402 lines
56 KiB
XML
402 lines
56 KiB
XML
<document id="0A43A0456F8760E4046E97118D4F5017" ID-DOI="10.1080/00222933.2018.1437231" ID-ISSN="1464-5262" ID-Zenodo-Dep="5178489" ID-ZooBank="9FC2A016-1295-4047-877E-BC9030BD2BD2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628626551251" checkinUser="felipe" docAuthor="Santarém, Maria Clara Alves, Borkent, Art, Spinelli, Gustavo & Felippe-Bauer, Maria Luiza" docDate="2018" docId="375B87BEFFC1A60526EFCA68FB31FEF1" docLanguage="en" docName="JNatHist.52.9-10.509-540.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (9 - 10)" docSource="http://dx.doi.org/10.1080/00222933.2018.1437231" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Downeshelea multilineata" docType="key" docVersion="3" lastPageNumber="513" masterDocId="CB62FFC6FFC3A600264FCC1AFFB5FFA5" masterDocTitle="New Neotropical species of Downeshelea Wirth and Grogan and redescription of D. multilineata (Lutz) (Diptera: Ceratopogonidae)" masterLastPageNumber="540" masterPageNumber="509" pageNumber="510" updateTime="1699048774547" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="84C19F5B6CC34670EC157BD1C1304D6B">
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<mods:title id="05366AD0D48C419268657906ABB647B6">New Neotropical species of Downeshelea Wirth and Grogan and redescription of D. multilineata (Lutz) (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="07E4F8E59E867D3894451D05857CF553">Santarém, Maria Clara Alves</mods:namePart>
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<mods:namePart id="1A0674BA1B376DCE769339382AD979D3">Borkent, Art</mods:namePart>
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<mods:namePart id="F1B986A76C64735C62E50D86F5D70AE2">Spinelli, Gustavo</mods:namePart>
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</mods:name>
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<mods:name id="9F6FB7408788F90E8A456F1CA545B851" type="personal">
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<mods:roleTerm id="B4F6ED953E367130467615AE461904C9">Author</mods:roleTerm>
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<mods:namePart id="C96CF2499823394211ACABE3AE6E08FB">Felippe-Bauer, Maria Luiza</mods:namePart>
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<mods:typeOfResource id="1992B95B3F99CBD11C580210D83800FF">text</mods:typeOfResource>
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<mods:title id="AAE885C2B3F86C4851E059B00D44C496">Journal of Natural History</mods:title>
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<mods:part id="9FD2031401E3E305F6CB6B6614A964FC">
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<mods:date id="151862E22D84EAD57BCCDE1B53CCFAC9">2018</mods:date>
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<mods:detail id="8334C3F04E062812BF453F3C97384413" type="series">
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<mods:title id="0757D1F9A4EEE1B4D7A74A21235FC458">J. Nat. Hist.</mods:title>
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<mods:number id="96A7A368C35A78728B8FA921ACA55ADE">2018-03-07</mods:number>
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<mods:detail id="4FA54946B263ADB04903B9770C59941A" type="volume">
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<mods:number id="1C562BE1F4AFAF57A801319E26E80C4C">52</mods:number>
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<mods:detail id="105320A574AA9FA8646841BFF787951C" type="issue">
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<mods:number id="8D5DF374EC657A50B4C9A2BD21024C33">9 - 10</mods:number>
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<mods:start id="A5F576B53BA4B4A3A16F322124DCE60D">509</mods:start>
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<mods:url id="876402BFF732DF98EBF011B2B6E7AB48">http://dx.doi.org/10.1080/00222933.2018.1437231</mods:url>
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<mods:classification id="5DC9373EA00914BCA1A2A09D17CDA9FF">journal article</mods:classification>
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<mods:identifier id="EC39B624B9EAC9FE6FB82986269AE030" type="DOI">10.1080/00222933.2018.1437231</mods:identifier>
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<mods:identifier id="60A910D7E9D415CBA2D2D1AC09992980" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="DB0F955CEFE170D5C8014426CC236AAF" type="Zenodo-Dep">5178489</mods:identifier>
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<mods:identifier id="C6F37014EA072189C8212360E399097C" type="ZooBank">9FC2A016-1295-4047-877E-BC9030BD2BD2</mods:identifier>
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<treatment id="375B87BEFFC1A60526EFCA68FB31FEF1" LSID="urn:lsid:plazi:treatment:375B87BEFFC1A60526EFCA68FB31FEF1" httpUri="http://treatment.plazi.org/id/375B87BEFFC1A60526EFCA68FB31FEF1" lastPageId="5" lastPageNumber="513" pageId="2" pageNumber="510">
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<subSubSection id="F7E86523FFC1A60226EFCA68FE1DF91A" pageId="2" pageNumber="510" type="nomenclature">
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<paragraph id="BF4D36A8FFC1A60226EFCA68FC80F92E" blockId="2.[160,821,1650,1675]" box="[160,821,1650,1675]" pageId="2" pageNumber="510">
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<heading id="E40581C4FFC1A60226EFCA68FC80F92E" bold="true" box="[160,821,1650,1675]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="510" reason="3">
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<emphasis id="8D86EABAFFC1A60226EFCA68FC80F92E" bold="true" box="[160,821,1650,1675]" pageId="2" pageNumber="510">
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<taxonomicName id="78F24D2BFFC1A60226EFCA68FE57F92E" ID-CoL="37HPF" baseAuthorityName="Lutz" baseAuthorityYear="1914" box="[160,482,1650,1675]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="510" phylum="Arthropoda" rank="species" species="multilineata">
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<emphasis id="8D86EABAFFC1A60226EFCA68FE57F92E" bold="true" box="[160,482,1650,1675]" italics="true" pageId="2" pageNumber="510">Downeshelea multilineata</emphasis>
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</taxonomicName>
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species group: new group
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="BF4D36A8FFC1A60226EFCABCFE1DF91A" blockId="2.[160,424,1702,1727]" box="[160,424,1702,1727]" pageId="2" pageNumber="510">
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<emphasis id="8D86EABAFFC1A60226EFCABCFE17F91A" box="[160,418,1702,1727]" italics="true" pageId="2" pageNumber="510">Abbreviated description</emphasis>
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.
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||
</paragraph>
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</subSubSection>
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<subSubSection id="F7E86523FFC0A603268FCC94FDA7FCDB" pageId="3" pageNumber="511" type="description">
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<paragraph id="BF4D36A8FFC0A603268FCC94FDA7FCDB" blockId="3.[160,1156,142,894]" pageId="3" pageNumber="511">
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Small- to medium-sized species (male wing length
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<quantity id="780A9B4DFFC0A60324B2CC94FC2BFF03" box="[765,926,142,166]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" metricValueMax="1.37" metricValueMin="0.85" pageId="3" pageNumber="511" unit="mm" value="1.11" valueMax="1.37" valueMin="0.85">0.85–1.37 mm</quantity>
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, female wing length
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<quantity id="780A9B4DFFC0A60326EFCCABFEFDFF6C" box="[160,328,177,201]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" metricValueMax="1.3" metricValueMin="0.92" pageId="3" pageNumber="511" unit="mm" value="1.11" valueMax="1.3" valueMin="0.92">0.92–1.30 mm</quantity>
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), brown in colouration. Male eyes narrowly separated by a distance shorter than one ommatidia or contiguous in lower area; female eyes contiguous. Male antenna pale, except base of flagellomere 1, distal portion of flagellomere 10 and flagellomeres 11–13 brown; female antenna brown, basal portion of flagellomeres 1–7 pale; flagellomeres 2–8 somewhat barrel-shaped, 9–13 cylindrical, elongate. Palpus brown, segment 3 swollen or slightly swollen on midportion, with broad, deep sensory pit. Male mandible unarmed, female mandible with 9–12 teeth. Legs brown or pale brown, hind leg darker (except in
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<taxonomicName id="78F24D2BFFC0A6032451CDB9FD0BFE19" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[542,702,419,444]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="511" phylum="Arthropoda" rank="species" species="casimirensis">
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<emphasis id="8D86EABAFFC0A6032451CDB9FD0BFE19" box="[542,702,419,444]" italics="true" pageId="3" pageNumber="511">D. casimirensis</emphasis>
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</taxonomicName>
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), hind tibial comb with six spines; apical spines of male hind tarsomeres 2–4: 1–1–1; male claws about 0.35–0.45 length of their respective tarsomere 5; female hind leg claw 0.89–1.53 length of tarsomere 5. Wing hyaline, with dark bristles on costa; two distinctive black spots, one on r-m, other in r
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<subScript id="237634EDFFC0A6032236CE0CFB36FD83" attach="left" box="[1145,1155,534,550]" fontSize="7" pageId="3" pageNumber="511">3</subScript>
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from apex of r
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<subScript id="237634EDFFC0A6032700CE23FEECFDEC" attach="left" box="[335,345,569,585]" fontSize="7" pageId="3" pageNumber="511">2</subScript>
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to M
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<subScript id="237634EDFFC0A60327D1CE23FE1DFDEC" attach="right" box="[414,424,569,585]" fontSize="7" pageId="3" pageNumber="511">1</subScript>
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; three distinctive greyish areas: first near apex of M
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<subScript id="237634EDFFC0A6032258CE23FB94FDEC" attach="left" box="[1047,1057,569,585]" fontSize="7" pageId="3" pageNumber="511">1</subScript>
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, second sigmoid-shaped near apex of M
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<subScript id="237634EDFFC0A6032447CE41FDA7FDCE" attach="left" box="[520,530,603,619]" fontSize="7" pageId="3" pageNumber="511">2</subScript>
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extending to wing margin in m
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<subScript id="237634EDFFC0A60325CFCE41FC3FFDCE" attach="left" box="[896,906,603,619]" fontSize="7" pageId="3" pageNumber="511">2</subScript>
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, third over CuA
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||
<subScript id="237634EDFFC0A6032270CE41FBFCFDCE" attach="right" box="[1087,1097,603,619]" fontSize="7" pageId="3" pageNumber="511">2</subScript>
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, also in CuA
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<quantity id="780A9B4DFFC0A60326BDCE64FE94FD2E" box="[242,289,627,654]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="511" unit="in" value="1.0">
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<subScript id="237634EDFFC0A60326BDCE64FF49FD2B" attach="left" box="[242,252,638,654]" fontSize="7" pageId="3" pageNumber="511">1</subScript>
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in
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||
some species. Male tergite 9 gradually tapering distally, distal ½ with sclerotised band laterally; gonocoxite moderately stout; gonostylus nearly straight, proximal ½ slightly pilose; parameres fused on basal portion, each with trilobed basal arm, heavily sclerotised; stem stout, straight or sinuous; distal portion abruptly bent ventrally, directed anteromesally, tapering to pointed tip, overlapped in some species; aedeagus subtriangular to rectangular, posterior portion with mesal excavation. Female with spermathecae variable in size, with short sclerotised ducts, third rudimentary spermathecae generally present.
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</paragraph>
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</subSubSection>
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<subSubSection id="F7E86523FFC0A60526EFCFB9FB31FEF1" lastPageId="5" lastPageNumber="513" pageId="3" pageNumber="511" type="key">
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<paragraph id="BF4D36A8FFC0A60326EFCFB9FCCEFC1E" blockId="3.[160,891,931,955]" box="[160,891,931,955]" pageId="3" pageNumber="511">
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<emphasis id="8D86EABAFFC0A60326EFCFB9FCCEFC1E" bold="true" box="[160,891,931,955]" pageId="3" pageNumber="511">
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Key to males of the
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<taxonomicName id="78F24D2BFFC0A60327D7CFB9FD7FFC1E" ID-CoL="37HPF" baseAuthorityName="Lutz" baseAuthorityYear="1914" box="[408,714,931,955]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="511" phylum="Arthropoda" rank="species" species="multilineata">
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<emphasis id="8D86EABAFFC0A60327D7CFB9FD7FFC1E" bold="true" box="[408,714,931,955]" italics="true" pageId="3" pageNumber="511">Downeshelea multilineata</emphasis>
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</taxonomicName>
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species group
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</emphasis>
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</paragraph>
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<key id="CB636199FFC0A60426EFCFE8FB31FAE4" lastPageId="4" lastPageNumber="512" pageId="3" pageNumber="511">
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<keyStep id="04062E0DFFC0A60326EFCFE8FB36FB7F" pageId="3" pageNumber="511">
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<paragraph id="BF4D36A8FFC0A60326EFCFE8FB36FB30" blockId="3.[160,1156,1010,1727]" pageId="3" pageNumber="511">
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<keyLead id="0403959DFFC0A60326EFCFE8FB36FB30" pageId="3" pageNumber="511">
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1. Legs brown or pale brown, without bands (although some with varying pigmentation); wing with three distinctive greyish areas (in addition to the two dark spots on r-m and in r
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<subScript id="237634EDFFC0A603268DC858FF79FBF7" attach="left" box="[194,204,1090,1106]" fontSize="7" pageId="3" pageNumber="511">3</subScript>
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from the apex of r
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<subScript id="237634EDFFC0A60327F4C858FE70FBF7" attach="left" box="[443,453,1090,1106]" fontSize="7" pageId="3" pageNumber="511">2</subScript>
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to M
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<subScript id="237634EDFFC0A6032443C858FDA3FBF7" attach="right" box="[524,534,1090,1106]" fontSize="7" pageId="3" pageNumber="511">1</subScript>
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) (
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<figureCitation id="27C92A2DFFC0A603247DC82DFD15FBEA" box="[562,672,1079,1103]" captionStart="Figure 1" captionStartId="6.[160,225,1575,1598]" captionTargetBox="[166,1142,144,1526]" captionTargetId="figure-1@6.[163,1152,142,1544]" captionTargetPageId="6" captionText="Figure 1. Downeshelea multilineata (Lutz). Male: (a) wing; arrows: greyish spots; (b) fore, mid, hind legs (bottom to top); (c) head, anterior view. Female: (d) wing; (e) eye separation, anterior view; (f) antenna; (g) palpus; (h) fore, hind tarsomeres (left to right; from different specimens); (i) apex of female abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5178491" httpUri="https://zenodo.org/record/5178491/files/figure.png" pageId="3" pageNumber="511">Figure 1a</figureCitation>
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); parameres distal portion abruptly bent ventrally, directed anteromesally, tapering to pointed tip, 0.50–0.75 of total paramere length ..................................................
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<taxonomicName id="78F24D2BFFC0A6032463C866FD1AFB30" ID-CoL="37HPF" baseAuthorityName="Lutz" baseAuthorityYear="1914" box="[556,687,1148,1173]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="511" phylum="Arthropoda" rank="species" species="multilineata">
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<emphasis id="8D86EABAFFC0A6032463C866FD1AFB30" box="[556,687,1148,1173]" italics="true" pageId="3" pageNumber="511">multilineata</emphasis>
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</taxonomicName>
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species group.................................................. 2
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</keyLead>
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</paragraph>
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<paragraph id="BF4D36A8FFC0A60326EFC885FB36FB7F" blockId="3.[160,1156,1010,1727]" pageId="3" pageNumber="511">
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<keyLead id="0403959DFFC0A60326EFC885FB36FB7F" pageId="3" pageNumber="511">
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– Legs various; wing with variable distinctive greyish areas; parameres distal portion not as above ........................................................................................ other
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<taxonomicName id="78F24D2BFFC0A60325D8C8DBFB36FB7F" box="[919,1155,1217,1242]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="511" phylum="Arthropoda" rank="species" species="undefined-2">
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<emphasis id="8D86EABAFFC0A60325D8C8DBFB9DFB7F" box="[919,1064,1217,1242]" italics="true" pageId="3" pageNumber="511">Downeshelea</emphasis>
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species
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="04062E0DFFC0A60326EFC8FFFB36FA0F" pageId="3" pageNumber="511">
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<paragraph id="BF4D36A8FFC0A60326EFC8FFFB36FAE7" blockId="3.[160,1156,1010,1727]" pageId="3" pageNumber="511">
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||
<keyLead id="0403959DFFC0A60326EFC8FFFB36FAE7" pageId="3" pageNumber="511">2. Legs brown, hind femur brown; medium- to large-sized species; midtarsomere 1 with seven to eight ventral spines. Parameres with a short posteromedian projection on the medial fused portion................................................................................................................................ 3</keyLead>
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</paragraph>
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<paragraph id="BF4D36A8FFC0A60326EFC956FB36FA0F" blockId="3.[160,1156,1010,1727]" pageId="3" pageNumber="511">
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||
<keyLead id="0403959DFFC0A60326EFC956FB36FA0F" pageId="3" pageNumber="511">– Legs brown or pale brown, hind femur darker subapically; small- to medium-sized species; midtarsomere 1 with two to six ventral spines. Parameres without a posteromedian projection on the medial fused portion....................................................................... 4</keyLead>
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</paragraph>
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||
</keyStep>
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<keyStep id="04062E0DFFC0A60426EFC9AFFB31FF6C" lastPageId="4" lastPageNumber="512" pageId="3" pageNumber="511">
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<paragraph id="BF4D36A8FFC0A60326EFC9AFFB31F9F2" blockId="3.[160,1156,1010,1727]" pageId="3" pageNumber="511">
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<keyLead id="0403959DFFC0A60326EFC9AFFB31F9F2" pageId="3" pageNumber="511">
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3. Tergite 9 with short, broad apicolateral process; paramere stem sinuous on midportion, expanded subapically forming a broad lobe laterally directed; distal portion 0.55– 0.84 of total paramere length (
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<figureCitation id="27C92A2DFFC0A6032476C9E0FD16F9B7" box="[569,675,1530,1554]" captionStart="Figure 8" captionStartId="23.[160,225,1645,1668]" captionTargetBox="[242,1072,145,1606]" captionTargetId="figure-1@23.[242,1073,142,1614]" captionTargetPageId="23" captionText="Figure 8. Downeshelea multilineata (Lutz), (a) male terminalia with aedeagus; (b) parameres. Downeshelea casimirensis sp. nov. (c) male terminalia; (d) aedeagus; (e) parameres. Downeshelea costaricensis sp. nov., (f) male terminalia with aedeagus; (g) parameres. Downeshelea eclectica sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178505" httpUri="https://zenodo.org/record/5178505/files/figure.png" pageId="3" pageNumber="511">Figure 8i</figureCitation>
|
||
); aedeagus without elliptical sclerotised anterior areas, distal portion terminating in sclerotised non-serrate process (
|
||
<figureCitation id="27C92A2DFFC0A603225ECA07FBCEF990" box="[1041,1147,1565,1589]" captionStart="Figure 8" captionStartId="23.[160,225,1645,1668]" captionTargetBox="[242,1072,145,1606]" captionTargetId="figure-1@23.[242,1073,142,1614]" captionTargetPageId="23" captionText="Figure 8. Downeshelea multilineata (Lutz), (a) male terminalia with aedeagus; (b) parameres. Downeshelea casimirensis sp. nov. (c) male terminalia; (d) aedeagus; (e) parameres. Downeshelea costaricensis sp. nov., (f) male terminalia with aedeagus; (g) parameres. Downeshelea eclectica sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178505" httpUri="https://zenodo.org/record/5178505/files/figure.png" pageId="3" pageNumber="511">Figure 8h</figureCitation>
|
||
) .....................................................................................................................................
|
||
<taxonomicName id="78F24D2BFFC0A60325E9CA24FB97F9F2" authority="Santarém & Borkent & Spinelli & Felippe-Bauer, 2018" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[934,1058,1598,1623]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="511" phylum="Arthropoda" rank="species" species="eclectica" status="sp. nov.">
|
||
<emphasis id="8D86EABAFFC0A60325E9CA24FB97F9F2" box="[934,1058,1598,1623]" italics="true" pageId="3" pageNumber="511">D. eclectica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="96B557C1FFC0A6032264CA25FB31F9F2" box="[1067,1156,1599,1623]" pageId="3" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="BF4D36A8FFC0A60426EFCA78FB31FF6C" blockId="3.[160,1156,1010,1727]" lastBlockId="4.[160,1156,142,1345]" lastPageId="4" lastPageNumber="512" pageId="3" pageNumber="511">
|
||
<keyLead id="0403959DFFC0A60426EFCA78FB31FF6C" lastPageId="4" lastPageNumber="512" pageId="3" pageNumber="511">
|
||
– Tergite 9 with long, slender apicolateral process; paramere stem sinuous, proximal 2/3 directed posterolaterally, distal 1/3 slender directed posteromesally; distal portion 0.50 of total paramere length (
|
||
<figureCitation id="27C92A2DFFC0A60327AACABDFDFFF91A" box="[485,586,1703,1727]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="3" pageNumber="511">Figure 9i</figureCitation>
|
||
); aedeagus with two elliptical sclerotised anterior areas; distal portion terminating in sclerotised serrate processes (
|
||
<figureCitation id="27C92A2DFFC7A60425EFCC94FBB9FF03" box="[928,1036,142,166]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="4" pageNumber="512">Figure 9h</figureCitation>
|
||
).................... ......................................................................................................................................
|
||
<taxonomicName id="78F24D2BFFC7A60425E6CCAAFB97FF6C" authority="Santarém & Borkent & Spinelli & Felippe-Bauer, 2018" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[937,1058,176,201]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="moravia" status="sp. nov.">
|
||
<emphasis id="8D86EABAFFC7A60425E6CCAAFB97FF6C" box="[937,1058,176,201]" italics="true" pageId="4" pageNumber="512">D. moravia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="96B557C1FFC7A6042264CCABFB31FF6C" box="[1067,1156,177,201]" pageId="4" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="04062E0DFFC7A60426EFCCCEFB36FEF1" pageId="4" pageNumber="512">
|
||
<paragraph id="BF4D36A8FFC7A60426EFCCCEFB36FEAB" blockId="4.[160,1156,142,1345]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFCCCEFB36FEAB" pageId="4" pageNumber="512">
|
||
4. CuA
|
||
<subScript id="237634EDFFC7A60426A6CCC4FF46FF4B" attach="left" box="[233,243,222,238]" fontSize="7" pageId="4" pageNumber="512">1</subScript>
|
||
pale; paramere distal portion short, not reaching the medial fused portion, 0.53– 0.62 of total paramere length ............................................................................................................ 5
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="BF4D36A8FFC7A60426EFCD03FB36FEF1" blockId="4.[160,1156,142,1345]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFCD03FB36FEF1" pageId="4" pageNumber="512">
|
||
– CuA
|
||
<subScript id="237634EDFFC7A60426A8CD39FF44FE96" attach="left" box="[231,241,291,307]" fontSize="7" pageId="4" pageNumber="512">1</subScript>
|
||
greyish; paramere distal portion elongate, reaching the medial fused portion, 0.62–0.80 of total paramere length .................................................................................................. 7
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="04062E0DFFC7A60426EFCD44FB31FD81" pageId="4" pageNumber="512">
|
||
<paragraph id="BF4D36A8FFC7A60426EFCD44FB36FE19" blockId="4.[160,1156,142,1345]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFCD44FB36FE19" pageId="4" pageNumber="512">5. Legs pale brown, foretibia pale or darker apically; halter knob pale; aedeagus subtriangular, with or without sclerotised anteromesal areas, not reaching midlength when present; basal arch extending to 0.22–0.56 of total length....................................... 6</keyLead>
|
||
</paragraph>
|
||
<paragraph id="BF4D36A8FFC7A60426EFCDDCFB31FD81" blockId="4.[160,1156,142,1345]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFCDDCFB31FD81" pageId="4" pageNumber="512">
|
||
– Legs brown (
|
||
<figureCitation id="27C92A2DFFC7A6042719CDDCFE73FE7B" box="[342,454,454,478]" captionStart="Figure 7" captionStartId="22.[160,225,1582,1605]" captionTargetBox="[163,1150,144,1550]" captionTargetId="figure-1@22.[163,1152,142,1551]" captionTargetPageId="22" captionText="Figure 7. Downeshelea marambaia sp. nov., male (a–d). (a) wing; (b) antenna; (c) head, anterior view; (d) fore, mid, hind legs (left to right). Downeshelea moravia sp. nov., male (e–h). (e) wing; (f) head, anterior view; (g) palpus; (h) fore, mid, hind legs (left to right)." figureDoi="http://doi.org/10.5281/zenodo.5178503" httpUri="https://zenodo.org/record/5178503/files/figure.png" pageId="4" pageNumber="512">Figure 7d</figureCitation>
|
||
); halter knob brown; aedeagus rectangular, with two large elliptical sclerotised anteromesal areas reaching midlength; basal arch extending to 0.17 of total length (
|
||
<figureCitation id="27C92A2DFFC7A60427E9CE16FDBEFD81" box="[422,523,524,548]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="4" pageNumber="512">Figure 9f</figureCitation>
|
||
)................................................................
