469 lines
95 KiB
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469 lines
95 KiB
XML
<document id="C4BCB2BB3929A9D43B164218F3C1C997" ID-DOI="10.11646/zootaxa.5476.1.29" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681921" ID-ZooBank="DD4EA1FE-A0BE-4BA4-9AD3-E460B131C518" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="felipe" checkinTime="1720375669801" checkinUser="plazi" docAuthor="Rahayu, Dwi Listyo & Ng, Peter K. L." docDate="2024" docId="03DD87DAFFB8124E28EEFBBC87BE1EA1" docLanguage="en" docName="zootaxa.5476.1.29.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.29" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyphira acutidens" docType="treatment" docVersion="5" lastPageNumber="368" masterDocId="FFE4FFA2FFBE12442879FFAF843C1F54" masterDocTitle="On the genus Lyphira Galil, 2009 (Crustacea: Brachyura: Leucosiidae), with descriptions of four new species from the West Pacific" masterLastPageNumber="392" masterPageNumber="358" pageNumber="364" updateTime="1721070200110" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="4710E450AFE0E9CDC454A646FAE27824">On the genus Lyphira Galil, 2009 (Crustacea: Brachyura: Leucosiidae), with descriptions of four new species from the West Pacific</mods:title>
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<mods:namePart id="847460BFBEFA0493ED1717EF41467663">Rahayu, Dwi Listyo</mods:namePart>
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<mods:affiliation id="EBD2DF1F9337742F07A03C27A47AB09E">Research Center for Marine and Land Bioindustry, National Research and Innovation Agency, Dusun Teluk Kodek, Pemenang, Lombok Utara 83756, NTB, Indonesia.</mods:affiliation>
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<mods:namePart id="E4B7A2EB9923E17EA09C90F8AD5E0557">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="DD4A5F85CF9B38D16B7027AC0D5B4E43">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore.</mods:affiliation>
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<mods:date id="12593ECAD4E07FEEFD142BA388C8CE63">2024</mods:date>
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<heading id="D08381A0FFB8124228EEFBBC86241B7A" bold="true" box="[151,536,1043,1070]" fontSize="11" level="1" pageId="6" pageNumber="364" reason="1">
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<taxonomicName id="4C744D4FFFB8124228EEFBBC86241B7A" ID-CoL="4GDTY" authority="(Chen, 1987)" baseAuthorityName="Chen" baseAuthorityYear="1987" box="[151,536,1043,1070]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="acutidens">
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<emphasis id="B900EADEFFB8124228EEFBBC86241B7A" bold="true" box="[151,536,1043,1070]" pageId="6" pageNumber="364">
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<emphasis id="B900EADEFFB8124228EEFBBC854E1B7A" bold="true" box="[151,370,1043,1070]" italics="true" pageId="6" pageNumber="364">Lyphira acutidens</emphasis>
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(
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<bibRefCitation id="EFE54B3DFFB8124229F8FBBB862C1B7A" author="Chen, H." box="[385,528,1044,1070]" pageId="6" pageNumber="364" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen, 1987</bibRefCitation>
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<paragraph id="8BCB36CCFFB8124228EEFB9787831B06" blockId="6.[151,959,1043,1107]" box="[151,959,1080,1107]" pageId="6" pageNumber="364">
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(
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<figureCitation id="134F2A49FFB8124228E6FB97853F1B06" box="[159,259,1080,1106]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Figs. 3D</figureCitation>
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,
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<figureCitation id="134F2A49FFB812422976FB97850E1B06" box="[271,306,1080,1106]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="6" pageNumber="364">6B</figureCitation>
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<figureCitation id="134F2A49FFB812422947FB96855E1B07" box="[318,354,1081,1107]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="6" pageNumber="364">7D</figureCitation>
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<figureCitation id="134F2A49FFB812422917FB9785AD1B06" box="[366,401,1080,1106]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="6" pageNumber="364">9B</figureCitation>
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<figureCitation id="134F2A49FFB8124229E4FB9785F11B06" box="[413,461,1080,1106]" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="6" pageNumber="364">10C</figureCitation>
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<figureCitation id="134F2A49FFB8124229A0FB9786351B06" box="[473,521,1080,1106]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="6" pageNumber="364">11C</figureCitation>
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<figureCitation id="134F2A49FFB812422A6CFB97865B1B06" box="[533,615,1080,1106]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="6" pageNumber="364">13B, C</figureCitation>
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<figureCitation id="134F2A49FFB812422A0BFB9786981B06" box="[626,676,1080,1106]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="6" pageNumber="364">14B</figureCitation>
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<figureCitation id="134F2A49FFB812422AD6FB9786DE1B07" box="[687,738,1080,1107]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="6" pageNumber="364">15D</figureCitation>
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<figureCitation id="134F2A49FFB812422A94FB9787221B06" box="[749,798,1080,1106]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="6" pageNumber="364">16C</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB812422B53FB9787471B06" box="[810,891,1080,1106]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="6" pageNumber="364">17H–L</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB812422BFEFB97878B1B06" box="[903,951,1080,1106]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="6" pageNumber="364">21B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6547FFB8124228EEFBD880181BFA" pageId="6" pageNumber="364" type="reference_group">
|
||
<paragraph id="8BCB36CCFFB8124228EEFBD880E31BDB" blockId="6.[151,1247,1143,1198]" box="[151,1247,1143,1168]" pageId="6" pageNumber="364">
|
||
<treatmentCitationGroup id="AB6411E2FFB8124228EEFBD880E31BDB" box="[151,1247,1143,1168]" pageId="6" pageNumber="364">
|
||
<treatmentCitation id="0AD510DDFFB8124228EEFBD885C71BDB" author="Chen, H." box="[151,507,1143,1167]" page="201" pageId="6" pageNumber="364" year="1987">
|
||
<taxonomicName id="4C744D4FFFB8124228EEFBD885C71BDB" authority="Chen, 1987: 201" authorityName="Chen" authorityPageNumber="201" authorityYear="1987" box="[151,507,1143,1167]" class="Malacostraca" family="Leucosiidae" genus="Phylira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="acutidens">
|
||
<emphasis id="B900EADEFFB8124228EEFBD885771BDB" box="[151,331,1143,1167]" italics="true" pageId="6" pageNumber="364">Phylira acutidens</emphasis>
|
||
<bibRefCitation id="EFE54B3DFFB812422928FBD885C71BDB" author="Chen, H." box="[337,507,1143,1167]" pageId="6" pageNumber="364" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen, 1987: 201</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, fig. 1;
|
||
<treatmentCitation id="0AD510DDFFB812422A30FBD887081BDB" author="Chen, H. & Sun, H." box="[585,820,1143,1167]" page="381" pageId="6" pageNumber="364" year="2002">
|
||
<bibRefCitation id="EFE54B3DFFB812422A30FBD887081BDB" author="Chen, H. & Sun, H." box="[585,820,1143,1167]" pageId="6" pageNumber="364" refId="ref19489" refString="Chen, H. & Sun, H. (2002) Brachyura marine primitive crabs. Fauna Sinica Invertebrata Arthropoda Crustacea. Vol. 30. Science Press, Beijing, xiii + 597 pp., 237 figs., 6 pls." type="book" year="2002">Chen & Sun 2002: 381</bibRefCitation>
|
||
</treatmentCitation>
|
||
, text fig. 170, pl. 15.1;
|
||
<treatmentCitation id="0AD510DDFFB812422C58FBD780E71BDB" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[1057,1243,1143,1168]" page="93" pageId="6" pageNumber="364" year="2008">
|
||
<bibRefCitation id="EFE54B3DFFB812422C58FBD780E71BDB" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[1057,1243,1143,1168]" pageId="6" pageNumber="364" pagination="1 - 286" refId="ref21130" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
|
||
Ng
|
||
<emphasis id="B900EADEFFB812422C3FFBD780451BDB" box="[1094,1145,1143,1167]" italics="true" pageId="6" pageNumber="364">et al.</emphasis>
|
||
2008: 93
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCB36CCFFB8124228EEFB3980181BFA" blockId="6.[151,1247,1143,1198]" box="[151,1060,1174,1198]" pageId="6" pageNumber="364">
|
||
<treatmentCitationGroup id="AB6411E2FFB8124228EEFB3980181BFA" box="[151,1060,1174,1198]" pageId="6" pageNumber="364">
|
||
<emphasis id="B900EADEFFB8124228EEFB3985BA1BFA" box="[151,390,1174,1198]" italics="true" pageId="6" pageNumber="364">
|
||
<taxonomicName id="4C744D4FFFB8124228EEFB3985541BFA" ID-CoL="3WTBK" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,360,1174,1198]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="heterograna">Lyphira heterograna</taxonomicName>
|
||
.—
|
||
</emphasis>
|
||
<treatmentCitation id="0AD510DDFFB8124229FFFB3986101BFA" author="Galil, B. S." box="[390,556,1174,1198]" httpUri="http://treatment.plazi.org/id/039D87BCFFBF4A26FD4BC4ADFDE1ECDD" page="300" pageId="6" pageNumber="364" year="2009">
|
||
<bibRefCitation id="EFE54B3DFFB8124229FFFB3986101BFA" author="Galil, B. S." box="[390,556,1174,1198]" pageId="6" pageNumber="364" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil, 2009: 300</bibRefCitation>
|
||
</treatmentCitation>
|
||
(part). (not
|
||
<taxonomicName id="4C744D4FFFB812422ADFFB3980251BFA" ID-CoL="76YMK" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[678,1049,1174,1198]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB812422ADFFB3987481BFA" box="[678,884,1174,1198]" italics="true" pageId="6" pageNumber="364">Philyra heterograna</emphasis>
|
||
<bibRefCitation id="EFE54B3DFFB812422B02FB3980251BFA" author="Ortmann, A." box="[891,1049,1174,1198]" pageId="6" pageNumber="364" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||
</taxonomicName>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6547FFB8124228EEFB79801F1A41" pageId="6" pageNumber="364" type="materials_examined">
|
||
<paragraph id="8BCB36CCFFB8124228EEFB79801F1A41" blockId="6.[151,1437,1238,2021]" pageId="6" pageNumber="364">
|
||
<emphasis id="B900EADEFFB8124228EEFB7985451BA4" bold="true" box="[151,377,1238,1264]" pageId="6" pageNumber="364">Material examined</emphasis>
|
||
.
|
||
<specimenCount id="9D72FD45FFB8124229FFFB7885EC1BA5" box="[390,464,1239,1265]" count="1" pageId="6" pageNumber="364" type="male">1 male</specimenCount>
|
||
(8.9 ×
|
||
<quantity id="4C8C9B29FFB812422A66FB7886441BA5" box="[543,632,1239,1265]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="6" pageNumber="364" unit="mm" value="8.6">8.6 mm</quantity>
|
||
),
|
||
<materialsCitation id="3B1C3C91FFB812422AF2FB7880231A41" collectingDate="1958-12-09" collectionCode="ZRC" location="Institute of Oceanography" municipality="Chinese Academy of Science" pageId="6" pageNumber="364" specimenCode="ZRC 2002.0504" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="9D72FD45FFB812422AF2FB7886D71BA5" box="[651,747,1239,1265]" count="1" pageId="6" pageNumber="364" type="female">1 female</specimenCount>
|
||
(13.4 ×
|
||
<quantity id="4C8C9B29FFB812422B31FB7887931BA5" box="[840,943,1239,1265]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="6" pageNumber="364" unit="mm" value="12.9">12.9 mm</quantity>
|
||
),
|
||
<specimenCode id="DBD29EB7FFB812422BB8FB7880441BA5" box="[961,1144,1238,1265]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="364">ZRC 2002.0504</specimenCode>
|
||
, station 4027, Yellow Sea, coll.
|
||
<location id="8EAB6017FFB8124228B6FB5485C81A41" LSID="urn:lsid:plazi:treatment:03DD87DAFFB8124E28EEFBBC87BE1EA1:8EAB6017FFB8124228B6FB5485C81A41" box="[207,500,1275,1301]" municipality="Chinese Academy of Science" name="Institute of Oceanography" pageId="6" pageNumber="364">Institute of Oceanography</location>
|
||
,
|
||
<collectingMunicipality id="6BAFACB6FFB812422A79FB5487761A41" box="[512,842,1275,1301]" pageId="6" pageNumber="364">Chinese Academy of Science</collectingMunicipality>
|
||
,
|
||
<date id="FFCA100CFFB812422B2CFB5480231A41" box="[853,1055,1274,1301]" pageId="6" pageNumber="364" value="1958-12-09">
|
||
<collectingDate id="EF8EE9E4FFB812422B2CFB5480231A41" box="[853,1055,1274,1301]" pageId="6" pageNumber="364" value="1958-12-09">9 December 1958</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6547FFB8124328BEFAB185921FAD" lastPageId="7" lastPageNumber="365" pageId="6" pageNumber="364" type="diagnosis">
|
||
<paragraph id="8BCB36CCFFB8124328BEFAB186591F81" blockId="6.[151,1437,1238,2021]" lastBlockId="7.[151,1436,151,249]" lastPageId="7" lastPageNumber="365" pageId="6" pageNumber="364">
|
||
<emphasis id="B900EADEFFB8124228BEFAB185061A6C" bold="true" box="[199,314,1310,1336]" pageId="6" pageNumber="364">Diagnosis</emphasis>
|
||
. Carapace slightly broader than long, dorsal surface of carapace covered with closely spaced minute granules, hepatic and intestinal regions with larger, more prominent granules (
|
||
<figureCitation id="134F2A49FFB812422C41FAEC809D1A09" box="[1080,1185,1347,1373]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Figs. 3D</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB812422CCAFAEC80EB1A09" box="[1203,1239,1347,1373]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="6" pageNumber="364">6B</figureCitation>
|
||
). Frontal margin granulated, postfrontal lobe slightly convex (
|
||
<figureCitation id="134F2A49FFB812422AE3FAC887371AD5" box="[666,779,1383,1409]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="6" pageNumber="364">Figs. 11C</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB812422B63FAC887531AD5" box="[794,879,1383,1409]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="6" pageNumber="364">13B, C</figureCitation>
|
||
). Endostomial spine absent (
|
||
<figureCitation id="134F2A49FFB812422CC0FAC881731AD5" box="[1209,1359,1383,1409]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="6" pageNumber="364">Figs. 13B, C</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB812422D24FAC881B31AD5" box="[1373,1423,1383,1409]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="6" pageNumber="364">14B</figureCitation>
|
||
). Lateral and posterolateral margin of carapace with row of small granules interspersed by several larger granules (
|
||
<figureCitation id="134F2A49FFB8124228E6FA0084C71A9D" box="[159,251,1455,1481]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Fig. 3D</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB812422970FA0085111A9D" box="[265,301,1455,1481]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="6" pageNumber="364">6B</figureCitation>
|
||
); posterior carapace margin gently concave with row of larger granule, medially slightly produced (
|
||
<figureCitation id="134F2A49FFB8124228E6FA7C85391AB9" box="[159,261,1491,1517]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Figs. 3D</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB81242296BFA7C85791AB9" box="[274,325,1491,1517]" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="6" pageNumber="364">10C</figureCitation>
|
||
). Third maxilliped exopod 2.9 times as long as broad, as broad as basal part of endopod, forming petaliform structure with convex outer margin, endopod with completely fused basis-ischium, as long as merus, with shallow submarginal sulcus near inner margin (
|
||
<figureCitation id="134F2A49FFB812422A9AF9B4876E1961" box="[739,850,1563,1589]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="6" pageNumber="364">Figs. 11C</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB812422B27F9B487AD1961" box="[862,913,1563,1589]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="6" pageNumber="364">17H</figureCitation>
|
||
). Chelipeds subequal; merus, carpus and chela surfaces smooth or covered by closely spaced, small rounded granules (
|
||
<figureCitation id="134F2A49FFB812422BB1F9908011190D" box="[968,1069,1599,1625]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Figs. 3D</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB812422C42F9908062190D" box="[1083,1118,1599,1625]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="6" pageNumber="364">6B</figureCitation>
|
||
); chela stout, fingers longer than length of palm, surface with longitudinal sulcus flanked by row of granules; pollex slightly bent, cutting edge lined with denticles; dactylus 1.2 times as long as palm along upper margin, row of small granules on upper margin, cutting edge with median large tooth lined with denticles, remaining of edge with small denticles (
|
||
<figureCitation id="134F2A49FFB812422D5AF90481B31991" box="[1315,1423,1706,1733]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="6" pageNumber="364">Fig. 15D</figureCitation>
|
||
). P2–P5 slender, short; merus distinctly longer than carpus and propodus; dactylus longer than propodus, lanceolate, terminating in corneous tips (
|
||
<figureCitation id="134F2A49FFB8124229A4F95C867C1859" box="[477,576,1779,1805]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="6" pageNumber="364">Figs. 3D</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB812422A32F95C86531859" box="[587,623,1779,1805]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="6" pageNumber="364">6B</figureCitation>
|
||
); fourth leg shortest with merus 4.1 times as long as broad, margins lined with small granules, more prominent on ventral margin (
|
||
<figureCitation id="134F2A49FFB812422B6FF8B887411865" box="[790,893,1815,1841]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="6" pageNumber="364">Fig. 16C</figureCitation>
|
||
). Thoracic sternites transversely narrow, surface finely granulated, larger granules laterally (
|
||
<figureCitation id="134F2A49FFB812422AE8F89486D31801" box="[657,751,1851,1877]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="6" pageNumber="364">Fig. 7D</figureCitation>
|
||
). Sternopleonal cavity deep, reaching to proximal distance between fused thoracic sternites 1–3. Pleon narrow, somites 2–6 fused, somite 6 longitudinally subrectangular, surface with one low granule medially, lateral margins gently convex; telson triangular (
|
||
<figureCitation id="134F2A49FFB812422C1CF82C80F418C9" box="[1125,1224,1923,1949]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="6" pageNumber="364">Figs. 7D</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB812422CAAF82C80C018C9" box="[1235,1276,1923,1949]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="6" pageNumber="364">17I</figureCitation>
|
||
). G1 elongate, slender, same size from proximal to distal area; distal process bilobed, tip rounded (
|
||
<figureCitation id="134F2A49FFB812422C10F80880D71895" box="[1129,1259,1959,1985]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="6" pageNumber="364">Fig. 17J‒L</figureCitation>
|
||
). Female pleon broad, convex, smooth, somites 2‒6 fused, suture between somites 2 and 3; telson as long as its basal width (
|
||
<figureCitation id="134F2A49FFB812432D16F86484861FE5" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" lastPageId="7" lastPageNumber="365" pageId="6" pageNumber="364">Fig. 9B</figureCitation>
|
||
); vulvae positioned closer to median part of sternite 6, each opening prominently crescent-shaped, relatively long, without opercular cover (
|
||
<figureCitation id="134F2A49FFB91243298BFF1486641F81" box="[498,600,187,213]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="7" pageNumber="365">Fig. 21B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCB36CCFFB9124328BEFF7185921FAD" blockId="7.[151,1436,151,249]" box="[199,430,222,249]" pageId="7" pageNumber="365">
|
||
<emphasis id="B900EADEFFB9124328BEFF7185271FAC" bold="true" box="[199,283,222,248]" pageId="7" pageNumber="365">Colour</emphasis>
|
||
. Not known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0B6644FFB9124328EEF8B5864318B2" ID-DOI="http://doi.org/10.5281/zenodo.12681932" ID-Zenodo-Dep="12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="7" pageNumber="365" startId="7.[151,250,1818,1842]" targetBox="[151,1435,273,1794]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8BCB36CCFFB9124328EEF8B5864318B2" blockId="7.[151,1436,1818,2022]" pageId="7" pageNumber="365">
|
||
<emphasis id="B900EADEFFB9124328EEF8B585291866" bold="true" box="[151,277,1818,1842]" pageId="7" pageNumber="365">FIGURE 3.</emphasis>
|
||
Overall dorsal habitus. A,
|
||
<taxonomicName id="4C744D4FFFB912432A56F8B587861866" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[559,954,1818,1842]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB912432A56F8B5873F1866" box="[559,771,1818,1842]" italics="true" pageId="7" pageNumber="365">Lyphira heterograna</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE54B3DFFB912432B6BF8B5878F1866" author="Ortmann, A." box="[786,947,1818,1842]" pageId="7" pageNumber="365" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B,
|
||
<taxonomicName id="4C744D4FFFB912432885F89086791802" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[252,581,1854,1878]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB912432885F89085AF1802" box="[252,403,1854,1878]" italics="true" pageId="7" pageNumber="365">L. heterograna</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE54B3DFFB9124329D9F89186021802" author="Ortmann, A." box="[416,574,1854,1878]" pageId="7" pageNumber="365" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of
|
||
<taxonomicName id="4C744D4FFFB912432CFCF89184F2182E" authority="Shen, 1932" authorityName="Shen" authorityYear="1932" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="peitaihoensis">
|
||
<emphasis id="B900EADEFFB912432CFCF89181611802" box="[1157,1373,1854,1878]" italics="true" pageId="7" pageNumber="365">Philyra peitaihoensis</emphasis>
|
||
<bibRefCitation id="EFE54B3DFFB912432D1AF89184F2182E" author="Shen, C. - J." pageId="7" pageNumber="365" pagination="1 - 300" refId="ref21817" refString="Shen, C. - J. (1932) The brachyuran Crustacea of North China. Zoologia Sinica, 9 (1), 1 - 300, pls. 1 - 10." type="journal article" year="1932">Shen, 1932</bibRefCitation>
|
||
</taxonomicName>
|
||
], China; C,
|
||
<taxonomicName id="4C744D4FFFB912432938F8CC86B8182E" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[321,644,1890,1914]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB912432938F8CC85E9182E" box="[321,469,1890,1914]" italics="true" pageId="7" pageNumber="365">L. heterograna</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE54B3DFFB912432998F8CD8641182E" author="Ortmann, A." box="[481,637,1890,1914]" pageId="7" pageNumber="365" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D,
|
||
<taxonomicName id="4C744D4FFFB912432CEDF8CC81A5182E" authority="(Chen, 1987)" baseAuthorityName="Chen" baseAuthorityYear="1987" box="[1172,1433,1890,1914]" class="Malacostraca" family="Leucosiidae" genus="Phylira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="acutidens">
|
||
<emphasis id="B900EADEFFB912432CEDF8CC8132182E" box="[1172,1294,1890,1914]" italics="true" pageId="7" pageNumber="365">L. acutidens</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE54B3DFFB912432D60F8CD81AD182E" author="Chen, H." box="[1305,1425,1890,1914]" pageId="7" pageNumber="365" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen, 1987</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E,
|
||
<taxonomicName id="4C744D4FFFB912432AAFF828870718CA" authorityName="Rahayu & Ng" authorityYear="2024" box="[726,827,1926,1950]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||
<emphasis id="B900EADEFFB912432AAFF828870718CA" box="[726,827,1926,1950]" italics="true" pageId="7" pageNumber="365">L. chomel</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EADEFFB912432B3BF829874118CA" bold="true" box="[834,893,1926,1950]" pageId="7" pageNumber="365">
|
||
<taxonomicNameLabel id="A23357A5FFB912432B3BF829874118CA" box="[834,893,1926,1950]" pageId="7" pageNumber="365" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F,
|
||
<taxonomicName id="4C744D4FFFB912432962F80485431896" authorityName="Rahayu & Ng" authorityYear="2024" box="[283,383,1962,1986]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||
<emphasis id="B900EADEFFB912432962F80485431896" box="[283,383,1962,1986]" italics="true" pageId="7" pageNumber="365">L. chomel</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EADEFFB9124329FEF80585FC1896" bold="true" box="[391,448,1962,1986]" pageId="7" pageNumber="365">
|
||
<taxonomicNameLabel id="A23357A5FFB9124329FEF80585FC1896" box="[391,448,1962,1986]" pageId="7" pageNumber="365" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G,
|
||
<taxonomicName id="4C744D4FFFB912432C22F80480831896" authorityName="Rahayu & Ng" authorityYear="2024" box="[1115,1215,1962,1986]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="chomel" status="n. sp.">
|
||
<emphasis id="B900EADEFFB912432C22F80480831896" box="[1115,1215,1962,1986]" italics="true" pageId="7" pageNumber="365">L. chomel</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EADEFFB912432CBFF805813C1896" bold="true" box="[1222,1280,1962,1986]" pageId="7" pageNumber="365">
|
||
<taxonomicNameLabel id="A23357A5FFB912432CBFF805813C1896" box="[1222,1280,1962,1986]" pageId="7" pageNumber="365" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0B6644FFB6124C28EEF8BE854418C1" ID-DOI="http://doi.org/10.5281/zenodo.12681934" ID-Zenodo-Dep="12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="8" pageNumber="366" startId="8.[151,250,1809,1833]" targetBox="[151,1435,181,1785]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8BCB36CCFFB6124C28EEF8BE854418C1" blockId="8.[151,1437,1809,1941]" pageId="8" pageNumber="366">
|
||
<emphasis id="B900EADEFFB6124C28EEF8BE8528187D" bold="true" box="[151,276,1809,1833]" pageId="8" pageNumber="366">FIGURE 4.</emphasis>
|
||
Overall dorsal habitus,
|
||
<taxonomicName id="4C744D4FFFB6124C2A70F8BE86A8187D" authorityName="Rahayu & Ng" authorityYear="2024" box="[521,660,1809,1833]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="366" phylum="Arthropoda" rank="species" species="linda" status="sp. nov.">
|
||
<emphasis id="B900EADEFFB6124C2A70F8BE86A8187D" box="[521,660,1809,1833]" italics="true" pageId="8" pageNumber="366">Lyphira linda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EADEFFB6124C2AE2F8BE86E9187D" bold="true" box="[667,725,1809,1833]" pageId="8" pageNumber="366">
|
||
<taxonomicNameLabel id="A23357A5FFB6124C2AE2F8BE86E9187D" box="[667,725,1809,1833]" pageId="8" pageNumber="366" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0B6644FFB7124D28EEFB7185701A36" ID-DOI="http://doi.org/10.5281/zenodo.12681936" ID-Zenodo-Dep="12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="9" pageNumber="367" startId="9.[151,250,1246,1270]" targetBox="[151,1435,181,1222]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8BCB36CCFFB7124D28EEFB7185701A36" blockId="9.[151,1437,1246,1378]" pageId="9" pageNumber="367">
|
||
<emphasis id="B900EADEFFB7124D28EEFB7185291BA2" bold="true" box="[151,277,1246,1270]" pageId="9" pageNumber="367">FIGURE 5.</emphasis>
|
||
Overall dorsal habitus. A,
|
||
<taxonomicName id="4C744D4FFFB7124D2A48FB7186E21BA2" authorityName="Rahayu & Ng" authorityYear="2024" box="[561,734,1246,1270]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="ngankee" status="sp. nov.">
|
||
<emphasis id="B900EADEFFB7124D2A48FB7186E21BA2" box="[561,734,1246,1270]" italics="true" pageId="9" pageNumber="367">Lyphira ngankee</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EADEFFB7124D2A9EFB7187141BA2" bold="true" box="[743,808,1246,1270]" pageId="9" pageNumber="367">
|
||
<taxonomicNameLabel id="A23357A5FFB7124D2A9EFB71871F1BA2" box="[743,803,1246,1270]" pageId="9" pageNumber="367" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B,
|
||
<taxonomicName id="4C744D4FFFB7124D2886FAAC85521A4E" authorityName="Rahayu & Ng" authorityYear="2024" box="[255,366,1282,1306]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
|
||
<emphasis id="B900EADEFFB7124D2886FAAC85521A4E" box="[255,366,1282,1306]" italics="true" pageId="9" pageNumber="367">L. ngankee</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357A5FFB7124D290CFAAD85921A4D" box="[373,430,1282,1306]" pageId="9" pageNumber="367" rank="species">
|
||
<emphasis id="B900EADEFFB7124D290CFAAD85941A4E" bold="true" box="[373,424,1282,1306]" pageId="9" pageNumber="367">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C,
|
||
<taxonomicName id="4C744D4FFFB7124D2C14FAAC80CB1A4E" authorityName="Rahayu & Ng" authorityYear="2024" box="[1133,1271,1282,1306]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||
<emphasis id="B900EADEFFB7124D2C14FAAC80CB1A4E" box="[1133,1271,1282,1306]" italics="true" pageId="9" pageNumber="367">L. bellagrana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EADEFFB7124D2C87FAAD810B1A4E" bold="true" box="[1278,1335,1282,1306]" pageId="9" pageNumber="367">
|
||
<taxonomicNameLabel id="A23357A5FFB7124D2C87FAAD810B1A4E" box="[1278,1335,1282,1306]" pageId="9" pageNumber="367" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D,
|
||
<taxonomicName id="4C744D4FFFB7124D2B65FA8887991A6A" authorityName="Rahayu & Ng" authorityYear="2024" box="[796,933,1318,1342]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="bellagrana" status="n. sp.">
|
||
<emphasis id="B900EADEFFB7124D2B65FA8887991A6A" box="[796,933,1318,1342]" italics="true" pageId="9" pageNumber="367">L. bellagrana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EADEFFB7124D2BD5FA8987D91A6A" bold="true" box="[940,997,1318,1342]" pageId="9" pageNumber="367">
|
||
<taxonomicNameLabel id="A23357A5FFB7124D2BD5FA8987D91A6A" box="[940,997,1318,1342]" pageId="9" pageNumber="367" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36E6547FFB7124E28BEFA3F800C1EF9" lastPageId="10" lastPageNumber="368" pageId="9" pageNumber="367" type="discussion">
|
||
<paragraph id="8BCB36CCFFB7124D28BEFA3F807A19F2" blockId="9.[151,1437,1424,2027]" pageId="9" pageNumber="367">
|
||
<emphasis id="B900EADEFFB7124D28BEFA3F85081AFE" bold="true" box="[199,308,1424,1450]" pageId="9" pageNumber="367">Remarks</emphasis>
|
||
.
|
||
<bibRefCitation id="EFE54B3DFFB7124D293FFA3F85EA1AFE" author="Chen, H." box="[326,470,1424,1450]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen (1987)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C744D4FFFB7124D2A2EFA3F871C1AFE" authorityName="Chen" authorityYear="1987" box="[599,800,1424,1450]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="acutidens">
|
||
<emphasis id="B900EADEFFB7124D2A2EFA3F871C1AFE" box="[599,800,1424,1450]" italics="true" pageId="9" pageNumber="367">Philyra acutidens</emphasis>
|
||
</taxonomicName>
|
||
from a large series of specimens from the East
|
||
<collectingCountry id="F363765CFFB7124D2D20FA3F81A01AFE" box="[1369,1436,1424,1450]" name="China" pageId="9" pageNumber="367">China</collectingCountry>
|
||
Sea and Yellow Sea (
|
||
<specimenCount id="9D72FD45FFB7124D29F5FA1B85CF1A9A" box="[396,499,1460,1486]" count="87" pageId="9" pageNumber="367" type="male">87 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D72FD45FFB7124D2A57FA1B86821A9A" box="[558,702,1460,1486]" count="101" pageId="9" pageNumber="367" type="female">101 females</specimenCount>
|
||
), and she compared it with
|
||
<taxonomicName id="4C744D4FFFB7124D2B87FA1A80981A9A" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[1022,1188,1460,1486]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB7124D2B87FA1A80981A9A" box="[1022,1188,1460,1486]" italics="true" pageId="9" pageNumber="367">P. heterograna</emphasis>
|
||
</taxonomicName>
|
||
, noting that it can be distinguished by its slightly smaller adult size (largest carapace length
|
||
<quantity id="4C8C9B29FFB7124D2BC4FA77801E1AA7" box="[957,1058,1496,1523]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="9" pageNumber="367" unit="mm" value="16.5">16.5 mm</quantity>
|
||
versus
|
||
<quantity id="4C8C9B29FFB7124D2C03FA7780D81AA6" box="[1146,1252,1496,1523]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="9" pageNumber="367" unit="mm" value="18.5">18.5 mm</quantity>
|
||
), slightly longer carapace, the presence of more granules on the carapace, the third maxilliped exopod is slightly narrower, the proportionately shorter fingers of the chela, the cutting margin of the fingers lacking a large tubercle, male pleonal somite 6 with median tooth, and the female telson is longer than broad. She figured a specimen of what she regarded was
|
||
<taxonomicName id="4C744D4FFFB7124D28B3F9C6854D19D6" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[202,369,1640,1666]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB7124D28B3F9C6854D19D6" box="[202,369,1640,1666]" italics="true" pageId="9" pageNumber="367">L. heterograna</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357A5FFB7124D2900F9C6858819D7" box="[377,436,1641,1667]" pageId="9" pageNumber="367" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
(
|
||
<bibRefCitation id="EFE54B3DFFB7124D29BAF9C7867F19D6" author="Chen, H." box="[451,579,1640,1666]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen 1987</bibRefCitation>
|
||
: fig. 2) but did not state where it was from. From her figures,
|
||
<bibRefCitation id="EFE54B3DFFB7124D2C87F9C781A519D6" author="Chen, H." box="[1278,1433,1640,1666]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen’s (1987</bibRefCitation>
|
||
: fig. 2) “
|
||
<taxonomicName id="4C744D4FFFB7124D2896F92385EF19F2" authorityName="Ortmann" authorityYear="1892" box="[239,467,1676,1702]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB7124D2896F92385EF19F2" box="[239,467,1676,1702]" italics="true" pageId="9" pageNumber="367">Philyra heterograna</emphasis>
|
||
</taxonomicName>
|
||
” agrees well with
|
||
<taxonomicName id="4C744D4FFFB7124D2ADCF922877019F2" authorityName="Ortmann" authorityYear="1892" box="[677,844,1676,1702]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB7124D2ADCF922877019F2" box="[677,844,1676,1702]" italics="true" pageId="9" pageNumber="367">L. heterograna</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357A5FFB7124D2B2AF92287B219F3" box="[851,910,1677,1703]" pageId="9" pageNumber="367" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
as defined here.
|
||
</paragraph>
|
||
<paragraph id="8BCB36CCFFB7124D28BEF91F8636182A" blockId="9.[151,1437,1424,2027]" pageId="9" pageNumber="367">
|
||
<bibRefCitation id="EFE54B3DFFB7124D28BEF91F8569199E" author="Galil, B. S." box="[199,341,1712,1738]" pageId="9" pageNumber="367" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil (2009)</bibRefCitation>
|
||
commented that
|
||
<taxonomicName id="4C744D4FFFB7124D2A5DF91E868D199E" authorityName="Chen" authorityYear="1987" box="[548,689,1712,1738]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="acutidens">
|
||
<emphasis id="B900EADEFFB7124D2A5DF91E868D199E" box="[548,689,1712,1738]" italics="true" pageId="9" pageNumber="367">L. acutidens</emphasis>
|
||
</taxonomicName>
|
||
is a junior synonym of
|
||
<taxonomicName id="4C744D4FFFB7124D2BACF91E80BC199E" authorityName="Ortmann" authorityYear="1892" box="[981,1152,1712,1738]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="367" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB7124D2BACF91E80BC199E" box="[981,1152,1712,1738]" italics="true" pageId="9" pageNumber="367">L. heterograna</emphasis>
|
||
</taxonomicName>
|
||
, arguing the differences observed were only due to their subadult nature. This is not the case; the species is not small, with the
|
||
<typeStatus id="54CF886EFFB7124D2C87F97B815C19BA" box="[1278,1376,1748,1774]" pageId="9" pageNumber="367" type="holotype">holotype</typeStatus>
|
||
male measuring 15.0 × 16.0 mm and
|
||
<typeStatus id="54CF886EFFB7124D2A7BF95786601846" box="[514,604,1784,1810]" pageId="9" pageNumber="367" type="allotype">allotype</typeStatus>
|
||
female 11.9 × 12.0 mm. The present male specimen (ZRC 2002.0504) is relatively small (8.9 ×
|
||
<quantity id="4C8C9B29FFB7124D29ECF8B385D01862" box="[405,492,1820,1846]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="9" pageNumber="367" unit="mm" value="8.6">8.6 mm</quantity>
|
||
) and does not appear fully adult; but the slightly larger female (13.4 ×
|
||
<quantity id="4C8C9B29FFB7124D2D7AF8B381541862" box="[1283,1384,1820,1846]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="9" pageNumber="367" unit="mm" value="12.9">12.9 mm</quantity>
|
||
) has the pleon round and domed and is clearly adult. Their characters, however, agree in most aspects with
|
||
<bibRefCitation id="EFE54B3DFFB7124D2C87F8EF81A0180E" author="Chen, H." box="[1278,1436,1856,1882]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen’s (1987)</bibRefCitation>
|
||
detailed descriptions and figures.
|
||
</paragraph>
|
||
<paragraph id="8BCB36CCFFB7124E28BEF826800C1EF9" blockId="9.[151,1437,1424,2027]" lastBlockId="10.[151,1437,151,501]" lastPageId="10" lastPageNumber="368" pageId="9" pageNumber="367">
|
||
We are of the opinion the two species are distinct. The differences in the adult size observed by
|
||
<bibRefCitation id="EFE54B3DFFB7124D2C91F827814818F6" author="Chen, H." box="[1256,1396,1928,1954]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen (1987)</bibRefCitation>
|
||
are actually not significant, and carapace proportions alone is not enough to justify recognizing two species. The authors have on hand two non-type topotypic specimens from the Yellow Sea but are part of
|
||
<bibRefCitation id="EFE54B3DFFB7124D2C29F87F80CC18BE" author="Chen, H." box="[1104,1264,2000,2026]" pageId="9" pageNumber="367" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen’s (1987)</bibRefCitation>
|
||
original series, and they do show the other differences highlighted by her. Most marked are the denser concentration of granules on the dorsal surface. The median tubercle on male pleonal somite 6 of
|
||
<taxonomicName id="4C744D4FFFB4124E2BDCFF1480131F81" authorityName="Chen" authorityYear="1987" box="[933,1071,187,213]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="acutidens">
|
||
<emphasis id="B900EADEFFB4124E2BDCFF1480131F81" box="[933,1071,187,213]" italics="true" pageId="10" pageNumber="368">L. acutidens</emphasis>
|
||
</taxonomicName>
|
||
is slightly sharper than those of
|
||
<taxonomicName id="4C744D4FFFB4124E28EEFF7085011FAD" authorityName="Ortmann" authorityYear="1892" box="[151,317,223,249]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB4124E28EEFF7085011FAD" box="[151,317,223,249]" italics="true" pageId="10" pageNumber="368">L. heterograna</emphasis>
|
||
</taxonomicName>
|
||
and is slightly more subdistal in position, just behind the distal margin (
|
||
<figureCitation id="134F2A49FFB4124E2C1AFF7080FA1FAD" box="[1123,1222,223,249]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="10" pageNumber="368">Figs. 7D</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB4124E2CA8FF7080C41FAD" box="[1233,1272,223,249]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="10" pageNumber="368">17I</figureCitation>
|
||
) whereas in
|
||
<taxonomicName id="4C744D4FFFB4124E2DFFFF70851C1E49" authorityName="Ortmann" authorityYear="1892" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB4124E2DFFFF70851C1E49" italics="true" pageId="10" pageNumber="368">L. heterograna</emphasis>
|
||
</taxonomicName>
|
||
, it is positioned a bit further back (
|
||
<figureCitation id="134F2A49FFB4124E2ADCFEAC87151E49" box="[677,809,259,285]" captionStart="FIGURE 7" captionStartId="12.[151,250,1749,1773]" captionTargetBox="[151,1435,181,1725]" captionTargetId="figure-17@12.[151,1435,181,1725]" captionTargetPageId="12" captionText="FIGURE 7. Male thoracic sternum and pleon. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5), China; C, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia." figureDoi="http://doi.org/10.5281/zenodo.12681940" httpUri="https://zenodo.org/record/12681940/files/figure.png" pageId="10" pageNumber="368">Figs. 7A‒C</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A49FFB4124E2B4DFEAC87591E49" box="[820,869,259,285]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="10" pageNumber="368">17B</figureCitation>
|
||
).
|
||
<bibRefCitation id="EFE54B3DFFB4124E2B00FEAC80241E49" author="Chen, H." box="[889,1048,259,285]" pageId="10" pageNumber="368" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen’s (1987)</bibRefCitation>
|
||
figure of the male pleon is slightly inaccurate, the median tubercle on somite 6 is not distal in position (
|
||
<bibRefCitation id="EFE54B3DFFB4124E2BE4FE8880221E15" author="Chen, H." box="[925,1054,295,321]" pageId="10" pageNumber="368" pagination="195 - 203" refId="ref19456" refString="Chen, H. (1987) On two new species of Leucosiidae (Crustacea, Brachyura) from the Chinese waters. Studia Marina Sinica, 28, 195 - 203." type="journal article" year="1987">Chen 1987</bibRefCitation>
|
||
: fig. 1.3) but is actually subdistal. Also significant is that the vulvae are differently structured, with the opening in
|
||
<taxonomicName id="4C744D4FFFB4124E2C6FFEE480811E31" authorityName="Ortmann" authorityYear="1892" box="[1046,1213,331,357]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="heterograna">
|
||
<emphasis id="B900EADEFFB4124E2C6FFEE480811E31" box="[1046,1213,331,357]" italics="true" pageId="10" pageNumber="368">L. heterograna</emphasis>
|
||
</taxonomicName>
|
||
is short and weakly crescent-shaped (
|
||
<figureCitation id="134F2A49FFB4124E2920FEC085821EDD" box="[345,446,367,393]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="10" pageNumber="368">Fig. 21A</figureCitation>
|
||
) whereas it is slit-like and distinctly more crescent-shaped in
|
||
<taxonomicName id="4C744D4FFFB4124E2C17FEC080CA1EDD" authorityName="Chen" authorityYear="1987" box="[1134,1270,367,393]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="acutidens">
|
||
<emphasis id="B900EADEFFB4124E2C17FEC080CA1EDD" box="[1134,1270,367,393]" italics="true" pageId="10" pageNumber="368">L. acutidens</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134F2A49FFB4124E2D7CFEC081571EDD" box="[1285,1387,367,393]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="10" pageNumber="368">Fig. 21B</figureCitation>
|
||
). As such, we believe it is better to recognize
|
||
<taxonomicName id="4C744D4FFFB4124E2A19FE3C86D41EF9" authorityName="Chen" authorityYear="1987" box="[608,744,403,429]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="acutidens">
|
||
<emphasis id="B900EADEFFB4124E2A19FE3C86D41EF9" box="[608,744,403,429]" italics="true" pageId="10" pageNumber="368">L. acutidens</emphasis>
|
||
</taxonomicName>
|
||
as a distinct species for now.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6547FFB4124E28BEFE1886801E85" box="[199,700,438,465]" pageId="10" pageNumber="368" type="materials_examined">
|
||
<paragraph id="8BCB36CCFFB4124E28BEFE1886801E85" blockId="10.[151,1437,151,501]" box="[199,700,438,465]" pageId="10" pageNumber="368">
|
||
<materialsCitation id="3B1C3C91FFB4124E28BEFE1886841E85" box="[199,696,438,465]" country="China" county="Sea" location="Yellow Sea" municipality="North" pageId="10" pageNumber="368" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B900EADEFFB4124E28BEFE18855C1E84" bold="true" box="[199,352,438,465]" pageId="10" pageNumber="368">
|
||
<typeStatus id="54CF886EFFB4124E28BEFE18853E1E85" box="[199,258,439,465]" pageId="10" pageNumber="368">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<location id="8EAB6017FFB4124E2914FE1885D21E85" LSID="urn:lsid:plazi:treatment:03DD87DAFFB8124E28EEFBBC87BE1EA1:8EAB6017FFB4124E2914FE1885D21E85" box="[365,494,439,465]" country="China" county="Sea" municipality="North" name="Yellow Sea" pageId="10" pageNumber="368">Yellow Sea</location>
|
||
,
|
||
<collectingMunicipality id="6BAFACB6FFB4124E2983FE1886071E85" box="[506,571,439,465]" pageId="10" pageNumber="368">North</collectingMunicipality>
|
||
<collectingCountry id="F363765CFFB4124E2A3BFE1886B91E85" box="[578,645,439,465]" name="China" pageId="10" pageNumber="368">China</collectingCountry>
|
||
<collectingCounty id="62AA4E40FFB4124E2AF5FE1886841E85" box="[652,696,439,465]" pageId="10" pageNumber="368">Sea</collectingCounty>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6547FFB4124E28BEFE7587BE1EA1" box="[199,898,474,501]" pageId="10" pageNumber="368" type="distribution">
|
||
<paragraph id="8BCB36CCFFB4124E28BEFE7587BE1EA1" blockId="10.[151,1437,151,501]" box="[199,898,474,501]" pageId="10" pageNumber="368">
|
||
<emphasis id="B900EADEFFB4124E28BEFE7585651EA0" bold="true" box="[199,345,474,500]" pageId="10" pageNumber="368">Distribution</emphasis>
|
||
. Bohai Sea, Yellow Sea and Northern
|
||
<collectingCountry id="F363765CFFB4124E2B71FE7487771EA1" box="[776,843,475,501]" name="China" pageId="10" pageNumber="368">China</collectingCountry>
|
||
Sea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |