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<document id="91B61158DBA5AF8509A81670EBDDC8BF" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A136D5312FDA6FAC9FB88FBBC" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus arauzae Moret 2020, sp. nov." docType="treatment" docUuid="B992F8BA-69F8-4591-A010-440432759483" docUuidSource="ZooBank" docVersion="8" lastPageNumber="40" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="38" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="8A1DF408801AA1B47D8F293CC4DCF30B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4F004AED2A1DD16ACE5DAE15E3FAED55">
<mods:title id="6E290A8922C34A7B602500FAD3D14FB2">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
</mods:titleInfo>
<mods:name id="329CFAC25D4C238CAC30B4010572D112" type="personal">
<mods:role id="8F8E26D4929CB4DCAEE5636D01EE8070">
<mods:roleTerm id="3F1AEA9CE073895DD84F49521A3EAE96">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A33B4A21C70179B12C5F07933FD71F31">Moret, Pierre</mods:namePart>
<mods:nameIdentifier id="7EDA563E58A800E6FB5B5971FD487C17" type="LSID">2E194645-D255-438B-819E-A2B6C39AD699</mods:nameIdentifier>
<mods:affiliation id="CBE8FFC5510E42FE7E2B9FFF3BB4B1FD">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="CAEAE08E91DA6D8BAD5487229F12D209" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="E765304234AA1304DCF252FFC87B4FF5" type="personal">
<mods:role id="340C50D5FB2EE19402F5BFFABAEDF8B7">
<mods:roleTerm id="10A21CFD676F9E841066C1A2F456C9D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02F569262FAD9F7F718DD73789C5E5F7">Murienne, Jérôme</mods:namePart>
<mods:nameIdentifier id="EB62F599126213569C2F1EA2BB163F16" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
<mods:affiliation id="84C066EF48474C0E8E42C900FF8033C9">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
<mods:nameIdentifier id="8EADB7F7F37DE43C42CC163481C1484B" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1894FEA7F4280A226DCD87035FF3590F">text</mods:typeOfResource>
<mods:relatedItem id="22EB4BD27C1872F8436987BCC8292065" type="host">
<mods:titleInfo id="337EE39E94667954A9AC6F793869AD3E">
<mods:title id="88BF0B0C8913B1ACF7A85263BDA4752C">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="4F7F2CFDD7FFD0524FC1CF950AC00016">
<mods:date id="9D852793D1E71478B660B591A5D0C860">2020</mods:date>
<mods:detail id="FAD128D895298C04416D124B3716CE5C" type="pubDate">
<mods:number id="F29B9FF46B806C5F2049064E3A6A8A57">2020-05-15</mods:number>
</mods:detail>
<mods:detail id="7C586E7ADCFC6C71C71A7C82F1115C9B" type="volume">
<mods:number id="0C47B387FBEC84743BA9BA8477405720">646</mods:number>
</mods:detail>
<mods:extent id="C25340BD866CB8695D3897AD4DB440E7" unit="page">
<mods:start id="7D5F074B7BE39A3E03E80B287D30FF22">1</mods:start>
<mods:end id="9033919BAD9700F6FFB5ABC0B6C924A5">55</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="74A4EC59A31E0A6F521DAE41793F444E">journal article</mods:classification>
<mods:identifier id="20B537D6E468DD8C78E2E98F50E5C758" type="CLB-Dataset">22092</mods:identifier>
<mods:identifier id="90CA790C5C86CAD9439469DABCC474C8" type="DOI">10.5852/ejt.2020.646</mods:identifier>
<mods:identifier id="B12227C9118E5A0178CC268D20FB8B6C" type="GBIF-Dataset">c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b</mods:identifier>
<mods:identifier id="103965B9DF2F8AA1833E80D773A14C17" type="Zenodo-Dep">3829682</mods:identifier>
<mods:identifier id="AACBC05B3AC90F497A0F4AF5FE82160E" type="ZooBank">4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B</mods:identifier>
</mods:mods>
<treatment id="03B7726A136D5312FDA6FAC9FB88FBBC" ID-DOI="http://doi.org/10.5281/zenodo.3848383" ID-GBIF-Taxon="164160392" ID-Zenodo-Dep="3848383" LSID="urn:lsid:zoobank.org:act:B992F8BA-69F8-4591-A010-440432759483" httpUri="http://treatment.plazi.org/id/03B7726A136D5312FDA6FAC9FB88FBBC" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
<subSubSection id="C30490F7136D5310FDA6FAC9FB3EFAF2" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8BA1C37C136D5310FDA6FAC9FC5BFACE" blockId="37.[380,1208,1331,1427]" box="[599,989,1331,1358]" pageId="37" pageNumber="38">
<heading id="D0E97410136D5310FDA6FAC9FC5BFACE" box="[599,989,1331,1358]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="38" reason="2">
<taxonomicName id="4C1EB8FF136D5310FDA6FAC9FCFDFACE" ID-CoL="8MMR9" authority="Moret" authorityName="Moret" authorityYear="2020" box="[599,891,1331,1358]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="arauzae" status="sp. nov.">
<emphasis id="B96A1F6E136D5310FDA6FAC9FCAAFACE" bold="true" box="[599,812,1331,1358]" italics="true" pageId="37" pageNumber="38">Dyscolus arauzae</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136D5310FC73FAC9FC5BFACE" box="[898,989,1331,1357]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BA1C37C136D5310FE8DFAACFB3EFAF2" blockId="37.[380,1208,1331,1427]" box="[380,1208,1366,1393]" pageId="37" pageNumber="38">
<uri id="FF8FCF7E136D5310FE8DFAACFB3EFAF2" box="[380,1208,1366,1393]" pageId="37" pageNumber="38">
urn:lsid:zoobank.org:act:
<uuid id="FFB8F9A9136D5310FD69FAACFB3EFAF2" box="[664,1208,1366,1393]" pageId="37" pageNumber="38">B992F8BA-69F8-4591-A010-440432759483</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30490F7136D5310FD29FA83FCDDFA10" box="[728,859,1401,1427]" pageId="37" pageNumber="38" type="description">
<paragraph id="8BA1C37C136D5310FD29FA83FCDDFA10" blockId="37.[380,1208,1331,1427]" box="[728,859,1401,1427]" pageId="37" pageNumber="38">
<figureCitation id="1325DFF9136D5310FD29FA83FCDDFA10" box="[728,859,1401,1427]" captionStart="Figs 36–41" captionStartId="38.[189,241,1854,1880]" captionTargetBox="[288,1297,277,1813]" captionTargetId="figure@38.[278,1309,265,1818]" captionTargetPageId="38" captionText="Figs 36–41. Dyscolus spp., habitus (36–38) and aedeagus, median lobe in lateral view (39–40) and apex of the median lobe (41). 36. Dyscolus arauzae Moret sp. nov., male holotype (QCAZ). 37. D. piscator Moret sp. nov., male holotype (QACZ). 38. D. placitus Moret sp. nov., male holotype (MNHN). 39. D. arauzae Moret sp. nov. 40. D. piscator Moret sp. nov. 41. D. placitus Moret sp. nov. (above) and D. piscator Moret sp. nov. (below)." figureDoi="http://doi.org/10.5281/zenodo.3829702" httpUri="https://zenodo.org/record/3829702/files/figure.png" pageId="37" pageNumber="38">Figs 36, 39</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30490F7136D5310FF4CFA41FECDF9A5" pageId="37" pageNumber="38" type="etymology">
<paragraph id="8BA1C37C136D5310FF4CFA41FEC2FA56" blockId="37.[189,324,1467,1493]" box="[189,324,1467,1493]" pageId="37" pageNumber="38">
<heading id="D0E97410136D5310FF4CFA41FEC2FA56" bold="true" box="[189,324,1467,1493]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B96A1F6E136D5310FF4CFA41FEC2FA56" bold="true" box="[189,324,1467,1493]" pageId="37" pageNumber="38">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C136D5310FF4CFA10FECDF9A5" blockId="37.[189,1398,1512,1574]" pageId="37" pageNumber="38">Noun in the genitive case, dedicated to María de los Ángeles Aráuz who took part in the discovery of this species.</paragraph>
</subSubSection>
<subSubSection id="C30490F7136D5312FF4CF9B4FB88FBBC" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="8BA1C37C136D5310FF4CF9B4FEEAF9EB" blockId="37.[189,364,1614,1640]" box="[189,364,1614,1640]" pageId="37" pageNumber="38">
<heading id="D0E97410136D5310FF4CF9B4FEEAF9EB" bold="true" box="[189,364,1614,1640]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B96A1F6E136D5310FF4CF9B4FEEAF9EB" bold="true" box="[189,364,1614,1640]" pageId="37" pageNumber="38">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C136D5310FF1BF987FEDAF914" blockId="37.[189,1399,1661,1758]" box="[234,348,1661,1687]" pageId="37" pageNumber="38">
<emphasis id="B96A1F6E136D5310FF1BF987FEDAF914" bold="true" box="[234,348,1661,1687]" pageId="37" pageNumber="38">
<typeStatus id="54A57DDE136D5310FF1BF987FEDAF914" box="[234,348,1661,1687]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA1C37C136D5310FF4CF95AFC36F95E" blockId="37.[189,1399,1661,1758]" pageId="37" pageNumber="38">
<materialsCitation id="3B76C921136D5310FF4CF95AFC36F95E" ID-GBIF-Occurrence="2620303352" collectingDate="2015-10-30" collectionCode="QCAZ" collectorName="Volcan Cayambe SW &amp; P. Moret &amp; M. Arauz" country="Ecuador" elevation="4405" latitude="0.006962" location="Waypoint 73" longLatPrecision="1" longitude="-78.02028" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Pichincha Province" typeStatus="holotype">
<collectingCountry id="F30983EC136D5310FF4CF95AFECDF939" box="[189,331,1696,1722]" name="Ecuador" pageId="37" pageNumber="38">ECUADOR</collectingCountry>
•
<specimenCount id="9D1808F5136D5310FE92F964FEFCF939" box="[355,378,1694,1722]" pageId="37" pageNumber="38" type="male">♂</specimenCount>
;
<collectingRegion id="49DA0D9E136D5310FE78F95AFDE9F939" box="[393,623,1696,1722]" country="Ecuador" name="Pichincha" pageId="37" pageNumber="38">Pichincha Province</collectingRegion>
,
<collectorName id="26EBA6AA136D5310FD8BF95AFCF2F939" box="[634,884,1696,1722]" pageId="37" pageNumber="38">Volcán Cayambe SW</collectorName>
,
<location id="8EC195A7136D5310FC71F95AFB9FF939" LSID="urn:lsid:plazi:treatment:03B7726A136D5312FDA6FAC9FB88FBBC:8EC195A7136D5310FC71F95AFB9FF939" box="[896,1049,1696,1722]" country="Ecuador" latitude="0.006962" longLatPrecision="1" longitude="-78.02028" name="Waypoint 73" pageId="37" pageNumber="38" stateProvince="Pichincha Province">Waypoint 73</location>
;
<geoCoordinate id="EE2AA5BB136D5310FBD4F95AFB3DF939" box="[1061,1211,1696,1722]" degrees="0.006962" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="1" value="0.006962">0.006962º N</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB136D5310FB36F95AFAF4F939" box="[1223,1394,1696,1722]" degrees="78.020278" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="1" value="-78.02028">78.020278º W</geoCoordinate>
;
