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<document id="C23CC9C7F0EA7BCE0A315CC411C24B0D" ID-DOI="10.11646/zootaxa.3694.4.2" ID-GBIF-Dataset="8ce29893-89ba-4b0e-8941-287814ff4a8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="220826" ID-ZooBank="03DDE60B-89C5-4B20-B14F-21C6CF69BF15" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460658596926" checkinUser="plazi" docAuthor="Yin, Zi-Wei, Coulon, Georges &amp; Li, Li-Zhen" docDate="2013" docId="03A2DE12975E713E34D9FDF672C0FCDB" docLanguage="en" docName="zt03694p342.pdf" docOrigin="Zootaxa 3694 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tmesiphodimerus malaysianus Coulon and Yin, new species" docType="treatment" docVersion="7" lastPageNumber="342" masterDocId="FF9BA66A975B7138344EFFD27679FFE3" masterDocTitle="Description of two new species of the genus Tmesiphodimerus gen. n. (Coleoptera: Staphylinidae: Pselaphinae) from South Asia" masterLastPageNumber="342" masterPageNumber="336" pageNumber="341" updateTime="1698347210820" updateUser="plazi">
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<mods:title id="2349AC9FFC3165E0F10ACF07BE2A5FC0">Description of two new species of the genus Tmesiphodimerus gen. n. (Coleoptera: Staphylinidae: Pselaphinae) from South Asia</mods:title>
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<mods:namePart id="4F7ED67E462AA3C79D550B6A090A94CE">Yin, Zi-Wei</mods:namePart>
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<mods:namePart id="718D6BA22981A0039C8938D47A749F00">Coulon, Georges</mods:namePart>
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<mods:namePart id="9E7F9A8D2FD6E57C1DB24A9EEF650A99">Li, Li-Zhen</mods:namePart>
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<mods:title id="FFCBDF50EE3DD4A5F9ACC890AA8A2D21">Zootaxa</mods:title>
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<mods:date id="FC037C1A60CA109EAA6DF30E8A7A847C">2013</mods:date>
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<treatment id="03A2DE12975E713E34D9FDF672C0FCDB" ID-DOI="http://doi.org/10.5281/zenodo.6154986" ID-GBIF-Taxon="119574232" ID-Zenodo-Dep="6154986" LSID="urn:lsid:plazi:treatment:03A2DE12975E713E34D9FDF672C0FCDB" httpUri="http://treatment.plazi.org/id/03A2DE12975E713E34D9FDF672C0FCDB" lastPageId="6" lastPageNumber="342" pageId="5" pageNumber="341">
<subSubSection id="C3113C8F975E713D34D9FDF67764FDBC" pageId="5" pageNumber="341" type="nomenclature">
<paragraph id="8BB46F04975E713D34D9FDF67500FDDD" blockId="5.[151,889,548,607]" box="[151,889,548,574]" pageId="5" pageNumber="341">
<heading id="D0FCD868975E713D34D9FDF67500FDDD" bold="true" box="[151,889,548,574]" fontSize="11" level="1" pageId="5" pageNumber="341" reason="1">
<emphasis id="B97FB316975E713D34D9FDF67500FDDD" bold="true" box="[151,889,548,574]" pageId="5" pageNumber="341">
<taxonomicName id="4C0B1487975E713D34D9FDF674A5FDDD" ID-CoL="579WM" authority="Coulon and Yin" authorityName="Coulon and Yin" box="[151,732,548,574]" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="341" phylum="Arthropoda" rank="species" species="malaysianus" status="sp. nov.">
<emphasis id="B97FB316975E713D34D9FDF67477FDDD" bold="true" box="[151,526,548,574]" italics="true" pageId="5" pageNumber="341">Tmesiphodimerus malaysianus</emphasis>
Coulon and Yin
</taxonomicName>
,
<taxonomicNameLabel id="A24C0E6D975E713D36A4FDF67500FDDD" box="[746,889,548,574]" pageId="5" pageNumber="341" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BB46F04975E713D34D9FD947764FDBC" blockId="5.[151,889,548,607]" box="[151,285,582,607]" pageId="5" pageNumber="341">
(
<figureCitation id="13307381975E713D34D1FD94769BFDBC" box="[159,226,582,607]" captionStart="FIGURE 1" captionStartId="1.[151,250,1941,1963]" captionTargetBox="[162,1429,771,1910]" captionTargetId="figure@1.[151,1436,762,1917]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Tmesiphodimerus sinensis (A) and T. malaysianus (B). Scales (mm): 0.5." httpUri="https://zenodo.org/record/220827/files/figure.png" pageId="5" pageNumber="341">Figs 1</figureCitation>
B, 4)
</paragraph>
</subSubSection>
<subSubSection id="C3113C8F975E713D34D9FD5C777EFD0C" pageId="5" pageNumber="341" type="materials_examined">
<paragraph id="8BB46F04975E713D34D9FD5C777EFD0C" blockId="5.[151,1436,654,751]" pageId="5" pageNumber="341">
<emphasis id="B97FB316975E713D34D9FD5C773BFD44" bold="true" box="[151,322,654,679]" pageId="5" pageNumber="341">
<typeStatus id="54B0D1A6975E713D34D9FD5C76ABFD44" box="[151,210,654,679]" pageId="5" pageNumber="341">Type</typeStatus>
material
</emphasis>
(
<specimenCount id="9D0DA48D975E713D3517FD5D77FEFD44" box="[345,391,655,679]" pageId="5" pageNumber="341" type="generic">1 ♂</specimenCount>
).
<emphasis id="B97FB316975E713D35ECFD5C74E3FD44" bold="true" box="[418,666,654,679]" pageId="5" pageNumber="341">
<typeStatus id="54B0D1A6975E713D35ECFD5C746AFD44" box="[418,531,654,679]" pageId="5" pageNumber="341" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F31C2F94975E713D366AFD5C74ECFD44" box="[548,661,654,679]" name="Malaysia" pageId="5" pageNumber="341">Malaysia</collectingCountry>
:
</emphasis>
♂, labeled West
<collectingCountry id="F31C2F94975E713D3738FD5C7598FD44" box="[886,993,654,679]" name="Malaysia" pageId="5" pageNumber="341">Malaysia</collectingCountry>
: Perak, / ca.
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N. Chenderiang, / 4°2146″N, 101°1509″E, / alt.
<quantity id="4CF3C2E1975E713D35BBFD61740EFD29" box="[501,631,691,715]" metricMagnitude="2" metricUnit="m" metricValue="3.65" metricValueMax="4.0" metricValueMin="3.3" pageId="5" pageNumber="341" unit="m" value="365.0" valueMax="400.0" valueMin="330.0">330400 m</quantity>
, lpoh, 1517, /
<date id="FFB549C4975E713D3769FD6075EAFD28" box="[807,915,690,715]" pageId="5" pageNumber="341" value="1995-01-22">22.i.1995</date>
, evergreen rain / forest, leg. P. Schwendinger (MHNG).
</paragraph>
</subSubSection>
<caption id="DF743F8C975E713D34D9F846727EF80C" httpUri="https://zenodo.org/record/220830/files/figure.png" pageId="5" pageNumber="341" targetBox="[160,1427,797,1925]" targetPageId="5">
<paragraph id="8BB46F04975E713D34D9F846727EF80C" blockId="5.[151,1436,1940,2031]" pageId="5" pageNumber="341">
<emphasis id="B97FB316975E713D34D9F846776BF84A" bold="true" box="[151,274,1940,1962]" pageId="5" pageNumber="341">FIGURE 4.</emphasis>
Diagnostic features of
<taxonomicName id="4C0B1487975E713D35B1F8467541F849" box="[511,824,1940,1962]" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="341" phylum="Arthropoda" rank="species" species="malaysianus">
<emphasis id="B97FB316975E713D35B1F8467541F849" box="[511,824,1940,1962]" italics="true" pageId="5" pageNumber="341">Tmesiphodimerus malaysianus</emphasis>
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. A. Antenna. B Subantennal sulcus. C. Maxillary palpus. D. Sternite IX. E. Aedeagus, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Parameres, in lateral view. I. Same, in ventral view. Scales (mm): A = 0.2; B, C, EG = 0.1 mm; D, H, I = 0.05 mm.
</paragraph>
</caption>
<subSubSection id="C3113C8F975D713E3489FF45728BFF14" pageId="6" pageNumber="342" type="diagnosis">
<paragraph id="8BB46F04975D713E3489FF45728BFF14" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="342">
<emphasis id="B97FB316975D713E3489FF457738FF53" bold="true" box="[199,321,151,176]" pageId="6" pageNumber="342">Diagnosis.</emphasis>
Reddish brown; length
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. Dorsal surfaces of head, pronotum, elytra and abdomen covered with dense, short setae. Apical four antennomeres enlarged, unmodified, antennomeres VIIIX of similar width. Maxillary palpomeres II only slightly projected apicolaterally. Aedeagus with triangular parameres.
</paragraph>
</subSubSection>
<subSubSection id="C3113C8F975D713E3489FED677FBFD2E" pageId="6" pageNumber="342" type="description">
<paragraph id="8BB46F04975D713E3489FED672EEFE33" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="342">
<emphasis id="B97FB316975D713E3489FED67720FEFE" bold="true" box="[199,345,260,285]" pageId="6" pageNumber="342">Description.</emphasis>
Male. Length
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. Head transverse, HL
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, HW
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; eyes each composed of more than 130 facets. Antennomeres VIIIXI (
<figureCitation id="13307381975D713E36E4FEF57497FEA3" box="[682,750,295,320]" captionStart="FIGURE 4" captionStartId="5.[151,250,1940,1962]" captionTargetBox="[160,1427,797,1925]" captionTargetId="figure@5.[151,1436,788,1929]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tmesiphodimerus malaysianus. A. Antenna. B Subantennal sulcus. C. Maxillary palpus. D. Sternite IX. E. Aedeagus, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Parameres, in lateral view. I. Same, in ventral view. Scales (mm): A = 0.2; B, C, E G = 0.1 mm; D, H, I = 0.05 mm." httpUri="https://zenodo.org/record/220830/files/figure.png" pageId="6" pageNumber="342">Fig. 4</figureCitation>
A) enlarged, simple, subcylindrical. Maxillary palpomeres II slightly projected apicolaterally (
<figureCitation id="13307381975D713E3652FE9E741FFE87" box="[540,614,332,357]" captionStart="FIGURE 4" captionStartId="5.[151,250,1940,1962]" captionTargetBox="[160,1427,797,1925]" captionTargetId="figure@5.[151,1436,788,1929]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tmesiphodimerus malaysianus. A. Antenna. B Subantennal sulcus. C. Maxillary palpus. D. Sternite IX. E. Aedeagus, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Parameres, in lateral view. I. Same, in ventral view. Scales (mm): A = 0.2; B, C, E G = 0.1 mm; D, H, I = 0.05 mm." httpUri="https://zenodo.org/record/220830/files/figure.png" pageId="6" pageNumber="342">Fig. 4</figureCitation>
C). Pronotum as long as wide, PL
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, PW
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, roundly expanded at lateral margins. Elytra wider than long, EL
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, EW
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. Abdomen broad at base and narrowed apically, AL
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, AW
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. Sternite IX as in
<figureCitation id="13307381975D713E3736FE4675C5FE4F" box="[888,956,404,429]" captionStart="FIGURE 4" captionStartId="5.[151,250,1940,1962]" captionTargetBox="[160,1427,797,1925]" captionTargetId="figure@5.[151,1436,788,1929]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tmesiphodimerus malaysianus. A. Antenna. B Subantennal sulcus. C. Maxillary palpus. D. Sternite IX. E. Aedeagus, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Parameres, in lateral view. I. Same, in ventral view. Scales (mm): A = 0.2; B, C, E G = 0.1 mm; D, H, I = 0.05 mm." httpUri="https://zenodo.org/record/220830/files/figure.png" pageId="6" pageNumber="342">Fig. 4</figureCitation>
D. Aedeagal length
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, median lobe (
<figureCitation id="13307381975D713E34D1FE65769CFE33" box="[159,229,439,464]" captionStart="FIGURE 4" captionStartId="5.[151,250,1940,1962]" captionTargetBox="[160,1427,797,1925]" captionTargetId="figure@5.[151,1436,788,1929]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tmesiphodimerus malaysianus. A. Antenna. B Subantennal sulcus. C. Maxillary palpus. D. Sternite IX. E. Aedeagus, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Parameres, in lateral view. I. Same, in ventral view. Scales (mm): A = 0.2; B, C, E G = 0.1 mm; D, H, I = 0.05 mm." httpUri="https://zenodo.org/record/220830/files/figure.png" pageId="6" pageNumber="342">Figs 4</figureCitation>
EG) nearly symmetric dorsoventrally, triangular parameres (
<figureCitation id="13307381975D713E37D9FE6575A4FE33" box="[919,989,439,464]" captionStart="FIGURE 4" captionStartId="5.[151,250,1940,1962]" captionTargetBox="[160,1427,797,1925]" captionTargetId="figure@5.[151,1436,788,1929]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Tmesiphodimerus malaysianus. A. Antenna. B Subantennal sulcus. C. Maxillary palpus. D. Sternite IX. E. Aedeagus, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Parameres, in lateral view. I. Same, in ventral view. Scales (mm): A = 0.2; B, C, E G = 0.1 mm; D, H, I = 0.05 mm." httpUri="https://zenodo.org/record/220830/files/figure.png" pageId="6" pageNumber="342">Figs 4</figureCitation>
HI) symmetric.
</paragraph>
<paragraph id="8BB46F04975D713E3489FE0E77E4FE16" blockId="6.[151,1437,151,824]" box="[199,413,476,501]" pageId="6" pageNumber="342">Female. Unknown.</paragraph>
<paragraph id="8BB46F04975D713E3489FE2D77FBFD2E" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="342">
<emphasis id="B97FB316975D713E3489FE2D77C8FDFB" bold="true" box="[199,433,511,536]" pageId="6" pageNumber="342">Comparative notes.</emphasis>
The new species can be separated from
<taxonomicName id="4C0B1487975D713E3735FDD37589FDFB" box="[891,1008,513,536]" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="342" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B97FB316975D713E3735FDD37589FDFB" box="[891,1008,513,536]" italics="true" pageId="6" pageNumber="342">T. sinensis</emphasis>
</taxonomicName>
by the smaller size (
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), dorsal surface of the body covered with short setae, the simple, subcylindrical antennomeres IX and XI, the less prominent apicolateral margins of maxillary palpomeres II, and the different aedeagal structures.
<taxonomicName id="4C0B1487975D713E309BFD957689FD67" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="342" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B97FB316975D713E309BFD957689FD67" italics="true" pageId="6" pageNumber="342">Tmesiphodimerus sinensis</emphasis>
</taxonomicName>
is greater in size (
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), the dorsal surface of the body is covered with longer hair, the antennomeres IX and XI each has a disc-shaped projection, and the maxillary palpomeres II is more prominent at apicolateral margins.
</paragraph>
</subSubSection>
<subSubSection id="C3113C8F975D713E3489FD05741AFD13" box="[199,611,727,752]" pageId="6" pageNumber="342" type="distribution">
<paragraph id="8BB46F04975D713E3489FD05741AFD13" blockId="6.[151,1437,151,824]" box="[199,611,727,752]" pageId="6" pageNumber="342">
<emphasis id="B97FB316975D713E3489FD057719FD13" bold="true" box="[199,352,727,752]" pageId="6" pageNumber="342">Distribution.</emphasis>
West
<collectingCountry id="F31C2F94975D713E35EBFD057469FD13" box="[421,528,727,752]" name="Malaysia" pageId="6" pageNumber="342">Malaysia</collectingCountry>
: Perak.
</paragraph>
</subSubSection>
<subSubSection id="C3113C8F975D713E3489FD2E75CCFCF6" box="[199,949,764,789]" pageId="6" pageNumber="342" type="biology_ecology">
<paragraph id="8BB46F04975D713E3489FD2E75CCFCF6" blockId="6.[151,1437,151,824]" box="[199,949,764,789]" pageId="6" pageNumber="342">
<emphasis id="B97FB316975D713E3489FD2E775EFCF6" bold="true" box="[199,295,764,789]" pageId="6" pageNumber="342">Biology.</emphasis>
The single adult was collected in an evergreen rain forest.
</paragraph>
</subSubSection>
<subSubSection id="C3113C8F975D713E3489FCCD72C0FCDB" box="[199,1209,799,824]" pageId="6" pageNumber="342" type="etymology">
<paragraph id="8BB46F04975D713E3489FCCD72C0FCDB" blockId="6.[151,1437,151,824]" box="[199,1209,799,824]" pageId="6" pageNumber="342">
<emphasis id="B97FB316975D713E3489FCCD7737FCDB" bold="true" box="[199,334,799,824]" pageId="6" pageNumber="342">Etymology.</emphasis>
The new species is named after the country where the
<typeStatus id="54B0D1A6975D713E37FAFCF2759DFCDB" box="[948,996,800,824]" pageId="6" pageNumber="342">type</typeStatus>
locality is located.
</paragraph>
</subSubSection>
</treatment>
</document>