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<document id="B5FB4BCF89E6C0ECB91CCC52AE26818C" ID-CLB-Dataset="101995" ID-DOI="10.11646/zootaxa.5168.2.8" ID-GBIF-Dataset="8d8035eb-1bc7-4ff3-97d9-ca9909da9298" ID-ISSN="1175-5326" ID-Zenodo-Dep="6877349" ID-ZooBank="62868315-AD98-4FF7-AA3B-EBADC9F04C6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658421154441" checkinUser="plazi" docAuthor="Shrestha, Biraj, Pandey, Binita, Joshi, Jyoti, Manandhar, Prajwol, Karmacharya, Dibesh, Ohler, Annemarie, Dubois, Alain &amp; OConnell, Kyle A." docDate="2022" docId="92169E001566602AFF3773772DF372A1" docLanguage="en" docName="zootaxa.5168.2.8.pdf" docOrigin="Zootaxa 5168 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paa ercepeae" docType="treatment" docVersion="7" lastPageNumber="230" masterDocId="6E2FE67815616022FFA0712C2960714C" masterDocTitle="Molecular and morphological identification of frog species collected at Rara Lake in Rara National Park, Nepal" masterLastPageNumber="236" masterPageNumber="222" pageNumber="229" updateTime="1699361566052" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F7C8054EC885E815C4EAEF8A02663B8C">Molecular and morphological identification of frog species collected at Rara Lake in Rara National Park, Nepal</mods:title>
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<mods:namePart id="7D525C0EAD9365F0084D337561AC909D">Shrestha, Biraj</mods:namePart>
<mods:affiliation id="54C364A875E9EDC955561C87ED025DED">Division of Physical and Biological Sciences, Coastal Science and Policy, University of California, Santa Cruz, California, USA Department of Environmental Science, Khwopa College, Dekocha, Bhaktapur, Nepal</mods:affiliation>
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<mods:namePart id="CBCAB82A4EB0E5BB6C0A76D7750335F7">Pandey, Binita</mods:namePart>
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<mods:affiliation id="3FEEDE324D9281C330E30F198E23FE59">binita. p 862 @ gmail. com; https: // orcid. org / 0000 - 0002 - 9810 - 1357</mods:affiliation>
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<mods:namePart id="BE128487B1EA1989DD4A45523E14475C">Joshi, Jyoti</mods:namePart>
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<mods:affiliation id="307E770D391305C1AD7EE4E947FCA7F2">Department of Wildlife Genetics, Center for Molecular Dynamics Nepal, Kathmandu, Nepal &amp; jtwinj 1988 @ gmail. com; https: // orcid. org / 0000 - 0002 - 3968 - 517 X</mods:affiliation>
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<mods:namePart id="E8CAC7C96B63A364DEB2F2C945889D8C">Manandhar, Prajwol</mods:namePart>
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<mods:affiliation id="E92C695AAD20A1D5137198634A461F4B">Department of Wildlife Genetics, Center for Molecular Dynamics Nepal, Kathmandu, Nepal &amp; prajwol. f. manandhar @ gmail. com; https: // orcid. org / 0000 - 0002 - 5259 - 0706</mods:affiliation>
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<mods:affiliation id="3A9335AF818B14023D25562734FB5C85">Department of Wildlife Genetics, Center for Molecular Dynamics Nepal, Kathmandu, Nepal &amp; dibesh @ cmdn. org; https: // orcid. org / 0000 - 0001 - 5093 - 6737</mods:affiliation>
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<mods:namePart id="F61E62096B782168774C98AA72BB9CA6">Ohler, Annemarie</mods:namePart>
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<mods:affiliation id="968924610F91B4FA2231C1A042191C5C">Département Systematique &amp; Evolution, Museum national dHistoire naturelle, Paris, France &amp; annemarie. ohler @ mnhn. fr; https: // orcid. org / 0000 - 0001 - 6531 - 464 X</mods:affiliation>
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<mods:namePart id="B05DBE0A7ADB9CA036E5468D748F16C0">Dubois, Alain</mods:namePart>
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<mods:affiliation id="D8C82FD769A5B7705FD6BD6B800C3016">Département Systematique &amp; Evolution, Museum national dHistoire naturelle, Paris, France &amp; sapo 421 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6463 - 3435</mods:affiliation>
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<mods:namePart id="F2EB3607F432E3B352679F48E7539141">OConnell, Kyle A.</mods:namePart>
<mods:affiliation id="346270656E46E15975FF3988A362E236">Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA Department of Biological Sciences, The George Washington University, Washington, DC, USA</mods:affiliation>
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<mods:date id="B1EFA3E27B2DDD0F656ABBC9E3E2E18E">2022</mods:date>
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<treatment id="92169E001566602AFF3773772DF372A1" ID-DOI="http://doi.org/10.5281/zenodo.6877231" ID-GBIF-Taxon="197837415" ID-Zenodo-Dep="6877231" LSID="urn:lsid:plazi:treatment:92169E001566602AFF3773772DF372A1" httpUri="http://treatment.plazi.org/id/92169E001566602AFF3773772DF372A1" lastPageId="8" lastPageNumber="230" pageId="7" pageNumber="229">
<subSubSection id="52A57C9D15666025FF3773772A0273FB" pageId="7" pageNumber="229" type="nomenclature">
<paragraph id="1A002F1615666025FF3773772B797338" blockId="7.[151,537,602,629]" box="[151,537,602,629]" pageId="7" pageNumber="229">
<heading id="4148987A15666025FF3773772B797338" bold="true" box="[151,537,602,629]" fontSize="11" level="1" pageId="7" pageNumber="229" reason="1">
<taxonomicName id="DDBF549515666025FF3773772B797338" ID-CoL="4BP3Q" authority="(Dubois, 1974)" baseAuthorityName="Dubois" baseAuthorityYear="1974" box="[151,537,602,629]" class="Amphibia" family="Dicroglossidae" genus="Paa" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Anura" pageId="7" pageNumber="229" phylum="Chordata" rank="species" species="ercepeae">
<emphasis id="28CBF30415666025FF3773772B797338" bold="true" box="[151,537,602,629]" pageId="7" pageNumber="229">
<emphasis id="28CBF30415666025FF37737729A67339" bold="true" box="[151,198,603,629]" italics="true" pageId="7" pageNumber="229">Paa</emphasis>
cf.
<emphasis id="28CBF30415666025FF537377283D7339" bold="true" box="[243,349,603,629]" italics="true" pageId="7" pageNumber="229">ercepeae</emphasis>
(Dubois, 1974)
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<paragraph id="1A002F1615666025FF3773B12A0273FB" blockId="7.[151,866,669,695]" box="[151,866,669,695]" pageId="7" pageNumber="229">
Specimens examined: One male adult (NHM 17A-0120);
<figureCitation id="8284339315666025FC8173B12A0273FB" box="[801,866,669,695]" captionStart="FIGURE 4" captionStartId="7.[151,250,1929,1953]" captionTargetBox="[208,1380,719,1906]" captionTargetId="figure-226@7.[208,1380,719,1906]" captionTargetPageId="7" captionText="FIGURE 4. The adult male specimen of Paa cf. ercepeae (NHM 17A-0120). (AF) specimen in preservative; (A &amp; D) dorsal view; (C &amp; E) ventral view; (B) frontal view; (F) left foot." figureDoi="http://doi.org/10.5281/zenodo.6877357" httpUri="https://zenodo.org/record/6877357/files/figure.png" pageId="7" pageNumber="229">Fig. 4</figureCitation>
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<caption id="4EC07F9E15666025FF3776A52B877689" ID-DOI="http://doi.org/10.5281/zenodo.6877357" ID-Zenodo-Dep="6877357" httpUri="https://zenodo.org/record/6877357/files/figure.png" pageId="7" pageNumber="229" startId="7.[151,250,1929,1953]" targetBox="[208,1380,719,1906]" targetPageId="7">
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<emphasis id="28CBF30415666025FF3776A5287376ED" bold="true" box="[151,275,1929,1953]" pageId="7" pageNumber="229">FIGURE 4.</emphasis>
The adult male specimen of
<taxonomicName id="DDBF549515666025FD9C76A62B8676ED" baseAuthorityName="Dubois" baseAuthorityYear="1974" box="[572,742,1929,1953]" class="Amphibia" family="Dicroglossidae" genus="Paa" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Anura" pageId="7" pageNumber="229" phylum="Chordata" rank="species" species="ercepeae">
<emphasis id="28CBF30415666025FD9C76A62B0576ED" box="[572,613,1930,1953]" italics="true" pageId="7" pageNumber="229">Paa</emphasis>
cf.
<emphasis id="28CBF30415666025FD2C76A62B8676ED" box="[652,742,1930,1953]" italics="true" pageId="7" pageNumber="229">ercepeae</emphasis>
</taxonomicName>
(NHM 17A-0120). (AF) specimen in preservative; (A &amp; D) dorsal view; (C &amp; E) ventral view; (B) frontal view; (F) left foot.
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Measurements: SVL =
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, and T5 =
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.
</paragraph>
<paragraph id="1A002F161569602AFF67700B2C297355" blockId="8.[151,1437,150,1005]" pageId="8" pageNumber="230">
Diagnostic features (measured in mm): Our field-collected specimen of
<taxonomicName id="DDBF54951569602AFC50700B2DCC700C" baseAuthorityName="Dubois" baseAuthorityYear="1974" box="[1008,1196,295,321]" class="Amphibia" family="Dicroglossidae" genus="Paa" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Anura" pageId="8" pageNumber="230" phylum="Chordata" rank="species" species="ercepeae">
<emphasis id="28CBF3041569602AFC50700B2D7D700C" box="[1008,1053,295,320]" italics="true" pageId="8" pageNumber="230">Paa</emphasis>
cf.
<emphasis id="28CBF3041569602AFBE8700B2DCC700C" box="[1096,1196,295,320]" italics="true" pageId="8" pageNumber="230">ercepeae</emphasis>
</taxonomicName>
is a robust adult frog (SVL 75.2) devoid of spines in the hand region, particularly the first and second fingers and in the pectoral region. The snout is rounded, and the head is broader than long (HW 30.3, HL 22.2). The canthus is weakly developed (blunt) and rounded, fully-fledged supra-tympanic fold but the tympanum is indistinct. Nostril is closer to the eye than to the snout (EN 3.6, NS 5.2). The loreal region is slightly concave and the nostril is dorsolaterally positioned. Eyes are large (EL 8.5, 28% of HW) with patchy iris and rhombus-shaped pupils. The interorbital distance is larger than the upper eyelid width (IOD 7.5, UEW 5.7). Vomerine teeth and one set of maxillary teeth are present.
</paragraph>
<paragraph id="1A002F161569602AFF67730F29857259" blockId="8.[151,1437,150,1005]" pageId="8" pageNumber="230">Finger/toe tips are rounded into small discs. No webbing in fingers but toes are fully webbed. The relative length of fingers follows the pattern I&lt;II&lt;IV&lt;III. Inner metacarpal tubercles and subarticular tubercles in fingers are distinct. Hindlimbs are long and sturdy but thighs are shorter than the tibia (FL 38.1, TL 41.3). The relative length of toes follows the style I&lt;II&lt;V&lt;III&lt;IV. The inner metatarsal tubercle is large, while the outer metatarsal tubercle is indistinct. Dorsal skin is entirely smooth, including the head and lateral parts of the body, and dorsolateral folds absent. The cloacal aperture is quite prominent. The lower legs region has occasionally placed minute keratinized spines.</paragraph>
<paragraph id="1A002F161569602AFF6772332DF372A1" blockId="8.[151,1437,150,1005]" pageId="8" pageNumber="230">Dorsum color is golden yellow interspersed with orchid pink to dark vermiculations at the flank region, upper and lower jaws, nares, forelimbs, and hindlimbs. A black line runs through the supratympanic fold reaching the posterior end of the jaws. Ventral sections of the head, body, and limbs are pinkish white. Black divergent streaks radiate outwards of the black pupil bordered by the golden iris and the translucent nictitating membrane is colored light pink. In ethanol, the dorsal skin color is faded to yellowish-grey, limbs are light gray, while all spots, lines, and vermiculations darkened. Nictitating membrane turned sky blue, and pupils faded to white.</paragraph>
</subSubSection>
</treatment>
</document>