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data/03/C4/52/03C4526D0331A042029626BCFA57F836.xml
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@ -95,7 +96,7 @@ KAHOʻOLAWE
575
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<emphasis id="B9193F690331A04200BA237CFEA4FA23" bold="true" box="[198,304,1405,1424]" pageId="5" pageNumber="575">Figure 3.</emphasis>
Adult
@ -199,7 +200,7 @@ almost entirely nearly white, with some dark scales
(
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1) (
<figureCitation id="1356FFFE0331A04204E220F4FB61F8B8" box="[1182,1269,1781,1803]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." pageId="5" pageNumber="575">Fig. 4B</figureCitation>
<figureCitation id="1356FFFE0331A04204E220F4FB61F8B8" box="[1182,1269,1781,1803]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." figureDoi="http://doi.org/10.5281/zenodo.14807147" httpUri="https://zenodo.org/record/14807147/files/figure.png" pageId="5" pageNumber="575">Fig. 4B</figureCitation>
). Valvae roughly present on femur. Forewing ground colour pale symmetrical, both with long and somewhat robust brown, with very dark brown band extending from arms, although left valva slightly larger than right. base of wing to termen; additional dark brown scales Both valvae bent nearly 90
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upward in the middle, in thin band along costal margin; fringe pale brown. with four heavy, long, sclerotized spur-like setae

127
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@ -102,7 +103,7 @@ KAHOʻOLAWE
(
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= 2) (
<figureCitation id="1356FFFE0339A04A026D26FEFDFEFEA7" box="[529,618,254,277]" captionStart="Figure 5" captionStartId="7.[198,277,1241,1260]" captionTargetBox="[428,1254,253,1205]" captionTargetId="figure-325@7.[427,1255,251,1205]" captionTargetPageId="7" captionText="Figure 5. Female genitalia and larval cases of Hyposmocoma. A, Hyposmocoma hooilo sp. nov., slide WPH042- 13; B, Hyposmocoma kamaula sp. nov., slide WPH050-13; C, Hyposmocoma kanaloa sp. nov., slide WPH123- 13; D, Hyposmocoma mahoepo sp. nov., slide WPH053-13; E, Hyposmocoma nohomeha sp. nov., slide WPH076-13; F, Hyposmocoma oolea sp. nov., slide WPH048-13; G, Hyposmocoma pahanalo sp. nov., slide WPH044-13; H, Hyposmocoma kanaloa sp. nov., larval case, dorsal view, length = 4 mm; I, Hyposmocoma oolea sp. nov., larval case, dorsal view, length = 6 mm. Abbreviations: a, anterior apophysis; cb, corpus bursa; db, ductus bursa; o, ostium-bearing process; p, posterior apophysis; pa, papillae anales." pageId="13" pageNumber="583">Fig. 5G</figureCitation>
<figureCitation id="1356FFFE0339A04A026D26FEFDFEFEA7" box="[529,618,254,277]" captionStart="Figure 5" captionStartId="7.[198,277,1241,1260]" captionTargetBox="[428,1254,253,1205]" captionTargetId="figure-325@7.[427,1255,251,1205]" captionTargetPageId="7" captionText="Figure 5. Female genitalia and larval cases of Hyposmocoma. A, Hyposmocoma hooilo sp. nov., slide WPH042- 13; B, Hyposmocoma kamaula sp. nov., slide WPH050-13; C, Hyposmocoma kanaloa sp. nov., slide WPH123- 13; D, Hyposmocoma mahoepo sp. nov., slide WPH053-13; E, Hyposmocoma nohomeha sp. nov., slide WPH076-13; F, Hyposmocoma oolea sp. nov., slide WPH048-13; G, Hyposmocoma pahanalo sp. nov., slide WPH044-13; H, Hyposmocoma kanaloa sp. nov., larval case, dorsal view, length = 4 mm; I, Hyposmocoma oolea sp. nov., larval case, dorsal view, length = 6 mm. Abbreviations: a, anterior apophysis; cb, corpus bursa; db, ductus bursa; o, ostium-bearing process; p, posterior apophysis; pa, papillae anales." figureDoi="http://doi.org/10.5281/zenodo.14807149" httpUri="https://zenodo.org/record/14807149/files/figure.png" pageId="13" pageNumber="583">Fig. 5G</figureCitation>
). Papillae anales Antennal flagellomeres with alternating rings of short. Apophyses thin and straight; posterior apophy- dark and pale brown scales; scape dark brown; ses very long, about two times length of anterior pecten absent. Thorax, tegula, and metascutellum apophyses. Ostium-bearing process heavily sclero- mottled pale brown to brown. Foreleg and midleg tized and very large, externally protruding, snail-shell coxa, femur, tibia, and tarsomeres mostly dark shaped and curled, with broad base. Ductus bursae brown; fringe brown. Hindleg entirely nearly white. long, about same length as anterior apophysis. Corpus Forewing ground colour pale brown, with three or bursae oval, about same length as anterior apophysis; four wide bands of brown scales extending more or signum absent. Inception of ductus seminalis situated less vertically from costa to anal margin; several at posterior end of corpus bursae. brown spots near cell; some scattered brown scales near anal margin. Hindwing and fringe uniformly
</paragraph>
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@ -121,7 +122,7 @@ Based on its position in the phylogenetic
(
<emphasis id="B9193F690339A04A044B2527FBDFFC88" box="[1079,1099,806,827]" italics="true" pageId="13" pageNumber="583">N</emphasis>
= 2) (
<figureCitation id="1356FFFE0339A04A04E82524FB77FC89" box="[1172,1251,805,827]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." pageId="13" pageNumber="583">Fig. 4I</figureCitation>
<figureCitation id="1356FFFE0339A04A04E82524FB77FC89" box="[1172,1251,805,827]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." figureDoi="http://doi.org/10.5281/zenodo.14807147" httpUri="https://zenodo.org/record/14807147/files/figure.png" pageId="13" pageNumber="583">Fig. 4I</figureCitation>
). Valvae asymmettree, we expect larvae of
<taxonomicName id="4C6D98F80339A04A027E2545FD05FCEB" authorityName="Doorenweerd &amp; Austin &amp; Rubinoff &amp; n &amp; n &amp; n &amp; n &amp; n" authorityYear="2023" box="[514,657,835,857]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="583" phylum="Arthropoda" rank="species" species="pahanalo">
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@ -131,7 +132,7 @@ to construct rical, left arm long and robust, enlarged apically, ‘flat purse
, based on its being sister to with four heavy, long, sclerotized spur-like setae proanother clade that makes this case
<typeStatus id="54D65DD90339A04A02232580FD07FC25" box="[607,659,897,918]" pageId="13" pageNumber="583">type</typeStatus>
. jecting dorsally near apex (lost in
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<figureCitation id="1356FFFE0339A04A04912581FAA8FC25" box="[1261,1340,896,918]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." figureDoi="http://doi.org/10.5281/zenodo.14807147" httpUri="https://zenodo.org/record/14807147/files/figure.png" pageId="13" pageNumber="583">Fig. 4I</figureCitation>
); right arm long and slender with no heavy setae, apex deeply
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(
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3I
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,
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<figureCitation id="1356FFFE0339A04A02EB2242FD24FBEE" box="[663,688,1091,1117]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." figureDoi="http://doi.org/10.5281/zenodo.14807147" httpUri="https://zenodo.org/record/14807147/files/figure.png" pageId="13" pageNumber="583">4I</figureCitation>
) enlarged. Phallus large, cylindrical, heavily sclerotized, curved near apex.
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@ -306,7 +307,7 @@ with the right valva purse’ case, based on its being nested within a clade dee
(
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= 4) (
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<figureCitation id="1356FFFE0339A04A01A32114FDBAF899" box="[479,558,1813,1835]" captionStart="Figure 3" captionStartId="5.[198,277,1405,1424]" captionTargetBox="[420,1247,251,1369]" captionTargetId="figure-28@5.[420,1248,251,1369]" captionTargetPageId="5" captionText="Figure 3. Adult Hyposmocoma. A, Hyposmocoma anoai sp. nov., holotype ♂, forewing = 4.5 mm; B, Hyposmocoma hooilo sp. nov., paratype ♀, forewing = 5.8 mm; C, Hyposmocoma kamaula sp. nov., paratype ♀, forewing = 4.5 mm; D, Hyposmocoma kanaloa sp. nov., holotype ♂, forewing = 3.2 mm; E, Hyposmocoma mahoepo sp. nov., paratype ♂, forewing = 5.6 mm; F, Hyposmocoma nohomeha sp. nov., holotype ♂, forewing = 4.3 mm; G, Hyposmocoma oolea sp. nov., holotype ♂, forewing = 5.0 mm; H, Hyposmocoma pahanalo sp. nov., holotype ♂, forewing = 5.8 mm; I, Hyposmocoma waauhi sp. nov., paratype ♂, forewing = 4.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807145" httpUri="https://zenodo.org/record/14807145/files/figure.png" pageId="13" pageNumber="583">Fig. 3I</figureCitation>
). Wingspan 9.2
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habitats, including decaying logs and leaf litter. They

129
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@ -229,7 +230,7 @@ W
(
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<figureCitation id="1356FFFE033FA04C018A24ABFDD9FD73" box="[502,589,682,704]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." figureDoi="http://doi.org/10.5281/zenodo.14807147" httpUri="https://zenodo.org/record/14807147/files/figure.png" pageId="11" pageNumber="581">Fig. 4F</figureCitation>
). Valvae symmetri- noff, WPH081-13, DNA extraction DN2187. Near cal, simple, slightly upcurved, enlarged and rounded Honokanaiʻa camp, N
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156.68657.
@ -258,7 +259,7 @@ genitalia:
(
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= 1) (
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<figureCitation id="1356FFFE033FA04C026D2524FDFCFC88" box="[529,616,805,827]" captionStart="Figure 5" captionStartId="7.[198,277,1241,1260]" captionTargetBox="[428,1254,253,1205]" captionTargetId="figure-325@7.[427,1255,251,1205]" captionTargetPageId="7" captionText="Figure 5. Female genitalia and larval cases of Hyposmocoma. A, Hyposmocoma hooilo sp. nov., slide WPH042- 13; B, Hyposmocoma kamaula sp. nov., slide WPH050-13; C, Hyposmocoma kanaloa sp. nov., slide WPH123- 13; D, Hyposmocoma mahoepo sp. nov., slide WPH053-13; E, Hyposmocoma nohomeha sp. nov., slide WPH076-13; F, Hyposmocoma oolea sp. nov., slide WPH048-13; G, Hyposmocoma pahanalo sp. nov., slide WPH044-13; H, Hyposmocoma kanaloa sp. nov., larval case, dorsal view, length = 4 mm; I, Hyposmocoma oolea sp. nov., larval case, dorsal view, length = 6 mm. Abbreviations: a, anterior apophysis; cb, corpus bursa; db, ductus bursa; o, ostium-bearing process; p, posterior apophysis; pa, papillae anales." figureDoi="http://doi.org/10.5281/zenodo.14807149" httpUri="https://zenodo.org/record/14807149/files/figure.png" pageId="11" pageNumber="581">Fig. 5E</figureCitation>
).
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anales DN2219. short.
@ -316,7 +317,7 @@ noho (reside) and meha (isolated/alone), so named because the phylogeny
(
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<figureCitation id="1356FFFE033FA04C04132292FB5CFB1B" box="[1135,1224,1171,1193]" captionStart="Figure 3" captionStartId="5.[198,277,1405,1424]" captionTargetBox="[420,1247,251,1369]" captionTargetId="figure-28@5.[420,1248,251,1369]" captionTargetPageId="5" captionText="Figure 3. Adult Hyposmocoma. A, Hyposmocoma anoai sp. nov., holotype ♂, forewing = 4.5 mm; B, Hyposmocoma hooilo sp. nov., paratype ♀, forewing = 5.8 mm; C, Hyposmocoma kamaula sp. nov., paratype ♀, forewing = 4.5 mm; D, Hyposmocoma kanaloa sp. nov., holotype ♂, forewing = 3.2 mm; E, Hyposmocoma mahoepo sp. nov., paratype ♂, forewing = 5.6 mm; F, Hyposmocoma nohomeha sp. nov., holotype ♂, forewing = 4.3 mm; G, Hyposmocoma oolea sp. nov., holotype ♂, forewing = 5.0 mm; H, Hyposmocoma pahanalo sp. nov., holotype ♂, forewing = 5.8 mm; I, Hyposmocoma waauhi sp. nov., paratype ♂, forewing = 4.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807145" httpUri="https://zenodo.org/record/14807145/files/figure.png" pageId="11" pageNumber="581">Fig. 3G</figureCitation>
).
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10.0
@ -397,7 +398,7 @@ genitalia:
(
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<figureCitation id="1356FFFE033FA04C04F220F7FB7CF8BF" box="[1166,1256,1782,1804]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." figureDoi="http://doi.org/10.5281/zenodo.14807147" httpUri="https://zenodo.org/record/14807147/files/figure.png" pageId="11" pageNumber="581">Fig. 4G</figureCitation>
).
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asymmetrical, left valva robust and with four heavy, long,
@ -413,14 +414,14 @@ asymmetrical, left valva robust and with four heavy, long,
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<figureCitation id="1356FFFE033FA04C02282121FDE3F889" box="[596,631,1824,1850]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." figureDoi="http://doi.org/10.5281/zenodo.14807147" httpUri="https://zenodo.org/record/14807147/files/figure.png" pageId="11" pageNumber="581">4G</figureCitation>
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<figureCitation id="1356FFFE033FA04C02F42121FD50F889" box="[648,708,1824,1851]" captionStart="Figure 5" captionStartId="7.[198,277,1241,1260]" captionTargetBox="[428,1254,253,1205]" captionTargetId="figure-325@7.[427,1255,251,1205]" captionTargetPageId="7" captionText="Figure 5. Female genitalia and larval cases of Hyposmocoma. A, Hyposmocoma hooilo sp. nov., slide WPH042- 13; B, Hyposmocoma kamaula sp. nov., slide WPH050-13; C, Hyposmocoma kanaloa sp. nov., slide WPH123- 13; D, Hyposmocoma mahoepo sp. nov., slide WPH053-13; E, Hyposmocoma nohomeha sp. nov., slide WPH076-13; F, Hyposmocoma oolea sp. nov., slide WPH048-13; G, Hyposmocoma pahanalo sp. nov., slide WPH044-13; H, Hyposmocoma kanaloa sp. nov., larval case, dorsal view, length = 4 mm; I, Hyposmocoma oolea sp. nov., larval case, dorsal view, length = 6 mm. Abbreviations: a, anterior apophysis; cb, corpus bursa; db, ductus bursa; o, ostium-bearing process; p, posterior apophysis; pa, papillae anales." figureDoi="http://doi.org/10.5281/zenodo.14807149" httpUri="https://zenodo.org/record/14807149/files/figure.png" pageId="11" pageNumber="581">5F, I</figureCitation>
) sclerotized spur-like setae projecting dorsally near apex of both valvae; one additional heavy seta pro-
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<document id="04BD9857590D02A2B42E09F6F3854E54" ID-ISSN="0024-4082" ID-ZooBank="7A6A530-3EBA-4AF2-94E2-E9B8A1D07459" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738713251128" checkinUser="plazi" docAuthor="Roggero, Angela, Dierkens, Michael, Barbero, Enrico &amp; Palestrini, Claudia" docDate="2017" docId="03DC6B6BFFEEFFC8FF37FF02FB50FD13" docLanguage="en" docName="zoj.12498.pdf" docOrigin="Zoological Journal of the Linnean Society 180" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Hamonthophagus acutus" docType="treatment" docVersion="2" lastPageNumber="313" masterDocId="FFE51313FFE1FFC7FFA1FFD5FF82FFF0" masterDocTitle="Combined phylogenetic analysis of two new Afrotropical genera of Onthophagini (Coleoptera, Scarabaeidae)" masterLastPageNumber="320" masterPageNumber="298" pageNumber="313" updateTime="1738713669289" updateUser="ExternalLinkService">
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<mods:title id="EDBA146A0262C901E2974EEFBB4AE892">Combined phylogenetic analysis of two new Afrotropical genera of Onthophagini (Coleoptera, Scarabaeidae)</mods:title>
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<mods:namePart id="52854CED1A29FC67DDF66C9D62420190">Roggero, Angela</mods:namePart>
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<mods:namePart id="D6F311236EF44200A0F6899A6BD0CF54">Dierkens, Michael</mods:namePart>
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<mods:namePart id="F713A9915002B4FE166A20EF309E6FE3">Barbero, Enrico</mods:namePart>
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<mods:namePart id="627077B474EFD0250D190CE17B1FF224">Palestrini, Claudia</mods:namePart>
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<mods:title id="F3798A219C2E83BCFBC3FB3553A298F0">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="0001CFF6AFAF8142B9CFF1A9F9CFC8F3">2017</mods:date>
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<treatment id="03DC6B6BFFEEFFC8FF37FF02FB50FD13" ID-DOI="http://doi.org/10.5281/zenodo.14807216" ID-Zenodo-Dep="14807216" LSID="urn:lsid:plazi:treatment:03DC6B6BFFEEFFC8FF37FF02FB50FD13" httpUri="http://treatment.plazi.org/id/03DC6B6BFFEEFFC8FF37FF02FB50FD13" lastPageNumber="313" pageId="15" pageNumber="313">
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<paragraph id="8BCADA7DFFEEFFC8FF37FF02FD7BFF1C" blockId="15.[150,761,210,267]" box="[150,761,210,237]" pageId="15" pageNumber="313">
<heading id="D0826D11FFEEFFC8FF37FF02FD7BFF1C" box="[150,761,210,237]" centered="true" fontSize="9" level="2" pageId="15" pageNumber="313" reason="2">
<taxonomicName id="4C75A1FEFFEEFFC8FF37FF02FD7BFF1C" ID-CoL="3JJ8S" authority="(D' ORBIGNY, 1908: 171)" baseAuthorityName="D' ORBIGNY" baseAuthorityPageNumber="171" baseAuthorityYear="1908" box="[150,761,210,237]" class="Insecta" family="Scarabaeidae" genus="Hamonthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="313" phylum="Arthropoda" rank="species" species="acutus">
<emphasis id="B901066FFFEEFFC8FF37FF02FE50FF1A" box="[150,466,215,236]" italics="true" pageId="15" pageNumber="313">
<smallCapsWord id="8D2C4CA1FFEEFFC8FF37FF02FEF0FF1A" baselines="231,230" box="[150,370,215,236]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Hamonthophagus" pageId="15" pageNumber="313">HAMONTHOPHAGUS</smallCapsWord>
<smallCapsWord id="8D2C4CA1FFEEFFC8FEDAFF0FFE50FF1A" baselines="230" box="[379,466,218,234]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="acutus" pageId="15" pageNumber="313">ACUTUS</smallCapsWord>
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(
<smallCapsWord id="8D2C4CA1FFEEFFC8FE47FF0FFE79FF1C" baselines="230" box="[486,507,210,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="d" pageId="15" pageNumber="313">D’</smallCapsWord>
<smallCapsWord id="8D2C4CA1FFEEFFC8FE5AFF01FDE3FF1B" baselines="230,231" box="[507,609,212,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Orbigny" pageId="15" pageNumber="313">ORBIGNY</smallCapsWord>
, 1908: 171)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C36F89F6FFEEFFC8FEFFFF25FB50FD13" pageId="15" pageNumber="313" type="description">
<paragraph id="8BCADA7DFFEEFFC8FEFFFF25FDB3FEFA" blockId="15.[150,761,210,267]" box="[350,561,240,267]" pageId="15" pageNumber="313">
(
<figureCitation id="134EC6F8FFEEFFC8FEC6FF27FE3AFEFA" box="[359,440,240,266]" captionStart="Figure 2" captionStartId="4.[162,241,1141,1160]" captionTargetBox="[391,1215,203,1113]" captionTargetId="figure-431@4.[391,1215,203,1113]" captionTargetPageId="4" captionText="Figure 2. Epipharynx of the species of the genera Hamonthophagus (AC = acutus, BI = bitubeculatus, DE = depressus, FA = fallax, LA = laceratus) and Morettius (PA = pallens, UT = utete). Scale bar = 0.5 mm." pageId="15" pageNumber="313">
<smallCapsWord id="8D2C4CA1FFEEFFC8FEC6FF27FE1EFEF9" baselines="260,261" box="[359,412,242,266]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="15" pageNumber="313">FIGS</smallCapsWord>
2
</figureCitation>
,
<figureCitation id="134EC6F8FFEEFFC8FE66FF24FE5BFEFB" box="[455,473,241,267]" captionStart="Figure 7" captionStartId="8.[162,241,917,936]" captionTargetBox="[391,1215,201,883]" captionTargetId="figure-27@8.[388,1212,198,884]" captionTargetPageId="8" captionText="Figure 7. Aedeagus of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA); see text for the codes. Scale bar = 0.5 mm." pageId="15" pageNumber="313">7</figureCitation>
,
<figureCitation id="134EC6F8FFEEFFC8FE49FF25FE7BFEFA" box="[488,505,240,266]" captionStart="Figure 8" captionStartId="8.[162,241,1850,1869]" captionTargetBox="[391,1215,1064,1819]" captionTargetId="figure-59@8.[388,1212,1062,1817]" captionTargetPageId="8" captionText="Figure 8. Primary lamella of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA). Scale bar = 0.5 mm." pageId="15" pageNumber="313">8</figureCitation>
,
<figureCitation id="134EC6F8FFEEFFC8FDA8FF25FDAAFEFA" box="[521,552,240,266]" captionStart="Figure 10" captionStartId="9.[148,227,1743,1762]" captionTargetBox="[373,1197,1135,1709]" captionTargetId="figure-119@9.[373,1197,1135,1709]" captionTargetPageId="9" captionText="Figure 10. Vagina and receptaculum seminis of the species of the genus Hamonthophagus (AC, BI, DE, FA and LA). Scale bar = 0.5 mm." pageId="15" pageNumber="313">10</figureCitation>
)
</paragraph>
<paragraph id="8BCADA7DFFEEFFC8FF35FE94FE47FDBB" blockId="15.[148,764,321,587]" pageId="15" pageNumber="313">
<emphasis id="B901066FFFEEFFC8FF35FE94FED5FEA6" box="[148,343,321,342]" italics="true" pageId="15" pageNumber="313">Type material:</emphasis>
<materialsCitation id="3B1DD020FFEEFFC8FEC7FE94FD75FE85" collectionCode="MNHN" country="Namibia" location="Okahandja" pageId="15" pageNumber="313" specimenCount="1" typeStatus="paralectotype">
<collectingCountry id="F3629AEDFFEEFFC8FEC7FE94FE58FEA6" box="[358,474,321,342]" name="Namibia" pageId="15" pageNumber="313">NAMIBIA</collectingCountry>
[Sud-Ouest africain allemand]:
<location id="8EAA8CA6FFEEFFC8FE8BFE8AFE2AFE85" LSID="urn:lsid:plazi:treatment:03DC6B6BFFEEFFC8FF37FF02FB50FD13:8EAA8CA6FFEEFFC8FE8BFE8AFE2AFE85" box="[298,424,351,373]" country="Namibia" name="Okahandja" pageId="15" pageNumber="313">Okahandja</location>
[
<collectionCode id="ED6442B8FFEEFFC8FE6AFEB5FDA4FE85" box="[459,550,352,373]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="313">MNHN</collectionCode>
].
<typeStatus id="54CE64DFFFEEFFC8FDEDFE8AFD75FE85" box="[588,759,351,373]" pageId="15" pageNumber="313" type="paralectotype">Paralectotypes</typeStatus>
</materialsCitation>
:
<materialsCitation id="3B1DD020FFEEFFC8FF35FEABFDB8FE63" box="[148,570,382,404]" collectionCode="MNHN" country="Botswana" location="lake Ngami" pageId="15" pageNumber="313" specimenCount="1" typeStatus="paralectotype">
<collectingCountry id="F3629AEDFFEEFFC8FF35FEABFEAEFE63" box="[148,300,382,403]" name="Botswana" pageId="15" pageNumber="313">BOTSWANA</collectingCountry>
:
<location id="8EAA8CA6FFEEFFC8FE9DFEABFE47FE64" LSID="urn:lsid:plazi:treatment:03DC6B6BFFEEFFC8FF37FF02FB50FD13:8EAA8CA6FFEEFFC8FE9DFEABFE47FE64" box="[316,453,382,404]" country="Botswana" name="lake Ngami" pageId="15" pageNumber="313">lake Ngami</location>
[
<collectionCode id="ED6442B8FFEEFFC8FE78FEABFDB6FE63" box="[473,564,382,403]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="313">MNHN</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="3B1DD020FFEEFFC8FDEAFEABFECDFE21" collectionCode="MNHN" collectorName="de Mpala" country="Democratic Republic of the Congo" location="Tanganyika" pageId="15" pageNumber="313" specimenCount="1" typeStatus="paralectotype">
<collectingCountry id="F3629AEDFFEEFFC8FDEAFEABFE41FE42" name="Democratic Republic of the Congo" pageId="15" pageNumber="313">DEMOCRATIC REPUBLIC OF CONGO</collectingCountry>
: [
<location id="8EAA8CA6FFEEFFC8FE7CFE48FDE9FE43" LSID="urn:lsid:plazi:treatment:03DC6B6BFFEEFFC8FF37FF02FB50FD13:8EAA8CA6FFEEFFC8FE7CFE48FDE9FE43" box="[477,619,413,435]" country="Democratic Republic of the Congo" name="Tanganyika" pageId="15" pageNumber="313">Tanganyika</location>
,] region
<collectorName id="2680BFABFFEEFFC8FD40FE48FF5EFE21" pageId="15" pageNumber="313">de Mpala</collectorName>
[
<collectionCode id="ED6442B8FFEEFFC8FF4FFE69FECBFE21" box="[238,329,444,465]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="313">MNHN</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="3B1DD020FFEEFFC8FEFFFE6EFEBBFE00" country="Malawi" location="Malawi Lake" pageId="15" pageNumber="313" specimenCount="1" typeStatus="paralectotype">
<collectingCountry id="F3629AEDFFEEFFC8FEFFFE6EFE4DFE20" box="[350,463,443,464]" name="Malawi" pageId="15" pageNumber="313">MALAWI</collectingCountry>
:
<location id="8EAA8CA6FFEEFFC8FE7CFE6EFDF4FE21" LSID="urn:lsid:plazi:treatment:03DC6B6BFFEEFFC8FF37FF02FB50FD13:8EAA8CA6FFEEFFC8FE7CFE6EFDF4FE21" box="[477,630,443,465]" country="Malawi" name="Malawi Lake" pageId="15" pageNumber="313">Malawi Lake</location>
[= Nyassa] [not located]
</materialsCitation>
.
<materialsCitation id="3B1DD020FFEEFFC8FEF8FE0FFD75FE00" box="[345,759,474,496]" country="Namibia" location="Salem" pageId="15" pageNumber="313" specimenCount="1" typeStatus="paralectotype">
<collectingCountry id="F3629AEDFFEEFFC8FEF8FE0FFE53FE1F" box="[345,465,474,495]" name="Namibia" pageId="15" pageNumber="313">NAMIBIA</collectingCountry>
:
<location id="8EAA8CA6FFEEFFC8FE51FE0FFDB5FE00" LSID="urn:lsid:plazi:treatment:03DC6B6BFFEEFFC8FF37FF02FB50FD13:8EAA8CA6FFEEFFC8FE51FE0FFDB5FE00" box="[496,567,474,496]" country="Namibia" name="Salem" pageId="15" pageNumber="313">Salem</location>
[not located]
</materialsCitation>
.
<materialsCitation id="3B1DD020FFEEFFC8FF35FE2DFE43FDBB" collectionCode="MNHN" collectorName="For" country="South Africa" location="Cafrerie" pageId="15" pageNumber="313" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paralectotype">
<collectingCountry id="F3629AEDFFEEFFC8FF35FE2DFEDFFDFD" box="[148,349,504,525]" name="South Africa" pageId="15" pageNumber="313">SOUTH AFRICA</collectingCountry>
:
<collectingRegion id="49B1149FFFEEFFC8FEC9FE2CFDECFDFE" box="[360,622,504,526]" country="South Africa" name="Eastern Cape" pageId="15" pageNumber="313">Eastern Cape province</collectingRegion>
[=
<location id="8EAA8CA6FFEEFFC8FD34FE2DFD77FDFE" LSID="urn:lsid:plazi:treatment:03DC6B6BFFEEFFC8FF37FF02FB50FD13:8EAA8CA6FFEEFFC8FD34FE2DFD77FDFE" box="[661,757,504,526]" country="South Africa" name="Cafrerie" pageId="15" pageNumber="313" stateProvince="Eastern Cape">Cafrerie</location>
] [
<collectionCode id="ED6442B8FFEEFFC8FF3AFDC2FF74FDDC" box="[155,246,535,556]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="313">MNHN</collectionCode>
].
<collectorName id="2680BFABFFEEFFC8FEA9FDC2FEB2FDDC" box="[264,304,535,556]" pageId="15" pageNumber="313">For</collectorName>
the morphological account, please refer to the original description
</materialsCitation>
.
</paragraph>
<paragraph id="8BCADA7DFFEEFFC8FF35FDA7FED5FB5E" blockId="15.[148,763,626,1198]" pageId="15" pageNumber="313">
<emphasis id="B901066FFFEEFFC8FF35FDA7FDD6FD78" box="[148,596,626,648]" italics="true" pageId="15" pageNumber="313">
Geographical distribution (
<figureCitation id="134EC6F8FFEEFFC8FE54FDA6FDC4FD78" box="[501,582,627,648]" captionStart="Figure 14" captionStartId="12.[162,241,1389,1408]" captionTargetBox="[391,1208,203,1361]" captionTargetId="figure-188@12.[388,1212,198,1356]" captionTargetPageId="12" captionText="Figure 14. Distribution of the species (see Appendix 3, Supporting Information for a list of localities) with the Olson et al. (2001) terrestrial biomes classification." pageId="15" pageNumber="313">Fig. 14</figureCitation>
):
</emphasis>
<materialsCitation id="3B1DD020FFEEFFC8FDC2FDA7FED1FB5E" collectionCode="MNHN, BMNH" collectorName="Nyassa &amp; The &amp; Collection Oberthur &amp; du British Museum' &amp; M. Barclay &amp; Neither &amp; As &amp; Also" country="Namibia" county="Tanganika area" location="The" municipality="Besides" pageId="15" pageNumber="313" specimenCount="1">
<location id="8EAA8CA6FFEEFFC8FDC2FDA7FD0DFD78" LSID="urn:lsid:plazi:treatment:03DC6B6BFFEEFFC8FF37FF02FB50FD13:8EAA8CA6FFEEFFC8FDC2FDA7FD0DFD78" box="[611,655,626,648]" country="Namibia" county="Tanganika area" municipality="Besides" name="The" pageId="15" pageNumber="313">The</location>
species distribution comprises
<collectingCountry id="F3629AEDFFEEFFC8FE49FD44FDCFFD57" box="[488,589,657,679]" name="Namibia" pageId="15" pageNumber="313">Namibia</collectingCountry>
, south-west
<collectingCountry id="F3629AEDFFEEFFC8FF35FD65FE86FD35" box="[148,260,688,709]" name="Botswana" pageId="15" pageNumber="313">Botswana</collectingCountry>
and north-west
<collectingCountry id="F3629AEDFFEEFFC8FDA9FD7AFD32FD35" box="[520,688,687,709]" name="South Africa" pageId="15" pageNumber="313">South Africa</collectingCountry>
(see
<location id="8EAA8CA6FFEEFFC8FF35FD1BFE82FD14" LSID="urn:lsid:plazi:treatment:03DC6B6BFFEEFFC8FF37FF02FB50FD13:8EAA8CA6FFEEFFC8FF35FD1BFE82FD14" box="[148,256,718,740]" country="Namibia" county="Tanganika area" municipality="Besides" name="Appendix" pageId="15" pageNumber="313">Appendix</location>
3,
<location id="8EAA8CA6FFEEFFC8FE93FD1BFDC5FD14" LSID="urn:lsid:plazi:treatment:03DC6B6BFFEEFFC8FF37FF02FB50FD13:8EAA8CA6FFEEFFC8FE93FD1BFDC5FD14" box="[306,583,718,740]" country="Namibia" county="Tanganika area" municipality="Besides" name="Supporting Information" pageId="15" pageNumber="313">Supporting Information</location>
for a detailed list of the localities).
<collectingMunicipality id="6BAE4007FFEEFFC8FE0BFD38FD81FCF3" box="[426,515,749,771]" pageId="15" pageNumber="313">Besides</collectingMunicipality>
, in the
<typeStatus id="54CE64DFFFEEFFC8FDD6FD38FD2AFCF2" box="[631,680,749,770]" pageId="15" pageNumber="313">type</typeStatus>
series
<bibRefCitation id="EFE4A78CFFEEFFC8FF35FCDEFEDFFCD0" author="d'Orbigny H" box="[148,349,779,801]" pageId="15" pageNumber="313" pagination="65 - 208" refId="ref13608" refString="d'Orbigny H. 1908. Descriptions d'especes nouvelles d'Onthophagides africains et notes synonimiques. Annales de la Societe Entomologique de France 77: 65-208." type="journal article" year="1908">d’Orbigny (1908)</bibRefCitation>
also included material from the
<collectingCounty id="62ABA2F1FFEEFFC8FF35FCFFFECEFCCF" box="[148,332,810,832]" pageId="15" pageNumber="313">Tanganika area</collectingCounty>
(Democratic Republic of Congo) and
<collectorName id="2680BFABFFEEFFC8FF35FC9CFF65FCAE" box="[148,231,841,862]" pageId="15" pageNumber="313">Nyassa</collectorName>
(i.e.
<collectingCountry id="F3629AEDFFEEFFC8FE98FC9DFE10FCAE" box="[313,402,840,862]" name="Malawi" pageId="15" pageNumber="313">Malawi</collectingCountry>
).
<collectorName id="2680BFABFFEEFFC8FE10FC9DFE5FFCAE" box="[433,477,840,862]" pageId="15" pageNumber="313">The</collectorName>
former specimen was reported in the
<collectorName id="2680BFABFFEEFFC8FEEAFCB2FDB0FC8D" box="[331,562,871,893]" pageId="15" pageNumber="313">Collection Oberthur</collectorName>
,
<collectionCode id="ED6442B8FFEEFFC8FD9EFCB2FD14FC8C" box="[575,662,871,892]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="313">MNHN</collectionCode>
, and the latter was reported in ‘coll.
<collectorName id="2680BFABFFEEFFC8FE74FC50FD3EFC6B" box="[469,700,901,923]" pageId="15" pageNumber="313">du British Museum’</collectorName>
(now
<collectionCode id="ED6442B8FFEEFFC8FF35FC71FF64FC49" box="[148,230,932,953]" pageId="15" pageNumber="313">BMNH</collectionCode>
) where, however, it has not been traced (
<collectorName id="2680BFABFFEEFFC8FF3DFC16FE9FFC28" box="[156,285,962,984]" pageId="15" pageNumber="313">M. Barclay</collectorName>
, pers. comm.).
<collectorName id="2680BFABFFEEFFC8FE6AFC17FDA1FC28" box="[459,547,962,984]" pageId="15" pageNumber="313">Neither</collectorName>
specimen could be examined by us.
<collectorName id="2680BFABFFEEFFC8FECAFC34FE0AFC06" box="[363,392,993,1014]" pageId="15" pageNumber="313">As</collectorName>
no other collection data from these areas were found within the studied material, these records were here regarded as uncertain until further confirmation.
<collectorName id="2680BFABFFEEFFC8FD39FBE8FD4EFBA3" box="[664,716,1085,1107]" pageId="15" pageNumber="313">Also</collectorName>
, a specimen from the
<collectionCode id="ED6442B8FFEEFFC8FE36FB89FE69FB81" box="[407,491,1116,1137]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="313">MNHN</collectionCode>
labelled as ‘
<collectingCountry id="F3629AEDFFEEFFC8FD01FB8EFD78FB81" box="[672,762,1115,1137]" name="Senegal" pageId="15" pageNumber="313">Senegal</collectingCountry>
provenance tres douteuse’ was not included in the present analysis
</materialsCitation>
.
</paragraph>
<paragraph id="8BCADA7DFFEEFFC8FF52FB30FDC4FAE9" blockId="15.[243,668,1253,1337]" pageId="15" pageNumber="313">
<taxonomicName id="4C75A1FEFFEEFFC8FF52FB30FDC4FAE9" authority="(OLIVIER, 1789: 131)" baseAuthorityName="Olivier" baseAuthorityPageNumber="131" baseAuthorityYear="1789" class="Insecta" family="Scarabaeidae" genus="Hamonthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="313" phylum="Arthropoda" rank="species" species="bituberculatus">
<emphasis id="B901066FFFEEFFC8FF52FB30FD1EFB09" box="[243,668,1253,1274]" italics="true" pageId="15" pageNumber="313">
<smallCapsWord id="8D2C4CA1FFEEFFC8FF52FB30FE4DFB09" baselines="1269,1269" box="[243,463,1253,1274]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Hamonthophagus" pageId="15" pageNumber="313">HAMONTHOPHAGUS</smallCapsWord>
<smallCapsWord id="8D2C4CA1FFEEFFC8FE78FB3CFD1EFB09" baselines="1269" box="[473,668,1257,1273]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="bituberculatus" pageId="15" pageNumber="313">BITUBERCULATUS</smallCapsWord>
</emphasis>
(
<smallCapsWord id="8D2C4CA1FFEEFFC8FEF3FAD4FE2CFAE7" baselines="1299,1299" box="[338,430,1281,1305]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Olivier" pageId="15" pageNumber="313">OLIVIER</smallCapsWord>
, 1789: 131)
</taxonomicName>
</paragraph>
<paragraph id="8BCADA7DFFEEFFC8FEFFFACBFDB3FAC8" blockId="15.[243,668,1253,1337]" box="[350,561,1310,1337]" pageId="15" pageNumber="313">
(
<figureCitation id="134EC6F8FFEEFFC8FEC6FAF5FE3AFAC8" box="[359,440,1310,1336]" captionStart="Figure 2" captionStartId="4.[162,241,1141,1160]" captionTargetBox="[391,1215,203,1113]" captionTargetId="figure-431@4.[391,1215,203,1113]" captionTargetPageId="4" captionText="Figure 2. Epipharynx of the species of the genera Hamonthophagus (AC = acutus, BI = bitubeculatus, DE = depressus, FA = fallax, LA = laceratus) and Morettius (PA = pallens, UT = utete). Scale bar = 0.5 mm." pageId="15" pageNumber="313">
<smallCapsWord id="8D2C4CA1FFEEFFC8FEC6FAF5FE1EFAC6" baselines="1330,1330" box="[359,412,1312,1336]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="15" pageNumber="313">FIGS</smallCapsWord>
2
</figureCitation>
,
<figureCitation id="134EC6F8FFEEFFC8FE66FACAFE5BFAC9" box="[455,473,1311,1337]" captionStart="Figure 7" captionStartId="8.[162,241,917,936]" captionTargetBox="[391,1215,201,883]" captionTargetId="figure-27@8.[388,1212,198,884]" captionTargetPageId="8" captionText="Figure 7. Aedeagus of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA); see text for the codes. Scale bar = 0.5 mm." pageId="15" pageNumber="313">7</figureCitation>
,
<figureCitation id="134EC6F8FFEEFFC8FE49FACBFE7BFAC8" box="[488,505,1310,1336]" captionStart="Figure 8" captionStartId="8.[162,241,1850,1869]" captionTargetBox="[391,1215,1064,1819]" captionTargetId="figure-59@8.[388,1212,1062,1817]" captionTargetPageId="8" captionText="Figure 8. Primary lamella of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA). Scale bar = 0.5 mm." pageId="15" pageNumber="313">8</figureCitation>
,
<figureCitation id="134EC6F8FFEEFFC8FDA8FACBFDAAFAC8" box="[521,552,1310,1336]" captionStart="Figure 10" captionStartId="9.[148,227,1743,1762]" captionTargetBox="[373,1197,1135,1709]" captionTargetId="figure-119@9.[373,1197,1135,1709]" captionTargetPageId="9" captionText="Figure 10. Vagina and receptaculum seminis of the species of the genus Hamonthophagus (AC, BI, DE, FA and LA). Scale bar = 0.5 mm." pageId="15" pageNumber="313">10</figureCitation>
)
</paragraph>
<paragraph id="8BCADA7DFFEEFFC8FF35FABBFE58FA52" blockId="15.[148,763,1390,1443]" pageId="15" pageNumber="313">
<emphasis id="B901066FFFEEFFC8FF35FABBFE93FA73" box="[148,273,1390,1411]" italics="true" pageId="15" pageNumber="313">Synonymy:</emphasis>
<taxonomicName id="4C75A1FEFFEEFFC8FEBEFABBFE80FA52" authority="(Olivier, 1789: 171)" baseAuthorityName="Olivier" baseAuthorityPageNumber="171" baseAuthorityYear="1789" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="313" phylum="Arthropoda" rank="species" species="discoideus">
<emphasis id="B901066FFFEEFFC8FEBEFABBFDE4FA73" box="[287,614,1390,1411]" italics="true" pageId="15" pageNumber="313">Onthophagus discoideus</emphasis>
(Olivier, 1789:171)
</taxonomicName>
<emphasis id="B901066FFFEEFFC8FEAAFA58FEBFFA52" box="[267,317,1421,1442]" italics="true" pageId="15" pageNumber="313">teste</emphasis>
Harold 1880.
</paragraph>
<paragraph id="8BCADA7DFFEEFFC8FF35FA1EFE98F89D" blockId="15.[148,763,1482,1901]" pageId="15" pageNumber="313">
<emphasis id="B901066FFFEEFFC8FF35FA1EFEC3FA2F" box="[148,321,1482,1504]" italics="true" pageId="15" pageNumber="313">Type material:</emphasis>
At present, the typical material of
<taxonomicName id="4C75A1FEFFEEFFC8FF35FA3CFEDFFA0D" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[148,349,1512,1534]" class="Insecta" family="Scarabaeidae" genus="Hamonthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="313" phylum="Arthropoda" rank="species" species="bituberculatus">
<emphasis id="B901066FFFEEFFC8FF35FA3CFEDFFA0D" box="[148,349,1512,1534]" italics="true" pageId="15" pageNumber="313">H. bituberculatus</emphasis>
</taxonomicName>
could not be found. Although various materials of the Olivier collection were traced in several museum collections over the years (
<bibRefCitation id="EFE4A78CFFEEFFC8FF3DF991FEA9F9AA" author="Bragg PE" box="[156,299,1604,1626]" pageId="15" pageNumber="313" pagination="105 - 115" refId="ref12693" refString="Bragg PE. 1996. Type specimens of Phasmida in the Nationaal Natuurhistorisch Museum, Leiden (Insecta: Phasmida). Zoologische Mededelingen 70: 105-115." type="journal article" year="1996">Bragg, 1996</bibRefCitation>
;
<bibRefCitation id="EFE4A78CFFEEFFC8FE99F991FD0AF9A9" author="Staines CL &amp; Whittington AE" box="[312,648,1604,1626]" pageId="15" pageNumber="313" pagination="1 - 8" refId="ref14208" refString="Staines CL, Whittington AE. 2003. Chrysomelidae (Coleoptera) types in the Royal Museum of Scotland Collection. Zootaxa 192: 1-8." type="journal article" year="2003">Staines &amp; Whittington, 2003</bibRefCitation>
; G ultekin
<emphasis id="B901066FFFEEFFC8FD0BF990FD34F9A8" bold="true" box="[682,694,1605,1624]" pageId="15" pageNumber="313">€</emphasis>
&amp;
<bibRefCitation id="EFE4A78CFFEEFFC8FF60F9B6FE0DF988" author="Korotyaev BA" box="[193,399,1635,1656]" pageId="15" pageNumber="313" pagination="101 - 111" refId="ref13226" refString="G ultekin € L, Korotyaev BA. 2011. Lixus petiolicola n. sp. from Northeastern Turkey and Lixus furcatus Olivier: comparative systematic and ecological study (Coleoptera: Curculionidae: Lixinae). Annales de la Societe Entomologique de France 47: 101-111." type="journal article" year="2011">Korotyaev, 2011</bibRefCitation>
), most specimens are still missing. The type material of this species was collected from ‘Senegal’ by Geoffroy de Villeneuve, as well as its synonym
<taxonomicName id="4C75A1FEFFEEFFC8FE1AF96AFDD6F923" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[443,596,1726,1748]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="313" phylum="Arthropoda" rank="species" species="discoideus">
<emphasis id="B901066FFFEEFFC8FE1AF96AFDD6F923" box="[443,596,1726,1748]" italics="true" pageId="15" pageNumber="313">O. discoideus</emphasis>
</taxonomicName>
(which was recorded also from
<collectingCountry id="F3629AEDFFEEFFC8FE38F908FDBEF903" box="[409,572,1757,1779]" name="Goree Island" pageId="15" pageNumber="313">Goree Island</collectingCountry>
). As the type material of this species could not be located at present, no
<typeStatus id="54CE64DFFFEEFFC8FEBFF8CFFE06F8C0" box="[286,388,1818,1840]" pageId="15" pageNumber="313" type="lectotype">lectotype</typeStatus>
could be designed here. For the morphological account, please refer to the original description.
</paragraph>
<paragraph id="8BCADA7DFFEEFFC8FC8AFF12FB50FD13" blockId="15.[811,1426,199,740]" pageId="15" pageNumber="313">
<emphasis id="B901066FFFEEFFC8FC8AFF12FB51FF2C" box="[811,1235,199,220]" italics="true" pageId="15" pageNumber="313">
Geographical distribution (
<figureCitation id="134EC6F8FFEEFFC8FBD5FF12FB47FF2C" box="[1140,1221,199,220]" captionStart="Figure 14" captionStartId="12.[162,241,1389,1408]" captionTargetBox="[391,1208,203,1361]" captionTargetId="figure-188@12.[388,1212,198,1356]" captionTargetPageId="12" captionText="Figure 14. Distribution of the species (see Appendix 3, Supporting Information for a list of localities) with the Olson et al. (2001) terrestrial biomes classification." pageId="15" pageNumber="313">Fig. 14</figureCitation>
):
</emphasis>
The species is widely distributed in the whole sub-Saharan area [
<collectingCountry id="F3629AEDFFEEFFC8FC93FED1FCF8FEEA" box="[818,890,260,282]" name="Benin" pageId="15" pageNumber="313">Benin</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FC31FED1FBB5FEE9" box="[912,1079,260,282]" name="Burkina Faso" pageId="15" pageNumber="313">Burkina Faso</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FBECFED1FB20FEEA" box="[1101,1186,260,282]" name="Eritrea" pageId="15" pageNumber="313">Eritrea</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FB19FED1FA9EFEEA" box="[1208,1308,260,282]" name="Ethiopia" pageId="15" pageNumber="313">Ethiopia</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FA93FED1FA0CFEEA" box="[1330,1422,260,282]" name="Gambia" pageId="15" pageNumber="313">Gambia</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FC8AFEF7FCF8FEC8" box="[811,890,290,312]" name="Ghana" pageId="15" pageNumber="313">Ghana</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FC3AFEF7FC73FEC8" box="[923,1009,290,312]" name="Guinea-Bissau" pageId="15" pageNumber="313">Guinea</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FBB3FEF7FBE7FEC8" box="[1042,1125,290,312]" name="Guinea-Bissau" pageId="15" pageNumber="313">Guinea</collectingCountry>
<collectingRegion id="49B1149FFFEEFFC8FB23FEF7FB53FEC8" box="[1154,1233,290,312]" country="Guinea-Bissau" name="Bissau" pageId="15" pageNumber="313">Bissau</collectingRegion>
,
<collectingCountry id="F3629AEDFFEEFFC8FB53FEF6FA0CFEC7" box="[1266,1422,290,312]" name="Ivory Coast" pageId="15" pageNumber="313">Ivory Coast</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FC8AFE94FC32FEA7" box="[811,944,321,343]" name="Mauritania" pageId="15" pageNumber="313">Mauritania</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FC6FFE94FB93FEA7" box="[974,1041,321,343]" name="Niger" pageId="15" pageNumber="313">Niger</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FB8EFE94FB05FEA7" box="[1071,1159,321,343]" name="Nigeria" pageId="15" pageNumber="313">Nigeria</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FB07FE94FA82FEA7" box="[1190,1280,321,343]" name="Senegal" pageId="15" pageNumber="313">Senegal</collectingCountry>
(the
<typeStatus id="54CE64DFFFEEFFC8FAC0FE94FA10FEA6" box="[1377,1426,321,342]" pageId="15" pageNumber="313">type</typeStatus>
locality),
<collectingCountry id="F3629AEDFFEEFFC8FC00FE8AFC68FE85" box="[929,1002,351,373]" name="Sudan" pageId="15" pageNumber="313">Sudan</collectingCountry>
and
<collectingCountry id="F3629AEDFFEEFFC8FB9AFEB5FBF4FE85" box="[1083,1142,352,373]" name="Togo" pageId="15" pageNumber="313">Togo</collectingCountry>
], and in Central and Eastern Africa (
<collectingCountry id="F3629AEDFFEEFFC8FC5AFEABFAB4FE64" box="[1019,1334,382,404]" name="Central African Republic" pageId="15" pageNumber="313">Central African Republic</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FAEEFEABFA0CFE64" box="[1359,1422,382,404]" name="Chad" pageId="15" pageNumber="313">Chad</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FC8AFE48FB38FE42" box="[811,1210,413,435]" name="Democratic Republic of the Congo" pageId="15" pageNumber="313">Democratic Republic of Congo</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FB76FE48FAA6FE43" box="[1239,1316,413,435]" name="Gabon" pageId="15" pageNumber="313">Gabon</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FAE0FE48FA0FFE42" box="[1345,1421,413,434]" name="Kenya" pageId="15" pageNumber="313">Kenya</collectingCountry>
,
<collectingCountry id="F3629AEDFFEEFFC8FC8AFE6EFBA9FE20" box="[811,1067,443,465]" name="Republic of the Congo" pageId="15" pageNumber="313">Republic of Congo</collectingCountry>
and
<collectingCountry id="F3629AEDFFEEFFC8FB39FE6EFB74FE21" box="[1176,1270,443,465]" name="Uganda" pageId="15" pageNumber="313">Uganda</collectingCountry>
], extending eastwards and southwards toward
<collectingCountry id="F3629AEDFFEEFFC8FB48FE0FFAD6FE00" box="[1257,1364,474,496]" name="Tanzania" pageId="15" pageNumber="313">Tanzania</collectingCountry>
and
<collectingCountry id="F3629AEDFFEEFFC8FC8AFE2DFC02FDFE" box="[811,896,504,526]" name="Malawi" pageId="15" pageNumber="313">Malawi</collectingCountry>
(see Appendix 3, Supporting Information for more details). The species is also recorded from
<collectingRegion id="49B1149FFFEEFFC8FC8AFDE0FCEBFDBB" box="[811,873,565,587]" country="Egypt" name="Al Qahirah" pageId="15" pageNumber="313">Cairo</collectingRegion>
(
<collectingCountry id="F3629AEDFFEEFFC8FCDCFDE3FC46FDBB" box="[893,964,566,587]" name="Egypt" pageId="15" pageNumber="313">Egypt</collectingCountry>
,
<bibRefCitation id="EFE4A78CFFEEFFC8FC72FDE0FB24FDBB" author="Schatzmayr A" box="[979,1190,565,587]" pageId="15" pageNumber="313" pagination="40 - 84" refId="ref14065" refString="Schatzmayr A. 1946. Gli scarabaeidi coprofagi della Libia e dell'Egitto. Atti della Societa italiana di Scienze Naturali 85: 40-84." type="journal article" year="1946">Schatzmayr, 1946</bibRefCitation>
;
<bibRefCitation id="EFE4A78CFFEEFFC8FB17FDE0FAD6FDBB" author="Baraud J" box="[1206,1364,565,587]" pageId="15" pageNumber="313" pagination="1 - 652" refId="ref12423" refString="Baraud J. 1985. Coleopteres Scarabaeoidea, Faune du nord de l'Afrique, du Maroc au Sina € i. Encyclopedie Entomologique 46: 1-652." type="journal article" year="1985">Baraud, 1985</bibRefCitation>
) and Arabia (
<bibRefCitation id="EFE4A78CFFEEFFC8FC2FFD81FBB4FD99" author="Paulian R" box="[910,1078,596,618]" pageId="15" pageNumber="313" pagination="141 - 154" refId="ref13685" refString="Paulian R. 1980. Insects of Saudi Arabia. Coleoptera: Scarabaeoidea (1 ere contribution). Fauna of Saudi Arabia 2: 141-154." type="journal article" year="1980">Paulian, 1980</bibRefCitation>
), but these data need to be confirmed. Accidental introduction is reported in Antilles (
<collectingCountry id="F3629AEDFFEEFFC8FC37FD44FB98FD57" box="[918,1050,657,679]" name="Martinique" pageId="15" pageNumber="313">Martinique</collectingCountry>
), where an anthropogenic cause was hypothesized to explain these findings (
<bibRefCitation id="EFE4A78CFFEEFFC8FC92FD1BFC6FFD14" author="Matthews EG" box="[819,1005,718,740]" pageId="15" pageNumber="313" pagination="1 - 133" refId="ref13369" refString="Matthews EG. 1966. A taxonomic and zoogeographic survey of the Scarabaeinae of the Antilles. Memoirs of the American Entomological Society 21: 1-133." type="journal article" year="1966">Matthews, 1966</bibRefCitation>
;
<bibRefCitation id="EFE4A78CFFEEFFC8FC5BFD1BFB44FD13" author="Chalumeau F" box="[1018,1222,718,740]" pageId="15" pageNumber="313" refId="ref12780" refString="Chalumeau F. 1983. Les Coleopteres Scarabaeides des Petites Antilles (Guadeloupe a Martinique). Taxonomie- Ethologie-Biogeographie. Paris: Editions Lechevalier." type="book" year="1983">Chalumeau, 1983</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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(
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, 1908: 172)
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(
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2
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,
<figureCitation id="134EC6F8FFF0FFD6FE66FA6DFE5BFA22" box="[455,473,1464,1490]" captionStart="Figure 7" captionStartId="8.[162,241,917,936]" captionTargetBox="[391,1215,201,883]" captionTargetId="figure-27@8.[388,1212,198,884]" captionTargetPageId="8" captionText="Figure 7. Aedeagus of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA); see text for the codes. Scale bar = 0.5 mm." pageId="17" pageNumber="315">7</figureCitation>
,
<figureCitation id="134EC6F8FFF0FFD6FE49FA62FE7BFA21" box="[488,505,1463,1489]" captionStart="Figure 8" captionStartId="8.[162,241,1850,1869]" captionTargetBox="[391,1215,1064,1819]" captionTargetId="figure-59@8.[388,1212,1062,1817]" captionTargetPageId="8" captionText="Figure 8. Primary lamella of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA). Scale bar = 0.5 mm." pageId="17" pageNumber="315">8</figureCitation>
,
<figureCitation id="134EC6F8FFF0FFD6FDA8FA62FDAAFA21" box="[521,552,1463,1489]" captionStart="Figure 11" captionStartId="10.[162,241,707,726]" captionTargetBox="[391,1215,198,674]" captionTargetId="figure-136@10.[391,1215,198,674]" captionTargetPageId="10" captionText="Figure 11. Vagina and receptaculum seminis of the species of the genus Morettius (PA and UT). Scale bar = 0.5 mm." pageId="17" pageNumber="315">11</figureCitation>
)
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FF35F9DDFEF8F946" blockId="17.[148,763,1543,1718]" pageId="17" pageNumber="315">
<emphasis id="B901066FFFF0FFD6FF35F9DDFEC4F9EC" box="[148,326,1543,1565]" italics="true" pageId="17" pageNumber="315">Type material:</emphasis>
<materialsCitation id="3B1DD020FFF0FFD6FEF4F9D2FD75F9A9" collectionCode="MNHN" country="Chad" location="Chad lake" municipality="Kiao-Kata" pageId="17" pageNumber="315" specimenCount="1" stateProvince="Moyen-Chari" typeStatus="lectotype">
<typeStatus id="54CE64DFFFF0FFD6FEF4F9D2FE41F9EC" box="[341,451,1543,1564]" pageId="17" pageNumber="315" type="lectotype">Lectotype</typeStatus>
here designated: (male)
<collectingCountry id="F3629AEDFFF0FFD6FF35F9F3FF61F9CB" box="[148,227,1574,1595]" name="Chad" pageId="17" pageNumber="315">CHAD</collectingCountry>
:
<collectingMunicipality id="6BAE4007FFF0FFD6FF51F9F0FEEAF9CB" box="[240,360,1573,1595]" pageId="17" pageNumber="315">Kiao-Kata</collectingMunicipality>
,
<collectingRegion id="49B1149FFFF0FFD6FED7F9F0FD8CF9CB" box="[374,526,1573,1595]" country="Chad" name="Moyen-Chari" pageId="17" pageNumber="315">Moyen-Chari</collectingRegion>
, south to
<location id="8EAA8CA6FFF0FFD6FD24F9F0FD79F9CB" LSID="urn:lsid:plazi:treatment:03DC6B6BFFF0FFD5FF1DFA48FE91F989:8EAA8CA6FFF0FFD6FD24F9F0FD79F9CB" box="[645,763,1573,1595]" country="Chad" municipality="Kiao-Kata" name="Chad lake" pageId="17" pageNumber="315" stateProvince="Moyen-Chari">Chad lake</location>
[=moyen Chari rives, Kiao-Kata] [
<collectionCode id="ED6442B8FFF0FFD6FD37F991FD73F9A9" box="[662,753,1604,1625]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="315">MNHN</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="3B1DD020FFF0FFD6FF35F9B6FEF4F946" collectionCode="MNHN" country="Chad" location="Chad lake" municipality="Kiao-Kata" pageId="17" pageNumber="315" specimenCount="1" stateProvince="Moyen-Chari" typeStatus="paralectotype">
<typeStatus id="54CE64DFFFF0FFD6FF35F9B6FEB6F989" box="[148,308,1635,1657]" pageId="17" pageNumber="315" type="paralectotype">Paralectotype</typeStatus>
: (female) same locality [
<collectionCode id="ED6442B8FFF0FFD6FDC1F9B6FD39F988" box="[608,699,1635,1656]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="315">MNHN</collectionCode>
]. For the morphological account, please refer to the original description
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.
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FF35F908FEAFF89D" blockId="17.[148,763,1757,1901]" pageId="17" pageNumber="315">
<emphasis id="B901066FFFF0FFD6FF35F908FDB2F902" box="[148,560,1757,1778]" italics="true" pageId="17" pageNumber="315">
Geographical distribution (
<figureCitation id="134EC6F8FFF0FFD6FE70F908FDA0F902" box="[465,546,1757,1778]" captionStart="Figure 14" captionStartId="12.[162,241,1389,1408]" captionTargetBox="[391,1208,203,1361]" captionTargetId="figure-188@12.[388,1212,198,1356]" captionTargetPageId="12" captionText="Figure 14. Distribution of the species (see Appendix 3, Supporting Information for a list of localities) with the Olson et al. (2001) terrestrial biomes classification." pageId="17" pageNumber="315">Fig. 14</figureCitation>
):
</emphasis>
The species was reported from
<collectingCountry id="F3629AEDFFF0FFD6FEEBF929FE43F8E1" box="[330,449,1788,1809]" name="Cameroon" pageId="17" pageNumber="315">Cameroon</collectingCountry>
,
<collectingCountry id="F3629AEDFFF0FFD6FE74F92EFD96F8E1" box="[469,532,1787,1809]" name="Chad" pageId="17" pageNumber="315">Chad</collectingCountry>
,
<collectingCountry id="F3629AEDFFF0FFD6FD89F92EFDFFF8E1" box="[552,637,1787,1809]" name="Nigeria" pageId="17" pageNumber="315">Nigeria</collectingCountry>
(southern border),
<collectingCountry id="F3629AEDFFF0FFD6FF5EF8CFFE62F8DF" box="[255,480,1818,1840]" name="Republic of the Congo" pageId="17" pageNumber="315">Republic of Congo</collectingCountry>
and
<collectingCountry id="F3629AEDFFF0FFD6FD8FF8CFFDF5F8C0" box="[558,631,1818,1840]" name="Sudan" pageId="17" pageNumber="315">Sudan</collectingCountry>
(collection localities are listed in Appendix 3, Supporting Information).
</paragraph>
<caption id="DF0A8AF5FFF0FFD6FC8AF8EFFBB5F89B" pageId="17" pageNumber="315" startId="17.[811,890,1850,1869]" targetBox="[890,1344,1160,1818]" targetPageId="17" targetType="figure">
<paragraph id="8BCADA7DFFF0FFD6FC8AF8EFFBB5F89B" blockId="17.[811,1426,1850,1899]" pageId="17" pageNumber="315">
<emphasis id="B901066FFFF0FFD6FC8AF8EFFB4DF8BD" bold="true" box="[811,1231,1850,1869]" pageId="17" pageNumber="315">
Figure 17.
<taxonomicName id="4C75A1FEFFF0FFD6FC0FF8EFFBE1F8BD" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[942,1123,1850,1869]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete" status="sp. nov.">
<emphasis id="B901066FFFF0FFD6FC0FF8EFFBE1F8BD" bold="true" box="[942,1123,1850,1869]" italics="true" pageId="17" pageNumber="315">Morettius utete</emphasis>
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<taxonomicNameLabel id="A232BB14FFF0FFD6FBD1F8EFFB4DF8BD" box="[1136,1231,1850,1869]" pageId="17" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
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, paratype female facies. Scale bar = 1 mm.
</paragraph>
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<paragraph id="8BCADA7DFFF3FFD5FF03FF12FED0FDFE" blockId="18.[162,777,199,1382]" pageId="18" pageNumber="316">covering more than half the pronotal length and prolonged narrowly on the sides to reach the anterior angles. Pronotal pubescence black and very short, only evident on the lateral edges. Pronotum unarmed, very weakly transverse (length/width ratio: 0.55). Base evenly curved, markedly bordered. Posterior angles not sinuate. Anterior angles strongly sinuate, sharply projected outwards. Surface reticulate. Area surrounding the callus not granulate; remaining pronotal surface entirely covered by small granules.</paragraph>
<paragraph id="8BCADA7DFFF3FFD5FF1BFDC2FF78FC29" blockId="18.[162,777,199,1382]" pageId="18" pageNumber="316">
Elytra yellow with black symmetrical spots, one basal on the fifth interstriae, one on the first third of the sixth and seventh interstriae, another four connected on the middle of the second to fifth interstriae forming a zig-zig pattern. Juxtasutural interstriae yellow
<emphasis id="B901066FFFF3FFD5FE22FD67FE0DFD34" box="[387,399,690,708]" italics="true" pageId="18" pageNumber="316">–</emphasis>
orange, darkened anteriorly. Basal carina of interstriae bronze. Pubescence yellow, very short and scattered, only evident posteriorly. Elytral ground reticulate. Interstriae bearing small yellow granules, arranged on the juxtasutural interstriae in a regular row. Interstriae weakly convex, basally carinate. Striae narrow, well marked, yellow. Punctures of striae never wider than the striae. The seventh stria sinuate basally.
</paragraph>
<paragraph id="8BCADA7DFFF3FFD5FF1BFC34FEC7FBC4" blockId="18.[162,777,199,1382]" pageId="18" pageNumber="316">Pygidium yellow almost smooth, finely microreticulate, with small, rare, hardly perceptible punctures. Base carinate.</paragraph>
<paragraph id="8BCADA7DFFF3FFD5FF1BFBE8FEEAFB5F" blockId="18.[162,777,199,1382]" pageId="18" pageNumber="316">Epipleura yellow. Sternal thoracic surface bronze, except for the base of propleurae bronze. Abdominal sternites yellow. Metasternal pubescence scattered, short and yellow.</paragraph>
<paragraph id="8BCADA7DFFF3FFD5FF1BFB62FEC3FA96" blockId="18.[162,777,199,1382]" pageId="18" pageNumber="316">Coxae yellow. Trochanters bronze. Femura yellow, apically bronze. Tibiae bronze, meso- and metatibiae apically yellowish. Tarsomeres weakly bronze. Pubescence yellow. Fore tibiae three-toothed. Tibial spur elongate, bent inward, apically rounded. Tarsi normally shaped.</paragraph>
<paragraph id="8BCADA7DFFF3FFD5FF03FA58FE0DF9ED" blockId="18.[162,776,1421,1565]" pageId="18" pageNumber="316">
<emphasis id="B901066FFFF3FFD5FF03FA58FE19FA53" box="[162,411,1421,1443]" italics="true" pageId="18" pageNumber="316">Individual variation:</emphasis>
<typeStatus id="54CE64DFFFF3FFD5FE0BFA5BFD97FA53" box="[426,533,1422,1443]" pageId="18" pageNumber="316" type="paratype">Paratype</typeStatus>
: the wide posterior spot of the right elytra is extended on the sixth interstria. Spot of the scond interstria obviously longer than in the hotolypus. Seventh elytral stria only weakly sinuate.
</paragraph>
<paragraph id="8BCADA7DFFF3FFD5FF03F991FE91F989" blockId="18.[162,776,1604,1657]" pageId="18" pageNumber="316">
<emphasis id="B901066FFFF3FFD5FF03F991FE25F9A9" box="[162,423,1604,1626]" italics="true" pageId="18" pageNumber="316">
Epipharynx (
<figureCitation id="134EC6F8FFF3FFD5FEF7F990FE1BF9A9" box="[342,409,1604,1626]" captionStart="Figure 2" captionStartId="4.[162,241,1141,1160]" captionTargetBox="[391,1215,203,1113]" captionTargetId="figure-431@4.[391,1215,203,1113]" captionTargetPageId="4" captionText="Figure 2. Epipharynx of the species of the genera Hamonthophagus (AC = acutus, BI = bitubeculatus, DE = depressus, FA = fallax, LA = laceratus) and Morettius (PA = pallens, UT = utete). Scale bar = 0.5 mm." pageId="18" pageNumber="316">Fig. 2</figureCitation>
):
</emphasis>
See the above generic diagnosis.
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<taxonomicName id="4C75A1FEFFF0FFD6FC19FF03FB14FF1A" ID-CoL="44D83" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[952,1174,214,235]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete" status="sp. nov.">
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.
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.
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</paragraph>
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<subSubSection id="C36F89F6FFF0FFD6FC5CFF25FD16FA96" pageId="17" pageNumber="315" type="description">
<paragraph id="8BCADA7DFFF0FFD6FC5CFF25FB3DFEFA" blockId="17.[952,1286,210,266]" box="[1021,1215,240,266]" pageId="17" pageNumber="315">
(
<figureCitation id="134EC6F8FFF0FFD6FBA6FF27FBDAFEFA" box="[1031,1112,240,266]" captionStart="Figure 2" captionStartId="4.[162,241,1141,1160]" captionTargetBox="[391,1215,203,1113]" captionTargetId="figure-431@4.[391,1215,203,1113]" captionTargetPageId="4" captionText="Figure 2. Epipharynx of the species of the genera Hamonthophagus (AC = acutus, BI = bitubeculatus, DE = depressus, FA = fallax, LA = laceratus) and Morettius (PA = pallens, UT = utete). Scale bar = 0.5 mm." pageId="17" pageNumber="315">
<smallCapsWord id="8D2C4CA1FFF0FFD6FBA6FF27FBBEFEF9" baselines="260,261" box="[1031,1084,242,266]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="17" pageNumber="315">FIGS</smallCapsWord>
2
</figureCitation>
,
<figureCitation id="134EC6F8FFF0FFD6FBC6FF25FB0AFEFA" box="[1127,1160,240,266]" captionStart="Figure 11" captionStartId="10.[162,241,707,726]" captionTargetBox="[391,1215,198,674]" captionTargetId="figure-136@10.[391,1215,198,674]" captionTargetPageId="10" captionText="Figure 11. Vagina and receptaculum seminis of the species of the genus Morettius (PA and UT). Scale bar = 0.5 mm." pageId="17" pageNumber="315">11</figureCitation>
,
<figureCitation id="134EC6F8FFF0FFD6FB36FF25FB34FEFA" box="[1175,1206,240,266]" captionStart="Figure 17" captionStartId="17.[811,890,1850,1869]" captionTargetBox="[890,1344,1160,1818]" captionTargetId="figure-521@17.[892,1346,1159,1817]" captionTargetPageId="17" captionText="Figure 17. Morettius utete sp. nov., paratype female facies. Scale bar = 1 mm." pageId="17" pageNumber="315">17</figureCitation>
)
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FC8AFE94FC7AFE85" blockId="17.[811,1426,321,373]" pageId="17" pageNumber="315">
<emphasis id="B901066FFFF0FFD6FC8AFE94FC2BFEA6" box="[811,937,321,342]" italics="true" pageId="17" pageNumber="315">Etymology:</emphasis>
The species was named after the collection locality.
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FC8AFE48FB0DFE1F" blockId="17.[811,1427,413,496]" pageId="17" pageNumber="315">
<emphasis id="B901066FFFF0FFD6FC8AFE48FC74FE42" box="[811,1014,413,434]" italics="true" pageId="17" pageNumber="315">Type material:</emphasis>
<materialsCitation id="3B1DD020FFF0FFD6FBA4FE48FB08FE21" collectionCode="MHNL" country="Tanzania" location="Female" municipality="Utete-Rufijikindwjivi" pageId="17" pageNumber="315" specimenCount="1" typeStatus="holotype">
<emphasis id="B901066FFFF0FFD6FBA4FE48FBEDFE42" box="[1029,1135,413,434]" italics="true" pageId="17" pageNumber="315">
<typeStatus id="54CE64DFFFF0FFD6FBA4FE48FBEBFE42" box="[1029,1129,413,434]" pageId="17" pageNumber="315" type="holotype">Holotype</typeStatus>
:
</emphasis>
<location id="8EAA8CA6FFF0FFD6FBDCFE48FB56FE43" LSID="urn:lsid:plazi:treatment:03DC6B6BFFF0FFD6FC19FF03FD16FA96:8EAA8CA6FFF0FFD6FBDCFE48FB56FE43" box="[1149,1236,413,435]" country="Tanzania" municipality="Utete-Rufijikindwjivi" name="Female" pageId="17" pageNumber="315">Female</location>
,
<collectingCountry id="F3629AEDFFF0FFD6FAA5FE48FA0CFE42" box="[1284,1422,413,434]" name="Tanzania" pageId="17" pageNumber="315">TANZANIA</collectingCountry>
:
<collectingMunicipality id="6BAE4007FFF0FFD6FC8AFE6EFB9EFE21" box="[811,1052,443,465]" pageId="17" pageNumber="315">Utete-Rufijikindwjivi</collectingMunicipality>
[
<collectionCode id="ED6442B8FFF0FFD6FB8CFE69FB06FE21" box="[1069,1156,444,465]" country="France" httpUri="http://grbio.org/cool/4jmm-iazx" name="Musee Guimet d'Histoire Naturelle de Lyon" pageId="17" pageNumber="315">MHNL</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="3B1DD020FFF0FFD6FB39FE69FB09FE1F" collectionCode="MHNL, EBCT" country="Tanzania" location="Female" municipality="Utete-Rufijikindwjivi" pageId="17" pageNumber="315" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<typeStatus id="54CE64DFFFF0FFD6FB39FE69FA8CFE21" box="[1176,1294,444,465]" pageId="17" pageNumber="315" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7311F4FFF0FFD6FABDFE6EFA0CFE21" box="[1308,1422,443,465]" count="2" pageId="17" pageNumber="315" type="female">2 females</specimenCount>
, same locality [
<collectionCode id="ED6442B8FFF0FFD6FC6EFE0FFBA1FE1F" box="[975,1059,474,495]" country="France" httpUri="http://grbio.org/cool/4jmm-iazx" name="Musee Guimet d'Histoire Naturelle de Lyon" pageId="17" pageNumber="315">MHNL</collectionCode>
] [
<collectionCode id="ED6442B8FFF0FFD6FB99FE0FFB06FE1F" box="[1080,1156,474,495]" pageId="17" pageNumber="315">EBCT</collectionCode>
]
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.
</paragraph>
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<emphasis id="B901066FFFF0FFD6FC8AFDC2FC2FFDDC" box="[811,941,535,556]" italics="true" pageId="17" pageNumber="315">Description</emphasis>
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FC8AFDE0FC72FDBB" blockId="17.[811,1008,535,587]" box="[811,1008,565,587]" pageId="17" pageNumber="315">
<emphasis id="B901066FFFF0FFD6FC8AFDE0FCE9FDBA" box="[811,875,565,586]" italics="true" pageId="17" pageNumber="315">Male:</emphasis>
Unknown.
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FC8AFDA7FBA7FC07" blockId="17.[811,1426,626,1076]" pageId="17" pageNumber="315">
<emphasis id="B901066FFFF0FFD6FC8AFDA7FC07FD77" box="[811,901,626,647]" italics="true" pageId="17" pageNumber="315">Female:</emphasis>
Length:
<quantity id="4C8D7798FFF0FFD6FBBBFDA7FB20FD78" box="[1050,1186,626,648]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="6.6" metricValueMin="6.1" pageId="17" pageNumber="315" unit="mm" value="6.35" valueMax="6.6" valueMin="6.1">
6.1
<emphasis id="B901066FFFF0FFD6FB9DFDA0FBCAFD77" box="[1084,1096,629,647]" italics="true" pageId="17" pageNumber="315">–</emphasis>
6.6 mm
</quantity>
. Head bronze, transverse (length/width ratio: 0.72), with maximum width just anteriorly to the eyes. Clypeus sinuate, with clypeal edge reddish. Clypeo-genal junction not sinuate. Frontal carina fine, weakly curved, placed at the mid-length of the head, short and low, occupying half of the interocular space. Surface markedly reticulate. Clypeus covered by flat and transverse granules, more or less merged. Genal granules large and round. Vertex unarmed, weakly concave, with round and fine granules. Antennal scape normally shaped, not dentate or serrulate. Antennal club yellow.
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FCE5FBD5FD12FB1C" blockId="17.[811,1426,626,1076]" lastBlockId="17.[148,763,1176,1260]" pageId="17" pageNumber="315">
Pronotum bronze, with hind angles bearing a bronze callus surrounded by a wide yellowish area
<emphasis id="B901066FFFF0FFD6FF35FB4DFEA4FB5D" box="[148,294,1176,1197]" italics="true" pageId="17" pageNumber="315">Distribution:</emphasis>
The genus
<taxonomicName id="4C75A1FEFFF0FFD6FE63FB4CFDADFB5E" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[450,559,1177,1198]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="B901066FFFF0FFD6FE63FB4CFDADFB5E" box="[450,559,1177,1198]" italics="true" pageId="17" pageNumber="315">Morettius</emphasis>
</taxonomicName>
is characterized by a disjoint distribution, being found in central west Africa, and south-eastern Africa (
<collectingCountry id="F3629AEDFFF0FFD6FDB4FB03FD06FB1C" box="[533,644,1238,1260]" name="Tanzania" pageId="17" pageNumber="315">Tanzania</collectingCountry>
).
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FF35FAC6FD16FA96" blockId="17.[148,763,1299,1382]" pageId="17" pageNumber="315">
<emphasis id="B901066FFFF0FFD6FF35FAC6FE90FAD8" box="[148,274,1299,1320]" italics="true" pageId="17" pageNumber="315">Etymology:</emphasis>
The genus is named after our colleague, the French entomologist Philippe Moretto, who works extensively on African Scarabaeoidea.
</paragraph>
</subSubSection>
</treatment>
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216
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species:
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<taxonomicName id="4C75A1FEFFF1FFD7FC7BFECEFA08FEC0" authority="d'Orbigny, 1908" authorityName="d'Orbigny" authorityYear="1908" box="[986,1418,283,305]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="species" species="pallens">
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(
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)
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, and
<taxonomicName id="4C75A1FEFFF1FFD7FC99FEA2FC14FE7C" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[824,918,375,396]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="species" species="utete" status="sp. nov.">
<emphasis id="B901066FFFF1FFD7FC99FEA2FC14FE7C" box="[824,918,375,396]" italics="true" pageId="16" pageNumber="314">M. utete</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232BB14FFF1FFD7FC3FFEA2FC75FE7C" box="[926,1015,375,396]" pageId="16" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BCADA7DFFF1FFD7FC99FE7AFB28FD49" blockId="16.[824,1439,430,697]" pageId="16" pageNumber="314">
<emphasis id="B901066FFFF1FFD7FC99FE7AFC32FE34" box="[824,944,431,452]" italics="true" pageId="16" pageNumber="314">Diagnosis:</emphasis>
Species of the genus
<taxonomicName id="4C75A1FEFFF1FFD7FB17FE7AFAA1FE34" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[1206,1315,431,452]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B901066FFFF1FFD7FB17FE7AFAA1FE34" box="[1206,1315,431,452]" italics="true" pageId="16" pageNumber="314">Morettius</emphasis>
</taxonomicName>
(
<figureCitation id="134EC6F8FFF1FFD7FA97FE7BFA0AFE33" box="[1334,1416,430,452]" captionStart="Figure 3" captionStartId="5.[148,227,1580,1599]" captionTargetBox="[161,1409,195,1544]" captionTargetId="figure-178@5.[161,1409,195,1544]" captionTargetPageId="5" captionText="Figure 3. Scatterplots of the RW 1 and 2 of head, pronotum, right elytron and mentum (semilandmarks method). Only the specimens employed to build the matrix are shown here. The deformation grids corresponding to the minimum and maximum values of the axes are shown for each anatomical trait. See text for the codes." pageId="16" pageNumber="314">Fig. S3</figureCitation>
<emphasis id="B901066FFFF1FFD7FA32FE64FA1DFE33" box="[1427,1439,433,451]" italics="true" pageId="16" pageNumber="314">–</emphasis>
Appendix 4, Supporting Information) are characterized by the mostly rounded and only slightly notched anterior margin of the clypeus, and the pronotum covered by granules, sometimes mixed to points. The pygidium is always smooth, with some rade points. The species show a moderate sexual dimorphism in the fore tibiae, and the pygidium is larger in males than in females.
</paragraph>
<paragraph id="8BCADA7DFFF1FFD7FC99FD08FAAFFB6F" blockId="16.[824,1440,733,1183]" pageId="16" pageNumber="314">
<emphasis id="B901066FFFF1FFD7FC99FD08FC47FD02" box="[824,965,733,754]" italics="true" pageId="16" pageNumber="314">Epipharynx:</emphasis>
The epipharynx (
<figureCitation id="134EC6F8FFF1FFD7FB01FD08FB66FD02" box="[1184,1252,733,755]" captionStart="Figure 2" captionStartId="4.[162,241,1141,1160]" captionTargetBox="[391,1215,203,1113]" captionTargetId="figure-431@4.[391,1215,203,1113]" captionTargetPageId="4" captionText="Figure 2. Epipharynx of the species of the genera Hamonthophagus (AC = acutus, BI = bitubeculatus, DE = depressus, FA = fallax, LA = laceratus) and Morettius (PA = pallens, UT = utete). Scale bar = 0.5 mm." pageId="16" pageNumber="314">Fig. 2</figureCitation>
) fore margin is arched, with a largely V-shaped notch in the middle. The corypha is reduced, the chaetopariae are arched with short, thick and almost equal-length setae. The pubescence of the haptomerum is dense. The proplegmatium is subequal along the whole length, with the posterior triangular sclerotization reaching at least as much or more than half the length of the anterior epitorma, which is rectilinear, well sclerotized and thin. The base of the triangular sclerotization reaches the small, thick and upwardturned apophyses. Laeotorma and dexiotorma are symmetrical, short and stout. Pternotormae are well sclerotized, and the crepis is short but evident, with a sharp apex. The plegmatic area is visible.
</paragraph>
<paragraph id="8BCADA7DFFF1FFD7FC99FB14FBE0FA3C" blockId="16.[824,1439,1217,1484]" pageId="16" pageNumber="314">
<emphasis id="B901066FFFF1FFD7FC99FB14FC7CFB26" box="[824,1022,1217,1238]" italics="true" pageId="16" pageNumber="314">Male genitalia:</emphasis>
Only the male genitalia of
<taxonomicName id="4C75A1FEFFF1FFD7FC99FB34FC30FB05" baseAuthorityName="d'Orbigny" baseAuthorityYear="1908" box="[824,946,1248,1270]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B901066FFFF1FFD7FC99FB34FC30FB05" box="[824,946,1248,1270]" italics="true" pageId="16" pageNumber="314">M. pallens</emphasis>
</taxonomicName>
could be examined. The phallobase (
<figureCitation id="134EC6F8FFF1FFD7FCE0FB2AFC06FAE4" box="[833,900,1279,1301]" captionStart="Figure 7" captionStartId="8.[162,241,917,936]" captionTargetBox="[391,1215,201,883]" captionTargetId="figure-27@8.[388,1212,198,884]" captionTargetPageId="8" captionText="Figure 7. Aedeagus of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA); see text for the codes. Scale bar = 0.5 mm." pageId="16" pageNumber="314">Fig. 7</figureCitation>
) is short, only slightly arched, slender and of equal size along the whole length. The parameres are symmetrical, and squared, with a small tip at the apex, and a small, rounded protrusion ventrally. The internal sac (
<figureCitation id="134EC6F8FFF1FFD7FB8EFAACFBF0FA7E" box="[1071,1138,1401,1423]" captionStart="Figure 8" captionStartId="8.[162,241,1850,1869]" captionTargetBox="[391,1215,1064,1819]" captionTargetId="figure-59@8.[388,1212,1062,1817]" captionTargetPageId="8" captionText="Figure 8. Primary lamella of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA). Scale bar = 0.5 mm." pageId="16" pageNumber="314">Fig. 8</figureCitation>
) is membranous, with various well-sclerotized parts differing greatly from
<taxonomicName id="4C75A1FEFFF1FFD7FC99FA63FBE0FA3C" box="[824,1122,1462,1484]" class="Insecta" family="Scarabaeidae" genus="Hamonthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B901066FFFF1FFD7FC99FA63FB80FA3B" box="[824,1026,1462,1483]" italics="true" pageId="16" pageNumber="314">Hamonthophagus</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="8BCADA7DFFF1FFD7FC99FA3BFAE2F907" blockId="16.[824,1439,1518,1784]" pageId="16" pageNumber="314">
<emphasis id="B901066FFFF1FFD7FC99FA3BFB89F9F3" box="[824,1035,1518,1539]" italics="true" pageId="16" pageNumber="314">Female genitalia:</emphasis>
The female genitalia are very peculiar (
<figureCitation id="134EC6F8FFF1FFD7FC0AF9D9FC7EF9D1" box="[939,1020,1548,1570]" captionStart="Figure 11" captionStartId="10.[162,241,707,726]" captionTargetBox="[391,1215,198,674]" captionTargetId="figure-136@10.[391,1215,198,674]" captionTargetPageId="10" captionText="Figure 11. Vagina and receptaculum seminis of the species of the genus Morettius (PA and UT). Scale bar = 0.5 mm." pageId="16" pageNumber="314">Fig. 11</figureCitation>
), as the vagina of both species is entirely membranous, and no sclerotization is present at all. Furthermore, the infundibular tube is lowered as in
<taxonomicName id="4C75A1FEFFF1FFD7FBACF9BDFB55F98D" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[1037,1239,1640,1661]" class="Insecta" family="Scarabaeidae" genus="Hamonthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B901066FFFF1FFD7FBACF9BDFB55F98D" box="[1037,1239,1640,1661]" italics="true" pageId="16" pageNumber="314">Hamonthophagus</emphasis>
</taxonomicName>
, but here an expanded portion is identifiable in the central part of the vagina, which is differently shaped in the two species. The receptaculum seminis is sickle-shaped, namely slim, arched and apically sharp (
<figureCitation id="134EC6F8FFF1FFD7FAAFF937FAD0F907" box="[1294,1362,1762,1784]" captionStart="Figure 9" captionStartId="9.[148,227,955,974]" captionTargetBox="[376,1200,195,918]" captionTargetId="figure-117@9.[376,1200,195,918]" captionTargetPageId="9" captionText="Figure 9. Above, the receptaculum seminis; the features that characterize the two models are clearly represented by H. bituberculatus (BI) and M. pallens (PA). Scale bar = 0.2 mm. Below, a generic example of the vagina sclerotization (H. bituberculatus), after being cleared from membranes and cut off. The various parts were numbered (see text for further details)." pageId="16" pageNumber="314">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BCADA7DFFF1FFD7FC99F8CFFB37F89D" blockId="16.[824,1439,1818,1901]" pageId="16" pageNumber="314">
<emphasis id="B901066FFFF1FFD7FC99F8CFFB98F8DF" box="[824,1050,1818,1839]" italics="true" pageId="16" pageNumber="314">Specific diagnosis:</emphasis>
These species can be easily distinguished on the basis of external morphology, epipharynx and female genitalia.
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FF0CFF12FE32FE21" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
The pronotum in
<taxonomicName id="4C75A1FEFFF0FFD6FEDFFF12FE75FF2C" baseAuthorityName="d'Orbigny" baseAuthorityYear="1908" box="[382,503,199,220]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B901066FFFF0FFD6FEDFFF12FE75FF2C" box="[382,503,199,220]" italics="true" pageId="17" pageNumber="315">M. pallens</emphasis>
</taxonomicName>
is covered by distinct rasping points mixed with smaller, yellow granules; the granules of the rasping points instead are large, darker than the background surface, and carry long, thick and light yellow setae. The pronotum of
<taxonomicName id="4C75A1FEFFF0FFD6FF35FEB5FF70FE85" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[148,242,352,373]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
<emphasis id="B901066FFFF0FFD6FF35FEB5FF70FE85" box="[148,242,352,373]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
</taxonomicName>
is covered by only a few rasping setigerous points with thin, yellow setae and many small granules, which are very thick, evenly coloured as the base and without points.
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FF0CFE0FFF7BFDDC" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
Elytral striae of
<taxonomicName id="4C75A1FEFFF0FFD6FE20FE0FFE78FE1F" baseAuthorityName="d'Orbigny" baseAuthorityYear="1908" box="[385,506,474,495]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B901066FFFF0FFD6FE20FE0FFE78FE1F" box="[385,506,474,495]" italics="true" pageId="17" pageNumber="315">M. pallens</emphasis>
</taxonomicName>
are as large as the points, being instead larger than the points in
<taxonomicName id="4C75A1FEFFF0FFD6FF35FDC2FF70FDDC" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[148,242,535,556]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
<emphasis id="B901066FFFF0FFD6FF35FDC2FF70FDDC" box="[148,242,535,556]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FF0CFDE0FDFEFD56" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
The smooth pygidium carries in
<taxonomicName id="4C75A1FEFFF0FFD6FDE4FDE3FD3CFDBA" baseAuthorityName="d'Orbigny" baseAuthorityYear="1908" box="[581,702,565,587]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B901066FFFF0FFD6FDE4FDE3FD3CFDBA" box="[581,702,565,587]" italics="true" pageId="17" pageNumber="315">M. pallens</emphasis>
</taxonomicName>
few, small, rade and deep setigerous points that are not granulated, and in
<taxonomicName id="4C75A1FEFFF0FFD6FE2DFDA6FE6BFD78" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[396,489,627,648]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
<emphasis id="B901066FFFF0FFD6FE2DFDA6FE6BFD78" box="[396,489,627,648]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
</taxonomicName>
only some large and superficial vanishing points without setae.
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FF0CFD7AFDA7FCB0" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
The epipharynx (see
<figureCitation id="134EC6F8FFF0FFD6FE69FD7AFD89FD35" box="[456,523,687,709]" captionStart="Figure 2" captionStartId="4.[162,241,1141,1160]" captionTargetBox="[391,1215,203,1113]" captionTargetId="figure-431@4.[391,1215,203,1113]" captionTargetPageId="4" captionText="Figure 2. Epipharynx of the species of the genera Hamonthophagus (AC = acutus, BI = bitubeculatus, DE = depressus, FA = fallax, LA = laceratus) and Morettius (PA = pallens, UT = utete). Scale bar = 0.5 mm." pageId="17" pageNumber="315">Fig. 2</figureCitation>
for a comparison among species) has the characteristic shape of
<taxonomicName id="4C75A1FEFFF0FFD6FD11FD1AFE9CFCF3" pageId="17" pageNumber="315">
<emphasis id="B901066FFFF0FFD6FD11FD1AFF3DFCF2" italics="true" pageId="17" pageNumber="315">Morettius</emphasis>
species
</taxonomicName>
, but can be distinguished from
<taxonomicName id="4C75A1FEFFF0FFD6FD3CFD38FD78FCF2" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[669,762,749,770]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
<emphasis id="B901066FFFF0FFD6FD3CFD38FD78FCF2" box="[669,762,749,770]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
</taxonomicName>
by the more developed apotormae, and the more slender laeotorma and dexiotorma.
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FF0CFC9CFD79FC29" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
In both species, the vagina is wholly desclerotized, but carries two globose symmetrical expansions that encircle the desclerotized and lowered infundibulum in
<taxonomicName id="4C75A1FEFFF0FFD6FF35FC70FE8EFC49" baseAuthorityName="d'Orbigny" baseAuthorityYear="1908" box="[148,268,932,954]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="B901066FFFF0FFD6FF35FC70FE8EFC49" box="[148,268,932,954]" italics="true" pageId="17" pageNumber="315">M. pallens</emphasis>
</taxonomicName>
, while in
<taxonomicName id="4C75A1FEFFF0FFD6FE29FC70FE67FC4A" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[392,485,933,954]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
<emphasis id="B901066FFFF0FFD6FE29FC70FE67FC4A" box="[392,485,933,954]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
</taxonomicName>
there is a single, large expansion (see
<figureCitation id="134EC6F8FFF0FFD6FE9AFC16FE09FC28" box="[315,395,963,985]" captionStart="Figure 11" captionStartId="10.[162,241,707,726]" captionTargetBox="[391,1215,198,674]" captionTargetId="figure-136@10.[391,1215,198,674]" captionTargetPageId="10" captionText="Figure 11. Vagina and receptaculum seminis of the species of the genus Morettius (PA and UT). Scale bar = 0.5 mm." pageId="17" pageNumber="315">Fig. 11</figureCitation>
for a comparison among species).
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FF0CFC34FE68FBE6" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
As the male of
<taxonomicName id="4C75A1FEFFF0FFD6FEFAFC37FE3AFC07" authorityName="Roggero &amp; Dierkens &amp; Barbero &amp; Palestrini" authorityYear="2017" box="[347,440,994,1015]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
<emphasis id="B901066FFFF0FFD6FEFAFC37FE3AFC07" box="[347,440,994,1015]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
</taxonomicName>
is unknown, no comparison can be made between species.
</paragraph>
<paragraph id="8BCADA7DFFF0FFD6FF35FBE8FED2FB81" blockId="17.[148,763,1085,1137]" pageId="17" pageNumber="315">
<emphasis id="B901066FFFF0FFD6FF35FBE8FEDCFBA2" box="[148,350,1085,1106]" italics="true" pageId="17" pageNumber="315">General remarks:</emphasis>
No preimaginal stages have been described so far.
</paragraph>
</subSubSection>
</treatment>
</document>

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(
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, 1913: 471)
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(
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2
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,
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,
<figureCitation id="134EC6F8FFF1FFD7FE54FDF7FD84FDCC" box="[501,518,546,572]" captionStart="Figure 8" captionStartId="8.[162,241,1850,1869]" captionTargetBox="[391,1215,1064,1819]" captionTargetId="figure-59@8.[388,1212,1062,1817]" captionTargetPageId="8" captionText="Figure 8. Primary lamella of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA). Scale bar = 0.5 mm." pageId="16" pageNumber="314">8</figureCitation>
,
<figureCitation id="134EC6F8FFF1FFD7FDB7FDF7FDB7FDCC" box="[534,565,546,572]" captionStart="Figure 10" captionStartId="9.[148,227,1743,1762]" captionTargetBox="[373,1197,1135,1709]" captionTargetId="figure-119@9.[373,1197,1135,1709]" captionTargetPageId="9" captionText="Figure 10. Vagina and receptaculum seminis of the species of the genus Hamonthophagus (AC, BI, DE, FA and LA). Scale bar = 0.5 mm." pageId="16" pageNumber="314">10</figureCitation>
)
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<emphasis id="B901066FFFF1FFD7FF03FDA6FED1FD77" box="[162,339,626,648]" italics="true" pageId="16" pageNumber="314">Type material:</emphasis>
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<typeStatus id="54CE64DFFFF1FFD7FEC3FDA6FE52FD78" box="[354,464,627,648]" pageId="16" pageNumber="314" type="lectotype">Lectotype</typeStatus>
here designated: (male)
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: [Nyassa
<collectingRegion id="49B1149FFFF1FFD7FE3DFD44FE6AFD57" box="[412,488,657,679]" country="Malawi" name="Zomba" pageId="16" pageNumber="314">Zomba</collectingRegion>
haut Chire=]
<taxonomicName id="4C75A1FEFFF1FFD7FD14FD44FF5DFD35" authority=", Shire" authorityName="Shire" class="Insecta" family="Scarabaeidae" genus="Zomba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="genus">Zomba, Shire</taxonomicName>
river upper course,
<location id="8EAA8CA6FFF1FFD7FE4CFD7AFD0EFD35" LSID="urn:lsid:plazi:treatment:03DC6B6BFFF1FFD7FF07FDDDFEB8FB60:8EAA8CA6FFF1FFD7FE4CFD7AFD0EFD35" box="[493,652,687,709]" country="Malawi" name="Malawi Lake" pageId="16" pageNumber="314" stateProvince="Zomba">Malawi Lake</location>
[
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.
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<typeStatus id="54CE64DFFFF1FFD7FF03FD1BFEC0FD14" box="[162,322,718,740]" pageId="16" pageNumber="314" type="paralectotype">Paralectotype</typeStatus>
: (female)
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[=Afrique Or Alem]:
<collectingRegion id="49B1149FFFF1FFD7FEBAFD39FE46FCF2" box="[283,452,748,770]" country="Tanzania" name="Dar es Salaam" pageId="16" pageNumber="314">Dar-es-Salaam</collectingRegion>
[
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]. For the morphological account, please refer to the original description
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.
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<emphasis id="B901066FFFF1FFD7FF03FCB2FDBEFC8C" box="[162,572,871,892]" italics="true" pageId="16" pageNumber="314">
Geographical distribution (
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):
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The species has been described from
<collectingCountry id="F3629AEDFFF1FFD7FE3EFC50FE76FC6B" box="[415,500,901,923]" name="Malawi" pageId="16" pageNumber="314">Malawi</collectingCountry>
and
<collectingCountry id="F3629AEDFFF1FFD7FD9BFC50FD25FC6B" box="[570,679,901,923]" name="Tanzania" pageId="16" pageNumber="314">Tanzania</collectingCountry>
, and at present is recorded from
<collectingCountry id="F3629AEDFFF1FFD7FDAEFC71FD00FC49" box="[527,642,932,953]" name="Botswana" pageId="16" pageNumber="314">Botswana</collectingCountry>
,
<collectingCountry id="F3629AEDFFF1FFD7FD00FC71FC81FC4A" box="[673,771,932,954]" name="Burundi" pageId="16" pageNumber="314">Burundi</collectingCountry>
,
<collectingCountry id="F3629AEDFFF1FFD7FF03FC17FDA8FC28" box="[162,554,962,984]" name="Democratic Republic of the Congo" pageId="16" pageNumber="314">Democratic Republic of Congo</collectingCountry>
,
<collectingCountry id="F3629AEDFFF1FFD7FDE4FC16FD13FC28" box="[581,657,963,984]" name="Kenya" pageId="16" pageNumber="314">Kenya</collectingCountry>
,
<collectingCountry id="F3629AEDFFF1FFD7FD0DFC17FC86FC28" box="[684,772,962,984]" name="Malawi" pageId="16" pageNumber="314">Malawi</collectingCountry>
,
<collectingCountry id="F3629AEDFFF1FFD7FF03FC34FE85FC07" box="[162,263,993,1015]" name="Namibia" pageId="16" pageNumber="314">Namibia</collectingCountry>
,
<collectingCountry id="F3629AEDFFF1FFD7FEBAFC34FE04FC07" box="[283,390,993,1015]" name="Tanzania" pageId="16" pageNumber="314">Tanzania</collectingCountry>
and
<collectingCountry id="F3629AEDFFF1FFD7FE70FC34FDA8FC07" box="[465,554,993,1015]" name="Zambia" pageId="16" pageNumber="314">Zambia</collectingCountry>
. The record from Graaf-Reinet (
<collectingRegion id="49B1149FFFF1FFD7FEEBFBD5FDD9FBE6" box="[330,603,1024,1046]" country="South Africa" name="Eastern Cape" pageId="16" pageNumber="314">Eastern Cape province</collectingRegion>
,
<collectingCountry id="F3629AEDFFF1FFD7FDCAFBD5FC83FBE6" box="[619,769,1024,1046]" name="South Africa" pageId="16" pageNumber="314">South Africa</collectingCountry>
) is very interesting, but needs to be confirmed by further records. A detailed list of the localities is given in the Appendix 3 (Supporting Information).
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Linnaeus, 1758
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<taxonomicName id="44865DFBFFB2FF9185C8902BFE75FAEA" ID-CoL="PC3" authority="Bourgeois, 1882: 97" authorityName="Bourgeois" authorityPageNumber="97" authorityYear="1882" box="[148,473,1329,1351]" class="Insecta" family="Lycidae" genus="Adoceta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus" status="syn. nov.">
<emphasis id="B1F2FA6AFFB2FF9185C8902BFF41FAEA" box="[148,237,1329,1350]" italics="true" pageId="6" pageNumber="53">Adoceta</emphasis>
<bibRefCitation id="E7175B89FFB2FF9185AA902BFE75FAEA" author="Bourgeois J" box="[246,473,1329,1351]" pageId="6" pageNumber="53" pagination="1 - 120" refId="ref10962" refString="Bourgeois J. 1882. Monographie des Lycides de l' ancienmonde. L' Abeille: Journal d' Entomologie 20: 1-120." type="journal article" year="1882">Bourgeois, 1882: 97</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<taxonomicNameLabel id="AAC14711FFB2FF9184B9902BFDE0FAEA" box="[485,588,1329,1350]" pageId="6" pageNumber="53">syn. nov.</taxonomicNameLabel>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CB9C75F3FFB2FF9185CF9075FB42FD0B" pageId="6" pageNumber="53" type="materials_examined">
<paragraph id="83392678FFB2FF9185CF9075FEBFFA0E" blockId="6.[147,762,1390,1442]" pageId="6" pageNumber="53">
<emphasis id="B1F2FA6AFFB2FF9185CF9075FE9DFA28" box="[147,305,1391,1412]" italics="true" pageId="6" pageNumber="53">
<typeStatus id="5C3D98DAFFB2FF9185CF9075FF65FA28" box="[147,201,1391,1412]" pageId="6" pageNumber="53">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="44865DFBFFB2FF91841C9074FD6FFA2F" authority="Bourgeois, 1882" authorityName="Bourgeois" authorityYear="1882" box="[320,707,1390,1412]" class="Insecta" family="Lycidae" genus="Adoceta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="caroli">
<emphasis id="B1F2FA6AFFB2FF91841C9074FE42FA2F" box="[320,494,1390,1411]" italics="true" pageId="6" pageNumber="53">Adoceta caroli</emphasis>
<bibRefCitation id="E7175B89FFB2FF91875E9074FD6FFA2F" author="Bourgeois J" box="[514,707,1390,1412]" pageId="6" pageNumber="53" pagination="1 - 120" refId="ref10962" refString="Bourgeois J. 1882. Monographie des Lycides de l' ancienmonde. L' Abeille: Journal d' Entomologie 20: 1-120." type="journal article" year="1882">Bourgeois, 1882</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
<paragraph id="83392678FFB2FF9185CF90D0FBFBFF57" blockId="6.[147,762,1482,1871]" lastBlockId="6.[810,1425,199,251]" pageId="6" pageNumber="53">
<emphasis id="B1F2FA6AFFB2FF9185CF90D0FDF9FA73" box="[147,597,1482,1503]" italics="true" pageId="6" pageNumber="53">
Differential diagnosis:
<taxonomicName id="44865DFBFFB2FF9184E090D0FDF9FA73" authorityName="Mulsant" authorityYear="1838" box="[444,597,1482,1503]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">Lygistopterus</taxonomicName>
</emphasis>
is widely distributed and differs from the Old World genera
<taxonomicName id="44865DFBFFB2FF9185CF931DFE88F9B0" box="[147,292,1543,1564]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9185CF931DFE88F9B0" box="[147,292,1543,1564]" italics="true" pageId="6" pageNumber="53">Micronychus</emphasis>
</taxonomicName>
and
<taxonomicName id="44865DFBFFB2FF9184D59312FE47F9B1" authorityName="Motyka &amp; Masek &amp; Bocak" authorityYear="2017" box="[393,491,1544,1565]" class="Insecta" family="Lycidae" genus="Parantis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B1F2FA6AFFB2FF9184D59312FE47F9B1" box="[393,491,1544,1565]" italics="true" pageId="6" pageNumber="53">Parantis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAC14711FFB2FF918755931DFD29F9B0" box="[521,645,1543,1564]" pageId="6" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
in the membranous internal sac (
<figureCitation id="1BBD3AFDFFB2FF9184AC933FFDCAF997" box="[496,614,1573,1595]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="6" pageNumber="53">Fig. 5I, J</figureCitation>
) and from
<taxonomicName id="44865DFBFFB2FF9185CF935EFE84F9F5" authorityName="Guerin-Meneville" authorityYear="1833" box="[147,296,1604,1625]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9185CF935EFE84F9F5" box="[147,296,1604,1625]" italics="true" pageId="6" pageNumber="53">Calochromus</emphasis>
</taxonomicName>
by the presence of sharp pronotal lateral edges. This genus occurs sympatrically with
<taxonomicName id="44865DFBFFB2FF9185CF9398FF5DF93B" box="[147,241,1666,1687]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9185CF9398FF5DF93B" box="[147,241,1666,1687]" italics="true" pageId="6" pageNumber="53">Lucaina</emphasis>
</taxonomicName>
and
<taxonomicName id="44865DFBFFB2FF918464939BFE73F93A" authorityName="Green" authorityYear="1954" box="[312,479,1665,1686]" class="Insecta" family="Lycidae" genus="Caloptognatha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF918464939BFE73F93A" box="[312,479,1665,1686]" italics="true" pageId="6" pageNumber="53">Caloptognatha</emphasis>
</taxonomicName>
in the Nearctic region and differs in the absence of long erected setae in the elytra. Neotropical
<taxonomicName id="44865DFBFFB2FF9184F693A4FD2EF97F" authorityName="Pic" authorityYear="1929" box="[426,642,1726,1747]" class="Insecta" family="Lycidae" genus="Macrolygistopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9184F693A4FD2EF97F" box="[426,642,1726,1747]" italics="true" pageId="6" pageNumber="53">Macrolygistopterus</emphasis>
</taxonomicName>
differs in the shape of male genitalia (
<figureCitation id="1BBD3AFDFFB2FF9184B693C7FDE8F95F" box="[490,580,1757,1779]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="6" pageNumber="53">Fig. 5N</figureCitation>
). The length of the rostrum is variable in
<taxonomicName id="44865DFBFFB2FF91849893E6FDF1F8BD" authorityName="Mulsant" authorityYear="1838" box="[452,605,1788,1809]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF91849893E6FDF1F8BD" box="[452,605,1788,1809]" italics="true" pageId="6" pageNumber="53">Lygistopterus</emphasis>
</taxonomicName>
; for example,
<taxonomicName id="44865DFBFFB2FF9185CF9201FE65F89C" authorityName="Linnaeus" authorityYear="1758" box="[147,457,1819,1840]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="sanguineus">
<emphasis id="B1F2FA6AFFB2FF9185CF9201FE65F89C" box="[147,457,1819,1840]" italics="true" pageId="6" pageNumber="53">Lygistopterus sanguineus</emphasis>
</taxonomicName>
has a well-developed rostrum, but the rostrum is absent in the closely related East Palearctic species or extremely short in the North African species.
</paragraph>
<paragraph id="83392678FFB2FF9186769438FAD1FE94" blockId="6.[810,1405,290,312]" box="[810,1405,290,312]" pageId="6" pageNumber="53">
<emphasis id="B1F2FA6AFFB2FF9186769438FC10FE9B" box="[810,956,290,311]" italics="true" pageId="6" pageNumber="53">Distribution:</emphasis>
Palearctic, Nearctic, Oriental regions.
</paragraph>
<paragraph id="83392678FFB2FF9186769445FB42FD0B" blockId="6.[810,1425,351,679]" pageId="6" pageNumber="53">
<emphasis id="B1F2FA6AFFB2FF9186769445FC3AFED8" box="[810,918,351,372]" italics="true" pageId="6" pageNumber="53">Remarks:</emphasis>
<taxonomicName id="44865DFBFFB2FF9186F89445FB6FFED9" authority="Bourgeois, 1882" authorityName="Bourgeois" authorityYear="1882" box="[932,1219,351,373]" class="Insecta" family="Lycidae" genus="Adoceta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9186F89445FC52FED8" box="[932,1022,351,372]" italics="true" pageId="6" pageNumber="53">Adoceta</emphasis>
<bibRefCitation id="E7175B89FFB2FF9181569445FB6FFED9" author="Bourgeois J" box="[1034,1219,351,373]" pageId="6" pageNumber="53" pagination="1 - 120" refId="ref10962" refString="Bourgeois J. 1882. Monographie des Lycides de l' ancienmonde. L' Abeille: Journal d' Entomologie 20: 1-120." type="journal article" year="1882">Bourgeois, 1882</bibRefCitation>
</taxonomicName>
is proposed as a junior synonym of
<taxonomicName id="44865DFBFFB2FF91814A9465FB03FE38" authorityName="Mulsant" authorityYear="1838" box="[1046,1199,383,404]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF91814A9465FB03FE38" box="[1046,1199,383,404]" italics="true" pageId="6" pageNumber="53">Lygistopterus</emphasis>
</taxonomicName>
as
<taxonomicName id="44865DFBFFB2FF9181BA9464FAE5FE3F" authorityName="Bourgeois" authorityYear="1882" box="[1254,1353,382,403]" class="Insecta" family="Lycidae" genus="Adoceta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="caroli">
<emphasis id="B1F2FA6AFFB2FF9181BA9464FAE5FE3F" box="[1254,1353,382,403]" italics="true" pageId="6" pageNumber="53">A. caroli</emphasis>
</taxonomicName>
(
<typeStatus id="5C3D98DAFFB2FF9180039464FA3DFE3F" box="[1375,1425,382,403]" pageId="6" pageNumber="53">type</typeStatus>
species) has a membranous internal sac and cannot be distinguished from
<taxonomicName id="44865DFBFFB2FF91816294A6FB7BFE7D" authorityName="Mulsant" authorityYear="1838" box="[1086,1239,444,465]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF91816294A6FB7BFE7D" box="[1086,1239,444,465]" italics="true" pageId="6" pageNumber="53">Lygistopterus</emphasis>
</taxonomicName>
. All sequenced species from sub-Saharan Africa represent a distant clade. They have a sclerotized internal sac and form a subclade within
<taxonomicName id="44865DFBFFB2FF918157970DFB30FD80" box="[1035,1180,535,556]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF918157970DFB30FD80" box="[1035,1180,535,556]" italics="true" pageId="6" pageNumber="53">Micronychus</emphasis>
</taxonomicName>
. Two North African species are placed in
<taxonomicName id="44865DFBFFB2FF91812C972CFAA5FDE7" authorityName="Mulsant" authorityYear="1838" box="[1136,1289,566,587]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF91812C972CFAA5FDE7" box="[1136,1289,566,587]" italics="true" pageId="6" pageNumber="53">Lygistopterus</emphasis>
</taxonomicName>
and the Neotropical species of
<taxonomicName id="44865DFBFFB2FF918173974FFACEFDC5" authority="Kleine, 1933" authorityName="Kleine" authorityYear="1933" box="[1071,1378,596,618]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF918173974FFB64FDC6" box="[1071,1224,597,618]" italics="true" pageId="6" pageNumber="53">Lygistopterus</emphasis>
<bibRefCitation id="E7175B89FFB2FF91818D974EFACEFDC5" author="Kleine R" box="[1233,1378,596,618]" pageId="6" pageNumber="53" refId="ref11706" refString="Kleine R. 1933. Coleopterorum Catalogus auspiciis et auxilio W. Junk editus S. Schenkling. Pars 128: Lycidae. Berlin: W. Junk." type="journal volume" year="1933">Kleine, 1933</bibRefCitation>
</taxonomicName>
are transferred to
<taxonomicName id="44865DFBFFB2FF91868F9768FB07FD2B" authorityName="Pic" authorityYear="1929" box="[979,1195,626,647]" class="Insecta" family="Lycidae" genus="Macrolygistopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF91868F9768FB07FD2B" box="[979,1195,626,647]" italics="true" pageId="6" pageNumber="53">Macrolygistopterus</emphasis>
</taxonomicName>
as earlier proposed by
<bibRefCitation id="E7175B89FFB2FF918611978BFC6FFD0A" author="Pic M" box="[845,963,657,679]" pageId="6" pageNumber="53" pagination="1 - 36" refId="ref12562" refString="Pic M. 1929. Nouveautes diverses. M elanges <20> Exotico-Entomologiques 53: 1-36." type="journal article" year="1929">Pic (1929)</bibRefCitation>
. See Appendix for details.
</paragraph>
</subSubSection>
</treatment>
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<paragraph id="83392678FFB2FF9186769611FAB8FC8D" blockId="6.[810,1300,779,801]" box="[810,1300,779,801]" pageId="6" pageNumber="53">
<heading id="D8719114FFB2FF9186769611FAB8FC8D" box="[810,1300,779,801]" centered="true" fontSize="9" level="2" pageId="6" pageNumber="53" reason="2">
<treatmentCitationGroup id="A3960156FFB2FF9186769611FAB8FC8D" box="[810,1300,779,801]" pageId="6" pageNumber="53">
<treatmentCitation id="02270069FFB2FF9186769611FABDFC8D" author="Guerin-Meneville FE" box="[810,1297,779,801]" page="158" pageId="6" pageNumber="53" year="1833">
<taxonomicName id="44865DFBFFB2FF9186769611FABDFC8D" ID-CoL="3FSN" authority="Guerin-Meneville, 1833: 158" authorityName="Guerin-Meneville" authorityPageNumber="158" authorityYear="1833" box="[810,1297,779,801]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9186769611FC13FC8C" box="[810,959,779,800]" italics="true" pageId="6" pageNumber="53">Calochromus</emphasis>
<bibRefCitation id="E7175B89FFB2FF9186949611FABDFC8D" author="Guerin-Meneville FE" box="[968,1297,779,801]" pageId="6" pageNumber="53" pagination="155 - 160" refId="ref11470" refString="Guerin-Meneville FE. 1833. Memoire sur deux nouveaux genres de l'ordre des Coleopteres et description des especes qui les composent. Annales de la Societe entomologique de France 2: 155-160." type="journal article" year="1833">Guerin-Meneville, 1833: 158</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="CB9C75F3FFB2FF9186769653FB3DFAC9" pageId="6" pageNumber="53" type="materials_examined">
<paragraph id="83392678FFB2FF9186769653FB3EFCD0" blockId="6.[810,1425,840,893]" pageId="6" pageNumber="53">
<emphasis id="B1F2FA6AFFB2FF9186769653FC63FCF2" box="[810,975,841,862]" italics="true" pageId="6" pageNumber="53">
<typeStatus id="5C3D98DAFFB2FF9186769653FCCCFCF2" box="[810,864,841,862]" pageId="6" pageNumber="53">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="44865DFBFFB2FF9186829652FC73FCD0" authority="Guerin-Meneville, 1833" authorityName="Guerin-Meneville" authorityYear="1833" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="glaucopterus">
<emphasis id="B1F2FA6AFFB2FF9186829652FAB2FCF1" box="[990,1310,840,861]" italics="true" pageId="6" pageNumber="53">Calochromus glaucopterus</emphasis>
<bibRefCitation id="E7175B89FFB2FF9180649652FC73FCD0" author="Guerin-Meneville FE" pageId="6" pageNumber="53" pagination="155 - 160" refId="ref11470" refString="Guerin-Meneville FE. 1833. Memoire sur deux nouveaux genres de l'ordre des Coleopteres et description des especes qui les composent. Annales de la Societe entomologique de France 2: 155-160." type="journal article" year="1833">Guerin-Meneville, 1833</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
<paragraph id="83392678FFB2FF91867696BEFB3DFAC9" blockId="6.[810,1425,932,1382]" pageId="6" pageNumber="53">
<emphasis id="B1F2FA6AFFB2FF91867696BEFB72FC15" box="[810,1246,932,953]" italics="true" pageId="6" pageNumber="53">
Differential diagnosis:
<taxonomicName id="44865DFBFFB2FF91811596BEFB72FC15" authorityName="Guerin-Meneville" authorityYear="1833" box="[1097,1246,932,953]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">Calochromus</taxonomicName>
</emphasis>
differs from other genera in the widely rounded obtuse lateral edge of the pronotum, relatively long parameres (
<figureCitation id="1BBD3AFDFFB2FF91866E96E5FC2AFBB9" box="[818,902,1023,1045]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="6" pageNumber="53">Fig. 5L</figureCitation>
) and the lateral process of the phallobase widely rounded (
<figureCitation id="1BBD3AFDFFB2FF9186A39104FBF9FB98" box="[1023,1109,1054,1076]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="6" pageNumber="53">Fig. 5L</figureCitation>
), unlike in
<taxonomicName id="44865DFBFFB2FF9181A49105FA3DFB98" authorityName="Kleine" authorityYear="1933" box="[1272,1425,1055,1076]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9181A49105FA3DFB98" box="[1272,1425,1055,1076]" italics="true" pageId="6" pageNumber="53">Lygistopterus</emphasis>
</taxonomicName>
where it is pointed.
<taxonomicName id="44865DFBFFB2FF9181699127FB66FBFE" authorityName="Guerin-Meneville" authorityYear="1833" box="[1077,1226,1085,1106]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9181699127FB66FBFE" box="[1077,1226,1085,1106]" italics="true" pageId="6" pageNumber="53">Calochromus</emphasis>
</taxonomicName>
has a compact cranium, unlike
<taxonomicName id="44865DFBFFB2FF91815D9146FBF3FBDD" box="[1025,1119,1116,1137]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF91815D9146FBF3FBDD" box="[1025,1119,1116,1137]" italics="true" pageId="6" pageNumber="53">Lucaina</emphasis>
</taxonomicName>
and
<taxonomicName id="44865DFBFFB2FF9181ED9141FA25FBDC" authorityName="Pic" authorityYear="1929" box="[1201,1417,1115,1136]" class="Insecta" family="Lycidae" genus="Macrolygistopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9181ED9141FA25FBDC" box="[1201,1417,1115,1136]" italics="true" pageId="6" pageNumber="53">Macrolygistopterus</emphasis>
</taxonomicName>
, each of which has a long rostrum, or some
<taxonomicName id="44865DFBFFB2FF9186769183FC6FFB02" authorityName="Kleine" authorityYear="1933" box="[810,963,1177,1198]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9186769183FC6FFB02" box="[810,963,1177,1198]" italics="true" pageId="6" pageNumber="53">Lygistopterus</emphasis>
</taxonomicName>
and
<taxonomicName id="44865DFBFFB2FF9186A39182FB3CFB01" box="[1023,1168,1176,1197]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9186A39182FB3CFB01" box="[1023,1168,1176,1197]" italics="true" pageId="6" pageNumber="53">Micronychus</emphasis>
</taxonomicName>
with a short rostrum.
<taxonomicName id="44865DFBFFB2FF91867691ADFC17FB60" box="[810,955,1207,1228]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF91867691ADFC17FB60" box="[810,955,1207,1228]" italics="true" pageId="6" pageNumber="53">Micronychus</emphasis>
</taxonomicName>
and
<taxonomicName id="44865DFBFFB2FF91814E91A2FBD8FB61" authorityName="Motyka &amp; Masek &amp; Bocak" authorityYear="2017" box="[1042,1140,1208,1229]" class="Insecta" family="Lycidae" genus="Parantis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B1F2FA6AFFB2FF91814E91A2FBD8FB61" box="[1042,1140,1208,1229]" italics="true" pageId="6" pageNumber="53">Parantis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAC14711FFB2FF9181D691ADFAADFB60" box="[1162,1281,1207,1228]" pageId="6" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
, previously placed in
<taxonomicName id="44865DFBFFB2FF9186F991CFFB96FB46" authorityName="Guerin-Meneville" authorityYear="1833" box="[933,1082,1237,1258]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9186F991CFFB96FB46" box="[933,1082,1237,1258]" italics="true" pageId="6" pageNumber="53">Calochromus</emphasis>
</taxonomicName>
, have a sclerotized internal sac (
<figureCitation id="1BBD3AFDFFB2FF91862C91EEFC47FAA5" box="[880,1003,1268,1290]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="6" pageNumber="53">Fig. 5I, J</figureCitation>
). We define three lineages in
<taxonomicName id="44865DFBFFB2FF9186769009FC13FA84" authorityName="Guerin-Meneville" authorityYear="1833" box="[810,959,1299,1320]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9186769009FC13FA84" box="[810,959,1299,1320]" italics="true" pageId="6" pageNumber="53">Calochromus</emphasis>
</taxonomicName>
with subgeneric rank:
<taxonomicName id="44865DFBFFB2FF9181889009FC32FAEB" authority="Pic, 1925" authorityName="Pic" authorityYear="1925" class="Insecta" family="Lycidae" genus="Flabellochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB2FF9181889009FA3DFA84" box="[1236,1425,1299,1320]" italics="true" pageId="6" pageNumber="53">Flabellochromus</emphasis>
<bibRefCitation id="E7175B89FFB2FF918676902BFC32FAEB" author="Pic M" box="[810,926,1329,1351]" pageId="6" pageNumber="53" pagination="5 - 8" refId="ref12547" refString="Pic M. 1925. Malacodermes exotiques. L'Echange 419: 5-8." type="journal article" year="1925">Pic, 1925</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="44865DFBFFB2FF9186EF902BFBE4FAEA" authorityName="Guerin-Meneville" authorityYear="1833" box="[947,1096,1329,1350]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B1F2FA6AFFB2FF9186EF902BFBE4FAEA" box="[947,1096,1329,1350]" italics="true" pageId="6" pageNumber="53">Calochromus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAC14711FFB2FF9181069028FBD2FAEB" box="[1114,1150,1330,1351]" pageId="6" pageNumber="53" sensu="stricto">
<emphasis id="B1F2FA6AFFB2FF9181069028FBDAFAEB" box="[1114,1142,1330,1351]" italics="true" pageId="6" pageNumber="53">s.s</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="44865DFBFFB2FF918196902BFA3CFAEA" authorityName="Motyka &amp; Masek &amp; Bocak" authorityYear="2017" box="[1226,1424,1329,1350]" pageId="6" pageNumber="53" rank="subGenus" status="subgen. nov." subGenus="Paracalochromus">
<emphasis id="B1F2FA6AFFB2FF918196902BFA3CFAEA" box="[1226,1424,1329,1350]" italics="true" pageId="6" pageNumber="53">Paracalochromus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAC14711FFB2FF918676904AFC16FAC9" box="[810,954,1360,1382]" pageId="6" pageNumber="53" rank="subGenus">subgen. nov.</taxonomicNameLabel>
(
<figureCitation id="1BBD3AFDFFB2FF918696904AFB29FAC9" box="[970,1157,1359,1382]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="6" pageNumber="53">
Fig. 5B
<emphasis id="B1F2FA6AFFB2FF9181429048FB86FAC8" box="[1054,1066,1362,1380]" italics="true" pageId="6" pageNumber="53">–</emphasis>
D, L
<emphasis id="B1F2FA6AFFB2FF9181019048FBC5FAC8" box="[1117,1129,1362,1380]" italics="true" pageId="6" pageNumber="53">–</emphasis>
M
</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<heading id="D8719114FFB3FF9085BB9635FD65FCE9" box="[231,713,811,838]" centered="true" fontSize="9" level="2" pageId="7" pageNumber="54" reason="2">
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<taxonomicName id="44865DFBFFB3FF9084359634FD65FCE9" authority="PIC, 1925" authorityName="Pic" authorityYear="1925" box="[361,713,811,838]" class="Insecta" family="Lycidae" genus="Flabellochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="genus">
<smallCapsWord id="85DFB0A4FFB3FF9084359634FDE4FCEF" baselines="831,831" box="[361,584,814,837]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Flabellochromus" pageId="7" pageNumber="54">
<emphasis id="B1F2FA6AFFB3FF9084359634FDE4FCEF" box="[361,584,814,837]" italics="true" pageId="7" pageNumber="54">FLABELLOCHROMUS</emphasis>
</smallCapsWord>
<bibRefCitation id="E7175B89FFB3FF90870E9637FD65FCE9" author="Pic M" box="[594,713,811,838]" pageId="7" pageNumber="54" pagination="5 - 8" refId="ref12547" refString="Pic M. 1925. Malacodermes exotiques. L'Echange 419: 5-8." type="journal article" year="1925">
<smallCapsWord id="85DFB0A4FFB3FF90870E9637FDD5FCEF" baselines="831,831" box="[594,633,813,837]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Pic" pageId="7" pageNumber="54">PIC</smallCapsWord>
, 1925
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</heading>
</paragraph>
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<emphasis id="B1F2FA6AFFB3FF9085F89666FEECFC3D" box="[164,320,892,913]" italics="true" pageId="7" pageNumber="54">
<typeStatus id="5C3D98DAFFB3FF9085F89666FF76FC3D" box="[164,218,892,913]" pageId="7" pageNumber="54">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="44865DFBFFB3FF9084139661FD7AFC3D" authority="Pic, 1925" authorityName="Pic" authorityYear="1925" box="[335,726,891,913]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="B1F2FA6AFFB3FF9084139661FDF8FC3C" box="[335,596,891,912]" italics="true" pageId="7" pageNumber="54">Calochromus pallidus</emphasis>
<bibRefCitation id="E7175B89FFB3FF9087399661FD7AFC3D" author="Pic M" box="[613,726,891,913]" pageId="7" pageNumber="54" pagination="5 - 8" refId="ref12547" refString="Pic M. 1925. Malacodermes exotiques. L'Echange 419: 5-8." type="journal article" year="1925">Pic, 1925</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
<paragraph id="83392678FFB3FF9085E196A2FEF3FC62" blockId="7.[164,779,891,974]" box="[189,351,952,974]" pageId="7" pageNumber="54">
<figureCitation id="1BBD3AFDFFB3FF9085E196A2FEF7FC62" box="[189,347,952,974]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="7" pageNumber="54">Figure 5C, M</figureCitation>
.
</paragraph>
<paragraph id="83392678FFB3FF9085F896EFFD82FBCA" blockId="7.[164,779,1013,1127]" pageId="7" pageNumber="54">
<emphasis id="B1F2FA6AFFB3FF9085F896EFFE09FBA6" box="[164,421,1013,1034]" italics="true" pageId="7" pageNumber="54">Differential diagnosis:</emphasis>
All males of
<taxonomicName id="44865DFBFFB3FF90871296EFFCA7FBA6" authorityName="Pic" authorityYear="1925" box="[590,779,1013,1034]" class="Insecta" family="Lycidae" genus="Flabellochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB3FF90871296EFFCA7FBA6" box="[590,779,1013,1034]" italics="true" pageId="7" pageNumber="54">Flabellochromus</emphasis>
</taxonomicName>
have flabellate antennae, unlike other Calochromine genera.
<taxonomicName id="44865DFBFFB3FF90845A9128FE6FFBEB" authorityName="Pic" authorityYear="1925" box="[262,451,1074,1095]" class="Insecta" family="Lycidae" genus="Flabellochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB3FF90845A9128FE6FFBEB" box="[262,451,1074,1095]" italics="true" pageId="7" pageNumber="54">Flabellochromus</emphasis>
</taxonomicName>
were recovered in a sister position to
<taxonomicName id="44865DFBFFB3FF908478914BFE15FBCA" authorityName="Guerin-Meneville" authorityYear="1833" box="[292,441,1105,1126]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB3FF908478914BFE15FBCA" box="[292,441,1105,1126]" italics="true" pageId="7" pageNumber="54">Calochromus</emphasis>
</taxonomicName>
(
<figureCitation id="1BBD3AFDFFB3FF908496914BFD8DFBCA" box="[458,545,1105,1127]" captionStart="Figure 1" captionStartId="3.[164,243,1304,1323]" captionTargetBox="[172,1438,198,1271]" captionTargetId="figure-200@3.[172,1438,198,1271]" captionTargetPageId="3" captionText="Figure 1. A, phylogenetic hypothesis for Calochromini based on the maximum-likelihood analysis of the MUSCLE alignment. Numbers at the branches are maximum-likelihood bootstrap values and Bayesian posterior probabilities. The complete tree with outgroups and full-branch lengths is shown in Fig. S8 (Supporting Information). B, discrete phylogeography reconstruction using a Bayesian approach as implemented in BEAST 2.1; pie charts represent posterior probabilities of biogeographical origins. C, geographical distribution chart of the Calochromini genera." figureDoi="http://doi.org/10.5281/zenodo.14807186" httpUri="https://zenodo.org/record/14807186/files/figure.png" pageId="7" pageNumber="54">Fig. 1A</figureCitation>
).
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<emphasis id="B1F2FA6AFFB3FF9085F89194FE9AFB0F" box="[164,310,1166,1187]" italics="true" pageId="7" pageNumber="54">Distribution:</emphasis>
Himalayas, Indo-Burma and the Great Sundas.
</paragraph>
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<taxonomicName id="44865DFBFFB3FF90866793C8FB0FF944" authority="Pic, 1929: 1" authorityName="Pic" authorityPageNumber="1" authorityYear="1929" box="[827,1187,1746,1768]" class="Insecta" family="Lycidae" genus="Macrolygistopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFB3FF90866793C8FBBFF94B" box="[827,1043,1746,1767]" italics="true" pageId="7" pageNumber="54">Macrolygistopterus</emphasis>
<bibRefCitation id="E7175B89FFB3FF90814093C8FB0FF944" author="Pic M" box="[1052,1187,1746,1768]" pageId="7" pageNumber="54" pagination="1 - 36" refId="ref12562" refString="Pic M. 1929. Nouveautes diverses. M elanges <20> Exotico-Entomologiques 53: 1-36." type="journal article" year="1929">Pic, 1929: 1</bibRefCitation>
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.
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<paragraph id="83392678FFB3FF908667920AFC17F8EF" blockId="7.[827,1442,1807,1891]" pageId="7" pageNumber="54">
<emphasis id="B1F2FA6AFFB3FF908667920AFC7AF889" box="[827,982,1808,1829]" italics="true" pageId="7" pageNumber="54">
<typeStatus id="5C3D98DAFFB3FF908667920AFCDDF889" box="[827,881,1808,1829]" pageId="7" pageNumber="54">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="44865DFBFFB3FF9086B9920AFAC2F889" authority="Latreille, 1811" authorityName="Latreille" authorityYear="1811" box="[997,1390,1807,1829]" class="Insecta" family="Lycidae" genus="Lycus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="succinetus">
<emphasis id="B1F2FA6AFFB3FF9086B9920AFB01F889" box="[997,1197,1808,1829]" italics="true" pageId="7" pageNumber="54">Lycus succinetus</emphasis>
Latreille, 1811
</taxonomicName>
(by monotypy).
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.
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<emphasis id="B1F2FA6AFFBCFF9F85CF90BCFF52FA15" bold="true" box="[147,254,1446,1465]" pageId="8" pageNumber="55">Figure 4.</emphasis>
A
<emphasis id="B1F2FA6AFFBCFF9F844690B2FE89FA14" box="[282,293,1448,1464]" italics="true" pageId="8" pageNumber="55">–</emphasis>
M,
<taxonomicName id="44865DFBFFBCFF9F841690BCFDF9FA14" authorityName="Linnaeus" authorityYear="1758" box="[330,597,1446,1464]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="sanguineus">
<emphasis id="B1F2FA6AFFBCFF9F841690BCFDF9FA14" box="[330,597,1446,1464]" italics="true" pageId="8" pageNumber="55">Lygistopterus sanguineus</emphasis>
</taxonomicName>
. A, head dorsal view; B, head ventral; C, antenna; D, labium; E, maxilla; F, mandible; G, labrum and hypopharynx; H, thorax ventrally; I, pronotum dorsally; J, pronotum ventrally; K, mesonotum; L, elytron; M, tarsus. Scale bars: A
<emphasis id="B1F2FA6AFFBCFF9F875A90F9FDBDFA5F" box="[518,529,1507,1523]" italics="true" pageId="8" pageNumber="55">–</emphasis>
C, I, J, L = 1 mm; D
<emphasis id="B1F2FA6AFFBCFF9F87B590F9FD58FA5F" box="[745,756,1507,1523]" italics="true" pageId="8" pageNumber="55">–</emphasis>
H, K, M = 0.25 mm.
</paragraph>
</caption>
<paragraph id="83392678FFBCFF9F85CF932FFE8BF8F1" blockId="8.[147,762,1589,1886]" pageId="8" pageNumber="55">
<emphasis id="B1F2FA6AFFBCFF9F85CF932FFD23F9E6" box="[147,655,1589,1610]" italics="true" pageId="8" pageNumber="55">
Differential diagnosis:
<taxonomicName id="44865DFBFFBCFF9F84EB932FFD23F9E6" authorityName="Pic" authorityYear="1929" box="[439,655,1589,1610]" class="Insecta" family="Lycidae" genus="Macrolygistopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="genus">Macrolygistopterus</taxonomicName>
</emphasis>
differs from other New World
<taxonomicName id="44865DFBFFBCFF9F84ED9349FDE6F9C5" box="[433,586,1619,1641]" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="58" phylum="Arthropoda" rank="tribe" tribe="Calochromini">Calochromini</taxonomicName>
genera in the slender basal part and widened apical third of parameres.
<taxonomicName id="44865DFBFFBCFF9F847E938BFE2CF90A" box="[290,384,1681,1702]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBCFF9F847E938BFE2CF90A" box="[290,384,1681,1702]" italics="true" pageId="8" pageNumber="55">Lucaina</emphasis>
</taxonomicName>
has the similar slender basal part of the parameres but the apex is characteristically modified (
<figureCitation id="1BBD3AFDFFBCFF9F848093D4FD9EF948" box="[476,562,1742,1764]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="8" pageNumber="55">Fig. 5O</figureCitation>
) and this genus has, similar to
<taxonomicName id="44865DFBFFBCFF9F840A93F6FE51F8AD" authorityName="Green" authorityYear="1954" box="[342,509,1772,1793]" class="Insecta" family="Lycidae" genus="Caloptognatha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBCFF9F840A93F6FE51F8AD" box="[342,509,1772,1793]" italics="true" pageId="8" pageNumber="55">Caloptognatha</emphasis>
</taxonomicName>
, long erect setae in the elytra.
<taxonomicName id="44865DFBFFBCFF9F847A9216FE13F88D" authorityName="Kleine" authorityYear="1933" box="[294,447,1804,1825]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBCFF9F847A9216FE13F88D" box="[294,447,1804,1825]" italics="true" pageId="8" pageNumber="55">Lygistopterus</emphasis>
</taxonomicName>
is distantly related and differs by its wide, parallel-sided parameres (
<figureCitation id="1BBD3AFDFFBCFF9F85C09252FEB7F8F1" box="[156,283,1864,1886]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="8" pageNumber="55">Fig. 5K, N</figureCitation>
).
</paragraph>
<paragraph id="83392678FFBCFF9F8676932FFC3DF9C5" blockId="8.[810,1425,1589,1641]" pageId="8" pageNumber="55">
<emphasis id="B1F2FA6AFFBCFF9F8676932FFC10F9E6" box="[810,956,1589,1610]" italics="true" pageId="8" pageNumber="55">Distribution:</emphasis>
<collectingCountry id="FB9166E8FFBCFF9F8697932FFBB2F9E7" box="[971,1054,1589,1611]" name="Mexico" pageId="8" pageNumber="55">Mexico</collectingCountry>
, Central America, South America.
</paragraph>
</subSubSection>
</treatment>
</document>

242
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<paragraph id="83392678FFBCFF9F867693D4FBF0F94F" blockId="8.[810,1116,1742,1763]" box="[810,1116,1742,1763]" pageId="8" pageNumber="55">
<heading id="D8719114FFBCFF9F867693D4FBF0F94F" box="[810,1116,1742,1763]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="55" reason="2">
<treatmentCitationGroup id="A3960156FFBCFF9F867693D4FBF0F94F" box="[810,1116,1742,1763]" pageId="8" pageNumber="55">
<treatmentCitation id="02270069FFBCFF9F867693D4FBF4F94F" author="Duges E" box="[810,1112,1742,1763]" page="180" pageId="8" pageNumber="55" year="1879">
<taxonomicName id="44865DFBFFBCFF9F867693D4FBF4F94F" ID-CoL="62Y3S" authority="Duges, 1879: 180" authorityName="Duges" authorityPageNumber="180" authorityYear="1879" box="[810,1112,1742,1763]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBCFF9F867693D4FC24F94F" box="[810,904,1742,1763]" italics="true" pageId="8" pageNumber="55">Lucaina</emphasis>
<bibRefCitation id="E7175B89FFBCFF9F86CC93D4FBF4F94F" author="Duges E" box="[912,1112,1742,1763]" pageId="8" pageNumber="55" pagination="167 - 188" refId="ref11283" refString="Duges E. 1879. Descripcion de coleopteros indigenas. La Naturaleza Mexicana 4: 167-188." type="journal article" year="1879">Duges, 1879: 180</bibRefCitation>
</taxonomicName>
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.
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<paragraph id="83392678FFBCFF9F86769211FC06F892" blockId="8.[810,1425,1803,1886]" pageId="8" pageNumber="55">
<emphasis id="B1F2FA6AFFBCFF9F86769211FC63F88C" box="[810,975,1803,1824]" italics="true" pageId="8" pageNumber="55">
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species:
</emphasis>
<taxonomicName id="44865DFBFFBCFF9F86829211FAFFF88C" authority="Duges, 1879" authorityName="Duges" authorityYear="1879" box="[990,1363,1803,1824]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="schini">
<emphasis id="B1F2FA6AFFBCFF9F86829211FB37F88C" box="[990,1179,1803,1824]" italics="true" pageId="8" pageNumber="55">Lucaina schini</emphasis>
<bibRefCitation id="E7175B89FFBCFF9F81E99211FAFFF88C" author="Duges E" box="[1205,1363,1803,1824]" pageId="8" pageNumber="55" pagination="167 - 188" refId="ref11283" refString="Duges E. 1879. Descripcion de coleopteros indigenas. La Naturaleza Mexicana 4: 167-188." type="journal article" year="1879">Duges, 1879</bibRefCitation>
</taxonomicName>
(by monotypy).
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<figureCitation id="1BBD3AFDFFBCFF9F861F9252FC70F8F1" box="[835,988,1864,1886]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="8" pageNumber="55">Figure 5E, O</figureCitation>
.
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<paragraph id="83392678FFBDFF9E85F89335FB25F979" blockId="9.[164,1442,1583,1749]" pageId="9" pageNumber="56">
<emphasis id="B1F2FA6AFFBDFF9E85F89335FEA2F9EE" bold="true" box="[164,270,1583,1602]" pageId="9" pageNumber="56">Figure 5.</emphasis>
A
<emphasis id="B1F2FA6AFFBDFF9E84779328FE9AF9EE" box="[299,310,1586,1602]" italics="true" pageId="9" pageNumber="56">–</emphasis>
H, general appearance: A,
<taxonomicName id="44865DFBFFBDFF9E873E932AFCDCF9EE" authorityName="Linnaeus" authorityYear="1758" box="[610,880,1584,1602]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="sanguineus">
<emphasis id="B1F2FA6AFFBDFF9E873E932AFCDCF9EE" box="[610,880,1584,1602]" italics="true" pageId="9" pageNumber="56">Lygistopterus sanguineus</emphasis>
</taxonomicName>
; B,
<taxonomicName id="44865DFBFFBDFF9E86F99335FB82F9EE" authorityName="Guerin-Meneville" authorityYear="1833" box="[933,1070,1583,1602]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBDFF9E86F99335FB82F9EE" box="[933,1070,1583,1602]" italics="true" pageId="9" pageNumber="56">Calochromus</emphasis>
</taxonomicName>
(
<emphasis id="B1F2FA6AFFBDFF9E811E932AFBF0F9EE" box="[1090,1116,1584,1602]" italics="true" pageId="9" pageNumber="56">s.s</emphasis>
.,
<taxonomicName id="44865DFBFFBDFF9E81299335FB56F9EE" authorityName="Guerin-Meneville" authorityYear="1833" box="[1141,1274,1583,1602]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="glaucopterus">
<emphasis id="B1F2FA6AFFBDFF9E81299335FB56F9EE" box="[1141,1274,1583,1602]" italics="true" pageId="9" pageNumber="56">glaucopterus</emphasis>
</taxonomicName>
; C,
<emphasis id="B1F2FA6AFFBDFF9E80739335FAEAF9EE" box="[1327,1350,1583,1602]" italics="true" pageId="9" pageNumber="56">C.</emphasis>
(
<taxonomicName id="44865DFBFFBDFF9E80059335FEBCF9F3" authorityName="Pic" authorityYear="1925" class="Insecta" family="Lycidae" genus="Flabellochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBDFF9E80059335FEBCF9F3" italics="true" pageId="9" pageNumber="56">Flabellochromus</emphasis>
</taxonomicName>
,
<taxonomicName id="44865DFBFFBDFF9E84439356FE02F9F3" box="[287,430,1612,1631]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="kelantanensis">
<emphasis id="B1F2FA6AFFBDFF9E84439356FE02F9F3" box="[287,430,1612,1631]" italics="true" pageId="9" pageNumber="56">kelantanensis</emphasis>
</taxonomicName>
; D,
<taxonomicName id="44865DFBFFBDFF9E84809357FC80F9F3" authorityName="Pic" authorityYear="1913" box="[476,812,1612,1632]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="giganteus" subGenus="Paracalochromus">
<emphasis id="B1F2FA6AFFBDFF9E84809357FE5FF9CC" box="[476,499,1613,1632]" italics="true" pageId="9" pageNumber="56">C.</emphasis>
(
<emphasis id="B1F2FA6AFFBDFF9E875E9356FD14F9F3" box="[514,696,1612,1631]" italics="true" pageId="9" pageNumber="56">Paracalochromus</emphasis>
)
<emphasis id="B1F2FA6AFFBDFF9E879B9357FC80F9F3" box="[711,812,1613,1631]" italics="true" pageId="9" pageNumber="56">giganteus</emphasis>
</taxonomicName>
; E,
<taxonomicName id="44865DFBFFBDFF9E86059357FB8BF9F3" authorityName="Horn" authorityYear="1885" box="[857,1063,1612,1631]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="discoidalis">
<emphasis id="B1F2FA6AFFBDFF9E86059357FB8BF9F3" box="[857,1063,1612,1631]" italics="true" pageId="9" pageNumber="56">Lucaina discoidalis</emphasis>
</taxonomicName>
; F,
<taxonomicName id="44865DFBFFBDFF9E810E9356FA37F9F3" authorityName="Motyka &amp; Masek &amp; Bocak" authorityYear="2017" baseAuthorityName="Leconte" baseAuthorityYear="1875" box="[1106,1435,1612,1631]" class="Insecta" family="Lycidae" genus="Macrolygistopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="rubripennis">
<emphasis id="B1F2FA6AFFBDFF9E810E9356FA37F9F3" box="[1106,1435,1612,1631]" italics="true" pageId="9" pageNumber="56">Macrolygistopterus rubripennis</emphasis>
</taxonomicName>
; G,
<taxonomicName id="44865DFBFFBDFF9E85949370FE76F9D1" authorityName="Motschoulsky" authorityYear="1861" box="[200,474,1642,1661]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B1F2FA6AFFBDFF9E85949370FE76F9D1" box="[200,474,1642,1661]" italics="true" pageId="9" pageNumber="56">Micronychus bimaculatus</emphasis>
</taxonomicName>
; H,
<taxonomicName id="44865DFBFFBDFF9E874D9370FD44F9D0" box="[529,744,1642,1660]" class="Insecta" family="Lycidae" genus="Parantis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="triguttatus">
<emphasis id="B1F2FA6AFFBDFF9E874D9370FD44F9D0" box="[529,744,1642,1660]" italics="true" pageId="9" pageNumber="56">Parantis triguttatus</emphasis>
</taxonomicName>
. I
<emphasis id="B1F2FA6AFFBDFF9E865F9376FCA2F9D0" box="[771,782,1644,1660]" italics="true" pageId="9" pageNumber="56">–</emphasis>
O, male genitalia, ventrally: I,
<taxonomicName id="44865DFBFFBDFF9E81369370FAD1F9D1" authorityName="Motschoulsky" authorityYear="1861" box="[1130,1405,1642,1661]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B1F2FA6AFFBDFF9E81369370FAD1F9D1" box="[1130,1405,1642,1661]" italics="true" pageId="9" pageNumber="56">Micronychus bimaculatus</emphasis>
</taxonomicName>
; J,
<taxonomicName id="44865DFBFFBDFF9E85F8939DFED4F935" box="[164,376,1671,1689]" class="Insecta" family="Lycidae" genus="Parantis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="triguttatus">
<emphasis id="B1F2FA6AFFBDFF9E85F8939DFED4F935" box="[164,376,1671,1689]" italics="true" pageId="9" pageNumber="56">Parantis triguttatus</emphasis>
</taxonomicName>
; K,
<taxonomicName id="44865DFBFFBDFF9E84FB939DFD1EF935" authorityName="Linnaeus" authorityYear="1758" box="[423,690,1671,1689]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="sanguineus">
<emphasis id="B1F2FA6AFFBDFF9E84FB939DFD1EF935" box="[423,690,1671,1689]" italics="true" pageId="9" pageNumber="56">Lygistopterus sanguineus</emphasis>
</taxonomicName>
; L,
<taxonomicName id="44865DFBFFBDFF9E8783939DFCC4F936" authorityName="Guerin-Meneville" authorityYear="1833" box="[735,872,1671,1690]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBDFF9E8783939DFCC4F936" box="[735,872,1671,1690]" italics="true" pageId="9" pageNumber="56">Calochromus</emphasis>
</taxonomicName>
(
<emphasis id="B1F2FA6AFFBDFF9E8625939DFC3FF935" box="[889,915,1671,1689]" italics="true" pageId="9" pageNumber="56">s.s</emphasis>
.),
<taxonomicName id="44865DFBFFBDFF9E86EC939DFB99F936" authorityName="Guerin-Meneville" authorityYear="1833" box="[944,1077,1671,1690]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="glaucopterus">
<emphasis id="B1F2FA6AFFBDFF9E86EC939DFB99F936" box="[944,1077,1671,1690]" italics="true" pageId="9" pageNumber="56">glaucopterus</emphasis>
</taxonomicName>
; M,
<taxonomicName id="44865DFBFFBDFF9E8134939DFF4AF91B" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="kelantanensis" subGenus="Flabellochromus">
<emphasis id="B1F2FA6AFFBDFF9E8134939DFBD3F936" box="[1128,1151,1671,1690]" italics="true" pageId="9" pageNumber="56">C.</emphasis>
(
<emphasis id="B1F2FA6AFFBDFF9E81D3939DFA91F936" box="[1167,1341,1671,1690]" italics="true" pageId="9" pageNumber="56">Flabellochromus</emphasis>
)
<emphasis id="B1F2FA6AFFBDFF9E8010939DFF4AF91B" italics="true" pageId="9" pageNumber="56">kelantanensis</emphasis>
</taxonomicName>
; N,
<taxonomicName id="44865DFBFFBDFF9E844993BEFDF1F91B" authorityName="Motyka &amp; Masek &amp; Bocak" authorityYear="2017" baseAuthorityName="Leconte" baseAuthorityYear="1875" box="[277,605,1700,1719]" class="Insecta" family="Lycidae" genus="Macrolygistopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="rubripennis">
<emphasis id="B1F2FA6AFFBDFF9E844993BEFDF1F91B" box="[277,605,1700,1719]" italics="true" pageId="9" pageNumber="56">Macrolygistopterus rubripennis</emphasis>
</taxonomicName>
; O,
<taxonomicName id="44865DFBFFBDFF9E87D093BFFC85F91B" authorityName="Duges" authorityYear="1879" box="[652,809,1700,1719]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="schini">
<emphasis id="B1F2FA6AFFBDFF9E87D093BFFC85F91B" box="[652,809,1700,1719]" italics="true" pageId="9" pageNumber="56">Lucaina schini</emphasis>
</taxonomicName>
. P,
<taxonomicName id="44865DFBFFBDFF9E860893BFFBF3F91B" authorityName="Linnaeus" authorityYear="1758" box="[852,1119,1701,1719]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="sanguineus">
<emphasis id="B1F2FA6AFFBDFF9E860893BFFBF3F91B" box="[852,1119,1701,1719]" italics="true" pageId="9" pageNumber="56">Lygistopterus sanguineus</emphasis>
</taxonomicName>
, terminal abdominal sclerites, ventrally. Q,
<taxonomicName id="44865DFBFFBDFF9E846D93D8FDA8F978" box="[305,516,1730,1748]" class="Insecta" family="Lycidae" genus="Parantis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="triguttatus">
<emphasis id="B1F2FA6AFFBDFF9E846D93D8FDA8F978" box="[305,516,1730,1748]" italics="true" pageId="9" pageNumber="56">Parantis triguttatus</emphasis>
</taxonomicName>
, antenna, detail. Scale bars: A
<emphasis id="B1F2FA6AFFBDFF9E861693DEFCF9F978" box="[842,853,1732,1748]" italics="true" pageId="9" pageNumber="56">–</emphasis>
H = 2.5 mm; I
<emphasis id="B1F2FA6AFFBDFF9E86B093DEFC5BF978" box="[1004,1015,1732,1748]" italics="true" pageId="9" pageNumber="56">–</emphasis>
Q = 0.25 mm.
</paragraph>
</caption>
<paragraph id="83392678FFBDFF9D85F893E1FD38FEB5" blockId="9.[164,779,1787,1901]" lastBlockId="10.[147,762,198,282]" lastPageId="10" lastPageNumber="57" pageId="9" pageNumber="56">
<emphasis id="B1F2FA6AFFBDFF9E85F893E1FDBAF8BD" box="[164,534,1787,1809]" italics="true" pageId="9" pageNumber="56">
Differential diagnosis:
<taxonomicName id="44865DFBFFBDFF9E84E493E6FDBAF8BD" authorityName="Duges" authorityYear="1879" box="[440,534,1788,1809]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">Lucaina</taxonomicName>
</emphasis>
and closely related
<taxonomicName id="44865DFBFFBDFF9E85F89200FED0F883" authorityName="Pic" authorityYear="1929" box="[164,380,1818,1839]" class="Insecta" family="Lycidae" genus="Macrolygistopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBDFF9E85F89200FED0F883" box="[164,380,1818,1839]" italics="true" pageId="9" pageNumber="56">Macrolygistopterus</emphasis>
</taxonomicName>
(
<figureCitation id="1BBD3AFDFFBDFF9E84D39200FE48F883" box="[399,484,1818,1840]" captionStart="Figure 1" captionStartId="3.[164,243,1304,1323]" captionTargetBox="[172,1438,198,1271]" captionTargetId="figure-200@3.[172,1438,198,1271]" captionTargetPageId="3" captionText="Figure 1. A, phylogenetic hypothesis for Calochromini based on the maximum-likelihood analysis of the MUSCLE alignment. Numbers at the branches are maximum-likelihood bootstrap values and Bayesian posterior probabilities. The complete tree with outgroups and full-branch lengths is shown in Fig. S8 (Supporting Information). B, discrete phylogeography reconstruction using a Bayesian approach as implemented in BEAST 2.1; pie charts represent posterior probabilities of biogeographical origins. C, geographical distribution chart of the Calochromini genera." figureDoi="http://doi.org/10.5281/zenodo.14807186" httpUri="https://zenodo.org/record/14807186/files/figure.png" pageId="9" pageNumber="56">Fig. 1A</figureCitation>
) share the rostrate head and they differ in the shape of the apical part of parameres (
<figureCitation id="1BBD3AFDFFBDFF9E846F924DFE06F8C0" box="[307,426,1879,1901]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="9" pageNumber="56">
Fig. 5N
<emphasis id="B1F2FA6AFFBDFF9E84D69240FE3AF8C0" box="[394,406,1882,1900]" italics="true" pageId="9" pageNumber="56">–</emphasis>
O
</figureCitation>
) and the long erect setae in the elytra of
<emphasis id="B1F2FA6AFFBDFF9E868893E6FB95F8BD" box="[980,1081,1788,1809]" italics="true" pageId="9" pageNumber="56">
<taxonomicName id="44865DFBFFBDFF9E868893E6FB99F8BD" authorityName="Duges" authorityYear="1879" box="[980,1077,1788,1809]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">Lucaina</taxonomicName>
.
</emphasis>
These genera have a vicariant distribution (Figs S8, S9, Supporting Information).
<taxonomicName id="44865DFBFFBDFF9E86679222FC4EF8E1" authorityName="Green" authorityYear="1954" box="[827,994,1848,1869]" class="Insecta" family="Lycidae" genus="Caloptognatha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBDFF9E86679222FC4EF8E1" box="[827,994,1848,1869]" italics="true" pageId="9" pageNumber="56">Caloptognatha</emphasis>
</taxonomicName>
has similar setae in the elytra, but does not have the prolonged rostrum and has long, slender mandibles (
<bibRefCitation id="E7175B89FFBEFF9D84DB95DDFD8CFF70" author="Green JW" box="[391,544,198,220]" pageId="10" pageNumber="57" pagination="55 - 57" refId="ref11442" refString="Green JW. 1954. An addition to the Lycid fauna of the United States (Coleoptera). The Coleopterists' Bulletin 8: 55-57." type="journal article" year="1954">Green, 1954</bibRefCitation>
).
<taxonomicName id="44865DFBFFBEFF9D876095DDFD79FF70" authorityName="Kleine" authorityYear="1933" box="[572,725,199,220]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D876095DDFD79FF70" box="[572,725,199,220]" italics="true" pageId="10" pageNumber="57">Lygistopterus</emphasis>
</taxonomicName>
is another genus distributed in the Nearctic region and differs by its wide, parallel-sided parameres.
</paragraph>
<paragraph id="83392678FFBEFF9D85CF945BFD15FEFB" blockId="10.[147,697,321,343]" box="[147,697,321,343]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D85CF945BFE89FEFA" box="[147,293,321,342]" italics="true" pageId="10" pageNumber="57">Distribution:</emphasis>
<collectingCountry id="FB9166E8FFBEFF9D8468945BFEC1FEFA" box="[308,365,321,342]" name="United States of America" pageId="10" pageNumber="57">USA</collectingCountry>
:
<collectingRegion id="4142E89AFFBEFF9D8424945BFE41FEFB" box="[376,493,321,343]" country="United States of America" name="California" pageId="10" pageNumber="57">California</collectingRegion>
,
<collectingRegion id="4142E89AFFBEFF9D84A6945BFDFAFEFB" box="[506,598,321,343]" country="United States of America" name="Arizona" pageId="10" pageNumber="57">Arizona</collectingRegion>
;
<collectingCountry id="FB9166E8FFBEFF9D873E945BFD1AFEFB" box="[610,694,321,343]" name="Mexico" pageId="10" pageNumber="57">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

169
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<heading id="D8719114FFBEFF9D861C95CCFAD0FF41" box="[832,1404,210,237]" centered="true" fontSize="9" level="2" pageId="10" pageNumber="57" reason="2">
<taxonomicName id="44865DFBFFBEFF9D861C95CCFB54FF47" authority="MOTYKA, MASEK &amp; BOCAK" authorityName="MOTYKA, MASEK &amp; BOCAK" authorityYear="2017" box="[832,1272,210,237]" class="Insecta" family="Lycidae" genus="Parantis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<smallCapsWord id="85DFB0A4FFBEFF9D861C95CCFC1AFF46" baselines="230,230" box="[832,950,214,235]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Parantis" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D861C95CCFC1AFF46" bold="true" box="[832,950,214,235]" italics="true" pageId="10" pageNumber="57">PARANTIS</emphasis>
</smallCapsWord>
<smallCapsWord id="85DFB0A4FFBEFF9D86E395CEFBB3FF47" baselines="230,231" box="[959,1055,212,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Motyka" pageId="10" pageNumber="57">MOTYKA</smallCapsWord>
,
<smallCapsWord id="85DFB0A4FFBEFF9D816D95CEFB2DFF47" baselines="230,231" box="[1073,1153,212,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Masek" pageId="10" pageNumber="57">MASEK</smallCapsWord>
&amp;
<smallCapsWord id="85DFB0A4FFBEFF9D81F795CEFB54FF47" baselines="230,231" box="[1195,1272,212,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Bocak" pageId="10" pageNumber="57">BOCAK</smallCapsWord>
</taxonomicName>
<emphasis id="B1F2FA6AFFBEFF9D805E95C3FAD0FF41" bold="true" box="[1282,1404,210,237]" pageId="10" pageNumber="57">
<taxonomicNameLabel id="AAC14711FFBEFF9D805E95C3FAD0FF41" box="[1282,1404,210,237]" pageId="10" pageNumber="57" rank="genus">
<smallCapsWord id="85DFB0A4FFBEFF9D805E95C3FA9EFF46" baselines="230" box="[1282,1330,217,234]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="gen" pageId="10" pageNumber="57">GEN</smallCapsWord>
.
<smallCapsWord id="85DFB0A4FFBEFF9D801F95C3FADFFF46" baselines="230" box="[1347,1395,217,234]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="10" pageNumber="57">NOV</smallCapsWord>
.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CB9C75F3FFBEFF9D86769439FBA4F8C1" pageId="10" pageNumber="57" type="materials_examined">
<paragraph id="83392678FFBEFF9D86769439FBE8FEFB" blockId="10.[810,1425,290,373]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D86769439FC6CFE94" box="[810,960,291,312]" italics="true" pageId="10" pageNumber="57">
<typeStatus id="5C3D98DAFFBEFF9D86769439FCCCFE94" box="[810,864,291,312]" pageId="10" pageNumber="57">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="44865DFBFFBEFF9D86939438FCFFFEFB" authority="Kleine, 1929: 260" authorityName="Kleine" authorityPageNumber="260" authorityYear="1929" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="triguttatus">
<emphasis id="B1F2FA6AFFBEFF9D86939438FB46FE94" box="[975,1258,290,312]" italics="true" pageId="10" pageNumber="57">Calochromus triguttatus</emphasis>
Kleine, 1929: 260
</taxonomicName>
(hereby designated).
</paragraph>
<paragraph id="83392678FFBEFF9D861F9445FC7BFED9" blockId="10.[810,1425,290,373]" box="[835,983,351,373]" pageId="10" pageNumber="57">
<figureCitation id="1BBD3AFDFFBEFF9D861F9445FC7BFED9" box="[835,983,351,373]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="10" pageNumber="57">Figure 5H, J</figureCitation>
</paragraph>
<paragraph id="83392678FFBEFF9D86769487FABDFD81" blockId="10.[810,1425,413,557]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D86769487FB02FE1E" box="[810,1198,413,434]" italics="true" pageId="10" pageNumber="57">
Differential diagnosis:
<taxonomicName id="44865DFBFFBEFF9D81109487FB02FE1E" authorityName="Motyka &amp; Masek &amp; Bocak" authorityYear="2017" box="[1100,1198,413,434]" class="Insecta" family="Lycidae" genus="Parantis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">Parantis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="AAC14711FFBEFF9D81909487FAE5FE1E" box="[1228,1353,413,434]" pageId="10" pageNumber="57" rank="genus">gen. nov.</taxonomicNameLabel>
and
<taxonomicName id="44865DFBFFBEFF9D867694A1FC17FE7C" authorityName="Motschoulsky" authorityYear="1861" box="[810,955,443,464]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D867694A1FC17FE7C" box="[810,955,443,464]" italics="true" pageId="10" pageNumber="57">Micronychus</emphasis>
</taxonomicName>
share a characteristic sclerotized internal sac (
<figureCitation id="1BBD3AFDFFBEFF9D868C94C0FBEDFE43" box="[976,1089,473,496]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="10" pageNumber="57">Fig. 5I, J</figureCitation>
).
<taxonomicName id="44865DFBFFBEFF9D810394C0FB6DFE43" authorityName="Motyka &amp; Masek &amp; Bocak" authorityYear="2017" box="[1119,1217,474,495]" class="Insecta" family="Lycidae" genus="Parantis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D810394C0FB6DFE43" box="[1119,1217,474,495]" italics="true" pageId="10" pageNumber="57">Parantis</emphasis>
</taxonomicName>
differs from the latter in the short antennomere 3 (
<figureCitation id="1BBD3AFDFFBEFF9D819394E2FA84FDA2" box="[1231,1320,504,526]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="10" pageNumber="57">Fig. 5Q</figureCitation>
), minute mouthparts and absent rostrum (
<figureCitation id="1BBD3AFDFFBEFF9D81F6970DFAA8FD81" box="[1194,1284,535,557]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="10" pageNumber="57">Fig. 5H</figureCitation>
).
</paragraph>
<paragraph id="83392678FFBEFF9D8676974EFB48F990" blockId="10.[810,1425,596,1657]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D8676974EFC56FDC5" box="[810,1018,596,617]" italics="true" pageId="10" pageNumber="57">Description: Male</emphasis>
: Body medium-sized,
<quantity id="447E8B9DFFBEFF9D807B974EFA3CFDC5" box="[1319,1424,596,617]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.8" metricValueMin="0.9" pageId="10" pageNumber="57" unit="mm" value="13.5" valueMax="18.0" valueMin="9.0">
9
<emphasis id="B1F2FA6AFFBEFF9D8069974DFAEDFDC5" box="[1333,1345,599,617]" italics="true" pageId="10" pageNumber="57">–</emphasis>
18 mm
</quantity>
long, parallel-sided, dorso-ventrally flattened. Head small, partly hidden by pronotum, non-rostrate, movable, prognathous to hypognathous. Eyes small, hemispherically prominent, frontal interocular distance three times as long as maximum eye diameter. Labrum sclerotized, transverse, with long setae at anterior margin; mandibles minute, curved in apical part, incisor margin simple; maxillary palpi four-segmented, slender, apical palpomere variable in shape, flat, more or less pointed apically. Antennal tubercles conspicuous, separated by narrow straight depression. Antennae 11-segmented, moderately long, surpassing humeral third of elytra, filiform, scapus pear-like, robust, pedicel small, antennomere 3 short, never more than twice as long as pedicel, antennomeres 4
<emphasis id="B1F2FA6AFFBEFF9D812C915AFBD0FBFE" box="[1136,1148,1088,1106]" italics="true" pageId="10" pageNumber="57">–</emphasis>
10 approximately same length, gradually shorter to apex of antennae, antennomere 11 longest, slender, pointed at apex. Pronotum transverse to quadrate, parallel-sided, with deep median depression, anterior angles prominent, posterior ones slightly pointed to weakly rounded. Scutellum short, nearly quadrate, posterior margin simple, at most with shallow incision. Elytra flat, parallel-sided, humeri rounded, elytra with four straight longitudinal costae, costae are moderately robust at humeri, very weak apically, transverse costae absent. Legs slender, flattened, tarsi five-segmented, tarsomeres 1
<emphasis id="B1F2FA6AFFBEFF9D811390B4FBF7FA6C" box="[1103,1115,1454,1472]" italics="true" pageId="10" pageNumber="57">–</emphasis>
4 with membranous pads, tarsomere 5 long, claws simple, slender. Male genitalia trilobate, phallus and parameres similar length, parameres slender apically, phallobase asymmetrical, internal sac sclerotized.
</paragraph>
<paragraph id="83392678FFBEFF9D861F935EFB31F9D4" blockId="10.[810,1425,596,1657]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D861F935EFC39F9F5" box="[835,917,1604,1625]" italics="true" pageId="10" pageNumber="57">Female</emphasis>
: Similar to male, without sexual polymorphism in the shape of antennae.
</paragraph>
<paragraph id="83392678FFBEFF9D867693BAFBE6F978" blockId="10.[810,1425,1696,1748]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D867693BAFC10F919" box="[810,956,1696,1717]" italics="true" pageId="10" pageNumber="57">Distribution:</emphasis>
<collectingCountry id="FB9166E8FFBEFF9D869793BAFBA6F91A" box="[971,1034,1696,1718]" name="India" pageId="10" pageNumber="57">India</collectingCountry>
,
<collectingCountry id="FB9166E8FFBEFF9D814B93BAFBCCF91A" box="[1047,1120,1696,1718]" name="China" pageId="10" pageNumber="57">China</collectingCountry>
:
<collectingRegion id="4142E89AFFBEFF9D813793BAFB65F919" box="[1131,1225,1696,1717]" country="China" name="Yunnan" pageId="10" pageNumber="57">Yunnan</collectingRegion>
,
<collectingRegion id="4142E89AFFBEFF9D818993BAFAB6F91A" box="[1237,1306,1696,1718]" country="China" name="Hubei" pageId="10" pageNumber="57">Hubei</collectingRegion>
(Fig. S11, Supporting Information).
</paragraph>
<paragraph id="83392678FFBEFF9D867693E6FBA4F8C1" blockId="10.[810,1425,1788,1901]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D867693E6FC04F8BD" box="[810,936,1788,1809]" italics="true" pageId="10" pageNumber="57">Etymology:</emphasis>
The name
<taxonomicName id="44865DFBFFBEFF9D811D93E6FB0FF8BD" authorityName="Motyka &amp; Masek &amp; Bocak" authorityYear="2017" box="[1089,1187,1788,1809]" class="Insecta" family="Lycidae" genus="Parantis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D811D93E6FB0FF8BD" box="[1089,1187,1788,1809]" italics="true" pageId="10" pageNumber="57">Parantis</emphasis>
</taxonomicName>
is a matronym in honour of Justinah
<taxonomicName id="44865DFBFFBEFF9D817F9200FB2BF89C" authorityName="Motyka &amp; Masek &amp; Bocak" authorityYear="2017" box="[1059,1159,1818,1840]" class="Insecta" family="Lycidae" genus="Parantis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="genus" status="gen. nov.">Parantis</taxonomicName>
, a biologist from the
<collectingRegion id="4142E89AFFBEFF9D86769223FCDDF8E3" box="[810,881,1849,1871]" country="Malaysia" name="Sabah" pageId="10" pageNumber="57">Sabah</collectingRegion>
National Park Authority, Poring,
<collectingCountry id="FB9166E8FFBEFF9D807E9223FA21F8E3" box="[1314,1421,1849,1871]" name="Malaysia" pageId="10" pageNumber="57">Malaysia</collectingCountry>
. Gender: masculine.
</paragraph>
</subSubSection>
<subSubSection id="CB9C75F3FFBFFF9C859895DDFD47FF72" box="[196,747,198,222]" pageId="11" pageNumber="58" type="key">
<paragraph id="83392678FFBFFF9C859895DDFD47FF72" blockId="11.[196,747,198,222]" box="[196,747,198,222]" pageId="11" pageNumber="58">
KEY TO THE
<taxonomicName id="44865DFBFFBFFF9C842795DCFDF2FF72" box="[379,606,198,222]" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="tribe" tribe="Calochromini">CALOCHROMINI</taxonomicName>
GENERA*
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="83392678FFBEFF9D85CF9182FDACFB02" blockId="10.[147,512,1176,1198]" box="[147,512,1176,1198]" pageId="10" pageNumber="57">
<heading id="D8719114FFBEFF9D85CF9182FDACFB02" box="[147,512,1176,1198]" centered="true" fontSize="9" level="2" pageId="10" pageNumber="57" reason="2">
<treatmentCitationGroup id="A3960156FFBEFF9D85CF9182FDACFB02" box="[147,512,1176,1198]" pageId="10" pageNumber="57">
<treatmentCitation id="02270069FFBEFF9D85CF9182FE50FB02" author="Green JW" box="[147,508,1176,1198]" page="55" pageId="10" pageNumber="57" year="1954">
<taxonomicName id="44865DFBFFBEFF9D85CF9182FE50FB02" ID-CoL="3FX3" authority="Green, 1954: 55" authorityName="Green" authorityPageNumber="55" authorityYear="1954" box="[147,508,1176,1198]" class="Insecta" family="Lycidae" genus="Caloptognatha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D85CF9182FE96FB01" box="[147,314,1176,1197]" italics="true" pageId="10" pageNumber="57">Caloptognatha</emphasis>
<bibRefCitation id="E7175B89FFBEFF9D841F9182FE50FB02" author="Green JW" box="[323,508,1176,1198]" pageId="10" pageNumber="57" pagination="55 - 57" refId="ref11442" refString="Green JW. 1954. An addition to the Lycid fauna of the United States (Coleoptera). The Coleopterists' Bulletin 8: 55-57." type="journal article" year="1954">Green, 1954: 55</bibRefCitation>
</taxonomicName>
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.
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="CB9C75F3FFBEFF9D85CF91CCFDB6F8C1" pageId="10" pageNumber="57" type="materials_examined">
<paragraph id="83392678FFBEFF9D85CF91CCFE1FFAA6" blockId="10.[147,762,1237,1290]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D85CF91CCFE82FB47" box="[147,302,1238,1259]" italics="true" pageId="10" pageNumber="57">
<typeStatus id="5C3D98DAFFBEFF9D85CF91CCFF65FB47" box="[147,201,1238,1259]" pageId="10" pageNumber="57">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="44865DFBFFBEFF9D846191CFFF02FAA6" authority="Green, 1954: 56" authorityName="Green" authorityPageNumber="56" authorityYear="1954" class="Insecta" family="Lycidae" genus="Caloptognatha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="beameri">
<emphasis id="B1F2FA6AFFBEFF9D846191CFFDFCFB46" box="[317,592,1237,1258]" italics="true" pageId="10" pageNumber="57">Caloptognatha beameri</emphasis>
<bibRefCitation id="E7175B89FFBEFF9D873C91CCFF02FAA6" author="Green JW" pageId="10" pageNumber="57" pagination="55 - 57" refId="ref11442" refString="Green JW. 1954. An addition to the Lycid fauna of the United States (Coleoptera). The Coleopterists' Bulletin 8: 55-57." type="journal article" year="1954">Green, 1954: 56</bibRefCitation>
</taxonomicName>
(original designation).
</paragraph>
<paragraph id="83392678FFBEFF9D85CF902BFEB8FA52" blockId="10.[147,762,1329,1534]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D85CF902BFDFFFAEA" box="[147,595,1329,1350]" italics="true" pageId="10" pageNumber="57">
Differential diagnosis:
<taxonomicName id="44865DFBFFBEFF9D84F0902BFDFFFAEA" authorityName="Green" authorityYear="1954" box="[428,595,1329,1350]" class="Insecta" family="Lycidae" genus="Caloptognatha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">Caloptognatha</taxonomicName>
</emphasis>
differs from other genera in having densely pubescent elytral costae. Additionally,
<taxonomicName id="44865DFBFFBEFF9D84FE9075FD97FA28" authorityName="Kleine" authorityYear="1933" box="[418,571,1391,1412]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D84FE9075FD97FA28" box="[418,571,1391,1412]" italics="true" pageId="10" pageNumber="57">Lygistopterus</emphasis>
</taxonomicName>
,
<taxonomicName id="44865DFBFFBEFF9D87069075FD14FA28" authorityName="Duges" authorityYear="1879" box="[602,696,1391,1412]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D87069075FD14FA28" box="[602,696,1391,1412]" italics="true" pageId="10" pageNumber="57">Lucaina</emphasis>
</taxonomicName>
and
<taxonomicName id="44865DFBFFBEFF9D85CF9097FEC7FA0E" authorityName="Pic" authorityYear="1929" box="[147,363,1421,1442]" class="Insecta" family="Lycidae" genus="Macrolygistopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D85CF9097FEC7FA0E" box="[147,363,1421,1442]" italics="true" pageId="10" pageNumber="57">Macrolygistopterus</emphasis>
</taxonomicName>
occur in the New World and the latter two have a long rostrum while
<taxonomicName id="44865DFBFFBEFF9D871990B6FD72FA6D" authorityName="Kleine" authorityYear="1933" box="[581,734,1452,1473]" class="Insecta" family="Lycidae" genus="Lygistopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D871990B6FD72FA6D" box="[581,734,1452,1473]" italics="true" pageId="10" pageNumber="57">Lygistopterus</emphasis>
</taxonomicName>
is easily distinguishable by wide, parallel-sided parameres.
</paragraph>
<paragraph id="83392678FFBEFF9D85CF933FFE75F997" blockId="10.[147,473,1573,1595]" box="[147,473,1573,1595]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D85CF933FFE89F996" box="[147,293,1573,1594]" italics="true" pageId="10" pageNumber="57">Distribution:</emphasis>
<collectingCountry id="FB9166E8FFBEFF9D8468933CFEC1F997" box="[308,365,1574,1595]" name="United States of America" pageId="10" pageNumber="57">USA</collectingCountry>
:
<collectingRegion id="4142E89AFFBEFF9D8424933FFE79F997" box="[376,469,1573,1595]" country="United States of America" name="Arizona" pageId="10" pageNumber="57">Arizona</collectingRegion>
.
</paragraph>
<paragraph id="83392678FFBEFF9D85CF9379FDB6F8C1" blockId="10.[147,762,1634,1901]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D85CF9379FF53F9D4" box="[147,255,1635,1656]" italics="true" pageId="10" pageNumber="57">Remarks:</emphasis>
<bibRefCitation id="E7175B89FFBEFF9D84529379FE09F9D4" author="Green JW" box="[270,421,1634,1656]" pageId="10" pageNumber="57" pagination="55 - 57" refId="ref11442" refString="Green JW. 1954. An addition to the Lycid fauna of the United States (Coleoptera). The Coleopterists' Bulletin 8: 55-57." type="journal article" year="1954">Green (1954)</bibRefCitation>
noted that
<taxonomicName id="44865DFBFFBEFF9D87689379FD77F9D4" authorityName="Green" authorityYear="1954" box="[564,731,1635,1656]" class="Insecta" family="Lycidae" genus="Caloptognatha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D87689379FD77F9D4" box="[564,731,1635,1656]" italics="true" pageId="10" pageNumber="57">Caloptognatha</emphasis>
</taxonomicName>
is unique among the Nearctic genera due to sparse and unequally distributed elytral pubescence, but this character is shared with
<taxonomicName id="44865DFBFFBEFF9D848693A5FD94F978" authorityName="Duges" authorityYear="1879" box="[474,568,1727,1748]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D848693A5FD94F978" box="[474,568,1727,1748]" italics="true" pageId="10" pageNumber="57">Lucaina</emphasis>
</taxonomicName>
. Unfortunately,
<taxonomicName id="44865DFBFFBEFF9D85CF93C7FE96F95E" authorityName="Green" authorityYear="1954" box="[147,314,1757,1778]" class="Insecta" family="Lycidae" genus="Caloptognatha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D85CF93C7FE96F95E" box="[147,314,1757,1778]" italics="true" pageId="10" pageNumber="57">Caloptognatha</emphasis>
</taxonomicName>
, despite its occurrence in western
<collectingCountry id="FB9166E8FFBEFF9D85CF93E1FF67F8BC" box="[147,203,1787,1808]" name="United States of America" pageId="10" pageNumber="57">USA</collectingCountry>
, has not been collected again. As the
<typeStatus id="5C3D98DAFFBEFF9D87CE93E6FD6FF8BD" box="[658,707,1788,1809]" pageId="10" pageNumber="57">type</typeStatus>
was inaccessible, we provisionally classify
<taxonomicName id="44865DFBFFBEFF9D870E9200FD55F883" authorityName="Green" authorityYear="1954" box="[594,761,1818,1839]" class="Insecta" family="Lycidae" genus="Caloptognatha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D870E9200FD55F883" box="[594,761,1818,1839]" italics="true" pageId="10" pageNumber="57">Caloptognatha</emphasis>
</taxonomicName>
as related to
<taxonomicName id="44865DFBFFBEFF9D846F9223FE3DF8E2" authorityName="Duges" authorityYear="1879" box="[307,401,1849,1870]" class="Insecta" family="Lycidae" genus="Lucaina" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D846F9223FE3DF8E2" box="[307,401,1849,1870]" italics="true" pageId="10" pageNumber="57">Lucaina</emphasis>
</taxonomicName>
and its definitive position can be resolved with further material.
</paragraph>
</subSubSection>
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139
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<heading id="D8719114FFBEFF9D85CF94A1FDE2FE7C" box="[147,590,443,465]" centered="true" fontSize="9" level="2" pageId="10" pageNumber="57" reason="2">
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<taxonomicName id="44865DFBFFBEFF9D85CF94A1FDE6FE7C" authority="Motschoulsky, 1861: 137" authorityName="Motschoulsky" authorityPageNumber="137" authorityYear="1861" box="[147,586,443,465]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D85CF94A1FE88FE7C" box="[147,292,443,464]" italics="true" pageId="10" pageNumber="57">Micronychus</emphasis>
<bibRefCitation id="E7175B89FFBEFF9D847194A1FDE6FE7C" author="Motschoulsky V" box="[301,586,443,465]" pageId="10" pageNumber="57" pagination="95 - 155" refId="ref12258" refString="Motschoulsky V. 1861. D'un catalogue des insecten de l'ile Ceylon. Bulletin of the Moscow Society of Naturalists 34: 95-155." type="book chapter" year="1861">Motschoulsky, 1861: 137</bibRefCitation>
</taxonomicName>
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.
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<paragraph id="83392678FFBEFF9D85CF94E3FD84FD80" blockId="10.[147,762,504,587]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D85CF94E3FED9FDA2" box="[147,373,505,526]" italics="true" pageId="10" pageNumber="57">
<typeStatus id="5C3D98DAFFBEFF9D85CF94E3FF65FDA2" box="[147,201,505,526]" pageId="10" pageNumber="57">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="44865DFBFFBEFF9D84D894E2FED9FD81" authority="Motschoulsky, 1861" authorityName="Motschoulsky" authorityYear="1861" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B1F2FA6AFFBEFF9D84D894E2FD56FDA1" box="[388,762,504,525]" italics="true" pageId="10" pageNumber="57">Micronychus bimaculatus</emphasis>
<bibRefCitation id="E7175B89FFBEFF9D85CF970DFED9FD81" author="Motschoulsky V" box="[147,373,535,557]" pageId="10" pageNumber="57" pagination="95 - 155" refId="ref12258" refString="Motschoulsky V. 1861. D'un catalogue des insecten de l'ile Ceylon. Bulletin of the Moscow Society of Naturalists 34: 95-155." type="book chapter" year="1861">Motschoulsky, 1861</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
<paragraph id="83392678FFBEFF9D85F0972FFEC8FDE6" blockId="10.[147,762,504,587]" box="[172,356,565,587]" pageId="10" pageNumber="57">
<figureCitation id="1BBD3AFDFFBEFF9D85F0972FFECCFDE6" box="[172,352,565,587]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="10" pageNumber="57">Figure 5G, I, Q</figureCitation>
.
</paragraph>
<paragraph id="83392678FFBEFF9D85CF9768FECDFC8D" blockId="10.[147,762,626,801]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D85CF9768FDECFD2B" box="[147,576,626,647]" italics="true" pageId="10" pageNumber="57">
Differential diagnosis:
<taxonomicName id="44865DFBFFBEFF9D84F39768FDECFD2B" authorityName="Motschoulsky" authorityYear="1861" box="[431,576,626,647]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">Micronychus</taxonomicName>
</emphasis>
and
<taxonomicName id="44865DFBFFBEFF9D87CB9769FD55FD24" authorityName="Motyka &amp; Masek &amp; Bocak" authorityYear="2017" box="[663,761,627,648]" class="Insecta" family="Lycidae" genus="Parantis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B1F2FA6AFFBEFF9D87CB9769FD55FD24" box="[663,761,627,648]" italics="true" pageId="10" pageNumber="57">Parantis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAC14711FFBEFF9D85CF978BFEA8FD0A" box="[147,260,657,678]" pageId="10" pageNumber="57" rank="genus">gen. nov.</taxonomicNameLabel>
are characterized by the presence of a sclerotized internal sac (
<figureCitation id="1BBD3AFDFFBEFF9D848C97B5FDEAFD69" box="[464,582,687,709]" captionStart="Figure 5" captionStartId="9.[164,243,1583,1602]" captionTargetBox="[175,1438,199,1550]" captionTargetId="figure-94@9.[172,1438,198,1550]" captionTargetPageId="9" captionText="Figure 5. A–H, general appearance: A, Lygistopterus sanguineus; B, Calochromus (s.s., glaucopterus; C, C. (Flabellochromus, kelantanensis; D, C. (Paracalochromus) giganteus; E, Lucaina discoidalis; F, Macrolygistopterus rubripennis; G, Micronychus bimaculatus; H, Parantis triguttatus. I–O, male genitalia, ventrally: I, Micronychus bimaculatus; J, Parantis triguttatus; K, Lygistopterus sanguineus; L, Calochromus (s.s.), glaucopterus; M, C. (Flabellochromus) kelantanensis; N, Macrolygistopterus rubripennis; O, Lucaina schini. P, Lygistopterus sanguineus, terminal abdominal sclerites, ventrally. Q, Parantis triguttatus, antenna, detail. Scale bars: A–H = 2.5 mm; I–Q = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14807196" httpUri="https://zenodo.org/record/14807196/files/figure.png" pageId="10" pageNumber="57">Fig. 5I, J</figureCitation>
).
<taxonomicName id="44865DFBFFBEFF9D873497B5FD55FD68" authorityName="Motschoulsky" authorityYear="1861" box="[616,761,687,708]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D873497B5FD55FD68" box="[616,761,687,708]" italics="true" pageId="10" pageNumber="57">Micronychus</emphasis>
</taxonomicName>
differs by antennomere 3 being twice as long as the pedicel, well-developed mouthparts and sometimes in the rostrate head.
</paragraph>
<paragraph id="83392678FFBEFF9D85CF9652FD02FCF2" blockId="10.[147,686,840,862]" box="[147,686,840,862]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D85CF9652FE89FCF1" box="[147,293,840,861]" italics="true" pageId="10" pageNumber="57">Distribution:</emphasis>
Oriental and Afrotropical region.
</paragraph>
<paragraph id="83392678FFBEFF9D85CF969FFD58FB9F" blockId="10.[147,762,901,1076]" pageId="10" pageNumber="57">
<emphasis id="B1F2FA6AFFBEFF9D85CF969FFF53FC36" box="[147,255,901,922]" italics="true" pageId="10" pageNumber="57">Remarks:</emphasis>
<bibRefCitation id="E7175B89FFBEFF9D8452969FFE1FFC36" author="Kleine R" box="[270,435,901,923]" pageId="10" pageNumber="57" refId="ref11706" refString="Kleine R. 1933. Coleopterorum Catalogus auspiciis et auxilio W. Junk editus S. Schenkling. Pars 128: Lycidae. Berlin: W. Junk." type="journal volume" year="1933">Kleine (1933)</bibRefCitation>
listed
<taxonomicName id="44865DFBFFBEFF9D8746969FFD07FC36" authorityName="Motschoulsky" authorityYear="1861" box="[538,683,901,922]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D8746969FFD07FC36" box="[538,683,901,922]" italics="true" pageId="10" pageNumber="57">Micronychus</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="44865DFBFFBEFF9D842B96BEFDA0FC15" authorityName="Guerin-Meneville" authorityYear="1833" box="[375,524,932,953]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D842B96BEFDA0FC15" box="[375,524,932,953]" italics="true" pageId="10" pageNumber="57">Calochromus</emphasis>
</taxonomicName>
, but we found that these genera are distantly related. Here, we resurrect the name
<taxonomicName id="44865DFBFFBEFF9D842096FBFDA1FC5A" authorityName="Motschoulsky" authorityYear="1861" box="[380,525,993,1014]" class="Insecta" family="Lycidae" genus="Micronychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D842096FBFDA1FC5A" box="[380,525,993,1014]" italics="true" pageId="10" pageNumber="57">Micronychus</emphasis>
</taxonomicName>
for Oriental species previously classified in
<taxonomicName id="44865DFBFFBEFF9D848096E5FDDDFBB8" authorityName="Guerin-Meneville" authorityYear="1833" box="[476,625,1023,1044]" class="Insecta" family="Lycidae" genus="Calochromus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D848096E5FDDDFBB8" box="[476,625,1023,1044]" italics="true" pageId="10" pageNumber="57">Calochromus</emphasis>
</taxonomicName>
and the Afrotropical species placed in
<taxonomicName id="44865DFBFFBEFF9D84B79104FD5CFB9F" authority="(Kleine, 1933)" baseAuthorityName="Kleine" baseAuthorityYear="1933" box="[491,752,1054,1076]" class="Insecta" family="Lycidae" genus="Adoceta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B1F2FA6AFFBEFF9D84B79104FDE8FB9F" box="[491,580,1054,1075]" italics="true" pageId="10" pageNumber="57">Adoceta</emphasis>
(
<bibRefCitation id="E7175B89FFBEFF9D87099104FD44FB9F" author="Kleine R" box="[597,744,1054,1076]" pageId="10" pageNumber="57" refId="ref11706" refString="Kleine R. 1933. Coleopterorum Catalogus auspiciis et auxilio W. Junk editus S. Schenkling. Pars 128: Lycidae. Berlin: W. Junk." type="journal volume" year="1933">Kleine, 1933</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>