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<mods:titleid="96B0B44D635369889930F78F372C005D">Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka</mods:title>
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<mods:titleid="CEEDB97FB450FF4CBCFCAF9FECD63B27">Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka</mods:title>
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<mods:titleid="DC183FE03D8969185E537F8814D4AB27">Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka</mods:title>
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0.39 long, front width 0.28, back width 0.32. Chelicerae 0.83 long Labium 0.30 long, 0.44 wide. Sternum yellow with shallow semi-circular pits, 1.06 long, 1.04 wide. Legs with proximal tubercle on anterior femora (
<figureCitationid="130FF487FF9DFFA5FD21FD12FC96FCEF"box="[763,846,763,789]"captionStart="FIGURE 12"captionStartId="17.[151,250,1133,1157]"captionText="FIGURE 12. Distribution map of known Mallinella species of Sri Lanka." pageId="10"pageNumber="110">Fig. 12</figureCitation>
<figureCitationid="130FF487FF9DFFA5FD21FD12FC96FCEF"box="[763,846,763,789]"captionStart="FIGURE 12"captionStartId="17.[151,250,1133,1157]"captionTargetId="figure-90@17.[426,1162,345,1110]"captionTargetPageId="17"captionText="FIGURE 12. Distribution map of known Mallinella species of Sri Lanka." figureDoi="http://doi.org/10.5281/zenodo.14753588"httpUri="https://zenodo.org/record/14753588/files/figure.png" pageId="10"pageNumber="110">Fig. 12</figureCitation>
<documentid="D0E53EC63C8636920DB0ED91588BFEF7"ID-DOI="10.11646/zootaxa.5570.1.5"ID-ISSN="1175-5326"ID-Zenodo-Dep="14734630"ID-ZooBank="8EF2BC5F-5939-440C-A053-A8924F07A42C"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="diego"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="diego"IM.treatments_approvedBy="diego"checkinTime="1737736316380"checkinUser="plazi"docAuthor="Zheng, Xinyi, Li, Yuang, Tan, Zhixiang & Wu, San’An"docDate="2025"docId="03CD8437232DFF82FF5C578DFD0BF8A6"docLanguage="en"docName="zootaxa.5570.1.5.pdf"docOrigin="Zootaxa 5570 (1)"docSource="https://doi.org/10.11646/zootaxa.5570.1.5"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Puto sinensis Zheng & Wu 2025, sp. n."docType="treatment"docVersion="1"lastPageNumber="133"masterDocId="FFF4FC4F2325FF8CFFCB537EFFB5FF8F"masterDocTitle="A new species of the genus Puto Signoret (Hemiptera: Coccomorpha: Putoidae) from China and redescription of Puto huangshanensis Wu"masterLastPageNumber="137"masterPageNumber="119"pageNumber="127"updateTime="1738059294444"updateUser="diego"zenodo-license-document="CLOSED">
<mods:titleid="71D637EF973B0FE8D40BFEDE9BD34755">A new species of the genus Puto Signoret (Hemiptera: Coccomorpha: Putoidae) from China and redescription of Puto huangshanensis Wu</mods:title>
<mods:affiliationid="F3058BD91DFC0B95A251C4A48B845641">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<mods:affiliationid="24A5277F12CB5A66AE104D178416AD10">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<mods:affiliationid="DBC20AC5B68460CF801743AC97DA3747">Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025, China</mods:affiliation>
<mods:affiliationid="03303358A0E321506519DBEF2C71D0F8">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<taxonomicNameid="4C644EA2232DFF84FF5C578DFE50FA82"authority="Zheng & Wu"authorityName="Zheng & Wu"authorityYear="2025"box="[151,485,1266,1293]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="8"pageNumber="127"phylum="Arthropoda"rank="species"species="sinensis"status="sp. nov.">
<materialsCitationid="3B0C3F7C232DFF84FE465645FA2CFAF6"collectedFrom="on Lindera communis (Lauraceae)"collectingDate="2023-05-01"collectorName="Zhixiang Tan"country="China"county="Kaiyang County"location="Nanjiang Grand Canyon"municipality="Guiyang City"pageId="8"pageNumber="127"specimenCount="2"specimenCount-adult="1"specimenCount-female="1"stateProvince="Guizhou Prov."typeStatus="holotype">
<locationid="8EBB63FA232DFF84FAFD5645FE8AFAF6"LSID="urn:lsid:plazi:treatment:03CD8437232DFF82FF5C578DFD0BF8A6:8EBB63FA232DFF84FAFD5645FE8AFAF6"country="China"county="Kaiyang County"municipality="Guiyang City"name="Nanjiang Grand Canyon"pageId="8"pageNumber="127"stateProvince="Guizhou Prov.">Nanjiang Grand Canyon</location>
<collectingDateid="EF9EEA09232DFF84FBE65621FB0FFAF6"box="[1069,1210,1375,1401]"pageId="8"pageNumber="127"value="2023-05-01">1 May 2023</collectingDate>
<materialsCitationid="3B0C3F7C232DFF84FBD35691FA23F9FA"collectedFrom="Photinia serratifolia (Rosaceae"collectingDate="2021-04-20"collectionCode="BFUC"collectorName="San-an Wu"country="China"county="Yao Autonomous County of Ruyuan"location="Nanling National Forest Park"municipality="Shaoguan City"pageId="8"pageNumber="127"specimenCount="5"specimenCount-female="2"specimenCount-male="3"stateProvince="Guangdong Prov.">
<collectingCountyid="62BA4DAD232DFF84FD2B5549FBCAF9DE"box="[736,1151,1591,1617]"pageId="8"pageNumber="127">Yao Autonomous County of Ruyuan</collectingCounty>
,
<locationid="8EBB63FA232DFF84FB405549FF78F9FA"LSID="urn:lsid:plazi:treatment:03CD8437232DFF82FF5C578DFD0BF8A6:8EBB63FA232DFF84FB405549FF78F9FA"country="China"county="Yao Autonomous County of Ruyuan"municipality="Shaoguan City"name="Nanling National Forest Park"pageId="8"pageNumber="127"stateProvince="Guangdong Prov.">Nanling National Forest Park</location>
<emphasisid="B910E933232DFF84FF5C5428FEC7F8FF"bold="true"box="[151,370,1878,1904]"pageId="8"pageNumber="127">Appearance in life</emphasis>
(
<figureCitationid="135F29A4232DFF84FE4A5429FD92F8FE"box="[385,551,1879,1905]"captionStart="FIGURE 1"captionStartId="3.[151,250,1113,1137]"captionTargetBox="[151,1436,234,1086]"captionTargetId="figure-45@3.[151,1436,234,1086]"captionTargetPageId="3"captionText="FIGURE 1. Habitus photographs. Puto huangshanensis Wu: A: dorsal view of adult female. Puto sinensis Zheng & Wu, sp. n.: B: habitat of the type samples; C: dorsal view of adult female (photographed by Rixin Jiang); D: adult male and white, felted cocoons on fallen leaves."figureDoi="http://doi.org/10.5281/zenodo.14734634"httpUri="https://zenodo.org/record/14734634/files/figure.png"pageId="8"pageNumber="127">Figs 1B and C</figureCitation>
). Body oval, dorsum covered with plates of dense white flocculent wax, venter thinly coated with mealy white wax.
<collectionCodeid="ED75ADE4232CFF85FDFA53E9FDE4FF3E"box="[561,593,151,177]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="9"pageNumber="128">VI</collectionCode>
, 140–155; VII, 138–148; VIII, 128–155; and IX, 168–208. Each segment nearly cylindrical, except apical segment ovoid; antenna with few hair-like setae, each 75–150 μm long. Scape (segment I) widest, 153–180 μm wide, with fewest hair-like setae (4 or 5); pedicel (segment II) (
<figureCitationid="135F29A4232CFF85FB1453A1FA8DFF76"box="[1247,1336,223,249]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4A</figureCitation>
) 68–103 μm wide, with 1 circular sensory pore and 1 or 2 coeloconic sensilla; remaining segments each 50–66 μm wide. Segments III, IV and
<collectionCodeid="ED75ADE4232CFF85FE4D5259FE11FECE"box="[390,420,295,321]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="9"pageNumber="128">VI</collectionCode>
each with a pair of intersegmental sensilla (
<figureCitationid="135F29A4232CFF85FC465259FC56FECE"box="[909,995,295,321]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4B</figureCitation>
) near distal end; segments VII and VIII each with a sensory seta near apex, 45–78 μm long; apical segment (IX) with 20–23 hair-like setae, 3 sensory setae, and sometimes with 1 coeloconic sensilla. Eyes protruding, each with stem sclerotized and apex membranous; eye prominence 155–185 μm long, situated posterior to scape. Clypeolabral shield 385–440 μm long, with anterior part sinking into derm, posterior part exposed and bearing a pair of hair-like setae, each seta about 185 μm long. Labium (
<figureCitationid="135F29A4232CFF85FF5452A5FF4DFE7A"box="[159,248,475,501]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4C</figureCitation>
) 3-segmented, 335–360 μm long; segment I poorly sclerotized, bearing 2 hair-like setae (each 58–130 μm long) and 1 spine-like seta 45–55 μm on each side; segments II and III sclerotized, segment II with 1 hair-like seta on each side; segment III with many hair-like setae, shorter setae near apex each 10–40 μm long. Legs well developed, lengths (in μm): foreleg shortest: entire length
; coxa 330–405; trochanter + femur 890–930; tibia 880–920; tarsus 240–275, and claw 60–88. Hind leg tibia + tarsus 1.30– 1.34 times as long as trochanter + femur; tibia 3.2–3.5 times as long as tarsus. Legs each with many hair-like setae, each 68–135 μm long; translucent pores absent. Trochanters each with 2 basal setae (about 5 μm long) and bearing 3 sensory pores on each surface plus a long hair-like seta, 273–350 μm long. Femora each with some spine-like setae on ventral side, each seta 35–55 μm long. Tibiae each with some short spine-like setae, each about 35 μm long. Tarsi each with 1 tarsal campaniform sensillum proximally; tarsal digitules each with pointed apex. Claws (
<figureCitationid="135F29A4232CFF85FAF65019FA21FC0E"box="[1341,1428,871,897]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4D</figureCitation>
) each with a plantar denticle and a pair of basal spurs; claw digitules each 60–68 μm long with capitate apex, longer than claw. Thoracic spiracles (
<figureCitationid="135F29A4232CFF85FE3B50D1FDF0FC46"box="[496,581,943,969]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4E</figureCitation>
) each with opening 105–125 μm in diameter, metathoracic spiracles larger. Circulus short and wide, 270–375 μm wide, more than three times as wide as long, without a medial constriction, situated inside posterior margin of abdominal segment III. Anterior and posterior ostioles each with inner edges of lips sclerotized, each lip with numerous lanceolate setae and trilocular pores, without oral collar tubular ducts. Anal ring (
<figureCitationid="135F29A4232CFF85FF1E5741FE9FFBD6"box="[213,298,1087,1113]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4F</figureCitation>
) 125–135 μm in diameter, with 3–5 rows of cells in a single anal ring, and bearing 6 sharply-tipped setae, each 275–340 μm long. Nearby is 1 pair of cisanal setae, each 160–245 μm long. Cerarii numbering 17 basic pairs (preocular cerarii (C
) absent); also a few supplementary cerarii usually present between thorax and abdominal segment IV. Cerarii each with many lanceolate setae (
<figureCitationid="135F29A4232CFF85FD3257D5FCE7FB4A"box="[761,850,1195,1221]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4G</figureCitation>
) (each seta 30–48 μm long and about 8 μm wide at widest point) and a compact group of trilocular pores (
<figureCitationid="135F29A4232CFF85FD3057B1FCE0FB66"box="[763,853,1231,1257]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4H</figureCitation>
), each pore 7–8 μm in diameter, but without tubular ducts. Cerarii each situated on a weakly sclerotized area with margin not clearly defined. Anal lobe cerarii each containing 25–30 lanceolate setae and 35–40 trilocular pores, all situated on a sclerotized area (145–200 μm wide) occupying most of lobe, with sclerotized area larger than anal ring. A few discoidal pores (
<figureCitationid="135F29A4232CFF85FBB55645FB7EFADA"box="[1150,1227,1339,1365]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4I</figureCitation>
) present near most cerarii, each about 3 μm in diameter.
