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<mods:title id="E8AEC37444AE935F64CC700B9AE49555">Desmosomatidae and Nannoniscidae (Crustacea, Isopoda, Asellota) from bathyal and abyssal depths oOE North Carolina and their evolution</mods:title>
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<treatment id="03E1535BFFA924543FF954D6FEC4FA6A" ID-DOI="http://doi.org/10.5281/zenodo.5275761" ID-Zenodo-Dep="5275761" LSID="urn:lsid:plazi:treatment:03E1535BFFA924543FF954D6FEC4FA6A" httpUri="http://treatment.plazi.org/id/03E1535BFFA924543FF954D6FEC4FA6A" lastPageId="15" lastPageNumber="1844" pageId="14" pageNumber="1843">
<subSubSection id="C352B1C6FFA924553FF954D6FD68F9B3" box="[421,721,1521,1546]" pageId="14" pageNumber="1843" type="nomenclature">
<paragraph id="8BF7E24DFFA924553FF954D6FD68F9B3" blockId="14.[421,721,1521,1546]" box="[421,721,1521,1546]" pageId="14" pageNumber="1843">
<heading id="D0BF5521FFA924553FF954D6FD68F9B3" box="[421,721,1521,1546]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="1843" reason="3">
<emphasis id="B93C3E5FFFA924553FF954D6FDC3F9B0" bold="true" box="[421,634,1521,1545]" pageId="14" pageNumber="1843">
Genus
<taxonomicName id="4C4899CEFFA924553FAF54D6FDC3F9B0" ID-CoL="41181132-8620-4b9a-afe4-b3f7719b7a6b" authority="George, 2001" authorityName="George" authorityYear="2001" box="[499,634,1521,1545]" class="Malacostraca" family="Nannoniscidae" genus="Leutziniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1843" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93C3E5FFFA924553FAF54D6FDC3F9B0" bold="true" box="[499,634,1521,1545]" italics="true" pageId="14" pageNumber="1843">Leutziniscus</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20F8324FFA924553CD854D5FD68F9B3" box="[644,721,1522,1546]" pageId="14" pageNumber="1843" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C352B1C6FFA924543EDA5700FEC4FA6A" lastPageId="15" lastPageNumber="1844" pageId="14" pageNumber="1843" type="diagnosis">
<paragraph id="8BF7E24DFFA924543EDA5700FF76FBBC" blockId="14.[93,1049,1574,1662]" lastBlockId="15.[93,1049,973,1491]" lastPageId="15" lastPageNumber="1844" pageId="14" pageNumber="1843">
<emphasis id="B93C3E5FFFA924553EDA5700FF41F987" box="[134,248,1575,1598]" italics="true" pageId="14" pageNumber="1843">Diagnosis.</emphasis>
<taxonomicName id="4C4899CEFFA924553F5F5701FECBF987" authorityName="George" authorityYear="2001" box="[259,370,1574,1598]" class="Malacostraca" family="Nannoniscidae" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="1852" phylum="Arthropoda" rank="subFamily" subFamily="Leutzinae">Leutzinae</taxonomicName>
with sixth and seventh pereonites free. Antennula with 11 articles, terminal article not bulbous. Body not ¯at or broad. Pleon in adult female with posterolateral spines. Female operculum lacking ventral medial spine or hook, and posterior margin broadly rounded and fringed with small setae. Uropods biramous.
</paragraph>
<caption id="DF37B2C5FFA824543E01520EFE8DFC10" ID-DOI="http://doi.org/10.5281/zenodo.5275972" ID-Zenodo-Dep="5275972" httpUri="https://zenodo.org/record/5275972/files/figure.png" pageId="15" pageNumber="1844" startId="15.[93,109,809,830]" targetBox="[102,1043,107,782]" targetPageId="15">
<paragraph id="8BF7E24DFFA824543E01520EFE8DFC10" blockId="15.[93,1049,809,937]" pageId="15" pageNumber="1844">
FIG. 6. Five genera of the subfamily
<taxonomicName id="4C4899CEFFA824543FA1520EFD2CFC86" authorityName="Hansen" authorityYear="1916" box="[509,661,809,831]" class="Malacostraca" family="Nannoniscidae" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="family">Nannoniscidae</taxonomicName>
. (A)
<taxonomicName id="4C4899CEFFA824543C85520DFCECFC86" authorityName="G. O. Sars" authorityYear="1870" box="[729,853,810,831]" class="Malacostraca" family="Nannoniscidae" genus="Nannoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543C85520DFCECFC86" box="[729,853,810,831]" italics="true" pageId="15" pageNumber="1844">Nannoniscus</emphasis>
</taxonomicName>
e.g.
<taxonomicName id="4C4899CEFFA824543DCD520DFE73FCE0" authority="(Menzies and George, 1972)" baseAuthorityName="Menzies and George" baseAuthorityYear="1972" class="Malacostraca" family="Nannoniscidae" genus="Nannoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="species" species="muscarius">
<emphasis id="B93C3E5FFFA824543DCD520DFBA0FC86" box="[913,1049,810,831]" italics="true" pageId="15" pageNumber="1844">N. muscarius</emphasis>
(
<bibRefCitation id="EFD99FBCFFA824543EF35264FE04FCE0" author="MENZIES, R. J. &amp; GEORGE, R. Y." box="[175,445,835,857]" pageId="15" pageNumber="1844" pagination="124" refId="ref12265" refString="MENZIES, R. J. and GEORGE, R. Y., 1972, Isopod Crustacea of the Peru-Chile Trench, Anton Bruun Report 9, 1 ± 124." type="journal article" year="1972">Menzies and George, 1972</bibRefCitation>
)
</taxonomicName>
. (B)
<taxonomicName id="4C4899CEFFA824543FA15264FDC5FCE0" authorityName="Siebenaller and Hessler" authorityYear="1981" box="[509,636,835,857]" class="Malacostraca" family="Nannoniscidae" genus="Regabellator" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543FA15264FDC5FCE0" box="[509,636,835,857]" italics="true" pageId="15" pageNumber="1844">Regabellator</emphasis>
</taxonomicName>
e.g.
<taxonomicName id="4C4899CEFFA824543CF05263FF6EFCCD" authority="Siebenaller and Hessler, 1881" authorityName="Siebenaller and Hessler" authorityYear="1881" class="Malacostraca" family="Nannoniscidae" genus="Regabellator" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="species" species="profugus">
<emphasis id="B93C3E5FFFA824543CF05263FCA6FCE0" box="[684,799,835,857]" italics="true" pageId="15" pageNumber="1844">R. profugus</emphasis>
Siebenaller and Hessler, 1881
</taxonomicName>
. (C)
<emphasis id="B93C3E5FFFA824543F495278FE3AFCCD" box="[277,387,863,884]" italics="true" pageId="15" pageNumber="1844">Raponiscus</emphasis>
e.g.
<taxonomicName id="4C4899CEFFA824543FE35278FD59FCCD" authority="(Hansen, 1916)" baseAuthorityName="Hansen" baseAuthorityYear="1916" box="[447,736,862,884]" class="Malacostraca" family="Nannoniscidae" genus="Regabellator" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="species" species="crasipes">
<emphasis id="B93C3E5FFFA824543FE35278FD97FCCD" box="[447,558,863,884]" italics="true" pageId="15" pageNumber="1844">R. crasipes</emphasis>
(
<bibRefCitation id="EFD99FBCFFA824543C185278FD6AFCCD" author="HANSEN, H. J." box="[580,723,862,884]" pageId="15" pageNumber="1844" pagination="1" refId="ref11946" refString="HANSEN, H. J., 1916, Crustacea Malacostraca III. Isopoda, Danish Ingolf Expedition, 3 (5), 1 ± 162." type="journal article" year="1916">Hansen, 1916</bibRefCitation>
)
</taxonomicName>
. (D)
<taxonomicName id="4C4899CEFFA824543D435279FC30FCCD" authorityName="Siebenaller and Hessler" authorityYear="1977" box="[799,905,862,884]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543D435279FC30FCCD" box="[799,905,862,884]" italics="true" pageId="15" pageNumber="1844">Hebefustis</emphasis>
</taxonomicName>
e.g.
<taxonomicName id="4C4899CEFFA824543D9E5278FE6CFC37" authority="Siebenaller and Hessler, 1977" authorityName="Siebenaller and Hessler" authorityYear="1977" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="species" species="vafer">
<emphasis id="B93C3E5FFFA824543D9E5278FBA0FCCD" box="[962,1049,862,884]" italics="true" pageId="15" pageNumber="1844">H. vafer</emphasis>
<bibRefCitation id="EFD99FBCFFA824543EF9525FFE6CFC37" author="SIEBENALLER, J. F. &amp; HESSLER, R. R." box="[165,469,888,910]" pageId="15" pageNumber="1844" pagination="17" refId="ref12635" refString="SIEBENALLER, J. F. and HESSLER, R. R., 1977, The Nannoniscidae (Isopoda, Asellota) Hebefestus n. gen. and Nannoniscoides Hansen, Transactions of San Diego Society of Natural History, 19 (2), 17 ± 44." type="journal article" year="1977">Siebenaller and Hessler, 1977</bibRefCitation>
</taxonomicName>
. (E)
<taxonomicName id="4C4899CEFFA824543C4E525FFD12FC37" authorityName="Hansen" authorityYear="1916" box="[530,683,888,910]" class="Malacostraca" family="Nannoniscidae" genus="Nannoniscoides" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543C4E525FFD12FC37" box="[530,683,888,910]" italics="true" pageId="15" pageNumber="1844">Nannoniscoides</emphasis>
</taxonomicName>
e.g.
<emphasis id="B93C3E5FFFA824543CBD525EFCCBFC37" box="[737,882,888,910]" italics="true" pageId="15" pageNumber="1844">N. latediOEusus</emphasis>
<bibRefCitation id="EFD99FBCFFA824543D26525FFE94FC10" author="SIEBENALLER, J. F. &amp; HESSLER, R. R." pageId="15" pageNumber="1844" pagination="17" refId="ref12635" refString="SIEBENALLER, J. F. and HESSLER, R. R., 1977, The Nannoniscidae (Isopoda, Asellota) Hebefestus n. gen. and Nannoniscoides Hansen, Transactions of San Diego Society of Natural History, 19 (2), 17 ± 44." type="journal article" year="1977">Siebenaller and Hessler, 1977</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8BF7E24DFFA824543EDA552CFD0AFBC6" blockId="15.[93,1049,973,1491]" pageId="15" pageNumber="1844">
<emphasis id="B93C3E5FFFA824543EDA552CFF4EFB9A" box="[134,247,1035,1059]" italics="true" pageId="15" pageNumber="1844">Etymology</emphasis>
. This new genus is named in honour of Dr James R. Leutze, Chancellor of the University of
<collectingRegion id="498C2CAFFFA824543F28550DFD88FBFB" box="[372,561,1066,1090]" country="United States of America" name="North Carolina" pageId="15" pageNumber="1844">North Carolina</collectingRegion>
at Wilmington, in recognition of his concern for the health of the environment and his contributions to international understanding through the`Globe Watch Program.
</paragraph>
<paragraph id="8BF7E24DFFA824543EDA55A1FC74FA81" blockId="15.[93,1049,973,1491]" pageId="15" pageNumber="1844">
<emphasis id="B93C3E5FFFA824543EDA55A1FF50FB27" box="[134,233,1158,1182]" italics="true" pageId="15" pageNumber="1844">Remarks</emphasis>
. This new genus
<taxonomicName id="4C4899CEFFA824543F8C55A0FDEEFB27" authorityName="George" authorityYear="2001" box="[464,599,1159,1182]" class="Malacostraca" family="Nannoniscidae" genus="Leutziniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543F8C55A0FDEEFB27" box="[464,599,1159,1182]" italics="true" pageId="15" pageNumber="1844">Leutziniscus</emphasis>
</taxonomicName>
belongs to a cluster of genera, i.e.
<taxonomicName id="4C4899CEFFA824543E015581FF51FB07" authorityName="Vanhoffen" authorityYear="1914" box="[93,232,1190,1214]" class="Malacostraca" family="Nannoniscidae" genus="Austroniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543E015581FF51FB07" box="[93,232,1190,1214]" italics="true" pageId="15" pageNumber="1844">Austroniscus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4899CEFFA824543F685581FE5EFB07" authorityName="Hessler" authorityYear="1970" box="[308,487,1190,1214]" class="Malacostraca" family="Nannoniscidae" genus="Thaumastosoma" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543F685581FE5EFB07" box="[308,487,1190,1214]" italics="true" pageId="15" pageNumber="1844">Thaumastosoma</emphasis>
</taxonomicName>
of the subfamily
<taxonomicName id="4C4899CEFFA824543C8D5581FCF9FB07" authorityName="George" authorityYear="2001" box="[721,832,1190,1214]" class="Malacostraca" family="Nannoniscidae" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="subFamily" subFamily="Leutzinae">Leutzinae</taxonomicName>
, characterized by antennul a lacking the bulbous terminal article and with ®ve or more than ®ve articles in the antennula. Among these genera
<taxonomicName id="4C4899CEFFA824543C4155C4FD11FB42" authorityName="Vanhoffen" authorityYear="1914" box="[541,680,1251,1275]" class="Malacostraca" family="Nannoniscidae" genus="Austroniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543C4155C4FD11FB42" box="[541,680,1251,1275]" italics="true" pageId="15" pageNumber="1844">Austroniscus</emphasis>
</taxonomicName>
is an exception to the rule with a ¯at and broad body. Sexual dimorphism is seen in the pleon con®guration with either males or females having posterolateral spines in some of these genera.
</paragraph>
<paragraph id="8BF7E24DFFA824543EDA5467FEC4FA6A" blockId="15.[93,1049,973,1491]" pageId="15" pageNumber="1844">
In this new genus the pleon of the female has posterolateral spines. Species of the genera
<taxonomicName id="4C4899CEFFA824543EBA5478FE20FACE" authorityName="Hessler" authorityYear="1970" box="[230,409,1375,1399]" class="Malacostraca" family="Nannoniscidae" genus="Thaumastosoma" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543EBA5478FE20FACE" box="[230,409,1375,1399]" italics="true" pageId="15" pageNumber="1844">Thaumastosoma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4899CEFFA824543FBD5447FDD1FACE" authorityName="George" authorityYear="2001" box="[481,616,1376,1399]" class="Malacostraca" family="Nannoniscidae" genus="Leutziniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543FBD5447FDD1FACE" box="[481,616,1376,1399]" italics="true" pageId="15" pageNumber="1844">Leutziniscus</emphasis>
</taxonomicName>
lack a ventral spine on the female operculum.
<taxonomicName id="4C4899CEFFA824543EB1545AFE19FA2C" authorityName="Hessler" authorityYear="1970" box="[237,416,1405,1429]" class="Malacostraca" family="Nannoniscidae" genus="Thaumastosoma" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543EB1545AFE19FA2C" box="[237,416,1405,1429]" italics="true" pageId="15" pageNumber="1844">Thaumastosoma</emphasis>
</taxonomicName>
is, however, distinguished from
<taxonomicName id="4C4899CEFFA824543D665459FC78FA2C" authorityName="George" authorityYear="2001" box="[826,961,1406,1429]" class="Malacostraca" family="Nannoniscidae" genus="Leutziniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="1844" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3E5FFFA824543D665459FC78FA2C" box="[826,961,1406,1429]" italics="true" pageId="15" pageNumber="1844">Leutziniscus</emphasis>
</taxonomicName>
in the female pleon con®guration, the former lacking posterolateral spines on the pleon and the latter with them.
</paragraph>
</subSubSection>
</treatment>
</document>