURE 31. Collection localities for Aspidoglossa subangulata (Chaudoir) in United States of America." httpUri="https://zenodo.org/record/173811/files/figure.png" pageId="8" pageNumber="9">Fig. 31</figureCitation>
) from Nebraska to Washington D.C., south to southern Florida (not yet recorded from the Keys) and northeastern
<collectingCountry id="47FB763BFF87FFC9D0503D829CCDE497" box="[564,657,1484,1510]" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingCountry>
, west to southeastern Arizona. I have also seen one specimen, in AMNH, labelled &quot;Oregon, Summit Lake [Klamath Co.],
<date id="4B52106BFF87FFC9D6393DBA9AB2E77F" box="[1117,1262,1524,1550]" pageId="8" pageNumber="9" value="1962-07-20">20.VII.1962</date>
, J.D. Vertrees&quot;, and another one, in CAS, labelled &quot;Montana. Gallatin Co., West Yellowstone,
<date id="4B52106BFF87FFC9D36C3E0A9FCBE72F" box="[264,407,1604,1630]" pageId="8" pageNumber="9" value="1949-07-06">6–VII–1949</date>
Durham Giuliani C.A.S. Accession&quot;. Whether or not these two specimens are mislabelled or represent legitimate records cannot be answered at the moment. This species has been also recorded from one locality in
<collectingCountry id="47FB763BFF87FFC9D1013EDA9DF4E7DF" box="[869,936,1684,1710]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
(
<bibRefCitation id="5B7D4B5AFF87FFC9D1D33EDA9A62E7DF" author="Habu" box="[951,1086,1684,1710]" pageId="8" pageNumber="28" refString="Habu, A. (1963) Notes on Aspidoglossa subangulata (Chaudoir) found in Japan [in Japanese]. Akitu, 11, 19 - 20." type="journal article" year="1963">Habu 1963</bibRefCitation>
).
</paragraph>
<paragraph id="3F5336ABFF87FFC8D35C3EF29A9FE047" blockId="8.[264,1325,283,1870]" lastBlockId="9.[264,1322,284,430]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="0D98EAB9FF87FFC9D35C3EF29FC4E7A7" bold="true" box="[312,408,1724,1750]" pageId="8" pageNumber="9">Habitat</emphasis>
.
<bibRefCitation id="5B7D4B5AFF87FFC9D3CF3EF29C2BE7A7" author="Ball" box="[427,631,1724,1750]" pageId="8" pageNumber="26" refString="Ball, G. E. (1960) Carabidae (Latreille, 1810). In: Arnett, R. H. Jr., The beetles of the United States (a manual for identification). Part 1. Archestomata and Adephaga. The Catholic University of America Press, Washington DC, pp. 55 - 181." type="book chapter" year="1960">Ball (1960: 109)</bibRefCitation>
reported that members of this species are found among sparse vegetation on wet clay near the margins of ponds and streams, and that they come to light on warm nights during the summer. They were collected on an eroding river bank composed of clay and sand in northern Florida (
<bibRefCitation id="5B7D4B5AFF87FFC9D12B3F7A9AF5E63F" author="Choate" box="[847,1193,1844,1870]" pageId="8" pageNumber="27" refString="Choate, P. M. &amp; Rogers, T. (1976) The occurrence of a troglophilic ground beetle in Florida (Coleoptera: Carabidae). The Coleopterists Bulletin, 30, 364." type="journal article" year="1976">Choate &amp; Rogers 1976: 364</bibRefCitation>
).
<bibRefCitation id="5B7D4B5AFF87FFC8D6DB3F7A9C0CE047" author="Ferguson" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="28" refString="Ferguson, H. J. &amp; McPherson, R. M. (1985) Abundance and diversity of adult Carabidae in four soybean cropping systems in Virginia. Journal of Entomological Science, 20,163 - 71." type="book" year="1985">Ferguson and McPherson (1985: 165)</bibRefCitation>
reported this species from soybean fields in Virginia.
</paragraph>
<paragraph id="3F5336ABFF86FFC8D35C390D9DADE0DF" blockId="9.[264,1322,284,430]" pageId="9" pageNumber="10">
<emphasis id="0D98EAB9FF86FFC8D35C390D9F2EE02C" bold="true" box="[312,370,323,349]" pageId="9" pageNumber="10">Note</emphasis>
. The male differs from the female by having the protarsomeres 2–4 larger and with long, flat, translucent setae underneath (
<figureCitation id="A7D72A2EFF86FFC8D17E39229D2FE0F7" box="[794,883,364,390]" captionStart="FIGURES 23 – 26. 23 – 24" captionStartId="16.[264,382,1139,1163]" captionTargetBox="[267,1323,326,1113]" captionTargetId="figure@16.[264,1323,326,1113]" captionTargetPageId="16" captionText="FIGURES 23 – 26. 23 – 24. Protarsomeres (ventral view). 23, Aspidoglossa subangulata ɗ; 24, Aspidoglossa subangulata Ψ. 25 – 26. Humeral region (dorsal view). 25, Aspidoglossa subangulata; 26, Semiardistomis viridis. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/173809/files/figure.png" pageId="9" pageNumber="10">Fig. 23</figureCitation>
). In the female the protarsomeres are narrower and have long but more or less regular setae (
<figureCitation id="A7D72A2EFF86FFC8D1EF39DA9DBFE0DF" box="[907,995,404,430]" captionStart="FIGURES 23 – 26. 23 – 24" captionStartId="16.[264,382,1139,1163]" captionTargetBox="[267,1323,326,1113]" captionTargetId="figure@16.[264,1323,326,1113]" captionTargetPageId="16" captionText="FIGURES 23 – 26. 23 – 24. Protarsomeres (ventral view). 23, Aspidoglossa subangulata ɗ; 24, Aspidoglossa subangulata Ψ. 25 – 26. Humeral region (dorsal view). 25, Aspidoglossa subangulata; 26, Semiardistomis viridis. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/173809/files/figure.png" pageId="9" pageNumber="10">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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