273 lines
24 KiB
XML
273 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-PMC="PMC3689139" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-PubMed="23794925" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="F5105916EA72DAD74E3BD79B3FF802C6" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Pterodactylus daviesii Owen 1874" docType="treatment" docVersion="6" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="36" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="35" updateTime="1668155938761" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rodrigues, Taissa</mods:namePart>
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<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
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<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-06-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>308</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>112</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
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<mods:identifier type="ZooBank">EEC31850AAAB4081B05AB80A2D944658</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFA2FFFEFFE74632FFB7FFC7D553CE26</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152044977" LSID="urn:lsid:plazi:treatment:F5105916EA72DAD74E3BD79B3FF802C6" httpUri="http://treatment.plazi.org/id/F5105916EA72DAD74E3BD79B3FF802C6" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
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<subSubSection pageId="34" pageNumber="35" type="nomenclature">
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<paragraph pageId="34" pageNumber="35">
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<taxonomicName LSID="F5105916-EA72-DAD7-4E3B-D79B3FF802C6" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pterodactylus daviesii" order="Pterosauria" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="daviesii">
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<pageBreakToken pageId="34" pageNumber="35" start="start">'</pageBreakToken>
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Pterodactylus ' daviesii Owen, 1874
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</taxonomicName>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A-D ' Pterodactylus' daviesii, holotype NHMUK PV 43074 (Albian, Gault Clay Formation), anterior part of the mandibular symphysis. A right lateral view B respective line drawing C dorsal view D respective line drawing. E-H ' Ornithocheirus' denticulatus, holotype? CAMSM B 54794 (Albian, Cambridge Greensand), anterior part of the rostrum E ventral view F respective line drawing G right lateral view H respective line drawing. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge, sul - sulcus. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. E and G from Seeley 1870. Photos A and C courtesy of The Natural History Museum." httpUri="https://binary.pensoft.net/fig/22081" pageId="34" pageNumber="35">Fig. 13A-D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="reference_group">
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<paragraph pageId="34" pageNumber="35">
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="daviesii">Pterodactylus daviesii</taxonomicName>
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Owen:
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<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B67" refString="Owen, R, 1874. Monograph on the fossil Reptilia of the Mesozoic Formations. Palaeontographical Society, London, 14 pp." title="Monograph on the fossil Reptilia of the Mesozoic Formations." year="1874">Owen 1874</bibRefCitation>
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: p. 2, pl. I, fig. 5-6
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</paragraph>
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<paragraph pageId="34" pageNumber="35">
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<taxonomicName class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="daviesii">Lonchodectes daviesii</taxonomicName>
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(Owen):
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
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: p. 538
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</paragraph>
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<paragraph pageId="34" pageNumber="35">
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<taxonomicName baseAuthorityName="Owen" baseAuthorityYear="1874" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="daviesii">Ornithocheirus daviesii</taxonomicName>
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(Owen):
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<bibRefCitation author="Arthaber, G" journalOrPublisher="Palaeontologische Zeitschrift" pageId="64" pageNumber="65" pagination="1 - 47" refId="B2" refString="Arthaber, G, 1922. Ueber Entwicklung, Ausbildung und Austerben der Flugsaurier. Palaeontologische Zeitschrift 4: 1 - 47" title="Ueber Entwicklung, Ausbildung und Austerben der Flugsaurier." volume="4" year="1922">Arthaber 1922</bibRefCitation>
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: p. 16
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</paragraph>
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<paragraph pageId="34" pageNumber="35">
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<taxonomicName authority="daviesi [sic]" class="Reptilia" family="Ornithocheiridae" genus="Ornithochirus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="34" pageNumber="35" phylum="Chordata" rank="variety" species="sic" subSpecies="daviesi" variety="sic">Ornithochirus [sic] daviesi [sic]</taxonomicName>
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(Owen):
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<bibRefCitation author="Lydekker, R" journalOrPublisher="W. Part I. Trustees of the British Museum, London" pageId="67" pageNumber="68" refId="B48" refString="Lydekker, R, 1888. Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History), Cromwell Road, S. W. Part I. Trustees of the British Museum, London, 309 pp." title="Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History), Cromwell Road, S." year="1888">Lydekker 1888</bibRefCitation>
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: p. 23
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</paragraph>
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<paragraph pageId="34" pageNumber="35">
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<taxonomicName authority="" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="34" pageNumber="35" phylum="Chordata" rank="subSpecies" species="daviesi" subSpecies="sic">Ornithocheirus daviesi [sic]</taxonomicName>
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(Owen):
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<bibRefCitation author="Wellnhofer, P" journalOrPublisher="Gustav Fischer Verlag, Stuttgart and New York" pageId="70" pageNumber="71" refId="B101" refString="Wellnhofer, P, 1978. Pterosauria. Handbuch der Palaeoherpetologie, Teil 19. Gustav Fischer Verlag, Stuttgart and New York, 82 pp." title="Pterosauria. Handbuch der Palaeoherpetologie, Teil 19." year="1978">Wellnhofer 1978</bibRefCitation>
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: p. 56, fig. 28
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</paragraph>
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<paragraph pageId="34" pageNumber="35">
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<taxonomicName baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="platystomus">Lonchodectes platystomus</taxonomicName>
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(Seeley):
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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: fig. 12C
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="holotype">
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<paragraph pageId="34" pageNumber="35">Holotype.</paragraph>
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<paragraph pageId="34" pageNumber="35">
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NHMUK PV 43074, partial mandibular symphysis (
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A-D ' Pterodactylus' daviesii, holotype NHMUK PV 43074 (Albian, Gault Clay Formation), anterior part of the mandibular symphysis. A right lateral view B respective line drawing C dorsal view D respective line drawing. E-H ' Ornithocheirus' denticulatus, holotype? CAMSM B 54794 (Albian, Cambridge Greensand), anterior part of the rostrum E ventral view F respective line drawing G right lateral view H respective line drawing. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge, sul - sulcus. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. E and G from Seeley 1870. Photos A and C courtesy of The Natural History Museum." httpUri="https://binary.pensoft.net/fig/22081" pageId="34" pageNumber="35">Fig. 13A-D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="type locality">
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<paragraph pageId="34" pageNumber="35">Type locality.</paragraph>
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<paragraph pageId="34" pageNumber="35">Folkestone, Kent, England.</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="type horizon">
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<paragraph pageId="34" pageNumber="35">Type horizon.</paragraph>
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<paragraph pageId="34" pageNumber="35">Gault Clay Formation (Albian).</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="diagnosis">
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<paragraph pageId="34" pageNumber="35">Diagnosis.</paragraph>
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<paragraph pageId="34" pageNumber="35">
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Pterodactyloid pterosaur with the following combination of characters that distinguishes it from other members of the clade (autapomorphies are marked with an asterisk): anterior expansion absent; mandibular crest absent; mandibular groove about 2.5 cm wide; mandibular groove with elevated margins; mandibular groove extends until the second pair of alveoli; alveoli of the anterior portion of the mandible without significant variation in size; alveoli equally spaced; first pair of teeth face anterodorsally; transverse section
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<normalizedToken originalValue="‘V-shaped’">'V-shaped'</normalizedToken>
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; approximately 4 alveoli per 3 cm of jaw margin*.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="description">
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<paragraph pageId="34" pageNumber="35">Description.</paragraph>
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<paragraph pageId="34" pageNumber="35">
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<taxonomicName lsidName="" pageId="34" pageNumber="35" rank="species" species="daviesii">
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<emphasis italics="true" pageId="34" pageNumber="35">
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<normalizedToken originalValue="‘Pterodactylus’">'Pterodactylus'</normalizedToken>
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daviesii
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</emphasis>
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</taxonomicName>
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is known from a partial mandibular symphysis from the Gault Clay Formation. In his description,
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<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B67" refString="Owen, R, 1874. Monograph on the fossil Reptilia of the Mesozoic Formations. Palaeontographical Society, London, 14 pp." title="Monograph on the fossil Reptilia of the Mesozoic Formations." year="1874">Owen (1874)</bibRefCitation>
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stressed the presence of a mandibular sulcus and that the first pair of alveoli is less elliptical than the remaining ones and directed forwards and upwards.
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<taxonomicName lsidName="" pageId="34" pageNumber="35" rank="species" species="daviesii">
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<emphasis italics="true" pageId="34" pageNumber="35">
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<normalizedToken originalValue="‘Pterodactylus’">'Pterodactylus'</normalizedToken>
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daviesii
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</emphasis>
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</taxonomicName>
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presents a unique mosaic of seemingly plesiomorphic features among pterodactyloids. It lacks an anterior expansion, a crest, or any other distinctive dental feature, which would permit referral to a known genus. Instead, it is the combination of these characters that makes this species distinct from all known pterodactyloid genera.
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</paragraph>
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<paragraph pageId="34" pageNumber="35">
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It shares with
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Lonchodraconidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="34" pageNumber="35" phylum="Chordata" rank="family">Lonchodraconidae</taxonomicName>
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the presence of alveoli of the anterior portion of the mandible without significant variation in size, but can be confidently excluded from this clade because its alveoli are slightly larger and are not located on elevated alveolar margins. Although the dentary sulcus is relatively wide, it is narrower than in
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="machaerorhynchus">
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<emphasis italics="true" pageId="34" pageNumber="35">Lonchodraco machaerorhynchus</emphasis>
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</taxonomicName>
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.
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<taxonomicName lsidName="" pageId="34" pageNumber="35" rank="species" species="daviesii">
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<emphasis italics="true" pageId="34" pageNumber="35">
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<normalizedToken originalValue="‘Pterodactylus’">'Pterodactylus'</normalizedToken>
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daviesii
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</emphasis>
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</taxonomicName>
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is also distinct from
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="giganteus">
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<emphasis italics="true" pageId="34" pageNumber="35">Lonchodraco giganteus</emphasis>
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</taxonomicName>
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. In the latter, the mandible is very wide, with divergent margins, and rounded anteriorly. Although not comparable to
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<taxonomicName baseAuthorityName="Owen" baseAuthorityYear="1861" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="simus">
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<emphasis italics="true" pageId="34" pageNumber="35">Ornithocheirus simus</emphasis>
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</taxonomicName>
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, it is unlikely that it represents this much larger pterosaur.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="remarks">
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<paragraph pageId="34" pageNumber="35">Remarks.</paragraph>
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<paragraph lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin (2001)</bibRefCitation>
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did not cite
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<taxonomicName lsidName="" pageId="35" pageNumber="36" rank="species" species="daviesii">
|
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<emphasis italics="true" pageId="35" pageNumber="36">
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<pageBreakToken pageId="35" pageNumber="36" start="start">'</pageBreakToken>
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<normalizedToken originalValue="Pterodactylus’">Pterodactylus'</normalizedToken>
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daviesii
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</emphasis>
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</taxonomicName>
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in the text of his review, but illustrated the holotype, referring to it as
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<taxonomicName authorityName="Hooley" authorityYear="1914" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="35" pageNumber="36" phylum="Chordata" rank="genus">
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||
<emphasis italics="true" pageId="35" pageNumber="36">Lonchodectes</emphasis>
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||
</taxonomicName>
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||
[=
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<taxonomicName genus="Ornithocheirus'" lsidName="" pageId="35" pageNumber="36" rank="genus">
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||
<emphasis italics="true" pageId="35" pageNumber="36">
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||
<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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</emphasis>
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</taxonomicName>
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]
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<emphasis italics="true" pageId="35" pageNumber="36">platystomus</emphasis>
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, and indicated that it bears a crest. We disagree that NHMUK PV 43074 represents a crested pterosaur; the ventral margin of the dentary is sharp but there is no evidence for a crest. Furthermore,
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<taxonomicName lsidName="" pageId="35" pageNumber="36" rank="species" species="daviesii">
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<emphasis italics="true" pageId="35" pageNumber="36">
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||
<normalizedToken originalValue="‘Pterodactylus’">'Pterodactylus'</normalizedToken>
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daviesii
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</emphasis>
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</taxonomicName>
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is not directly comparable to
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<taxonomicName lsidName="" pageId="35" pageNumber="36" rank="species" species="platystomus">
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<emphasis italics="true" pageId="35" pageNumber="36">
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<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
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platystomus
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</emphasis>
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</taxonomicName>
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because the latter is known only from an upper jaw.
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</paragraph>
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<paragraph pageId="35" pageNumber="36">
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<taxonomicName lsidName="" pageId="35" pageNumber="36" rank="species" species="daviesii">
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<emphasis italics="true" pageId="35" pageNumber="36">
|
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<normalizedToken originalValue="‘Pterodactylus’">'Pterodactylus'</normalizedToken>
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daviesii
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</emphasis>
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</taxonomicName>
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thus possibly represents a distinct taxon from the Gault Clay Formation. However, we refrain from naming it until more complete material becomes available and refer it using its original designation in single quotation marks.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/22081" pageId="35" pageNumber="36" start="Figure 13" startId="F13">
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<paragraph pageId="35" pageNumber="36">
|
||
<emphasis bold="true" pageId="35" pageNumber="36">Figure 13.</emphasis>
|
||
<emphasis bold="true" pageId="35" pageNumber="36">A-D</emphasis>
|
||
<taxonomicName lsidName="" pageId="35" pageNumber="36" rank="species" species="daviesii">
|
||
<emphasis italics="true" pageId="35" pageNumber="36">
|
||
<normalizedToken originalValue="‘Pterodactylus’">'Pterodactylus'</normalizedToken>
|
||
daviesii
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, holotype NHMUK PV 43074 (Albian, Gault Clay Formation), anterior part of the mandibular symphysis.
|
||
<emphasis bold="true" pageId="35" pageNumber="36">A</emphasis>
|
||
right lateral view
|
||
<emphasis bold="true" pageId="35" pageNumber="36">B</emphasis>
|
||
respective line drawing
|
||
<emphasis bold="true" pageId="35" pageNumber="36">C</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="35" pageNumber="36">D</emphasis>
|
||
respective line drawing.
|
||
<emphasis bold="true" pageId="35" pageNumber="36">E-H</emphasis>
|
||
<taxonomicName lsidName="" pageId="35" pageNumber="36" rank="species" species="denticulatus">
|
||
<emphasis italics="true" pageId="35" pageNumber="36">
|
||
<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
|
||
denticulatus
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, holotype?CAMSM B 54794 (Albian, Cambridge Greensand), anterior part of the rostrum
|
||
<emphasis bold="true" pageId="35" pageNumber="36">E</emphasis>
|
||
ventral view
|
||
<emphasis bold="true" pageId="35" pageNumber="36">F</emphasis>
|
||
respective line drawing
|
||
<emphasis bold="true" pageId="35" pageNumber="36">G</emphasis>
|
||
right lateral view
|
||
<emphasis bold="true" pageId="35" pageNumber="36">H</emphasis>
|
||
respective line drawing. Abbreviations:
|
||
<emphasis bold="true" pageId="35" pageNumber="36">m</emphasis>
|
||
- maxillae,
|
||
<emphasis bold="true" pageId="35" pageNumber="36">pm</emphasis>
|
||
- premaxillae,
|
||
<emphasis bold="true" pageId="35" pageNumber="36">prid</emphasis>
|
||
- palatal ridge,
|
||
<emphasis bold="true" pageId="35" pageNumber="36">sul</emphasis>
|
||
- sulcus. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm.
|
||
<emphasis bold="true" pageId="35" pageNumber="36">E</emphasis>
|
||
and
|
||
<emphasis bold="true" pageId="35" pageNumber="36">G</emphasis>
|
||
from
|
||
<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
|
||
. Photos
|
||
<emphasis bold="true" pageId="35" pageNumber="36">A</emphasis>
|
||
and
|
||
<emphasis bold="true" pageId="35" pageNumber="36">C</emphasis>
|
||
courtesy of The Natural History Museum.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |