treatments-xml/data/F5/52/87/F55287D4FFA53F5FFC23FCE8FEA0F961.xml

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<mods:title id="2136F3053BCF9375F8FD75AD1A11CE51">Scorpions of the Horn of Africa (Arachnida Scorpiones) Part XXI Parabuthus (Buthidae) (Part II), with description of five new species from Somaliland and Ethiopia</mods:title>
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<taxonomicName id="BAFB4D41FFA53F58FC23FCE8FB36FC20" ID-CoL="4CN5H" authority="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský, 2019" authorityName="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský" authorityYear="2019" box="[997,1226,874,898]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="mazuchi" status="sp. nov.">Parabuthus mazuchi</taxonomicName>
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(
<figureCitation id="E5C02A47FFA53F58FCA0FC09FBFFFC00" box="[870,1027,906,931]" captionStart="Figures 150–160" captionStartId="30.[113,190,1886,1908]" captionTargetBox="[128,1454,258,1831]" captionTargetId="figure-18@30.[113,1474,248,1851]" captionTargetPageId="30" captionText="Figures 150–160: Hemispermatophores. Figures 150–155. Parabuthus kabateki sp. n., male paratype 1549, right hemispermatophore, distal trunk, capsule and flagellum (150), capsule in posterior (151), convex (152), convex compressed (153), anterior (154) views, and whole hemispermatophore (155). Scale bars: 1 mm (150), 500 μm (151–154), 4 mm (155). Figures 156–160. Parabuthus mazuchi sp. n., male holotype, left hemispermatophore, whole hemispermatophore (156), capsule in posterior (157), convex (158), convex compressed (159), and anterior (160) views. Scale bars: 4 mm (156), 500μm (157–160)." pageId="33" pageNumber="32">Figs. 156–160</figureCitation>
, 
<figureCitation id="E5C02A47FFA53F58FBCBFC09FB91FC01" box="[1037,1133,907,931]" captionStart-0="Figures 163–164" captionStart-1="Figures 165–166" captionStart-2="Figures 167–171" captionStart-3="Figures 172–187" captionStart-4="Figures 188–189" captionStartId-0="32.[115,192,2011,2033]" captionStartId-1="34.[113,190,1574,1596]" captionStartId-2="35.[115,192,1988,2010]" captionStartId-3="36.[116,193,1936,1958]" captionStartId-4="37.[113,190,2020,2042]" captionTargetBox-0="[115,1476,226,1981]" captionTargetBox-1="[129,1463,236,1546]" captionTargetBox-2="[115,1476,215,1960]" captionTargetBox-3="[115,1469,244,1906]" captionTargetBox-4="[113,1474,234,2004]" captionTargetId-0="figure-18@32.[115,1476,226,1983]" captionTargetId-1="figure-228@34.[113,1474,215,1573]" captionTargetId-2="figure-9@35.[115,1476,213,1970]" captionTargetId-3="figure-19@36.[113,1474,234,1915]" captionTargetId-4="figure-9@37.[113,1474,234,2006]" captionTargetPageId-0="32" captionTargetPageId-1="34" captionTargetPageId-2="35" captionTargetPageId-3="36" captionTargetPageId-4="37" captionText-0="Figures 163–164. Somaliland, locality 18SJ, type locality of Parabuthus kabateki sp. n. and P. mazuchi sp. n." captionText-1="Figures 165–166. Parabuthus mazuchi sp. n., male holotype in dorsal (165) and ventral (166) views. Scale bar: 10 mm." captionText-2="Figures 167–171: Parabuthus mazuchi sp. n., male holotype, posterior coxosternal area and sternite III (167) carapace and tergites I–III (168), metasoma and telson in lateral (169), ventral (170), and dorsal (171) views. Scale bar: 10 mm (169–171)." captionText-3="Figures 172–187: Parabuthus mazuchi sp. n., male holotype. Figures 172–181. Pedipalp chela dorsal (172), external (173) and ventral (174) views, pedipalp patella dorsal (175), external (176) and ventral (177) views, pedipalp femur and trochanter internal (178) and dorsal (179) views, pedipalp movable (180) and fixed (181) finger dentition. Trichobothrial pattern is indicated in Figures 173–176, 178–179. Figures 182–183. Left chelicera ventral (182) and dorsal (183) views. Figures 184–187. Left legs I–IV, retrolateral aspect, respectively." captionText-4="Figures 188–189. Parabuthus mazuchi sp. n., male holotype in vivo habitus at type locality." pageId="33" pageNumber="32">163–189</figureCitation>
, 
<figureCitation id="E5C02A47FFA53F58FBBEFC09FB24FC01" box="[1144,1240,907,931]" captionStart="Figures 266–292" captionStartId="59.[113,190,1844,1866]" captionTargetBox="[118,1461,239,1828]" captionTargetId="figure-9@59.[113,1471,233,1828]" captionTargetPageId="59" captionText="Figures 266–292: Parabuthus chelae, dorsal and external views. Figures 266–267. P. heterurus, male holotype (266) and male from Somalia, Afgooye (267). Figure 268. P. kabateki sp. n., male holotype. Figure 269. P. kajibu, male holotype. Figure 270–271. P. robustus sp. n., male holotype (270) and paratype from locality 11SQ (271). Figures 272–273. P. somalilandus sp. n., male holotype (272) and paratype from Laas Gel (273). Figures 274–277. P. abyssinicus, male (274–275) and female (276–277) from locality 17SR. Figures 278–279. P. erigavoensis sp. n., female holotype. Figures 280–283. P. hamar, male (280–281) and female (282–283) paratypes. Figures 284–285. P. cimrmani, male holotype. Figures 286–287. P. eritreaensis, male holotype. Figures 288–289. P. mazuchi sp. n., male holotype. Figures 290–292. P. granimanus, male from locality 18ST (290–291) and male lektotype (292)." pageId="33" pageNumber="32">288–289</figureCitation>
, 
<figureCitation id="E5C02A47FFA53F58FB24FC08FAF0FC00" box="[1250,1292,906,930]" captionStart="Figure 305" captionStartId="62.[113,181,1336,1358]" captionTargetBox="[117,1471,237,1321]" captionTargetId="figure-216@62.[113,1474,232,1321]" captionTargetPageId="62" captionText="Figure 305. Map showing confirmed distribution of Parabuthus spp. in Djibouti, Eritrea, Ethiopia, Somalia, and Somaliland. Most points indicate sites sampled during 2011–2019 expeditions. We have not included in the map locality data for old specimens mainly from Somalia deposited in Italian museums which include large collections from this region (KovařÍk &amp; Whitman, 2005: 110–112), because these specimens are often mislabeled. Without corroboration from recently collected specimens, it is problematic to verify which of the locality labels are correct." pageId="33" pageNumber="32">305</figureCitation>
, 
<tableCitation id="30790379FFA53F58FAD1FC09FA8CFC01" box="[1303,1392,907,931]" captionStart="Table 1" captionStartId="6.[116,172,1082,1103]" captionText="Table 1. Comparative measurements of adults of Parabuthus erigavoensis sp. n. and P. mazuchi sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="33" pageNumber="32">Tables 1</tableCitation>
, 
<tableCitation id="30790379FFA53F58FAB8FC09FA70FC01" box="[1406,1420,907,931]" captionStart="Table 4" captionStartId="58.[113,169,705,726]" captionText="Table 4. Comparison among species (males) from Parabuthus heterurus complex and P. mazuchi sp. n., based upon selected morphometric ratios of adult males.Abbreviations: length (L), width (W), depth (D)." pageId="33" pageNumber="32">4</tableCitation>
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ETYMOLOGY. The name honors Czech herpetologist and friend TomÁš Mazuch who visited the 
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locality with us and helped to collect scorpions by hand without any accident.
</paragraph>
</subSubSection>
<subSubSection id="35E16549FFA53F58FCF6FA2EFA9CF966" pageId="33" pageNumber="32" type="diagnosis">
<paragraph id="7D4436C2FFA53F58FCF6FA2EFA9CF966" blockId="33.[816,1475,1452,1732]" pageId="33" pageNumber="32">
<materialsCitation id="CD933C9FFFA53F58FCF6FA2EFAA0F966" collectionCode="V" collectorName="Tarsomere I" location="Stridulatory area" pageId="33" pageNumber="32" specimenCount="2" typeStatus="holotype">
DIAGNOSIS. Adult male 
<typeStatus id="A2408860FFA53F58FB83FA2EFB5EFA66" box="[1093,1186,1452,1476]" pageId="33" pageNumber="32" type="holotype">holotype</typeStatus>
is 
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long, female unknown. Base color reddish brown to black, metasoma IV– 
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and telson black. Pectine teeth number 
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male. 
<location id="78246019FFA53F58FA85FA6EFCA0F986" LSID="urn:lsid:plazi:treatment:F55287D4FFA53F5FFC23FCE8FEA0F961:78246019FFA53F58FA85FA6EFCA0F986" name="Stridulatory area" pageId="33" pageNumber="32">Stridulatory area</location>
present on dorsal surface of first and second segments, reduced on third segment. Metasoma densely hirsute. Pedipalp movable finger with 14 rows of granules which include external and internal granules. Manus of pedipalp smooth and very narrow, pedipalp chela length/ width ratio 
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male 
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. 
<collectorName id="D00E5314FFA53F58FC5DF92FFBE6F966" box="[923,1050,1709,1732]" pageId="33" pageNumber="32">Tarsomere I</collectorName>
of all legs with bristle-combs
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="35E16549FFA53F5FFCF6F96EFF23F981" lastPageId="38" lastPageNumber="37" pageId="33" pageNumber="32" type="description">
<paragraph id="7D4436C2FFA53F58FCF6F96EFBC7F8C6" blockId="33.[816,1474,1772,2021]" pageId="33" pageNumber="32">
DESCRIPTION. The adult male 
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is 
<quantity id="BA039B27FFA53F58FB2DF96EFAC9F8A6" box="[1259,1333,1772,1796]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="33" pageNumber="32" unit="mm" value="63.0">63 mm</quantity>
long, female is unknown. The habitus is shown in 
<figureCitation id="E5C02A47FFA53F58FB2CF88EFA70F886" box="[1258,1420,1804,1828]" captionStart="Figures 165–166" captionStartId="34.[113,190,1574,1596]" captionTargetBox="[129,1463,236,1546]" captionTargetId="figure-228@34.[113,1474,215,1573]" captionTargetPageId="34" captionText="Figures 165–166. Parabuthus mazuchi sp. n., male holotype in dorsal (165) and ventral (166) views. Scale bar: 10 mm." pageId="33" pageNumber="32">Figs. 165–166</figureCitation>
. For position and distribution of trichobothria of pedipalps see 
<figureCitation id="E5C02A47FFA53F58FCF6F8CEFBCBF8C6" box="[816,1079,1868,1892]" captionStart="Figures 172–187" captionStartId="36.[116,193,1936,1958]" captionTargetBox="[115,1469,244,1906]" captionTargetId="figure-19@36.[113,1474,234,1915]" captionTargetPageId="36" captionText="Figures 172–187: Parabuthus mazuchi sp. n., male holotype. Figures 172–181. Pedipalp chela dorsal (172), external (173) and ventral (174) views, pedipalp patella dorsal (175), external (176) and ventral (177) views, pedipalp femur and trochanter internal (178) and dorsal (179) views, pedipalp movable (180) and fixed (181) finger dentition. Trichobothrial pattern is indicated in Figures 173–176, 178–179. Figures 182–183. Left chelicera ventral (182) and dorsal (183) views. Figures 184–187. Left legs I–IV, retrolateral aspect, respectively." pageId="33" pageNumber="32">Figs. 173–176, 178–179</figureCitation>
.
</paragraph>
<paragraph id="7D4436C2FFA53F5BFCF6F8EEFDBBF962" blockId="33.[816,1474,1772,2021]" lastBlockId="34.[111,769,1641,2017]" lastPageId="34" lastPageNumber="33" pageId="33" pageNumber="32">
<emphasis id="4F8FEAD0FFA53F58FCF6F8EEFC57F826" bold="true" box="[816,939,1900,1924]" pageId="33" pageNumber="32">Coloration</emphasis>
(
<figureCitation id="E5C02A47FFA53F58FC04F8EEFB9AF826" box="[962,1126,1900,1924]" captionStart="Figures 165–166" captionStartId="34.[113,190,1574,1596]" captionTargetBox="[129,1463,236,1546]" captionTargetId="figure-228@34.[113,1474,215,1573]" captionTargetPageId="34" captionText="Figures 165–166. Parabuthus mazuchi sp. n., male holotype in dorsal (165) and ventral (166) views. Scale bar: 10 mm." pageId="33" pageNumber="32">Figs. 165–166</figureCitation>
, 
<figureCitation id="E5C02A47FFA53F58FBBFF8EEFB26F826" box="[1145,1242,1900,1924]" captionStart="Figures 188–189" captionStartId="37.[113,190,2020,2042]" captionTargetBox="[113,1474,234,2004]" captionTargetId="figure-9@37.[113,1474,234,2006]" captionTargetPageId="37" captionText="Figures 188–189. Parabuthus mazuchi sp. n., male holotype in vivo habitus at type locality." pageId="33" pageNumber="32">188–189</figureCitation>
). The base color is reddish brown to black. The pedipalps and the femora of legs are grey to reddish brown, tarsomeres and patella of legs are grey to white. The metasoma I–II are reddish brown, lighter ventrally, metasoma III is reddish brown to black, metasoma IV–V and telson are black. Carapace and tergites are greyish black with several lighter spots on carapace.
</paragraph>
<caption id="2984664AFFA63F5BFFB7F9A4FB0FF999" box="[113,1267,1574,1596]" pageId="34" pageNumber="33" startId="34.[113,190,1574,1596]" targetBox="[129,1463,236,1546]" targetPageId="34">
<paragraph id="7D4436C2FFA63F5BFFB7F9A4FB0FF999" blockId="34.[113,1267,1574,1596]" box="[113,1267,1574,1596]" pageId="34" pageNumber="33">
<emphasis id="4F8FEAD0FFA63F5BFFB7F9A4FEE4F999" bold="true" box="[113,280,1574,1596]" pageId="34" pageNumber="33">Figures 165–166</emphasis>
. 
<taxonomicName id="BAFB4D41FFA63F5BFEE2F9A4FE16F999" authority="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský, 2019" authorityName="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský" authorityYear="2019" box="[292,490,1574,1595]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="mazuchi" status="sp. nov.">
<emphasis id="4F8FEAD0FFA63F5BFEE2F9A4FE16F999" box="[292,490,1574,1595]" italics="true" pageId="34" pageNumber="33">Parabuthus mazuchi</emphasis>
</taxonomicName>
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<emphasis id="4F8FEAD0FFA63F5BFE36F9A4FDFBF999" bold="true" box="[496,519,1574,1595]" pageId="34" pageNumber="33">sp</emphasis>
. 
<emphasis id="4F8FEAD0FFA63F5BFDD5F9A4FDDCF999" bold="true" box="[531,544,1574,1595]" pageId="34" pageNumber="33">n</emphasis>
.
</taxonomicNameLabel>
, male holotype in dorsal (165) and ventral (166) views. Scale bar: 10 mm.
