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<document id="14929BF6F0BDD535BB30BC0FB935AA39" ID-DOI="10.11646/phytotaxa.675.3.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="14522048" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734562248782" checkinUser="felipe" docAuthor="Rajeshkumar, Kunhiraman C., Paraparath, Sruthi O., Ashtamoorthy, Sreejith K., Robi, Aloor J., Gautam, Ajay Kumar, Verma, Rajnish Kumar, Jeewon, Rajesh, Ansil, Parayelil A., Karunarathna, Samantha C. &amp; Kavileveettil, Sankaran" docDate="2024" docId="03B1F245FFC2FF80FF370CCAFBD4D525" docLanguage="en" docName="phytotaxa.675.3.233-246.pdf" docOrigin="Phytotaxa 675 (3)" docSource="https://doi.org/10.11646/phytotaxa.675.3.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Striaticonidium kfriense Rajeshk., O. P. Sruthi, S. K. Ashtamoorthy &amp; K. V. Sankaran 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="239" masterDocId="FF888A3DFFC7FF86FFBF0F1EFF84D24A" masterDocTitle="Morpho-molecular systematics of Striaticonidium kfriense (Stachybotryaceae), a new synnematous species from the Western Ghats, India" masterLastPageNumber="246" masterPageNumber="233" pageNumber="238" updateTime="1734567033511" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B79BEF5EB6BD2D6351ECAD4D8783DECF">Morpho-molecular systematics of Striaticonidium kfriense (Stachybotryaceae), a new synnematous species from the Western Ghats, India</mods:title>
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<mods:namePart id="D668F9AA39FD2EA9DC7ECEC5FE53EBC3">Gautam, Ajay Kumar</mods:namePart>
<mods:affiliation id="3F59426F06383F2B16E8C9FB8BD62EDC">Patanjali Herbal Research Department, Patanjali Research Institute, Haridwar 249405, Uttarakhand, India</mods:affiliation>
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<mods:namePart id="C81DB2837A9BD1995F92AE292BE511F5">Karunarathna, Samantha C.</mods:namePart>
<mods:affiliation id="5E2D88FFA6696F6744C1C261C0628980">Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan, 655011, P. R. China</mods:affiliation>
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<mods:namePart id="316255990CC461EB882AD5C99D27F373">Kavileveettil, Sankaran</mods:namePart>
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<mods:title id="18198F43F50D9A6108B7F918D9A8D63D">Phytotaxa</mods:title>
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<mods:part id="449FBC7A0E99D8D80CC391F300E9B5E8">
<mods:date id="BD71300AC19CC6D3C55215E25E19A019">2024</mods:date>
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<mods:number id="AB13A265CD55C89A27D96BDE2F0D41B4">2024-12-04</mods:number>
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<mods:number id="E2D2A5A8E5E9B9395B06B16BE1ECF8F8">675</mods:number>
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<mods:number id="DEB4EF938B50ADF6F0EAF163F066A6A2">3</mods:number>
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<mods:url id="AE2F35FCB63C1F06752ABCD7EFC6E833">https://doi.org/10.11646/phytotaxa.675.3.3</mods:url>
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<mods:identifier id="39545396F74027898A6402F00012E74A" type="DOI">10.11646/phytotaxa.675.3.3</mods:identifier>
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<taxonomicName id="4C1838D0FFC2FF83FF370CCAFBCED1A4" authority="Rajeshk., O.P. Sruthi, S. K. Ashtamoorthy &amp; K. V. Sankaran" authorityName="Rajeshk., O. P. Sruthi, S. K. Ashtamoorthy &amp; K. V. Sankaran" authorityYear="2024" box="[136,1098,980,1007]" class="Sordariomycetes" family="Stachybotryaceae" genus="Striaticonidium" kingdom="Fungi" order="Hypocreales" pageId="5" pageNumber="238" phylum="Ascomycota" rank="species" species="kfriense" status="sp. nov.">
<emphasis id="B96C9F41FFC2FF83FF370CCAFE25D1A5" bold="true" box="[136,417,980,1007]" italics="true" pageId="5" pageNumber="238">Striaticonidium kfriense</emphasis>
Rajeshk., O.P. Sruthi, S. K. Ashtamoorthy &amp; K. V. Sankaran
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</emphasis>
(Figures 3,4)
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</paragraph>
<paragraph id="8BA74353FFC2FF83FF370CE6FDA7D659" blockId="5.[136,1452,1016,2015]" box="[136,547,1016,1043]" pageId="5" pageNumber="238">Index Fungorum number: IF 902552</paragraph>
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<emphasis id="B96C9F41FFC2FF83FF020B02FEBCD67D" bold="true" box="[189,312,1052,1079]" italics="true" pageId="5" pageNumber="238">Etymology</emphasis>
:—
</emphasis>
The specific
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epithet
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,
</emphasis>
named after the
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Forest Research Institute (KFRI),
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, on the occasion of the golden jubilee celebration of the Institution and in recognition of contribution of the institute to the Indian mycology, tropical biodiversity inventory and conservation.
</paragraph>
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:—
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.
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,
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,
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Division
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,
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,
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, on a decaying leaf of
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<emphasis id="B96C9F41FFC2FF83FF660BB2FE6FD68C" box="[217,491,1196,1222]" italics="true" pageId="5" pageNumber="238">Ochlandra travancorica</emphasis>
</taxonomicName>
,
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</date>
,
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<collectorName id="26ED2685FFC2FF83FD4F0BB2FCB0D68C" box="[752,820,1196,1222]" pageId="5" pageNumber="238">Sruthi</collectorName>
and
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,
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, ex-type culture
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.
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<paragraph id="8BA74353FFC2FF83FF020BEAFBEBD7D4" blockId="5.[136,1452,1016,2015]" pageId="5" pageNumber="238">
Diagnosis: The synnemata of
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<emphasis id="B96C9F41FFC2FF83FDB20BEAFC98D744" box="[525,796,1268,1294]" italics="true" pageId="5" pageNumber="238">Striaticonidium kfriense</emphasis>
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are longer and wider (144515 × 2297 µm) compared to the morphologically similar
<taxonomicName id="4C1838D0FFC2FF83FE720A06FDF3D778" box="[461,631,1304,1330]" class="Sordariomycetes" family="Stachybotryaceae" genus="Striaticonidium" kingdom="Fungi" order="Hypocreales" pageId="5" pageNumber="238" phylum="Ascomycota" rank="species" species="synnematum">
<emphasis id="B96C9F41FFC2FF83FE720A06FDF3D778" box="[461,631,1304,1330]" italics="true" pageId="5" pageNumber="238">S. synnematum</emphasis>
</taxonomicName>
(5085 × 3585 μm). The conidiogenous cells were slightly larger and with verrucose apex in
<taxonomicName id="4C1838D0FFC2FF83FE280A22FD8AD71C" authorityName="Rajeshk., O. P. Sruthi, S. K. Ashtamoorthy &amp; K. V. Sankaran" authorityYear="2024" box="[407,526,1340,1366]" class="Sordariomycetes" family="Stachybotryaceae" genus="Striaticonidium" kingdom="Fungi" order="Hypocreales" pageId="5" pageNumber="238" phylum="Ascomycota" rank="species" species="kfriense">
<emphasis id="B96C9F41FFC2FF83FE280A22FD8AD71C" box="[407,526,1340,1366]" italics="true" pageId="5" pageNumber="238">S. kfriense</emphasis>
</taxonomicName>
(1736 × 1.52 μm) in both
