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<document id="D7B7FE31F7A5F6455B5D769B27789246" ID-DOI="10.1111/j.1096-3642.2009.00628.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5754697" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1630446086990" checkinUser="valdenar" docAuthor="Turvey, Samuel T., Weksler, Marcelo, Morris, Elaine L. &amp; Nokkert, Mark" docDate="2010" docId="7C4E878AAB7CFF966302E44508EF8F12" docLanguage="en" docName="j.1096-3642.2009.00628.x.pdf" docOrigin="Zoological Journal of the Linnean Society 160 (4)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2009.00628.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Megalomys TROUESSART 1881" docType="treatment" docVersion="4" lastPageNumber="754" masterDocId="8077FFF2AB7FFF916076E1650950894F" masterDocTitle="Taxonomy, phylogeny, and diversity of the extinct Lesser Antillean rice rats (Sigmodontinae: Oryzomyini), with description of a new genus and species" masterLastPageNumber="772" masterPageNumber="748" pageNumber="751" updateTime="1727983355689" updateUser="juliana">
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<mods:title id="A5966FCBA713AE83DA79D1D393730677">Taxonomy, phylogeny, and diversity of the extinct Lesser Antillean rice rats (Sigmodontinae: Oryzomyini), with description of a new genus and species</mods:title>
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<mods:namePart id="98D35BE8F473868B65DBB8511916A9C5">Turvey, Samuel T.</mods:namePart>
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<mods:namePart id="9F072C9E75BA5E6B98681AC713E8D5C9">Morris, Elaine L.</mods:namePart>
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<treatment id="7C4E878AAB7CFF966302E44508EF8F12" LSID="urn:lsid:plazi:treatment:7C4E878AAB7CFF966302E44508EF8F12" httpUri="http://treatment.plazi.org/id/7C4E878AAB7CFF966302E44508EF8F12" lastPageId="7" lastPageNumber="754" pageId="3" pageNumber="751">
<subSubSection id="BCFD6517AB7CFF926302E4450C6F8C7A" box="[884,1343,1312,1333]" pageId="3" pageNumber="751" type="nomenclature">
<paragraph id="F458369CAB7CFF926302E4450C6F8C7A" blockId="3.[884,1343,1312,1333]" box="[884,1343,1312,1333]" pageId="3" pageNumber="751">
<heading id="AF1081F0AB7CFF926302E4450C6F8C7A" box="[884,1343,1312,1333]" centered="true" fontSize="8" level="2" pageId="3" pageNumber="751" reason="8">
GENUS
<taxonomicName id="33E74D1FAB7CFF9263BBE4450C6F8C7A" ID-CoL="62YZZ" authority="TROUESSART, 1881" authorityName="TROUESSART" authorityYear="1881" box="[973,1343,1312,1333]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="751" phylum="Chordata" rank="genus">
<emphasis id="C693EA8EAB7CFF9263BBE4450D0C8C7B" box="[973,1116,1312,1333]" italics="true" pageId="3" pageNumber="751">MEGALOMYS</emphasis>
TROUESSART, 1881
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BCFD6517AB7CFF926313E4230C1F8C13" box="[869,1359,1350,1372]" pageId="3" pageNumber="751" type="type_taxon">
<paragraph id="F458369CAB7CFF926313E4230C1F8C13" blockId="3.[869,1359,1350,1372]" box="[869,1359,1350,1372]" pageId="3" pageNumber="751">
(
<typeStatus id="2B5C883EAB7CFF92631BE42F0AF48C15" box="[877,932,1354,1370]" pageId="3" pageNumber="751">TYPE</typeStatus>
SPECIES
<taxonomicName id="33E74D1FAB7CFF926464E4220C178C13" authority="SAY, 1822" authorityName="SAY" authorityYear="1822" box="[1042,1351,1350,1372]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="751" phylum="Chordata" rank="species" species="pilorides">
<emphasis id="C693EA8EAB7CFF926464E4220D968C15" box="[1042,1222,1351,1372]" italics="true" pageId="3" pageNumber="751">MUS PILORIDES</emphasis>
SAY, 1822
</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="BCFD6517AB7CFF926350E4080ABF8F37" pageId="3" pageNumber="751" type="materials_examined">
<paragraph id="F458369CAB7CFF926350E4080C248C90" blockId="3.[806,1421,1389,1656]" pageId="3" pageNumber="751">
<emphasis id="C693EA8EAB7CFF926350E4080D428CCD" box="[806,1042,1389,1410]" italics="true" pageId="3" pageNumber="751">Examined material:</emphasis>
<taxonomicName id="33E74D1FAB7CFF926455E4080C668CCD" authority=": NHM" authorityName="NHM" box="[1059,1334,1389,1410]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="751" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB7CFF926455E4080C668CCD" box="[1059,1334,1389,1410]" italics="true" pageId="3" pageNumber="751">Megalomys desmarestii</emphasis>
</taxonomicName>
:
<materialsCitation id="448F3CC1AB7CFF92653BE4080C248C90" collectionCode="NHM" pageId="3" pageNumber="751" specimenCode="55.12.24.201, 50.11.30.5, 50.11.30.6" specimenCount="1">
<collectionCode id="92F6AE59AB7CFF92653BE4080CDD8CCD" box="[1357,1421,1389,1410]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="3" pageNumber="751">NHM</collectionCode>
<specimenCode id="A4419EE7AB7CFF926349E4EE0A9E8CEE" box="[831,974,1419,1441]" pageId="3" pageNumber="751">55.12.24.201</specimenCode>
(skin and skull);
<collectionCode id="92F6AE59AB7CFF9264B1E4E90C578CEE" box="[1223,1287,1420,1441]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="3" pageNumber="751">NHM</collectionCode>
<specimenCode id="A4419EE7AB7CFF92656CE4EE0CDD8CEE" box="[1306,1421,1419,1441]" pageId="3" pageNumber="751">50.11.30.5</specimenCode>
(skin), and
<collectionCode id="92F6AE59AB7CFF9263B0E4CE0D568C8F" box="[966,1030,1451,1472]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="3" pageNumber="751">NHM</collectionCode>
<specimenCode id="A4419EE7AB7CFF926466E4CF0DD38C8F" box="[1040,1155,1450,1472]" pageId="3" pageNumber="751">50.11.30.6</specimenCode>
(skull and postcranial skeleton) – these represent the same individual.
</materialsCitation>
</paragraph>
<paragraph id="F458369CAB7CFF926350E4820D378F53" blockId="3.[806,1421,1389,1656]" pageId="3" pageNumber="751">
<taxonomicName id="33E74D1FAB7CFF926350E4820D578CB3" authority=": NHM" authorityName="NHM" box="[806,1031,1511,1532]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="751" phylum="Chordata" rank="species" species="luciae">
<emphasis id="C693EA8EAB7CFF926350E4820D578CB3" box="[806,1031,1511,1532]" italics="true" pageId="3" pageNumber="751">Megalomys luciae</emphasis>
</taxonomicName>
:
<materialsCitation id="448F3CC1AB7CFF92645AE48D0D338F53" collectionCode="NHM" pageId="3" pageNumber="751" specimenCode="53.12.16.2" specimenCount="1" typeStatus="holotype">
<collectionCode id="92F6AE59AB7CFF92645AE48D0D3C8CB2" box="[1068,1132,1512,1533]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="3" pageNumber="751">NHM</collectionCode>
<specimenCode id="A4419EE7AB7CFF9264FDE4820DAF8CB2" box="[1163,1279,1511,1533]" pageId="3" pageNumber="751">53.12.16.2</specimenCode>
(
<typeStatus id="2B5C883EAB7CFF926553E4820CD98CB2" box="[1317,1417,1511,1533]" pageId="3" pageNumber="751" type="holotype">holotype</typeStatus>
; damaged skin and skull)
</materialsCitation>
.
</paragraph>
<paragraph id="F458369CAB7CFF926350E7410ABF8F37" blockId="3.[806,1421,1389,1656]" pageId="3" pageNumber="751">
<taxonomicName id="33E74D1FAB7CFF926350E7410D4B8F76" authority=": NHM M" authorityName="NHM M" box="[806,1051,1572,1593]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="751" phylum="Chordata" rank="species" species="audreyae">
<emphasis id="C693EA8EAB7CFF926350E7410D4B8F76" box="[806,1051,1572,1593]" italics="true" pageId="3" pageNumber="751">Megalomys audreyae</emphasis>
</taxonomicName>
:
<materialsCitation id="448F3CC1AB7CFF926447E7400ABF8F37" collectionCode="NHM" pageId="3" pageNumber="751" specimenCode="M7406a, M7406b" specimenCount="1" typeStatus="holotype">
<collectionCode id="92F6AE59AB7CFF926447E7400D218F75" box="[1073,1137,1573,1594]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="3" pageNumber="751">NHM</collectionCode>
<specimenCode id="A4419EE7AB7CFF926409E7410D8B8F75" box="[1151,1243,1572,1594]" pageId="3" pageNumber="751">M7406a</specimenCode>
(
<typeStatus id="2B5C883EAB7CFF926487E7410C058F75" box="[1265,1365,1572,1594]" pageId="3" pageNumber="751" type="holotype">holotype</typeStatus>
; left mandibular ramus lacking m1);
<collectionCode id="92F6AE59AB7CFF9264A9E7210C4F8F16" box="[1247,1311,1604,1625]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="3" pageNumber="751">NHM</collectionCode>
<specimenCode id="A4419EE7AB7CFF926547E7260CDD8F16" box="[1329,1421,1603,1625]" pageId="3" pageNumber="751">M7406b</specimenCode>
(upper incisor).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BCFD6517AB7CFF956350E7C70A5B8F14" lastPageId="4" lastPageNumber="752" pageId="3" pageNumber="751" type="diagnosis">
<paragraph id="F458369CAB7CFF956350E7C70A5B8F14" blockId="3.[806,1421,1697,1903]" lastBlockId="4.[164,779,1116,1627]" lastPageId="4" lastPageNumber="752" pageId="3" pageNumber="751">
<emphasis id="C693EA8EAB7CFF926350E7C70D778FF9" box="[806,1063,1697,1719]" italics="true" pageId="3" pageNumber="751">
Diagnosis:
<taxonomicName id="33E74D1FAB7CFF9263DEE7C40D778FF9" authorityName="TROUESSART" authorityYear="1881" box="[936,1063,1697,1718]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="751" phylum="Chordata" rank="genus">Megalomys</taxonomicName>
</emphasis>
can be differentiated from all other sigmodontine rodents by the combination of its extreme body size, oryzomyine synapomorphies (e.g. long palate, absence of both suspensory process of squamosal and alisphenoid struts), and several integumental, cranial, and dental traits, including nine synapomorphies (see below). Important characters include: pes without well-developed natatory fringes and interdigital webs, and with interdigital pads large and fleshy; robust skull with interorbital region convergent anteriorly, with well-developed supraorbital crests and squared braincase; zygomatic plate lacks an anterodorsal spinous process; small incisive foramina, not extending posteriorly between M1 alveoli; large posterolateral palatal pits recessed in deep fossae; stapedial foramen and posterior opening of alisphenoid canal small, and squamosal– alisphenoid groove and sphenofrontal foramen absent. Well-developed capsular process of lower incisor alveolus; and superior and inferior masseteric ridges joined anteriorly as an open chevron. Molars are bunodont, with M1 anterocone divided by a shallow anteromedian flexus; accessory rootlets are present on M1 and m1; m2 and m3 have three roots.