</paragraph>
</caption>
<paragraph id="7D4436C2FFA63F5BFFA9F94AFBCFF963" blockId="34.[111,769,1641,2017]" lastBlockId="34.[814,1472,1641,2017]" pageId="34" pageNumber="33">
<emphasis id="4F8FEAD0FFA63F5BFFA9F94AFE5FF943" bold="true" box="[111,419,1736,1761]" pageId="34" pageNumber="33">Carapace and mesosoma</emphasis>
(
<figureCitation id="E5C02A47FFA63F5BFE78F94BFD95F942" box="[446,617,1736,1761]" captionStart-0="Figures 165–166" captionStart-1="Figures 167–171" captionStartId-0="34.[113,190,1574,1596]" captionStartId-1="35.[115,192,1988,2010]" captionTargetBox-0="[129,1463,236,1546]" captionTargetBox-1="[115,1476,215,1960]" captionTargetId-0="figure-228@34.[113,1474,215,1573]" captionTargetId-1="figure-9@35.[115,1476,213,1970]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionText-0="Figures 165–166. Parabuthus mazuchi sp. n., male holotype in dorsal (165) and ventral (166) views. Scale bar: 10 mm." captionText-1="Figures 167–171: Parabuthus mazuchi sp. n., male holotype, posterior coxosternal area and sternite III (167) carapace and tergites I–III (168), metasoma and telson in lateral (169), ventral (170), and dorsal (171) views. Scale bar: 10 mm (169–171)." pageId="34" pageNumber="33">Figs. 165–168</figureCitation>
). The entire carapace is covered with large granules. Carinae are absent. The anterior margin of the carapace is almost straight, medially weakly convex, and bears 10–12 symmetrically distributed short, stout spiniform macrosetae. The tergites are granulated. Tergite VII is pentacarinate, with lateral pairs of carinae strong, serratocrenulate. The pectinal tooth count is 36– 
<quantity id="BA039B27FFA63F5BFF70F82BFF0EF863" box="[182,242,1961,1985]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" pageId="34" pageNumber="33" unit="in" value="36.0">36 in</quantity>
male the 
<typeStatus id="A2408860FFA63F5BFEA8F82BFE33F863" box="[366,463,1961,1985]" pageId="34" pageNumber="33" type="holotype">holotype</typeStatus>
. The pectine marginal tips extend to the midpoint of the fourth sternite in the female. The pectines have three marginal lamellae and 10–12 middle lamellae. The lamellae and fulcra bear numerous dark setae. All sternites are smooth.
</paragraph>
<paragraph id="7D4436C2FFA63F5FFCE8F94BFEACFE41" blockId="34.[814,1472,1641,2017]" lastBlockId="38.[113,772,235,1315]" lastPageId="38" lastPageNumber="37" pageId="34" pageNumber="33">
<emphasis id="4F8FEAD0FFA63F5BFCE8F94BFBD5F943" bold="true" box="[814,1065,1737,1761]" pageId="34" pageNumber="33">Metasoma and telson</emphasis>
(
<figureCitation id="E5C02A47FFA63F5BFBFAF94BFB22F943" box="[1084,1246,1737,1761]" captionStart="Figures 167–171" captionStartId="35.[115,192,1988,2010]" captionTargetBox="[115,1476,215,1960]" captionTargetId="figure-9@35.[115,1476,213,1970]" captionTargetPageId="35" captionText="Figures 167–171: Parabuthus mazuchi sp. n., male holotype, posterior coxosternal area and sternite III (167) carapace and tergites I–III (168), metasoma and telson in lateral (169), ventral (170), and dorsal (171) views. Scale bar: 10 mm (169–171)." pageId="34" pageNumber="33">Figs. 169–171</figureCitation>
). The first to fourth metasomal segments each bear a total of 10 granulated carinae. The fifth segment has five carinae, and its ventral and lateral surfaces are strongly granulated. The strong lateral and ventral granulation present also on metasoma II–IV. Dorsolateral carinae of the third and fourth segments terminate in stronger denticles, of which the posterior-most denticle is enlarged. Two parallel short rows of three pointed denticles present on ventral surface of metasoma V. The stridulatory area is located on the dorsal surface of the first and second segments. On the third segment it is reduced and on fourth and fifth segments the stridulatory area is absent. The entire metasoma and the telson are densely pilose with long hairs. The ventral surface of the telson is strongly granulated. The metasomal segment V length/ width ratio is 
<quantity id="BA039B27FFA23F5FFEB4FE09FE41FE01" box="[370,445,395,419]" metricMagnitude="-2" metricUnit="m" metricValue="4.2164" pageId="38" pageNumber="37" unit="in" value="1.66">1.66 in</quantity>
the male 
<typeStatus id="A2408860FFA23F5FFDEEFE09FD75FE01" box="[552,649,395,419]" pageId="38" pageNumber="37" type="holotype">holotype</typeStatus>
. The telson is rather elongate, with the aculeus approximately the same length as the vesicle.
</paragraph>
<caption id="2984664AFFA73F5AFFB5F846FB9AF855" pageId="35" pageNumber="34" startId="35.[115,192,1988,2010]" targetBox="[115,1476,215,1960]" targetPageId="35">
<paragraph id="7D4436C2FFA73F5AFFB5F846FB9AF855" blockId="35.[115,1473,1988,2039]" pageId="35" pageNumber="34">
<emphasis id="4F8FEAD0FFA73F5AFFB5F846FEE5F87B" bold="true" box="[115,281,1988,2010]" pageId="35" pageNumber="34">Figures 167–171</emphasis>
: 
<taxonomicName id="BAFB4D41FFA73F5AFEE3F846FE16F87B" authority="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský, 2019" authorityName="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský" authorityYear="2019" box="[293,490,1988,2009]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="mazuchi" status="sp. nov.">
<emphasis id="4F8FEAD0FFA73F5AFEE3F846FE16F87B" box="[293,490,1988,2009]" italics="true" pageId="35" pageNumber="34">Parabuthus mazuchi</emphasis>
</taxonomicName>
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<emphasis id="4F8FEAD0FFA73F5AFE29F846FDFAF87B" bold="true" box="[495,518,1988,2009]" pageId="35" pageNumber="34">sp</emphasis>
. 
<emphasis id="4F8FEAD0FFA73F5AFDD7F846FDE2F87B" bold="true" box="[529,542,1988,2009]" pageId="35" pageNumber="34">n</emphasis>
.
</taxonomicNameLabel>
, male holotype, posterior coxosternal area and sternite III (167) carapace and tergites I–III (168), metasoma and telson in lateral (169), ventral (170), and dorsal (171) views. Scale bar: 10 mm (169–171).
</paragraph>
</caption>
<caption id="2984664AFFA03F5DFFB2F812FAC1F85F" pageId="36" pageNumber="35" startId="36.[116,193,1936,1958]" targetBox="[115,1469,244,1906]" targetPageId="36">
<paragraph id="7D4436C2FFA03F5DFFB2F812FAC1F85F" blockId="36.[116,1474,1936,2045]" pageId="36" pageNumber="35">
<emphasis id="4F8FEAD0FFA03F5DFFB2F812FEE7F804" bold="true" box="[116,283,1936,1958]" pageId="36" pageNumber="35">Figures 172–187</emphasis>
: 
<taxonomicName id="BAFB4D41FFA03F5DFEE1F813FE11F804" authority="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský, 2019" authorityName="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský" authorityYear="2019" box="[295,493,1937,1958]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="mazuchi" status="sp. nov.">
<emphasis id="4F8FEAD0FFA03F5DFEE1F813FE11F804" box="[295,493,1937,1958]" italics="true" pageId="36" pageNumber="35">Parabuthus mazuchi</emphasis>
</taxonomicName>
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<emphasis id="4F8FEAD0FFA03F5DFE35F813FDF6F804" bold="true" box="[499,522,1937,1958]" pageId="36" pageNumber="35">sp</emphasis>
. 
<emphasis id="4F8FEAD0FFA03F5DFDD0F813FDDFF804" bold="true" box="[534,547,1937,1958]" pageId="36" pageNumber="35">n</emphasis>
.
</taxonomicNameLabel>
, male holotype. 
<emphasis id="4F8FEAD0FFA03F5DFD0CF812FC8DF804" bold="true" box="[714,881,1936,1958]" pageId="36" pageNumber="35">Figures 172–181</emphasis>
. Pedipalp chela dorsal (172), external (173) and ventral (174) views, pedipalp patella dorsal (175), external (176) and ventral (177) views, pedipalp femur and trochanter internal (178) and dorsal (179) views, pedipalp movable (180) and fixed (181) finger dentition. Trichobothrial pattern is indicated in Figures 173–176, 178–179. 
<emphasis id="4F8FEAD0FFA03F5DFAB3F848FF34F85E" bold="true" pageId="36" pageNumber="35">Figures 182–183</emphasis>
. Left chelicera ventral (182) and dorsal (183) views. 
<emphasis id="4F8FEAD0FFA03F5DFD0EF865FC93F85E" bold="true" box="[712,879,2023,2045]" pageId="36" pageNumber="35">Figures 184–187</emphasis>
. Left legs I–IV, retrolateral aspect, respectively.
</paragraph>
</caption>
<caption id="2984664AFFA13F5CFFB7F866FC16F858" box="[113,1002,2020,2042]" pageId="37" pageNumber="36" startId="37.[113,190,2020,2042]" targetBox="[113,1474,234,2004]" targetPageId="37">
<paragraph id="7D4436C2FFA13F5CFFB7F866FC16F858" blockId="37.[113,1002,2020,2042]" box="[113,1002,2020,2042]" pageId="37" pageNumber="36">
<emphasis id="4F8FEAD0FFA13F5CFFB7F866FEE4F85B" bold="true" box="[113,280,2020,2042]" pageId="37" pageNumber="36">Figures 188–189</emphasis>
. 
<taxonomicName id="BAFB4D41FFA13F5CFEE2F866FE16F85B" authority="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský, 2019" authorityName="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský" authorityYear="2019" box="[292,490,2020,2041]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="mazuchi" status="sp. nov.">
<emphasis id="4F8FEAD0FFA13F5CFEE2F866FE16F85B" box="[292,490,2020,2041]" italics="true" pageId="37" pageNumber="36">Parabuthus mazuchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54BC57ABFFA13F5CFE36F866FDD9F85B" box="[496,549,2020,2041]" pageId="37" pageNumber="36" rank="species">
<emphasis id="4F8FEAD0FFA13F5CFE36F866FDFBF85B" bold="true" box="[496,519,2020,2041]" pageId="37" pageNumber="36">sp</emphasis>
. 
<emphasis id="4F8FEAD0FFA13F5CFDD5F866FDDCF85B" bold="true" box="[531,544,2020,2041]" pageId="37" pageNumber="36">n</emphasis>
.
</taxonomicNameLabel>
, male holotype in vivo habitus at type locality.
</paragraph>
</caption>
<paragraph id="7D4436C2FFA23F5FFFB7FE68FDA8FD41" blockId="38.[113,772,235,1315]" pageId="38" pageNumber="37">
<emphasis id="4F8FEAD0FFA23F5FFFB7FE68FF1CFDA0" bold="true" box="[113,224,490,514]" pageId="38" pageNumber="37">Pedipalps</emphasis>
(
<figureCitation id="E5C02A47FFA23F5FFF32FE69FE6AFDA1" box="[244,406,491,515]" captionStart="Figures 172–187" captionStartId="36.[116,193,1936,1958]" captionTargetBox="[115,1469,244,1906]" captionTargetId="figure-19@36.[113,1474,234,1915]" captionTargetPageId="36" captionText="Figures 172–187: Parabuthus mazuchi sp. n., male holotype. Figures 172–181. Pedipalp chela dorsal (172), external (173) and ventral (174) views, pedipalp patella dorsal (175), external (176) and ventral (177) views, pedipalp femur and trochanter internal (178) and dorsal (179) views, pedipalp movable (180) and fixed (181) finger dentition. Trichobothrial pattern is indicated in Figures 173–176, 178–179. Figures 182–183. Left chelicera ventral (182) and dorsal (183) views. Figures 184–187. Left legs I–IV, retrolateral aspect, respectively." pageId="38" pageNumber="37">Figs. 172–181</figureCitation>
). The entire pedipalps are rather sparsely hirsute with long setae. The femur bears five carinae. The chela is smooth without carinae. The patella is finely granulated without carinae. The movable and fixed fingers of pedipalp bear 14 rows of granules, all with external and internal accessory granules. The fingers of pedipalps of with inner side of base smooth, tubercle absent. The pedipalp chela length/ width ratio 
<quantity id="BA039B27FFA23F5FFEFBFD48FE7BFD41" box="[317,391,714,739]" metricMagnitude="-1" metricUnit="m" metricValue="1.7907" pageId="38" pageNumber="37" unit="in" value="7.05">7.05 in</quantity>
the male 
<typeStatus id="A2408860FFA23F5FFE36FD49FDADFD41" box="[496,593,715,739]" pageId="38" pageNumber="37" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="7D4436C2FFA23F5FFFB7FD69FEF4FC61" blockId="38.[113,772,235,1315]" pageId="38" pageNumber="37">
<emphasis id="4F8FEAD0FFA23F5FFFB7FD69FF5AFCA1" bold="true" box="[113,166,747,771]" pageId="38" pageNumber="37">Legs</emphasis>
(
<figureCitation id="E5C02A47FFA23F5FFF7BFD69FEA3FCA1" box="[189,351,747,771]" captionStart="Figures 172–187" captionStartId="36.[116,193,1936,1958]" captionTargetBox="[115,1469,244,1906]" captionTargetId="figure-19@36.[113,1474,234,1915]" captionTargetPageId="36" captionText="Figures 172–187: Parabuthus mazuchi sp. n., male holotype. Figures 172–181. Pedipalp chela dorsal (172), external (173) and ventral (174) views, pedipalp patella dorsal (175), external (176) and ventral (177) views, pedipalp femur and trochanter internal (178) and dorsal (179) views, pedipalp movable (180) and fixed (181) finger dentition. Trichobothrial pattern is indicated in Figures 173–176, 178–179. Figures 182–183. Left chelicera ventral (182) and dorsal (183) views. Figures 184–187. Left legs I–IV, retrolateral aspect, respectively." pageId="38" pageNumber="37">Figs. 184–187</figureCitation>
). Legs III and IV bear tibial spurs. Retrolateral and prolateral pedal spurs are present on all legs. All legs without distinct carinae, smooth and densely pilose. The tarsomeres bear two rows of macrosetae on the ventral surface and other macrosetae on the other surfaces. Bristlecombs are present on all legs, although slightly reduced on the fourth leg.
</paragraph>
<paragraph id="7D4436C2FFA23F5FFFB7FC48FE54FA81" blockId="38.[113,772,235,1315]" pageId="38" pageNumber="37">
<emphasis id="4F8FEAD0FFA23F5FFFB7FC48FEAAFC40" bold="true" box="[113,342,970,994]" pageId="38" pageNumber="37">Hemispermatophore</emphasis>
(
<figureCitation id="E5C02A47FFA23F5FFEAEFC49FDF9FC40" box="[360,517,970,995]" captionStart="Figures 150–160" captionStartId="30.[113,190,1886,1908]" captionTargetBox="[128,1454,258,1831]" captionTargetId="figure-18@30.[113,1474,248,1851]" captionTargetPageId="30" captionText="Figures 150–160: Hemispermatophores. Figures 150–155. Parabuthus kabateki sp. n., male paratype 1549, right hemispermatophore, distal trunk, capsule and flagellum (150), capsule in posterior (151), convex (152), convex compressed (153), anterior (154) views, and whole hemispermatophore (155). Scale bars: 1 mm (150), 500 μm (151–154), 4 mm (155). Figures 156–160. Parabuthus mazuchi sp. n., male holotype, left hemispermatophore, whole hemispermatophore (156), capsule in posterior (157), convex (158), convex compressed (159), and anterior (160) views. Scale bars: 4 mm (156), 500μm (157–160)." pageId="38" pageNumber="37">Figs. 156–160</figureCitation>
). Flagelliform, elongate and slender, trunk ca. 10 times length of capsule region. Flagellum emanating from posterior lobe of capsule; pars recta narrow, hyaline, same length as capsule; pars reflecta thicker, opaque white, ca. 3× length of capsule (damaged in dissection, not illustrated). Capsule region with 3 lobes: broad posterior lobe with angulate apical margin and strong anterior carina; small anterior lobe with narrowed apical tip; and robust basal lobe with sharp pointed tip. Hemispermatophore is similar to that reported for other 
<taxonomicName id="BAFB4D41FFA23F5FFE93FB69FDE2FAA1" box="[341,542,1259,1283]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4F8FEAD0FFA23F5FFE93FB69FE32FAA1" box="[341,462,1259,1283]" italics="true" pageId="38" pageNumber="37">Parabuthus</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="196A4B33FFA23F5FFDEFFB69FD04FAA1" author="KOVARIK" box="[553,760,1259,1285]" pageId="38" pageNumber="37" pagination="1 - 58" refId="ref22013" refString="KOVARIK, F., G. LOWE, J. PLISKOVA &amp; F. STAHLAVSKY. 2016. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VII. Parabuthus Pocock, 1890 (Buthidae), with description of P. hamar sp. n. and P. kajibu sp. n. from Ethiopia. Euscorpius, 228: 1 - 58." type="journal article" year="2016">KovařÍk et al., 2016</bibRefCitation>
). 
<emphasis id="4F8FEAD0FFA23F5FFFB7FA89FEE4FA81" bold="true" box="[113,280,1291,1315]" pageId="38" pageNumber="37">Measurements</emphasis>
. See 
<tableCitation id="30790379FFA23F5FFE94FA89FE5EFA81" box="[338,418,1291,1315]" captionStart="Table 1" captionStartId="6.[116,172,1082,1103]" captionText="Table 1. Comparative measurements of adults of Parabuthus erigavoensis sp. n. and P. mazuchi sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="38" pageNumber="37">Table 1</tableCitation>
.
</paragraph>
<paragraph id="7D4436C2FFA23F5FFFB7FAC9FF23F981" blockId="38.[113,772,1355,1571]" pageId="38" pageNumber="37">
AFFINITIES. The described features distinguish 
<taxonomicName id="BAFB4D41FFA23F5FFDA1FAC9FD20FAC1" authority="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský, 2019" authorityName="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský" authorityYear="2019" box="[615,732,1355,1379]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="mazuchi" status="sp. nov.">
<emphasis id="4F8FEAD0FFA23F5FFDA1FAC9FD20FAC1" box="[615,732,1355,1379]" italics="true" pageId="38" pageNumber="37">P. mazuchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54BC57ABFFA23F5FFD25FAC9FF7BFA20" pageId="38" pageNumber="37" rank="species">
<emphasis id="4F8FEAD0FFA23F5FFD25FAC9FD00FAC1" bold="true" box="[739,764,1355,1379]" pageId="38" pageNumber="37">sp</emphasis>
. 
<emphasis id="4F8FEAD0FFA23F5FFFB7FAE9FF7CFA21" bold="true" box="[113,128,1387,1411]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
from all other species of the genus. They are recounted in the key below. 
<taxonomicName id="BAFB4D41FFA23F5FFEC8FA09FE7CFA01" authority="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský, 2019" authorityName="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský" authorityYear="2019" box="[270,384,1419,1443]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="mazuchi" status="sp. nov.">
<emphasis id="4F8FEAD0FFA23F5FFEC8FA09FE7CFA01" box="[270,384,1419,1443]" italics="true" pageId="38" pageNumber="37">P. mazuchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54BC57ABFFA23F5FFE43FA09FE3CFA00" box="[389,448,1419,1443]" pageId="38" pageNumber="37" rank="species">
<emphasis id="4F8FEAD0FFA23F5FFE43FA09FE62FA01" bold="true" box="[389,414,1419,1443]" pageId="38" pageNumber="37">sp</emphasis>
. 
<emphasis id="4F8FEAD0FFA23F5FFE6DFA09FE46FA01" bold="true" box="[427,442,1419,1443]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
is associated with 
<taxonomicName id="BAFB4D41FFA23F5FFD42FA09FCFFFA00" authorityName="KovarIk" authorityYear="2004" box="[644,771,1419,1442]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="cimrmani">
<emphasis id="4F8FEAD0FFA23F5FFD42FA09FCFFFA00" box="[644,771,1419,1442]" italics="true" pageId="38" pageNumber="37">P. cimrmani</emphasis>
</taxonomicName>
and 
<taxonomicName id="BAFB4D41FFA23F5FFF58FA29FEC9FA60" authorityName="KovarIk" authorityYear="2003" box="[158,309,1451,1474]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="eritreaensis">
<emphasis id="4F8FEAD0FFA23F5FFF58FA29FEC9FA60" box="[158,309,1451,1474]" italics="true" pageId="38" pageNumber="37">P. eritreaensis</emphasis>
</taxonomicName>
, as all three species share characteristically elongated pedipalp fingers (
<figureCitation id="E5C02A47FFA23F5FFE58FA49FDC6FA41" box="[414,570,1483,1507]" captionStart="Figures 266–292" captionStartId="59.[113,190,1844,1866]" captionTargetBox="[118,1461,239,1828]" captionTargetId="figure-9@59.[113,1471,233,1828]" captionTargetPageId="59" captionText="Figures 266–292: Parabuthus chelae, dorsal and external views. Figures 266–267. P. heterurus, male holotype (266) and male from Somalia, Afgooye (267). Figure 268. P. kabateki sp. n., male holotype. Figure 269. P. kajibu, male holotype. Figure 270–271. P. robustus sp. n., male holotype (270) and paratype from locality 11SQ (271). Figures 272–273. P. somalilandus sp. n., male holotype (272) and paratype from Laas Gel (273). Figures 274–277. P. abyssinicus, male (274–275) and female (276–277) from locality 17SR. Figures 278–279. P. erigavoensis sp. n., female holotype. Figures 280–283. P. hamar, male (280–281) and female (282–283) paratypes. Figures 284–285. P. cimrmani, male holotype. Figures 286–287. P. eritreaensis, male holotype. Figures 288–289. P. mazuchi sp. n., male holotype. Figures 290–292. P. granimanus, male from locality 18ST (290–291) and male lektotype (292)." pageId="38" pageNumber="37">Figs. 284–289</figureCitation>