<elevation id="0033244F136D5310FF4CF939FED0F95E" box="[189,342,1731,1758]" metricMagnitude="3" metricUnit="m" metricValue="4.404999999999999" pageId="37" pageNumber="38" unit="m" value="4405.0">
<quantity id="4CE66E99136D5310FF4CF939FE90F95E" box="[189,278,1731,1758]" metricMagnitude="3" metricUnit="m" metricValue="4.404999999999999" pageId="37" pageNumber="38" unit="m" value="4405.0">4405 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC136D5310FE92F939FE79F95D" box="[355,511,1731,1758]" pageId="37" pageNumber="38" value="2015-10-30">
<collectingDate id="EFE41C54136D5310FE92F939FE79F95D" box="[355,511,1731,1758]" pageId="37" pageNumber="38" value="2015-10-30">30 Oct. 2015</collectingDate>
</date>
;
<collectorName id="26EBA6AA136D5310FDFAF939FDE9F95E" box="[523,623,1731,1757]" pageId="37" pageNumber="38">P. Moret</collectorName>
and
<collectorName id="26EBA6AA136D5310FD56F939FC91F95E" box="[679,791,1731,1757]" pageId="37" pageNumber="38">M. Aráuz</collectorName>
leg.;
<collectionCode id="ED0F5BB9136D5310FCA9F939FC2AF95E" box="[856,940,1731,1757]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="37" pageNumber="38" type="Museum">QCAZ</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA1C37C136D5310FF1BF8FFFD90F89C" blockId="37.[189,1399,1795,1999]" box="[234,534,1795,1823]" pageId="37" pageNumber="38">
<emphasis id="B96A1F6E136D5310FF1BF8FFFEE1F89C" bold="true" box="[234,359,1797,1823]" pageId="37" pageNumber="38">
<typeStatus id="54A57DDE136D5310FF1BF8FFFEE1F89C" box="[234,359,1797,1823]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
</emphasis>
(6 ♂♂,
<specimenCount id="9D1808F5136D5310FE38F8FEFD8FF89C" box="[457,521,1796,1823]" pageId="37" pageNumber="38" type="female">6 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BA1C37C136D5312FF4CF8D2FE6FFE0C" blockId="37.[189,1399,1795,1999]" lastBlockId="39.[189,1399,265,399]" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
<materialsCitation id="3B76C921136D5310FF4CF8D2FB67F8C1" ID-GBIF-Occurrence="2620303320" box="[189,1249,1830,1858]" collectingDate="2015-10-30" collectionCode="MNHN" collectorName="Volcan Cayambe SW &amp; P. Moret &amp; M. Arauz" country="Ecuador" elevation="4405" latitude="0.006962" location="Pichincha Province" longLatPrecision="1" longitude="-78.02028" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha Province">
<collectingCountry id="F30983EC136D5310FF4CF8D2FECDF8C1" box="[189,331,1832,1858]" name="Ecuador" pageId="37" pageNumber="38">ECUADOR</collectingCountry>
–
<emphasis id="B96A1F6E136D5310FE9CF8DDFDE7F8C2" bold="true" box="[365,609,1831,1857]" pageId="37" pageNumber="38">
<collectingRegion id="49DA0D9E136D5310FE9CF8DDFDE7F8C2" box="[365,609,1831,1857]" country="Ecuador" name="Pichincha" pageId="37" pageNumber="38">Pichincha Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D1808F5136D5310FD71F8D2FD36F8C1" box="[640,688,1830,1858]" pageId="37" pageNumber="38" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB9136D5310FB76F8D2FB67F8C1" box="[1159,1249,1832,1858]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="37" pageNumber="38">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921136D5310FB0EF8D2FD3BF8E6" ID-GBIF-Occurrence="2620303369" collectingDate="2015-10-30" collectionCode="CPM" collectorName="Volcan Cayambe SW &amp; P. Moret &amp; M. Arauz" country="Ecuador" elevation="4405" latitude="0.006962" location="Waypoint" longLatPrecision="1" longitude="-78.02028" pageId="37" pageNumber="38" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Pichincha">
<specimenCount id="9D1808F5136D5310FB0EF8D2FAA8F8C1" box="[1279,1326,1830,1858]" pageId="37" pageNumber="38" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1808F5136D5310FAB1F8D2FAE9F8C1" box="[1344,1391,1832,1858]" pageId="37" pageNumber="38" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB9136D5310FD8EF8B1FD3BF8E6" box="[639,701,1867,1893]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="37" pageNumber="38">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921136D5310FD2BF8B1FD6BF80B" ID-GBIF-Occurrence="2620303410" collectingDate="2015-10-30" collectionCode="CPM" collectorName="Volcan Cayambe SW &amp; P. Moret &amp; M. Arauz" country="Ecuador" elevation="4405" latitude="0.006962" location="Waypoint" longLatPrecision="1" longitude="-78.02028" pageId="37" pageNumber="38" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha">
<specimenCount id="9D1808F5136D5310FD2BF8B1FC8FF8E6" box="[730,777,1867,1893]" pageId="37" pageNumber="38" type="female">1 ♀</specimenCount>
; same collection data as for holotype; COI voucher PM073-03, BOLD sequence SUM020-18;
<collectionCode id="ED0F5BB9136D5310FD5EF894FD6BF80B" box="[687,749,1902,1928]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="37" pageNumber="38">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921136D5310FCF3F894FDEEF84D" ID-GBIF-Occurrence="2620303356" collectingDate="2015-10-30" collectionCode="CPM" collectorName="Volcan Cayambe SW &amp; P. Moret &amp; M. Arauz" country="Ecuador" elevation="4300" latitude="0.007724" location="Waypoint 77" longLatPrecision="1" longitude="-78.02335" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha">
<specimenCount id="9D1808F5136D5310FCF3F894FCA8F80B" box="[770,814,1900,1928]" pageId="37" pageNumber="38" type="male">1 ♂</specimenCount>
;
<collectorName id="26EBA6AA136D5310FCCBF894FBB7F80B" box="[826,1073,1902,1928]" pageId="37" pageNumber="38">Volcán Cayambe SW</collectorName>
,
<location id="8EC195A7136D5310FBCAF894FB55F80A" LSID="urn:lsid:plazi:treatment:03B7726A136D5312FDA6FAC9FB88FBBC:8EC195A7136D5310FBCAF894FB55F80A" box="[1083,1235,1902,1929]" country="Ecuador" latitude="0.007724" longLatPrecision="1" longitude="-78.02335" name="Waypoint 77" pageId="37" pageNumber="38" stateProvince="Pichincha">Waypoint 77</location>
;
<geoCoordinate id="EE2AA5BB136D5310FB2CF894FAF4F80A" box="[1245,1394,1902,1929]" degrees="0.007724" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="1" value="0.007724">0.007724º N</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB136D5310FF4CF86BFEEAF82F" box="[189,364,1937,1964]" degrees="78.023350" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="1" value="-78.02335">78.023350º W</geoCoordinate>
;
<elevation id="0033244F136D5310FE8DF86BFD9BF828" box="[380,541,1937,1964]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="37" pageNumber="38" unit="m" value="4300.0">
<quantity id="4CE66E99136D5310FE8DF86BFE5FF82F" box="[380,473,1937,1964]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="37" pageNumber="38" unit="m" value="4300.0">4300 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC136D5310FDDEF86BFD55F82F" box="[559,723,1937,1964]" pageId="37" pageNumber="38" value="2015-10-30">
<collectingDate id="EFE41C54136D5310FDDEF86BFD55F82F" box="[559,723,1937,1964]" pageId="37" pageNumber="38" value="2015-10-30">30 Oct. 2015</collectingDate>
</date>
;
<collectorName id="26EBA6AA136D5310FD12F868FCCDF82F" box="[739,843,1938,1964]" pageId="37" pageNumber="38">P. Moret</collectorName>
and
<collectorName id="26EBA6AA136D5310FC7DF868FB86F828" box="[908,1024,1937,1964]" pageId="37" pageNumber="38">M. Aráuz</collectorName>
leg.; COI voucher PM077-01, BOLD sequence SUM099-18;
<collectionCode id="ED0F5BB9136D5310FDDBF84EFDEEF84D" box="[554,616,1972,1998]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="37" pageNumber="38">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921136D5312FD73F84EFBBBFEA6" ID-GBIF-Occurrence="2620303402" collectingDate="2017-03-04" collectionCode="CPM" collectorName="Volcan Cayambe SW &amp; P. Moret &amp; M. Gobbi" country="Ecuador" elevation="4294" lastPageId="39" lastPageNumber="40" latitude="0.0069166664" location="Waypoint 267" longLatPrecision="1" longitude="-78.0223" pageId="37" pageNumber="38" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Pichincha">
<specimenCount id="9D1808F5136D5310FD73F84EFD43F84C" box="[642,709,1971,1999]" pageId="37" pageNumber="38" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D1808F5136D5310FD22F84EFC91F84C" box="[723,791,1972,1999]" pageId="37" pageNumber="38" type="female">2 ♀♀</specimenCount>
;
<collectorName id="26EBA6AA136D5310FCD4F84EFBA6F84D" box="[805,1056,1972,1998]" pageId="37" pageNumber="38">Volcán Cayambe SW</collectorName>
,
<location id="8EC195A7136D5310FBDDF84EFB52F84D" LSID="urn:lsid:plazi:treatment:03B7726A136D5312FDA6FAC9FB88FBBC:8EC195A7136D5310FBDDF84EFB52F84D" box="[1068,1236,1972,1998]" country="Ecuador" latitude="0.0069166664" longLatPrecision="1" longitude="-78.0223" name="Waypoint 267" pageId="37" pageNumber="38" stateProvince="Pichincha">Waypoint 267</location>
;
<geoCoordinate id="EE2AA5BB136D5310FB10F849FAF4F84C" box="[1249,1394,1971,1999]" degrees="0" direction="north" minutes="00.415" orientation="latitude" pageId="37" pageNumber="38" precision="1" value="0.0069166664">0°00.415′ N</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB136F5312FF4CFEF3FED2FEA5" box="[189,340,265,294]" degrees="78" direction="west" minutes="1.338" orientation="longitude" pageId="39" pageNumber="40" precision="1" value="-78.0223">78°1.338′ W</geoCoordinate>
;
<elevation id="0033244F136F5312FE90FEF1FE7DFEA6" box="[353,507,267,294]" metricMagnitude="3" metricUnit="m" metricValue="4.294" pageId="39" pageNumber="40" unit="m" value="4294.0">
<quantity id="4CE66E99136F5312FE90FEF1FE3DFEA5" box="[353,443,267,294]" metricMagnitude="3" metricUnit="m" metricValue="4.294" pageId="39" pageNumber="40" unit="m" value="4294.0">4294 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC136F5312FDFBFEF1FD1BFEA6" box="[522,669,267,294]" pageId="39" pageNumber="40" value="2017-03-04">
<collectingDate id="EFE41C54136F5312FDFBFEF1FD1BFEA6" box="[522,669,267,294]" pageId="39" pageNumber="40" value="2017-03-04">4 Mar. 2017</collectingDate>
</date>
;
<collectorName id="26EBA6AA136F5312FD5BFEF6FC89FEA5" box="[682,783,268,294]" pageId="39" pageNumber="40">P. Moret</collectorName>
and
<collectorName id="26EBA6AA136F5312FCB8FEF6FC3AFEA6" box="[841,956,267,294]" pageId="39" pageNumber="40">M. Gobbi</collectorName>
leg.;
<collectionCode id="ED0F5BB9136F5312FC0EFEF1FBBBFEA6" box="[1023,1085,267,293]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="39" pageNumber="40">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921136F5312FBA9FEF1FE38FEEF" ID-GBIF-Occurrence="2620303376" collectingDate="2016-07-18" collectionCode="CPM" collectorName="Volcan Cayambe SW &amp; P. Moret &amp; E. Moreno" country="Ecuador" elevation="4390" latitude="0.00686" location="Waypoint 76" longLatPrecision="1" longitude="-78.02071" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha">
<specimenCount id="9D1808F5136F5312FBA9FEF1FB03FEA5" box="[1112,1157,267,294]" pageId="39" pageNumber="40" type="female">1 ♀</specimenCount>
(in ethanol);
<collectorName id="26EBA6AA136F5312FAD7FEF1FEE4FECA" pageId="39" pageNumber="40">Volcán Cayambe SW</collectorName>
,
<location id="8EC195A7136F5312FE80FED4FD88FECB" LSID="urn:lsid:plazi:treatment:03B7726A136D5312FDA6FAC9FB88FBBC:8EC195A7136F5312FE80FED4FD88FECB" box="[369,526,302,328]" country="Ecuador" latitude="0.00686" longLatPrecision="1" longitude="-78.02071" name="Waypoint 76" pageId="39" pageNumber="40" stateProvince="Pichincha">Waypoint 76</location>
;
<geoCoordinate id="EE2AA5BB136F5312FDECFED4FD30FECA" box="[541,694,302,329]" degrees="0.006860" direction="north" orientation="latitude" pageId="39" pageNumber="40" precision="1" value="0.00686">0.006860º N</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB136F5312FD34FED5FCF5FECA" box="[709,883,303,329]" degrees="78.020714" direction="west" orientation="longitude" pageId="39" pageNumber="40" precision="1" value="-78.02071">78.020714º W</geoCoordinate>
;
<elevation id="0033244F136F5312FC73FED5FBA7FECB" box="[898,1057,302,329]" metricMagnitude="3" metricUnit="m" metricValue="4.39" pageId="39" pageNumber="40" unit="m" value="4390.0">
<quantity id="4CE66E99136F5312FC73FED5FC58FECA" box="[898,990,303,329]" metricMagnitude="3" metricUnit="m" metricValue="4.39" pageId="39" pageNumber="40" unit="m" value="4390.0">4390 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC136F5312FBC3FED5FB4AFECB" box="[1074,1228,302,329]" pageId="39" pageNumber="40" value="2016-07-18">
<collectingDate id="EFE41C54136F5312FBC3FED5FB4AFECB" box="[1074,1228,302,329]" pageId="39" pageNumber="40" value="2016-07-18">18 Jul. 2016</collectingDate>
</date>
;
<collectorName id="26EBA6AA136F5312FB2AFED5FAC4FECA" box="[1243,1346,303,329]" pageId="39" pageNumber="40">P. Moret</collectorName>
and
<collectorName id="26EBA6AA136F5312FF4CFEA8FEBAFEEF" box="[189,316,338,364]" pageId="39" pageNumber="40">E. Moreno</collectorName>
leg.;
<collectionCode id="ED0F5BB9136F5312FE71FEA8FE38FEEF" box="[384,446,338,364]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="39" pageNumber="40">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921136F5312FE29FEA8FE63FE0C" ID-GBIF-Occurrence="2620303425" collectingDate="2017-03-04" collectionCode="CPM" collectorName="P. Moret &amp; M. Gobbi" country="Ecuador" elevation="4390" latitude="0.00686" location="Waypoint" longLatPrecision="1" longitude="-78.02071" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha">
<specimenCount id="9D1808F5136F5312FE29FEA8FD83FEEF" box="[472,517,338,364]" pageId="39" pageNumber="40" type="female">1 ♀</specimenCount>
(in ethanol); same collection data as for preceding;
<date id="FFA0E5BC136F5312FB80FEA8FA83FEEF" box="[1137,1285,338,364]" pageId="39" pageNumber="40" value="2017-03-04">
<collectingDate id="EFE41C54136F5312FB80FEA8FA83FEEF" box="[1137,1285,338,364]" pageId="39" pageNumber="40" value="2017-03-04">4 Mar. 2017</collectingDate>
</date>
;
<collectorName id="26EBA6AA136F5312FAE3FEA8FAF1FEEF" box="[1298,1399,338,364]" pageId="39" pageNumber="40">P. Moret</collectorName>
and
<collectorName id="26EBA6AA136F5312FF1EFE8FFEE6FE0C" box="[239,352,373,399]" pageId="39" pageNumber="40">M. Gobbi</collectorName>
leg.;
<collectionCode id="ED0F5BB9136F5312FE52FE8FFE63FE0C" box="[419,485,373,399]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="39" pageNumber="40">CPM</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="DF6193F4136E5313FF4CF8C4FDDAF865" ID-DOI="http://doi.org/10.5281/zenodo.3829702" ID-Zenodo-Dep="3829702" httpUri="https://zenodo.org/record/3829702/files/figure.png" pageId="38" pageNumber="39" startId="38.[189,241,1854,1880]" targetBox="[288,1297,277,1813]" targetPageId="38">
<paragraph id="8BA1C37C136E5313FF4CF8C4FDDAF865" blockId="38.[189,1399,1854,2022]" pageId="38" pageNumber="39">
<emphasis id="B96A1F6E136E5313FF4CF8C4FECEF8DB" bold="true" box="[189,328,1854,1880]" pageId="38" pageNumber="39">Figs 36–41.</emphasis>
<taxonomicName id="4C1EB8FF136E5313FEBEF8C4FE3DF8DB" authorityName="Dejean" authorityYear="1831" box="[335,443,1854,1880]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B96A1F6E136E5313FEBEF8C4FE3DF8DB" box="[335,443,1854,1880]" italics="true" pageId="38" pageNumber="39">Dyscolus</emphasis>
</taxonomicName>
spp., habitus (36–38) and aedeagus, median lobe in lateral view (39–40) and apex of the median lobe (41).
<emphasis id="B96A1F6E136E5313FE13F89BFE79F8F8" bold="true" box="[482,511,1889,1915]" pageId="38" pageNumber="39">36</emphasis>
.
<taxonomicName id="4C1EB8FF136E5313FDFEF89BFCB7F8FF" authority="Moret" authorityName="Moret" authorityYear="2020" box="[527,817,1889,1916]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="arauzae" status="sp. nov.">
<emphasis id="B96A1F6E136E5313FDFEF89BFD67F8F8" box="[527,737,1889,1915]" italics="true" pageId="38" pageNumber="39">Dyscolus arauzae</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136E5313FCC8F898FC10F8FF" box="[825,918,1890,1916]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (QCAZ).
<emphasis id="B96A1F6E136E5313FB34F89BFB64F8F8" bold="true" box="[1221,1250,1889,1915]" pageId="38" pageNumber="39">37</emphasis>
.
<taxonomicName id="4C1EB8FF136E5313FB03F898FE83F81C" authority="Moret" authorityName="Moret" authorityYear="2020" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="piscator" status="sp. nov.">
<emphasis id="B96A1F6E136E5313FB03F898FAF0F8F8" box="[1266,1398,1890,1915]" italics="true" pageId="38" pageNumber="39">D. piscator</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136E5313FEE3F87FFEF3F81C" box="[274,373,1925,1951]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (QACZ).
<emphasis id="B96A1F6E136E5313FD48F87EFD50F81D" bold="true" box="[697,726,1924,1950]" pageId="38" pageNumber="39">38</emphasis>
.
<taxonomicName id="4C1EB8FF136E5313FD1BF87FFC40F81C" authority="Moret" authorityName="Moret" authorityYear="2020" box="[746,966,1924,1951]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E136E5313FD1BF87FFCF7F81D" box="[746,881,1924,1950]" italics="true" pageId="38" pageNumber="39">D. placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136E5313FC22F87FFBB0F81C" box="[979,1078,1925,1951]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (MNHN).
<emphasis id="B96A1F6E136E5313FF4CF85DFF5CF842" bold="true" box="[189,218,1959,1985]" pageId="38" pageNumber="39">39</emphasis>
.
<taxonomicName id="4C1EB8FF136E5313FF18F852FE3CF841" authority="Moret" authorityName="Moret" authorityYear="2020" box="[233,442,1960,1986]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="arauzae" status="sp. nov.">
<emphasis id="B96A1F6E136E5313FF18F852FEEDF842" box="[233,363,1960,1985]" italics="true" pageId="38" pageNumber="39">D. arauzae</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136E5313FE30F852FD9DF841" box="[449,539,1960,1986]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B96A1F6E136E5313FDD3F85DFDB9F842" bold="true" box="[546,575,1959,1985]" pageId="38" pageNumber="39">40</emphasis>
.
<taxonomicName id="4C1EB8FF136E5313FDBFF852FCA6F841" authority="Moret" authorityName="Moret" authorityYear="2020" box="[590,800,1960,1986]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="piscator" status="sp. nov.">
<emphasis id="B96A1F6E136E5313FDBFF852FD57F842" box="[590,721,1960,1985]" italics="true" pageId="38" pageNumber="39">D. piscator</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136E5313FCD6F852FC04F841" box="[807,898,1960,1986]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B96A1F6E136E5313FC79F85DFC23F842" bold="true" box="[904,933,1959,1985]" pageId="38" pageNumber="39">41</emphasis>
.
<taxonomicName id="4C1EB8FF136E5313FC45F852FB02F841" authority="Moret" authorityName="Moret" authorityYear="2020" box="[948,1156,1960,1986]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E136E5313FC45F852FBB3F841" box="[948,1077,1960,1986]" italics="true" pageId="38" pageNumber="39">D. placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136E5313FB7BF852FB63F841" box="[1162,1253,1960,1986]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
(above) and
<taxonomicName id="4C1EB8FF136E5313FF4CF836FE16F866" authority="Moret" authorityName="Moret" authorityYear="2020" box="[189,400,1995,2021]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="piscator" status="sp. nov.">
<emphasis id="B96A1F6E136E5313FF4CF836FEC7F866" box="[189,321,1996,2021]" italics="true" pageId="38" pageNumber="39">D. piscator</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136E5313FE66F831FE74F866" box="[407,498,1995,2021]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
(below).