</paragraph>
</subSubSection>
<caption id="A0986614AB7BFF9560D2E1AD0C458994" box="[164,1301,200,219]" pageId="4" pageNumber="752" startId="4.[164,228,200,219]" targetBox="[164,1441,257,979]" targetIsTable="true" targetPageId="4">
<paragraph id="F458369CAB7BFF9560D2E1AD0C458994" blockId="4.[164,1301,200,219]" box="[164,1301,200,219]" pageId="4" pageNumber="752">
<emphasis id="C693EA8EAB7BFF9560D2E1AD09AE8994" bold="true" box="[164,254,200,219]" pageId="4" pageNumber="752">Table 1.</emphasis>
Measurement data for
<taxonomicName id="33E74D1FAB7BFF956277E1AD0BA78994" box="[513,759,200,219]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB7BFF956277E1AD0BA78994" box="[513,759,200,219]" italics="true" pageId="4" pageNumber="752">Megalomys desmarestii</emphasis>
</taxonomicName>
,
<taxonomicName id="33E74D1FAB7BFF956370E1AD0A928994" box="[774,962,200,219]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="luciae">
<emphasis id="C693EA8EAB7BFF956370E1AD0A928994" box="[774,962,200,219]" italics="true" pageId="4" pageNumber="752">Megalomys luciae</emphasis>
</taxonomicName>
, and
<taxonomicName id="33E74D1FAB7BFF956474E1AD0D848995" baseAuthorityName="THOMAS" baseAuthorityYear="1898" box="[1026,1236,200,219]" class="Mammalia" family="Cricetidae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="victus">
<emphasis id="C693EA8EAB7BFF956474E1AD0D848995" box="[1026,1236,200,219]" italics="true" pageId="4" pageNumber="752">Oligoryzomys victus</emphasis>
</taxonomicName>
(mm)
</paragraph>
</caption>
<paragraph id="F458369CAB7BFF9560D2E07A0C598A9D" pageId="4" pageNumber="752">
<table id="86E7C43CAB7B006E60D2E0640CF18A9C" box="[164,1441,257,979]" gridcols="5" gridrows="24" pageId="4" pageNumber="752">
<tr id="4AD734DEAB7B006E60D2E0640CF1885A" box="[164,1441,257,277]" gridrow="0" isHeader="true" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E06408AE887D" box="[164,510,257,306]" gridcol="0" gridrow="0" pageId="4" pageNumber="752" rowspan="2" rowspanBelow="1">
<emphasis id="C693EA8EAB7BFF9560D2E07A0860887E" box="[164,304,287,305]" italics="true" pageId="4" pageNumber="752">Measurement</emphasis>
</th>
<th id="09065DA2AB7B006E6268E0640BBF885A" box="[542,751,257,277]" gridcol="1" gridrow="0" pageId="4" pageNumber="752">
<taxonomicName id="33E74D1FAB7BFF956268E0670BEB885B" box="[542,699,257,276]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB7BFF956268E0670BEB885B" box="[542,699,257,276]" italics="true" pageId="4" pageNumber="752">M. desmarestii</emphasis>
</taxonomicName>
</th>
<th id="09065DA2AB7B006E6379E0640AB6885A" box="[783,998,257,277]" gridcol="2" gridrow="0" pageId="4" pageNumber="752">
<taxonomicName id="33E74D1FAB7BFF956379E0670AFB885B" box="[783,939,257,276]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB7BFF956379E0670AFB885B" box="[783,939,257,276]" italics="true" pageId="4" pageNumber="752">M. desmarestii</emphasis>
</taxonomicName>
</th>
<th id="09065DA2AB7B006E6470E0640D94885A" box="[1030,1220,257,277]" gridcol="3" gridrow="0" pageId="4" pageNumber="752">
<taxonomicName id="33E74D1FAB7BFF956470E0670D38885B" box="[1030,1128,257,276]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="luciae">
<emphasis id="C693EA8EAB7BFF956470E0670D38885B" box="[1030,1128,257,276]" italics="true" pageId="4" pageNumber="752">M. luciae</emphasis>
</taxonomicName>
</th>
<th id="09065DA2AB7B006E6495E0640CF1885A" box="[1251,1441,257,277]" gridcol="4" gridrow="0" pageId="4" pageNumber="752">
<taxonomicName id="33E74D1FAB7BFF956495E0670C11885B" baseAuthorityName="THOMAS" baseAuthorityYear="1898" box="[1251,1345,258,277]" class="Mammalia" family="Cricetidae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="victus">
<emphasis id="C693EA8EAB7BFF956495E0670C11885B" box="[1251,1345,258,277]" italics="true" pageId="4" pageNumber="752">O. victus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="4AD734DEAB7B006E60D2E07A0CF1887D" box="[164,1441,287,306]" gridrow="1" isHeader="true" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E6268E07A0BBF887D" box="[542,751,287,306]" gridcol="1" gridrow="1" pageId="4" pageNumber="752">
(
<collectionCode id="92F6AE59AB7BFF956253E07A0B30887D" box="[549,608,287,306]" pageId="4" pageNumber="752">NHM</collectionCode>
50.11.30.5-6)
</th>
<th id="09065DA2AB7B006E6379E07A0AB6887D" box="[783,998,287,306]" gridcol="2" gridrow="1" pageId="4" pageNumber="752">
(
<collectionCode id="92F6AE59AB7BFF956363E07A0A00887D" box="[789,848,287,306]" pageId="4" pageNumber="752">NHM</collectionCode>
55.12.24.201)
</th>
<th id="09065DA2AB7B006E6470E07A0D94887D" box="[1030,1220,287,306]" gridcol="3" gridrow="1" pageId="4" pageNumber="752">
(
<collectionCode id="92F6AE59AB7BFF95647AE07A0D18887D" box="[1036,1096,287,306]" pageId="4" pageNumber="752">NHM</collectionCode>
53.12.16.2)
</th>
<th id="09065DA2AB7B006E6495E07A0CF1887D" box="[1251,1441,287,306]" gridcol="4" gridrow="1" pageId="4" pageNumber="752">
(
<collectionCode id="92F6AE59AB7BFF95649CE07A0C75887D" box="[1258,1317,287,306]" pageId="4" pageNumber="752">NHM</collectionCode>
97.12.26.1)
</th>
</tr>
<tr id="4AD734DEAB7B006E60D2E03C0CF18823" box="[164,1441,345,364]" gridrow="2" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E03C08AE8823" box="[164,510,345,364]" gridcol="0" gridrow="2" pageId="4" pageNumber="752">Greatest skull length</th>
<td id="09065DA2AB7B006E6268E03C0BBF8823" box="[542,751,345,364]" gridcol="1" gridrow="2" pageId="4" pageNumber="752">55.2</td>
<td id="09065DA2AB7B006E6379E03C0AB68823" box="[783,998,345,364]" gridcol="2" gridrow="2" pageId="4" pageNumber="752">–</td>
<td id="09065DA2AB7B006E6470E03C0D948823" box="[1030,1220,345,364]" gridcol="3" gridrow="2" pageId="4" pageNumber="752">–</td>
<td id="09065DA2AB7B006E6495E03C0CF18823" box="[1251,1441,345,364]" gridcol="4" gridrow="2" pageId="4" pageNumber="752">–</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E0130CF188C6" box="[164,1441,374,393]" gridrow="3" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E01308AE88C6" box="[164,510,374,393]" gridcol="0" gridrow="3" pageId="4" pageNumber="752">Condylo-incisive length</th>
<td id="09065DA2AB7B006E6268E0130BBF88C6" box="[542,751,374,393]" gridcol="1" gridrow="3" pageId="4" pageNumber="752">53.6</td>
<td id="09065DA2AB7B006E6379E0130AB688C6" box="[783,998,374,393]" gridcol="2" gridrow="3" pageId="4" pageNumber="752">–</td>
<td id="09065DA2AB7B006E6470E0130D9488C6" box="[1030,1220,374,393]" gridcol="3" gridrow="3" pageId="4" pageNumber="752">47.5</td>
<td id="09065DA2AB7B006E6495E0130CF188C6" box="[1251,1441,374,393]" gridcol="4" gridrow="3" pageId="4" pageNumber="752">–</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E0F60CF188E8" box="[164,1441,403,423]" gridrow="4" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E0F608AE88E8" box="[164,510,403,423]" gridcol="0" gridrow="4" pageId="4" pageNumber="752">Breadth of occipital condyles</th>
<td id="09065DA2AB7B006E6268E0F60BBF88E8" box="[542,751,403,423]" gridcol="1" gridrow="4" pageId="4" pageNumber="752">9.6</td>
<td id="09065DA2AB7B006E6379E0F60AB688E8" box="[783,998,403,423]" gridcol="2" gridrow="4" pageId="4" pageNumber="752">–</td>
<td id="09065DA2AB7B006E6470E0F60D9488E8" box="[1030,1220,403,423]" gridcol="3" gridrow="4" pageId="4" pageNumber="752">9.8</td>
<td id="09065DA2AB7B006E6495E0F60CF188E8" box="[1251,1441,403,423]" gridcol="4" gridrow="4" pageId="4" pageNumber="752">–</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E0D50CF1888B" box="[164,1441,432,452]" gridrow="5" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E0D508AE888B" box="[164,510,432,452]" gridcol="0" gridrow="5" pageId="4" pageNumber="752">Length of diastema</th>
<td id="09065DA2AB7B006E6268E0D50BBF888B" box="[542,751,432,452]" gridcol="1" gridrow="5" pageId="4" pageNumber="752">17.8</td>
<td id="09065DA2AB7B006E6379E0D50AB6888B" box="[783,998,432,452]" gridcol="2" gridrow="5" pageId="4" pageNumber="752">16.9</td>
<td id="09065DA2AB7B006E6470E0D50D94888B" box="[1030,1220,432,452]" gridcol="3" gridrow="5" pageId="4" pageNumber="752">15.6</td>
<td id="09065DA2AB7B006E6495E0D50CF1888B" box="[1251,1441,432,452]" gridcol="4" gridrow="5" pageId="4" pageNumber="752">7.8</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E0AB0CF188AE" box="[164,1441,462,481]" gridrow="6" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E0AB08AE88AE" box="[164,510,462,481]" gridcol="0" gridrow="6" pageId="4" pageNumber="752">Palatal bridge</th>
<td id="09065DA2AB7B006E6268E0AB0BBF88AE" box="[542,751,462,481]" gridcol="1" gridrow="6" pageId="4" pageNumber="752">14.0</td>
<td id="09065DA2AB7B006E6379E0AB0AB688AE" box="[783,998,462,481]" gridcol="2" gridrow="6" pageId="4" pageNumber="752">12.8</td>
<td id="09065DA2AB7B006E6470E0AB0D9488AE" box="[1030,1220,462,481]" gridcol="3" gridrow="6" pageId="4" pageNumber="752">13.0</td>
<td id="09065DA2AB7B006E6495E0AB0CF188AE" box="[1251,1441,462,481]" gridcol="4" gridrow="6" pageId="4" pageNumber="752">6.2</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E08E0CF188B1" box="[164,1441,491,510]" gridrow="7" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E08E08AE88B1" box="[164,510,491,510]" gridcol="0" gridrow="7" pageId="4" pageNumber="752">Length of incisive foramen</th>
<td id="09065DA2AB7B006E6268E08E0BBF88B1" box="[542,751,491,510]" gridcol="1" gridrow="7" pageId="4" pageNumber="752">9.4</td>
<td id="09065DA2AB7B006E6379E08E0AB688B1" box="[783,998,491,510]" gridcol="2" gridrow="7" pageId="4" pageNumber="752">8.3</td>
<td id="09065DA2AB7B006E6470E08E0D9488B1" box="[1030,1220,491,510]" gridcol="3" gridrow="7" pageId="4" pageNumber="752">7.8</td>
<td id="09065DA2AB7B006E6495E08E0CF188B1" box="[1251,1441,491,510]" gridcol="4" gridrow="7" pageId="4" pageNumber="752">5.4</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E36D0CF18B54" box="[164,1441,520,539]" gridrow="8" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E36D08AE8B54" box="[164,510,520,539]" gridcol="0" gridrow="8" pageId="4" pageNumber="752">Breadth of incisive foramen</th>
<td id="09065DA2AB7B006E6268E36D0BBF8B54" box="[542,751,520,539]" gridcol="1" gridrow="8" pageId="4" pageNumber="752">3.3</td>
<td id="09065DA2AB7B006E6379E36D0AB68B54" box="[783,998,520,539]" gridcol="2" gridrow="8" pageId="4" pageNumber="752">3.1</td>
<td id="09065DA2AB7B006E6470E36D0D948B54" box="[1030,1220,520,539]" gridcol="3" gridrow="8" pageId="4" pageNumber="752">2.7</td>
<td id="09065DA2AB7B006E6495E36D0CF18B54" box="[1251,1441,520,539]" gridcol="4" gridrow="8" pageId="4" pageNumber="752">2.1</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E3400CF18B76" box="[164,1441,549,569]" gridrow="9" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E34008AE8B76" box="[164,510,549,569]" gridcol="0" gridrow="9" pageId="4" pageNumber="752">Length of maxillary molars</th>
<td id="09065DA2AB7B006E6268E3400BBF8B76" box="[542,751,549,569]" gridcol="1" gridrow="9" pageId="4" pageNumber="752">8.2</td>
<td id="09065DA2AB7B006E6379E3400AB68B76" box="[783,998,549,569]" gridcol="2" gridrow="9" pageId="4" pageNumber="752">9.4</td>
<td id="09065DA2AB7B006E6470E3400D948B76" box="[1030,1220,549,569]" gridcol="3" gridrow="9" pageId="4" pageNumber="752">7.7</td>
<td id="09065DA2AB7B006E6495E3400CF18B76" box="[1251,1441,549,569]" gridcol="4" gridrow="9" pageId="4" pageNumber="752">4.5</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E3260CF18B19" box="[164,1441,579,598]" gridrow="10" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E32608AE8B19" box="[164,510,579,598]" gridcol="0" gridrow="10" pageId="4" pageNumber="752">Breadth of first maxillary molar</th>
<td id="09065DA2AB7B006E6268E3260BBF8B19" box="[542,751,579,598]" gridcol="1" gridrow="10" pageId="4" pageNumber="752">2.5</td>
<td id="09065DA2AB7B006E6379E3260AB68B19" box="[783,998,579,598]" gridcol="2" gridrow="10" pageId="4" pageNumber="752">2.6</td>
<td id="09065DA2AB7B006E6470E3260D948B19" box="[1030,1220,579,598]" gridcol="3" gridrow="10" pageId="4" pageNumber="752">2.5</td>
<td id="09065DA2AB7B006E6495E3260CF18B19" box="[1251,1441,579,598]" gridcol="4" gridrow="10" pageId="4" pageNumber="752">1.4</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E3050CF18B3C" box="[164,1441,608,627]" gridrow="11" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E30508AE8B3C" box="[164,510,608,627]" gridcol="0" gridrow="11" pageId="4" pageNumber="752">External alveolar breadth</th>
<td id="09065DA2AB7B006E6268E3050BBF8B3C" box="[542,751,608,627]" gridcol="1" gridrow="11" pageId="4" pageNumber="752">10.7</td>
<td id="09065DA2AB7B006E6379E3050AB68B3C" box="[783,998,608,627]" gridcol="2" gridrow="11" pageId="4" pageNumber="752">10.5</td>
<td id="09065DA2AB7B006E6470E3050D948B3C" box="[1030,1220,608,627]" gridcol="3" gridrow="11" pageId="4" pageNumber="752">10.4</td>
<td id="09065DA2AB7B006E6495E3050CF18B3C" box="[1251,1441,608,627]" gridcol="4" gridrow="11" pageId="4" pageNumber="752">5.6</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E3180CF18BDF" box="[164,1441,637,656]" gridrow="12" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E31808AE8BDF" box="[164,510,637,656]" gridcol="0" gridrow="12" pageId="4" pageNumber="752">Cranial height</th>
<td id="09065DA2AB7B006E6268E3180BBF8BDF" box="[542,751,637,656]" gridcol="1" gridrow="12" pageId="4" pageNumber="752">15.7</td>
<td id="09065DA2AB7B006E6379E3180AB68BDF" box="[783,998,637,656]" gridcol="2" gridrow="12" pageId="4" pageNumber="752">14.6</td>
<td id="09065DA2AB7B006E6470E3180D948BDF" box="[1030,1220,637,656]" gridcol="3" gridrow="12" pageId="4" pageNumber="752">13.7</td>
<td id="09065DA2AB7B006E6495E3180CF18BDF" box="[1251,1441,637,656]" gridcol="4" gridrow="12" pageId="4" pageNumber="752">9.3</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E3FE0CF18BE1" box="[164,1441,667,686]" gridrow="13" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E3FE08AE8BE1" box="[164,510,667,686]" gridcol="0" gridrow="13" pageId="4" pageNumber="752">Rostrum length</th>
<td id="09065DA2AB7B006E6268E3FE0BBF8BE1" box="[542,751,667,686]" gridcol="1" gridrow="13" pageId="4" pageNumber="752">21.0</td>
<td id="09065DA2AB7B006E6379E3FE0AB68BE1" box="[783,998,667,686]" gridcol="2" gridrow="13" pageId="4" pageNumber="752">–</td>
<td id="09065DA2AB7B006E6470E3FE0D948BE1" box="[1030,1220,667,686]" gridcol="3" gridrow="13" pageId="4" pageNumber="752">–</td>
<td id="09065DA2AB7B006E6495E3FE0CF18BE1" box="[1251,1441,667,686]" gridcol="4" gridrow="13" pageId="4" pageNumber="752">11.2</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E3DD0CF18B84" box="[164,1441,696,715]" gridrow="14" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E3DD08AE8B84" box="[164,510,696,715]" gridcol="0" gridrow="14" pageId="4" pageNumber="752">Rostrum breadth</th>
<td id="09065DA2AB7B006E6268E3DD0BBF8B84" box="[542,751,696,715]" gridcol="1" gridrow="14" pageId="4" pageNumber="752">11.4</td>
<td id="09065DA2AB7B006E6379E3DD0AB68B84" box="[783,998,696,715]" gridcol="2" gridrow="14" pageId="4" pageNumber="752">10.6</td>
<td id="09065DA2AB7B006E6470E3DD0D948B84" box="[1030,1220,696,715]" gridcol="3" gridrow="14" pageId="4" pageNumber="752">9.0</td>
