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<document id="E9E1B5089F5A9E5AE23B5ABF5860351C" ID-CLB-Dataset="307873" ID-DOI="10.11646/zootaxa.5570.1.4" ID-GBIF-Dataset="eb726389-0502-4416-a346-222915e72a20" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734655" ID-ZooBank="1995C18B-1993-4DC4-B8AB-E4891F148A1E" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1737736390657" checkinUser="plazi" docAuthor="Dayananda, Naruwan &amp; Benjamin, Suresh P." docDate="2025" docId="039D5914FF9DFFA3FF4DFC8FFC8DFD7F" docLanguage="en" docName="zootaxa.5570.1.4.pdf" docOrigin="Zootaxa 5570 (1)" docSource="https://doi.org/10.11646/zootaxa.5570.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mallinella truffles Dayananda &amp; Benjamin, 2025, sp. nov." docType="treatment" docVersion="5" lastPageNumber="112" masterDocId="FFA4216CFF97FFAFFFDAFFE9FFD8FFFA" masterDocTitle="Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka" masterLastPageNumber="118" masterPageNumber="100" pageNumber="110" updateTime="1738059402187" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8F6B09D281F787B10CD892C6D8880026">Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka</mods:title>
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<mods:namePart id="C974B40CBA251B79EEB7E5CC5AEBE84B">Dayananda, Naruwan</mods:namePart>
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<mods:namePart id="DA045E4A6BD0B055EA58733C47CCA5D5">Benjamin, Suresh P.</mods:namePart>
<mods:affiliation id="D8ED9B25875ED1AFEA1BB09411C7B9DC">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka.</mods:affiliation>
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<mods:date id="39EF52D63B1176338D91BDD1033F73B0">2025</mods:date>
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<treatment id="039D5914FF9DFFA3FF4DFC8FFC8DFD7F" ID-DOI="http://doi.org/10.5281/zenodo.14753592" ID-Zenodo-Dep="14753592" LSID="urn:lsid:plazi:treatment:039D5914FF9DFFA3FF4DFC8FFC8DFD7F" httpUri="http://treatment.plazi.org/id/039D5914FF9DFFA3FF4DFC8FFC8DFD7F" lastPageId="12" lastPageNumber="112" pageId="10" pageNumber="110" scope_family="Zodariidae" scope_order="Araneae">
<subSubSection id="C32EBB89FF9DFFA5FF4DFC8FFE00FC7B" box="[151,472,870,897]" pageId="10" pageNumber="110" type="nomenclature">
<paragraph id="8B8BE802FF9DFFA5FF4DFC8FFE00FC7B" blockId="10.[151,489,870,933]" box="[151,472,870,897]" pageId="10" pageNumber="110">
<heading id="D0C35F6EFF9DFFA5FF4DFC8FFE00FC7B" bold="true" box="[151,472,870,897]" fontSize="11" level="1" pageId="10" pageNumber="110" reason="1">
<emphasis id="B9403410FF9DFFA5FF4DFC8FFE00FC7B" bold="true" box="[151,472,870,897]" pageId="10" pageNumber="110">
<taxonomicName id="4C349381FF9DFFA5FF4DFC8FFEABFC7B" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[151,371,870,897]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="110" phylum="Arthropoda" rank="species" species="truffles" status="sp. nov.">
<emphasis id="B9403410FF9DFFA5FF4DFC8FFEABFC7B" bold="true" box="[151,371,870,897]" italics="true" pageId="10" pageNumber="110">Mallinella truffles</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273896BFF9DFFA5FEA1FC8EFE00FC7B" box="[379,472,871,897]" pageId="10" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C32EBB89FF9DFFA5FF4DFC62FE31FC5F" box="[151,489,906,933]" pageId="10" pageNumber="110" type="description">
<paragraph id="8B8BE802FF9DFFA5FF4DFC62FE31FC5F" blockId="10.[151,489,870,933]" box="[151,489,906,933]" pageId="10" pageNumber="110">
<figureCitation id="130FF487FF9DFFA5FF4DFC62FF06FC5F" box="[151,222,906,933]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">
Figs
<geoCoordinate id="EE008EC5FF9DFFA5FF14FC63FF06FC5F" box="[206,222,906,933]" degrees="5, 6" direction="east" orientation="longitude" pageId="10" pageNumber="110" precision="5555" value="5.6">5</geoCoordinate>
</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FF30FC62FEEAFC5F" box="[234,306,907,933]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="10" pageNumber="110">6E, 6J</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FEE4FC62FE58FC5F" box="[318,384,907,933]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="10" pageNumber="110">7E,7J</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FE51FC62FE75FC5F" box="[395,429,907,933]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="10" pageNumber="110">8E</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FE63FC62FE31FC5F" box="[441,489,907,933]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="10" pageNumber="110">9IJ</figureCitation>
</paragraph>
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<subSubSection id="C32EBB89FF9DFFA5FF4DFC3AFE1AFBCF" pageId="10" pageNumber="110" type="materials_examined">
<paragraph id="8B8BE802FF9DFFA5FF4DFC3AFE1AFBCF" blockId="10.[151,1437,978,2013]" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF4DFC3AFE9DFC16" bold="true" box="[151,325,978,1005]" pageId="10" pageNumber="110">Type material.</emphasis>
<materialsCitation id="3B5CE25FFF9DFFA5FE96FC3AFC34FBEB" collectingDate="2010-07-13" country="Sri Lanka" elevation="1480" latitude="7.145833" location="Loolecondera Estate" longLatPrecision="21" longitude="80.69806" municipality="Kandy District" pageId="10" pageNumber="110" specimenCount="1" specimenCount-male="1" stateProvince="Central" typeStatus="holotype">
<emphasis id="B9403410FF9DFFA5FE96FC3AFCB8FC17" bold="true" box="[332,864,978,1008]" pageId="10" pageNumber="110">
<typeStatus id="548F56A0FF9DFFA5FE96FC3AFE61FC17" box="[332,441,979,1005]" pageId="10" pageNumber="110" type="holotype">Holotype</typeStatus>
<specimenCount id="9D32238BFF9DFFA5FE1AFC3DFE0FFC0A" box="[448,471,980,1008]" count="1" pageId="10" pageNumber="110" type="male"></specimenCount>
:
<collectingCountry id="F323A892FF9DFFA5FE3CFC3BFD5DFC16" box="[486,645,978,1004]" name="Sri Lanka" pageId="10" pageNumber="110">SRI LANKA</collectingCountry>
:
<emphasis id="B9403410FF9DFFA5FD48FC3BFCB8FC17" bold="true" box="[658,864,978,1005]" italics="true" pageId="10" pageNumber="110">
<collectingRegion id="49F026E0FF9DFFA5FD48FC3BFC81FC17" box="[658,857,978,1005]" country="Sri Lanka" name="Central" pageId="10" pageNumber="110">Central Province</collectingRegion>
:
</emphasis>
</emphasis>
<collectingMunicipality id="6BEF7278FF9DFFA5FCBDFC3AFBC8FC17" box="[871,1040,979,1005]" pageId="10" pageNumber="110">Kandy District</collectingMunicipality>
,
<location id="8EEBBED9FF9DFFA5FBC1FC3AFADBFC17" LSID="urn:lsid:plazi:treatment:039D5914FF9DFFA3FF4DFC8FFC8DFD7F:8EEBBED9FF9DFFA5FBC1FC3AFADBFC17" box="[1051,1283,979,1005]" country="Sri Lanka" latitude="7.145833" longLatPrecision="21" longitude="80.69806" municipality="Kandy District" name="Loolecondera Estate" pageId="10" pageNumber="110" stateProvince="Central">Loolecondera Estate</location>
,
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<geoCoordinate id="EE008EC5FF9DFFA5FF4DFC1FFEFBFBEB" box="[151,291,1014,1041]" degrees="80" direction="east" minutes="41" orientation="longitude" pageId="10" pageNumber="110" precision="15" seconds="53" value="80.69806">80°4153E</geoCoordinate>
,
<quantity id="4CCC45E7FF9DFFA5FEF5FC1EFE5EFBEB" box="[303,390,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="10" pageNumber="110" unit="m" value="1480.0">
<elevation id="00190F31FF9DFFA5FEF5FC1EFE5EFBEB" box="[303,390,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="10" pageNumber="110" unit="m" value="1480.0">1480 m</elevation>
</quantity>
,
<date id="FF8ACEC2FF9DFFA5FE48FC1EFDFDFBEB" box="[402,549,1015,1041]" pageId="10" pageNumber="110" value="2010-07-13">
<collectingDate id="EFCE372AFF9DFFA5FE48FC1EFDFDFBEB" box="[402,549,1015,1041]" pageId="10" pageNumber="110" value="2010-07-13">13 July 2010</collectingDate>
</date>
, leg. SP. Benjamin
<emphasis id="B9403410FF9DFFA5FD27FC1EFCF6FBEB" box="[765,814,1015,1041]" italics="true" pageId="10" pageNumber="110">et al</emphasis>
. (IFS_ZOD_43)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF9DFFA5FC2CFC1EFE67FBCF" collectingDate="2010-07-13" country="Sri Lanka" elevation="1480" latitude="7.145833" location="Loolecondera Estate" longLatPrecision="21" longitude="80.69806" municipality="Kandy District" pageId="10" pageNumber="110" specimenCount="1" specimenCount-female="1" stateProvince="Paratypes" typeStatus="paratype">
<emphasis id="B9403410FF9DFFA5FC2CFC1EFBAFFBEB" bold="true" box="[1014,1143,1015,1041]" pageId="10" pageNumber="110">
<typeStatus id="548F56A0FF9DFFA5FC2CFC1EFBAAFBEB" box="[1014,1138,1015,1041]" pageId="10" pageNumber="110" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D32238BFF9DFFA5FBA4FC1EFB79FBEB" box="[1150,1185,1015,1041]" count="1" pageId="10" pageNumber="110" type="female">1♀</specimenCount>
, with same data as for holotype (IFS_ZOD_230)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9DFFA5FF1DFBD6FAC2FBA3" box="[199,1306,1087,1113]" pageId="10" pageNumber="110" type="etymology">
<paragraph id="8B8BE802FF9DFFA5FF1DFBD6FAC2FBA3" blockId="10.[151,1437,978,2013]" box="[199,1306,1087,1113]" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF1DFBD6FE96FBA3" bold="true" box="[199,334,1087,1113]" pageId="10" pageNumber="110">Etymology.</emphasis>
The new species is named after the chocolate Truffles. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9DFFA5FF1DFB8BFE11FA83" pageId="10" pageNumber="110" type="diagnosis">
<paragraph id="8B8BE802FF9DFFA5FF1DFB8BFE11FA83" blockId="10.[151,1437,978,2013]" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF1DFB8BFE9AFB86" bold="true" box="[199,322,1122,1148]" pageId="10" pageNumber="110">Diagnosis.</emphasis>
<taxonomicName id="4C349381FF9DFFA5FE94FB8AFDC7FB87" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[334,543,1123,1149]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="110" phylum="Arthropoda" rank="species" species="truffles" status="sp. nov.">
<emphasis id="B9403410FF9DFFA5FE94FB8AFDC7FB87" box="[334,543,1123,1149]" italics="true" pageId="10" pageNumber="110">Mallinella truffles</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9DFFA5FDF6FB8AFD53FB87" bold="true" box="[556,651,1123,1149]" pageId="10" pageNumber="110">
<taxonomicNameLabel id="A273896BFF9DFFA5FDF6FB8AFD53FB87" box="[556,651,1123,1149]" pageId="10" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
share features such as unbranched embolus, simple rostrated TA, longitudinal furrow of the epigyne with the members of the tuberculata group. Within tuberculata group, males of
<taxonomicName id="4C349381FF9DFFA5FF4DFB42FED3FB3F" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[151,267,1195,1221]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="110" phylum="Arthropoda" rank="species" species="truffles" status="sp. nov.">
<emphasis id="B9403410FF9DFFA5FF4DFB42FED3FB3F" box="[151,267,1195,1221]" italics="true" pageId="10" pageNumber="110">M. truffles</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9DFFA5FEC8FB42FEB4FB3F" bold="true" box="[274,364,1195,1221]" pageId="10" pageNumber="110">
<taxonomicNameLabel id="A273896BFF9DFFA5FEC8FB42FEB4FB3F" box="[274,364,1195,1221]" pageId="10" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are most similar to the males of
<taxonomicName id="4C349381FF9DFFA5FD02FB42FCB2FB3F" authorityName="Dankittipakul, Jocque &amp; Singtripop" authorityYear="2012" box="[728,874,1195,1221]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="110" phylum="Arthropoda" rank="species" species="brachiata">
<emphasis id="B9403410FF9DFFA5FD02FB42FCB2FB3F" box="[728,874,1195,1221]" italics="true" pageId="10" pageNumber="110">M. brachiata</emphasis>
</taxonomicName>
as both share digitiform RTA, cymbial fold which is less than half the length of the cymbium, triangular apico prolateral process but can be separated from it by long narrow unbranched embolus. (cf.
<bibRefCitation id="EFA595F3FF9DFFA5FDC0FB1AFCE5FAF7" author="Dankittipakul, P. &amp; Jocque, R. &amp; Singtripop, T." box="[538,829,1267,1293]" pageId="10" pageNumber="110" pagination="1 - 327" refId="ref11715" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from south East Asia (Araneae, Zodariidae). Zootaxa, 3369 (1), 1 - 327. https: // doi. org / 10.11646 / zootaxa. 3369.1.1" type="journal article" year="2012">
Dankittipakul
<emphasis id="B9403410FF9DFFA5FD64FB1AFD29FAF7" box="[702,753,1267,1293]" italics="true" pageId="10" pageNumber="110">et al</emphasis>
. 2012
</bibRefCitation>
: figs 147151 and
<figureCitation id="130FF487FF9DFFA5FBC1FB1AFB4EFAF7" box="[1051,1174,1266,1293]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Figs 5FH</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FB7EFB1AFB1FFAF7" box="[1188,1223,1267,1293]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="10" pageNumber="110">8E</figureCitation>
). Females are also most similar to the females of
<taxonomicName id="4C349381FF9DFFA5FE3FFAFEFD52FACB" authorityName="Dankittipakul, Jocque &amp; Singtripop" authorityYear="2012" box="[485,650,1303,1329]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="110" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B9403410FF9DFFA5FE3FFAFEFD52FACB" box="[485,650,1303,1329]" italics="true" pageId="10" pageNumber="110">M. tuberculata</emphasis>
</taxonomicName>
as both share sub-rectangular epigynal plate, and deep anterior median depression, but can be separated from it by globular spermathecae (cf.
<bibRefCitation id="EFA595F3FF9DFFA5FC68FAD2FB16FAAF" author="Dankittipakul, P. &amp; Jocque, R. &amp; Singtripop, T." box="[946,1230,1339,1365]" pageId="10" pageNumber="110" pagination="1 - 327" refId="ref11715" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from south East Asia (Araneae, Zodariidae). Zootaxa, 3369 (1), 1 - 327. https: // doi. org / 10.11646 / zootaxa. 3369.1.1" type="journal article" year="2012">
Dankittipakul
<emphasis id="B9403410FF9DFFA5FB89FAD2FB5CFAAF" box="[1107,1156,1339,1365]" italics="true" pageId="10" pageNumber="110">et al</emphasis>
. 2012
</bibRefCitation>
: figs 152155 and
<figureCitation id="130FF487FF9DFFA5FF4DFAB6FEC3FA83" box="[151,283,1374,1401]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Figs. 5CD</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FEFCFAB6FEB6FA83" box="[294,366,1375,1401]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="10" pageNumber="110">7E, 7J</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FEA0FAB6FE65FA83" box="[378,445,1375,1401]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="10" pageNumber="110">9I9J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9DFFA3FF1DFA6BFA9EFD9B" lastPageId="12" lastPageNumber="112" pageId="10" pageNumber="110" type="description">
<paragraph id="8B8BE802FF9DFFA5FF1DFA6BFB45F88B" blockId="10.[151,1437,978,2013]" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF1DFA6BFE81FA66" bold="true" box="[199,345,1410,1436]" pageId="10" pageNumber="110">Description.</emphasis>
<emphasis id="B9403410FF9DFFA5FE84FA6AFE40FA67" box="[350,408,1411,1437]" italics="true" pageId="10" pageNumber="110">Male</emphasis>
in alcohol (
<typeStatus id="548F56A0FF9DFFA5FDCCFA6AFDA5FA67" box="[534,637,1411,1437]" pageId="10" pageNumber="110" type="holotype">holotype</typeStatus>
;
<figureCitation id="130FF487FF9DFFA5FD5DFA6AFD06FA67" box="[647,734,1410,1437]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Fig. 5A</figureCitation>
). Body length 4.96; carapace 2.66 long, 2.20 wide; opisthosoma 2.30 long, 1.60 wide. Habitus as in
<figureCitation id="130FF487FF9DFFA5FDF8FA4EFDA2FA3B" box="[546,634,1447,1473]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="10" pageNumber="110">Fig. 7A</figureCitation>
. Carapace ovoid, smooth, red-brown; fovea red-brown. Clypeus brown, 0.57 high. Eye sizes and inter-distances: AME 0.16, ALE 0.10, PME 0.12, PLE 0.10; AMEAME 0.10, AMEPME 0.13, AMEALE 0.12, ALEALE 0.55, PMEPME 0.12, PMEPLE 0.25, PLEPLE 0.76, ALEPLE 0.06. MOA 0.44 long, front width 0.38, back width 0.41. Chelicerae 0.89 long, light brown, without teeth. Endites yellow. Labium triangular, yellow-brown, 0.20 long, 0.35 wide. Sternum yellowish, furnished with sparse black setae, 0.89 long, 0.74 wide. Lateral margins of sternum with small semi-circular pits in front of each coxa of third and fourth pair of legs. Legs yellowish, with proximal tubercle. Measurements of palp and legs: pedipalp (right) 2.36 (0.83 + 0.40 + 0.44 + 0.69), I 6.10 (1.60 + 0.65 + 1.40 + 1.45+ 1.00), II 5.45 (1.45 + 0.65 + 1.10 + 1.25 + 1.00), III 6.00 (1.60 + 0.65 + 1.25 + 1.50 + 1.00), IV 8.00 (2.00 + 0.70+ 1.60 + 2.35 + 1.35). Leg formula: 4132. Opisthosoma oval; pattern on dorsum of opisthosoma provided large pale patches, roughly oval first pair of patches connected vertically with nearly round second pair, third and fourth pairs of nearly round patches connected medially and vertically with each other (
<figureCitation id="130FF487FF9DFFA5FE0BF8DAFDF7F8B7" box="[465,559,1842,1869]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Fig. 5A</figureCitation>
). Dorsum of opisthosoma with large ovoid scutum. Posterior ventral spines elongated, gradually tapering, with acute apices (
<figureCitation id="130FF487FF9DFFA5FD67F8BEFCCFF88B" box="[701,791,1878,1905]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Fig. 5A</figureCitation>
). Spinnerets pale yellow (
<figureCitation id="130FF487FF9DFFA5FBE3F8BEFB48F88B" box="[1081,1168,1878,1905]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Fig. 5E</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF9DFFA4FF1DF892FD71FF2F" blockId="10.[151,1437,978,2013]" lastBlockId="11.[151,1436,150,213]" lastPageId="11" lastPageNumber="111" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF1DF892FF24F86F" box="[199,252,1915,1941]" italics="true" pageId="10" pageNumber="110">Palp</emphasis>
(
<figureCitation id="130FF487FF9DFFA5FED5F892FE57F86F" box="[271,399,1914,1941]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Figs 5FH</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FE45F892FE1AF86F" box="[415,450,1915,1941]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="10" pageNumber="110">8E</figureCitation>
). VTA elevated subtriangular hump (
<figureCitation id="130FF487FF9DFFA5FCACF892FC00F86F" box="[886,984,1914,1941]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Figs 5G</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FC32F892FBD3F86F" box="[1000,1035,1915,1941]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="10" pageNumber="110">8E</figureCitation>
); RTA digitiform, tapered towards blunt apex (
<figureCitation id="130FF487FF9DFFA5FEFBF876FE74F843" box="[289,428,1950,1977]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Figs. 5GH</figureCitation>
); Cymbium with deep retrolateral fold, less than half the length of cymbium (
<figureCitation id="130FF487FF9DFFA5FAE9F876FA57F843" box="[1331,1423,1950,1977]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Fig. 5G</figureCitation>
), apicoprolateral process triangular with blunt apex, TT large triangular with pointed apex. Embolic base aligned in diagonally, directed posteriorly, with narrow membranous area (
<figureCitation id="130FF487FF9CFFA4FCB8FF7EFC64FF4B" box="[866,956,150,177]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="11" pageNumber="111">Fig. 5G</figureCitation>
). Embolus, long, narrow, unbranched, with blunt apex. Conductor triangular, pointed apex.
</paragraph>
<caption id="DF4BB88AFF9CFFA4FF4DF922FD8DF825" ID-DOI="http://doi.org/10.5281/zenodo.14734672" ID-Zenodo-Dep="14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="11" pageNumber="111" startId="11.[151,250,1739,1763]" targetBox="[275,1312,241,1710]" targetPageId="11" targetType="figure">
<paragraph id="8B8BE802FF9CFFA4FF4DF922FD8DF825" blockId="11.[151,1437,1739,2016]" pageId="11" pageNumber="111">
<emphasis id="B9403410FF9CFFA4FF4DF922FECAF91E" bold="true" box="[151,274,1739,1764]" pageId="11" pageNumber="111">FIGURE 5.</emphasis>
<taxonomicName id="4C349381FF9CFFA4FEC2F922FE08F919" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[280,464,1739,1763]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="111" phylum="Arthropoda" rank="species" species="truffles" status="sp. nov.">
<emphasis id="B9403410FF9CFFA4FEC2F922FE08F919" box="[280,464,1739,1763]" italics="true" pageId="11" pageNumber="111">Mallinella truffles</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9CFFA4FE0FF925FDFDF91E" bold="true" box="[469,549,1740,1764]" pageId="11" pageNumber="111">
<taxonomicNameLabel id="A273896BFF9CFFA4FE0FF925FDFDF91E" box="[469,549,1740,1764]" pageId="11" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8BE802FF9BFFA3FF1DFF7EFEDDFE2B" blockId="12.[151,1437,150,645]" pageId="12" pageNumber="112">
<emphasis id="B9403410FF9BFFA3FF1DFF7EFEC3FF4B" box="[199,283,151,177]" italics="true" pageId="12" pageNumber="112">Female</emphasis>
in alcohol (
<typeStatus id="548F56A0FF9BFFA3FE73FF7EFDD7FF4B" box="[425,527,151,177]" pageId="12" pageNumber="112" type="paratype">paratype</typeStatus>
;
<figureCitation id="130FF487FF9BFFA3FDC6FF7EFDAEFF4B" box="[540,630,150,177]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Fig. 5B</figureCitation>
); Body length 6.46; carapace 2.97 long, 2.01 wide; opisthosoma 3.49 long, 2.46 wide. Habitus and details as in male except for the following (
<figureCitation id="130FF487FF9BFFA3FC1AFF52FBC0FF2F" box="[960,1048,186,213]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Fig. 5B</figureCitation>
). Clypeus 0.34 high. Eye sizes and inter-distances:
<collectionCode id="ED2570C7FF9BFFA3FE92FF36FE5EFF03" box="[328,390,223,249]" pageId="12" pageNumber="112">AME</collectionCode>
0.13,
<collectionCode id="ED2570C7FF9BFFA3FE11FF36FDD9FF03" box="[459,513,223,249]" pageId="12" pageNumber="112">ALE</collectionCode>
0.11,
<collectionCode id="ED2570C7FF9BFFA3FD92FF36FD5AFF03" box="[584,642,223,249]" pageId="12" pageNumber="112">PME</collectionCode>
0.12,
<collectionCode id="ED2570C7FF9BFFA3FD12FF36FD22FF03" box="[712,762,223,249]" pageId="12" pageNumber="112">PLE</collectionCode>
0.11;
<collectionCode id="ED2570C7FF9BFFA3FCE5FF36FCA5FF03" box="[831,893,223,249]" pageId="12" pageNumber="112">AME</collectionCode>
<collectionCode id="ED2570C7FF9BFFA3FC51FF36FC11FF03" box="[907,969,223,249]" pageId="12" pageNumber="112">AME</collectionCode>
0.09,
<collectionCode id="ED2570C7FF9BFFA3FBD5FF36FB95FF03" box="[1039,1101,223,249]" pageId="12" pageNumber="112">AME</collectionCode>
<collectionCode id="ED2570C7FF9BFFA3FB81FF36FB4DFF03" box="[1115,1173,223,249]" pageId="12" pageNumber="112">PME</collectionCode>
0.18,
<collectionCode id="ED2570C7FF9BFFA3FB00FF36FAC0FF03" box="[1242,1304,223,249]" pageId="12" pageNumber="112">AME</collectionCode>
<collectionCode id="ED2570C7FF9BFFA3FAFCFF36FA84FF03" box="[1318,1372,223,249]" pageId="12" pageNumber="112">ALE</collectionCode>
0.13,
<collectionCode id="ED2570C7FF9BFFA3FF4DFEEAFF15FEE7" box="[151,205,259,285]" pageId="12" pageNumber="112">ALE</collectionCode>
<collectionCode id="ED2570C7FF9BFFA3FF06FEEAFECAFEE7" box="[220,274,259,285]" pageId="12" pageNumber="112">ALE</collectionCode>
0.60,
<collectionCode id="ED2570C7FF9BFFA3FE80FEEAFE4CFEE7" box="[346,404,259,285]" pageId="12" pageNumber="112">PME</collectionCode>
<collectionCode id="ED2570C7FF9BFFA3FE78FEEAFE04FEE7" box="[418,476,259,285]" pageId="12" pageNumber="112">PME</collectionCode>
0.13,
<collectionCode id="ED2570C7FF9BFFA3FDFEFEEAFD85FEE7" box="[548,605,259,285]" pageId="12" pageNumber="112">PME</collectionCode>
<collectionCode id="ED2570C7FF9BFFA3FDB1FEEAFD45FEE7" box="[619,669,259,285]" pageId="12" pageNumber="112">PLE</collectionCode>
0.26,
<collectionCode id="ED2570C7FF9BFFA3FD3FFEEAFCCFFEE7" box="[741,791,259,285]" pageId="12" pageNumber="112">PLE</collectionCode>
<collectionCode id="ED2570C7FF9BFFA3FCFFFEEAFC8FFEE7" box="[805,855,259,285]" pageId="12" pageNumber="112">PLE</collectionCode>
0.78,
<collectionCode id="ED2570C7FF9BFFA3FC44FEEAFC0CFEE7" box="[926,980,259,285]" pageId="12" pageNumber="112">ALE</collectionCode>
<collectionCode id="ED2570C7FF9BFFA3FC38FEEAFBCCFEE7" box="[994,1044,259,285]" pageId="12" pageNumber="112">PLE</collectionCode>
0.04.
<collectionCode id="ED2570C7FF9BFFA3FB86FEEAFB44FEE7" box="[1116,1180,259,285]" pageId="12" pageNumber="112">MOA</collectionCode>
0.45 long, front width 0.36, back width 0.38. Labium 0.34 long, 0.47 wide. Sternum with shallow semi-circular pits, 1.25 long, 1.08 wide. Measurements of palp and legs: pedipalp (right) 2.25 (0.81 + 0.37 + 0.40 + 0.67) I 6.65 (1.80 + 0.75 + 1.60 + 1.55 + 0.95), II 6.60 (1.65 + 0.75 + 1.40 + 1.55 + 1.25), III 6.70 (1.70 + 0.75+ 1.50 + 1.75 + 1.00), IV 8.05 (1.95 + 0.75 + 1.75 + 2.25 + 1.35). Dorsum pattern of opisthosoma, indistinct, covered with thick black and white pubescence (
<figureCitation id="130FF487FF9BFFA3FF45FE5EFF20FE2B" box="[159,248,438,465]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF9BFFA3FF1DFE32FA9EFD9B" blockId="12.[151,1437,150,645]" pageId="12" pageNumber="112">
<emphasis id="B9403410FF9BFFA3FF1DFE32FEE9FE0F" box="[199,305,475,501]" italics="true" pageId="12" pageNumber="112">Genitalia</emphasis>
(
<figureCitation id="130FF487FF9BFFA3FE9BFE32FE19FE0F" box="[321,449,474,501]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Figs 5CD</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FE14FE32FDC0FE0F" box="[462,536,475,501]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="12" pageNumber="112">7E, 7J</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FDFFFE32FDB0FE0F" box="[549,616,475,501]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="12" pageNumber="112">9I9J</figureCitation>
). Epigynal plate wide, sub rectangular, with curved sclerotized epigynal furrows directed towards posteriorly (
<figureCitation id="130FF487FF9BFFA3FD8EFE16FD6EFDE3" box="[596,694,510,537]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Figs 5C</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FD1DFE16FD3AFDE3" box="[711,738,511,537]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="12" pageNumber="112">9I</figureCitation>
). Lateral borders sub-triangular, terminally acute (
<figureCitation id="130FF487FF9BFFA3FAECFE16FA40FDE3" box="[1334,1432,510,537]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Figs 5C</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FF4DFDCAFF6AFDC7" box="[151,178,547,573]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="12" pageNumber="112">9I</figureCitation>
). Insemination ducts short, thick, distally diverging (
<figureCitation id="130FF487FF9BFFA3FCC5FDCAFC59FDC7" box="[799,897,546,573]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Figs 5D</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FC4AFDCAFC05FDC7" box="[912,989,547,573]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="12" pageNumber="112">7E, 7J</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FC37FDCAFBD1FDC7" box="[1005,1033,547,573]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="12" pageNumber="112">9J</figureCitation>
), longer than spermathecal length. Spermathecae, globular in shape distally with coiled internal ducts (
<figureCitation id="130FF487FF9BFFA3FC55FDAEFC07FD9B" box="[911,991,583,609]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="12" pageNumber="112">Fig. 9J</figureCitation>
). Fertilization ducts as in fig 7J.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9BFFA3FF1DFD83FC8DFD7F" box="[199,853,618,645]" pageId="12" pageNumber="112" type="distribution">
<paragraph id="8B8BE802FF9BFFA3FF1DFD83FC8DFD7F" blockId="12.[151,1437,150,645]" box="[199,853,618,645]" pageId="12" pageNumber="112">
<emphasis id="B9403410FF9BFFA3FF1DFD83FEB9FD7E" bold="true" box="[199,353,618,644]" pageId="12" pageNumber="112">Distribution.</emphasis>
Known only from the
<typeStatus id="548F56A0FF9BFFA3FDBAFD82FD48FD7F" box="[608,656,619,645]" pageId="12" pageNumber="112">type</typeStatus>
locality (
<figureCitation id="130FF487FF9BFFA3FD21FD82FC90FD7F" box="[763,840,619,645]" captionStart="FIGURE 12" captionStartId="17.[151,250,1133,1157]" captionTargetId="figure-90@17.[426,1162,345,1110]" captionTargetPageId="17" captionText="FIGURE 12. Distribution map of known Mallinella species of Sri Lanka." figureDoi="http://doi.org/10.5281/zenodo.14753588" httpUri="https://zenodo.org/record/14753588/files/figure.png" pageId="12" pageNumber="112">Fig.12</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>