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<document id="F999481E517B8E1E90B471E250CB048F" ID-CLB-Dataset="3888" ID-DOI="10.1080/0022293021000028252" ID-GBIF-Dataset="79cf214d-c7d5-4053-a49c-07cfd914f6ad" ID-ISSN="1464-5262" ID-Zenodo-Dep="5258614" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629911030116" checkinUser="felipe" docAuthor="Wang, Hongzhu &amp; Erséus, Christer" docDate="2004" docId="704ECD36FFF32173FDEAEE97CE43DA2C" docLanguage="en" docName="JNatHist.38.3.269-299.pdf" docOrigin="Journal of Natural History 38 (3)" docSource="http://www.tandfonline.com/doi/abs/10.1080/0022293021000028252" docStyle="DocumentStyle:443144A73DAE3E092C51A378E4CD0138.2:JNatHist.2003-2004.journal_article.1cover" docStyleId="443144A73DAE3E092C51A378E4CD0138" docStyleName="JNatHist.2003-2004.journal_article.1cover" docStyleVersion="2" docTitle="Doliodrilus tener Erseus 1984" docType="treatment" docVersion="8" lastPageNumber="274" masterDocId="8C77B54EFFF72175FF87EE22CC33DB61" masterDocTitle="New species of Doliodrilus and other Limnodriloidinae (Oligochaeta, Tubificidae) from Hainan and other parts of the north-west Pacific Ocean" masterLastPageNumber="299" masterPageNumber="269" pageNumber="272" updateTime="1699144775815" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D8BFD27343172FA8F4EAA0A1A9A33A3F">New species of Doliodrilus and other Limnodriloidinae (Oligochaeta, Tubificidae) from Hainan and other parts of the north-west Pacific Ocean</mods:title>
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<mods:name id="F406DD5CCA6E3480337F566CED5A6AD2" type="personal">
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<mods:namePart id="140EBDB20EE34A575AC0D2B678D77296">Wang, Hongzhu</mods:namePart>
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<mods:namePart id="283931980728F10446B286352BEF3BF6">Erséus, Christer</mods:namePart>
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<mods:title id="66E43B1A824D2620440DF00B3AC8E175">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="2032061B3B3B818E68A975E7EAF9CB81">
<mods:date id="7457AF7E38247D0EA81E301D66952020">2004</mods:date>
<mods:detail id="86BD23C62AF43A42A53D5CEAD995C573" type="pubDate">
<mods:number id="5CE4D31A9C2751F1A508EB4C82F7BC64">2004-10-10</mods:number>
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<mods:detail id="209E177D700D61C330F8921DB07BC689" type="volume">
<mods:number id="CF4DB27314AAE15F45544BE447D886F1">38</mods:number>
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<mods:number id="4ACC2D220A93242AB9F432108C095CFF">3</mods:number>
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<mods:start id="9B9399CABEA2971AC21768FAF26E6723">269</mods:start>
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<mods:identifier id="D70B34C89CA81F8998C0734B526A4E08" type="CLB-Dataset">3888</mods:identifier>
<mods:identifier id="418B32668D5A00030D5969F0ED267DFC" type="DOI">10.1080/0022293021000028252</mods:identifier>
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<treatment id="704ECD36FFF32173FDEAEE97CE43DA2C" ID-DOI="http://doi.org/10.5281/zenodo.5597134" ID-GBIF-Taxon="189354021" ID-Zenodo-Dep="5597134" LSID="urn:lsid:plazi:treatment:704ECD36FFF32173FDEAEE97CE43DA2C" httpUri="http://treatment.plazi.org/id/704ECD36FFF32173FDEAEE97CE43DA2C" lastPageId="6" lastPageNumber="274" pageId="4" pageNumber="272">
<subSubSection id="B0FD2FABFFF32171FDEAEE97CF78DB8C" pageId="4" pageNumber="272" type="nomenclature">
<paragraph id="F8587C20FFF32171FDEAEE97CFF3DBAC" blockId="4.[621,960,181,205]" box="[621,960,181,205]" pageId="4" pageNumber="272">
<heading id="A310CB4CFFF32171FDEAEE97CFF3DBAC" box="[621,960,181,205]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="272" reason="2">
<taxonomicName id="3FE707A3FFF32171FDEAEE97CFF3DBAC" authority="Erseus, 1984" authorityName="Erseus" authorityYear="1984" box="[621,960,181,205]" class="Clitellata" family="Naididae" genus="Doliodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="272" phylum="Annelida" rank="species" species="tener">
<emphasis id="CA93A032FFF32171FDEAEE97CF1BDBAD" bold="true" box="[621,808,181,205]" italics="true" pageId="4" pageNumber="272">Doliodrilus tener</emphasis>
Erséus, 1984
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F8587C20FFF32171FD65EEF7CF78DB8C" blockId="4.[313,1268,213,295]" box="[738,843,213,237]" pageId="4" pageNumber="272">(figure 1)</paragraph>
</subSubSection>
<subSubSection id="B0FD2FABFFF32173FEBEEED5CE43DA2C" lastPageId="6" lastPageNumber="274" pageId="4" pageNumber="272" type="description">
<paragraph id="F8587C20FFF32171FEBEEED5C88BDA47" blockId="4.[313,1268,213,295]" pageId="4" pageNumber="272">
<emphasis id="CA93A032FFF32171FEBEEED5CDEDDA6D" box="[313,478,247,268]" italics="true" pageId="4" pageNumber="272">
<author id="205A94C5FFF32171FEBEEED5CD94DA6D" box="[313,423,247,268]" fns="*" inm="true" npo="*" pageId="4" pageNumber="272">Doliodrilus</author>
<title id="8AE39716FFF32171FE2BEED5CDEDDA6D" box="[428,478,247,268]" jopScore="0.008289838" pageId="4" pageNumber="272" type="_" typeScore="6">tener</title>
</emphasis>
Erséus, 1984: 159–160, figure 15; Erséus,
<year id="8C4F3D73FFF32171FC07EED5CF82DA6D" box="[896,945,247,268]" pageId="4" pageNumber="272">1990</year>
a (
<emphasis id="CA93A032FFF32171FC4BEED5C83DDA6D" box="[972,1038,247,268]" italics="true" pageId="4" pageNumber="272">partim</emphasis>
): 287–288, figure 11A, B, E–G, I, J; Erséus
<emphasis id="CA93A032FFF32171FDBCEF30CE54DA47" box="[571,615,273,295]" italics="true" pageId="4" pageNumber="272">et al</emphasis>
., 1990: 113, figure 2K;
<journalOrPublisher id="222676ECFFF32171FCDCEF33C821DA46" box="[859,1042,273,295]" jopScore="0.04681627" pageId="4" pageNumber="272">Erséus and Hsieh</journalOrPublisher>
,
<part id="8C43D6BDFFF32171FB9AEF33C863DA47" box="[1053,1104,273,294]" pageId="4" pageNumber="272" type="volume">1997</part>
:
<pagination id="C12E82D4FFF32171FBDBEF33C887DA47" box="[1116,1204,273,294]" pageId="4" pageNumber="272" type="pagination">100–101</pagination>
.
</paragraph>
<caption id="AC982CA8FFF32171FEBEE8B5C815DDA6" ID-DOI="http://doi.org/10.5281/zenodo.5258616" ID-Zenodo-Dep="5258616" httpUri="https://zenodo.org/record/5258616/files/figure.png" pageId="4" pageNumber="272" startId="4.[313,329,1687,1708]" targetBox="[320,1262,368,1659]" targetPageId="4">
<paragraph id="F8587C20FFF32171FEBEE8B5C815DDA6" blockId="4.[313,1268,1687,1736]" pageId="4" pageNumber="272">
F. 1.
<taxonomicName id="3FE707A3FFF32171FE12E8B5CEFADDCD" authority="Erseus, 1984" authorityName="Erseus" authorityYear="1984" box="[405,713,1687,1709]" class="Clitellata" family="Naididae" genus="Doliodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="272" phylum="Annelida" rank="species" species="tener">
<emphasis id="CA93A032FFF32171FE12E8B5CE0EDDCC" box="[405,573,1687,1709]" italics="true" pageId="4" pageNumber="272">Doliodrilus tener</emphasis>
Erséus, 1984
</taxonomicName>
. Male ducts of two specimens, with triangular (A) or hemispherical (B) prostatic projections, respectively.
</paragraph>
</caption>
<paragraph id="F8587C20FFF22170FEE6EE94C8E1DA4C" blockId="5.[313,1269,181,1736]" pageId="5" pageNumber="273">
<emphasis id="CA93A032FFF22170FEE6EE94CE36DBAC" box="[353,517,181,205]" italics="true" pageId="5" pageNumber="273">New material.</emphasis>
<materialsCitation id="488F767DFFF22170FD95EE94CF3BDBAC" ID-GBIF-Occurrence="3392780336" box="[530,776,181,205]" collectionCode="IHB" pageId="5" pageNumber="273" specimenCode="HANA2000003" specimenCount="1">
<collectionCode id="9EF6E4E5FFF22170FD95EE94CE77DBAC" box="[530,580,182,205]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="5" pageNumber="273">IHB</collectionCode>
<specimenCode id="A841D45BFFF22170FDD5EE97CF3BDBAC" box="[594,776,181,205]" collectionCode="HANA" pageId="5" pageNumber="273">HANA2000003</specimenCode>
</materialsCitation>
b–e,
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<specimenCode id="A841D45BFFF22170FCC5EE97CFC4DBAC" box="[834,1015,181,205]" collectionCode="HANA" pageId="5" pageNumber="273">HANA2000017</specimenCode>
</materialsCitation>
e,
<materialsCitation id="488F767DFFF22170FB93EE97C8F8DBAC" ID-GBIF-Occurrence="3392780302" box="[1044,1227,181,205]" collectionCode="HANA" pageId="5" pageNumber="273" specimenCode="HANA2000022" specimenCount="1">
<specimenCode id="A841D45BFFF22170FB93EE97C8F8DBAC" box="[1044,1227,181,205]" collectionCode="HANA" pageId="5" pageNumber="273">HANA2000022</specimenCode>
</materialsCitation>
a–e,
<materialsCitation id="488F767DFFF22170FEBEEEF7CDDEDB8C" ID-GBIF-Occurrence="3392780329" box="[313,493,213,237]" collectionCode="HANA" pageId="5" pageNumber="273" specimenCode="HANA2000029" specimenCount="1">
<specimenCode id="A841D45BFFF22170FEBEEEF7CDDEDB8C" box="[313,493,213,237]" collectionCode="HANA" pageId="5" pageNumber="273">HANA2000029</specimenCode>
</materialsCitation>
c–f,
<specimenCount id="EEE1B7A9FFF22170FD9BEEF7CE80DB8C" box="[540,691,213,237]" pageId="5" pageNumber="273" type="generic">14 specimens</specimenCount>
: four from SY00-2A, one from SY00-9C, five from Station HU00-14A, four from HU00-19A.
<materialsCitation id="488F767DFFF22170FCC7EED7C8FDDA4C" ID-GBIF-Occurrence="3392780328" collectionCode="SMNH" pageId="5" pageNumber="273" specimenCount="13">
<collectionCode id="9EF6E4E5FFF22170FCC7EED7CFA6DA6C" box="[832,917,245,269]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="5" pageNumber="273">SMNH</collectionCode>
Main Coll. 43665–43677,
<specimenCount id="EEE1B7A9FFF22170FB5DEED7CD9CDA4C" pageId="5" pageNumber="273" type="generic">13 specimens</specimenCount>
: three from SY00-2A, seven from HU00-14A, three from HU00-19A
</materialsCitation>
.
</paragraph>
<paragraph id="F8587C20FFF22170FEE6EF17C86ADF0B" blockId="5.[313,1269,181,1736]" pageId="5" pageNumber="273">
<emphasis id="CA93A032FFF22170FEE6EF17CEE3DA2C" box="[353,720,309,333]" italics="true" pageId="5" pageNumber="273">Brief description of new material.</emphasis>
Six complete specimens
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long, 28–45 segments. Diameter at XI about
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. Prostomium usually conical. Clitellum extending over XI–XII. Chaetae
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50–75
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</quantity>
m long, about
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2.5
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m thick, with upper teeth 1–1.5 times as long as, and thinner than or as thick as, lower; two to five per bundle anteriorly, one to three per bundle in post-clitellar segments. Chloragogen cells from VI onwards. Oesophagus in IX barrel-shaped, thick-walled and granulated, with or without chloragogen cells; semi-embedded blood plexus permeating dorsal region, with regular transverse vessels and less regular longitudinal ones. Vasa deferentia (figure 1A: vd) about
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135
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m long,
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12–19
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</quantity>
m wide. Atria totally
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200– 220
<emphasis id="CA93A032FFF22170FEE0EC76CD45D90A" bold="true" box="[359,374,596,619]" italics="true" pageId="5" pageNumber="273">m</emphasis>
</quantity>
m long,
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14–36
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</quantity>
m wide; ental end of atrial ampullae thin-walled and distinctly dilated; ventral projections on ampullae either (1) triangular (figure 1A), each with small (only
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10–17
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</quantity>
m long) prostatic pad (ppd) at bottom, and with numerous spindle-shaped nuclei (ncc) along both sides of pad (especially ental to pad), or (2) hemispherical (figure 1B), with large (
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29–34
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</quantity>
m long) prostatic pad (ppd), and with dense, but smaller patches of slender nuclei (ncc) around pad. Dense cluster of nuclei also present in dorsal wall of ampulla opposite to prostatic pad; these nuclei slender and regularly arranged when ventral projection triangular (figure 1A: nco), but oval and less regular when projection hemispherical (figure 1B: nco). Prostate glands (figure 1A, B: pr) usually medium to large, sometimes small, with small nuclei and large nucleus-like bodies, latter oblong to round, maximally
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12
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</quantity>
m long,
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11
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</quantity>
m wide. Atrial ducts (figure 1A, B: ad) each with (1) posterior blind sac (bs),
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25–50
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</quantity>
m long,
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15–30
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</quantity>
m wide, and (2) efferent duct (ed),
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25–35
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</quantity>
m long,
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19–27
<emphasis id="CA93A032FFF22170FB0EEDF1C8ABD88B" bold="true" box="[1161,1176,979,1002]" italics="true" pageId="5" pageNumber="273">m</emphasis>
</quantity>
m wide. Sperm sac in one to five segments within IX–XIII, when developed at all. Egg sac in one to three segments within XI–XIII, when developed. Spermathecal ducts
<quantity id="3F1FD1C5FFF22170FEBEEA13CDA0DF28" box="[313,403,1073,1098]" metricMagnitude="1" metricUnit="m" metricValue="7.0" metricValueMax="9.5" metricValueMin="4.5" pageId="5" pageNumber="273" unit="m" value="70.0" valueMax="95.0" valueMin="45.0">
45–95
<emphasis id="CA93A032FFF22170FE03EA10CDA0DF28" bold="true" box="[388,403,1074,1097]" italics="true" pageId="5" pageNumber="273">m</emphasis>
</quantity>
m long,
<quantity id="3F1FD1C5FFF22170FE7DEA10CE67DF28" box="[506,596,1074,1098]" metricMagnitude="1" metricUnit="m" metricValue="3.4" metricValueMax="4.4" metricValueMin="2.4" pageId="5" pageNumber="273" unit="m" value="34.0" valueMax="44.0" valueMin="24.0">
24–44
<emphasis id="CA93A032FFF22170FDC2EA10CE67DF28" bold="true" box="[581,596,1074,1097]" italics="true" pageId="5" pageNumber="273">m</emphasis>
</quantity>
m wide, with ectal vestibules; ampullae
<quantity id="3F1FD1C5FFF22170FBB5EA13C8AADF28" box="[1074,1177,1073,1098]" metricMagnitude="2" metricUnit="m" metricValue="1.3" metricValueMax="1.75" metricValueMin="0.85" pageId="5" pageNumber="273" unit="m" value="130.0" valueMax="175.0" valueMin="85.0">
85–175
<emphasis id="CA93A032FFF22170FB0DEA10C8AADF28" bold="true" box="[1162,1177,1074,1097]" italics="true" pageId="5" pageNumber="273">m</emphasis>
</quantity>
m long,
<quantity id="3F1FD1C5FFF22170FEBEEA73CDA0DF08" box="[313,403,1105,1130]" metricMagnitude="1" metricUnit="m" metricValue="6.25" metricValueMax="9.5" metricValueMin="3.0" pageId="5" pageNumber="273" unit="m" value="62.5" valueMax="95.0" valueMin="30.0">
30–95
<emphasis id="CA93A032FFF22170FE03EA70CDA0DF08" bold="true" box="[388,403,1106,1129]" italics="true" pageId="5" pageNumber="273">m</emphasis>
</quantity>
m wide, with sperm arranged in bundles or masses in lumina.
</paragraph>
<paragraph id="F8587C20FFF22170FEE6EA50CDE0DE48" blockId="5.[313,1269,181,1736]" pageId="5" pageNumber="273">
<emphasis id="CA93A032FFF22170FEE6EA50CDFFDFEB" box="[353,460,1138,1162]" italics="true" pageId="5" pageNumber="273">
<author id="205A94C5FFF22170FEE6EA50CDFBDFEB" box="[353,456,1138,1162]" fns="*" inm="true" npo="*" pageId="5" pageNumber="273">Remarks</author>
.
</emphasis>
<title id="8AE39716FFF22170FE54EA50C8C3DFEB" box="[467,1264,1138,1162]" jopScore="0.07906442" pageId="5" pageNumber="273">This species was originally described from Hong Kong (Erséus, 1984)</title>
. In 1990,
<volumeTitle id="474B1E10FFF22170FE21EAB0C853DE68" jopScore="0.30799273" pageId="5" pageNumber="273">
Erséus reported it from Hong Kong again, noting some morphological variation, but a form with unusually long upper teeth on the chaetae is now regarded as a separate taxon (see
<emphasis id="CA93A032FFF22170FDD0EAF0CF23DF88" box="[599,784,1233,1257]" italics="true" pageId="5" pageNumber="273">D. longidentatus</emphasis>
sp. n. below). The other known distributional areas of
<emphasis id="CA93A032FFF22170FE6EEAD0CE75DE68" box="[489,582,1266,1289]" italics="true" pageId="5" pageNumber="273">D. tener</emphasis>
are Qingdao (Erséus
<emphasis id="CA93A032FFF22170FCC1EAD0CF4BDE68" box="[838,888,1265,1289]" italics="true" pageId="5" pageNumber="273">et al</emphasis>
., 1990) and Taiwan
</volumeTitle>
(
<journalOrPublisher id="222676ECFFF22170FBF1EAD3C8C6DE68" box="[1142,1269,1265,1289]" jopScore="0.070280224" pageId="5" pageNumber="273">Erséus and</journalOrPublisher>
<editor id="3E7B9423FFF22170FEBEEB33CD4EDE48" box="[313,381,1297,1321]" fns="*" inm="true" npo="*" pageId="5" pageNumber="273">Hsieh</editor>
,
<year id="8C4F3D73FFF22170FE0CEB33CDF4DE48" box="[395,455,1297,1321]" pageId="5" pageNumber="273">1997</year>
).
</paragraph>
<paragraph id="F8587C20FFF22170FEE6EB13CDF5DDE9" blockId="5.[313,1269,181,1736]" pageId="5" pageNumber="273">
<title id="8AE39716FFF22170FEE6EB13CF5ADEA8" jopScore="0.29281205" pageId="5" pageNumber="273">
The most prominent character of
<emphasis id="CA93A032FFF22170FD62EB10CF72DE28" box="[741,833,1330,1353]" italics="true" pageId="5" pageNumber="273">D. tener</emphasis>
, not specifically noted in the previous descriptions, is the conspicuous thickening of the dorsal wall opposite to the prostatic pad in the atrium, and the clusters of nuclei in this wall and around the pad. We observed this character in all the material from Hainan, as well as in numerous old specimens from Hong Kong [treated by Erséus
</title>
, 1984,
<year id="8C4F3D73FFF22170FC47EB93CFC5DEA8" box="[960,1014,1457,1481]" pageId="5" pageNumber="273">1990</year>
a (
<emphasis id="CA93A032FFF22170FB9DEB93C850DEA9" box="[1050,1123,1457,1480]" italics="true" pageId="5" pageNumber="273">partim</emphasis>
)].
<author id="205A94C5FFF22170FB00EB93C8C3DEA9" box="[1159,1264,1457,1480]" fns="*" inm="true" npo="*" pageId="5" pageNumber="273">However</author>
,
<journalOrPublisher id="222676ECFFF22170FEBEEBF2CDC0DD69" jopScore="0.06503752" pageId="5" pageNumber="273" type="_" typeScore="4">there are two different appearances of the ventral projections on the atria; they are either triangular</journalOrPublisher>
(figure
<part id="8C43D6BDFFF22170FDCCEBD2CE66DD69" box="[587,597,1520,1544]" pageId="5" pageNumber="273" type="volume">1</part>
A) or hemispherical (figure
<pagination id="C12E82D4FFF22170FC13EBD2CFADDD69" box="[916,926,1520,1544]" pageId="5" pageNumber="273" type="pagination">1</pagination>
B) (see above description). In the former state, the small prostatic pads seem to have discharged secretion into the atria, and the nuclei around the pads are widely distributed. In the latter state, the large pads appear to be full of secretion, and the nuclei are more restricted in distribution.
</paragraph>
<paragraph id="F8587C20FFF22173FEE6E8B2CF11DB8C" blockId="5.[313,1269,181,1736]" lastBlockId="6.[313,1269,181,333]" lastPageId="6" lastPageNumber="274" pageId="5" pageNumber="273">
<author id="205A94C5FFF22170FEE6E8B2CDBEDDC9" box="[353,397,1680,1704]" fns="*" inm="true" npo="*" pageId="5" pageNumber="273">The</author>
<title id="8AE39716FFF22170FE12E8B2C8BDDDC9" box="[405,1166,1680,1704]" jopScore="0.06767113" pageId="5" pageNumber="273">prostate glands were stated to be small in the Hong Kong material</title>
(
<journalOrPublisher id="222676ECFFF22170FB25E8B2C8C3DDC9" box="[1186,1264,1680,1704]" jopScore="0.0048053036" pageId="5" pageNumber="273" type="_" typeScore="6">Erséus</journalOrPublisher>
,
<year id="8C4F3D73FFF22170FEBEE892CD42DDA9" box="[313,369,1712,1736]" pageId="5" pageNumber="273">1984</year>
, 1990a), but a re-examination shows that the glands in the specimens from Hong Kong and Taiwan (Erséus and Hsieh, 1997), as well as in the new material, vary considerably, i.e. from small to large.
</paragraph>
<paragraph id="F8587C20FFF12173FEE6EED7CE43DA2C" blockId="6.[313,1269,181,333]" pageId="6" pageNumber="274">
<emphasis id="CA93A032FFF12173FEE6EED7CE45DA6C" box="[353,630,245,269]" italics="true" pageId="6" pageNumber="274">Distribution and habitat.</emphasis>
Known only from
<collectingCountry id="80F03CB0FFF12173FCDFEED7CFAEDA6C" box="[856,925,245,269]" name="China" pageId="6" pageNumber="274">China</collectingCountry>
[
<collectingRegion id="3A23B2C2FFF12173FC37EED7C836DA6C" box="[944,1029,245,269]" country="China" name="Hainan" pageId="6" pageNumber="274">Hainan</collectingRegion>
(new record),
<collectingCountry id="80F03CB0FFF12173FB33EED7CD4FDA4C" name="Hong Kong" pageId="6" pageNumber="274">Hong Kong</collectingCountry>
,
<collectingCountry id="80F03CB0FFF12173FE0EEF37CE17DA4C" box="[393,548,277,301]" name="China" pageId="6" pageNumber="274">Jiaozhou Bay</collectingCountry>
(at Qingdao),
<collectingCountry id="80F03CB0FFF12173FD57EF37CF1ADA4C" box="[720,809,277,301]" name="Taiwan" pageId="6" pageNumber="274">Taiwan</collectingCountry>
]. Brackish-water, intertidal and subtidal soft mud and muddy sand.
</paragraph>
</subSubSection>
</treatment>
</document>
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