<figureCitationid="135F29A4232CFF85FDF856FDFD37FA12"box="[563,642,1411,1437]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4J</figureCitation>
) each 20–40 μm long, smaller than a cerarian seta; forming group on head and loose transverse bands across other segments. Multilocular disc-pores absent. Trilocular pores (
<figureCitationid="135F29A4232CFF85FAF056D9FA21FA4E"box="[1339,1428,1447,1473]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4K</figureCitation>
) each 6–7 μm in diameter, numerous, forming broad transverse bands across each segment. Submarginal line of relatively bare cuticle with only a few lanceolate setae and trilocular pores. Oral collar tubular ducts (
<figureCitationid="135F29A4232CFF85FAD55691FAC1F986"box="[1310,1396,1519,1545]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4L</figureCitation>
, each outer ductule 23–25 μm long and 8–10 μm wide; with 2–5 ducts present submedially on head, on submargins of thorax, and forming groups (of up to 4 ducts) posterior to most cerarii.
Lanceolate setae absent. With slender flagellate setae (
<figureCitationid="135F29A4232CFF85FC5B5525FC47F9FA"box="[912,1010,1627,1653]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4M</figureCitation>
), each 75–230 μm long, scattered on head, around legs and abdominal spiracles, and on median area of thorax; also forming transverse band across each abdominal segment, and in small groups around vulva and on submargins of each segment. With multilocular disc-pores of 2 sizes: (i) small disc-pores (
<figureCitationid="135F29A4232CFF85FDF255B9FD26F96E"box="[569,659,1735,1761]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4N</figureCitation>
), each 10–12 μm in diameter, with 5, 7 or 9 (mostly 7) circular loculi; numerous from head to abdominal segment
<collectionCodeid="ED75ADE4232CFF85FD4C5595FD12F88A"box="[647,679,1771,1797]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="9"pageNumber="128">VI</collectionCode>
, reaching almost to margins except on abdominal segment
<collectionCodeid="ED75ADE4232CFF85FA895595FAD6F88A"box="[1346,1379,1771,1797]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="9"pageNumber="128">VI</collectionCode>
; also forming group near each thoracic spiracle; and (ii) large disc-pores (
<figureCitationid="135F29A4232CFF85FC575471FC43F8A6"box="[924,1014,1807,1833]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4O</figureCitation>
), each about 13 μm in diameter, with 5 or 7 (mostly 7) kidney-shaped loculi; present on posteriormost 3 abdominal segments, numerous around vulva, not reaching to margins. Trilocular pores (
<figureCitationid="135F29A4232CFF85FDB35429FD7BF8FE"box="[632,718,1879,1905]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4P</figureCitation>
), each about 5 μm in diameter (smaller than those on dorsum), distributed similarly to that of setae but more numerous on margins and posterior areas of abdomen. Oral collar tubular ducts of
<figureCitationid="135F29A4232CFF85FD8C54E1FD29F836"box="[583,668,1951,1977]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4L</figureCitation>
), same size as ducts on dorsum, few on margins of posterior abdomen; (ii) small ducts (
<figureCitationid="135F29A4232CFF85FE9854BDFE1BF852"box="[339,430,1987,2013]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="9"pageNumber="128">Fig. 4Q</figureCitation>
), each outer ductule 19–23 μm long and 5–7 μm wide; forming groups on margins, also with few near each thoracic spiracle and pro- and meso-coxa, and also forming short submedial transverse bands on abdominal segments IV–VII; and (iii) slender ducts (
<figureCitationid="135F29A4232EFF87FD2453C5FCFCFF5A"box="[751,841,187,213]"captionStart="FIGURE 4"captionStartId="10.[151,250,1856,1880]"captionTargetBox="[269,1316,183,1828]"captionTargetId="figure-23@10.[269,1318,181,1832]"captionTargetPageId="10"captionText="FIGURE 4. Adult female of Puto sinensis Zheng & Wu, sp. n. A: antennal segment II; B: intersegmental sensillum; C: labium; D: claw; E: thoracic spiracle; F: anal ring; G: lanceolate cerarian seta; H: trilocular pore found in cerarii; I. discoidal pore; J: lanceolate seta on dorsum; K: trilocular pore on dorsum; L: large oral collar tubular duct; M: slender flagellate seta; N: small multilocular disc-pores; O: large multilocular disc-pore; P: trilocular pore on venter; Q: small oral collar tubular duct; R: slender oral collar tubular duct."figureDoi="http://doi.org/10.5281/zenodo.14734642"httpUri="https://zenodo.org/record/14734642/files/figure.png"pageId="11"pageNumber="130">Fig. 4R</figureCitation>
), each outer ductule 25–28 μm long and about 3 μm wide, sometimes deformed; with about 10 ducts anterior to clypeolabral shield. Each anal lobe with 1 anal lobe seta, 290–350 μm long.
<taxonomicNameid="4C644EA2232FFF86FE77543FFD78F8D7"authority="Zheng & Wu"authorityName="Zheng & Wu"authorityYear="2025"box="[444,717,1856,1880]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="10"pageNumber="129"phylum="Arthropoda"rank="species"species="sinensis"status="sp. nov.">
(i) circulus on abdominal segment III complete, more than three times as wide as long, without a constriction; (ii) anal lobe cerarius containing 25–30 lanceolate setae; (iii) abundant small multilocular disc-pores present from head to abdominal segment
<collectionCodeid="ED75ADE4232EFF87FD335211FCACFE06"box="[760,793,367,393]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="11"pageNumber="130">VI</collectionCode>
; (iv) numerous large multilocular disc-pores present on abdominal segments
<collectionCodeid="ED75ADE4232EFF87FE4352EDFE13FE22"box="[392,422,403,429]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="11"pageNumber="130">VI</collectionCode>
–VIII; (v) ostioles with setae on lips; (vi) anal ring bearing six acute setae; (vii) dorsum with a few large oral collar tubular ducts posterior to sclerotized areas of most cerarii (not within or immediately adjacent to sclerotized areas), also with a few present submedially on head and submargins of thorax; (viii) a few large oral collar tubular ducts ventrally on margins of posterior abdomen; (ix) venter with small oral collar tubular ducts forming groups on margins, and short submedial transverse bands on abdominal segments IV–VII; (x) claw digitules clubbed, extending beyond apex of claw; (xi) cerarii numbering 17 basic pairs, supplementary cerarii usually present also, between thorax and abdominal segment IV; (xii) a few slender oral collar tubular ducts present anterior to clypeolabral shield; and (xiii) venter without lanceolate setae.
<taxonomicNameid="4C644EA2232EFF87FDF851CDFD1FFD43"authorityName="Zheng & Wu"authorityYear="2025"box="[563,682,691,716]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="11"pageNumber="130"phylum="Arthropoda"rank="species"species="sinensis">
<bibRefCitationid="EFF548D0232EFF87FB8751CDFB4EFD42"author="Williams, D. J."box="[1100,1275,691,717]"pageId="11"pageNumber="130"refId="ref20067"refString="Williams, D. J. (2004) Mealybugs of Southern Asia. The Natural History Museum, London. Southdene SDN. BHD, Kuala Lumpur, 896 pp."type="book"year="2004">
in having the following character states: (i) a single circulus situated on venter of abdominal segment III; (ii) anal ring bearing sharply-tipped setae; (iii) oral collar tubular ducts absent from cerarii; (iv) dorsum without additional dorsal cerarii; (v) wide oral collar tubular ducts present on dorsum of head and thorax, and (vi) wide tubular ducts present ventrally on margins.
<taxonomicNameid="4C644EA2232EFF87FEEB503DFE01FCD3"authorityName="Zheng & Wu"authorityYear="2025"box="[288,436,835,860]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="11"pageNumber="130"phylum="Arthropoda"rank="species"species="sinensis">
given in parentheses): (i) multilocular disc-pores of 2 sizes, each pore with 5, 7 or 9 subcentral loculi only (with multilocular disc-pores of only 1 size, each pores with 8 subcentral loculi); and (ii) a few slender oral collar ducts present anterior to clypeolabral shield (with large oral collar ducts anterior to clypeolabral shield) (data of
<bibRefCitationid="EFF548D0232EFF87FF1F50ADFE2DFC62"author="Williams, D. J."box="[212,408,979,1005]"pageId="11"pageNumber="130"refId="ref20067"refString="Williams, D. J. (2004) Mealybugs of Southern Asia. The Natural History Museum, London. Southdene SDN. BHD, Kuala Lumpur, 896 pp."type="book"year="2004">Williams (2004))</bibRefCitation>
<taxonomicNameid="4C644EA2232EFF87FE6E5089FDACFB9F"authorityName="Zheng & Wu"authorityYear="2025"box="[421,537,1015,1040]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="11"pageNumber="130"phylum="Arthropoda"rank="species"species="sinensis">
<bibRefCitationid="EFF548D0232EFF87FC595089FB9DFB9E"author="Danzig, E. M."box="[914,1064,1015,1041]"pageId="11"pageNumber="130"pagination="124 - 132"refId="ref19179"refString="Danzig, E. M. (1978) New species of mealybugs (Homoptera, Coccoidea) from Siberia and the Far East. In " New Species of Animals ". Trudy Akademii Nauk SSR Zoologicheskogo Instituta. St. Petersburg, 61, 124 - 132."type="journal article"year="1978">
in having the following character states: (i) a single circulus situated on venter of abdominal segment III; (ii) anal ring bearing sharply-tipped setae; (iii) oral colar tubular ducts absent from cerarii; (iv) dorsum without additional dorsal cerarii; (v) wide oral collar tubular ducts present on dorsum of head and thorax; (vi) multilocular disc-pores of 2 sizes; and (vii) a few slender oral collar ducts present anterior to clypeolabral shield, these sometimes deformed.
<taxonomicNameid="4C644EA2232EFF87FB9C57F9FB5BFB2F"authorityName="Zheng & Wu"authorityYear="2025"box="[1111,1262,1159,1184]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="11"pageNumber="130"phylum="Arthropoda"rank="species"species="sinensis">
are given in parentheses): (i) multilocular disc-pores each with 5, 7 or 9 subcentral loculi (pores each with 9 or 10 subcentral loculi, quinquelocular pores absent); and (ii) wide oral collar tubular ducts present ventrally on margins throughout (wide tubular ducts absent from margins of head and thorax) (data of
<bibRefCitationid="EFF548D0232EFF87FCD15669FB2BFABE"author="Danzig, E. M. & Gavrilov-Zimin, I. A."box="[794,1182,1303,1329]"pageId="11"pageNumber="130"refId="ref19265"refString="Danzig, E. M. & Gavrilov-Zimin, I. A. (2014) Palaearctic mealybugs (Homoptera: Coccinea: Pseudococcidae), Part 1: Subfamily Phenacoccinae. Russian Academy of Sciences, Zoological Institute, St. Petersburg, 678 pp."type="book"year="2014">Danzig & Gavrilov-Zimin (2014))</bibRefCitation>
<emphasisid="B910E933232EFF87FF5C56B4FEC9FA6B"bold="true"box="[151,380,1482,1508]"pageId="11"pageNumber="130">Appearance in life</emphasis>
(
<figureCitationid="135F29A4232EFF87FE5B56B5FDF3FA6A"box="[400,582,1483,1509]"captionStart="FIGURE 3"captionStartId="6.[151,250,1196,1220]"captionTargetBox="[151,1435,345,1166]"captionTargetId="figure-97@6.[151,1435,342,1170]"captionTargetPageId="6"captionText="FIGURE 3. Adult male of Puto huangshanensis Wu, 2001. A: loculate pores; B: small simple pore; C: long hair-like seta with satellite setae; D: hair-like seta; E: sensory seta; F: apex of antennal segment X; G: antennal intersegmental sensillum; H: prosternum; I: tarsus and claw; J: claw; K: glandular pouch; L: penial sheath; M: penial sheath seta; N: spine-like setae on sclerotized framework; O: penial sheath sensillum."figureDoi="http://doi.org/10.5281/zenodo.14734640"httpUri="https://zenodo.org/record/14734640/files/figure.png"pageId="11"pageNumber="130">Figs 3B and D</figureCitation>
). Body midge-like; forewings smoky gray, with anterior sclerotized area black; mesothorax and head coated with white wax; a pair of long, stout caudal pencils of wax present at end of abdomen.
long. Ventral surface of ocular sclerites with sclerotization restricted to near eyes; ocular sclerites each with 7 simple eyes. Antennae slightly longer than body, each with long hair-like setae and satellite setae present. Body with many long hair-like setae (hs). Loculate pores (lp) (
<figureCitationid="135F29A4232EFF87FF5455DDFF43F932"box="[159,246,1698,1725]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="11"pageNumber="130">Fig. 5A</figureCitation>
) present, each 5–8 μm in diameter, with 4 or 5 loculi (rarely 6); small simple pores (smp) each about 2.5 μm in diameter, few, present near glandular pouches and posterior to each mesothoracic spiracle. Legs well developed, long hair-like setae, satellite setae and spine-like setae present but without bifurcated setae; tarsi 2 segmented; claws each with a pair of basal spurs, plantar denticle weak or absent, plus a weak denticle near apex; claw digitules with pointed apices. Wings without alar setae or sensoria; a pair of ostioles present between abdominal segments
<collectionCodeid="ED75ADE4232EFF87FA84544DFAD8F8C2"box="[1359,1389,1843,1869]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="11"pageNumber="130">VI</collectionCode>
and VII; and a pair of glandular pouches present on abdominal segment VIII. Aedeagus slender.
<taxonomicNameid="4C644EA22329FF80FE71577AFD6DFB95"authority="Zheng & Wu"authorityName="Zheng & Wu"authorityYear="2025"box="[442,728,1027,1051]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="12"pageNumber="131"phylum="Arthropoda"rank="species"species="sinensis"status="sp. nov.">
dorsal mid-cranial ridge (dmcr) extending full length of head, fusing with transverse postoccipital suture (pos) posteriorly. Sclerotization extending from eyes to postoccipital suture (pos); membrane on each side of sclerotized area around dmcr; with 9–11 dorsal head setae (dhs) present on each side of dmcr, lp absent. Dorsum of ocular sclerites (ocs) with 10–17 dhs, each 10–15 μm long.
se near mid-cranial ridge larger, 58–60 μm in diameter; lateral simple eyes smaller, 30–43 μm in diameter; simple eyes prominent. With a single ocellus (o) laterally on each side, about 25 μm in diameter, difficult to observe.
sclerotization of ocs restricted to near eyes, small sclerotized area near preoral ridges and postocular ridges. Venter of each ocs with 30–40 ventral head setae (vhs), some setae with basal sclerotization. Ventral arm of midcranial ridge (vmcr) extending to fuse with preoral ridges (pror) posteriorly; preocular ridge (procr) fusing with vmcr medially. Membranous area anterior to preocular ridge with 9–13 vhs and 9–13 lp. Postocular ridge (pocr) developed, extending from preoral ridges posteriorly along posterior margin of ocs ventrally. Preoral ridge (pror) sclerotized, extending from posterior end of vmcr posterolaterally, articulating with cervical sclerites (pepcv). Area posterior to pror membranous, without setae. Mouth (m) present. Hair-like setae present on head, each 35–60 μm long.
, ratio of total body length to antennal length 1: 1.0–1.1. Segment lengths (in µm): I, 120–135; II, 75–95; III, 390–460; IV, 425–500; V, 390–440;
<collectionCodeid="ED75ADE42329FF80FAB35588FA2DF89F"box="[1400,1432,1782,1808]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="12"pageNumber="131">VI</collectionCode>
, 413–505; VII, 370–430; VIII, 360–390; IX, 270–365; and X, 280–345. Scape (scp) (segment I) stout, 35–153 µm wide, with base sclerotized and apex membranous; bearing 2–5 short hair-like setae, each 35–58 μm long. Pedicel (pdc) (segment II) stout, 83–90µm wide, completely sclerotized, with 4–6 long hair-like setae and 1–3 short hair-like setae plus 1 campaniform sensillum (camp) dorsally. Each segment of flagellum parallel-sided, 28–65 μm wide, with many long hair-like setae (
<figureCitationid="135F29A42329FF80FD8954D7FD2FF84B"box="[578,666,1961,1988]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="12"pageNumber="131">Fig. 5B</figureCitation>
), each seta 290–465 μm long, mostly with 1 or 2 satellite setae (
<figureCitationid="135F29A42329FF80FAA454D7FF0EF867"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="12"pageNumber="131">Fig. 5B</figureCitation>
); also with 0–3 hair-like setae (
<figureCitationid="135F29A42329FF80FDD354B3FDDAF867"box="[536,623,1997,2024]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="12"pageNumber="131">Fig. 5C</figureCitation>
), each seta 100–133 μm long. Segments VIII and IX each with 1 sensory seta (
<figureCitationid="135F29A42328FF81FF1F53E9FE98FF3E"box="[212,301,150,177]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="13"pageNumber="132">Fig. 5D</figureCitation>
) near apex; and apical segment (segment X) with 2 sensory setae and 5 hair-like setae subapically. Antennal intersegmental sensilla (
<figureCitationid="135F29A42328FF81FDDF53C5FDDCFF5A"box="[532,617,186,213]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="13"pageNumber="132">Fig. 5E</figureCitation>
) present between segments III–IV, IV–V and VII–VIII, with 2 sensilla at each location.
: pronotal ridges (prnr) developed and uninterrupted, fused medially, extending ventrally and nearly touching proepisternum + cervical sclerite (pepcv). With 0–3 hair-like setae anteriorly on each side between prnr and postoccipital suture (pos). Pronotal sclerite (prn) developed, each with a group of 8–11 lateral pronotal setae (lpns) and about 11 lp. Post-tergite (pt) present, 115–125 μm wide, lying transversely, each with a group of 4–7 post-tergital setae (pts) and 6–8 lp. With 0–4 medial pronotal setae (mpns) anterior to prescutum.
: pepcv sclerotized and interrupted medially; anterior part articulating with preoral ridge (pror), posterior end with a triangular sclerotization, articulating with prnr; posterior part becoming broad near articulation with coxa. With a large group of 13–17 long propleural setae (pl
prescutum (prsc) sclerotized but not separated from scutum, prescutal ridge (pscr) and suture (pscs) absent or indistinct; without prescutal setae (prscs). Scutum (sct) 690–720 μm wide; with a small triangular membranous area anterior to scutellum, 80–115 μm long and 100–135 μm wide. Scutum with 2 longitudinal bands of 12–20 scutal setae (scts) on each side, also 1–3 setae along each lateral margin. Scutellum (scl) 295–350 µm wide and 165–175 µm long, nearly rectangular, with developed marginal ridges and 16–25 scutellar setae (scls).
) either absent or fused to tergites of abdominal segment I; with a short band of 6–9 metatergal setae (mts) and 1–5 lp on each side; and with 5–7 dorsospiracular setae (dss) with 5–13 lp.
) developed, extending ventrally 142–185 µm; an additional short sclerotization present close to each articulation of metacoxa, 65–125 μm long. Metepimeron (epm
wide; ratio of length to width 1: 0.4; ratio of total body length to wing length 1: 0.96; covered with microtrichia (
<figureCitationid="135F29A42328FF81FB11556DFA86F9A2"box="[1242,1331,1554,1581]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="13"pageNumber="132">Fig. 5G</figureCitation>
); without alar setae or sensoria. Costal thickening (CT) slightly sclerotized. Subcosta (Sc) and radius (R) present along CT from the wing base, almost reach the apex. Cubitus anterior (CuA) originates from Sc at about 1/5 along wing length and runs obliquely to posterior wing margin. Alar lobe fold (alf) sclerotized along proximal posterior margin, with a membranous pocket; anal fold (af) transparent. Hamulohalteres (h) (
<figureCitationid="135F29A42328FF81FC7655DDFBA3F932"box="[957,1046,1698,1725]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="13"pageNumber="132">Fig. 5H</figureCitation>
) with anterior margin sclerotized, 233–285 µm long and 60–65 µm wide; with 3 or 4 hamuli (ham) at tip, each 105–115 µm long; each hamulus with apex curved.
; coxa 240–250; trochanter + femur 710–810; tibia 955–1,100; tarsus 290–345, and claw 74–75. Ratio of length of metatibia to length of metatarsus 1: 0.30–0.32. Legs setose, each with many hair-like setae, each seta 150–290 µm long; coxa, trochanter, femur and tibia with satellite setae; each coxa and trochanter also with some short hair-like setae (30–40 μm long). Trochanters each with 2 basal spine-like setae (20–23 μm long), and with 3 or 4 (mostly 4) sensory pores on each surface. Tibia with some inner-side spine-like setae (
<figureCitationid="135F29A4232BFF82FB6953C5FB45FF5A"box="[1186,1264,186,213]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="14"pageNumber="133">Fig. 5I</figureCitation>
) on distal half, each seta 45–55 μm long. Tarsi each 2 segmented, with proximal segment triangular, without setae; segment II with a tarsal campaniform sensillum proximally, inner side with spine-like setae, each 25–40 μm long; outer side with hair-like setae, each 38–88 μm long; tarsal digitules (tdgt) with pointed apices, each 63–65 μm long. Claws (c) (
<figureCitationid="135F29A4232BFF82FAA45259FF04FEEA"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="14"pageNumber="133">Fig. 5J</figureCitation>
) each with a pair of basal spurs (each about 10 μm long), plantar denticle weak or absent; plus a weak denticle near apex; claw digitules (cdgt) conical, each 30–35 μm, shorter than claw.
. Segments I–VII: tergites (at) represented by pairs of small sclerotizations on anterior margins of all segments, smaller posteriorly, at on segments I–
<collectionCodeid="ED75ADE4232BFF82FD7C52C9FD61FE5E"box="[695,724,439,465]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="14"pageNumber="133">VI</collectionCode>
reticulated. Sternites (as) similar to at but on segments II–VII, reticulated on segments II–V. Caudal extensions (ce) absent. Each segment with dorsal abdominal setae (ads), dorsopleural setae (dps), ventral pleural setae (vps) and ventral abdominal setae (avs), accompanied by lp; with more lp on margins. A pair of distinct ostioles present laterally between segments
<collectionCodeid="ED75ADE4232BFF82FBF7515DFBEFFDB2"box="[1084,1114,547,573]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="14"pageNumber="133">VI</collectionCode>
and VII. Segment VIII with at strongly sclerotized and cambered across segment, and a group of ads posterior to at; also with a pair of large sclerotized as, a group of avs posterior to each as. Hair-like setae forming groups on margins. Glandular pouches (gp) (
<figureCitationid="135F29A4232BFF82FF1E51F1FE98FD26"box="[213,301,654,681]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="14"pageNumber="133">Fig. 5K</figureCitation>
) large and shallow, each with about 240 lp, each pore (
<figureCitationid="135F29A4232BFF82FC5C51F1FC5EFD26"box="[919,1003,654,681]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="14"pageNumber="133">Fig. 5L</figureCitation>
) with 4–7 loculi; each gp containing 2 rather thick glandular pouch setae (gls), each 275–290 µm long, plus 11–15 shorter setae, each 25–98 μm long; each pair of gls with a large group of duct-like pores around their bases. Abdominal spiracles not detected.
. Anal opening (an) present on membranous dorsum of penial sheath, 50–58 μm wide. Penial sheath (ps) (
<figureCitationid="135F29A4232BFF82FEE85061FE34FCB6"box="[291,385,798,825]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="14"pageNumber="133">Fig. 5M</figureCitation>
) extending posteriorly from segment VIII, with base broad and tapering to a point, 390–445 µm long and 220–320 μm wide basally; mostly membranous but with sclerotized framework. Venter of membrane with 7–9 hair-like penial sheath setae, each 33–55 μm long; dorsum with 6 or 7 setae on membrane; sclerotized framework with some hair-like setae anteriorly, becoming spine-like (
<figureCitationid="135F29A4232BFF82FC6B50F5FC43FC2A"box="[928,1014,906,933]"captionStart="FIGURE 5"captionStartId="12.[151,250,1027,1051]"captionTargetBox="[155,1431,182,1000]"captionTargetId="figure-23@12.[151,1435,179,1001]"captionTargetPageId="12"captionText="FIGURE 5. Adult male of Puto sinensis Zheng & Wu, sp. n. A: loculate pores; B: long hair-like seta with satellite setae; C: hair-like seta; D: sensory seta; E: antennal intersegmental sensillum; F: fore wing; G: microtrichia; H: hamulohaltere; I: spine-like seta; J: claw; K: glandular pouch; L: loculate pores near glandular pouch; M: penial sheath; N: spine-like seta on sclerotized framework."figureDoi="http://doi.org/10.5281/zenodo.14734644"httpUri="https://zenodo.org/record/14734644/files/figure.png"pageId="14"pageNumber="133">Fig. 5N</figureCitation>
) and shorter posteriorly; with a group of sensoria (psp) near tip. Basal rod (bra) strongly sclerotized, 200–255 μm long, attached ventrally to anterior margin of penial sheath. Aedeagus (aed) sclerotized, 405–475 µm long, long and C-shaped, broadest at base (about 58 µm wide), tapering posteriorly (apex about 13 µm wide); attached to bra basally; apex not bilobate.
(i) fore-wings developed; (ii) claw with a weak denticle near apex and a pair of basal spurs, with or without a weak plantar denticle medially; (iii) tarsal digitules with pointed apices; (iv) with 9–13 loculate pores on each side of membranous area anterior to ocular sclerite; (v) with seven eyes on each side of head; (vi) antennal segment II with elongate setae and satellite setae; (vii) glandular pouches each containing about 240 loculate pores, pores each with 4–7 loculi, mostly with 5 or 6 loculi; (viii) both sides of abdomen with loculate pores, these fewer on venter; (ix) long hair-like setae on antenna each 290–465 μm long; (x) tegula with about 3 tegular setae; (xi) apex of aedeagus not bilobate; (xii) sclerotization of ocular sclerites restricted to near eyes; and (xiii) small simple pores very few, present near glandular pouches and posterior to each mesothoracic spiracle.
<taxonomicNameid="4C644EA2232BFF82FDC75645FD35FADB"authorityName="Zheng & Wu"authorityYear="2025"box="[524,640,1339,1364]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="14"pageNumber="133"phylum="Arthropoda"rank="species"species="sinensis">
are given in parentheses): (i) sclerotization of ocular sclerites restricted to near eyes (sclerotization extending to postocular ridge); (ii) claw without or with a weak plantar denticle medially (with a strong plantar denticle medially); (iii) glandular pouches each containing about 240 loculate pores (with about 100 loculate pores); (iv) sixth antennal segment longest (fifth antennal segment longest); and (v) a few small simple pores present near glandular pouches and posterior to each mesothoracic spiracle (present throughout whole body).
<taxonomicNameid="4C644EA2232BFF82FE5F556DFDBDF9A3"authorityName="Zheng & Wu"authorityYear="2025"box="[404,520,1555,1580]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="14"pageNumber="133"phylum="Arthropoda"rank="species"species="sinensis">
<bibRefCitationid="EFF548D0232BFF82FCA6556DFBAFF9A2"author="Coleman, G. A."box="[877,1050,1555,1581]"pageId="14"pageNumber="133"pagination="197 - 198"refId="ref19033"refString="Coleman, G. A. (1908) Coccidae of the Coniferae. Supplement No. 1. Description of two new species. Journal of the New York Entomological Society, 16, 197 - 198."type="journal article"year="1908">Coleman, 1908</bibRefCitation>
)
</taxonomicName>
in having the following character states: (i) aedeagus blunt apically; (ii) each side of head with seven eyes plus one ocellus; (iii) antennal segment III more than 350 µm long; and (iv) antennal segment IV or
<collectionCodeid="ED75ADE4232BFF82FCD15525FC8DF9FA"box="[794,824,1627,1653]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="14"pageNumber="133">VI</collectionCode>
longest.
<taxonomicNameid="4C644EA2232BFF82FC555525FB86F9FB"authorityName="Zheng & Wu"authorityYear="2025"box="[926,1075,1627,1652]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="14"pageNumber="133"phylum="Arthropoda"rank="species"species="sinensis">
using morphological characters, because the descriptions of the latter are all too simple (
<bibRefCitationid="EFF548D0232BFF82FB255501FA22F916"author="Coleman, G. A."box="[1262,1431,1663,1689]"pageId="14"pageNumber="133"pagination="409 - 420"refId="ref18989"refString="Coleman, G. A. (1901) The redwood mealy bug (Dactylopius sequoiae, sp. nov.). Proceedings of the California Academy of Sciences (ser. 3, Zoology), 2, 409 - 420."type="journal article"year="1901">Coleman 1901</bibRefCitation>
,
<bibRefCitationid="EFF548D0232BFF82FF5C55DDFF67F932"author="Coleman, G. A."box="[151,210,1699,1725]"pageId="14"pageNumber="133"pagination="197 - 198"refId="ref19033"refString="Coleman, G. A. (1908) Coccidae of the Coniferae. Supplement No. 1. Description of two new species. Journal of the New York Entomological Society, 16, 197 - 198."type="journal article"year="1908">1908</bibRefCitation>
,
<bibRefCitationid="EFF548D0232BFF82FF1655DDFED4F932"author="Ferris, G. F."box="[221,353,1699,1725]"pageId="14"pageNumber="133"refId="ref19312"refString="Ferris, G. F. (1918) The California species of mealy bugs. Leland Stanford Junior University Publications, University Series. Palo Alto, 78 pp."type="book"year="1918">Ferris 1918</bibRefCitation>
). However, they can be distinguished by their distributions and host plant preferences:
<taxonomicNameid="4C644EA2232BFF82FAE255DDFA28F933"authorityName="Zheng & Wu"authorityYear="2025"box="[1321,1437,1699,1724]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="14"pageNumber="133"phylum="Arthropoda"rank="species"species="sinensis">
<documentid="D1FFA26074789C3AF724404ED43E02AA"ID-DOI="10.11646/zootaxa.5570.1.5"ID-ISSN="1175-5326"ID-Zenodo-Dep="14734630"ID-ZooBank="8EF2BC5F-5939-440C-A053-A8924F07A42C"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1737736316380"checkinUser="plazi"docAuthor="Zheng, Xinyi, Li, Yuang, Tan, Zhixiang & Wu, San’An"docDate="2025"docId="03CD8437232DFF84FF5C578DFC21F932"docLanguage="en"docName="zootaxa.5570.1.5.pdf"docOrigin="Zootaxa 5570 (1)"docSource="https://doi.org/10.11646/zootaxa.5570.1.5"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Puto sinensis Zheng & Wu 2025, sp. n."docType="treatment"docVersion="2"lastPageNumber="127"masterDocId="FFF4FC4F2325FF8CFFCB537EFFB5FF8F"masterDocTitle="A new species of the genus Puto Signoret (Hemiptera: Coccomorpha: Putoidae) from China and redescription of Puto huangshanensis Wu"masterLastPageNumber="137"masterPageNumber="119"pageNumber="127"updateTime="1737736742446"updateUser="ExternalLinkService"zenodo-license-document="CLOSED">
<mods:titleid="6B36BABDD495D14D11B7C6272E8B49AB">A new species of the genus Puto Signoret (Hemiptera: Coccomorpha: Putoidae) from China and redescription of Puto huangshanensis Wu</mods:title>
<mods:affiliationid="CCBE0AF08D5278240C088CD009B88EC7">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<mods:affiliationid="336146914B57EC8DEBFBDC064FF33236">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<mods:affiliationid="0ECC771B407F544DA06780270383747A">Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025, China</mods:affiliation>
<mods:affiliationid="8E7B51E7CE719D28E03FC2607EA8F7D8">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<taxonomicNameid="4C644EA2232DFF84FF5C578DFE50FA82"authority="Zheng & Wu"authorityName="Zheng & Wu"authorityYear="2025"box="[151,485,1266,1293]"class="Insecta"family="Pseudococcidae"genus="Puto"kingdom="Animalia"order="Hemiptera"pageId="8"pageNumber="127"phylum="Arthropoda"rank="species"species="sinensis"status="sp. nov.">