<emphasis id="B96C9F41FFC2FF83FCDB0A23FC33D71C" box="[868,951,1341,1366]" italics="true" pageId="5" pageNumber="238">in vitro</emphasis>
and
<emphasis id="B96C9F41FFC2FF83FC4D0A23FBC4D71C" box="[1010,1088,1341,1366]" italics="true" pageId="5" pageNumber="238">in vivo</emphasis>
conditions than
<taxonomicName id="4C1838D0FFC2FF83FABE0A22FA2FD71C" box="[1281,1451,1340,1366]" class="Sordariomycetes" family="Stachybotryaceae" genus="Striaticonidium" kingdom="Fungi" order="Hypocreales" pageId="5" pageNumber="238" phylum="Ascomycota" rank="species" species="synnematum">
<emphasis id="B96C9F41FFC2FF83FABE0A22FA2FD71C" box="[1281,1451,1340,1366]" italics="true" pageId="5" pageNumber="238">S. synnematum</emphasis>
</taxonomicName>
(1830 × 12 μm
<emphasis id="B96C9F41FFC2FF83FEEB0A7FFE25D730" box="[340,417,1377,1402]" italics="true" pageId="5" pageNumber="238">in vivo</emphasis>
and 510 × 12 μm
<emphasis id="B96C9F41FFC2FF83FD320A7FFD5AD730" box="[653,734,1377,1402]" italics="true" pageId="5" pageNumber="238">in vitro</emphasis>
). Likewise, conidia were larger and with a prominent truncated base
<emphasis id="B96C9F41FFC2FF83FF7E0A9BFE96D7D4" box="[193,274,1413,1438]" italics="true" pageId="5" pageNumber="238">in vitro</emphasis>
in
<taxonomicName id="4C1838D0FFC2FF83FE8A0A9AFE2ED7D4" authorityName="Rajeshk., O. P. Sruthi, S. K. Ashtamoorthy &amp; K. V. Sankaran" authorityYear="2024" box="[309,426,1412,1438]" class="Sordariomycetes" family="Stachybotryaceae" genus="Striaticonidium" kingdom="Fungi" order="Hypocreales" pageId="5" pageNumber="238" phylum="Ascomycota" rank="species" species="kfriense">
<emphasis id="B96C9F41FFC2FF83FE8A0A9AFE2ED7D4" box="[309,426,1412,1438]" italics="true" pageId="5" pageNumber="238">S. kfriense</emphasis>
</taxonomicName>
(612 × 24 μm) compared to
<taxonomicName id="4C1838D0FFC2FF83FCB70A9AFC34D7D4" box="[776,944,1412,1438]" class="Sordariomycetes" family="Stachybotryaceae" genus="Striaticonidium" kingdom="Fungi" order="Hypocreales" pageId="5" pageNumber="238" phylum="Ascomycota" rank="species" species="synnematum">
<emphasis id="B96C9F41FFC2FF83FCB70A9AFC34D7D4" box="[776,944,1412,1438]" italics="true" pageId="5" pageNumber="238">S. synnematum</emphasis>
</taxonomicName>
(78 × 23 μm).
</paragraph>
<paragraph id="8BA74353FFC2FF83FF020AB6FDACD561" blockId="5.[136,1452,1016,2015]" pageId="5" pageNumber="238">
Saprobic on decaying leaf of
<emphasis id="B96C9F41FFC2FF83FE450AB6FC94D788" box="[506,784,1448,1474]" italics="true" pageId="5" pageNumber="238">
<taxonomicName id="4C1838D0FFC2FF83FE450AB6FC8FD788" box="[506,779,1448,1474]" class="Liliopsida" family="Poaceae" genus="Ochlandra" kingdom="Plantae" order="Poales" pageId="5" pageNumber="238" phylum="Tracheophyta" rank="species" species="travancorica">Ochlandra travancorica</taxonomicName>
.
</emphasis>
Sexual morph:Undetermined.Asexual morph:
<emphasis id="B96C9F41FFC2FF83FAA60AB6FA2FD788" box="[1305,1451,1448,1474]" italics="true" pageId="5" pageNumber="238">Conidiomata</emphasis>
synnematous, solitary, 144515 × 2297 µm (
<emphasis id="B96C9F41FFC2FF83FD220AD3FD2ED7AC" box="[669,682,1485,1510]" italics="true" pageId="5" pageNumber="238"></emphasis>
= 337 × 19.8 µm, n = 17); synnemata head 4397 × 57115 µm, cylindrical, broadened towards apex, unbranched, consisting of bundles of parallel, longitudinal, closely compacted septate hyphae, terminating in whorls of conidiogenous cells enclosed by marginal thick-walled hyphae in a spirally twisted manner.
<emphasis id="B96C9F41FFC2FF83FEFC0926FD82D418" box="[323,518,1592,1618]" italics="true" pageId="5" pageNumber="238">Marginal hyphae</emphasis>
2.22.8 µm wide (
<emphasis id="B96C9F41FFC2FF83FD5E0927FD6AD418" box="[737,750,1593,1618]" italics="true" pageId="5" pageNumber="238"></emphasis>
= 2.5 µm, n = 30), branched or unbranched, sinuous, hyaline to olivaceous green, verrucose with a curved blunt apex.
<emphasis id="B96C9F41FFC2FF83FC9B0942FCF0D43C" box="[804,884,1628,1654]" italics="true" pageId="5" pageNumber="238">Stroma</emphasis>
well-developed of
<emphasis id="B96C9F41FFC2FF83FBE80943FA9BD43C" box="[1111,1311,1628,1654]" italics="true" pageId="5" pageNumber="238">textura globulosa</emphasis>
and
<emphasis id="B96C9F41FFC2FF83FAE30943FF77D4D0" italics="true" pageId="5" pageNumber="238">textura angularis</emphasis>
.
<emphasis id="B96C9F41FFC2FF83FEBE099EFE22D4D0" box="[257,422,1664,1690]" italics="true" pageId="5" pageNumber="238">Conidiophores</emphasis>
terminating in whorls of 24 conidiogenous cells.
<emphasis id="B96C9F41FFC2FF83FC64099EFB3AD4D0" box="[987,1214,1664,1690]" italics="true" pageId="5" pageNumber="238">Conidiogenous cells</emphasis>
phialidic, cylindrical to subcylindrical, hyaline, smooth or slightly verrucose towards the apex, 1736 × 1.52 μm (
<emphasis id="B96C9F41FFC2FF83FB0409BBFB43D4F4" box="[1211,1223,1701,1726]" italics="true" pageId="5" pageNumber="238"></emphasis>
= 25 × 1.9 µm, n = 30),
<emphasis id="B96C9F41FFC2FF83FF0409D7FE82D4A8" box="[187,262,1737,1762]" italics="true" pageId="5" pageNumber="238">in vivo</emphasis>
with conspicuous collarettes and periclinal thickenings.
<emphasis id="B96C9F41FFC2FF83FCC109D6FC5CD4A8" box="[894,984,1736,1762]" italics="true" pageId="5" pageNumber="238">Conidia</emphasis>
aseptate, longitudinally striate, hyaline to olivaceous green, fusiform to ellipsoidal, 612 × 24 μm (
<emphasis id="B96C9F41FFC2FF83FCB309F3FC9DD54C" box="[780,793,1773,1798]" italics="true" pageId="5" pageNumber="238"></emphasis>
= 8.5 × 3 μm), with a distinct hilum, truncated base is more prominent in conidia formed
<emphasis id="B96C9F41FFC2FF83FE6F080FFDA5D560" box="[464,545,1809,1834]" italics="true" pageId="5" pageNumber="238">in vitro</emphasis>
.
</paragraph>
<paragraph id="8BA74353FFC2FF83FF02082AFD6DD595" blockId="5.[136,1452,1016,2015]" pageId="5" pageNumber="238">
<emphasis id="B96C9F41FFC2FF83FF02082AFE5CD504" bold="true" box="[189,472,1844,1870]" pageId="5" pageNumber="238">Culture characteristics:</emphasis>
—Colonies on MEA at 25 ± 2 ºC after 7 d fast growing,
<quantity id="4CE0EEB6FFC2FF83FBF9082AFB3AD505" box="[1094,1214,1844,1871]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="5" pageNumber="238" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">2832 mm</quantity>
diam., initially white (1
<collectionCode id="ED09DB96FFC2FF83FF220846FF3CD538" box="[157,184,1880,1906]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="5" pageNumber="238" type="Herbarium">A</collectionCode>
1), when mature colonies are light grey (1
<collectionCode id="ED09DB96FFC2FF83FD3B0846FD18D538" box="[644,668,1880,1906]" country="Denmark" name="University of Copenhagen" pageId="5" pageNumber="238" type="Herbarium">C</collectionCode>
1) to greenish grey (1
<collectionCode id="ED09DB96FFC2FF83FC3A0846FC1ED538" box="[901,922,1880,1906]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="5" pageNumber="238" type="Herbarium">B</collectionCode>
2) when mature, reverse yellowish white (4
<collectionCode id="ED09DB96FFC2FF83FAC00846FA11D538" box="[1407,1429,1880,1906]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="5" pageNumber="238" type="Herbarium">A</collectionCode>
2) to white (3
<collectionCode id="ED09DB96FFC2FF83FEBE0862FE9FD5DC" box="[257,283,1916,1942]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="5" pageNumber="238" type="Herbarium">A</collectionCode>
1). Margin regular, brownish grey to grey (6D2 to 6
<collectionCode id="ED09DB96FFC2FF83FCE90862FCEFD5DC" box="[854,875,1916,1942]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="5" pageNumber="238" type="Herbarium">E</collectionCode>
1), slightly umbonate at center, greenish grey (1
<collectionCode id="ED09DB96FFC2FF83FAC80862FA09D5DC" box="[1399,1421,1916,1942]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="5" pageNumber="238" type="Herbarium">B</collectionCode>
2). Floccose colony with diurnal pattern. Sporulating on media and forming synnemata on the surface of colonies, covered by slimy herbage to olivaceous green conidial masses.
</paragraph>
<caption id="DF6713DBFFC1FF80FF370926FB09D4DC" ID-DOI="http://doi.org/10.5281/zenodo.14522056" ID-Zenodo-Dep="14522056" httpUri="https://zenodo.org/record/14522056/files/figure.png" pageId="6" pageNumber="239" startId="6.[136,229,1592,1614]" targetBox="[151,1436,187,1554]" targetPageId="6" targetType="figure">
<paragraph id="8BA74353FFC1FF80FF370926FB09D4DC" blockId="6.[136,1451,1592,1687]" pageId="6" pageNumber="239">
<emphasis id="B96C9F41FFC1FF80FF370926FF78D404" bold="true" box="[136,252,1592,1614]" pageId="6" pageNumber="239">FIGURE 3.</emphasis>
<taxonomicName id="4C1838D0FFC1FF80FF400927FE61D404" authorityName="Rajeshk., O. P. Sruthi, S. K. Ashtamoorthy &amp; K. V. Sankaran" authorityYear="2024" box="[255,485,1593,1614]" class="Sordariomycetes" family="Stachybotryaceae" genus="Striaticonidium" kingdom="Fungi" order="Hypocreales" pageId="6" pageNumber="239" phylum="Ascomycota" rank="species" species="kfriense">
<emphasis id="B96C9F41FFC1FF80FF400927FE61D404" box="[255,485,1593,1614]" italics="true" pageId="6" pageNumber="239">Striaticonidium kfriense</emphasis>
</taxonomicName>
(AMH 10769).AC Synnemata on natural substrate. D Upper portion of Synnemata. E Conidiogenous cells releasing conidia. F Conidiogenous cell with conspicuous collarette and periclinal thickening. G Striated conidia. H Colonies on MEA obverse and reverse. Scale bars: A = 100 μm. B = 200 μm. C = 100 μm. D = 20 μm. E, F, G = 10 μm.
</paragraph>
</caption>
<paragraph id="8BA74353FFC1FF80FF0209F4FD59D506" blockId="6.[136,1452,1769,1903]" pageId="6" pageNumber="239">
Notes: Phylogenetic analyses delineated
<taxonomicName id="4C1838D0FFC1FF80FD2F09F7FC83D549" authorityName="Rajeshk., O. P. Sruthi, S. K. Ashtamoorthy &amp; K. V. Sankaran" authorityYear="2024" box="[656,775,1769,1795]" class="Sordariomycetes" family="Stachybotryaceae" genus="Striaticonidium" kingdom="Fungi" order="Hypocreales" pageId="6" pageNumber="239" phylum="Ascomycota" rank="species" species="kfriense">
<emphasis id="B96C9F41FFC1FF80FD2F09F7FC83D549" box="[656,775,1769,1795]" italics="true" pageId="6" pageNumber="239">S. kfriense</emphasis>
</taxonomicName>
as a new lineage sister to
<taxonomicName id="4C1838D0FFC1FF80FB8209F7FB63D549" box="[1085,1255,1769,1795]" class="Sordariomycetes" family="Stachybotryaceae" genus="Striaticonidium" kingdom="Fungi" order="Hypocreales" pageId="6" pageNumber="239" phylum="Ascomycota" rank="species" species="synnematum">
<emphasis id="B96C9F41FFC1FF80FB8209F7FB63D549" box="[1085,1255,1769,1795]" italics="true" pageId="6" pageNumber="239">S. synnematum</emphasis>
</taxonomicName>
with a statistical support of ML-BS = 67% and PP = 0.95, respectively. In addition, PHI of 0.99, indicates no significant genetic recombination between
<taxonomicName id="4C1838D0FFC1FF80FE2A082FFD8ED501" authorityName="Rajeshk., O. P. Sruthi, S. K. Ashtamoorthy &amp; K. V. Sankaran" authorityYear="2024" box="[405,522,1841,1867]" class="Sordariomycetes" family="Stachybotryaceae" genus="Striaticonidium" kingdom="Fungi" order="Hypocreales" pageId="6" pageNumber="239" phylum="Ascomycota" rank="species" species="kfriense">
<emphasis id="B96C9F41FFC1FF80FE2A082FFD8ED501" box="[405,522,1841,1867]" italics="true" pageId="6" pageNumber="239">S. kfriense</emphasis>
</taxonomicName>
and its allied taxa.
</paragraph>
<paragraph id="8BA74353FFC1FF80FF02084BFBD4D525" blockId="6.[136,1452,1769,1903]" box="[189,1104,1877,1903]" pageId="6" pageNumber="239">
<collectionCode id="ED09DB96FFC1FF80FF02084BFF54D525" box="[189,208,1877,1903]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="239" type="Herbarium">A</collectionCode>
synopsis of the five species reported under
<taxonomicName id="4C1838D0FFC1FF80FD04084BFCEDD525" authorityName="L.Lombard &amp; Crous" authorityYear="2016" box="[699,873,1877,1903]" class="Sordariomycetes" family="Stachybotryaceae" genus="Striaticonidium" kingdom="Fungi" order="Hypocreales" pageId="6" pageNumber="239" phylum="Ascomycota" rank="genus">
<emphasis id="B96C9F41FFC1FF80FD04084BFCEDD525" box="[699,873,1877,1903]" italics="true" pageId="6" pageNumber="239">Striaticonidium</emphasis>
</taxonomicName>
is shown in
<tableCitation id="C69A76E8FFC1FF80FC48084BFBC9D525" box="[1015,1101,1877,1903]" captionStart="TABLE 2" captionStartId="8.[136,216,499,523]" captionText="TABLE 2. Synopsis of the comparative morphology of the species in Striaticonidium." pageId="6" pageNumber="239">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>