<td id="09065DA2AB7B006E6495E3DD0CF18B84" box="[1251,1441,696,715]" gridcol="4" gridrow="14" pageId="4" pageNumber="752">6.1</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E3B00CF18BA6" box="[164,1441,725,745]" gridrow="15" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E3B008AE8BA6" box="[164,510,725,745]" gridcol="0" gridrow="15" pageId="4" pageNumber="752">Least interorbital breadth</th>
<td id="09065DA2AB7B006E6268E3B00BBF8BA6" box="[542,751,725,745]" gridcol="1" gridrow="15" pageId="4" pageNumber="752">9.1</td>
<td id="09065DA2AB7B006E6379E3B00AB68BA6" box="[783,998,725,745]" gridcol="2" gridrow="15" pageId="4" pageNumber="752">9.7</td>
<td id="09065DA2AB7B006E6470E3B00D948BA6" box="[1030,1220,725,745]" gridcol="3" gridrow="15" pageId="4" pageNumber="752">9.4</td>
<td id="09065DA2AB7B006E6495E3B00CF18BA6" box="[1251,1441,725,745]" gridcol="4" gridrow="15" pageId="4" pageNumber="752">5.1</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E3970CF18A49" box="[164,1441,754,774]" gridrow="16" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E39708AE8A49" box="[164,510,754,774]" gridcol="0" gridrow="16" pageId="4" pageNumber="752">Orbital length</th>
<td id="09065DA2AB7B006E6268E3970BBF8A49" box="[542,751,754,774]" gridcol="1" gridrow="16" pageId="4" pageNumber="752">17.4</td>
<td id="09065DA2AB7B006E6379E3970AB68A49" box="[783,998,754,774]" gridcol="2" gridrow="16" pageId="4" pageNumber="752">17.5</td>
<td id="09065DA2AB7B006E6470E3970D948A49" box="[1030,1220,754,774]" gridcol="3" gridrow="16" pageId="4" pageNumber="752">15.6</td>
<td id="09065DA2AB7B006E6495E3970CF18A49" box="[1251,1441,754,774]" gridcol="4" gridrow="16" pageId="4" pageNumber="752">11.0</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E2750CF18A6C" box="[164,1441,784,803]" gridrow="17" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E27508AE8A6C" box="[164,510,784,803]" gridcol="0" gridrow="17" pageId="4" pageNumber="752">Zygomatic breadth</th>
<td id="09065DA2AB7B006E6268E2750BBF8A6C" box="[542,751,784,803]" gridcol="1" gridrow="17" pageId="4" pageNumber="752">30.6</td>
<td id="09065DA2AB7B006E6379E2750AB68A6C" box="[783,998,784,803]" gridcol="2" gridrow="17" pageId="4" pageNumber="752">–</td>
<td id="09065DA2AB7B006E6470E2750D948A6C" box="[1030,1220,784,803]" gridcol="3" gridrow="17" pageId="4" pageNumber="752">26.9</td>
<td id="09065DA2AB7B006E6495E2750CF18A6C" box="[1251,1441,784,803]" gridcol="4" gridrow="17" pageId="4" pageNumber="752">16.7</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E2480CF18A0F" box="[164,1441,813,832]" gridrow="18" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E24808AE8A0F" box="[164,510,813,832]" gridcol="0" gridrow="18" pageId="4" pageNumber="752">Breadth of braincase</th>
<td id="09065DA2AB7B006E6268E2480BBF8A0F" box="[542,751,813,832]" gridcol="1" gridrow="18" pageId="4" pageNumber="752">16.9</td>
<td id="09065DA2AB7B006E6379E2480AB68A0F" box="[783,998,813,832]" gridcol="2" gridrow="18" pageId="4" pageNumber="752">–</td>
<td id="09065DA2AB7B006E6470E2480D948A0F" box="[1030,1220,813,832]" gridcol="3" gridrow="18" pageId="4" pageNumber="752">15.1</td>
<td id="09065DA2AB7B006E6495E2480CF18A0F" box="[1251,1441,813,832]" gridcol="4" gridrow="18" pageId="4" pageNumber="752">11.4</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E22F0CF18A12" box="[164,1441,842,861]" gridrow="19" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E22F08AE8A12" box="[164,510,842,861]" gridcol="0" gridrow="19" pageId="4" pageNumber="752">Breadth of zygomatic plate</th>
<td id="09065DA2AB7B006E6268E22F0BBF8A12" box="[542,751,842,861]" gridcol="1" gridrow="19" pageId="4" pageNumber="752">7.8</td>
<td id="09065DA2AB7B006E6379E22F0AB68A12" box="[783,998,842,861]" gridcol="2" gridrow="19" pageId="4" pageNumber="752">6.1</td>
<td id="09065DA2AB7B006E6470E22F0D948A12" box="[1030,1220,842,861]" gridcol="3" gridrow="19" pageId="4" pageNumber="752">6.1</td>
<td id="09065DA2AB7B006E6495E22F0CF18A12" box="[1251,1441,842,861]" gridcol="4" gridrow="19" pageId="4" pageNumber="752">3.9</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E2020CF18A34" box="[164,1441,871,891]" gridrow="20" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E20208AE8A34" box="[164,510,871,891]" gridcol="0" gridrow="20" pageId="4" pageNumber="752">Head–body length</th>
<td id="09065DA2AB7B006E6268E2020BBF8A34" box="[542,751,871,891]" gridcol="1" gridrow="20" pageId="4" pageNumber="752">257</td>
<td id="09065DA2AB7B006E6379E2020AB68A34" box="[783,998,871,891]" gridcol="2" gridrow="20" pageId="4" pageNumber="752">312</td>
<td id="09065DA2AB7B006E6470E2020D948A34" box="[1030,1220,871,891]" gridcol="3" gridrow="20" pageId="4" pageNumber="752">261</td>
<td id="09065DA2AB7B006E6495E2020CF18A34" box="[1251,1441,871,891]" gridcol="4" gridrow="20" pageId="4" pageNumber="752">94</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E2E00CF18AD7" box="[164,1441,901,920]" gridrow="21" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E2E008AE8AD7" box="[164,510,901,920]" gridcol="0" gridrow="21" pageId="4" pageNumber="752">Tail length</th>
<td id="09065DA2AB7B006E6268E2E00BBF8AD7" box="[542,751,901,920]" gridcol="1" gridrow="21" pageId="4" pageNumber="752">254</td>
<td id="09065DA2AB7B006E6379E2E00AB68AD7" box="[783,998,901,920]" gridcol="2" gridrow="21" pageId="4" pageNumber="752">278</td>
<td id="09065DA2AB7B006E6470E2E00D948AD7" box="[1030,1220,901,920]" gridcol="3" gridrow="21" pageId="4" pageNumber="752">145*</td>
<td id="09065DA2AB7B006E6495E2E00CF18AD7" box="[1251,1441,901,920]" gridcol="4" gridrow="21" pageId="4" pageNumber="752">121</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E2C70CF18AFA" box="[164,1441,930,949]" gridrow="22" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E2C708AE8AFA" box="[164,510,930,949]" gridcol="0" gridrow="22" pageId="4" pageNumber="752">Hindfoot length (with claw)</th>
<td id="09065DA2AB7B006E6268E2C70BBF8AFA" box="[542,751,930,949]" gridcol="1" gridrow="22" pageId="4" pageNumber="752">45</td>
<td id="09065DA2AB7B006E6379E2C70AB68AFA" box="[783,998,930,949]" gridcol="2" gridrow="22" pageId="4" pageNumber="752">59</td>
<td id="09065DA2AB7B006E6470E2C70D948AFA" box="[1030,1220,930,949]" gridcol="3" gridrow="22" pageId="4" pageNumber="752">50</td>
<td id="09065DA2AB7B006E6495E2C70CF18AFA" box="[1251,1441,930,949]" gridcol="4" gridrow="22" pageId="4" pageNumber="752">25</td>
</tr>
<tr id="4AD734DEAB7B006E60D2E2DA0CF18A9C" box="[164,1441,959,979]" gridrow="23" pageId="4" pageNumber="752">
<th id="09065DA2AB7B006E60D2E2DA08AE8A9C" box="[164,510,959,979]" gridcol="0" gridrow="23" pageId="4" pageNumber="752">Length of internal side of ear</th>
<td id="09065DA2AB7B006E6268E2DA0BBF8A9C" box="[542,751,959,979]" gridcol="1" gridrow="23" pageId="4" pageNumber="752">18</td>
<td id="09065DA2AB7B006E6379E2DA0AB68A9C" box="[783,998,959,979]" gridcol="2" gridrow="23" pageId="4" pageNumber="752">20</td>
<td id="09065DA2AB7B006E6470E2DA0D948A9C" box="[1030,1220,959,979]" gridcol="3" gridrow="23" pageId="4" pageNumber="752">18</td>
<td id="09065DA2AB7B006E6495E2DA0CF18A9C" box="[1251,1441,959,979]" gridcol="4" gridrow="23" pageId="4" pageNumber="752">14</td>
</tr>
</table>
</paragraph>
<paragraph id="F458369CAB7BFF9560D2E29F0B248D42" blockId="4.[164,628,1018,1037]" box="[164,628,1018,1037]" pageId="4" pageNumber="752">
<tableNote id="09013712AB7BFF9560D2E29F0B248D42" box="[164,628,1018,1037]" pageId="4" pageNumber="752" targetBox="[164,1441,257,979]" targetPageId="4">
*The tail of
<collectionCode id="92F6AE59AB7BFF95615FE29F08348D42" box="[297,356,1018,1037]" pageId="4" pageNumber="752">NHM</collectionCode>
53.12.16.2 is incomplete.
</tableNote>
</paragraph>
<subSubSection id="BCFD6517AB7BFF9760D2E7E60D0C8BAB" lastPageId="6" lastPageNumber="754" pageId="4" pageNumber="752" type="description">
<paragraph id="F458369CAB7BFF9560D2E7E60CF28F99" blockId="4.[164,779,1667,1903]" lastBlockId="4.[826,1442,1116,1903]" pageId="4" pageNumber="752">
<emphasis id="C693EA8EAB7BFF9560D2E7E609B38FF9" italics="true" pageId="4" pageNumber="752">
Description (based on
<taxonomicName id="33E74D1FAB7BFF9561DBE7E60B0A8FD7" box="[429,602,1667,1688]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="desmarestii">M. desmarestii</taxonomicName>
and
<taxonomicName id="33E74D1FAB7BFF9562EBE7E60A5A8FD7" box="[669,778,1667,1688]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="luciae">M. luciae</taxonomicName>
only):
</emphasis>
Very large rats, as large or larger than any extant oryzomyines (
<tableCitation id="B9650327AB7BFF9561D2E7A508A78F99" box="[420,503,1728,1750]" captionStart="Table 1" captionStartId="4.[164,228,200,219]" captionTargetPageId="4" captionText="Table 1. Measurement data for Megalomys desmarestii, Megalomys luciae, and Oligoryzomys victus (mm)" pageId="4" pageNumber="752">Table 1</tableCitation>
). Dorsal pelage dark, blackish brown (almost black) (tan brown in
<specimenCount id="E2E1FD15AB7BFF956295E7BA085E8E5C" pageId="4" pageNumber="752" type="generic">one specimen</specimenCount>
in which the colour might have faded); ventral pelage abruptly paler in
<taxonomicName id="33E74D1FAB7BFF956258E6790B8B8E7E" box="[558,731,1820,1841]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB7BFF956258E6790B8B8E7E" box="[558,731,1820,1841]" italics="true" pageId="4" pageNumber="752">M. desmarestii</emphasis>
</taxonomicName>
, all white, except for a few darker guard hairs; in
<taxonomicName id="33E74D1FAB7BFF95629AE65E09B78E21" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="luciae">
<emphasis id="C693EA8EAB7BFF95629AE65E09B78E21" italics="true" pageId="4" pageNumber="752">M. luciae</emphasis>
</taxonomicName>
, general ventral colour is dark, with plumbeous base, but there are distinct paler areas under the chin and throat and under the lower thorax, the two regions being separated by a streak of darker fur (
<figureCitation id="6CDC2A19AB7BFF956334E5DD0AD48D81" box="[834,900,1208,1230]" captionStart="Figure 3" captionStartId="5.[144,223,1321,1340]" captionTargetBox="[367,1199,196,1291]" captionTargetId="figure-222@5.[367,1199,196,1291]" captionTargetPageId="5" captionText="Figure 3. Dorsal, lateral, and ventral views of Megalomys skins. Top, Megalomys desmarestii (NHM 55.12.24.201); bottom, Megalomys luciae (NHM 53.12.16.2). Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.5754703" httpUri="https://zenodo.org/record/5754703/files/figure.png" pageId="4" pageNumber="752">Fig. 3</figureCitation>
). Soft fur, subauricular patches absent. Pinnae small, not reaching eye when laid forward. Mystacial and superciliary vibrissae not extending posteriorly beyond pinnae when laid back. Manual claws small and unkeeled. Hindfoot without well-developed natatory fringes and interdigital webs; tufts of ungual hairs at bases of claws on dII–dV vestigial or absent; plantar surface covered with distinct squamae distal to thenar pad; hypothenar pad absent or vestigial in
<taxonomicName id="33E74D1FAB7BFF95634CE4A90AFA8CAF" box="[826,938,1483,1505]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="luciae">
<emphasis id="C693EA8EAB7BFF95634CE4A90AFA8CAF" box="[826,938,1483,1505]" italics="true" pageId="4" pageNumber="752">M. luciae</emphasis>
</taxonomicName>
, present but reduced in
<taxonomicName id="33E74D1FAB7BFF95649CE4A90CCA8CAF" box="[1258,1434,1483,1505]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB7BFF95649CE4A90CCA8CAF" box="[1258,1434,1483,1505]" italics="true" pageId="4" pageNumber="752">M. desmarestii</emphasis>
</taxonomicName>
; interdigital pads large and fleshy, pads 1 and 4 contiguous with 2 and 3; claw of digit I (dI) extending just beyond first interphalangeal joint of dII; claw of dV extending to middle of second phalange of dIV (
<figureCitation id="6CDC2A19AB7BFF956334E7010AD58F35" box="[834,901,1636,1658]" captionStart="Figure 4" captionStartId="6.[164,243,938,957]" captionTargetBox="[282,660,196,908]" captionTargetId="figure-592@6.[282,660,196,908]" captionTargetPageId="6" captionText="Figure 4. Plantar view of left hindfoot of Megalomys desmarestii (NHM 50.11.30.6). Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5754705" httpUri="https://zenodo.org/record/5754705/files/figure.png" pageId="4" pageNumber="752">Fig. 4</figureCitation>
). Tail about same length as head and body, sparsely haired and covered with conspicuous epidermal scales; without tuft of long terminal hairs, and unicoloured (dark). Mammary complement unknown.
</paragraph>
<paragraph id="F458369CAB7BFF976325E7BB09AC8C07" blockId="4.[826,1442,1116,1903]" lastBlockId="6.[164,779,1085,1902]" lastPageId="6" lastPageNumber="754" pageId="4" pageNumber="752">
Skull with stout rostrum flanked by moderate or deep zygomatic notches; interorbital region convergent anteriorly with well-developed supraorbital crests (symmetrically constricted in an older adult specimen of
<taxonomicName id="33E74D1FAB7BFF9563BCE63C0D248E21" box="[970,1140,1881,1902]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="752" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB7BFF9563BCE63C0D248E21" box="[970,1140,1881,1902]" italics="true" pageId="4" pageNumber="752">M. desmarestii</emphasis>
</taxonomicName>
); braincase squared, with well-developed temporal crests; lambdoidal and nuchal crests well developed in older adults. Posterior margin of zygomatic plate dorsal to M1 alveolus; zygomatic plate lacks an anterodorsal spinous process. Jugal present and small (the maxillary and squamosal zygomatic processes overlap). Nasal bones with acutely angled posterior nasal margins; extending posteriorly behind lacrimals; lacrimals sutured mainly to maxillary. Posterior wall of the orbit with postorbital ridge (faint in
<taxonomicName id="33E74D1FAB7AFF9461B2E7C70B608FF9" box="[452,560,1697,1719]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="753" phylum="Chordata" rank="species" species="luciae">
<emphasis id="C693EA8EAB7AFF9461B2E7C70B608FF9" box="[452,560,1697,1719]" italics="true" pageId="5" pageNumber="753">M. luciae</emphasis>
</taxonomicName>
). Frontosquamosal suture anterior to frontoparietal suture. Parietals with broad lateral expansions. Incisive foramina small, not extending posteriorly between M1 alveoli, spindle-shaped. Bony palate between the molar rows is smooth or weakly sculpted. Posterolateral palatal pits large, complex, and recessed in deep fossae; mesopterygoid fossa penetrating anteriorly between maxillae; bony roof of mesopterygoid fossa completely ossified in an older adult specimen of
<taxonomicName id="33E74D1FAB7AFF9464ABE4A90CD68CAF" box="[1245,1414,1483,1505]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="753" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB7AFF9464ABE4A90CD68CAF" box="[1245,1414,1483,1505]" italics="true" pageId="5" pageNumber="753">M. desmarestii</emphasis>
</taxonomicName>
, perforated by small (slit-like) sphenopalatine vacuities in a younger specimen of
<taxonomicName id="33E74D1FAB7AFF9464F8E76C0C698F52" box="[1166,1337,1544,1566]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="753" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB7AFF9464F8E76C0C698F52" box="[1166,1337,1544,1566]" italics="true" pageId="5" pageNumber="753">M. desmarestii</emphasis>
</taxonomicName>
and in
<taxonomicName id="33E74D1FAB7AFF946350E7420ACC8F73" box="[806,924,1575,1596]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="753" phylum="Chordata" rank="species" species="luciae">
<emphasis id="C693EA8EAB7AFF946350E7420ACC8F73" box="[806,924,1575,1596]" italics="true" pageId="5" pageNumber="753">M. luciae</emphasis>
</taxonomicName>
. Alisphenoid strut absent (buccinator– masticatory foramen and accessory foramen ovale confluent). Alisphenoid canal with large anterior opening. Stapedial foramen and posterior opening of alisphenoid canal small; squamosal–alisphenoid groove and sphenofrontal foramen absent; secondary anastomosis of internal carotid crosses dorsal surface of pterygoid plate (= carotid circulatory pattern 3 of
<bibRefCitation id="90764B6DAB7AFF946350E6790AFD8E7D" author="Voss RS" box="[806,941,1820,1842]" pageId="5" pageNumber="753" pagination="259 - 493" refId="ref16452" refString="Voss RS. 1988. Systematics and ecology of ichthyomyine rodents (Muroidea): patterns of morphological evolution in a small adaptive radiation. Bulletin of the American Museum of Natural History 188: 259 - 493." type="journal article" year="1988">Voss, 1988</bibRefCitation>
). Posterior suspensory process of the squamosal absent. Postglenoid foramen small and dorsoventrally compressed, covered by tegmen tympani; subsquamosal fenestra vestigial or absent. Periotic exposed posteromedially, between ectotympanic and basioccipital, extending anteriorly to carotid canal; mastoid unfenestrated. Capsular process of lower incisor alveolus well developed in adult specimens (smaller in
<taxonomicName id="33E74D1FAB79FF976184E5B20B0D8DA4" box="[498,605,1238,1260]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="species" species="luciae">
<emphasis id="C693EA8EAB79FF976184E5B20B0D8DA4" box="[498,605,1238,1260]" italics="true" pageId="6" pageNumber="754">M. luciae</emphasis>
</taxonomicName>
); superior and inferior masseteric ridges joined anteriorly as open chevron below m1 (slightly anterior in
<taxonomicName id="33E74D1FAB79FF97622DE4710A538C67" box="[603,771,1299,1321]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB79FF97622DE4710A538C67" box="[603,771,1299,1321]" italics="true" pageId="6" pageNumber="754">M. desmarestii</emphasis>
</taxonomicName>
) (
<figureCitation id="6CDC2A19AB79FF9760DAE45709BE8C07" box="[172,238,1330,1352]" captionStart="Figure 5" captionStartId="7.[144,223,1302,1321]" captionTargetBox="[144,1425,196,1273]" captionTargetId="figure-220@7.[144,1425,196,1273]" captionTargetPageId="7" captionText="Figure 5. Cranium and mandible of Megalomys. Left to right: Megalomys desmarestii (NHM 50.11.30.6) and Megalomys luciae (NHM 53.12.16.2). Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5754707" httpUri="https://zenodo.org/record/5754707/files/figure.png" pageId="6" pageNumber="754">Fig. 5</figureCitation>
).
</paragraph>
<caption id="A0986614AB7AFF9460E6E44C0A758C15" ID-DOI="http://doi.org/10.5281/zenodo.5754703" ID-Zenodo-Dep="5754703" httpUri="https://zenodo.org/record/5754703/files/figure.png" pageId="5" pageNumber="753" startId="5.[144,223,1321,1340]" targetBox="[367,1199,196,1291]" targetPageId="5">
<paragraph id="F458369CAB7AFF9460E6E44C0A758C15" blockId="5.[144,1421,1321,1370]" pageId="5" pageNumber="753">
<emphasis id="C693EA8EAB7AFF9460E6E44C09A88C73" bold="true" box="[144,248,1321,1340]" pageId="5" pageNumber="753">Figure 3.</emphasis>
Dorsal, lateral, and ventral views of
<taxonomicName id="33E74D1FAB7AFF9462D3E44C0A4A8C73" authorityName="TROUESSART" authorityYear="1881" box="[677,794,1321,1340]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="753" phylum="Chordata" rank="genus">
<emphasis id="C693EA8EAB7AFF9462D3E44C0A4A8C73" box="[677,794,1321,1340]" italics="true" pageId="5" pageNumber="753">Megalomys</emphasis>
</taxonomicName>
skins. Top,
<taxonomicName id="33E74D1FAB7AFF9463DEE44C0DF18C73" box="[936,1185,1321,1340]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="753" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB7AFF9463DEE44C0DF18C73" box="[936,1185,1321,1340]" italics="true" pageId="5" pageNumber="753">Megalomys desmarestii</emphasis>
</taxonomicName>
(
<collectionCode id="92F6AE59AB7AFF9464C2E44C0DA08C73" box="[1204,1264,1321,1340]" pageId="5" pageNumber="753">NHM</collectionCode>
55.12.24.201); bottom,
<taxonomicName id="33E74D1FAB7AFF94609EE42308F48C16" box="[232,420,1350,1369]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="753" phylum="Chordata" rank="species" species="luciae">
<emphasis id="C693EA8EAB7AFF94609EE42308F48C16" box="[232,420,1350,1369]" italics="true" pageId="5" pageNumber="753">Megalomys luciae</emphasis>
</taxonomicName>
(
<collectionCode id="92F6AE59AB7AFF9461C5E42208BE8C15" box="[435,494,1351,1370]" pageId="5" pageNumber="753">NHM</collectionCode>
53.12.16.2). Scale bar: 5 cm.
</paragraph>
</caption>
<caption id="A0986614AB79FF9760D2E2CF0B2A8A94" ID-DOI="http://doi.org/10.5281/zenodo.5754705" ID-Zenodo-Dep="5754705" httpUri="https://zenodo.org/record/5754705/files/figure.png" pageId="6" pageNumber="754" startId="6.[164,243,938,957]" targetBox="[282,660,196,908]" targetPageId="6">
<paragraph id="F458369CAB79FF9760D2E2CF0B2A8A94" blockId="6.[164,778,938,987]" pageId="6" pageNumber="754">
<emphasis id="C693EA8EAB79FF9760D2E2CF085C8AF2" bold="true" box="[164,268,938,957]" pageId="6" pageNumber="754">Figure 4.</emphasis>
Plantar view of left hindfoot of
<taxonomicName id="33E74D1FAB79FF976213E2CF09AA8A95" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB79FF976213E2CF09AA8A95" italics="true" pageId="6" pageNumber="754">Megalomys desmarestii</emphasis>
</taxonomicName>
(
<collectionCode id="92F6AE59AB79FF97617CE2A208158A95" box="[266,325,967,986]" pageId="6" pageNumber="754">NHM</collectionCode>
50.11.30.6). Scale bar: 1 cm.
</paragraph>
</caption>
<paragraph id="F458369CAB79FF9760CAE43408518E21" blockId="6.[164,779,1085,1902]" pageId="6" pageNumber="754">
Upper incisors with smoothly rounded enamel bands. Maxillary tooth rows parallel. Molars bunodont and with labial flexi enclosed by a cingulum; labial and lingual flexi of M1 and M2 meet at midline, enamel overlaps. M1 anterocone divided into anterolabial and anterolingual conules by shallow anteromedian flexus; anteroloph well developed and fused with anterostyle on labial cingulum, separated from anterocone by persistent anteroflexus; protostyle absent; mesolophs present on all upper molars; paracone connected by enamel bridge to anterior or middle moiety of protocone; median mure connected to protocone. M2 protoflexus absent; mesoflexus present as single internal fossette; paracone without an accessory loph. M3 with posteroloph and developed hypoflexus (the latter remaining excavated with moderate to heavy wear) (
<figureCitation id="6CDC2A19AB79FF97612FE65F08978E1F" box="[345,455,1850,1872]" captionStart="Figure 6" captionStartId="8.[164,243,954,973]" captionTargetBox="[323,1283,196,924]" captionTargetId="figure-406@8.[323,1283,196,924]" captionTargetPageId="8" captionText="Figure 6. Maxillary molars of West Indian rice rats. A, Megalomys desmarestii (NHM 55.12.24.201). B, Megalomys luciae (NHM 53.12.16.2). C, Oligoryzomys victus (NHM 97.12.26.1). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5754709" httpUri="https://zenodo.org/record/5754709/files/figure.png" pageId="6" pageNumber="754">Fig 6A, B</figureCitation>
). Accessory labial root of M1 present.
</paragraph>
<paragraph id="F458369CAB79FF976325E1A30C2988FE" blockId="6.[826,1442,198,740]" pageId="6" pageNumber="754">
m1 anteroconid without an anteromedian flexid; anterolabial cingulum present on all lower molars; ectolophid absent on m1 and m2; anterolophid present on m2 and m3; mesolophid distinct on unworn m1 and m2; m2 hypoflexid short; posteroflexid present on m3 (
<figureCitation id="6CDC2A19AB79FF976441E03A0DDE883A" box="[1079,1166,351,373]" captionStart="Figure 7" captionStartId="9.[144,223,1042,1061]" captionTargetBox="[303,1263,197,1011]" captionTargetId="figure-359@9.[303,1263,197,1012]" captionTargetPageId="9" captionText="Figure 7. Mandibular molars of West Indian rice rats. A, Megalomys desmarestii (NHM 55.12.24.201). B, Megalomys audreyae (NHM M7406a) (reversed for comparison). C, Oligoryzomys victus (NHM 97.12.26.1). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5754711" httpUri="https://zenodo.org/record/5754711/files/figure.png" pageId="6" pageNumber="754">Fig. 7A</figureCitation>
). Accessory lingual and labial roots of m1 present; m2 and m3 each with two small anterior roots and one large posterior root.
</paragraph>
<paragraph id="F458369CAB79FF976325E0DE0DB68BC7" blockId="6.[826,1442,198,740]" pageId="6" pageNumber="754">First rib with double articulation. Humerus without entepicondylar and supratrochlear foramina. Twelve ribs (false thirteenth rib present in older adult specimen). Fifth lumbar (seventeenth thoracicolumbar) vertebra with well-developed anapophysis. Hemal arch between second and third caudal vertebrae, with posterior spinous process.</paragraph>
<paragraph id="F458369CAB79FF976325E3F40D0C8BAB" blockId="6.[826,1442,198,740]" pageId="6" pageNumber="754">Conditions of soft anatomy (stomach, liver, and male accessory reproductive glands) and glans penis characters are unknown.</paragraph>
</subSubSection>
<subSubSection id="BCFD6517AB79FF96634CE26E08EF8F12" lastPageId="7" lastPageNumber="755" pageId="6" pageNumber="754" type="discussion">
<paragraph id="F458369CAB79FF97634CE26E0DAB8F51" blockId="6.[826,1441,779,1903]" pageId="6" pageNumber="754">
<emphasis id="C693EA8EAB79FF97634CE26E0AF68A6F" box="[826,934,779,800]" italics="true" pageId="6" pageNumber="754">Remarks:</emphasis>
The complex early taxonomic history of
<taxonomicName id="33E74D1FAB79FF97634CE24F0AE98A70" authorityName="TROUESSART" authorityYear="1881" box="[826,953,810,831]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="genus">
<emphasis id="C693EA8EAB79FF97634CE24F0AE98A70" box="[826,953,810,831]" italics="true" pageId="6" pageNumber="754">Megalomys</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="90764B6DAB79FF9764E0E24F0C7B8A0F" author="Tate GHH" box="[1174,1323,810,832]" pageId="6" pageNumber="754" pagination="1 - 19" refId="ref16152" refString="Tate GHH. 1932 a. The taxonomic history of the Neotropical cricetid genera Holochilus, Nectomys, Scapteromys, Megalomys, Tylomys and Ototylomys. American Museum Novitates 562: 1 - 19." type="journal article" year="1932">Tate (1932a)</bibRefCitation>
. Although historical museum specimens of
<taxonomicName id="33E74D1FAB79FF9764C8E22C0C3B8A12" box="[1214,1387,840,862]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="species" species="desmarestii">
<emphasis id="C693EA8EAB79FF9764C8E22C0C3B8A12" box="[1214,1387,840,862]" italics="true" pageId="6" pageNumber="754">M. desmarestii</emphasis>
</taxonomicName>
and
<taxonomicName id="33E74D1FAB79FF97634CE20D0AF68A33" box="[826,934,871,893]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="species" species="luciae">
<emphasis id="C693EA8EAB79FF97634CE20D0AF68A33" box="[826,934,871,893]" italics="true" pageId="6" pageNumber="754">M. luciae</emphasis>
</taxonomicName>
have permitted redescription of the genus and comparison of
<taxonomicName id="33E74D1FAB79FF976460E2E30DC58AD4" authorityName="TROUESSART" authorityYear="1881" box="[1046,1173,902,923]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="genus">
<emphasis id="C693EA8EAB79FF976460E2E30DC58AD4" box="[1046,1173,902,923]" italics="true" pageId="6" pageNumber="754">Megalomys</emphasis>
</taxonomicName>
with other extinct rice rats from the Windward and Leeward Islands, the status of other species that have been assigned to the genus requires further clarification through future research. The limited
<typeStatus id="2B5C883EAB79FF976432E5650D258D5A" box="[1092,1141,1024,1045]" pageId="6" pageNumber="754">type</typeStatus>
material of
<taxonomicName id="33E74D1FAB79FF976579E5640CF18D5A" box="[1295,1441,1024,1046]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="species" species="audreyae">
<emphasis id="C693EA8EAB79FF976579E5640CF18D5A" box="[1295,1441,1024,1046]" italics="true" pageId="6" pageNumber="754">M. audreyae</emphasis>
</taxonomicName>
(
<figureCitation id="6CDC2A19AB79FF976334E57A0AC58D7B" box="[834,917,1055,1077]" captionStart="Figure 7" captionStartId="9.[144,223,1042,1061]" captionTargetBox="[303,1263,197,1011]" captionTargetId="figure-359@9.[303,1263,197,1012]" captionTargetPageId="9" captionText="Figure 7. Mandibular molars of West Indian rice rats. A, Megalomys desmarestii (NHM 55.12.24.201). B, Megalomys audreyae (NHM M7406a) (reversed for comparison). C, Oligoryzomys victus (NHM 97.12.26.1). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5754711" httpUri="https://zenodo.org/record/5754711/files/figure.png" pageId="6" pageNumber="754">Fig. 7B</figureCitation>
) is difficult to evaluate, and the genus-level identity of this species remains unclear. Abundant fossil and zooarchaeological rice rat material is known from the
<collectingCountry id="8CF0760CAB79FF97638CE51F0D068DDF" box="[1018,1110,1146,1168]" name="Antigua and Barbuda" pageId="6" pageNumber="754">Antigua</collectingCountry>
–
<collectingCountry id="8CF0760CAB79FF976413E51F0D978DDF" box="[1125,1223,1146,1168]" name="Antigua and Barbuda" pageId="6" pageNumber="754">Barbuda</collectingCountry>
Bank (
<bibRefCitation id="90764B6DAB79FF97656BE51F0A238DE0" author="Wing ES &amp; Hoffman CA Jr &amp; Ray CE" pageId="6" pageNumber="754" pagination="123 - 139" refId="ref16950" refString="Wing ES, Hoffman CA Jr, Ray CE. 1968. Vertebrate remains from Indian sites on Antigua, West Indies. Caribbean Journal of Science 8: 123 - 139." type="journal article" year="1968">
Wing
<emphasis id="C693EA8EAB79FF976515E51E0CCA8DC0" box="[1379,1434,1146,1168]" italics="true" pageId="6" pageNumber="754">et al.</emphasis>
, 1968
</bibRefCitation>
;
<bibRefCitation id="90764B6DAB79FF9763F1E5FC0DCE8DE1" author="Steadman DW &amp; Pregill GK &amp; Olson SL" box="[903,1182,1177,1199]" pageId="6" pageNumber="754" pagination="4448 - 4451" refId="ref15964" refString="Steadman DW, Pregill GK, Olson SL. 1984 a. Fossil vertebrates from Antigua, Lesser Antilles: evidence for late Holocene human-caused extinctions in the West Indies. Proceedings of the National Academy of Sciences of the United States of America 81: 4448 - 4451." type="journal article" year="1984">
Steadman
<emphasis id="C693EA8EAB79FF97647CE5FF0D128DE1" box="[1034,1090,1177,1199]" italics="true" pageId="6" pageNumber="754">et al.</emphasis>
, 1984a
</bibRefCitation>
;
<bibRefCitation id="90764B6DAB79FF9764C4E5FC0C118DE0" author="Jones AR" box="[1202,1345,1177,1199]" pageId="6" pageNumber="754" pagination="518 - 536" refId="ref14293" refString="Jones AR. 1985. Dietary change and human population at Indian Creek, Antigua. American Antiquity 50: 518 - 536." type="journal article" year="1985">Jones, 1985</bibRefCitation>
;
<bibRefCitation id="90764B6DAB79FF976523E5FC0AE88D82" author="Pregill GK &amp; Steadman DW &amp; Watters DR" pageId="6" pageNumber="754" pagination="1 - 51" refId="ref15611" refString="Pregill GK, Steadman DW, Watters DR. 1994. Late Quaternary vertebrate faunas of the Lesser Antilles: historical components of Caribbean biogeography. Bulletin of the Carnegie Museum of Natural History 30: 1 - 51." type="journal article" year="1994">
Pregill
<emphasis id="C693EA8EAB79FF97634CE5DD0A228D82" box="[826,882,1208,1229]" italics="true" pageId="6" pageNumber="754">et al.</emphasis>
, 1994
</bibRefCitation>
;
<bibRefCitation id="90764B6DAB79FF9763B5E5DD0C578D82" author="MacPhee RDE &amp; Flemming C" box="[963,1287,1208,1230]" pageId="6" pageNumber="754" pagination="333 - 371" refId="ref14967" refString="MacPhee RDE, Flemming C. 1999. Requiem aeternam: the last five hundred years of mammalian species extinctions. In: MacPhee RDE, ed. Extinctions in near time: causes, contexts, and consequences. New York: Kluwer Academic / Plenum, 333 - 371." type="book chapter" year="1999">MacPhee &amp; Flemming, 1999</bibRefCitation>
;
<bibRefCitation id="90764B6DAB79FF976564E5DD0AAD8DA3" author="Steadman DW &amp; Hilgartner WB" pageId="6" pageNumber="754" pagination="75 - 83" refId="ref15928" refString="Steadman DW, Hilgartner WB. 1999. A new species of extinct barn owl (Aves: Tyto) from Barbuda, Lesser Antilles. Smithsonian Contributions to Paleobiology 89: 75 - 83." type="journal article" year="1999">Steadman &amp; Hilgartner, 1999</bibRefCitation>
), and fossil material from
<collectingCountry id="8CF0760CAB79FF976548E5B30CF18DA3" box="[1342,1441,1238,1260]" name="Antigua and Barbuda" pageId="6" pageNumber="754">Barbuda</collectingCountry>
has been named ‘
<emphasis id="C693EA8EAB79FF976469E5900C628C45" box="[1055,1330,1269,1290]" italics="true" pageId="6" pageNumber="754">Ekbletomys hypenemus</emphasis>
’ by
<bibRefCitation id="90764B6DAB79FF976502E5900AD48C66" author="Ray CE" pageId="6" pageNumber="754" refId="ref15681" refString="Ray CE. 1962. Oryzomyine rodents of the Antillean subregion. Unpublished PhD thesis, Harvard University." type="book" year="1962">Ray (1962)</bibRefCitation>
; however, this taxon remains a
<emphasis id="C693EA8EAB79FF97648CE4710CF18C67" box="[1274,1441,1299,1321]" italics="true" pageId="6" pageNumber="754">nomen nudum</emphasis>
because Ray’s work has never been formally published. Although further material apparently referable to
<taxonomicName id="33E74D1FAB79FF9763ACE4150D218CCB" box="[986,1137,1391,1413]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="species" species="audreyae">
<emphasis id="C693EA8EAB79FF9763ACE4150AA18CCA" box="[986,1009,1392,1413]" italics="true" pageId="6" pageNumber="754">M</emphasis>
.
<emphasis id="C693EA8EAB79FF97647CE40A0D218CCB" box="[1034,1137,1391,1412]" italics="true" pageId="6" pageNumber="754">audreyae</emphasis>
</taxonomicName>
has been reported and radiometrically dated by
<bibRefCitation id="90764B6DAB79FF9764F3E4EB0AD48C8D" author="MacPhee RDE &amp; Flemming C" pageId="6" pageNumber="754" pagination="333 - 371" refId="ref14967" refString="MacPhee RDE, Flemming C. 1999. Requiem aeternam: the last five hundred years of mammalian species extinctions. In: MacPhee RDE, ed. Extinctions in near time: causes, contexts, and consequences. New York: Kluwer Academic / Plenum, 333 - 371." type="book chapter" year="1999">MacPhee &amp; Flemming (1999)</bibRefCitation>
, the identity of rice rats from the
<collectingCountry id="8CF0760CAB79FF976541E4C90CC28C8D" box="[1335,1426,1452,1474]" name="Antigua and Barbuda" pageId="6" pageNumber="754">Antigua</collectingCountry>
–
<collectingCountry id="8CF0760CAB79FF97634CE4AE0ACD8CAE" box="[826,925,1483,1505]" name="Antigua and Barbuda" pageId="6" pageNumber="754">Barbuda</collectingCountry>
Bank cannot be determined in the absence of further investigation of this material, which is beyond the scope of the present study.
</paragraph>
<paragraph id="F458369CAB79FF966325E74208EF8F12" blockId="6.[826,1441,779,1903]" lastBlockId="7.[144,759,1393,1629]" lastPageId="7" lastPageNumber="755" pageId="6" pageNumber="754">
Two other large-bodied extinct insular cricetid rodents,
<taxonomicName id="33E74D1FAB79FF9763D3E7200C0E8F14" authority="Niethammer, 1964" authorityName="Niethammer" authorityYear="1964" box="[933,1374,1605,1627]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="species" species="curioi">
<emphasis id="C693EA8EAB79FF9763D3E7200D248F14" box="[933,1140,1605,1627]" italics="true" pageId="6" pageNumber="754">Megalomys curioi</emphasis>
Niethammer, 1964
</taxonomicName>
from the Galápagos Islands and
<taxonomicName id="33E74D1FAB79FF9764ECE7010A8C8FD7" authority="Hooijer, 1959" authorityName="Hooijer" authorityYear="1959" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="species" species="curazensis">
<emphasis id="C693EA8EAB79FF9764ECE7010CF18F35" box="[1178,1441,1636,1658]" italics="true" pageId="6" pageNumber="754">Megalomys curazensis</emphasis>
<bibRefCitation id="90764B6DAB79FF97634CE7E70A8C8FD7" author="Hooijer DA" box="[826,988,1666,1688]" pageId="6" pageNumber="754" pagination="1 - 27" refId="ref14028" refString="Hooijer DA. 1959. Fossil rodents from Curacao and Bonaire. Studies on the Fauna of Curacao and other Caribbean Islands 9: 1 - 27." type="journal article" year="1959">Hooijer, 1959</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="8CF0760CAB79FF976443E7E60DC68FD7" box="[1077,1174,1667,1688]" name="Curaçao" pageId="6" pageNumber="754">Curaçao</collectingCountry>
, were also originally referred to the genus.
<taxonomicName id="33E74D1FAB79FF976422E7C40C738FF8" authorityName="Niethammer" authorityYear="1964" box="[1108,1315,1697,1719]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="species" species="curioi">
<emphasis id="C693EA8EAB79FF976422E7C40C738FF8" box="[1108,1315,1697,1719]" italics="true" pageId="6" pageNumber="754">Megalomys curioi</emphasis>
</taxonomicName>
has since been transferred to the new genus
<taxonomicName id="33E74D1FAB79FF97648DE7A50D2B8FBB" authority="(Lenglet &amp; Coppois, 1979)" baseAuthorityName="Lenglet &amp; Coppois" baseAuthorityYear="1979" class="Mammalia" family="Cricetidae" genus="Megaoryzomys" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="754" phylum="Chordata" rank="genus">
<emphasis id="C693EA8EAB79FF97648DE7A50CF18F9A" box="[1275,1441,1728,1749]" italics="true" pageId="6" pageNumber="754">Megaoryzomys</emphasis>
(
<bibRefCitation id="90764B6DAB79FF976334E7BB0D228FBB" author="Lenglet G &amp; Coppois G" box="[834,1138,1758,1780]" pageId="6" pageNumber="754" pagination="632 - 648" refId="ref14564" refString="Lenglet G, Coppois G. 1979. Description du crane et de quelques ossements d'un genre nouveau eteint de Cricetidae (Mammalia-Rodentia) geant des Galapagos: Megaoryzomys (gen. nov.). Bulletin de l'Academie Royale de Belgique (Classe des Sciences), 5 th Series 65: 632 - 648." type="book chapter" year="1979">Lenglet &amp; Coppois, 1979</bibRefCitation>
)
</taxonomicName>
and reinterpreted as a thomasomyine (
<bibRefCitation id="90764B6DAB79FF976399E7980DA68E5C" author="Steadman DW &amp; Ray CE" box="[1007,1270,1789,1811]" pageId="6" pageNumber="754" pagination="1 - 23" refId="ref16014" refString="Steadman DW, Ray CE. 1982. The relationships of Megaoryzomys curioi, an extinct cricetine rodent (Muroidea: Muridae) from the Galapagos Islands, Ecuador. Smithsonian Contributions to Paleobiology 51: 1 - 23." type="journal article" year="1982">Steadman &amp; Ray, 1982</bibRefCitation>
). The southern
<collectingCountry id="8CF0760CAB79FF97634CE67E0D678E7E" box="[826,1079,1819,1841]" name="Curaçao" pageId="6" pageNumber="754">Netherlands Antilles</collectingCountry>
show little biogeographical affinity with the Windward and Leeward Islands, and have close faunal and floral similarities to mainland South America (
<bibRefCitation id="90764B6DAB78FF96612DE4140B508CC8" author="Hooijer DA" box="[347,512,1393,1415]" pageId="7" pageNumber="755" pagination="1 - 27" refId="ref14028" refString="Hooijer DA. 1959. Fossil rodents from Curacao and Bonaire. Studies on the Fauna of Curacao and other Caribbean Islands 9: 1 - 27." type="journal article" year="1959">Hooijer, 1959</bibRefCitation>
,
<bibRefCitation id="90764B6DAB78FF966260E4140B1F8CC8" author="Hooijer DA" box="[534,591,1393,1415]" pageId="7" pageNumber="755" pagination="531 - 532" refId="ref14056" refString="Hooijer DA. 1966. Fossil mammals of the Netherlands Antilles. Archives Neerlandaises de Zoologie 16: 531 - 532." type="journal article" year="1966">1966</bibRefCitation>
,
<bibRefCitation id="90764B6DAB78FF966213E4140BCE8CC8" author="Hooijer DA" box="[613,670,1393,1415]" pageId="7" pageNumber="755" pagination="399 - 406" refId="ref14078" refString="Hooijer DA. 1967. Pleistocene vertebrates of the Netherlands Antilles. In: Martin PS, Wright HE Jr, eds. Pleistocene extinctions: the search for a cause. New Haven: Yale University Press, 399 - 406." type="book chapter" year="1967">1967</bibRefCitation>
; Trejo-Torres &amp; Ackerman, 2001;
<bibRefCitation id="90764B6DAB78FF9661A9E4EA09998C8C" author="Vazquez-Miranda H &amp; Navarro-Siguenza AG &amp; Morrone JJ" pageId="7" pageNumber="755" pagination="180 - 200" refId="ref16419" refString="Vazquez-Miranda H, Navarro-Siguenza AG, Morrone JJ. 2007. Biogeographical patterns of the avifaunas of the Caribbean Basin Islands: a parsimony perspective. Cladistics 23: 180 - 200." type="journal article" year="2007">
Vázquez-Miranda
<emphasis id="C693EA8EAB78FF9662CEE4F50BA08CEB" box="[696,752,1423,1445]" italics="true" pageId="7" pageNumber="755">et al.</emphasis>
, 2007
</bibRefCitation>
;
<bibRefCitation id="90764B6DAB78FF9660A3E4CA08808C8B" author="Voss RS &amp; Weksler M" box="[213,464,1454,1476]" pageId="7" pageNumber="755" pagination="1 - 7" refId="ref16491" refString="Voss RS, Weksler M. 2009. On the taxonomic status of Oryzomys curasoae McFarlane and Debrot, 2001, with remarks on the phylogenetic relationships of O. gorgasi Hershkovitz, 1971. Caribbean Journal of Science 45: 1 - 7." type="journal article" year="2009">Voss &amp; Weksler, 2009</bibRefCitation>
), and it has been hypothesized that
<taxonomicName id="33E74D1FAB78FF96616FE4A808E78CAD" authorityName="Hooijer" authorityYear="1959" box="[281,439,1485,1506]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="755" phylum="Chordata" rank="species" species="curazensis">
<emphasis id="C693EA8EAB78FF96616FE4A808608CAD" box="[281,304,1485,1506]" italics="true" pageId="7" pageNumber="755">M</emphasis>
.
<emphasis id="C693EA8EAB78FF966137E4A808E78CAD" box="[321,439,1485,1506]" italics="true" pageId="7" pageNumber="755">curazensis</emphasis>
</taxonomicName>
is derived from a different mainland oryzomyine ancestor (
<bibRefCitation id="90764B6DAB78FF96627CE48E085B8F6F" author="McFarlane DA &amp; Lundberg J" pageId="7" pageNumber="755" pagination="278 - 281" refId="ref14849" refString="McFarlane DA, Lundberg J. 2002. A Middle Pleistocene age and biogeography for the extinct rodent Megalomys curazensis from Curacao, Netherlands Antilles. Caribbean Journal of Science 38: 278 - 281." type="journal article" year="2002">McFarlane &amp; Lundberg, 2002</bibRefCitation>
). A redescription of
<taxonomicName id="33E74D1FAB78FF96619AE76E0BD88F6F" authorityName="Hooijer" authorityYear="1959" box="[492,648,1546,1568]" class="Mammalia" family="Cricetidae" genus="Megalomys" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="755" phylum="Chordata" rank="species" species="curazensis">
<emphasis id="C693EA8EAB78FF96619AE76E0B538F6F" box="[492,515,1547,1568]" italics="true" pageId="7" pageNumber="755">M</emphasis>
.
<emphasis id="C693EA8EAB78FF966264E76E0BD88F6F" box="[530,648,1547,1568]" italics="true" pageId="7" pageNumber="755">curazensis</emphasis>
</taxonomicName>
will form the basis for a future study in this series, and it is not considered further herein.
</paragraph>
</subSubSection>
</treatment>
</document>
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