) that are absent in all other known 
<taxonomicName id="BAFB4D41FFA23F5FFEEFFA69FE59F9A1" authorityName=": Pocock" authorityYear="1895" box="[297,421,1515,1539]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="4F8FEAD0FFA23F5FFEEFFA69FE59F9A1" box="[297,421,1515,1539]" italics="true" pageId="38" pageNumber="37">Parabuthus</emphasis>
</taxonomicName>
from the Horn of Africa (
<figureCitation id="E5C02A47FFA23F5FFD0BFA69FF2FF981" captionStart="Figures 266–292" captionStartId="59.[113,190,1844,1866]" captionTargetBox="[118,1461,239,1828]" captionTargetId="figure-9@59.[113,1471,233,1828]" captionTargetPageId="59" captionText="Figures 266–292: Parabuthus chelae, dorsal and external views. Figures 266–267. P. heterurus, male holotype (266) and male from Somalia, Afgooye (267). Figure 268. P. kabateki sp. n., male holotype. Figure 269. P. kajibu, male holotype. Figure 270–271. P. robustus sp. n., male holotype (270) and paratype from locality 11SQ (271). Figures 272–273. P. somalilandus sp. n., male holotype (272) and paratype from Laas Gel (273). Figures 274–277. P. abyssinicus, male (274–275) and female (276–277) from locality 17SR. Figures 278–279. P. erigavoensis sp. n., female holotype. Figures 280–283. P. hamar, male (280–281) and female (282–283) paratypes. Figures 284–285. P. cimrmani, male holotype. Figures 286–287. P. eritreaensis, male holotype. Figures 288–289. P. mazuchi sp. n., male holotype. Figures 290–292. P. granimanus, male from locality 18ST (290–291) and male lektotype (292)." pageId="38" pageNumber="37">Figs. 266–283</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="35E16549FFA23F5FFFB7F9C9FEA0F961" pageId="38" pageNumber="37" type="discussion">
<paragraph id="7D4436C2FFA23F5FFFB7F9C9FEA0F961" blockId="38.[113,775,1611,1731]" pageId="38" pageNumber="37">
COMMENTS ON LOCALITIES AND LIFE STRATEGY. The 
<typeStatus id="A2408860FFA23F5FFD13F9C9FCFFF9C1" box="[725,771,1611,1635]" pageId="38" pageNumber="37">type</typeStatus>
locality, 
<emphasis id="4F8FEAD0FFA23F5FFF16F9E8FEFBF920" bold="true" box="[208,263,1642,1666]" pageId="38" pageNumber="37">18SJ</emphasis>
is also the 
<typeStatus id="A2408860FFA23F5FFE4AF9E9FE46F921" box="[396,442,1643,1667]" pageId="38" pageNumber="37">type</typeStatus>
locality for here described 
<taxonomicName id="BAFB4D41FFA23F5FFD29F9E9FF37F901" authority="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský, 2019" authorityName="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský" authorityYear="2019" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="kabateki" status="sp. nov.">
<emphasis id="4F8FEAD0FFA23F5FFD29F9E9FF37F901" italics="true" pageId="38" pageNumber="37">P. kabateki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54BC57ABFFA23F5FFF1EF909FEE5F900" box="[216,281,1675,1699]" pageId="38" pageNumber="37" rank="species">
<emphasis id="4F8FEAD0FFA23F5FFF1EF909FF0DF901" bold="true" box="[216,241,1675,1699]" pageId="38" pageNumber="37">sp</emphasis>
. 
<emphasis id="4F8FEAD0FFA23F5FFEC2F909FEEFF901" bold="true" box="[260,275,1675,1699]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
, see comments under the description of 
<taxonomicName id="BAFB4D41FFA23F5FFD29F909FF37F961" authority="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský, 2019" authorityName="Kovařík &amp; Lowe &amp; Elmi &amp; Šťáhlavský" authorityYear="2019" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="kabateki" status="sp. nov.">
<emphasis id="4F8FEAD0FFA23F5FFD29F909FF37F961" italics="true" pageId="38" pageNumber="37">P. kabateki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54BC57ABFFA23F5FFF14F929FEF3F960" box="[210,271,1707,1731]" pageId="38" pageNumber="37" rank="species">
<emphasis id="4F8FEAD0FFA23F5FFF14F929FF17F961" bold="true" box="[210,235,1707,1731]" pageId="38" pageNumber="37">sp</emphasis>
. 
<emphasis id="4F8FEAD0FFA23F5FFF3FF929FEF4F961" bold="true" box="[249,264,1707,1731]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
above.
</paragraph>
</subSubSection>
</treatment>
</document>