Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes Author Moret, Pierre 2E194645-D255-438B-819E-A2B6C39AD699 Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France. pierre.moret@univ-tlse2.fr&pierre.moret@univ-tlse2.fr Author Murienne, Jérôme 3A6964D4-A7D6-46DB-8129-B762D14BC8F9 Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France. jerome.murienne@univ-tlse3.fr text European Journal of Taxonomy 2020 2020-05-15 646 1 55 journal article 22092 10.5852/ejt.2020.646 c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b 3829682 4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B Dyscolus arauzae Moret sp. nov. urn:lsid:zoobank.org:act: B992F8BA-69F8-4591-A010-440432759483 Figs 36, 39 Etymology Noun in the genitive case, dedicated to María de los Ángeles Aráuz who took part in the discovery of this species. Type material Holotype ECUADOR • ♂ ; Pichincha Province , Volcán Cayambe SW , Waypoint 73 ; 0.006962º N , 78.020278º W ; 4405 m a.s.l. ; 30 Oct. 2015 ; P. Moret and M. Aráuz leg.; QCAZ . Paratypes (6 ♂♂, 6 ♀♀ ) ECUADOR – Pichincha Province • 1 ♂ ; same collection data as for holotype; MNHN • 1 ♂ , 1 ♀ ; same collection data as for holotype; CPM • 1 ♀ ; same collection data as for holotype; COI voucher PM073-03, BOLD sequence SUM020-18; CPM • 1 ♂ ; Volcán Cayambe SW , Waypoint 77 ; 0.007724º N , 78.023350º W ; 4300 m a.s.l. ; 30 Oct. 2015 ; P. Moret and M. Aráuz leg.; COI voucher PM077-01, BOLD sequence SUM099-18; CPM • 3 ♂♂ , 2 ♀♀ ; Volcán Cayambe SW , Waypoint 267 ; 0°00.415′ N , 78°1.338′ W ; 4294 m a.s.l. ; 4 Mar. 2017 ; P. Moret and M. Gobbi leg.; CPM • 1 ♀ (in ethanol); Volcán Cayambe SW , Waypoint 76 ; 0.006860º N , 78.020714º W ; 4390 m a.s.l. ; 18 Jul. 2016 ; P. Moret and E. Moreno leg.; CPM • 1 ♀ (in ethanol); same collection data as for preceding; 4 Mar. 2017 ; P. Moret and M. Gobbi leg.; CPM . Figs 36–41. Dyscolus spp., habitus (36–38) and aedeagus, median lobe in lateral view (39–40) and apex of the median lobe (41). 36 . Dyscolus arauzae Moret sp. nov. , male holotype (QCAZ). 37 . D. piscator Moret sp. nov. , male holotype (QACZ). 38 . D. placitus Moret sp. nov. , male holotype (MNHN). 39 . D. arauzae Moret sp. nov. 40 . D. piscator Moret sp. nov. 41 . D. placitus Moret sp. nov. (above) and D. piscator Moret sp. nov. (below). Diagnostic description Habitus: Fig. 36 . Wingless. Body length: 8.5–10.9 mm . Antennae unicoloured, testaceous, as in D. hirsutus (apical part of the antennomeres darker in D. piscator Moret sp. nov. and D. placitus Moret sp. nov. ). Colour of the ventral face variable. Dimensions of the head intermediate between D. piscator Moret sp. nov. and D. placitus Moret sp. nov. (length of the right mandible / width of the frons between the anterior setae = 1.05). Apex of the labrum straight. Lateroposterior seta of the pronotum close to the hind angle (slightly shifted forward in D. piscator Moret sp. nov. and D. placitus Moret sp. nov. ). Third elytral interval with 4–7 setae, fifth interval with 2–4 setae, seventh interval normally without setae, only one paratype with one seta; 15–17 umbilicate setae along the lateral margin ( 11–13 in D. piscator Moret sp. nov. , 17–20 in D. placitus Moret sp. nov. ). Male genitalia : Fig. 39 . Apex of the median lobe bent downwards, resembling that of D. placitus Moret sp. nov. ; apical part of the endophallus covered with sclerotized squamae (unarmed in D. piscator Moret sp. nov. and D. placitus Moret sp. nov. ). Habitat Riparian along glacier-fed streams in the humid superpáramo at 4300–4400 m a.s.l. Geographic distribution Microendemic species, presumably restricted to the Cayambe mountain.