161 lines
18 KiB
XML
161 lines
18 KiB
XML
<document id="D5C67030ABA4C60065DE42EAB78FBF38" ID-ISSN="0024-4082" ID-ZooBank="7A6A530-3EBA-4AF2-94E2-E9B8A1D07459" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738713251128" checkinUser="plazi" docAuthor="Roggero, Angela, Dierkens, Michael, Barbero, Enrico & Palestrini, Claudia" docDate="2017" docId="03DC6B6BFFF0FFD5FF1DFA48FE91F989" docLanguage="en" docName="zoj.12498.pdf" docOrigin="Zoological Journal of the Linnean Society 180" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Morettius pallens" docType="treatment" docVersion="1" lastPageNumber="316" masterDocId="FFE51313FFE1FFC7FFA1FFD5FF82FFF0" masterDocTitle="Combined phylogenetic analysis of two new Afrotropical genera of Onthophagini (Coleoptera, Scarabaeidae)" masterLastPageNumber="320" masterPageNumber="298" pageNumber="315" updateTime="1738713343532" updateUser="GgImagineBatch">
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<mods:title id="BB22C549C84F89BCF15DE317BC904705">Combined phylogenetic analysis of two new Afrotropical genera of Onthophagini (Coleoptera, Scarabaeidae)</mods:title>
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<mods:namePart id="BF5A1199F1F1E0F21888EFEB8C1B4B80">Roggero, Angela</mods:namePart>
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<mods:namePart id="B1118F3346A102226B94E712DCA10E9D">Dierkens, Michael</mods:namePart>
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<mods:namePart id="E25F5DB78E9F30C6CD456F3B6901F4ED">Barbero, Enrico</mods:namePart>
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<mods:namePart id="014700A3F5AAC9747347111F96C60B92">Palestrini, Claudia</mods:namePart>
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<mods:title id="9160E8B595BD6F0E107BB508B2899698">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="AC11749A9EA872FDAFFA39D7128726F1">2017</mods:date>
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<mods:number id="8282564BE6C6B8F7EC42094B614B3040">180</mods:number>
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<treatment id="03DC6B6BFFF0FFD5FF1DFA48FE91F989" LSID="urn:lsid:plazi:treatment:03DC6B6BFFF0FFD5FF1DFA48FE91F989" httpUri="http://treatment.plazi.org/id/03DC6B6BFFF0FFD5FF1DFA48FE91F989" lastPageId="18" lastPageNumber="316" pageId="17" pageNumber="315">
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<subSubSection id="C36F89F6FFF0FFD6FF1DFA48FD51FA42" box="[188,723,1432,1459]" pageId="17" pageNumber="315" type="nomenclature">
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<paragraph id="8BCADA7DFFF0FFD6FF1DFA48FD51FA42" blockId="17.[188,723,1432,1490]" box="[188,723,1432,1459]" pageId="17" pageNumber="315">
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<heading id="D0826D11FFF0FFD6FF1DFA48FD51FA42" box="[188,723,1432,1459]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="315" reason="2">
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<taxonomicName id="4C75A1FEFFF0FFD6FF1DFA48FD51FA42" ID-CoL="44D82" authority="(D' ORBIGNY, 1908: 172)" baseAuthorityName="D' ORBIGNY" baseAuthorityPageNumber="172" baseAuthorityYear="1908" box="[188,723,1432,1459]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="pallens">
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<emphasis id="B901066FFFF0FFD6FF1DFA48FE2EFA40" box="[188,428,1437,1458]" italics="true" pageId="17" pageNumber="315">
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<smallCapsWord id="8D2C4CA1FFF0FFD6FF1DFA48FEC0FA40" baselines="1453,1452" box="[188,322,1437,1458]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Morettius" pageId="17" pageNumber="315">MORETTIUS</smallCapsWord>
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<smallCapsWord id="8D2C4CA1FFF0FFD6FEEAFA75FE2EFA40" baselines="1452" box="[331,428,1440,1456]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="pallens" pageId="17" pageNumber="315">PALLENS</smallCapsWord>
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</emphasis>
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(
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<smallCapsWord id="8D2C4CA1FFF0FFD6FE1EFA75FE57FA42" baselines="1452" box="[447,469,1432,1458]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="d" pageId="17" pageNumber="315">D’</smallCapsWord>
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<smallCapsWord id="8D2C4CA1FFF0FFD6FE75FA4FFDB8FA41" baselines="1452,1453" box="[468,570,1434,1458]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Orbigny" pageId="17" pageNumber="315">ORBIGNY</smallCapsWord>
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, 1908: 172)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F89F6FFF0FFD5FEFFFA62FE91F989" lastPageId="18" lastPageNumber="316" pageId="17" pageNumber="315" type="description">
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<paragraph id="8BCADA7DFFF0FFD6FEFFFA62FDB0FA21" blockId="17.[188,723,1432,1490]" box="[350,562,1463,1490]" pageId="17" pageNumber="315">
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(
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<figureCitation id="134EC6F8FFF0FFD6FEC6FA6CFE3AFA21" box="[359,440,1463,1489]" captionStart="Figure 2" captionStartId="4.[162,241,1141,1160]" captionTargetBox="[391,1215,203,1113]" captionTargetId="figure-431@4.[391,1215,203,1113]" captionTargetPageId="4" captionText="Figure 2. Epipharynx of the species of the genera Hamonthophagus (AC = acutus, BI = bitubeculatus, DE = depressus, FA = fallax, LA = laceratus) and Morettius (PA = pallens, UT = utete). Scale bar = 0.5 mm." pageId="17" pageNumber="315">
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<smallCapsWord id="8D2C4CA1FFF0FFD6FEC6FA6CFE1EFA3F" baselines="1483,1483" box="[359,412,1465,1489]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="17" pageNumber="315">FIGS</smallCapsWord>
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2
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</figureCitation>
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,
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<figureCitation id="134EC6F8FFF0FFD6FE66FA6DFE5BFA22" box="[455,473,1464,1490]" captionStart="Figure 7" captionStartId="8.[162,241,917,936]" captionTargetBox="[391,1215,201,883]" captionTargetId="figure-27@8.[388,1212,198,884]" captionTargetPageId="8" captionText="Figure 7. Aedeagus of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA); see text for the codes. Scale bar = 0.5 mm." pageId="17" pageNumber="315">7</figureCitation>
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,
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<figureCitation id="134EC6F8FFF0FFD6FE49FA62FE7BFA21" box="[488,505,1463,1489]" captionStart="Figure 8" captionStartId="8.[162,241,1850,1869]" captionTargetBox="[391,1215,1064,1819]" captionTargetId="figure-59@8.[388,1212,1062,1817]" captionTargetPageId="8" captionText="Figure 8. Primary lamella of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA). Scale bar = 0.5 mm." pageId="17" pageNumber="315">8</figureCitation>
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,
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<figureCitation id="134EC6F8FFF0FFD6FDA8FA62FDAAFA21" box="[521,552,1463,1489]" captionStart="Figure 11" captionStartId="10.[162,241,707,726]" captionTargetBox="[391,1215,198,674]" captionTargetId="figure-136@10.[391,1215,198,674]" captionTargetPageId="10" captionText="Figure 11. Vagina and receptaculum seminis of the species of the genus Morettius (PA and UT). Scale bar = 0.5 mm." pageId="17" pageNumber="315">11</figureCitation>
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)
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</paragraph>
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<paragraph id="8BCADA7DFFF0FFD6FF35F9DDFEF8F946" blockId="17.[148,763,1543,1718]" pageId="17" pageNumber="315">
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<emphasis id="B901066FFFF0FFD6FF35F9DDFEC4F9EC" box="[148,326,1543,1565]" italics="true" pageId="17" pageNumber="315">Type material:</emphasis>
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<materialsCitation id="3B1DD020FFF0FFD6FEF4F9D2FD75F9A9" collectionCode="MNHN" country="Chad" location="Chad lake" municipality="Kiao-Kata" pageId="17" pageNumber="315" specimenCount="1" stateProvince="Moyen-Chari" typeStatus="lectotype">
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<typeStatus id="54CE64DFFFF0FFD6FEF4F9D2FE41F9EC" box="[341,451,1543,1564]" pageId="17" pageNumber="315" type="lectotype">Lectotype</typeStatus>
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here designated: (male)
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<collectingCountry id="F3629AEDFFF0FFD6FF35F9F3FF61F9CB" box="[148,227,1574,1595]" name="Chad" pageId="17" pageNumber="315">CHAD</collectingCountry>
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:
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<collectingMunicipality id="6BAE4007FFF0FFD6FF51F9F0FEEAF9CB" box="[240,360,1573,1595]" pageId="17" pageNumber="315">Kiao-Kata</collectingMunicipality>
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,
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<collectingRegion id="49B1149FFFF0FFD6FED7F9F0FD8CF9CB" box="[374,526,1573,1595]" country="Chad" name="Moyen-Chari" pageId="17" pageNumber="315">Moyen-Chari</collectingRegion>
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, south to
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<location id="8EAA8CA6FFF0FFD6FD24F9F0FD79F9CB" LSID="urn:lsid:plazi:treatment:03DC6B6BFFF0FFD5FF1DFA48FE91F989:8EAA8CA6FFF0FFD6FD24F9F0FD79F9CB" box="[645,763,1573,1595]" country="Chad" municipality="Kiao-Kata" name="Chad lake" pageId="17" pageNumber="315" stateProvince="Moyen-Chari">Chad lake</location>
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[=moyen Chari rives, Kiao-Kata] [
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<collectionCode id="ED6442B8FFF0FFD6FD37F991FD73F9A9" box="[662,753,1604,1625]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="315">MNHN</collectionCode>
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]
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</materialsCitation>
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.
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<materialsCitation id="3B1DD020FFF0FFD6FF35F9B6FEF4F946" collectionCode="MNHN" country="Chad" location="Chad lake" municipality="Kiao-Kata" pageId="17" pageNumber="315" specimenCount="1" stateProvince="Moyen-Chari" typeStatus="paralectotype">
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<typeStatus id="54CE64DFFFF0FFD6FF35F9B6FEB6F989" box="[148,308,1635,1657]" pageId="17" pageNumber="315" type="paralectotype">Paralectotype</typeStatus>
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: (female) same locality [
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<collectionCode id="ED6442B8FFF0FFD6FDC1F9B6FD39F988" box="[608,699,1635,1656]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="315">MNHN</collectionCode>
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]. For the morphological account, please refer to the original description
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BCADA7DFFF0FFD6FF35F908FEAFF89D" blockId="17.[148,763,1757,1901]" pageId="17" pageNumber="315">
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<emphasis id="B901066FFFF0FFD6FF35F908FDB2F902" box="[148,560,1757,1778]" italics="true" pageId="17" pageNumber="315">
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Geographical distribution (
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<figureCitation id="134EC6F8FFF0FFD6FE70F908FDA0F902" box="[465,546,1757,1778]" captionStart="Figure 14" captionStartId="12.[162,241,1389,1408]" captionTargetBox="[391,1208,203,1361]" captionTargetId="figure-188@12.[388,1212,198,1356]" captionTargetPageId="12" captionText="Figure 14. Distribution of the species (see Appendix 3, Supporting Information for a list of localities) with the Olson et al. (2001) terrestrial biomes classification." pageId="17" pageNumber="315">Fig. 14</figureCitation>
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):
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</emphasis>
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The species was reported from
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<collectingCountry id="F3629AEDFFF0FFD6FEEBF929FE43F8E1" box="[330,449,1788,1809]" name="Cameroon" pageId="17" pageNumber="315">Cameroon</collectingCountry>
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,
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<collectingCountry id="F3629AEDFFF0FFD6FE74F92EFD96F8E1" box="[469,532,1787,1809]" name="Chad" pageId="17" pageNumber="315">Chad</collectingCountry>
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,
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<collectingCountry id="F3629AEDFFF0FFD6FD89F92EFDFFF8E1" box="[552,637,1787,1809]" name="Nigeria" pageId="17" pageNumber="315">Nigeria</collectingCountry>
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(southern border),
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<collectingCountry id="F3629AEDFFF0FFD6FF5EF8CFFE62F8DF" box="[255,480,1818,1840]" name="Republic of the Congo" pageId="17" pageNumber="315">Republic of Congo</collectingCountry>
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and
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<collectingCountry id="F3629AEDFFF0FFD6FD8FF8CFFDF5F8C0" box="[558,631,1818,1840]" name="Sudan" pageId="17" pageNumber="315">Sudan</collectingCountry>
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(collection localities are listed in Appendix 3, Supporting Information).
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</paragraph>
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<caption id="DF0A8AF5FFF0FFD6FC8AF8EFFBB5F89B" pageId="17" pageNumber="315" startId="17.[811,890,1850,1869]" targetBox="[890,1344,1160,1818]" targetPageId="17" targetType="figure">
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<paragraph id="8BCADA7DFFF0FFD6FC8AF8EFFBB5F89B" blockId="17.[811,1426,1850,1899]" pageId="17" pageNumber="315">
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<emphasis id="B901066FFFF0FFD6FC8AF8EFFB4DF8BD" bold="true" box="[811,1231,1850,1869]" pageId="17" pageNumber="315">
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Figure 17.
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<taxonomicName id="4C75A1FEFFF0FFD6FC0FF8EFFBE1F8BD" authorityName="Roggero & Dierkens & Barbero & Palestrini" authorityYear="2017" box="[942,1123,1850,1869]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete" status="sp. nov.">
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<emphasis id="B901066FFFF0FFD6FC0FF8EFFBE1F8BD" bold="true" box="[942,1123,1850,1869]" italics="true" pageId="17" pageNumber="315">Morettius utete</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A232BB14FFF0FFD6FBD1F8EFFB4DF8BD" box="[1136,1231,1850,1869]" pageId="17" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, paratype female facies. Scale bar = 1 mm.
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</paragraph>
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</caption>
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<paragraph id="8BCADA7DFFF3FFD5FF03FF12FED0FDFE" blockId="18.[162,777,199,1382]" pageId="18" pageNumber="316">covering more than half the pronotal length and prolonged narrowly on the sides to reach the anterior angles. Pronotal pubescence black and very short, only evident on the lateral edges. Pronotum unarmed, very weakly transverse (length/width ratio: 0.55). Base evenly curved, markedly bordered. Posterior angles not sinuate. Anterior angles strongly sinuate, sharply projected outwards. Surface reticulate. Area surrounding the callus not granulate; remaining pronotal surface entirely covered by small granules.</paragraph>
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<paragraph id="8BCADA7DFFF3FFD5FF1BFDC2FF78FC29" blockId="18.[162,777,199,1382]" pageId="18" pageNumber="316">
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Elytra yellow with black symmetrical spots, one basal on the fifth interstriae, one on the first third of the sixth and seventh interstriae, another four connected on the middle of the second to fifth interstriae forming a zig-zig pattern. Juxtasutural interstriae yellow
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<emphasis id="B901066FFFF3FFD5FE22FD67FE0DFD34" box="[387,399,690,708]" italics="true" pageId="18" pageNumber="316">–</emphasis>
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orange, darkened anteriorly. Basal carina of interstriae bronze. Pubescence yellow, very short and scattered, only evident posteriorly. Elytral ground reticulate. Interstriae bearing small yellow granules, arranged on the juxtasutural interstriae in a regular row. Interstriae weakly convex, basally carinate. Striae narrow, well marked, yellow. Punctures of striae never wider than the striae. The seventh stria sinuate basally.
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</paragraph>
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<paragraph id="8BCADA7DFFF3FFD5FF1BFC34FEC7FBC4" blockId="18.[162,777,199,1382]" pageId="18" pageNumber="316">Pygidium yellow almost smooth, finely microreticulate, with small, rare, hardly perceptible punctures. Base carinate.</paragraph>
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<paragraph id="8BCADA7DFFF3FFD5FF1BFBE8FEEAFB5F" blockId="18.[162,777,199,1382]" pageId="18" pageNumber="316">Epipleura yellow. Sternal thoracic surface bronze, except for the base of propleurae bronze. Abdominal sternites yellow. Metasternal pubescence scattered, short and yellow.</paragraph>
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<paragraph id="8BCADA7DFFF3FFD5FF1BFB62FEC3FA96" blockId="18.[162,777,199,1382]" pageId="18" pageNumber="316">Coxae yellow. Trochanters bronze. Femura yellow, apically bronze. Tibiae bronze, meso- and metatibiae apically yellowish. Tarsomeres weakly bronze. Pubescence yellow. Fore tibiae three-toothed. Tibial spur elongate, bent inward, apically rounded. Tarsi normally shaped.</paragraph>
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<paragraph id="8BCADA7DFFF3FFD5FF03FA58FE0DF9ED" blockId="18.[162,776,1421,1565]" pageId="18" pageNumber="316">
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<emphasis id="B901066FFFF3FFD5FF03FA58FE19FA53" box="[162,411,1421,1443]" italics="true" pageId="18" pageNumber="316">Individual variation:</emphasis>
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<typeStatus id="54CE64DFFFF3FFD5FE0BFA5BFD97FA53" box="[426,533,1422,1443]" pageId="18" pageNumber="316" type="paratype">Paratype</typeStatus>
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: the wide posterior spot of the right elytra is extended on the sixth interstria. Spot of the scond interstria obviously longer than in the hotolypus. Seventh elytral stria only weakly sinuate.
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</paragraph>
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<paragraph id="8BCADA7DFFF3FFD5FF03F991FE91F989" blockId="18.[162,776,1604,1657]" pageId="18" pageNumber="316">
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<emphasis id="B901066FFFF3FFD5FF03F991FE25F9A9" box="[162,423,1604,1626]" italics="true" pageId="18" pageNumber="316">
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Epipharynx (
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<figureCitation id="134EC6F8FFF3FFD5FEF7F990FE1BF9A9" box="[342,409,1604,1626]" captionStart="Figure 2" captionStartId="4.[162,241,1141,1160]" captionTargetBox="[391,1215,203,1113]" captionTargetId="figure-431@4.[391,1215,203,1113]" captionTargetPageId="4" captionText="Figure 2. Epipharynx of the species of the genera Hamonthophagus (AC = acutus, BI = bitubeculatus, DE = depressus, FA = fallax, LA = laceratus) and Morettius (PA = pallens, UT = utete). Scale bar = 0.5 mm." pageId="18" pageNumber="316">Fig. 2</figureCitation>
|
||
):
|
||
</emphasis>
|
||
See the above generic diagnosis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |