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<mods:title id="DA05F246E763658EFFBA7E3AE0BE316F">Revision of the Nearctic Species of the Shore-Fly Genus Scatophila Becker (Diptera: Ephydridae)</mods:title>
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<mods:namePart id="5A1A91F09AFE37C8C74C537691E016B6">Zatwarnicki, Tadeusz</mods:namePart>
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<mods:affiliation id="1CD648FF67271A96CC72DFE8625E123D">Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143</mods:affiliation>
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<mods:namePart id="F77E6FFE603BB346AD6F8908124EA16F">Mathis, Wayne N.</mods:namePart>
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<mods:affiliation id="6ABF04DC80AF83979CEE3171F6BC6F54">Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143 &amp; Department of Entomology, PO BOX 37012, MRC 169, Smithsonian Institution, Washington, DC 20013 - 7012, USA mathisw @ si. edu; https: // orcid. org / 0000 - 0002 - 5012 - 1762 &amp; Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143</mods:affiliation>
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3. 
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<emphasis id="B90CEAB33A407B40FF6CFDB956F7FD82" bold="true" box="[181,413,605,632]" italics="true" pageId="19" pageNumber="20">Scatophila angusta</emphasis>
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<taxonomicNameLabel id="A23F57C83A407B40FE7DFDBA5568FD82" box="[420,514,606,632]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BC736A13A407B40FF4EFD6656AEFD66" blockId="19.[151,514,605,668]" box="[151,452,642,668]" pageId="19" pageNumber="20">
<figureCitation id="13432A243A407B40FF4EFD665674FD66" box="[151,286,642,668]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1564,1588]" captionTargetBox="[172,1436,346,1540]" captionTargetId="figure-98@16.[151,1436,345,1540]" captionTargetPageId="16" captionText="FIGURES 14–17. Head of Scatophila. 14. S. exilis Cresson, latero-oblique view. 15. Same, lateral view. 16. Scatophila angusta sp. nov., latero-oblique view. 17. Same, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210781" httpUri="https://zenodo.org/record/13210781/files/figure.png" pageId="19" pageNumber="20">Figs. 16–17</figureCitation>
, 
<figureCitation id="13432A243A407B40FEF0FD665618FD66" box="[297,370,642,668]" captionStart="FIGURES 22–26" captionStartId="20.[151,264,1536,1560]" captionTargetBox="[151,1435,205,1511]" captionTargetId="figure-30@20.[151,1435,181,1512]" captionTargetPageId="20" captionText="FIGURES 22–26. Head, thorax and wing of Scatophila. 22. Scatophila angusta sp. nov., head and thorax, dorsal view. 23. Same, wing, dorsal view. 24. Wing of S. variabilis Cresson, dorsal view. 25–26. Head of Scatophila variabilis Cresson. 25. Male, latero-oblique view. 26. Male, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210787" httpUri="https://zenodo.org/record/13210787/files/figure.png" pageId="19" pageNumber="20">22–23</figureCitation>
, 
<figureCitation id="13432A243A407B40FEA4FD6656AEFD66" box="[381,452,642,668]" captionStart-0="FIGURES 27–29" captionStart-1="FIGURE 30" captionStartId-0="21.[151,264,1176,1200]" captionStartId-1="22.[151,250,1023,1047]" captionTargetBox-0="[159,1427,189,1144]" captionTargetBox-1="[151,1435,181,999]" captionTargetId-0="figure-27@21.[151,1435,181,1152]" captionTargetId-1="figure-30@22.[151,1435,181,999]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURES 27–29. Male terminalia of Scatophila angusta sp. nov. 27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm." captionText-1="FIGURE 30. Distribution map of Scatophila angusta sp. nov. (circles), S. conifera Sturtevant &amp; Wheeler (triangle), and S. contaminata (Stenhammar) (squares) in Nearctic Region." figureDoi-0="http://doi.org/10.5281/zenodo.13210789" figureDoi-1="http://doi.org/10.5281/zenodo.13210791" httpUri-0="https://zenodo.org/record/13210789/files/figure.png" httpUri-1="https://zenodo.org/record/13210791/files/figure.png" pageId="19" pageNumber="20">27–30</figureCitation>
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<taxonomicName id="4C784D223A407B40FF4EFD245603FD22" ID-CoL="4V455" box="[151,361,704,728]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B90CEAB33A407B40FF4EFD245603FD22" box="[151,361,704,728]" italics="true" pageId="19" pageNumber="20">Scatophila variabilis</emphasis>
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of authors, not 
<treatmentCitation id="0AD910B03A407B40FDDAFD2455D4FD22" author="Cresson, E. T., Jr." box="[515,702,704,728]" page="341" pageId="19" pageNumber="20" year="1917">
<bibRefCitation id="EFE94B503A407B40FDDAFD2455D4FD22" author="Cresson, E. T., Jr." box="[515,702,704,728]" pageId="19" pageNumber="20" pagination="340 - 341" refId="ref69446" refString="Cresson, E. T., Jr. (1917) Descriptions of new genera and species of the dipterous family Ephydridae. IV. Entomological News, 28 (8), 340 - 341." type="journal article" year="1917">Cresson 1917: 341</bibRefCitation>
</treatmentCitation>
[misidentification in part].— 
<treatmentCitation id="0AD910B03A407B40FC39FD2453ADFD22" author="Zatwarnicki, T." box="[992,1223,704,728]" page="292" pageId="19" pageNumber="20" year="1987">
<bibRefCitation id="EFE94B503A407B40FC39FD2453ADFD22" author="Zatwarnicki, T." box="[992,1223,704,728]" pageId="19" pageNumber="20" pagination="277 - 298" refId="ref71260" refString="Zatwarnicki, T. (1987) New synonyms and records of Palearctic Scatophila (Diptera, Ephydridae). Polskie Pismo Entomologiczne, 57 (2), 277 - 298." type="journal article" year="1987">Zatwarnicki 1987: 292</bibRefCitation>
</treatmentCitation>
[checklist].— 
<treatmentCitation id="0AD910B03A407B40FA8DFD2456B3FD0C" author="Mathis, W. N. &amp; Zatwarnicki, T." page="287" pageId="19" pageNumber="20" year="1995">
<bibRefCitation id="EFE94B503A407B40FA8DFD2456B3FD0C" author="Mathis, W. N. &amp; Zatwarnicki, T." pageId="19" pageNumber="20" pagination="1 - 423" refId="ref70301" refString="Mathis, W. N. &amp; Zatwarnicki, T. (1995) A world catalog of the shore flies (Diptera: Ephydridae). International Memoirs on Entomology, 4, i - vi + 1 - 423." type="journal article" year="1995">Mathis and Zatwarnicki 1995: 287</bibRefCitation>
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[world catalog].
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<taxonomicName id="4C784D223A407B40FF4EFD19560EFCEF" ID-CoL="4V43T" baseAuthorityName="Haliday" baseAuthorityYear="1839" box="[151,356,765,789]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="despecta">
<emphasis id="B90CEAB33A407B40FF4EFD19560EFCEF" box="[151,356,765,789]" italics="true" pageId="19" pageNumber="20">Scatophila despecta</emphasis>
</taxonomicName>
of authors, not (
<bibRefCitation id="EFE94B503A407B40FDCBFD1955F4FCEF" author="Haliday, A. H." box="[530,670,765,789]" pageId="19" pageNumber="20" pagination="401 - 411" refId="ref69860" refString="Haliday, A. H. (1839) Remarks on the generic distribution of the British Hydromyzidae. Annals of Natural History, Series 1, 3, 217 - 224 + 401 - 411. https: // doi. org / 10.1080 / 03745483909443252" type="journal article" year="1839">Haliday 1839</bibRefCitation>
) [misidentification].— 
<treatmentCitation id="0AD910B03A407B40FC51FD195339FCEF" author="Sturtevant, A. H." box="[904,1107,765,789]" page="11" pageId="19" pageNumber="20" year="1923">
<bibRefCitation id="EFE94B503A407B40FC51FD195339FCEF" author="Sturtevant, A. H." box="[904,1107,765,789]" pageId="19" pageNumber="20" pagination="1 - 12" refId="ref70800" refString="Sturtevant, A. H. (1923) New species and notes on synonymy and distribution of Muscidae Acalypteratae (Diptera). American Museum Novitates, 76, 1 - 12." type="journal article" year="1923">Sturtevant 1923: 11</bibRefCitation>
</treatmentCitation>
[taxonomic notes].— 
<treatmentCitation id="0AD910B03A407B40FAE9FD19568CFCCE" author="Sturtevant, A. H. &amp; Wheeler, M. R." page="190" pageId="19" pageNumber="20" year="1954">
<bibRefCitation id="EFE94B503A407B40FAE9FD19568CFCCE" author="Sturtevant, A. H. &amp; Wheeler, M. R." pageId="19" pageNumber="20" pagination="151 - 257" refId="ref70834" refString="Sturtevant, A. H. &amp; Wheeler, M. R. (1954) Synopses of Nearctic Ephydridae (Diptera). Transactions of the American Entomological Society, 79, 151 - 257." type="journal article" year="1954">Sturtevant and Wheeler 1954: 190–191</bibRefCitation>
</treatmentCitation>
[review].—
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<treatmentCitation id="0AD910B03A407B40FDBBFCF85463FCC9" author="Wirth, W. W." box="[610,777,795,820]" page="758" pageId="19" pageNumber="20" year="1965">
<bibRefCitation id="EFE94B503A407B40FDBBFCF85463FCC9" author="Wirth, W. W." box="[610,777,795,820]" pageId="19" pageNumber="20" pagination="734 - 759" refId="ref70987" refString="Wirth, W. W. (1965) Ephydridae. In: Stone, A., Sabrosky, C. W., Wirth, W. W., Foote, R. W. &amp; Coulson, J. R. (Eds.), A Catalog of the Diptera of America North of Mexico. Handbook 276. U. S. Department of Agriculture, Washington, D. C., pp. 734 - 759." type="book chapter" year="1965">Wirth 1965: 758</bibRefCitation>
</treatmentCitation>
[
<taxonomicName id="4C784D223A407B40FCCCFCF85404FCCE" box="[789,878,796,820]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="97" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
catalog].
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</paragraph>
</subSubSection>
<subSubSection id="C362652A3A407B46FF4EFCBF5209FA73" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BC736A13A407B40FF4EFCBF5525FC18" blockId="19.[151,1437,859,2038]" pageId="19" pageNumber="20">
Diagnosis.—This species is a member of the 
<taxonomicName id="4C784D223A407B40FD79FCBF5468FC8F" baseAuthorityName="Haliday" baseAuthorityYear="1839" box="[672,770,859,885]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="despecta">
<emphasis id="B90CEAB33A407B40FD79FCBF5468FC8F" box="[672,770,859,885]" italics="true" pageId="19" pageNumber="20">despecta</emphasis>
</taxonomicName>
species group and is distinguished from congeners by the following combination of characters: Body with tan to brown and usually with some lighter markings, scutellar disc with partially to mostly dark but often with basal and sometimes with apical areas gray; small to moderately small shore flies, body length 
<quantity id="4C809B443A407B40FE7CFC235521FC18" box="[421,587,967,994]" metricMagnitude="-3" metricUnit="m" metricValue="1.875" metricValueMax="2.1" metricValueMin="1.65" pageId="19" pageNumber="20" unit="mm" value="1.875" valueMax="2.1" valueMin="1.65">1.65–2.10 mm</quantity>
.
</paragraph>
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<emphasis id="B90CEAB33A407B40FF1EFC0F566EFBFF" box="[199,260,1003,1029]" italics="true" pageId="19" pageNumber="20">Head</emphasis>
(
<figureCitation id="13432A243A407B40FECDFC0F56F5FBFF" box="[276,415,1003,1029]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1564,1588]" captionTargetBox="[172,1436,346,1540]" captionTargetId="figure-98@16.[151,1436,345,1540]" captionTargetPageId="16" captionText="FIGURES 14–17. Head of Scatophila. 14. S. exilis Cresson, latero-oblique view. 15. Same, lateral view. 16. Scatophila angusta sp. nov., latero-oblique view. 17. Same, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210781" httpUri="https://zenodo.org/record/13210781/files/figure.png" pageId="19" pageNumber="20">Figs. 16–17</figureCitation>
): Lateroclinate fronto-orbital seta inserted far forward, 2–3X as far from medial vertical as the latter is from the lateral vertical seta. Face gray, face of 
<specimenCount id="9D7EFD283A407B40FCC3FBF45445FBD0" box="[794,815,1040,1066]" count="1" pageId="19" pageNumber="20" type="male">♂</specimenCount>
normally projected, transversely arched, without medial area shallowly depressed; ventral margin of male face nearly flat, not distinctly emarginate; face bearing porrect to slightly dorsally curved seta lateroventrally. Gena high, height slightly greater than height of basal flagellomere; gena-to-eye ratio 0.34–037.
</paragraph>
<paragraph id="8BC736A13A407B40FF1EFB7B562DFA4F" blockId="19.[151,1437,859,2038]" pageId="19" pageNumber="20">
<emphasis id="B90CEAB33A407B40FF1EFB7B5672FB43" box="[199,280,1183,1209]" italics="true" pageId="19" pageNumber="20">Thorax</emphasis>
(
<figureCitation id="13432A243A407B40FEFCFB7B56C9FB43" box="[293,419,1183,1209]" captionStart="FIGURES 22–26" captionStartId="20.[151,264,1536,1560]" captionTargetBox="[151,1435,205,1511]" captionTargetId="figure-30@20.[151,1435,181,1512]" captionTargetPageId="20" captionText="FIGURES 22–26. Head, thorax and wing of Scatophila. 22. Scatophila angusta sp. nov., head and thorax, dorsal view. 23. Same, wing, dorsal view. 24. Wing of S. variabilis Cresson, dorsal view. 25–26. Head of Scatophila variabilis Cresson. 25. Male, latero-oblique view. 26. Male, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210787" httpUri="https://zenodo.org/record/13210787/files/figure.png" pageId="19" pageNumber="20">Fig. 22–23</figureCitation>
): Mesonotum (
<figureCitation id="13432A243A407B40FD9EFB7B55FCFB43" box="[583,662,1183,1209]" captionStart="FIGURES 22–26" captionStartId="20.[151,264,1536,1560]" captionTargetBox="[151,1435,205,1511]" captionTargetId="figure-30@20.[151,1435,181,1512]" captionTargetPageId="20" captionText="FIGURES 22–26. Head, thorax and wing of Scatophila. 22. Scatophila angusta sp. nov., head and thorax, dorsal view. 23. Same, wing, dorsal view. 24. Wing of S. variabilis Cresson, dorsal view. 25–26. Head of Scatophila variabilis Cresson. 25. Male, latero-oblique view. 26. Male, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210787" httpUri="https://zenodo.org/record/13210787/files/figure.png" pageId="19" pageNumber="20">Fig. 22</figureCitation>
) mostly brown with some grayish stripes; acrostichal setae irregular in size, arranged in 2 regular rows; dorsocentral setae 2, anterior seta sutural or just postsutural, presutural dorsocentral setae lacking; scutum mostly brown but with extensive pattern of gray areas anterior, laterad of acrostichal tracks and especially laterad of dorsocentral track and over postpronotum; medial area brown, becoming wider posteriorly. Wing (
<figureCitation id="13432A243A407B40FF3BFACB5658FAB3" box="[226,306,1327,1353]" captionStart="FIGURES 22–26" captionStartId="20.[151,264,1536,1560]" captionTargetBox="[151,1435,205,1511]" captionTargetId="figure-30@20.[151,1435,181,1512]" captionTargetPageId="20" captionText="FIGURES 22–26. Head, thorax and wing of Scatophila. 22. Scatophila angusta sp. nov., head and thorax, dorsal view. 23. Same, wing, dorsal view. 24. Wing of S. variabilis Cresson, dorsal view. 25–26. Head of Scatophila variabilis Cresson. 25. Male, latero-oblique view. 26. Male, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210787" httpUri="https://zenodo.org/record/13210787/files/figure.png" pageId="19" pageNumber="20">Fig. 23</figureCitation>
) moderately dark with evident pattern or pale spots although these not prominent; cell r 
<subScript id="17FC34E43A407B40FB21FAA6526AFAAA" attach="left" box="[1272,1280,1346,1360]" fontSize="6" pageId="19" pageNumber="20">1</subScript>
lacking white spots; large, quadrate white spot of cell r 
<subScript id="17FC34E43A407B40FD8EFA825535FA8E" attach="left" box="[599,607,1382,1396]" fontSize="6" pageId="19" pageNumber="20">3</subScript>
lacking a dark center; costal vein ratio 0.20–0.21; M 
<subScript id="17FC34E43A407B40FB71FA8253DAFA8E" attach="left" box="[1192,1200,1382,1396]" fontSize="6" pageId="19" pageNumber="20">1</subScript>
vein ratio 0.50–0.60. Knob of halter yellow. Midfemur of 
<specimenCount id="9D7EFD283A407B40FDE8FA9C552CFA68" box="[561,582,1400,1426]" count="1" pageId="19" pageNumber="20" type="male">♂</specimenCount>
bearing row of spine-like setae posteroventrally on basal half; tarsi blackish brown to black.
</paragraph>
<paragraph id="8BC736A13A407B40FF1EFA5B5431F80F" blockId="19.[151,1437,859,2038]" pageId="19" pageNumber="20">
<emphasis id="B90CEAB33A407B40FF1EFA5B5650FA23" box="[199,314,1471,1497]" italics="true" pageId="19" pageNumber="20">Abdomen:</emphasis>
Tergites mostly gray to grayish brown or black, rather heavily microtomentose, rarely subshiny, tergites darker anteriorly and laterally. Male terminalia: (
<figureCitation id="13432A243A407B40FCEDFA0754A9FA07" box="[820,963,1507,1533]" captionStart="FIGURES 27–29" captionStartId="21.[151,264,1176,1200]" captionTargetBox="[159,1427,189,1144]" captionTargetId="figure-27@21.[151,1435,181,1152]" captionTargetPageId="21" captionText="FIGURES 27–29. Male terminalia of Scatophila angusta sp. nov. 27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210789" httpUri="https://zenodo.org/record/13210789/files/figure.png" pageId="19" pageNumber="20">Figs. 27–29</figureCitation>
): Epandrium in posterior view (
<figureCitation id="13432A243A407B40FAE7FA0752FEFA07" box="[1342,1428,1507,1533]" captionStart="FIGURES 27–29" captionStartId="21.[151,264,1176,1200]" captionTargetBox="[159,1427,189,1144]" captionTargetId="figure-27@21.[151,1435,181,1152]" captionTargetPageId="21" captionText="FIGURES 27–29. Male terminalia of Scatophila angusta sp. nov. 27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210789" httpUri="https://zenodo.org/record/13210789/files/figure.png" pageId="19" pageNumber="20">Fig. 27</figureCitation>
) generally oval in shape, almost trapezoidal, with dorsal margin moderately narrowly rounded, dorsal portion of lateral margin sloped laterally, almost straight, ventral portion of lateral margin shallowly sinuous, ventral margin emarginate, width equal to cercal cavity width, ventral ¼ bearing long setulae, especially laterally, otherwise setulae scattered, cercal cavity as wide as high, cercal height about ¼ epandrial height; cerci in posterior view (
<figureCitation id="13432A243A407B40FA99F99752FEF977" box="[1344,1428,1651,1677]" captionStart="FIGURES 27–29" captionStartId="21.[151,264,1176,1200]" captionTargetBox="[159,1427,189,1144]" captionTargetId="figure-27@21.[151,1435,181,1152]" captionTargetPageId="21" captionText="FIGURES 27–29. Male terminalia of Scatophila angusta sp. nov. 27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210789" httpUri="https://zenodo.org/record/13210789/files/figure.png" pageId="19" pageNumber="20">Fig. 27</figureCitation>
) semihemispherical, acutely pointed dorsally; aedeagus in lateral view (
<figureCitation id="13432A243A407B40FC64F973537AF94B" box="[957,1040,1687,1713]" captionStart="FIGURES 27–29" captionStartId="21.[151,264,1176,1200]" captionTargetBox="[159,1427,189,1144]" captionTargetId="figure-27@21.[151,1435,181,1152]" captionTargetPageId="21" captionText="FIGURES 27–29. Male terminalia of Scatophila angusta sp. nov. 27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210789" httpUri="https://zenodo.org/record/13210789/files/figure.png" pageId="19" pageNumber="20">Fig. 29</figureCitation>
) more or less irregularly quadrate, broadly rounded ventrally, deeply excavated dorsally, with a crossbar-like process, roundly pointed at posteroventral corner, in ventral view (
<figureCitation id="13432A243A407B40FE71F93B5691F903" box="[424,507,1759,1785]" captionStart="FIGURES 27–29" captionStartId="21.[151,264,1176,1200]" captionTargetBox="[159,1427,189,1144]" captionTargetId="figure-27@21.[151,1435,181,1152]" captionTargetPageId="21" captionText="FIGURES 27–29. Male terminalia of Scatophila angusta sp. nov. 27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210789" httpUri="https://zenodo.org/record/13210789/files/figure.png" pageId="19" pageNumber="20">Fig. 28</figureCitation>
) obovate, wider basally, anterior margin rounded, thereafter irregularly tapered to posterior, pointed apex; ventral aedeagal process lacking; phallapodeme in lateral view (
<figureCitation id="13432A243A407B40FBA0F8E753A1F8E7" box="[1145,1227,1795,1821]" captionStart="FIGURES 27–29" captionStartId="21.[151,264,1176,1200]" captionTargetBox="[159,1427,189,1144]" captionTargetId="figure-27@21.[151,1435,181,1152]" captionTargetPageId="21" captionText="FIGURES 27–29. Male terminalia of Scatophila angusta sp. nov. 27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210789" httpUri="https://zenodo.org/record/13210789/files/figure.png" pageId="19" pageNumber="20">Fig. 29</figureCitation>
) elongate, narrow, rod-like, nearly straight, in ventral view (
<figureCitation id="13432A243A407B40FDB2F8C355D4F8BB" box="[619,702,1831,1857]" captionStart="FIGURES 27–29" captionStartId="21.[151,264,1176,1200]" captionTargetBox="[159,1427,189,1144]" captionTargetId="figure-27@21.[151,1435,181,1152]" captionTargetPageId="21" captionText="FIGURES 27–29. Male terminalia of Scatophila angusta sp. nov. 27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210789" httpUri="https://zenodo.org/record/13210789/files/figure.png" pageId="19" pageNumber="20">Fig. 28</figureCitation>
) with a hour-glass shape, constricted medially, both anterior and posterior margins about same width, anterior surface very shallowly emarginate, posterior margin truncate; postgonite in lateral view (
<figureCitation id="13432A243A407B40FE8BF88B56CDF873" box="[338,423,1903,1929]" captionStart="FIGURES 27–29" captionStartId="21.[151,264,1176,1200]" captionTargetBox="[159,1427,189,1144]" captionTargetId="figure-27@21.[151,1435,181,1152]" captionTargetPageId="21" captionText="FIGURES 27–29. Male terminalia of Scatophila angusta sp. nov. 27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210789" httpUri="https://zenodo.org/record/13210789/files/figure.png" pageId="19" pageNumber="20">Fig. 29</figureCitation>
) irregularly linear with medial enlargements, posterior extension slightly tapered, apex pointed, midlength anterior, thumb-like process, anterior extension digitiform, in ventral view (
<figureCitation id="13432A243A407B40FB6FF877526DF857" box="[1206,1287,1939,1965]" captionStart="FIGURES 27–29" captionStartId="21.[151,264,1176,1200]" captionTargetBox="[159,1427,189,1144]" captionTargetId="figure-27@21.[151,1435,181,1152]" captionTargetPageId="21" captionText="FIGURES 27–29. Male terminalia of Scatophila angusta sp. nov. 27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210789" httpUri="https://zenodo.org/record/13210789/files/figure.png" pageId="19" pageNumber="20">Fig. 28</figureCitation>
) more or less H-shaped, lateral structures narrow, tapered, wider anteriorly, becoming very narrow posteriorly; neohypandrium in lateral view (
<figureCitation id="13432A243A407B40FEF3F83F5611F80F" box="[298,379,2011,2037]" captionStart="FIGURES 27–29" captionStartId="21.[151,264,1176,1200]" captionTargetBox="[159,1427,189,1144]" captionTargetId="figure-27@21.[151,1435,181,1152]" captionTargetPageId="21" captionText="FIGURES 27–29. Male terminalia of Scatophila angusta sp. nov. 27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210789" httpUri="https://zenodo.org/record/13210789/files/figure.png" pageId="19" pageNumber="20">Fig. 29</figureCitation>
) elongate, narrow, rod-like nearly straight.
</paragraph>
<caption id="DF0766293A477B47FF4EF9E45434F99A" ID-DOI="http://doi.org/10.5281/zenodo.13210787" ID-Zenodo-Dep="13210787" httpUri="https://zenodo.org/record/13210787/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,264,1536,1560]" targetBox="[151,1435,205,1511]" targetPageId="20" targetType="figure">
<paragraph id="8BC736A13A477B47FF4EF9E45434F99A" blockId="20.[151,1437,1536,1632]" pageId="20" pageNumber="21">
<emphasis id="B90CEAB33A477B47FF4EF9E45625F9E2" bold="true" box="[151,335,1536,1560]" pageId="20" pageNumber="21">FIGURES 22–26</emphasis>
. Head, thorax and wing of 
<taxonomicName id="4C784D223A477B47FDA8F9E455B7F9E2" box="[625,733,1536,1560]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB33A477B47FDA8F9E455B7F9E2" box="[625,733,1536,1560]" italics="true" pageId="20" pageNumber="21">Scatophila</emphasis>
</taxonomicName>
. 22. 
<taxonomicName id="4C784D223A477B47FCCDF9E454B3F9E2" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[788,985,1536,1560]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="angusta" status="sp. nov.">
<emphasis id="B90CEAB33A477B47FCCDF9E454B3F9E2" box="[788,985,1536,1560]" italics="true" pageId="20" pageNumber="21">Scatophila angusta</emphasis>
</taxonomicName>
<emphasis id="B90CEAB33A477B47FC3BF9E4535EF9E2" bold="true" box="[994,1076,1536,1560]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="A23F57C83A477B47FC3BF9E4535EF9E2" box="[994,1076,1536,1560]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, head and thorax, dorsal view. 23. Same, wing, dorsal view. 24. Wing of 
<taxonomicName id="4C784D223A477B47FDF3F9C15469F9C6" authority="Cresson" authorityName="Cresson" box="[554,771,1572,1596]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B90CEAB33A477B47FDF3F9C155CCF9C6" box="[554,678,1572,1596]" italics="true" pageId="20" pageNumber="21">S. variabilis</emphasis>
Cresson
</taxonomicName>
, dorsal view. 25–26. Head of 
<taxonomicName id="4C784D223A477B47FBE7F9C0521FF9C6" authority="Cresson." authorityName="Cresson." box="[1086,1397,1572,1596]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B90CEAB33A477B47FBE7F9C05279F9C6" box="[1086,1299,1572,1596]" italics="true" pageId="20" pageNumber="21">Scatophila variabilis</emphasis>
Cresson.
</taxonomicName>
25. Male, latero-oblique view. 26. Male, lateral view. Scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BC736A13A477B47FF1EF9575369F8A6" blockId="20.[151,1437,1714,1957]" pageId="20" pageNumber="21">
Type Material.— 
<materialsCitation id="3B103CFC3A477B47FE53F9565495F8A6" collectingDate="2006-06-07" collectionCode="USNM" collectorName="D. &amp; W. N. Mathis" country="United States of America" elevation="825" latitude="44.82" location="The" longLatPrecision="92" longitude="-117.463165" pageId="20" pageNumber="21" specimenCode="ENT 00250326" specimenCount="25" specimenCount-female="2" specimenCount-male="11" typeStatus="holotype">
<location id="8EA7607A3A477B47FE53F95656DCF936" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A477B47FE53F95656DCF936" box="[394,438,1714,1740]" country="United States of America" latitude="44.82" longLatPrecision="92" longitude="-117.463165" name="The" pageId="20" pageNumber="21">The</location>
<typeStatus id="54C388033A477B47FE19F9565548F936" box="[448,546,1714,1740]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
male of 
<taxonomicName id="4C784D223A477B47FD54F9565402F936" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[653,872,1714,1740]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="angusta" status="sp. nov.">
<emphasis id="B90CEAB33A477B47FD54F9565402F936" box="[653,872,1714,1740]" italics="true" pageId="20" pageNumber="21">Scatophila angusta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57C83A477B47FCA8F9575491F936" box="[881,1019,1714,1741]" pageId="20" pageNumber="21" rank="species">new species</taxonomicNameLabel>
is labeled “ 
<collectingCountry id="F36F76313A477B47FB51F95753AEF937" box="[1160,1220,1715,1741]" name="United States of America" pageId="20" pageNumber="21">USA</collectingCountry>
. OR[egon]. Baker: Goose Creek (
<geoCoordinate id="EE4C50663A477B47FEE5F93356DBF90B" box="[316,433,1751,1777]" degrees="44" direction="north" minutes="49.2" orientation="latitude" pageId="20" pageNumber="21" precision="92" value="44.82">44°49.2'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A477B47FE66F933553AF90B" box="[447,592,1751,1777]" degrees="117" direction="west" minutes="27.79" orientation="longitude" pageId="20" pageNumber="21" precision="9" value="-117.463165">117°27.79'W</geoCoordinate>
; 
<quantity id="4C809B443A477B47FDB8F93255C7F90B" box="[609,685,1750,1777]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="20" pageNumber="21" unit="m" value="825.0">
<elevation id="0055D1923A477B47FDB8F93255C7F90B" box="[609,685,1750,1777]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="20" pageNumber="21" unit="m" value="825.0">825 m</elevation>
</quantity>
), 
<date id="FFC610613A477B47FD1DF933542CF90A" box="[708,838,1750,1777]" pageId="20" pageNumber="21" value="2006-06-07">
<collectingDate id="EF82E9893A477B47FD1DF933542CF90A" box="[708,838,1750,1777]" pageId="20" pageNumber="21" value="2006-06-07">7 Jun 2006</collectingDate>
</date>
, 
<collectorName id="268D53773A477B47FC8AF9335345F90A" box="[851,1071,1750,1777]" pageId="20" pageNumber="21">D. &amp; W. N. Mathis</collectorName>
/ 
<collectionCode id="ED69AE643A477B47FBE1F93353E3F90B" box="[1080,1161,1751,1777]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
<specimenCode id="DBDE9EDA3A477B47FB4BF933522BF90B" box="[1170,1345,1750,1777]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">ENT 00250326</specimenCode>
[plastic bar code label]/HOLOTYPE 
<specimenCount id="9D7EFD283A477B47FE3CF91F5690F8EF" box="[485,506,1787,1813]" count="1" pageId="20" pageNumber="21" type="male">♂</specimenCount>
<taxonomicName id="4C784D223A477B47FDDAF91E5351F8EF" authority="Zatwarnicki &amp; Mathis USNM" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[515,1083,1786,1813]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="angusta">
<emphasis id="B90CEAB33A477B47FDDAF91E55B6F8EE" box="[515,732,1786,1812]" italics="true" pageId="20" pageNumber="21">Scatophila angusta</emphasis>
Zatwarnicki &amp; Mathis USNM
</taxonomicName>
[red].” The holotype is double mounted (minute nadel in block of plastic elastomere), is in excellent condition, and is deposited in the 
<collectionCode id="ED69AE643A477B47FA9DF8FB52F3F8C3" box="[1348,1433,1823,1849]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
. 
<specimenCount id="9D7EFD283A477B47FF4EF8A65631F8A7" box="[151,347,1858,1885]" count="12" pageId="20" pageNumber="21" type="generic" typeStatus="paratype">Twelve paratypes</specimenCount>
(
<specimenCount id="9D7EFD283A477B47FEB2F8A756F5F8A7" box="[363,415,1859,1885]" count="10" pageId="20" pageNumber="21" type="male">10♂</specimenCount>
, 
<specimenCount id="9D7EFD283A477B47FE72F8A756BAF8A7" box="[427,464,1859,1885]" count="2" pageId="20" pageNumber="21" type="female">2♀</specimenCount>
; 
<collectionCode id="ED69AE643A477B47FE04F8A7555AF8A7" box="[477,560,1859,1885]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
) bear the same label data as the holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A477B47FF1EF8825358F85F" blockId="20.[151,1437,1714,1957]" pageId="20" pageNumber="21">
Other Specimens Examined.— 
<materialsCitation id="3B103CFC3A477B47FDF0F88355F6F85F" collectingDate="2007-08-03" country="United States of America" county="Dolores" elevation="1955" latitude="37.648335" location="Dolores River" longLatPrecision="117" longitude="-108.735" municipality="Cahone" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Colorado">
<collectingCountry id="F36F76313A477B47FDF0F883546AF87B" box="[553,768,1895,1921]" name="United States of America" pageId="20" pageNumber="21">UNITED STATES</collectingCountry>
. 
<collectingRegion id="49BCF8433A477B47FCD4F883545FF85F" country="United States of America" name="Colorado" pageId="20" pageNumber="21">
<emphasis id="B90CEAB33A477B47FCD4F883537FF87A" box="[781,1045,1894,1921]" italics="true" pageId="20" pageNumber="21">
COLORADO. 
<collectingCounty id="62A64E2D3A477B47FC6AF8825365F87A" box="[947,1039,1894,1920]" pageId="20" pageNumber="21">Dolores</collectingCounty>
:
</emphasis>
<collectingMunicipality id="6BA3ACDB3A477B47FBC6F882531FF87A" box="[1055,1141,1894,1920]" pageId="20" pageNumber="21">Cahone</collectingMunicipality>
(
<quantity id="4C809B443A477B47FB51F88353AEF87A" box="[1160,1220,1894,1921]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="21" unit="km" value="9.0">9 km</quantity>
N; 
<location id="8EA7607A3A477B47FB2DF88252F2F87A" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A477B47FB2DF88252F2F87A" box="[1268,1432,1894,1920]" country="United States of America" county="Dolores" latitude="37.648335" longLatPrecision="117" longitude="-108.735" municipality="Cahone" name="Dolores River" pageId="20" pageNumber="21" stateProvince="Colorado">Dolores River</location>
; 
<geoCoordinate id="EE4C50663A477B47FF4EF86F5661F85F" box="[151,267,1931,1957]" degrees="37" direction="north" minutes="38.9" orientation="latitude" pageId="20" pageNumber="21" precision="92" value="37.648335">37°38.9'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A477B47FECEF86F56F5F85F" box="[279,415,1931,1957]" degrees="108" direction="west" minutes="44.1" orientation="longitude" pageId="20" pageNumber="21" precision="92" value="-108.735">108°44.1'W</geoCoordinate>
; 
<quantity id="4C809B443A477B47FE73F86E5568F85F" box="[426,514,1930,1957]" metricMagnitude="3" metricUnit="m" metricValue="1.955" pageId="20" pageNumber="21" unit="m" value="1955.0">
<elevation id="0055D1923A477B47FE73F86E5568F85F" box="[426,514,1930,1957]" metricMagnitude="3" metricUnit="m" metricValue="1.955" pageId="20" pageNumber="21" unit="m" value="1955.0">1955 m</elevation>
</quantity>
), 
<date id="FFC610613A477B47FDCFF86F55F6F85F" box="[534,668,1931,1957]" pageId="20" pageNumber="21" value="2007-08-03">
<collectingDate id="EF82E9893A477B47FDCFF86F55F6F85F" box="[534,668,1931,1957]" pageId="20" pageNumber="21" value="2007-08-03">3 Aug 2007</collectingDate>
</date>
, D. and 
<materialsCitation id="3B103CFC3A477B47FD21F86F5344F85F" box="[760,1070,1930,1957]" collectionCode="USNM" country="United States of America" location="Mathis" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="W">
W. N. 
<location id="8EA7607A3A477B47FC99F86E54E4F85E" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A477B47FC99F86E54E4F85E" box="[832,910,1930,1956]" country="United States of America" name="Mathis" pageId="20" pageNumber="21" stateProvince="W">Mathis</location>
(
<specimenCount id="9D7EFD283A477B47FC46F86F54A9F85F" box="[927,963,1931,1957]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
; 
<collectionCode id="ED69AE643A477B47FC09F86F534CF85F" box="[976,1062,1931,1957]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
)
</materialsCitation>
</collectingRegion>
</materialsCitation>
.
</paragraph>
<caption id="DF0766293A467B46FF4EFB7C523EFB2E" ID-DOI="http://doi.org/10.5281/zenodo.13210789" ID-Zenodo-Dep="13210789" httpUri="https://zenodo.org/record/13210789/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,264,1176,1200]" targetBox="[159,1427,189,1144]" targetPageId="21" targetType="figure">
<paragraph id="8BC736A13A467B46FF4EFB7C523EFB2E" blockId="21.[151,1436,1176,1236]" pageId="21" pageNumber="22">
<emphasis id="B90CEAB33A467B46FF4EFB7C5624FB4A" bold="true" box="[151,334,1176,1200]" pageId="21" pageNumber="22">FIGURES 27–29</emphasis>
. Male terminalia of 
<taxonomicName id="4C784D223A467B46FDFBFB7C558FFB4A" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[546,741,1176,1200]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="angusta" status="sp. nov.">
<emphasis id="B90CEAB33A467B46FDFBFB7C558FFB4A" box="[546,741,1176,1200]" italics="true" pageId="21" pageNumber="22">Scatophila angusta</emphasis>
</taxonomicName>
<emphasis id="B90CEAB33A467B46FD34FB7C5454FB4A" bold="true" box="[749,830,1176,1200]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="A23F57C83A467B46FD34FB7C5454FB4A" box="[749,830,1176,1200]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
27. Epandrium and cerci, posterior view. 28. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 29. Same, lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BC736A13A467B46FF1EFAC35209FA73" blockId="21.[151,1437,1318,2028]" pageId="21" pageNumber="22">
<emphasis id="B90CEAB33A467B46FF1EFAC3556BFABA" box="[199,513,1319,1345]" italics="true" pageId="21" pageNumber="22">
<materialsCitation id="3B103CFC3A467B46FF1EFAC35691FABA" box="[199,507,1319,1345]" collectorName="Benson" location="NORTH DAKOTA" pageId="21" pageNumber="22" specimenCount="1" stateProvince="North Dakota">
<collectingRegion id="49BCF8433A467B46FF1EFAC356F1FABB" box="[199,411,1319,1345]" country="United States of America" name="North Dakota" pageId="21" pageNumber="22">NORTH DAKOTA</collectingRegion>
. 
<collectorName id="268D53773A467B46FE7FFAC35691FABA" box="[422,507,1319,1344]" pageId="21" pageNumber="22">Benson</collectorName>
</materialsCitation>
:
</emphasis>
<materialsCitation id="3B103CFC3A467B46FDD3FAC252F3FABB" box="[522,1433,1318,1345]" collectingDate="1969-07-05" collectionCode="USNM" collectorName="W. W. Wirth" latitude="48.288334" location="Leeds" longLatPrecision="110" longitude="-99.43833" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Leeds">
<collectingRegion id="49BCF8433A467B46FDD3FAC25527FABA" box="[522,589,1318,1344]" country="United Kingdom" name="Leeds" pageId="21" pageNumber="22">Leeds</collectingRegion>
(
<geoCoordinate id="EE4C50663A467B46FD87FAC355B9FABB" box="[606,723,1319,1345]" degrees="48" direction="north" minutes="17.3" orientation="latitude" pageId="21" pageNumber="22" precision="92" value="48.288334">48°17.3'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FD39FAC2543CFABB" box="[736,854,1318,1345]" degrees="99" direction="west" minutes="26.3" orientation="longitude" pageId="21" pageNumber="22" precision="92" value="-99.43833">99°26.3'W</geoCoordinate>
; alkali lake), 
<date id="FFC610613A467B46FC2FFAC2531AFABA" box="[1014,1136,1318,1345]" pageId="21" pageNumber="22" value="1969-07-05">
<collectingDate id="EF82E9893A467B46FC2FFAC2531AFABA" box="[1014,1136,1318,1345]" pageId="21" pageNumber="22" value="1969-07-05">5 Jul 1969</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FBA4FAC35260FABA" box="[1149,1290,1318,1345]" pageId="21" pageNumber="22">W. W. Wirth</collectorName>
(1; 
<collectionCode id="ED69AE643A467B46FAE2FAC352FBFABB" box="[1339,1425,1319,1345]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B103CFC3A467B46FF4EFAAF565AFA73" collectingDate="2008-06-19" collectionCode="USNM" collectorName="D. and W. N. Mathis" county="Bowman" elevation="840" latitude="45.991665" location="Reservoir" longLatPrecision="112" longitude="-103.25667" municipality="Bowman-Haley Dam" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="Leeds">
<emphasis id="B90CEAB33A467B46FF4EFAAF5668FA9E" box="[151,258,1355,1380]" italics="true" pageId="21" pageNumber="22">
<collectingCounty id="62A64E2D3A467B46FF4EFAAF5796FA9E" box="[151,252,1355,1380]" pageId="21" pageNumber="22">Bowman</collectingCounty>
:
</emphasis>
<collectingMunicipality id="6BA3ACDB3A467B46FED3FAAE5692FA9F" box="[266,504,1354,1381]" pageId="21" pageNumber="22">Bowman-Haley Dam</collectingMunicipality>
and 
<location id="8EA7607A3A467B46FDF6FAAE55F6FA9E" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A467B46FDF6FAAE55F6FA9E" box="[559,668,1354,1380]" county="Bowman" latitude="45.991665" longLatPrecision="112" longitude="-103.25667" municipality="Bowman-Haley Dam" name="Reservoir" pageId="21" pageNumber="22" stateProvince="Leeds">Reservoir</location>
(
<geoCoordinate id="EE4C50663A467B46FD72FAAE544AFA9F" box="[683,800,1354,1381]" degrees="45" direction="north" minutes="59.5" orientation="latitude" pageId="21" pageNumber="22" precision="92" value="45.991665">45°59.5'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FCF5FAAE54DAFA9F" box="[812,944,1354,1381]" degrees="103" direction="west" minutes="15.4" orientation="longitude" pageId="21" pageNumber="22" precision="92" value="-103.25667">103°15.4'W</geoCoordinate>
; 
<quantity id="4C809B443A467B46FC66FAAF5363FA9F" box="[959,1033,1355,1381]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="21" pageNumber="22" unit="m" value="840.0">
<elevation id="0055D1923A467B46FC66FAAF5363FA9F" box="[959,1033,1355,1381]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="21" pageNumber="22" unit="m" value="840.0">840 m</elevation>
</quantity>
), 
<date id="FFC610613A467B46FBC7FAAF53C0FA9F" box="[1054,1194,1355,1381]" pageId="21" pageNumber="22" value="2008-06-19">
<collectingDate id="EF82E9893A467B46FBC7FAAF53C0FA9F" box="[1054,1194,1355,1381]" pageId="21" pageNumber="22" value="2008-06-19">19 Jun 2008</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FB6CFAAF52F6FA9E" box="[1205,1436,1354,1381]" pageId="21" pageNumber="22">D. and W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A467B46FF7AFA8B57AFFA73" box="[163,197,1391,1417]" count="1" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
; 
<collectionCode id="ED69AE643A467B46FF0BFA8B5642FA73" box="[210,296,1391,1417]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B103CFC3A467B46FEE2FA8A5235FA73" box="[315,1375,1390,1417]" collectingDate="1969-06-08" collectionCode="USNM" collectorName="W. W. Wirth" latitude="48.56333" location="Powers Lake" longLatPrecision="110" longitude="-102.64833" municipality="Mountrail" pageId="21" pageNumber="22" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Leeds">
<emphasis id="B90CEAB33A467B46FEE2FA8A56DFFA72" box="[315,437,1390,1416]" italics="true" pageId="21" pageNumber="22">
<collectingMunicipality id="6BA3ACDB3A467B46FEE2FA8A56DAFA72" box="[315,432,1390,1416]" pageId="21" pageNumber="22">Mountrail</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A467B46FE65FA8B5524FA72" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A467B46FE65FA8B5524FA72" box="[444,590,1390,1417]" latitude="48.56333" longLatPrecision="110" longitude="-102.64833" municipality="Mountrail" name="Powers Lake" pageId="21" pageNumber="22" stateProvince="Leeds">Powers Lake</location>
(
<geoCoordinate id="EE4C50663A467B46FD84FA8B55B8FA73" box="[605,722,1391,1417]" degrees="48" direction="north" minutes="33.8" orientation="latitude" pageId="21" pageNumber="22" precision="92" value="48.56333">48°33.8'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FD07FA8B540DFA73" box="[734,871,1391,1417]" degrees="102" direction="west" minutes="38.9" orientation="longitude" pageId="21" pageNumber="22" precision="92" value="-102.64833">102°38.9'W</geoCoordinate>
), 
<date id="FFC610613A467B46FCA2FA8B5492FA72" box="[891,1016,1390,1417]" pageId="21" pageNumber="22" value="1969-06-08">
<collectingDate id="EF82E9893A467B46FCA2FA8B5492FA72" box="[891,1016,1390,1417]" pageId="21" pageNumber="22" value="1969-06-08">8 Jun 1969</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FBDAFA8B53E4FA72" box="[1027,1166,1390,1417]" pageId="21" pageNumber="22">W. W. Wirth</collectorName>
(
<specimenCount id="9D7EFD283A467B46FB47FA8B53AEFA73" box="[1182,1220,1391,1417]" count="2" pageId="21" pageNumber="22" type="male">2♂</specimenCount>
, 
<specimenCount id="9D7EFD283A467B46FB16FA8B539EFA73" box="[1231,1268,1391,1417]" count="3" pageId="21" pageNumber="22" type="female">3♀</specimenCount>
; 
<collectionCode id="ED69AE643A467B46FAD8FA8B523DFA73" box="[1281,1367,1391,1417]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C362652A3A467B45FF1EFA7755CDFACD" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="8BC736A13A467B46FF1EFA775793FA2B" blockId="21.[151,1437,1318,2028]" pageId="21" pageNumber="22">
<materialsCitation id="3B103CFC3A467B46FF1EFA77579FFA2B" collectingDate="2005-08-03" collectionCode="USNM" collectorName="D. and W. N. Mathis" latitude="45.215" location="Cape Kiwanda" longLatPrecision="113" longitude="-123.971664" municipality="Tillamook" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="Oregon">
<emphasis id="B90CEAB33A467B46FF1EFA7756D3FA56" box="[199,441,1426,1453]" italics="true" pageId="21" pageNumber="22">
<collectingRegion id="49BCF8433A467B46FF1EFA775653FA57" box="[199,313,1427,1453]" country="United States of America" name="Oregon" pageId="21" pageNumber="22">OREGON</collectingRegion>
<collectingMunicipality id="6BA3ACDB3A467B46FE99FA7656D9FA56" box="[320,435,1426,1452]" pageId="21" pageNumber="22">Tillamook</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A467B46FE18FA77550DFA56" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A467B46FE18FA77550DFA56" box="[449,615,1426,1453]" latitude="45.215" longLatPrecision="113" longitude="-123.971664" municipality="Tillamook" name="Cape Kiwanda" pageId="21" pageNumber="22" stateProvince="Oregon">Cape Kiwanda</location>
(
<geoCoordinate id="EE4C50663A467B46FDAEFA765586FA57" box="[631,748,1426,1453]" degrees="45" direction="north" minutes="12.9" orientation="latitude" pageId="21" pageNumber="22" precision="92" value="45.215">45°12.9'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FD21FA7654EBFA57" box="[760,897,1426,1453]" degrees="123" direction="west" minutes="58.3" orientation="longitude" pageId="21" pageNumber="22" precision="92" value="-123.971664">123°58.3'W</geoCoordinate>
; beach), 
<date id="FFC610613A467B46FC3CFA775306FA57" box="[997,1132,1426,1453]" pageId="21" pageNumber="22" value="2005-08-03">
<collectingDate id="EF82E9893A467B46FC3CFA775306FA57" box="[997,1132,1426,1453]" pageId="21" pageNumber="22" value="2005-08-03">3 Aug 2005</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FBA1FA77520BFA56" box="[1144,1377,1426,1453]" pageId="21" pageNumber="22">D. and W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A467B46FAADFA7752FFFA57" box="[1396,1429,1427,1453]" count="1" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
; 
<collectionCode id="ED69AE643A467B46FF4EFA535787FA2B" box="[151,237,1463,1489]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A467B46FF1EFA3F5793F9C7" blockId="21.[151,1437,1318,2028]" pageId="21" pageNumber="22">
<materialsCitation id="3B103CFC3A467B46FF1EFA3F565CF9E3" collectingDate="2008-06-17" collectionCode="USNM" collectorName="D. and W. N. Mathis" elevation="1410" latitude="44.385" location="Deadwood" longLatPrecision="113" longitude="-103.68667" municipality="Lawrence" pageId="21" pageNumber="22" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="South Dakota">
<emphasis id="B90CEAB33A467B46FF1EFA3F557DFA0E" box="[199,535,1499,1525]" italics="true" pageId="21" pageNumber="22">
<collectingRegion id="49BCF8433A467B46FF1EFA3F56FDFA0F" box="[199,407,1499,1525]" country="United States of America" name="South Dakota" pageId="21" pageNumber="22">SOUTH DAKOTA</collectingRegion>
. 
<collectingMunicipality id="6BA3ACDB3A467B46FE79FA3F557BFA0E" box="[416,529,1499,1524]" pageId="21" pageNumber="22">Lawrence</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A467B46FDC4FA3E55FCFA0E" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A467B46FDC4FA3E55FCFA0E" box="[541,662,1498,1524]" latitude="44.385" longLatPrecision="113" longitude="-103.68667" municipality="Lawrence" name="Deadwood" pageId="21" pageNumber="22" stateProvince="South Dakota">Deadwood</location>
(
<geoCoordinate id="EE4C50663A467B46FD7CFA3F5470FA0F" box="[677,794,1499,1525]" degrees="44" direction="north" minutes="21.9" orientation="latitude" pageId="21" pageNumber="22" precision="92" value="44.365">44°21.9'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FCFCFA3F54C3FA0F" box="[805,937,1499,1525]" degrees="103" direction="west" minutes="43.9" orientation="longitude" pageId="21" pageNumber="22" precision="92" value="-103.73167">103°43.9'W</geoCoordinate>
; 
<quantity id="4C809B443A467B46FC61FA3F5365FA0F" box="[952,1039,1499,1525]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="21" pageNumber="22" unit="m" value="1410.0">
<elevation id="0055D1923A467B46FC61FA3F5365FA0F" box="[952,1039,1499,1525]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="21" pageNumber="22" unit="m" value="1410.0">1410 m</elevation>
</quantity>
), 
<date id="FFC610613A467B46FBFAFA3F53C7FA0F" box="[1059,1197,1499,1525]" pageId="21" pageNumber="22" value="2008-06-17">
<collectingDate id="EF82E9893A467B46FBFAFA3F53C7FA0F" box="[1059,1197,1499,1525]" pageId="21" pageNumber="22" value="2008-06-17">17 Jun 2008</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FB61FA3F52F6FA0E" box="[1208,1436,1498,1525]" pageId="21" pageNumber="22">D. and W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A467B46FF7AFA1B57AFF9E3" box="[163,197,1535,1561]" count="1" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
; 
<collectionCode id="ED69AE643A467B46FF0CFA1B5641F9E3" box="[213,299,1535,1561]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B103CFC3A467B46FE99FA1A579FF9C7" collectingDate="2008-06-17" collectionCode="USNM" collectorName="D. and W. N. Mathis" elevation="1410" latitude="44.385" location="Deadwood" longLatPrecision="113" longitude="-103.68667" municipality="Lawrence" pageId="21" pageNumber="22" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="South Dakota">
Deadwood (
<quantity id="4C809B443A467B46FE12FA1B5576F9E2" box="[459,540,1534,1561]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="21" pageNumber="22" unit="km" value="3.2">3.2 km</quantity>
E; 
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, 
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; 
<quantity id="4C809B443A467B46FC86FA1B54D0F9E3" box="[863,954,1535,1561]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="21" pageNumber="22" unit="m" value="1440.0">
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</quantity>
), 
<date id="FFC610613A467B46FC09FA1B530BF9E3" box="[976,1121,1535,1561]" pageId="21" pageNumber="22" value="2008-06-17">
<collectingDate id="EF82E9893A467B46FC09FA1B530BF9E3" box="[976,1121,1535,1561]" pageId="21" pageNumber="22" value="2008-06-17">17 Jun 2008</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FBB7FA1B5235F9E2" box="[1134,1375,1534,1561]" pageId="21" pageNumber="22">D. and W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A467B46FAABFA1B52FDF9E3" box="[1394,1431,1535,1561]" count="1" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
; 
<collectionCode id="ED69AE643A467B46FF4EF9C75787F9C7" box="[151,237,1571,1597]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A467B46FF1EF9A35357F937" blockId="21.[151,1437,1318,2028]" pageId="21" pageNumber="22">
<materialsCitation id="3B103CFC3A467B46FF1EF9A3560FF97F" collectingDate="2007-05-14" collectionCode="USNM" collectorName="D. and W. N. Mathis" elevation="1890" latitude="40.486668" location="Butterfield Canyon" longLatPrecision="116" longitude="-112.136665" municipality="Salt Lake" pageId="21" pageNumber="22" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Utah">
<emphasis id="B90CEAB33A467B46FF1EF9A356FBF99A" box="[199,401,1606,1633]" italics="true" pageId="21" pageNumber="22">
<collectingRegion id="49BCF8433A467B46FF1EF9A35664F99B" box="[199,270,1607,1633]" country="United States of America" name="Utah" pageId="21" pageNumber="22">UTAH</collectingRegion>
. 
<collectingMunicipality id="6BA3ACDB3A467B46FEC7F9A256E6F99A" box="[286,396,1606,1632]" pageId="21" pageNumber="22">Salt Lake</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A467B46FE43F9A25519F99B" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A467B46FE43F9A25519F99B" box="[410,627,1606,1633]" latitude="40.486668" longLatPrecision="116" longitude="-112.136665" municipality="Salt Lake" name="Butterfield Canyon" pageId="21" pageNumber="22" stateProvince="Utah">Butterfield Canyon</location>
(
<geoCoordinate id="EE4C50663A467B46FD5AF9A35593F99B" box="[643,761,1607,1633]" degrees="40" direction="north" minutes="29.2" orientation="latitude" pageId="21" pageNumber="22" precision="92" value="40.486668">40°29.2'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FCDCF9A354E2F99B" box="[773,904,1607,1633]" degrees="112" direction="west" minutes="08.2" orientation="longitude" pageId="21" pageNumber="22" precision="92" value="-112.136665">112°08.2'W</geoCoordinate>
; 
<quantity id="4C809B443A467B46FC40F9A35498F99B" box="[921,1010,1607,1633]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="21" pageNumber="22" unit="m" value="1890.0">
<elevation id="0055D1923A467B46FC40F9A35498F99B" box="[921,1010,1607,1633]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="21" pageNumber="22" unit="m" value="1890.0">1890 m</elevation>
</quantity>
), 
<date id="FFC610613A467B46FBD1F9A353C9F99B" box="[1032,1187,1607,1633]" pageId="21" pageNumber="22" value="2007-05-14">
<collectingDate id="EF82E9893A467B46FBD1F9A353C9F99B" box="[1032,1187,1607,1633]" pageId="21" pageNumber="22" value="2007-05-14">14 May 2007</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FB69F9A352F6F99A" box="[1200,1436,1606,1633]" pageId="21" pageNumber="22">D. and W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A467B46FF78F98F57ACF97F" box="[161,198,1643,1669]" count="3" pageId="21" pageNumber="22" type="male">3♂</specimenCount>
, 
<specimenCount id="9D7EFD283A467B46FF0AF98F579DF97F" box="[211,247,1643,1669]" count="1" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
; 
<collectionCode id="ED69AE643A467B46FEDEF98F5637F97F" box="[263,349,1643,1669]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B103CFC3A467B46FEA8F98E579FF953" collectingDate="1992-08-04" collectionCode="USNM" collectorName="D. and W. N. Mathis" latitude="39.933334" location="Goshen Warm Springs" longLatPrecision="928" longitude="-111.495" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="Utah">
<emphasis id="B90CEAB33A467B46FEA8F98E56D8F97E" box="[369,434,1642,1668]" italics="true" pageId="21" pageNumber="22">
<collectingRegion id="49BCF8433A467B46FEA8F98E56C6F97E" box="[369,428,1642,1668]" country="United States of America" name="Utah" pageId="21" pageNumber="22">Utah</collectingRegion>
:
</emphasis>
<location id="8EA7607A3A467B46FE62F98E55D1F97E" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A467B46FE62F98E55D1F97E" box="[443,699,1642,1669]" latitude="39.933334" longLatPrecision="928" longitude="-111.495" name="Goshen Warm Springs" pageId="21" pageNumber="22" stateProvince="Utah">Goshen Warm Springs</location>
(
<geoCoordinate id="EE4C50663A467B46FD12F98E5441F97F" box="[715,811,1642,1669]" degrees="39" direction="north" minutes="56" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="39.933334">39°56'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FCE0F98F54A8F97F" box="[825,962,1643,1669]" degrees="111" direction="west" minutes="29.7" orientation="longitude" pageId="21" pageNumber="22" precision="92" value="-111.495">111°29.7'W</geoCoordinate>
;), 
<date id="FFC610613A467B46FC05F98F530FF97F" box="[988,1125,1643,1669]" pageId="21" pageNumber="22" value="1992-08-04">
<collectingDate id="EF82E9893A467B46FC05F98F530FF97F" box="[988,1125,1643,1669]" pageId="21" pageNumber="22" value="1992-08-04">4 Aug 1992</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FBABF98F520AF97E" box="[1138,1376,1642,1669]" pageId="21" pageNumber="22">D. and W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A467B46FAADF98F52FFF97F" box="[1396,1429,1643,1669]" count="1" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
; 
<collectionCode id="ED69AE643A467B46FF4EF96B5787F953" box="[151,237,1679,1705]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B103CFC3A467B46FED8F96A5229F953" box="[257,1347,1678,1705]" collectingDate="2007-05-11" collectionCode="USNM" collectorName="D. and W. N. Mathis" elevation="1370" latitude="40.115" location="Lake Shore" longLatPrecision="116" longitude="-111.69666" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Utah">
<location id="8EA7607A3A467B46FED8F96A56E8F952" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A467B46FED8F96A56E8F952" box="[257,386,1678,1704]" latitude="40.115" longLatPrecision="116" longitude="-111.69666" name="Lake Shore" pageId="21" pageNumber="22" stateProvince="Utah">Lake Shore</location>
(
<geoCoordinate id="EE4C50663A467B46FE4BF96A556DF952" box="[402,519,1678,1704]" degrees="40" direction="north" minutes="06.9" orientation="latitude" pageId="21" pageNumber="22" precision="92" value="40.115">40°06.9'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FDCAF96B55F3F953" box="[531,665,1679,1705]" degrees="111" direction="west" minutes="41.8" orientation="longitude" pageId="21" pageNumber="22" precision="92" value="-111.69666">111°41.8'W</geoCoordinate>
; 
<quantity id="4C809B443A467B46FD7CF96B5594F953" box="[677,766,1679,1705]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="21" pageNumber="22" unit="m" value="1370.0">
<elevation id="0055D1923A467B46FD7CF96B5594F953" box="[677,766,1679,1705]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="21" pageNumber="22" unit="m" value="1370.0">1370 m</elevation>
</quantity>
), 
<date id="FFC610613A467B46FCCAF96B54C6F953" box="[787,940,1679,1705]" pageId="21" pageNumber="22" value="2007-05-11">
<collectingDate id="EF82E9893A467B46FCCAF96B54C6F953" box="[787,940,1679,1705]" pageId="21" pageNumber="22" value="2007-05-11">11 May 2007</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FC61F96B53C8F952" box="[952,1186,1678,1705]" pageId="21" pageNumber="22">D. and W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A467B46FB6DF96B53B2F953" box="[1204,1240,1679,1705]" count="1" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
; 
<collectionCode id="ED69AE643A467B46FB3CF96B5251F953" box="[1253,1339,1679,1705]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B103CFC3A467B46FA96F96A5353F937" collectingDate="2007-05-11" collectionCode="USNM" collectorName="D. and W. N. Mathis" elevation="1530" latitude="40.006668" location="Thistle" longLatPrecision="116" longitude="-111.495" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Utah">
<location id="8EA7607A3A467B46FA96F96A52F7F952" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A467B46FA96F96A52F7F952" box="[1359,1437,1678,1704]" latitude="40.006668" longLatPrecision="116" longitude="-111.495" name="Thistle" pageId="21" pageNumber="22" stateProvince="Utah">Thistle</location>
(
<geoCoordinate id="EE4C50663A467B46FF46F957566CF937" box="[159,262,1715,1741]" degrees="40" direction="north" minutes="0.4" orientation="latitude" pageId="21" pageNumber="22" precision="92" value="40.006668">40°0.4'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FECBF95756F2F937" box="[274,408,1715,1741]" degrees="111" direction="west" minutes="29.7" orientation="longitude" pageId="21" pageNumber="22" precision="92" value="-111.495">111°29.7'W</geoCoordinate>
; 
<quantity id="4C809B443A467B46FE7AF9565691F937" box="[419,507,1714,1741]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="21" pageNumber="22" unit="m" value="1530.0">
<elevation id="0055D1923A467B46FE7AF9565691F937" box="[419,507,1714,1741]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="21" pageNumber="22" unit="m" value="1530.0">1530 m</elevation>
</quantity>
), 
<date id="FFC610613A467B46FDD6F95755CDF937" box="[527,679,1715,1741]" pageId="21" pageNumber="22" value="2007-05-11">
<collectingDate id="EF82E9893A467B46FDD6F95755CDF937" box="[527,679,1715,1741]" pageId="21" pageNumber="22" value="2007-05-11">11 May 2007</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FD6BF95754F3F936" box="[690,921,1714,1741]" pageId="21" pageNumber="22">D. and W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A467B46FC73F95754A4F937" box="[938,974,1715,1741]" count="1" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
; 
<collectionCode id="ED69AE643A467B46FC02F957535BF937" box="[987,1073,1715,1741]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A467B46FF1EF933565EF8EF" blockId="21.[151,1437,1318,2028]" pageId="21" pageNumber="22">
<materialsCitation id="3B103CFC3A467B46FF1EF933565AF8EF" collectingDate="2007-10-29" collectionCode="USNM" collectorName="D. and W. N. Mathis" latitude="46.798332" location="Grayland" longLatPrecision="640" longitude="-124.1" municipality="Grays Harbor" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Washington">
<emphasis id="B90CEAB33A467B46FF1EF933555AF90A" box="[199,560,1750,1777]" italics="true" pageId="21" pageNumber="22">
<collectingRegion id="49BCF8433A467B46FF1EF9335610F90B" box="[199,378,1751,1777]" country="United States of America" name="Washington" pageId="21" pageNumber="22">WASHINGTON</collectingRegion>
. 
<collectingMunicipality id="6BA3ACDB3A467B46FE5FF9335540F90A" box="[390,554,1750,1777]" pageId="21" pageNumber="22">Grays Harbor</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A467B46FDE0F93255CBF90A" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A467B46FDE0F93255CBF90A" box="[569,673,1750,1776]" latitude="46.798332" longLatPrecision="640" longitude="-124.1" municipality="Grays Harbor" name="Grayland" pageId="21" pageNumber="22" stateProvince="Washington">Grayland</location>
(
<geoCoordinate id="EE4C50663A467B46FD6BF9325442F90A" box="[690,808,1750,1776]" degrees="46" direction="north" minutes="47.9" orientation="latitude" pageId="21" pageNumber="22" precision="92" value="46.798332">46°47.9'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FCECF93254CEF90B" box="[821,932,1750,1777]" degrees="124" direction="west" minutes="06" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="-124.1">124°06'W</geoCoordinate>
; beach), 
<date id="FFC610613A467B46FBC9F93353CBF90B" box="[1040,1185,1751,1777]" pageId="21" pageNumber="22" value="2007-10-29">
<collectingDate id="EF82E9893A467B46FBC9F93353CBF90B" box="[1040,1185,1751,1777]" pageId="21" pageNumber="22" value="2007-10-29">29 Oct 2007</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FB77F93352F6F90A" box="[1198,1436,1750,1777]" pageId="21" pageNumber="22">D. and W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A467B46FF78F91F57ACF8EF" box="[161,198,1787,1813]" count="1" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
; 
<collectionCode id="ED69AE643A467B46FF0BF91F5642F8EF" box="[210,296,1787,1813]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A467B46FF1EF8FB5793F8A7" blockId="21.[151,1437,1318,2028]" pageId="21" pageNumber="22">
<materialsCitation id="3B103CFC3A467B46FF1EF8FB579FF8A7" collectingDate="2008-06-20" collectionCode="USNM" collectorName="D. and W. N. Mathis" elevation="1330" latitude="44.643333" location="Alva" longLatPrecision="113" longitude="-104.38" municipality="Crook" pageId="21" pageNumber="22" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Wyoming">
<emphasis id="B90CEAB33A467B46FF1EF8FB56CCF8C2" box="[199,422,1822,1849]" italics="true" pageId="21" pageNumber="22">
<collectingRegion id="49BCF8433A467B46FF1EF8FB5627F8C3" box="[199,333,1823,1849]" country="United States of America" name="Wyoming" pageId="21" pageNumber="22">WYOMING</collectingRegion>
. 
<collectingMunicipality id="6BA3ACDB3A467B46FE81F8FA56CAF8C2" box="[344,416,1822,1848]" pageId="21" pageNumber="22">Crook</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A467B46FE75F8FA5688F8C2" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A467B46FE75F8FA5688F8C2" box="[428,482,1822,1848]" latitude="44.643333" longLatPrecision="113" longitude="-104.38" municipality="Crook" name="Alva" pageId="21" pageNumber="22" stateProvince="Wyoming">Alva</location>
(near; 
<geoCoordinate id="EE4C50663A467B46FDEBF8FA55CCF8C2" box="[562,678,1822,1848]" degrees="44" direction="north" minutes="38.6" orientation="latitude" pageId="21" pageNumber="22" precision="92" value="44.643333">44°38.6'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FD68F8FB5453F8C3" box="[689,825,1823,1849]" degrees="104" direction="west" minutes="22.8" orientation="longitude" pageId="21" pageNumber="22" precision="92" value="-104.38">104°22.8'W</geoCoordinate>
; 
<quantity id="4C809B443A467B46FC9CF8FB54F7F8C3" box="[837,925,1823,1849]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="21" pageNumber="22" unit="m" value="1330.0">
<elevation id="0055D1923A467B46FC9CF8FB54F7F8C3" box="[837,925,1823,1849]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="21" pageNumber="22" unit="m" value="1330.0">1330 m</elevation>
</quantity>
), 
<date id="FFC610613A467B46FC68F8FB5357F8C3" box="[945,1085,1823,1849]" pageId="21" pageNumber="22" value="2008-06-20">
<collectingDate id="EF82E9893A467B46FC68F8FB5357F8C3" box="[945,1085,1823,1849]" pageId="21" pageNumber="22" value="2008-06-20">20 Jun 2008</collectingDate>
</date>
, 
<collectorName id="268D53773A467B46FB91F8FB525AF8C2" box="[1096,1328,1822,1849]" pageId="21" pageNumber="22">D. and W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A467B46FA99F8FB520CF8C3" box="[1344,1382,1823,1849]" count="3" pageId="21" pageNumber="22" type="male">3♂</specimenCount>
, 
<specimenCount id="9D7EFD283A467B46FAA8F8FB52FCF8C3" box="[1393,1430,1823,1849]" count="2" pageId="21" pageNumber="22" type="female">2♀</specimenCount>
; 
<collectionCode id="ED69AE643A467B46FF4EF8A75787F8A7" box="[151,237,1859,1885]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A467B46FF1EF8835785F85F" blockId="21.[151,1437,1318,2028]" pageId="21" pageNumber="22">
<materialsCitation id="3B103CFC3A467B46FF1EF8835781F85F" country="United States of America" elevation="825" latitude="44.82" location="Goose Creek" longLatPrecision="92" longitude="-117.463165" municipality="Baker" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Oregon" typeStatus="holotype">
<typeStatus id="54C388033A467B46FF1EF8835795F87B" box="[199,255,1895,1921]" pageId="21" pageNumber="22">Type</typeStatus>
Locality.— 
<collectingCountry id="F36F76313A467B46FE5FF882554BF87B" box="[390,545,1894,1921]" name="United States of America" pageId="21" pageNumber="22">United States</collectingCountry>
. 
<collectingRegion id="49BCF8433A467B46FDF7F88355ECF87B" box="[558,646,1895,1921]" country="United States of America" name="Oregon" pageId="21" pageNumber="22">Oregon</collectingRegion>
. 
<collectingMunicipality id="6BA3ACDB3A467B46FD48F88255B2F87A" box="[657,728,1894,1920]" pageId="21" pageNumber="22">Baker</collectingMunicipality>
: 
<location id="8EA7607A3A467B46FD3CF8835412F87A" LSID="urn:lsid:plazi:treatment:03D187B73A407B45FF4EFDBA55CDFACD:8EA7607A3A467B46FD3CF8835412F87A" box="[741,888,1894,1921]" country="United States of America" latitude="44.82" longLatPrecision="92" longitude="-117.463165" municipality="Baker" name="Goose Creek" pageId="21" pageNumber="22" stateProvince="Oregon">Goose Creek</location>
(
<locationDeviation id="9A752F833A467B46FC50F882531FF87A" box="[905,1141,1894,1921]" location="Baker City" pageId="21" pageNumber="22">
<quantity id="4C809B443A467B46FC50F88254B9F87A" box="[905,979,1894,1921]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="22" unit="km" value="35.0">35 km</quantity>
E Baker City
</locationDeviation>
; 
<geoCoordinate id="EE4C50663A467B46FB5BF883539CF87B" box="[1154,1270,1895,1921]" degrees="44" direction="north" minutes="49.2" orientation="latitude" pageId="21" pageNumber="22" precision="92" value="44.82">44°49.2'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A467B46FADAF88352FEF87B" box="[1283,1428,1895,1921]" degrees="117" direction="west" minutes="27.79" orientation="longitude" pageId="21" pageNumber="22" precision="9" value="-117.463165">117°27.79'W</geoCoordinate>
; 
<quantity id="4C809B443A467B46FF4EF86E5788F85F" box="[151,226,1930,1957]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="21" pageNumber="22" unit="m" value="825.0">
<elevation id="0055D1923A467B46FF4EF86E5788F85F" box="[151,226,1930,1957]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="21" pageNumber="22" unit="m" value="825.0">825 m</elevation>
</quantity>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A467B46FF1EF84A56C3F816" blockId="21.[151,1437,1318,2028]" pageId="21" pageNumber="22">
Distribution (
<figureCitation id="13432A243A467B46FEBDF84A56D6F833" box="[356,444,1966,1993]" captionStart="FIGURE 30" captionStartId="22.[151,250,1023,1047]" captionTargetBox="[151,1435,181,999]" captionTargetId="figure-30@22.[151,1435,181,999]" captionTargetPageId="22" captionText="FIGURE 30. Distribution map of Scatophila angusta sp. nov. (circles), S. conifera Sturtevant &amp; Wheeler (triangle), and S. contaminata (Stenhammar) (squares) in Nearctic Region." figureDoi="http://doi.org/10.5281/zenodo.13210791" httpUri="https://zenodo.org/record/13210791/files/figure.png" pageId="21" pageNumber="22">Fig. 30</figureCitation>
).— 
<emphasis id="B90CEAB33A467B46FE31F84B553AF832" box="[488,592,1967,1992]" italics="true" pageId="21" pageNumber="22">
<taxonomicName id="4C784D223A467B46FE31F84B5520F832" box="[488,586,1967,1992]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
:
</emphasis>
<collectingCountry id="F36F76313A467B46FD87F84A5590F833" box="[606,762,1966,1993]" name="United States of America" pageId="21" pageNumber="22">United States</collectingCountry>
(
<collectingRegion id="49BCF8433A467B46FCC9F84A5417F832" box="[784,893,1966,1992]" country="United States of America" name="Colorado" pageId="21" pageNumber="22">Colorado</collectingRegion>
, 
<collectingRegion id="49BCF8433A467B46FC57F84A535BF832" box="[910,1073,1966,1992]" country="United States of America" name="North Dakota" pageId="21" pageNumber="22">North Dakota</collectingRegion>
, 
<collectingRegion id="49BCF8433A467B46FB9BF84B53F3F833" box="[1090,1177,1967,1993]" country="United States of America" name="Oregon" pageId="21" pageNumber="22">Oregon</collectingRegion>
, 
<collectingRegion id="49BCF8433A467B46FB72F84A5227F832" box="[1195,1357,1966,1992]" country="United States of America" name="South Dakota" pageId="21" pageNumber="22">South Dakota</collectingRegion>
, 
<collectingRegion id="49BCF8433A467B46FA86F84A52F2F832" box="[1375,1432,1966,1992]" country="United States of America" name="Utah" pageId="21" pageNumber="22">Utah</collectingRegion>
, 
<collectingRegion id="49BCF8433A467B46FF4EF8365675F816" box="[151,287,2002,2028]" country="United States of America" name="Washington" pageId="21" pageNumber="22">Washington</collectingRegion>
, 
<collectingRegion id="49BCF8433A467B46FEF3F83656F4F816" box="[298,414,2002,2028]" country="United States of America" name="Wyoming" pageId="21" pageNumber="22">Wyoming</collectingRegion>
).
</paragraph>
<caption id="DF0766293A457B45FF4EFC1B55B7FBC1" ID-DOI="http://doi.org/10.5281/zenodo.13210791" ID-Zenodo-Dep="13210791" httpUri="https://zenodo.org/record/13210791/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1023,1047]" targetBox="[151,1435,181,999]" targetPageId="22" targetType="figure">
<paragraph id="8BC736A13A457B45FF4EFC1B55B7FBC1" blockId="22.[151,1437,1023,1083]" pageId="22" pageNumber="23">
<emphasis id="B90CEAB33A457B45FF4EFC1B5671FBED" bold="true" box="[151,283,1023,1047]" pageId="22" pageNumber="23">FIGURE 30</emphasis>
. Distribution map of 
<taxonomicName id="4C784D223A457B45FDD9FC1B55AFFBED" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[512,709,1023,1047]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="angusta" status="sp. nov.">
<emphasis id="B90CEAB33A457B45FDD9FC1B55AFFBED" box="[512,709,1023,1047]" italics="true" pageId="22" pageNumber="23">Scatophila angusta</emphasis>
</taxonomicName>
<emphasis id="B90CEAB33A457B45FD14FC1B544AFBED" bold="true" box="[717,800,1023,1047]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A23F57C83A457B45FD14FC1B544AFBED" box="[717,800,1023,1047]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(circles), 
<taxonomicName id="4C784D223A457B45FC52FC1B538CFBED" authority="Sturtevant &amp; Wheeler" authorityName="Sturtevant &amp; Wheeler" box="[907,1254,1023,1047]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="conifera">
<emphasis id="B90CEAB33A457B45FC52FC1B5491FBED" box="[907,1019,1023,1047]" italics="true" pageId="22" pageNumber="23">S. conifera</emphasis>
Sturtevant &amp; Wheeler
</taxonomicName>
(triangle), and 
<taxonomicName id="4C784D223A457B45FA50FC1B56C7FBC1" authority="(Stenhammar)" baseAuthorityName="Stenhammar" baseAuthorityYear="1844" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="contaminata">
<emphasis id="B90CEAB33A457B45FA50FC1B567CFBC1" italics="true" pageId="22" pageNumber="23">S. contaminata</emphasis>
(Stenhammar)
</taxonomicName>
(squares) in 
<taxonomicName id="4C784D223A457B45FDE8FBC755EDFBC1" box="[561,647,1059,1083]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="97" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
Region.
</paragraph>
</caption>
<paragraph id="8BC736A13A457B45FF1EFB695478FB31" blockId="22.[151,1436,1165,1335]" pageId="22" pageNumber="23">
Etymology.—The species epithet, 
<emphasis id="B90CEAB33A457B45FD88FB6A55DAFB5D" box="[593,688,1166,1191]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="4C784D223A457B45FD88FB6A55C6FB5D" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[593,684,1166,1191]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
,
</emphasis>
is of Latin derivation and means narrow, referring to the narrow bifurcation of the ventral portion of the male epandrium.
</paragraph>
<paragraph id="8BC736A13A457B45FF1EFB3155CDFACD" blockId="22.[151,1436,1165,1335]" pageId="22" pageNumber="23">
Remarks.—This species is placed in the 
<taxonomicName id="4C784D223A457B45FD49FB315598FB15" baseAuthorityName="Haliday" baseAuthorityYear="1839" box="[656,754,1237,1263]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="despecta">
<emphasis id="B90CEAB33A457B45FD49FB315598FB15" box="[656,754,1237,1263]" italics="true" pageId="22" pageNumber="23">despecta</emphasis>
</taxonomicName>
group where it is similar and apparently closely related to 
<emphasis id="B90CEAB33A457B45FA5EFB325660FAE9" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="4C784D223A457B45FA5EFB32566CFAE9" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="variabilis">S. variabilis</taxonomicName>
.
</emphasis>
It differs from the latter species primarily by the narrower ventral half of the epandrium, especially the shallower bifurcation along the ventral margin.
</paragraph>
</subSubSection>
</treatment>
</document>