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<document id="28F3054E8450E1E5677BB816A113EFF0" ID-DOI="10.1051/acarologia/20111995" ID-ISSN="2107-7207" ID-Zenodo-Dep="5404294" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630342961213" checkinUser="marcus" docAuthor="Karg, W. &amp; Schorlemmer, A." docDate="2011" docId="DA69636EFFEAFFE7FC78FDCF88B6866A" docLanguage="en" docName="Acarologia.2011.51.1.3-29.pdf" docOrigin="Acarologia 51 (1)" docSource="http://dx.doi.org/10.1051/acarologia/20111995" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Pyrosejus verticis Karg &amp; Schorlemmer 2011, n. sp." docType="treatment" docUuid="DDCF9E1B-CDC3-4E38-A491-929EBF6137D2" docUuidSource="ZooBank" docVersion="5" lastPageNumber="10" masterDocId="26501B16FFEFFFEFFFA2FFCD885A804D" masterDocTitle="New Insights Into The Systematics Of Parasitiformes (Acarina) With New Species From South America" masterLastPageNumber="29" masterPageNumber="3" pageNumber="7" updateTime="1699236270688" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:title id="FDF7F51B962B8BB6CCB2C0FD07E4B19C">New Insights Into The Systematics Of Parasitiformes (Acarina) With New Species From South America</mods:title>
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<mods:namePart id="1042108C4833697C544A5BEAE95EFD09">Karg, W.</mods:namePart>
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<mods:title id="26153454A822054343CB1DCEB1386448">Acarologia</mods:title>
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<mods:date id="8392B479B886E1F0853EF9E1A32A2ACC">2011</mods:date>
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<mods:number id="B6F99B88C6462B636BF9DD5B6C8CC1DA">2011-03-30</mods:number>
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<paragraph id="527FD278FFEAFFEAFC78FDCF8CAF8257" blockId="5.[986,1269,514,538]" box="[986,1269,514,538]" pageId="5" pageNumber="7">
<heading id="09376514FFEAFFEAFC78FDCF8CAF8257" bold="true" box="[986,1269,514,538]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="7" reason="2">
<emphasis id="60B40E6AFFEAFFEAFC78FDCF8CAF8257" bold="true" box="[986,1269,514,538]" pageId="5" pageNumber="7">
<taxonomicName id="95C0A9FBFFEAFFEAFC78FDCF8CF48257" authority="Karg &amp; Schorlemmer, 2011" authorityName="Karg &amp; Schorlemmer" authorityYear="2011" box="[986,1198,514,538]" class="Arachnida" family="Pyrosejidae" genus="Pyrosejus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="7" phylum="Arthropoda" rank="species" species="verticis" status="sp. nov.">
<emphasis id="60B40E6AFFEAFFEAFC78FDCF8CF48257" bold="true" box="[986,1198,514,538]" italics="true" pageId="5" pageNumber="7">Pyrosejus verticis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7B87B311FFEAFFEAFB16FDCF8CAF8257" box="[1204,1269,514,538]" pageId="5" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="1ADA81F3FFEAFFE7FCC2FDF688B6866A" lastPageId="8" lastPageNumber="10" pageId="5" pageNumber="7" type="materials_examined">
<paragraph id="527FD278FFEAFFEAFCC2FDF68C3782F6" blockId="5.[826,1429,571,1948]" pageId="5" pageNumber="7">
<materialsCitation id="E2A8D825FFEAFFEAFCC2FDF68C3782F6" country="Ecuador" elevation="4100" location="Jumble" municipality="Province" pageId="5" pageNumber="7" specimenCount="1" stateProvince="Pichincha" typeStatus="holotype">
<typeStatus id="8D7B6CDAFFEAFFEAFCC2FDF68B95821E" box="[864,975,571,595]" pageId="5" pageNumber="7" type="holotype">Holotype</typeStatus>
<emphasis id="60B40E6AFFEAFFEAFC59FDF08C48821F" bold="true" box="[1019,1042,573,594]" pageId="5" pageNumber="7">~</emphasis>
<collectingCountry id="2AD792E8FFEAFFEAFBB9FDF68C24821E" box="[1051,1150,571,595]" name="Ecuador" pageId="5" pageNumber="7">Ecuador</collectingCountry>
, 1989,
<collectingMunicipality id="B21B4802FFEAFFEAFB73FDF68D62821E" box="[1233,1336,571,595]" pageId="5" pageNumber="7">Province</collectingMunicipality>
<collectingRegion id="90041C9AFFEAFFEAFAE0FDF68B3F823B" country="Ecuador" name="Pichincha" pageId="5" pageNumber="7">Pichincha</collectingRegion>
,
<locationDeviation id="43CDCB5AFFEAFFEAFCCDFD938CE6823B" box="[879,1212,606,630]" firstLocation="Pifo" pageId="5" pageNumber="7" secondLocation="Papellacta">between Pifo and Papellacta</locationDeviation>
,
<elevation id="D9ED354BFFEAFFEAFB65FD938D0E823B" box="[1223,1364,606,630]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="5" pageNumber="7" unit="m" value="4100.0">
<quantity id="95387F9DFFEAFFEAFB65FD938D438238" box="[1223,1305,606,630]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="5" pageNumber="7" unit="m" value="4100.0">4100 m</quantity>
a.s.l.
</elevation>
from felt-like debris under
<location id="571F84A3FFEAFFEAFB9BFD4C8CD682D4" LSID="urn:lsid:plazi:treatment:DA69636EFFEAFFE7FC78FDCF88B6866A:571F84A3FFEAFFEAFB9BFD4C8CD682D4" box="[1081,1164,641,665]" country="Ecuador" municipality="Province" name="Jumble" pageId="5" pageNumber="7" stateProvince="Pichincha">Jumble</location>
of dicotyledon, creeping to
<quantity id="95387F9DFFEAFFEAFC26FD698BF382F1" box="[900,937,676,700]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="7" unit="m" value="2.0">
<elevation id="D9ED354BFFEAFFEAFC26FD698BF382F1" box="[900,937,676,700]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="7" unit="m" value="2.0">2m</elevation>
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height on a tree.
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</paragraph>
<paragraph id="527FD278FFEAFFEAFCC2FD1C8DCF82A5" blockId="5.[826,1429,571,1948]" box="[864,1429,720,745]" pageId="5" pageNumber="7">
<typeStatus id="8D7B6CDAFFEAFFEAFCC2FD1C8B8F82A4" box="[864,981,721,745]" pageId="5" pageNumber="7" type="paratype">Paratypes</typeStatus>
— 9
<emphasis id="60B40E6AFFEAFFEAFBAEFD1E8C7982A5" bold="true" box="[1036,1059,723,744]" pageId="5" pageNumber="7">~</emphasis>
, collectors I. Loksa and A. Zicsi.
</paragraph>
<paragraph id="527FD278FFEAFFEAFCC2FD308D4A8375" blockId="5.[826,1429,571,1948]" pageId="5" pageNumber="7">Etymology — As to the well-developed vertex with relative long setae i1, z1, z2 and s1.</paragraph>
<paragraph id="527FD278FFEAFFEAFCC2FC818CDD83BD" blockId="5.[826,1429,571,1948]" pageId="5" pageNumber="7">
Diagnosis — Most dorsal setae remarkably long: 60 to 102 µm, on the post-dorsum setae I2 and Z2 absent,
<emphasis id="60B40E6AFFEAFFEAFC31FC588BF083E7" bold="true" box="[915,938,917,938]" pageId="5" pageNumber="7">~</emphasis>
between mesogynial shield and a median incision of the ventrianal shield a single distinct triangular shield (
<figureCitation id="CAFBCEFDFFEAFFEAFC53FC148C0583BC" box="[1009,1119,985,1009]" captionStart="FIGURE" captionStartId="6.[464,476,1883,1902]" captionTargetBox="[264,1336,358,1848]" captionTargetId="figure-5@6.[249,1338,340,1852]" captionTargetPageId="6" captionText="FIGURE 3: Pyrosejus verticis n. sp. Female. Dorsum, 348 374 µm long." figureDoi="http://doi.org/10.5281/zenodo.5404300" httpUri="https://zenodo.org/record/5404300/files/figure.png" pageId="5" pageNumber="7">Figures 3</figureCitation>
,
<figureCitation id="CAFBCEFDFFEAFFEAFBC9FC158C2083BD" box="[1131,1146,984,1008]" captionStart="FIGURE" captionStartId="9.[304,316,1883,1902]" captionTargetBox="[371,1177,357,1837]" captionTargetId="figure-4@9.[370,1218,340,1852]" captionTargetPageId="9" captionText="FIGURE 5: Holostethus longosetis n. sp. Female. a Dorsum, 957 1092 µm long; b Tectum, 88 µm wide." figureDoi="http://doi.org/10.5281/zenodo.5404304" httpUri="https://zenodo.org/record/5404304/files/figure.png" pageId="5" pageNumber="7">5</figureCitation>
).
</paragraph>
<paragraph id="527FD278FFEAFFEAFCC2FBC88B98840D" blockId="5.[826,1429,571,1948]" pageId="5" pageNumber="7">
Description — Idiosoma
<emphasis id="60B40E6AFFEAFFEAFB24FBC58CC78450" bold="true" box="[1158,1181,1032,1053]" pageId="5" pageNumber="7">~</emphasis>
348 374 x 244 270 µm, brown.
</paragraph>
<paragraph id="527FD278FFEAFFEAFCC2FB988CEE8586" blockId="5.[826,1429,571,1948]" pageId="5" pageNumber="7">
<emphasis id="60B40E6AFFEAFFEAFCC2FB988BE38421" box="[864,953,1109,1132]" italics="true" pageId="5" pageNumber="7">Dorsum</emphasis>
: Divided into a pro-dorsum, a postdorsum and backwards setae z3 separate marginal shields, dorsal shields and the posterior part of venter with scale like patterns, posterior margin of the post-dorsum crenellated, most of the dorsal setae remarkably long, longest setae I1, I3 and s4: = 102 µm, i2 = 65, i3 = 60, i4 = i5 = 85, z1 = 48, z4 = 77, s3 = 92, s5 = 78, r3 = 70, I4 = 75, Z1 = Z3 = 95 µm long, setae at the vertex and at the posterior margin shorter however not reduced: i1 = 46, z1 = 65, s1 = 18, z2 = r2 = 20, I5 = 31 µm long.
</paragraph>
<paragraph id="527FD278FFEAFFE7FCC2FA2D8A37822A" blockId="5.[826,1429,571,1948]" lastBlockId="8.[159,762,311,616]" lastPageId="8" lastPageNumber="10" pageId="5" pageNumber="7">
<emphasis id="60B40E6AFFEAFFEAFCC2FA2D8BFD85BA" box="[864,935,1504,1527]" italics="true" pageId="5" pageNumber="7">Venter</emphasis>
: With mesogynial-shield well developed and nearly rectangular, posterior part of the latigynial-plates broadened and with a second seta, the first seta between coxae III and IV, the first sternal setae on separate triangular plates, the second pair on an oval shield, also the third pair on a smaller oval shield, the fourth pair free on the soft cuticle, ventrianal-shield fused with the dorsum and with lateral and peritrematal shields, median of the ventrianal shield with a deep incision which is broadened posterior of the mesogynialshield and accommodates for a single triangular plate, surface of the ventrianal shield similar to the dorsum with scale like patterns, setae of the anterior part of the venter remarkably short, st1 = 34 µm long, the other 18 to 26 µm long, setae of the posterior part mostly longer, Vi2 = Vi3 = 34, Vi4 = 37, Vz1 = Vz2 = Vz3 = 50, Vz4 = 44 µm long, a strip of the marginal shields ventrally visible bearing the following dorsal setae: S3 = S5 = 80, Z4= 71, Z5 = 60 µm long, the anus remarkably small and with a pair of short adanal setae: Vi5 = 25 µm long.
</paragraph>
<caption id="06BF82F0FFE9FFE9FE72F8968C3F8723" ID-DOI="http://doi.org/10.5281/zenodo.5404300" ID-Zenodo-Dep="5404300" box="[464,1125,1882,1902]" httpUri="https://zenodo.org/record/5404300/files/figure.png" pageId="6" pageNumber="8" startId="6.[464,476,1883,1902]" targetBox="[264,1336,358,1848]" targetPageId="6">
<paragraph id="527FD278FFE9FFE9FE72F8968C3F8723" blockId="6.[464,1125,1882,1902]" box="[464,1125,1882,1902]" pageId="6" pageNumber="8">
FIGURE 3:
<taxonomicName id="95C0A9FBFFE9FFE9FD97F8968A978720" authority="Karg &amp; Schorlemmer, 2011" authorityName="Karg &amp; Schorlemmer" authorityYear="2011" box="[565,717,1883,1901]" class="Arachnida" family="Pyrosejidae" genus="Pyrosejus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="8" phylum="Arthropoda" rank="species" species="verticis" status="sp. nov.">
<emphasis id="60B40E6AFFE9FFE9FD97F8968A978720" box="[565,717,1883,1901]" italics="true" pageId="6" pageNumber="8">Pyrosejus verticis</emphasis>
</taxonomicName>
<emphasis id="60B40E6AFFE9FFE9FD70F8968B5C8723" bold="true" box="[722,774,1883,1902]" pageId="6" pageNumber="8">
<taxonomicNameLabel id="7B87B311FFE9FFE9FD70F8968B5C8723" box="[722,774,1883,1902]" pageId="6" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Female. Dorsum, 348 374 µm long.
</paragraph>
</caption>
<caption id="06BF82F0FFE8FFE8FE93F8968D5E8723" ID-DOI="http://doi.org/10.5281/zenodo.5404302" ID-Zenodo-Dep="5404302" box="[305,1284,1882,1902]" httpUri="https://zenodo.org/record/5404302/files/figure.png" pageId="7" pageNumber="9" startId="7.[305,317,1883,1902]" targetBox="[255,1346,350,1830]" targetPageId="7">
<paragraph id="527FD278FFE8FFE8FE93F8968D5E8723" blockId="7.[305,1284,1882,1902]" box="[305,1284,1882,1902]" pageId="7" pageNumber="9">
FIGURE 4:
<taxonomicName id="95C0A9FBFFE8FFE8FE34F8968A748720" authority="Karg &amp; Schorlemmer, 2011" authorityName="Karg &amp; Schorlemmer" authorityYear="2011" box="[406,558,1883,1901]" class="Arachnida" family="Pyrosejidae" genus="Pyrosejus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="9" phylum="Arthropoda" rank="species" species="verticis" status="sp. nov.">
<emphasis id="60B40E6AFFE8FFE8FE34F8968A748720" box="[406,558,1883,1901]" italics="true" pageId="7" pageNumber="9">Pyrosejus verticis</emphasis>
</taxonomicName>
<emphasis id="60B40E6AFFE8FFE8FD91F8968A3D8723" bold="true" box="[563,615,1883,1902]" pageId="7" pageNumber="9">
<taxonomicNameLabel id="7B87B311FFE8FFE8FD91F8968A3D8723" box="[563,615,1883,1902]" pageId="7" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Female. a Venter; b Tectum, 34 µm wide; c Chelicera, 35 µm long.
</paragraph>
</caption>
<paragraph id="527FD278FFE7FFE7FF67FD54899683A0" blockId="8.[159,762,665,1005]" pageId="8" pageNumber="10">
<taxonomicName id="95C0A9FBFFE7FFE7FF67FD5489AE82FC" authority=": Tritosternum" authorityName="Tritosternum" box="[197,500,665,689]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="8" pageNumber="10" phylum="Nematoda" rank="genus">
<emphasis id="60B40E6AFFE7FFE7FF67FD54891182FC" box="[197,331,665,689]" italics="true" pageId="8" pageNumber="10">Gnathosoma</emphasis>
: Tritosternum
</taxonomicName>
with two laciniae recognizable, anterior margin of tectum denticulate with teeth of different size, margins of hypostome fused, hypostomatic setae on the palpal coxae 9 to 20 µm long: C1 = 20, C2 = 18, C3 = 13, C4 = 9 µm long, paralaciniae 16 µm long with a 8 µm wide dented basis, chelicerae strong and stout, 22 µm wide, digitus mobilis 34 µm long, with a big proximal tooth and 9 10 small teeth, digitus fixus with 7 8 medium-sized teeth.
</paragraph>
<paragraph id="527FD278FFE7FFE7FF67FBED8AA384ED" blockId="8.[159,761,1055,1184]" pageId="8" pageNumber="10">
<emphasis id="60B40E6AFFE7FFE7FF67FBED88AC847A" box="[197,246,1056,1079]" italics="true" pageId="8" pageNumber="10">Legs</emphasis>
: I = 322, II = 261, III = 261, IV = 356 µm long appropriate an idiosoma 348 x 244 µm, tarsi I to IV with paired claws, at leg I longer however without pretarsus, at leg II to IV shorter and with pretarsus.
</paragraph>
<paragraph id="527FD278FFE7FFE7FF67FB1E88B6866A" blockId="8.[159,762,1234,1575]" pageId="8" pageNumber="10">
Taxonomy —
<bibRefCitation id="3651AF89FFE7FFE7FED4FB1F8A9C84A6" author="Lindquist E. E. &amp; Moraza M. L." box="[374,710,1234,1259]" pageId="8" pageNumber="10" pagination="283 - 307" refId="ref10691" refString="Lindquist E. E., Moraza M. L. 1993 - Pyrosejidae, a new family of Trigynaspid Mites (Acari: Mesostigmata: Cercomegistina) from Middle America - Acarologia, 34 (4): 283 - 307." type="journal article" year="1993">Lindquist &amp; Moraza (1993)</bibRefCitation>
described one new species of the new genus
<emphasis id="60B40E6AFFE7FFE7FD33FB3B8AA38540" box="[657,761,1270,1293]" italics="true" pageId="8" pageNumber="10">Pyroseius</emphasis>
and a second form, which they consider to be a second new genus. However, our new species represents beyond its own characteristics also features of both,
<taxonomicName id="95C0A9FBFFE7FFE7FF43FA4E8AC085D7" authority="Lindquist et Moraza, 1993" authorityName="Lindquist et Moraza" authorityYear="1993" box="[225,666,1410,1434]" class="Arachnida" family="Pyrosejidae" genus="Pyrosejus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="10" phylum="Arthropoda" rank="species" species="prionotus">
<emphasis id="60B40E6AFFE7FFE7FF43FA4E893885D4" box="[225,354,1410,1434]" italics="true" pageId="8" pageNumber="10">P. prionotus</emphasis>
<bibRefCitation id="3651AF89FFE7FFE7FECBFA4F8AC085D7" author="Lindquist E. E. &amp; Moraza M. L." box="[361,666,1410,1434]" pageId="8" pageNumber="10" pagination="283 - 307" refId="ref10691" refString="Lindquist E. E., Moraza M. L. 1993 - Pyrosejidae, a new family of Trigynaspid Mites (Acari: Mesostigmata: Cercomegistina) from Middle America - Acarologia, 34 (4): 283 - 307." type="journal article" year="1993">Lindquist et Moraza, 1993</bibRefCitation>
</taxonomicName>
and the assumed second new genus (
<tableCitation id="1F42E7C3FFE7FFE7FE5EFA688A0B85F0" box="[508,593,1445,1469]" captionStart="TABLE" captionStartId="8.[563,576,1634,1653]" captionText="TABLE 1: Diagnostic features of Pyrosejus species." pageId="8" pageNumber="10">Table 1</tableCitation>
). That means we have found an intermediate species. The three forms therefore are separate species of the same genus.
</paragraph>
</subSubSection>
</treatment>
</document>