</paragraph>
</caption>
<paragraph id="8BA1C37C136F5312FF4CFE4CFE52FE53" blockId="39.[189,468,438,464]" box="[189,468,438,464]" pageId="39" pageNumber="40">
<heading id="D0E97410136F5312FF4CFE4CFE52FE53" bold="true" box="[189,468,438,464]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B96A1F6E136F5312FF4CFE4CFE52FE53" bold="true" box="[189,468,438,464]" pageId="39" pageNumber="40">Diagnostic description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C136F5312FF4CFE1FFB71FCDC" blockId="39.[189,1399,485,863]" pageId="39" pageNumber="40">
Habitus:
<figureCitation id="1325DFF9136F5312FEDEFE1FFE08FE7C" box="[303,398,485,511]" captionStart="Figs 36–41" captionStartId="38.[189,241,1854,1880]" captionTargetBox="[288,1297,277,1813]" captionTargetId="figure@38.[278,1309,265,1818]" captionTargetPageId="38" captionText="Figs 36–41. Dyscolus spp., habitus (36–38) and aedeagus, median lobe in lateral view (39–40) and apex of the median lobe (41). 36. Dyscolus arauzae Moret sp. nov., male holotype (QCAZ). 37. D. piscator Moret sp. nov., male holotype (QACZ). 38. D. placitus Moret sp. nov., male holotype (MNHN). 39. D. arauzae Moret sp. nov. 40. D. piscator Moret sp. nov. 41. D. placitus Moret sp. nov. (above) and D. piscator Moret sp. nov. (below)." figureDoi="http://doi.org/10.5281/zenodo.3829702" httpUri="https://zenodo.org/record/3829702/files/figure.png" pageId="39" pageNumber="40">Fig. 36</figureCitation>
. Wingless. Body length:
<quantity id="4CE66E99136F5312FD21FE1FFCF1FE7C" box="[720,887,485,512]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" metricValueMax="10.9" metricValueMin="8.5" pageId="39" pageNumber="40" unit="mm" value="9.7" valueMax="10.9" valueMin="8.5">8.5–10.9 mm</quantity>
. Antennae unicoloured, testaceous, as in
<taxonomicName id="4C1EB8FF136F5312FF4CFDF3FEC6FDA1" authorityName="Moret" authorityYear="2005" box="[189,320,520,546]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B96A1F6E136F5312FF4CFDF3FEC6FDA1" box="[189,320,520,546]" italics="true" pageId="39" pageNumber="40">D. hirsutus</emphasis>
</taxonomicName>
(apical part of the antennomeres darker in
<taxonomicName id="4C1EB8FF136F5312FCC8FDF3FB8AFDA0" authority="Moret" authorityName="Moret" authorityYear="2020" box="[825,1036,521,547]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="piscator" status="sp. nov.">
<emphasis id="B96A1F6E136F5312FCC8FDF3FC3BFDA1" box="[825,957,521,546]" italics="true" pageId="39" pageNumber="40">D. piscator</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136F5312FBE2FDF3FBE8FDA0" box="[1043,1134,521,547]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1EB8FF136F5312FB56FDF3FAF1FDA0" authority="Moret" authorityName="Moret" authorityYear="2020" box="[1191,1399,520,547]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E136F5312FB56FDF3FAA1FDA1" box="[1191,1319,520,546]" italics="true" pageId="39" pageNumber="40">D. placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136F5312FF4CFDD6FE9EFDC5" box="[189,280,556,582]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
). Colour of the ventral face variable. Dimensions of the head intermediate between
<taxonomicName id="4C1EB8FF136F5312FB03FDD6FE83FDEA" authority="Moret" authorityName="Moret" authorityYear="2020" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="piscator" status="sp. nov.">
<emphasis id="B96A1F6E136F5312FB03FDD6FAF0FDC6" box="[1266,1398,556,581]" italics="true" pageId="39" pageNumber="40">D. piscator</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136F5312FEFDFDB5FEE0FDEA" box="[268,358,591,617]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1EB8FF136F5312FE6FFDB5FDE8FDEA" authority="Moret" authorityName="Moret" authorityYear="2020" box="[414,622,591,617]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E136F5312FE6FFDB5FD99FDEA" box="[414,543,591,617]" italics="true" pageId="39" pageNumber="40">D. placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136F5312FD84FDB5FD49FDEA" box="[629,719,591,617]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
(length of the right mandible / width of the frons between the anterior setae = 1.05). Apex of the labrum straight. Lateroposterior seta of the pronotum close to the hind angle (slightly shifted forward in
<taxonomicName id="4C1EB8FF136F5312FD70FD6CFCD2FD2C" authority="Moret" authorityName="Moret" authorityYear="2020" box="[641,852,661,687]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="piscator" status="sp. nov.">
<emphasis id="B96A1F6E136F5312FD70FD6CFC82FD2C" box="[641,772,662,687]" italics="true" pageId="39" pageNumber="40">D. piscator</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136F5312FCABFD6FFC33FD2C" box="[858,949,661,687]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1EB8FF136F5312FC1CFD6CFB3BFD2C" authority="Moret" authorityName="Moret" authorityYear="2020" box="[1005,1213,661,687]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E136F5312FC1CFD6CFBE8FD2C" box="[1005,1134,661,687]" italics="true" pageId="39" pageNumber="40">D. placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136F5312FB35FD6FFA98FD2C" box="[1220,1310,661,687]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
). Third elytral interval with 4–7 setae, fifth interval with 2–4 setae, seventh interval normally without setae, only one
<typeStatus id="54A57DDE136F5312FF1FFD26FED5FD75" box="[238,339,732,758]" pageId="39" pageNumber="40" type="paratype">paratype</typeStatus>
with one seta; 15–17 umbilicate setae along the lateral margin (
<quantity id="4CE66E99136F5312FBC9FD21FB19FD76" box="[1080,1183,731,757]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.302" metricValueMin="2.794" pageId="39" pageNumber="40" unit="in" value="12.0" valueMax="13.0" valueMin="11.0">11–13 in</quantity>
<taxonomicName id="4C1EB8FF136F5312FB54FD26FAF1FD75" authority="Moret" authorityName="Moret" authorityYear="2020" box="[1189,1399,732,758]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="piscator" status="sp. nov.">
<emphasis id="B96A1F6E136F5312FB54FD26FAAEFD76" box="[1189,1320,732,757]" italics="true" pageId="39" pageNumber="40">D. piscator</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136F5312FF4CFD05FE9AFC9A" box="[189,284,767,793]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
,
<quantity id="4CE66E99136F5312FED8FD05FE15FC9B" box="[297,403,766,793]" metricMagnitude="-1" metricUnit="m" metricValue="4.699" metricValueMax="5.08" metricValueMin="4.318" pageId="39" pageNumber="40" unit="in" value="18.5" valueMax="20.0" valueMin="17.0">17–20 in</quantity>
<taxonomicName id="4C1EB8FF136F5312FE6DFD05FDF6FC9A" authority="Moret" authorityName="Moret" authorityYear="2020" box="[412,624,767,793]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E136F5312FE6DFD05FD99FC9A" box="[412,543,767,793]" italics="true" pageId="39" pageNumber="40">D. placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136F5312FD89FD05FD53FC9A" box="[632,725,767,793]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
).
<emphasis id="B96A1F6E136F5312FD1EFD05FC1AFC9A" box="[751,924,767,793]" italics="true" pageId="39" pageNumber="40">Male genitalia</emphasis>
:
<figureCitation id="1325DFF9136F5312FC5FFD04FB80FC9A" box="[942,1030,766,793]" captionStart="Figs 36–41" captionStartId="38.[189,241,1854,1880]" captionTargetBox="[288,1297,277,1813]" captionTargetId="figure@38.[278,1309,265,1818]" captionTargetPageId="38" captionText="Figs 36–41. Dyscolus spp., habitus (36–38) and aedeagus, median lobe in lateral view (39–40) and apex of the median lobe (41). 36. Dyscolus arauzae Moret sp. nov., male holotype (QCAZ). 37. D. piscator Moret sp. nov., male holotype (QACZ). 38. D. placitus Moret sp. nov., male holotype (MNHN). 39. D. arauzae Moret sp. nov. 40. D. piscator Moret sp. nov. 41. D. placitus Moret sp. nov. (above) and D. piscator Moret sp. nov. (below)." figureDoi="http://doi.org/10.5281/zenodo.3829702" httpUri="https://zenodo.org/record/3829702/files/figure.png" pageId="39" pageNumber="40">Fig. 39</figureCitation>
. Apex of the median lobe bent downwards, resembling that of
<taxonomicName id="4C1EB8FF136F5312FDC2FCD9FC82FCBF" authority="Moret" authorityName="Moret" authorityYear="2020" box="[563,772,802,828]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E136F5312FDC2FCD9FD32FCBF" box="[563,692,802,828]" italics="true" pageId="39" pageNumber="40">D. placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136F5312FCFDFCD8FCE1FCBF" box="[780,871,802,828]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
; apical part of the endophallus covered with sclerotized squamae (unarmed in
<taxonomicName id="4C1EB8FF136F5312FDB6FCBCFC9DFCDC" authority="Moret" authorityName="Moret" authorityYear="2020" box="[583,795,837,863]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="piscator" status="sp. nov.">
<emphasis id="B96A1F6E136F5312FDB6FCBCFD4DFCDC" box="[583,715,838,863]" italics="true" pageId="39" pageNumber="40">D. piscator</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136F5312FCD3FCBFFCFAFCDC" box="[802,892,837,863]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1EB8FF136F5312FC44FCBCFB03FCDC" authority="Moret" authorityName="Moret" authorityYear="2020" box="[949,1157,837,863]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E136F5312FC44FCBCFBB0FCDC" box="[949,1078,837,863]" italics="true" pageId="39" pageNumber="40">D. placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136F5312FB7DFCBFFB60FCDC" box="[1164,1254,837,863]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
<paragraph id="8BA1C37C136F5312FF4CFC7CFE9BFC23" blockId="39.[189,285,902,928]" box="[189,285,902,928]" pageId="39" pageNumber="40">
<heading id="D0E97410136F5312FF4CFC7CFE9BFC23" bold="true" box="[189,285,902,928]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B96A1F6E136F5312FF4CFC7CFE9BFC23" bold="true" box="[189,285,902,928]" pageId="39" pageNumber="40">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C136F5312FF4CFC4FFB16FC4C" blockId="39.[189,1168,949,975]" box="[189,1168,949,975]" pageId="39" pageNumber="40">
Riparian along glacier-fed streams in the humid superpáramo at
<quantity id="4CE66E99136F5312FC41FC4FFBD4FC4C" box="[944,1106,949,975]" metricMagnitude="3" metricUnit="m" metricValue="4.35" metricValueMax="4.4" metricValueMin="4.3" pageId="39" pageNumber="40" unit="m" value="4350.0" valueMax="4400.0" valueMin="4300.0">4300–4400 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8BA1C37C136F5312FF4CFC0CFE6DFB93" blockId="39.[189,491,1014,1040]" box="[189,491,1014,1040]" pageId="39" pageNumber="40">
<heading id="D0E97410136F5312FF4CFC0CFE6DFB93" bold="true" box="[189,491,1014,1040]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B96A1F6E136F5312FF4CFC0CFE6DFB93" bold="true" box="[189,491,1014,1040]" pageId="39" pageNumber="40">Geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C136F5312FF4CFBDFFB88FBBC" blockId="39.[189,1038,1061,1087]" box="[189,1038,1061,1087]" pageId="39" pageNumber="40">Microendemic species, presumably restricted to the Cayambe mountain.</paragraph>
</subSubSection>
</treatment>
</document>
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