|
||
<taxonomicName id="78F24D2BFFC7A60425CBCE11FB97FD81" authority="Santarém & Borkent & Spinelli & Felippe-Bauer, 2018" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[900,1058,523,548]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="marambaia" status="sp. nov.">
|
||
<emphasis id="8D86EABAFFC7A60425CBCE11FB97FD81" box="[900,1058,523,548]" italics="true" pageId="4" pageNumber="512">D. marambaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="96B557C1FFC7A6042264CE16FB31FD81" box="[1067,1156,524,548]" pageId="4" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="04062E0DFFC7A60426EFCE34FB31FCFE" pageId="4" pageNumber="512">
|
||
<paragraph id="BF4D36A8FFC7A60426EFCE34FB31FD0B" blockId="4.[160,1156,142,1345]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFCE34FB31FD0B" pageId="4" pageNumber="512">
|
||
6. Hind tibia pale (
|
||
<figureCitation id="27C92A2DFFC7A6042725CE34FE66FDE3" box="[362,467,558,582]" captionStart="Figure 2" captionStartId="9.[160,225,1672,1695]" captionTargetBox="[165,1150,1197,1640]" captionTargetId="figure-279@9.[163,1152,1196,1641]" captionTargetPageId="9" captionText="Figure 2. Downeshelea casimirensis sp. nov., male. (a) Wing; (b) eye separation, anterior view; (c) head, anterior view; (d) fore, mid, hind legs (top to bottom)." figureDoi="http://doi.org/10.5281/zenodo.5178493" httpUri="https://zenodo.org/record/5178493/files/figure.png" pageId="4" pageNumber="512">Figure 2d</figureCitation>
|
||
); gonostylus 0.81 length of gonocoxite (
|
||
<figureCitation id="27C92A2DFFC7A60425DCCE34FC4FFDE3" box="[915,1018,558,582]" captionStart="Figure 8" captionStartId="23.[160,225,1645,1668]" captionTargetBox="[242,1072,145,1606]" captionTargetId="figure-1@23.[242,1073,142,1614]" captionTargetPageId="23" captionText="Figure 8. Downeshelea multilineata (Lutz), (a) male terminalia with aedeagus; (b) parameres. Downeshelea casimirensis sp. nov. (c) male terminalia; (d) aedeagus; (e) parameres. Downeshelea costaricensis sp. nov., (f) male terminalia with aedeagus; (g) parameres. Downeshelea eclectica sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178505" httpUri="https://zenodo.org/record/5178505/files/figure.png" pageId="4" pageNumber="512">Figure 8c</figureCitation>
|
||
); parameres stem straight, gradually swollen to apex; distal portion deeply curved (
|
||
<figureCitation id="27C92A2DFFC7A6042248CE4BFBC0FDCC" box="[1031,1141,593,617]" captionStart="Figure 8" captionStartId="23.[160,225,1645,1668]" captionTargetBox="[242,1072,145,1606]" captionTargetId="figure-1@23.[242,1073,142,1614]" captionTargetPageId="23" captionText="Figure 8. Downeshelea multilineata (Lutz), (a) male terminalia with aedeagus; (b) parameres. Downeshelea casimirensis sp. nov. (c) male terminalia; (d) aedeagus; (e) parameres. Downeshelea costaricensis sp. nov., (f) male terminalia with aedeagus; (g) parameres. Downeshelea eclectica sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178505" httpUri="https://zenodo.org/record/5178505/files/figure.png" pageId="4" pageNumber="512">Figure 8e</figureCitation>
|
||
); aedeagus with pair of submedian sclerotised stripes and oval areas, basal arch extending to 0.56 of total length (
|
||
<figureCitation id="27C92A2DFFC7A60427A6CE8CFDE1FD0B" box="[489,596,662,686]" captionStart="Figure 8" captionStartId="23.[160,225,1645,1668]" captionTargetBox="[242,1072,145,1606]" captionTargetId="figure-1@23.[242,1073,142,1614]" captionTargetPageId="23" captionText="Figure 8. Downeshelea multilineata (Lutz), (a) male terminalia with aedeagus; (b) parameres. Downeshelea casimirensis sp. nov. (c) male terminalia; (d) aedeagus; (e) parameres. Downeshelea costaricensis sp. nov., (f) male terminalia with aedeagus; (g) parameres. Downeshelea eclectica sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178505" httpUri="https://zenodo.org/record/5178505/files/figure.png" pageId="4" pageNumber="512">Figure 8d</figureCitation>
|
||
)....................................................
|
||
<taxonomicName id="78F24D2BFFC7A60425C9CE8FFB93FD0B" authority="Santarém & Borkent & Spinelli & Felippe-Bauer, 2018" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[902,1062,661,686]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="casimirensis" status="sp. nov.">
|
||
<emphasis id="8D86EABAFFC7A60425C9CE8FFB93FD0B" box="[902,1062,661,686]" italics="true" pageId="4" pageNumber="512">D. casimirensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="96B557C1FFC7A6042262CE8CFB31FD0B" box="[1069,1156,662,686]" pageId="4" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="BF4D36A8FFC7A60426EFCEA3FB31FCFE" blockId="4.[160,1156,142,1345]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFCEA3FB31FCFE" pageId="4" pageNumber="512">
|
||
– Hind tibia darker apically and on proximal third (
|
||
<figureCitation id="27C92A2DFFC7A6042545CEA3FCC2FD74" box="[778,887,697,721]" captionStart="Figure 5" captionStartId="19.[160,225,1556,1579]" captionTargetBox="[163,1151,146,1524]" captionTargetId="figure-1@19.[163,1152,142,1525]" captionTargetPageId="19" captionText="Figure 5. Downeshelea jarina sp. nov. Male: (a) wing; (b) head, anterior view; (c) fore, mid, hind legs (left to right). Female: (d) wing; (e) head, anterior view; (f) fore, mid, hind tarsomeres (top to bottom); (g) palpus; (h) apex of female abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5178499" httpUri="https://zenodo.org/record/5178499/files/figure.png" pageId="4" pageNumber="512">Figure 5c</figureCitation>
|
||
); gonostylus 0.58–0.64 length of gonocoxite (
|
||
<figureCitation id="27C92A2DFFC7A6042781CEC1FD8AFD56" box="[462,575,731,755]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="4" pageNumber="512">Figure 9a</figureCitation>
|
||
); parameres stem basolaterally expanded, more slender and convergent distally, distal portion nearly straight (
|
||
<figureCitation id="27C92A2DFFC7A60425DECEE4FC4AFCB3" box="[913,1023,766,790]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="4" pageNumber="512">Figure 9b</figureCitation>
|
||
); aedeagus without sclerotised stripes and oval areas, basal arch extending to 0.22–0.28 of total length (
|
||
<figureCitation id="27C92A2DFFC7A604275CCF59FECBFCFE" box="[275,382,835,859]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="4" pageNumber="512">Figure 9a</figureCitation>
|
||
) ....................................................................................................
|
||
<taxonomicName id="78F24D2BFFC7A604258BCF58FB96FCFE" authority="Santarém & Borkent & Spinelli & Felippe-Bauer, 2018" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[964,1059,834,859]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="jarina" status="sp. nov.">
|
||
<emphasis id="8D86EABAFFC7A604258BCF58FB96FCFE" box="[964,1059,834,859]" italics="true" pageId="4" pageNumber="512">D. jarina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="96B557C1FFC7A6042264CF59FB31FCFE" box="[1067,1156,835,859]" pageId="4" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="04062E0DFFC7A60426EFCF7CFB36FB8E" pageId="4" pageNumber="512">
|
||
<paragraph id="BF4D36A8FFC7A60426EFCF7CFB36FC66" blockId="4.[160,1156,142,1345]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFCF7CFB36FC66" pageId="4" pageNumber="512">
|
||
7. Sternite 9 with large prominent convex median lobe (
|
||
<figureCitation id="27C92A2DFFC7A604254ACF7CFCDFFCDB" box="[773,874,870,894]" captionStart="Figure 8" captionStartId="23.[160,225,1645,1668]" captionTargetBox="[242,1072,145,1606]" captionTargetId="figure-1@23.[242,1073,142,1614]" captionTargetPageId="23" captionText="Figure 8. Downeshelea multilineata (Lutz), (a) male terminalia with aedeagus; (b) parameres. Downeshelea casimirensis sp. nov. (c) male terminalia; (d) aedeagus; (e) parameres. Downeshelea costaricensis sp. nov., (f) male terminalia with aedeagus; (g) parameres. Downeshelea eclectica sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178505" httpUri="https://zenodo.org/record/5178505/files/figure.png" pageId="4" pageNumber="512">Figure 8a</figureCitation>
|
||
); paramere stem straight, distal portion slightly curved (
|
||
<figureCitation id="27C92A2DFFC7A6042459CF93FD36FC04" box="[534,643,905,929]" captionStart="Figure 8" captionStartId="23.[160,225,1645,1668]" captionTargetBox="[242,1072,145,1606]" captionTargetId="figure-1@23.[242,1073,142,1614]" captionTargetPageId="23" captionText="Figure 8. Downeshelea multilineata (Lutz), (a) male terminalia with aedeagus; (b) parameres. Downeshelea casimirensis sp. nov. (c) male terminalia; (d) aedeagus; (e) parameres. Downeshelea costaricensis sp. nov., (f) male terminalia with aedeagus; (g) parameres. Downeshelea eclectica sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178505" httpUri="https://zenodo.org/record/5178505/files/figure.png" pageId="4" pageNumber="512">Figure 8b</figureCitation>
|
||
); aedeagus subtriangular, with two elliptical sclerotised anterior areas (
|
||
<figureCitation id="27C92A2DFFC7A6042795CFB1FDF5FC66" box="[474,576,939,963]" captionStart="Figure 8" captionStartId="23.[160,225,1645,1668]" captionTargetBox="[242,1072,145,1606]" captionTargetId="figure-1@23.[242,1073,142,1614]" captionTargetPageId="23" captionText="Figure 8. Downeshelea multilineata (Lutz), (a) male terminalia with aedeagus; (b) parameres. Downeshelea casimirensis sp. nov. (c) male terminalia; (d) aedeagus; (e) parameres. Downeshelea costaricensis sp. nov., (f) male terminalia with aedeagus; (g) parameres. Downeshelea eclectica sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178505" httpUri="https://zenodo.org/record/5178505/files/figure.png" pageId="4" pageNumber="512">Figure 8a</figureCitation>
|
||
) ..............................................................
|
||
<taxonomicName id="78F24D2BFFC7A60425EECFB0FB36FC66" ID-CoL="37HPF" authority="(Lutz)" baseAuthorityName="Lutz" baseAuthorityYear="1914" box="[929,1155,938,963]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="multilineata">
|
||
<emphasis id="8D86EABAFFC7A60425EECFB0FB8AFC66" box="[929,1087,938,963]" italics="true" pageId="4" pageNumber="512">D. multilineata</emphasis>
|
||
(Lutz)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="BF4D36A8FFC7A60426EFCFD4FB36FB8E" blockId="4.[160,1156,142,1345]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFCFD4FB36FB8E" pageId="4" pageNumber="512">– Sternite 9 with moderately convex median lobe; paramere stem and distal portion various; aedeagus rectangular or subrectangular, without elliptical sclerotised anterior areas ............................................................................................................................................................... 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="04062E0DFFC7A60426EFC82CFB31FAE4" pageId="4" pageNumber="512">
|
||
<paragraph id="BF4D36A8FFC7A60426EFC82CFB31FB13" blockId="4.[160,1156,142,1345]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFC82CFB31FB13" pageId="4" pageNumber="512">
|
||
8. Halter knob pale; midtibia paler basally (
|
||
<figureCitation id="27C92A2DFFC7A60424D7C82CFCB6FBEB" box="[664,771,1078,1102]" captionStart="Figure 3" captionStartId="11.[160,225,1595,1618]" captionTargetBox="[167,1149,145,1564]" captionTargetId="figure-1@11.[163,1152,142,1564]" captionTargetPageId="11" captionText="Figure 3. Downeshelea costaricensis sp. nov. Male: (a) wing; (b) head, anterior view; (c) fore, mid, hind legs (left to right). Female: (d) wing; (e) palpus; (f) fore, mid, hind tarsomeres (bottom to top); (g) head, anterior view; (h) apex of female abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5178495" httpUri="https://zenodo.org/record/5178495/files/figure.png" pageId="4" pageNumber="512">Figure 3c</figureCitation>
|
||
); paramere stem broad, sinuous, expanded apicolaterally, distal portion gradually curved (
|
||
<figureCitation id="27C92A2DFFC7A604251BC843FC76FBD4" box="[852,963,1113,1137]" captionStart="Figure 8" captionStartId="23.[160,225,1645,1668]" captionTargetBox="[242,1072,145,1606]" captionTargetId="figure-1@23.[242,1073,142,1614]" captionTargetPageId="23" captionText="Figure 8. Downeshelea multilineata (Lutz), (a) male terminalia with aedeagus; (b) parameres. Downeshelea casimirensis sp. nov. (c) male terminalia; (d) aedeagus; (e) parameres. Downeshelea costaricensis sp. nov., (f) male terminalia with aedeagus; (g) parameres. Downeshelea eclectica sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178505" httpUri="https://zenodo.org/record/5178505/files/figure.png" pageId="4" pageNumber="512">
|
||
Figure
|
||
<quantity id="780A9B4DFFC7A60425E9C843FC76FBD4" box="[934,963,1113,1137]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="4" pageNumber="512" unit="g" value="8.0">8g</quantity>
|
||
</figureCitation>
|
||
); aedeagus subrectangular, basal arch extending to 0.22–0.26 of total length (
|
||
<figureCitation id="27C92A2DFFC7A60425C9C861FC5EFB36" box="[902,1003,1147,1171]" captionStart="Figure 8" captionStartId="23.[160,225,1645,1668]" captionTargetBox="[242,1072,145,1606]" captionTargetId="figure-1@23.[242,1073,142,1614]" captionTargetPageId="23" captionText="Figure 8. Downeshelea multilineata (Lutz), (a) male terminalia with aedeagus; (b) parameres. Downeshelea casimirensis sp. nov. (c) male terminalia; (d) aedeagus; (e) parameres. Downeshelea costaricensis sp. nov., (f) male terminalia with aedeagus; (g) parameres. Downeshelea eclectica sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178505" httpUri="https://zenodo.org/record/5178505/files/figure.png" pageId="4" pageNumber="512">Figure 8f</figureCitation>
|
||
).......................... .............................................................................................................................
|
||
<taxonomicName id="78F24D2BFFC7A6042534C887FB96FB13" authority="Santarém & Borkent & Spinelli & Felippe-Bauer, 2018" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[891,1059,1181,1206]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="costaricensis" status="sp. nov.">
|
||
<emphasis id="8D86EABAFFC7A6042534C887FB96FB13" box="[891,1059,1181,1206]" italics="true" pageId="4" pageNumber="512">D. costaricensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="96B557C1FFC7A6042264C884FB31FB13" box="[1067,1156,1182,1206]" pageId="4" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="BF4D36A8FFC7A60426EFC8DBFB31FAE4" blockId="4.[160,1156,142,1345]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFC8DBFB31FAE4" pageId="4" pageNumber="512">
|
||
– Halter knob brown; midtibia brown (
|
||
<figureCitation id="27C92A2DFFC7A6042416C8DBFD74FB7C" box="[601,705,1217,1241]" captionStart="Figure 6" captionStartId="21.[160,225,1579,1602]" captionTargetBox="[164,1149,144,1545]" captionTargetId="figure-1@21.[163,1152,142,1548]" captionTargetPageId="21" captionText="Figure 6. Downeshelea litorale sp. nov. Male: (a) wing; (b) fore, mid, hind legs (left to right); (c) head, anterior view. Female: (d) wing; (e) head, anterior view; (f) fore, mid, hind, tarsomeres (bottom to top); (g) palpus; (h) apex of female abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5178501" httpUri="https://zenodo.org/record/5178501/files/figure.png" pageId="4" pageNumber="512">Figure 6c</figureCitation>
|
||
); paramere stem nearly straight basally, expanded distally in inner portion, distal portion anteromesally directed abruptly curved to tip (
|
||
<figureCitation id="27C92A2DFFC7A6042728C91CFE60FABB" box="[359,469,1286,1310]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="4" pageNumber="512">Figure 9d</figureCitation>
|
||
); aedeagus rectangular, basal arch extending to 0.34–0.42 of total length (
|
||
<figureCitation id="27C92A2DFFC7A6042720C933FE62FAE4" box="[367,471,1321,1345]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="4" pageNumber="512">Figure 9c</figureCitation>
|
||
)......................................... .........................................
|
||
<taxonomicName id="78F24D2BFFC7A60425F7C932FB97FAE4" authority="Santarém & Borkent & Spinelli & Felippe-Bauer, 2018" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[952,1058,1320,1345]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="litorale" status="sp. nov.">
|
||
<emphasis id="8D86EABAFFC7A60425F7C932FB97FAE4" box="[952,1058,1320,1345]" italics="true" pageId="4" pageNumber="512">D. litorale</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="96B557C1FFC7A6042264C933FB31FAE4" box="[1067,1156,1321,1345]" pageId="4" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
<paragraph id="BF4D36A8FFC7A60426EFC974FC27FA23" blockId="4.[160,914,1390,1414]" box="[160,914,1390,1414]" pageId="4" pageNumber="512">
|
||
<emphasis id="8D86EABAFFC7A60426EFC974FC27FA23" bold="true" box="[160,914,1390,1414]" pageId="4" pageNumber="512">
|
||
Key to females of the
|
||
<taxonomicName id="78F24D2BFFC7A60427E0C974FD54FA23" ID-CoL="37HPF" baseAuthorityName="Lutz" baseAuthorityYear="1914" box="[431,737,1390,1414]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="multilineata">
|
||
<emphasis id="8D86EABAFFC7A60427E0C974FD54FA23" bold="true" box="[431,737,1390,1414]" italics="true" pageId="4" pageNumber="512">Downeshelea multilineata</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="BF4D36A8FFC7A60426EFC9BBFE89FA5B" blockId="4.[160,1155,1441,1534]" pageId="4" pageNumber="512">
|
||
Female members of the multilineata group can only be recognised as such if associated with males of the group. The females of
|
||
<taxonomicName id="78F24D2BFFC7A6042435C9D8FD41FA7E" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[634,756,1474,1499]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="moravia">
|
||
<emphasis id="8D86EABAFFC7A6042435C9D8FD41FA7E" box="[634,756,1474,1499]" italics="true" pageId="4" pageNumber="512">D. moravia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="78F24D2BFFC7A604254BC9D8FC16FA79" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[772,931,1474,1500]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="marambaia">
|
||
<emphasis id="8D86EABAFFC7A604254BC9D8FC16FA79" box="[772,931,1474,1500]" italics="true" pageId="4" pageNumber="512">D. marambaia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="78F24D2BFFC7A60425AEC9D9FB36FA79" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[993,1155,1475,1500]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="casimirensis">
|
||
<emphasis id="8D86EABAFFC7A60425AEC9D9FB36FA79" box="[993,1155,1475,1500]" italics="true" pageId="4" pageNumber="512">D. casimirensis</emphasis>
|
||
</taxonomicName>
|
||
are unknown.
|
||
</paragraph>
|
||
<key id="CB636199FFC7A60526EFCA21FB31FEF1" lastPageId="5" lastPageNumber="513" pageId="4" pageNumber="512">
|
||
<keyStep id="04062E0DFFC7A60426EFCA21FB36F9D3" pageId="4" pageNumber="512">
|
||
<paragraph id="BF4D36A8FFC7A60426EFCA21FB31F9F6" blockId="4.[160,1156,1594,1723]" box="[160,1156,1594,1619]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFCA21FB31F9F6" box="[160,1156,1594,1619]" pageId="4" pageNumber="512">
|
||
1. Legs pale brown; hind tibia darker apically and on proximal third..... .....
|
||
<taxonomicName id="78F24D2BFFC7A6042582CA20FB92F9F6" authority="Santarém & Borkent & Spinelli & Felippe-Bauer, 2018" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[973,1063,1594,1619]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="jarina" status="sp. nov.">
|
||
<emphasis id="8D86EABAFFC7A6042582CA20FB92F9F6" box="[973,1063,1594,1619]" italics="true" pageId="4" pageNumber="512">D. jarina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="96B557C1FFC7A6042260CA21FB31F9F6" box="[1071,1156,1595,1619]" pageId="4" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="BF4D36A8FFC7A60426EFCA44FB36F9D3" blockId="4.[160,1156,1594,1723]" box="[160,1155,1630,1654]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFCA44FB36F9D3" box="[160,1155,1630,1654]" pageId="4" pageNumber="512">– Legs brown; hind tibia dark brown.................................................................................................. 2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="BF4D36A8FFC7A60426EFCA9BFB36F91E" blockId="4.[160,1156,1594,1723]" pageId="4" pageNumber="512">
|
||
<keyLead id="0403959DFFC7A60426EFCA9BFB36F91E" pageId="4" pageNumber="512">
|
||
<keyStep id="04062E0DFFC7A60426EFCA9BFB36F91E" pageId="4" pageNumber="512">
|
||
2. Halter pale; mandible with 10 teeth......................... .........................
|
||
<taxonomicName id="78F24D2BFFC7A6042534CA9AFB96F93C" authority="Santarém & Borkent & Spinelli & Felippe-Bauer, 2018" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[891,1059,1664,1689]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="512" phylum="Arthropoda" rank="species" species="costaricensis" status="sp. nov.">
|
||
<emphasis id="8D86EABAFFC7A6042534CA9AFB96F93C" box="[891,1059,1664,1689]" italics="true" pageId="4" pageNumber="512">D. costaricensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="96B557C1FFC7A6042264CA9BFB31F93C" box="[1067,1156,1665,1689]" pageId="4" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Halter brown; mandible with 11–12 teeth.................................................................................... 3
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<keyStep id="04062E0DFFC6A60526EFCC94FB36FF6C" pageId="5" pageNumber="513">
|
||
<paragraph id="BF4D36A8FFC6A60526EFCC94FB31FF03" blockId="5.[160,1156,141,340]" box="[160,1156,141,166]" pageId="5" pageNumber="513">
|
||
<keyLead id="0403959DFFC6A60526EFCC94FB31FF03" box="[160,1156,141,166]" pageId="5" pageNumber="513">
|
||
3.Midtibia paler basally ............................................ ............................................
|
||
<taxonomicName id="78F24D2BFFC6A60525E7CC97FB31FF03" ID-CoL="37HPF" authority="Lutz" authorityName="Lutz" box="[936,1156,141,166]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="513" phylum="Arthropoda" rank="species" species="multilineata">
|
||
<emphasis id="8D86EABAFFC6A60525E7CC97FBFEFF03" box="[936,1099,141,166]" italics="true" pageId="5" pageNumber="513">D. multilineata</emphasis>
|
||
Lutz
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="BF4D36A8FFC6A60526EFCCABFB36FF6C" blockId="5.[160,1156,141,340]" box="[160,1155,177,201]" pageId="5" pageNumber="513">
|
||
<keyLead id="0403959DFFC6A60526EFCCABFB36FF6C" box="[160,1155,177,201]" pageId="5" pageNumber="513">– Midtibia brown ........................................................................................................................................... 4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="04062E0DFFC6A60526EFCCCEFB31FEF1" pageId="5" pageNumber="513">
|
||
<paragraph id="BF4D36A8FFC6A60526EFCCCEFB31FEAB" blockId="5.[160,1156,141,340]" pageId="5" pageNumber="513">
|
||
<keyLead id="0403959DFFC6A60526EFCCCEFB31FEAB" pageId="5" pageNumber="513">
|
||
4.Hind femur brown; midtarsomere 1 with six to nine ventral spines .................................. .....................................................................................................................................
|
||
<taxonomicName id="78F24D2BFFC6A60525EDCCEFFBAAFEAB" authority="Santarém & Borkent & Spinelli & Felippe-Bauer, 2018" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[930,1055,245,270]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="513" phylum="Arthropoda" rank="species" species="eclectica" status="sp. nov.">
|
||
<emphasis id="8D86EABAFFC6A60525EDCCEFFBAAFEAB" box="[930,1055,245,270]" italics="true" pageId="5" pageNumber="513">D. eclectica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="96B557C1FFC6A6052265CCECFB31FEAB" box="[1066,1156,246,270]" pageId="5" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="BF4D36A8FFC6A60526EFCD03FB31FEF1" blockId="5.[160,1156,141,340]" pageId="5" pageNumber="513">
|
||
<keyLead id="0403959DFFC6A60526EFCD03FB31FEF1" pageId="5" pageNumber="513">
|
||
– Hind femur darker subapically; midtarsomere 1 with four to five ventral spines............. .........................................................................................................................................
|
||
<taxonomicName id="78F24D2BFFC6A60525F7CD21FB97FEF1" authority="Santarém & Borkent & Spinelli & Felippe-Bauer, 2018" authorityName="Santarém & Borkent & Spinelli & Felippe-Bauer" authorityYear="2018" box="[952,1058,315,340]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="513" phylum="Arthropoda" rank="species" species="litorale" status="sp. nov.">
|
||
<emphasis id="8D86EABAFFC6A60525F7CD21FB97FEF1" box="[952,1058,315,340]" italics="true" pageId="5" pageNumber="513">D. litorale</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="96B557C1FFC6A6052264CD26FB31FEF1" box="[1067,1156,316,340]" pageId="5" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |