727 lines
127 KiB
XML
727 lines
127 KiB
XML
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<mods:title id="F4AE3AF66698F5E603100D0A52DB5AFE">A review of the records of Pachycheles rugimanus A. Milne-Edwards, 1880 (Crustacea: Decapoda: Anomura: Porcellanidae) from Brazil, with the description of a new species</mods:title>
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<mods:namePart id="1311F5CB4D9974B2718DA3C06CBB979D">Ferreira, Luciane Augusto De Azevedo</mods:namePart>
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<mods:namePart id="06CCAF485F94923535B61ED359DF404F">Tavares, Marcos</mods:namePart>
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<emphasis id="B8D8EAA3FFD8FFB341C7FDEDFE1BFD9C" bold="true" box="[151,496,546,572]" pageId="1" pageNumber="186">
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<taxonomicName id="4DAC4D32FFD8FFB341C7FDEDFE67FD9C" ID-CoL="ed3e9c81-eb9f-409e-ac83-ded6704f39a4" authority="Ferreira & Tavares, 2019" authorityName="Ferreira & Tavares" authorityYear="2019" box="[151,396,546,572]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="coelhoi" status="sp. nov.">
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<emphasis id="B8D8EAA3FFD8FFB341C7FDEDFE67FD9C" bold="true" box="[151,396,546,572]" italics="true" pageId="1" pageNumber="186">Pachycheles coelhoi</emphasis>
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<taxonomicNameLabel id="A3EB57D8FFD8FFB340C3FDEDFE1BFD9C" box="[403,496,546,572]" pageId="1" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C2B6653AFFD8FFB341C7FD8AFE2DFDFE" box="[151,454,581,606]" pageId="1" pageNumber="186" type="description">
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<paragraph id="8A1336B1FFD8FFB341C7FD8AFE2DFDFE" blockId="1.[151,496,546,606]" box="[151,454,581,606]" pageId="1" pageNumber="186">
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(
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<figureCitation id="12972A34FFD8FFB341CFFD8AFF0CFDFE" box="[159,231,581,606]" captionStart="FIGURE 1" captionStartId="3.[151,250,1888,1910]" captionTargetBox="[329,1288,217,1862]" captionTargetId="figure@3.[208,1381,195,1871]" captionTargetPageId="3" captionText="FIGURE 1. Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (A) carapace, dorsal view. (B) frontal-orbital region of carapace, frontal view. (C) carapace, lateral view, emphasizing the branchiostegite. (D) right antennule, ventral view. (E) right antennal peduncle, dorsal view (flagellum omitted). (F) left Mxp 3, ventral view (exopod flagellum not shown). (G) left minor P1, dorsal view. (H) right major P1, dorsal view. (I) right P2, lateral view. (J) telson, external view. (K) thoracic sternum, ventral view. Setae all omitted. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2599189/files/figure.png" pageId="1" pageNumber="186">Figs. 1</figureCitation>
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A–K, 2A–C, 4A,B)
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</paragraph>
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</subSubSection>
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<subSubSection id="C2B6653AFFD8FFB341C7FD49FAD6FD1B" pageId="1" pageNumber="186" type="reference_group">
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<paragraph id="8A1336B1FFD8FFB341C7FD49FAD6FD1B" blockId="1.[151,1436,646,699]" pageId="1" pageNumber="186">
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<treatmentCitationGroup id="AABC119FFFD8FFB341C7FD49FAD6FD1B" pageId="1" pageNumber="186">
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<taxonomicName id="4DAC4D32FFD8FFB341C7FD49FE62FD3D" ID-CoL="2303263a-0456-4070-83d3-5f1c7275e6cc" authorityName="A. Milne-Edwards" authorityYear="1880" box="[151,393,646,669]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="rugimanus">
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<emphasis id="B8D8EAA3FFD8FFB341C7FD49FE62FD3D" box="[151,393,646,669]" italics="true" pageId="1" pageNumber="186">Pachycheles rugimanus</emphasis>
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</taxonomicName>
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.
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<emphasis id="B8D8EAA3FFD8FFB340C0FD47FE4DFD3D" box="[400,422,648,669]" italics="true" pageId="1" pageNumber="186">—</emphasis>
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Coelho 1964: 255;
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<treatmentCitation id="0B0D10A0FFD8FFB3433FFD49FC94FD3C" author="Veloso, V. G. & Melo, G. A. S." box="[623,895,646,668]" page="173" pageId="1" pageNumber="186" year="1993">
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<bibRefCitation id="EE3D4B40FFD8FFB3433FFD49FCA7FD3C" author="Veloso, V. G. & Melo, G. A. S." box="[623,844,646,668]" pageId="1" pageNumber="186" pagination="171 - 186" refId="ref4382" refString="Veloso, V. G. & Melo, G. A. S. (1993) Taxonomia e distribuicao da familia Porcellanidae (Crustacea: Decapoda: Anomura) no litoral brasileiro. Iheringia, Serie Zoologia, 75, 171 - 186." type="journal article" year="1993">Veloso & Melo 1993</bibRefCitation>
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: 173
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</treatmentCitation>
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, 178 (in part);
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<treatmentCitation id="0B0D10A0FFD8FFB3454DFD49FB2EFD3C" author="Melo, G. A. S." box="[1053,1221,646,668]" page="227" pageId="1" pageNumber="186" year="1999">
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<bibRefCitation id="EE3D4B40FFD8FFB3454DFD49FB7AFD3C" author="Melo, G. A. S." box="[1053,1169,646,668]" pageId="1" pageNumber="186" refId="ref4112" refString="Melo, G. A. S. (1999) Manual de identificacao dos Crustacea Decapoda do litoral brasileiro: Anomura, Thalassinidea, Palinuridea, Astacidea. Pleiade Fapesp, Sao Paulo." type="book" year="1999">Melo 1999</bibRefCitation>
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: 227
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||
</treatmentCitation>
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||
, 242, figs. 163, 164;
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<treatmentCitation id="0B0D10A0FFD8FFB34197FD6AFE77FD1B" author="Coelho, P. A. & Almeida, O. A. & Bezerra, L. E. A. & Souza-Filho, J. F." box="[199,412,677,699]" page="1" pageId="1" pageNumber="186" year="2007">
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<bibRefCitation id="EE3D4B40FFD8FFB34197FD6AFE6DFD1B" author="Coelho, P. A. & Almeida, O. A. & Bezerra, L. E. A. & Souza-Filho, J. F." box="[199,390,677,699]" pageId="1" pageNumber="186" pagination="1 - 16" refId="ref3782" refString="Coelho, P. A., Almeida, O. A., Bezerra, L. E. A. & Souza-Filho, J. F. (2007) An updated checklist of decapod crustaceans (infraorders Astacidea, Thalassinidea, Polychelida, Palinura and Anomura) from the northern and northeastern Brazilian coast. Zootaxa, 1519, 1 - 16. https: // doi. org / 10.11646 / zootaxa. 1519.1.1" type="journal article" year="2007">
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||
Coelho
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||
<emphasis id="B8D8EAA3FFD8FFB34046FD69FEA1FD1B" box="[278,330,677,699]" italics="true" pageId="1" pageNumber="186">et al.</emphasis>
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2007
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</bibRefCitation>
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: 1
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</treatmentCitation>
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, tab 4;
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<treatmentCitation id="0B0D10A0FFD8FFB340B8FD6AFD16FD1B" ID-DOI="http://doi.org/10.5281/zenodo.3503919" ID-Zenodo-Dep="3503919" author="Ferreira, L. A. A. & Melo, G. A. S." box="[488,765,677,699]" httpUri="http://treatment.plazi.org/id/03B887D23E77FFC7FF2FE0DF68359D17" page="183" pageId="1" pageNumber="186" year="2016">
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<bibRefCitation id="EE3D4B40FFD8FFB340B8FD6AFD20FD1B" author="Ferreira, L. A. A. & Melo, G. A. S." box="[488,715,677,699]" pageId="1" pageNumber="186" pagination="175 - 194" refId="ref3860" refString="Ferreira, L. A. A. & Melo, G. A. S. (2016) Porcelain crabs from Brazil (Crustacea: Decapoda: Anomura: Porcellanidae). Zootaxa, 4092 (2), 175 - 194. https: // doi. org / 10.11646 / zootaxa. 4092.2.2" type="journal article" year="2016">Ferreira & Melo 2016</bibRefCitation>
|
||
: 183
|
||
</treatmentCitation>
|
||
. [Not
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||
<taxonomicName id="4DAC4D32FFD8FFB3426BFD6AFAD8FD1B" ID-CoL="2303263a-0456-4070-83d3-5f1c7275e6cc" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" box="[827,1331,677,699]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="rugimanus">
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||
<emphasis id="B8D8EAA3FFD8FFB3426BFD6AFBC6FD1B" box="[827,1069,677,699]" italics="true" pageId="1" pageNumber="186">Pachycheles rugimanus</emphasis>
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<bibRefCitation id="EE3D4B40FFD8FFB34563FD6AFAD8FD1B" author="Milne-Edwards, A." box="[1075,1331,677,699]" pageId="1" pageNumber="186" pagination="1 - 68" refId="ref4147" refString="Milne-Edwards, A. (1880) Reports on the results of dredging under the supervision of Alexander Agassiz, in the Gulf of Mexico and in the Caribbean Sea, 1877, 78, 79, by the U. S. Coast Survey Steamer " Blake ", Lieut. - Commander C. D. Sigsbee, U. S. N. and Commander J. R. Bartlett, U. S. N. commanding. VIII. Etudes preliminaires sur les Crustaces, 1 ere Partie. Bulletin of the Museum of Comparative Zoology, 8, 1 - 68." type="journal article" year="1880">A. Milne-Edwards, 1880</bibRefCitation>
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</taxonomicName>
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].
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</paragraph>
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<paragraph id="8A1336B1FFD8FFB341C7FD28FD0FFCE8" blockId="1.[151,1437,743,2029]" pageId="1" pageNumber="186">
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<emphasis id="B8D8EAA3FFD8FFB341C7FD28FEACFCA0" bold="true" box="[151,327,743,768]" pageId="1" pageNumber="186">Type material.</emphasis>
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<materialsCitation id="3AC43CECFFD8FFB34000FD28FBB3FC84" ID-GBIF-Occurrence="2013824446" collectionCode="III" country="Brazil" county="Cabo Orange" elevation="1971118" latitude="4.866667" location="Stn" longLatPrecision="925" longitude="-50.525" municipality="Project Geomar" pageId="1" pageNumber="186" specimenCode="MOUFPE 7491" specimenCount="1" stateProvince="Amapa" typeStatus="holotype">
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<typeStatus id="55178813FFD8FFB34000FD28FE56FCA0" box="[336,445,743,768]" pageId="1" pageNumber="186" type="holotype">Holotype</typeStatus>
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: male cl
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, cw
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(
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<specimenCode id="DA0A9ECAFFD8FFB34271FD27FC31FCA0" box="[801,986,744,768]" collectionCode="MOUFPE" country="Brazil" httpUri="http://grbio.org/cool/x21b-a7z4" name="Oceanographic Museum of the Federal University of Pernambuco" pageId="1" pageNumber="186">MOUFPE 7491</specimenCode>
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),
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<collectingCountry id="F2BB7621FFD8FFB342A0FD28FBDCFCA0" box="[1008,1079,743,768]" name="Brazil" pageId="1" pageNumber="186">Brazil</collectingCountry>
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,
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<collectingRegion id="4868F853FFD8FFB34515FD27FB72FCA0" box="[1093,1177,744,768]" country="Brazil" name="Amapa" pageId="1" pageNumber="186">Amapá</collectingRegion>
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,
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<collectingCounty id="63724E3DFFD8FFB345F6FD28FAD4FCA0" box="[1190,1343,743,768]" pageId="1" pageNumber="186">Cabo Orange</collectingCounty>
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,
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<collectingMunicipality id="6A77ACCBFFD8FFB3441DFD28FF1AFC84" pageId="1" pageNumber="186">Project Geomar</collectingMunicipality>
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<collectionCode id="ECBDAE74FFD8FFB341A8FCC2FEFDFC84" box="[248,278,781,804]" pageId="1" pageNumber="186">III</collectionCode>
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, R/V "Almirante Saldanha",
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<location id="8F73606AFFD8FFB3430EFCC3FD68FC84" LSID="urn:lsid:plazi:treatment:020587A7FFD8FFB541C7FDEDFBE4FE70:8F73606AFFD8FFB3430EFCC3FD68FC84" box="[606,643,780,804]" country="Brazil" county="Cabo Orange" latitude="4.866667" longLatPrecision="925" longitude="-50.525" municipality="Project Geomar" name="Stn" pageId="1" pageNumber="186" stateProvince="Amapa">Stn</location>
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208,
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:
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<geoCoordinate id="EF985076FFD8FFB3427EFCC3FC53FC85" box="[814,952,780,805]" direction="west" orientation="longitude" pageId="1" pageNumber="186" precision="15" value="-50.525">50°31'30"W</geoCoordinate>
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,
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<elevation id="0181D182FFD8FFB34294FCC3FBBFFC84" box="[964,1108,780,804]" metricMagnitude="6" metricUnit="m" metricValue="1.971118" pageId="1" pageNumber="186" unit="m" value="1971118.0">1971, 118 m</elevation>
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</quantity>
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.
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</materialsCitation>
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<materialsCitation id="3AC43CECFFD8FFB3450FFCC2FD0AFCE8" ID-GBIF-Occurrence="2013824447" collectionCode="MZUSP" country="Brazil" county="Cabo Orange" elevation="1971118" latitude="4.866667" location="Stn" longLatPrecision="925" longitude="-50.525" municipality="Project Geomar" pageId="1" pageNumber="186" specimenCode="MZUSP 39043" specimenCount="1" stateProvince="Amapa" typeStatus="paratype">
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<typeStatus id="55178813FFD8FFB3450FFCC2FB2EFC85" box="[1119,1221,781,805]" pageId="1" pageNumber="186" type="paratype">Paratype</typeStatus>
|
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: female cl
|
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<quantity id="4D549B54FFD8FFB3446FFCC3FA73FC84" box="[1343,1432,780,804]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="186" unit="mm" value="4.7">4.7 mm</quantity>
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||
, cw
|
||
<quantity id="4D549B54FFD8FFB341EEFCFFFEFFFCE7" box="[190,276,816,840]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="186" unit="mm" value="4.4">4.4 mm</quantity>
|
||
(
|
||
<specimenCode id="DA0A9ECAFFD8FFB34072FCFFFE38FCE8" box="[290,467,816,840]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="186">MZUSP 39043</specimenCode>
|
||
), same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8A1336B1FFD8FFB34197FC9BFEAEFC14" blockId="1.[151,1437,743,2029]" pageId="1" pageNumber="186">
|
||
<emphasis id="B8D8EAA3FFD8FFB34197FC9BFE3CFCCD" bold="true" box="[199,471,852,877]" pageId="1" pageNumber="186">Comparative material.</emphasis>
|
||
<taxonomicName id="4DAC4D32FFD8FFB3408DFC9BFBF5FCCC" authority="A. Milne-Edwards, 1880: 2" authorityName="A. Milne-Edwards" authorityPageNumber="2" authorityYear="1880" box="[477,1054,852,877]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="ackleianus">
|
||
<emphasis id="B8D8EAA3FFD8FFB3408DFC9BFD0CFCCC" box="[477,743,852,876]" italics="true" pageId="1" pageNumber="186">Pachycheles ackleianus</emphasis>
|
||
<bibRefCitation id="EE3D4B40FFD8FFB343BDFC9BFBEDFCCC" author="Milne-Edwards, A." box="[749,1030,852,877]" pageId="1" pageNumber="186" pagination="1 - 68" refId="ref4147" refString="Milne-Edwards, A. (1880) Reports on the results of dredging under the supervision of Alexander Agassiz, in the Gulf of Mexico and in the Caribbean Sea, 1877, 78, 79, by the U. S. Coast Survey Steamer " Blake ", Lieut. - Commander C. D. Sigsbee, U. S. N. and Commander J. R. Bartlett, U. S. N. commanding. VIII. Etudes preliminaires sur les Crustaces, 1 ere Partie. Bulletin of the Museum of Comparative Zoology, 8, 1 - 68." type="journal article" year="1880">A. Milne-Edwards, 1880</bibRefCitation>
|
||
:
|
||
<specimenCount id="9CAAFD38FFD8FFB34540FC9BFBF5FCCC" box="[1040,1054,852,876]" pageId="1" pageNumber="186" type="male">2</specimenCount>
|
||
</taxonomicName>
|
||
males,
|
||
<materialsCitation id="3AC43CECFFD8FFB34524FC9BFEAEFC14" ID-GBIF-Occurrence="2013824450" collectionCode="MZUSP, MOUFPE" country="Brazil" elevation="26" latitude="-6.6" location="Stn" longLatPrecision="1304" longitude="-34.783333" municipality="Projeto Algas" pageId="1" pageNumber="186" specimenCode="MZUSP 12892, MOUFPE 3082" specimenCount="26" stateProvince="Paraiba">
|
||
cl
|
||
<quantity id="4D549B54FFD8FFB345DEFC9BFB0DFCCC" box="[1166,1254,852,877]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="186" unit="mm" value="3.6">3.6 mm</quantity>
|
||
, cw
|
||
<quantity id="4D549B54FFD8FFB34448FC9BFA86FCCC" box="[1304,1389,852,876]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="186" unit="mm" value="4.2">4.2 mm</quantity>
|
||
and cl
|
||
<quantity id="4D549B54FFD8FFB341E5FCB7FEFBFC2F" box="[181,272,888,912]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="186" unit="mm" value="3.1">3.1 mm</quantity>
|
||
, cw
|
||
<quantity id="4D549B54FFD8FFB34019FCB7FE49FC2F" box="[329,418,888,912]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="186" unit="mm" value="3.8">3.8 mm</quantity>
|
||
(
|
||
<specimenCode id="DA0A9ECAFFD8FFB340E2FCB7FD8EFC30" box="[434,613,888,912]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="186">MZUSP 12892</specimenCode>
|
||
, ex-
|
||
<specimenCode id="DA0A9ECAFFD8FFB343F0FCB7FCBCFC30" box="[672,855,888,912]" collectionCode="MOUFPE" country="Brazil" httpUri="http://grbio.org/cool/x21b-a7z4" name="Oceanographic Museum of the Federal University of Pernambuco" pageId="1" pageNumber="186">MOUFPE 3082</specimenCode>
|
||
),
|
||
<collectingCountry id="F2BB7621FFD8FFB3423EFCB8FC5EFC30" box="[878,949,887,912]" name="Brazil" pageId="1" pageNumber="186">Brazil</collectingCountry>
|
||
,
|
||
<collectingRegion id="4868F853FFD8FFB34293FCB8FBF1FC30" box="[963,1050,887,912]" country="Brazil" name="Paraiba" pageId="1" pageNumber="186">Paraíba</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6A77ACCBFFD8FFB34578FCB8FB2DFC30" box="[1064,1222,887,912]" pageId="1" pageNumber="186">Projeto Algas</collectingMunicipality>
|
||
,
|
||
<location id="8F73606AFFD8FFB34585FCB7FB11FC30" LSID="urn:lsid:plazi:treatment:020587A7FFD8FFB541C7FDEDFBE4FE70:8F73606AFFD8FFB34585FCB7FB11FC30" box="[1237,1274,888,912]" country="Brazil" latitude="-6.6" longLatPrecision="1304" longitude="-34.783333" municipality="Projeto Algas" name="Stn" pageId="1" pageNumber="186" stateProvince="Paraiba">Stn</location>
|
||
86A,
|
||
<geoCoordinate id="EF985076FFD8FFB34414FCB8FA77FC30" box="[1348,1436,887,912]" direction="south" orientation="latitude" pageId="1" pageNumber="186" precision="925" value="-6.6">6°36'S–</geoCoordinate>
|
||
<geoCoordinate id="EF985076FFD8FFB341C7FC53FF12FC14" box="[151,249,924,948]" direction="west" orientation="longitude" pageId="1" pageNumber="186" precision="925" value="-34.783333">34°47'W</geoCoordinate>
|
||
,
|
||
<specimenCount id="9CAAFD38FFD8FFB34055FC53FEAEFC14" box="[261,325,924,949]" pageId="1" pageNumber="186" type="generic">
|
||
<quantity id="4D549B54FFD8FFB34055FC53FEAAFC14" box="[261,321,924,949]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="186" unit="m" value="26.0">
|
||
<elevation id="0181D182FFD8FFB34055FC53FEAAFC14" box="[261,321,924,949]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="186" unit="m" value="26.0">26 m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C2B6653AFFD8FFB34197FC0FFAA7FB74" pageId="1" pageNumber="186" type="reference_group">
|
||
<paragraph id="8A1336B1FFD8FFB34197FC0FFEACFB2C" blockId="1.[151,1437,743,2029]" pageId="1" pageNumber="186">
|
||
<treatmentCitationGroup id="AABC119FFFD8FFB34197FC0FFEACFB2C" infered="true" pageId="1" pageNumber="186">
|
||
<taxonomicName id="4DAC4D32FFD8FFB34197FC0FFCE6FC78" ID-CoL="2303263a-0456-4070-83d3-5f1c7275e6cc" authority="A. Milne-Edwards, 1880: 3" authorityName="A. Milne-Edwards" authorityPageNumber="3" authorityYear="1880" box="[199,781,959,984]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFD8FFB34197FC0FFE39FC77" box="[199,466,960,984]" italics="true" pageId="1" pageNumber="186">Pachycheles rugimanus</emphasis>
|
||
<treatmentCitation id="0B0D10A0FFD8FFB34089FC0FFCE6FC78" author="Milne-Edwards, A." box="[473,781,959,984]" page="3" pageId="1" pageNumber="186" year="1880">
|
||
<bibRefCitation id="EE3D4B40FFD8FFB34089FC0FFD1FFC78" author="Milne-Edwards, A." box="[473,756,959,984]" pageId="1" pageNumber="186" pagination="1 - 68" refId="ref4147" refString="Milne-Edwards, A. (1880) Reports on the results of dredging under the supervision of Alexander Agassiz, in the Gulf of Mexico and in the Caribbean Sea, 1877, 78, 79, by the U. S. Coast Survey Steamer " Blake ", Lieut. - Commander C. D. Sigsbee, U. S. N. and Commander J. R. Bartlett, U. S. N. commanding. VIII. Etudes preliminaires sur les Crustaces, 1 ere Partie. Bulletin of the Museum of Comparative Zoology, 8, 1 - 68." type="journal article" year="1880">A. Milne-Edwards, 1880</bibRefCitation>
|
||
:
|
||
<specimenCount id="9CAAFD38FFD8FFB343AFFC0FFCE6FC78" box="[767,781,960,984]" pageId="1" pageNumber="186" type="male">3</specimenCount>
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
males,
|
||
<materialsCitation id="3AC43CECFFD8FFB34234FC70FB2EFB80" ID-GBIF-Occurrence="2013824449" collectingDate="1979-09-21" collectionCode="USNM" collectorName="V Dolphin" country="United States of America" latitude="32.835" location="Stn" longLatPrecision="20" longitude="-78.604996" pageId="1" pageNumber="186" specimenCode="USNM 188445" specimenCount="35" specimenCount-male="35" stateProvince="South Carolina">
|
||
largest cl
|
||
<quantity id="4D549B54FFD8FFB34280FC0FFBC3FC77" box="[976,1064,960,984]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="1" pageNumber="186" unit="mm" value="8.1">8.1 mm</quantity>
|
||
, cw
|
||
<quantity id="4D549B54FFD8FFB3450CFC0FFB5EFC77" box="[1116,1205,960,984]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="1" pageNumber="186" unit="mm" value="7.8">7.8 mm</quantity>
|
||
; smallest cl
|
||
<quantity id="4D549B54FFD8FFB3446FFC0FFA73FC77" box="[1343,1432,960,984]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="186" unit="mm" value="7.3">7.3 mm</quantity>
|
||
, cw
|
||
<quantity id="4D549B54FFD8FFB34192FC2BFEF5FC5C" box="[194,286,996,1020]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="186" unit="mm" value="7.1">7.1 mm</quantity>
|
||
; 1 ovigerous female, cl
|
||
<quantity id="4D549B54FFD8FFB3436AFC2BFD7EFC5C" box="[570,661,996,1020]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="186" unit="mm" value="7.9">7.9 mm</quantity>
|
||
, cw
|
||
<quantity id="4D549B54FFD8FFB3439FFC2BFCC2FC5C" box="[719,809,996,1020]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="1" pageNumber="186" unit="mm" value="8.4">8.4 mm</quantity>
|
||
(
|
||
<specimenCode id="DA0A9ECAFFD8FFB3426AFC2BFC04FC5D" box="[826,1007,996,1021]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="186" type="Museum">USNM 188445</specimenCode>
|
||
),
|
||
<collectingCountry id="F2BB7621FFD8FFB34556FC2BFB4AFC5C" box="[1030,1185,996,1021]" name="United States of America" pageId="1" pageNumber="186">United States</collectingCountry>
|
||
,
|
||
<collectingRegion id="4868F853FFD8FFB345E0FC2BFAB4FC5D" box="[1200,1375,996,1021]" country="United States of America" name="South Carolina" pageId="1" pageNumber="186">South Carolina</collectingRegion>
|
||
, R/
|
||
<collectorName id="27595367FFD8FFB344D8FC2BFF1DFB80" pageId="1" pageNumber="186">V Dolphin</collectorName>
|
||
,
|
||
<location id="8F73606AFFD8FFB34051FBC7FECDFB80" LSID="urn:lsid:plazi:treatment:020587A7FFD8FFB541C7FDEDFBE4FE70:8F73606AFFD8FFB34051FBC7FECDFB80" box="[257,294,1032,1056]" country="United States of America" latitude="32.835" longLatPrecision="20" longitude="-78.604996" name="Stn" pageId="1" pageNumber="186" stateProvince="South Carolina">Stn</location>
|
||
MARMAP 791041,
|
||
<geoCoordinate id="EF985076FFD8FFB34344FBC8FD43FB80" box="[532,680,1031,1056]" direction="north" orientation="latitude" pageId="1" pageNumber="186" precision="15" value="32.835">32°50'06"N–</geoCoordinate>
|
||
<geoCoordinate id="EF985076FFD8FFB343F8FBC8FCC4FB80" box="[680,815,1031,1056]" direction="west" orientation="longitude" pageId="1" pageNumber="186" precision="15" value="-78.604996">78°36'18"W</geoCoordinate>
|
||
,
|
||
<date id="FE121071FFD8FFB3426BFBC7FBF2FB80" box="[827,1049,1031,1056]" pageId="1" pageNumber="186" value="1979-09-21">
|
||
<collectingDate id="EE56E999FFD8FFB3426BFBC7FBF2FB80" box="[827,1049,1031,1056]" pageId="1" pageNumber="186" value="1979-09-21">21 September 1979</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9CAAFD38FFD8FFB34574FBC8FB2EFB80" box="[1060,1221,1031,1056]" pageId="1" pageNumber="186" type="male">35 m. 2 males</specimenCount>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3AC43CECFFD8FFB34581FBC8FEACFB2C" ID-GBIF-Occurrence="2013824445" collectingDate="1963-09-12" collectionCode="USNM" collectorName="M. Gray" country="United States of America" county="Sapelo Island" elevation="59" latitude="30.67861" location="Sapelo Island" longLatPrecision="20" longitude="-80.101944" municipality="Sea Buoy" pageId="1" pageNumber="186" specimenCode="USNM 150684" specimenCount="59" stateProvince="Georgia">
|
||
cl
|
||
<quantity id="4D549B54FFD8FFB345BDFBC8FAAEFBBF" box="[1261,1349,1031,1056]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="1" pageNumber="186" unit="mm" value="6.2">6.2 mm</quantity>
|
||
, cw
|
||
<quantity id="4D549B54FFD8FFB34429FBC8FF28FBE4" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="1" pageNumber="186" unit="mm" value="5.9">5.9 mm</quantity>
|
||
and cl 5.8, cw
|
||
<quantity id="4D549B54FFD8FFB3402EFBE3FE32FBE4" box="[382,473,1068,1093]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="186" unit="mm" value="5.6">5.6 mm</quantity>
|
||
, 2 ovigerous females, cl
|
||
<quantity id="4D549B54FFD8FFB34250FBE3FCB0FBE4" box="[768,859,1068,1092]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="186" unit="mm" value="7.1">7.1 mm</quantity>
|
||
, cw
|
||
<quantity id="4D549B54FFD8FFB342C5FBE3FC05FBE4" box="[917,1006,1068,1092]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="1" pageNumber="186" unit="mm" value="7.2">7.2 mm</quantity>
|
||
and cl
|
||
<quantity id="4D549B54FFD8FFB3451AFBE3FB4EFBE4" box="[1098,1189,1068,1092]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="1" pageNumber="186" unit="mm" value="7.2">7.2 mm</quantity>
|
||
, cw
|
||
<quantity id="4D549B54FFD8FFB3458FFBE3FAD3FBE4" box="[1247,1336,1068,1092]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="186" unit="mm" value="7.4">7.4 mm</quantity>
|
||
(
|
||
<specimenCode id="DA0A9ECAFFD8FFB3441AFBE3FF04FBC8" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="186" type="Museum">
|
||
USNM
|
||
<date id="FE121071FFD8FFB341C7FB80FF04FBC8" box="[151,239,1103,1128]" pageId="1" pageNumber="186" value="84-06-15">150684</date>
|
||
</specimenCode>
|
||
),
|
||
<collectingRegion id="4868F853FFD8FFB34054FB80FE8AFBC8" box="[260,353,1103,1128]" pageId="1" pageNumber="186">Georgia</collectingRegion>
|
||
,
|
||
<location id="8F73606AFFD8FFB3403FFB80FDE7FBC8" LSID="urn:lsid:plazi:treatment:020587A7FFD8FFB541C7FDEDFBE4FE70:8F73606AFFD8FFB3403FFB80FDE7FBC8" box="[367,524,1103,1128]" country="Georgia" county="Sapelo Island" latitude="30.67861" longLatPrecision="20" longitude="-80.101944" municipality="Sea Buoy" name="Sapelo Island" pageId="1" pageNumber="186" stateProvince="Georgia">Sapelo Island</location>
|
||
, off Sea Buoy, Stn 373,
|
||
<geoCoordinate id="EF985076FFD8FFB34279FB80FC57FBC8" box="[809,956,1103,1128]" direction="north" orientation="latitude" pageId="1" pageNumber="186" precision="15" value="30.67861">30°40'43"N–</geoCoordinate>
|
||
<geoCoordinate id="EF985076FFD8FFB342ECFB80FBAEFBC8" box="[956,1093,1103,1128]" direction="west" orientation="longitude" pageId="1" pageNumber="186" precision="15" value="-80.101944">80°06'07"W</geoCoordinate>
|
||
,
|
||
<collectorName id="27595367FFD8FFB34503FB9FFB5FFBC8" box="[1107,1204,1104,1128]" pageId="1" pageNumber="186">M. Gray</collectorName>
|
||
coll.,
|
||
<date id="FE121071FFD8FFB345AFFB9FFF39FB2D" pageId="1" pageNumber="186" value="1963-09-12">
|
||
<collectingDate id="EE56E999FFD8FFB345AFFB9FFF39FB2D" pageId="1" pageNumber="186" value="1963-09-12">12 September 1963</collectingDate>
|
||
</date>
|
||
, 50–
|
||
<specimenCount id="9CAAFD38FFD8FFB34058FBBBFEACFB2C" box="[264,327,1140,1165]" pageId="1" pageNumber="186" type="generic">
|
||
<quantity id="4D549B54FFD8FFB34058FBBBFEA8FB2C" box="[264,323,1140,1165]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="1" pageNumber="186" unit="m" value="59.0">59 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</materialsCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8A1336B1FFD8FFB34197FB57FAA7FB74" blockId="1.[151,1437,743,2029]" pageId="1" pageNumber="186">
|
||
<treatmentCitationGroup id="AABC119FFFD8FFB34197FB57FAA7FB74" infered="true" pageId="1" pageNumber="186">
|
||
<taxonomicName id="4DAC4D32FFD8FFB34197FB57FDBAFB10" ID-CoL="62cff5ae-ba17-4d59-a49e-75ab01d6ab7b" authority="Haig, 1960: 1" authorityName="Haig" authorityPageNumber="1" authorityYear="1960" box="[199,593,1175,1200]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFD8FFB34197FB57FE45FB10" box="[199,430,1176,1200]" italics="true" pageId="1" pageNumber="186">Pachycheles velerae</emphasis>
|
||
<treatmentCitation id="0B0D10A0FFD8FFB340E5FB58FDBAFB10" author="Haig, J." box="[437,593,1175,1200]" page="1" pageId="1" pageNumber="186" year="1960">
|
||
<bibRefCitation id="EE3D4B40FFD8FFB340E5FB58FDDCFB10" author="Haig, J." box="[437,567,1175,1200]" pageId="1" pageNumber="186" pagination="1 - 440" refId="ref3943" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig, 1960</bibRefCitation>
|
||
:
|
||
<specimenCount id="9CAAFD38FFD8FFB34313FB57FDBAFB10" box="[579,593,1176,1200]" pageId="1" pageNumber="186" type="male">1</specimenCount>
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
male,
|
||
<materialsCitation id="3AC43CECFFD8FFB343CFFB58FAA7FB74" ID-GBIF-Occurrence="2013824448" collectingDate="1938-01-14" collectionCode="USNM" collectorName="Allan Hancock Expedition, R & V Velero" country="Costa Rica" elevation="86" latitude="5.5972223" location="Cocos Island" longLatPrecision="21" longitude="-86.98194" municipality="Chatham Bay" pageId="1" pageNumber="186" specimenCode="USNM 110551" specimenCount="86">
|
||
cl
|
||
<quantity id="4D549B54FFD8FFB343ECFB58FCFEFB0F" box="[700,789,1175,1200]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="1" pageNumber="186" unit="mm" value="5.4">5.4 mm</quantity>
|
||
, cw
|
||
<quantity id="4D549B54FFD8FFB3421BFB57FC49FB0F" box="[843,930,1176,1200]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="1" pageNumber="186" unit="mm" value="4.9">4.9 mm</quantity>
|
||
(
|
||
<specimenCode id="DA0A9ECAFFD8FFB342E1FB57FB88FB10" box="[945,1123,1175,1200]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="186" type="Museum">
|
||
USNM
|
||
<date id="FE121071FFD8FFB3455CFB58FB88FB10" box="[1036,1123,1175,1200]" pageId="1" pageNumber="186" value="51-11-05">110551</date>
|
||
</specimenCode>
|
||
),
|
||
<collectingCountry id="F2BB7621FFD8FFB34528FB57FB1DFB10" box="[1144,1270,1175,1200]" name="Costa Rica" pageId="1" pageNumber="186">Costa Rica</collectingCountry>
|
||
,
|
||
<location id="8F73606AFFD8FFB34453FB57FA73FB10" LSID="urn:lsid:plazi:treatment:020587A7FFD8FFB541C7FDEDFBE4FE70:8F73606AFFD8FFB34453FB57FA73FB10" box="[1283,1432,1175,1200]" country="Costa Rica" latitude="5.5972223" longLatPrecision="21" longitude="-86.98194" municipality="Chatham Bay" name="Cocos Island" pageId="1" pageNumber="186">Cocos Island</location>
|
||
,
|
||
<collectingMunicipality id="6A77ACCBFFD8FFB341C7FB73FEDAFB75" box="[151,305,1212,1237]" pageId="1" pageNumber="186">Chatham Bay</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="EF985076FFD8FFB3406CFB73FE2BFB75" box="[316,448,1212,1237]" direction="north" orientation="latitude" pageId="1" pageNumber="186" precision="15" value="5.5972223">5°35'50"N–</geoCoordinate>
|
||
<geoCoordinate id="EF985076FFD8FFB34090FB73FDA1FB75" box="[448,586,1212,1237]" direction="west" orientation="longitude" pageId="1" pageNumber="186" precision="15" value="-86.98194">86°58'55"W</geoCoordinate>
|
||
,
|
||
<collectorName id="27595367FFD8FFB34306FB73FC4AFB74" box="[598,929,1212,1237]" pageId="1" pageNumber="186">Allan Hancock Expedition, R</collectorName>
|
||
/
|
||
<collectorName id="27595367FFD8FFB342F9FB73FBFBFB75" box="[937,1040,1212,1237]" pageId="1" pageNumber="186">V Velero</collectorName>
|
||
,
|
||
<date id="FE121071FFD8FFB3454BFB73FB3CFB74" box="[1051,1239,1212,1237]" pageId="1" pageNumber="186" value="1938-01-14">
|
||
<collectingDate id="EE56E999FFD8FFB3454BFB73FB3CFB74" box="[1051,1239,1212,1237]" pageId="1" pageNumber="186" value="1938-01-14">14 January 1938</collectingDate>
|
||
</date>
|
||
, 73–
|
||
<specimenCount id="9CAAFD38FFD8FFB3445DFB73FAA7FB74" box="[1293,1356,1212,1237]" pageId="1" pageNumber="186" type="generic">
|
||
<quantity id="4D549B54FFD8FFB3445DFB73FAA3FB74" box="[1293,1352,1212,1237]" metricMagnitude="1" metricUnit="m" metricValue="8.6" pageId="1" pageNumber="186" unit="m" value="86.0">
|
||
<elevation id="0181D182FFD8FFB3445DFB73FAA3FB74" box="[1293,1352,1212,1237]" metricMagnitude="1" metricUnit="m" metricValue="8.6" pageId="1" pageNumber="186" unit="m" value="86.0">86 m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
</materialsCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C2B6653AFFD8FFB34197FB10FC17FB58" box="[199,1020,1247,1272]" pageId="1" pageNumber="186" type="distribution">
|
||
<paragraph id="8A1336B1FFD8FFB34197FB10FC17FB58" blockId="1.[151,1437,743,2029]" box="[199,1020,1247,1272]" pageId="1" pageNumber="186">
|
||
<emphasis id="B8D8EAA3FFD8FFB34197FB10FE8BFB58" bold="true" box="[199,352,1247,1272]" pageId="1" pageNumber="186">Distribution.</emphasis>
|
||
Currently known only from Cabo Orange,
|
||
<collectingRegion id="4868F853FFD8FFB34215FB2FFC7DFB58" box="[837,918,1248,1272]" country="Brazil" name="Amapa" pageId="1" pageNumber="186">Amapá</collectingRegion>
|
||
(
|
||
<collectingCountry id="F2BB7621FFD8FFB342F5FB10FC04FB58" box="[933,1007,1247,1272]" name="Brazil" pageId="1" pageNumber="186">Brazil</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C2B6653AFFD8FFB04197FACBFE26FC98" lastPageId="2" lastPageNumber="187" pageId="1" pageNumber="186" type="description">
|
||
<paragraph id="8A1336B1FFD8FFB34197FACBFB87F99D" blockId="1.[151,1437,743,2029]" pageId="1" pageNumber="186">
|
||
<emphasis id="B8D8EAA3FFD8FFB34197FACBFEB2FABD" bold="true" box="[199,345,1284,1309]" pageId="1" pageNumber="186">Description.</emphasis>
|
||
Carapace subcircular, longer than broad in both males and females, quite rugose dorsally, with scattered flattened tubercles dorsally. Frontal region prominent, rugose, slightly depressed on midline. Frontal width less than half of maximum carapace width. Frontal margin distinctly granulate, trilobate in dorsal and frontal views, devoid of setae; median lobe prominent, rounded, overreaching lateral lobes; lateral lobes broad, rounded. Outer orbital angle terminating in well-developed, long teeth. Orbit with rugose margin. Carapace lateral margin well-defined, wrinkled. Epibranchial region slightly swollen. Cervical groove well defined, ending in distinct notch anteriorly. Posterior branchial region with long, transverse rugae; posterolateral margin gently rounded, devoid of setae. Branchiostegite distinctly rugose, not pubescent, divided into calcified plates of different sizes; anterior plate distinctly larger than posterior; sometimes with several much smaller plates posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8A1336B1FFD8FFB34197F988FC70F908" blockId="1.[151,1437,743,2029]" pageId="1" pageNumber="186">Thoracic sternite 3 trilobate; median lobe broad, rounded, unarmed, not overreaching lateral lobes; lateral lobes narrow, subtriangular. Sternites 4–7 broad, sutures between sternites incomplete, distinct only laterally. Sternite 8 anterior margin slightly convex, posterior margin concave.</paragraph>
|
||
<paragraph id="8A1336B1FFD8FFB34197F97BFCC9F96D" blockId="1.[151,1437,743,2029]" box="[199,802,1716,1741]" pageId="1" pageNumber="186">Ocular peduncle short; cornea large, rounded, dilated.</paragraph>
|
||
<paragraph id="8A1336B1FFD8FFB34197F918FE0FF8B5" blockId="1.[151,1437,743,2029]" pageId="1" pageNumber="186">Antennular basal segment large, with 2 small distomesial spines. Segments 2–3 unarmed, subcylindrical; segment 3 broadened distally.</paragraph>
|
||
<paragraph id="8A1336B1FFD8FFB34197F8D0FB50F820" blockId="1.[151,1437,743,2029]" pageId="1" pageNumber="186">Antennal peduncular segments slightly rugose, short. Segment 1 subtriangular, membranous dorsally, calcified ventrally. Segments 2–3 about 2/3 length of segment 3, with well-developed distomesial tubercle. Segment 4 mesial surface with 2 or 3 well-developed granules. Segment 5 about half length of segment 4.</paragraph>
|
||
<paragraph id="8A1336B1FFD8FFB34197F843FC8AF84D" blockId="1.[151,1437,743,2029]" pageId="1" pageNumber="186">Mxp3 rugose on external surface. Exopod reniform, reaching about half of total length of merus; flagellum well developed. Ischium lateral lobe distinct, well-developed, separated by deep V-shaped incision. Merus as long as ischium; mesial lobe broad, tip rounded, with serrate margin.</paragraph>
|
||
<paragraph id="8A1336B1FFDBFFB04197FF57FD95FE0D" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="187">P1 similar between male and female; right and left sides unequal in size but similar in armature, robust, eroded, distinctly granulate dorsally; ventral surface with flattened tubercles. Merus about 1/3 length of carpus; mesial lobe subtriangular, well-developed, granulate; 2–4 distal spines well-developed ventrally. Carpus as long as palm; mesial margin with 3–4 granulate, strongly projecting teeth, progressively decreasing in size distally. Dorsal surface of carpus and propodus with 3 pronounced, longitudinal carinae and series of transverse, discrete ridges between the longitudinal carinae. Palm with large, granular tubercle near base of pollex. Fixed finger about 1/3 length of propodus. Dactylus about 2/3 length of propodus. Cutting edges of fingers furrowed, granulate, devoid of setae. Tip of fingers curved, closing tightly.</paragraph>
|
||
<paragraph id="8A1336B1FFDBFFB04197FE78FA76FDC0" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="187">Walking legs decreasing in size from P2–4, robust, rugose, fringed with dense, short setae on margins, especially on carpi and propodi, and with scattered long setae. Carpi about 2/3 length of P2–4 meri, lateral surface with 2 low, longitudinal, parallel carinae dorsally; dorsal margin with 1 to 3 discrete, dorsodistal spines. Propodi as long as meri, each with 4 spines on ventral surface: 2 spines along ventral margin, 2 on distoventral margin near articulation with dactylus. Dactyli about half length of propodi, each with row of 4 well-developed spines ventrally.</paragraph>
|
||
<paragraph id="8A1336B1FFDBFFB04197FDA3FD56FD08" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="187">P5 short, ischium 1/4 of length of merus; merus slightly longer than carpus; propodus about 1/3 of length of carpus; dactylus about 1/3 of length of propodus.</paragraph>
|
||
<paragraph id="8A1336B1FFDBFFB04197FD7BFE26FC98" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="187">Male and female pleon subrectangular, surface of each somite smooth dorsally, lateral margins with short, plumose setae. Male gonopods well developed on somite 2. Female pleopods well developed on somites 3–5. Uropods broad, reaching to distal margin of telson. Telson broader than long, subdivided into 5 plates, nearly smooth on external surface.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C2B6653AFFDBFFB04197FC8BFE63FC05" pageId="2" pageNumber="187" type="etymology">
|
||
<paragraph id="8A1336B1FFDBFFB04197FC8BFE63FC05" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="187">
|
||
<emphasis id="B8D8EAA3FFDBFFB04197FC8BFEA5FCFD" bold="true" box="[199,334,836,861]" pageId="2" pageNumber="187">Etymology.</emphasis>
|
||
The new species is named in honour of the late Prof. Petrônio Alves Coelho (Universidade Federal de
|
||
<collectingRegion id="4868F853FFDBFFB04075FCA8FE58FC20" box="[293,435,871,896]" country="Brazil" name="Pernambuco" pageId="2" pageNumber="187">Pernambuco</collectingRegion>
|
||
) in recognition of his outstanding contributions to the taxonomy of the decapod crustaceans in
|
||
<collectingCountry id="F2BB7621FFDBFFB0406CFC43FE6FFC05" box="[316,388,908,933]" name="Brazil" pageId="2" pageNumber="187">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C2B6653AFFDBFFB54197FC60FBE4FE70" lastPageId="7" lastPageNumber="192" pageId="2" pageNumber="187" type="discussion">
|
||
<paragraph id="8A1336B1FFDBFFB04197FC60FE30FB64" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="187">
|
||
<emphasis id="B8D8EAA3FFDBFFB04197FC60FED0FC68" bold="true" box="[199,315,943,968]" pageId="2" pageNumber="187">Remarks.</emphasis>
|
||
<taxonomicName id="4DAC4D32FFDBFFB04016FC7FFDDBFC68" authority="Ferreira & Tavares, 2019" authorityName="Ferreira & Tavares" authorityYear="2019" box="[326,560,944,968]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="coelhoi" status="sp. nov.">
|
||
<emphasis id="B8D8EAA3FFDBFFB04016FC7FFDDBFC68" box="[326,560,944,968]" italics="true" pageId="2" pageNumber="187">Pachycheles coelhoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B8D8EAA3FFDBFFB0436CFC60FD71FC68" bold="true" box="[572,666,943,968]" pageId="2" pageNumber="187">
|
||
<taxonomicNameLabel id="A3EB57D8FFDBFFB0436CFC60FD71FC68" box="[572,666,943,968]" pageId="2" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
superficially resembles
|
||
<taxonomicName id="4DAC4D32FFDBFFB042E8FC7EFBBBFC67" authorityName="A. Milne-Edwards" authorityYear="1880" box="[952,1104,944,968]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDBFFB042E8FC7EFBBBFC67" box="[952,1104,944,968]" italics="true" pageId="2" pageNumber="187">P. rugimanus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="12972A34FFDBFFB04534FC60FB5FFC68" box="[1124,1204,943,968]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="2" pageNumber="187">Figs. 2</figureCitation>
|
||
D–F, 4C, D) and
|
||
<taxonomicName id="4DAC4D32FFDBFFB044D7FC7EFF01FC4C" authorityName="Haig" authorityYear="1960" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDBFFB044D7FC7EFF01FC4C" italics="true" pageId="2" pageNumber="187">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="12972A34FFDBFFB041A9FC1BFEAFFC4C" box="[249,324,980,1005]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1931]" captionTargetBox="[201,1388,193,1893]" captionTargetId="figure@5.[201,1388,193,1893]" captionTargetPageId="5" captionText="FIGURE 3. (A–C) Pachycheles ackleianus (A. Milne-Edwards, 1880), male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D)." httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="2" pageNumber="187">Figs. 3</figureCitation>
|
||
D–F, 4E, F). However, the new species can be separated from both,
|
||
<taxonomicName id="4DAC4D32FFDBFFB04510FC1AFB3FFC4C" authorityName="A. Milne-Edwards" authorityYear="1880" box="[1088,1236,981,1004]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDBFFB04510FC1AFB3FFC4C" box="[1088,1236,981,1004]" italics="true" pageId="2" pageNumber="187">P. rugimanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4DAC4D32FFDBFFB0445AFC1AFA93FC4C" authorityName="Haig" authorityYear="1960" box="[1290,1400,980,1004]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDBFFB0445AFC1AFA93FC4C" box="[1290,1400,980,1004]" italics="true" pageId="2" pageNumber="187">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
, in that: (1) the major and minor P1 fingers are similar in shape; the fingers have nearly straight cutting edges and are interlocked subdistally, with the dactyli closing tightly against the fixed fingers,
|
||
<figureCitation id="12972A34FFDBFFB04543FBD3FB7FFB94" box="[1043,1172,1052,1077]" captionStart="FIGURE 1" captionStartId="3.[151,250,1888,1910]" captionTargetBox="[329,1288,217,1862]" captionTargetId="figure@3.[208,1381,195,1871]" captionTargetPageId="3" captionText="FIGURE 1. Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (A) carapace, dorsal view. (B) frontal-orbital region of carapace, frontal view. (C) carapace, lateral view, emphasizing the branchiostegite. (D) right antennule, ventral view. (E) right antennal peduncle, dorsal view (flagellum omitted). (F) left Mxp 3, ventral view (exopod flagellum not shown). (G) left minor P1, dorsal view. (H) right major P1, dorsal view. (I) right P2, lateral view. (J) telson, external view. (K) thoracic sternum, ventral view. Setae all omitted. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2599189/files/figure.png" pageId="2" pageNumber="187">Figs. 1G, H</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB045CEFBD3FB2AFB94" box="[1182,1217,1052,1076]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="2" pageNumber="187">2C</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB0459CFBD3FAE4FB94" box="[1228,1295,1052,1076]" captionStart="FIGURE 4" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[151,1436,661,1843]" captionTargetId="figure@6.[151,1436,661,1843]" captionTargetPageId="6" captionText="FIGURE 4. Major chelae. (A, B) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E)." httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="2" pageNumber="187">4A, B</figureCitation>
|
||
(in the other two species, the major and minor P1 fingers are unequal in shape; the major P1 fingers leave a broad hiatus when closed and are not interlocked distally, with the dactylus strongly curved to ventral side,
|
||
<figureCitation id="12972A34FFDBFFB04521FBABFB26FBDC" box="[1137,1229,1124,1149]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="2" pageNumber="187">Figs. 2F</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB04588FBABFB1DFBDC" box="[1240,1270,1124,1148]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1931]" captionTargetBox="[201,1388,193,1893]" captionTargetId="figure@5.[201,1388,193,1893]" captionTargetPageId="5" captionText="FIGURE 3. (A–C) Pachycheles ackleianus (A. Milne-Edwards, 1880), male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D)." httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="2" pageNumber="187">3F</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB04451FBABFAE4FBDC" box="[1281,1295,1124,1148]" captionStart="FIGURE 4" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[151,1436,661,1843]" captionTargetId="figure@6.[151,1436,661,1843]" captionTargetPageId="6" captionText="FIGURE 4. Major chelae. (A, B) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E)." httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="2" pageNumber="187">4</figureCitation>
|
||
C-F); (2) the dorsal surface of the major P1 finger is heavily tuberculated rather than nearly smooth in
|
||
<taxonomicName id="4DAC4D32FFDBFFB045E3FB46FAA1FB3F" authorityName="A. Milne-Edwards" authorityYear="1880" box="[1203,1354,1160,1184]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDBFFB045E3FB46FAA1FB3F" box="[1203,1354,1160,1184]" italics="true" pageId="2" pageNumber="187">P. rugimanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4DAC4D32FFDBFFB044D7FB46FF01FB64" authorityName="Haig" authorityYear="1960" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDBFFB044D7FB46FF01FB64" italics="true" pageId="2" pageNumber="187">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="12972A34FFDBFFB041A9FB63FEBEFB64" box="[249,341,1196,1221]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="2" pageNumber="187">Figs. 2F</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB04031FB63FE94FB64" box="[353,383,1196,1220]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1931]" captionTargetBox="[201,1388,193,1893]" captionTargetId="figure@5.[201,1388,193,1893]" captionTargetPageId="5" captionText="FIGURE 3. (A–C) Pachycheles ackleianus (A. Milne-Edwards, 1880), male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D)." httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="2" pageNumber="187">3F</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB040DAFB63FE25FB64" box="[394,462,1196,1220]" captionStart="FIGURE 4" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[151,1436,661,1843]" captionTargetId="figure@6.[151,1436,661,1843]" captionTargetPageId="6" captionText="FIGURE 4. Major chelae. (A, B) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E)." httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="2" pageNumber="187">4C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8A1336B1FFDBFFB04197FB1FFBE9FA44" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="187">
|
||
<taxonomicName id="4DAC4D32FFDBFFB04197FB1FFE5AFB48" authority="Ferreira & Tavares, 2019" authorityName="Ferreira & Tavares" authorityYear="2019" box="[199,433,1232,1256]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="coelhoi" status="sp. nov.">
|
||
<emphasis id="B8D8EAA3FFDBFFB04197FB1FFE5AFB48" box="[199,433,1232,1256]" italics="true" pageId="2" pageNumber="187">Pachycheles coelhoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B8D8EAA3FFDBFFB040ECFB00FDF0FB48" bold="true" box="[444,539,1231,1256]" pageId="2" pageNumber="187">
|
||
<taxonomicNameLabel id="A3EB57D8FFDBFFB040ECFB00FDF0FB48" box="[444,539,1231,1256]" pageId="2" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is further differentiated from
|
||
<taxonomicName id="4DAC4D32FFDBFFB042D0FB1EFBF2FB47" authorityName="A. Milne-Edwards" authorityYear="1880" box="[896,1049,1232,1256]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDBFFB042D0FB1EFBF2FB47" box="[896,1049,1232,1256]" italics="true" pageId="2" pageNumber="187">P. rugimanus</emphasis>
|
||
</taxonomicName>
|
||
by: (1) the carapace front being slightly deflexed and markedly trilobate in dorsal view; the median lobe being strongly projected, distinctly overreaching the tip of cornea in dorsal view and separated from the lateral lobes by broad concave margins (
|
||
<figureCitation id="12972A34FFDBFFB04434FAD8FF50FAF4" captionStart="FIGURE 1" captionStartId="3.[151,250,1888,1910]" captionTargetBox="[329,1288,217,1862]" captionTargetId="figure@3.[208,1381,195,1871]" captionTargetPageId="3" captionText="FIGURE 1. Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (A) carapace, dorsal view. (B) frontal-orbital region of carapace, frontal view. (C) carapace, lateral view, emphasizing the branchiostegite. (D) right antennule, ventral view. (E) right antennal peduncle, dorsal view (flagellum omitted). (F) left Mxp 3, ventral view (exopod flagellum not shown). (G) left minor P1, dorsal view. (H) right major P1, dorsal view. (I) right P2, lateral view. (J) telson, external view. (K) thoracic sternum, ventral view. Setae all omitted. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2599189/files/figure.png" pageId="2" pageNumber="187">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB0419EFAF3FF19FAF4" box="[206,242,1340,1364]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="2" pageNumber="187">2A</figureCitation>
|
||
) (versus carapace front markedly deflexed, scarcely trilobate; the median lobe is very low, poorly projected, barely overreaching the tip of cornea and separated from the lateral lobes by nearly straight margins in
|
||
<taxonomicName id="4DAC4D32FFDBFFB044D7FAAEFEE4FA3C" authorityName="A. Milne-Edwards" authorityYear="1880" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDBFFB044D7FAAEFEE4FA3C" italics="true" pageId="2" pageNumber="187">P. rugimanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB0404DFA4BFE93FA3C" box="[285,376,1412,1437]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="2" pageNumber="187">Fig. 2D</figureCitation>
|
||
); (2) the median lobe of the thoracic sternite 3 is rounded and separated from the lateral lobes by shallow, gently concave margins (
|
||
<figureCitation id="12972A34FFDBFFB0431AFA68FD59FA60" box="[586,690,1447,1472]" captionStart="FIGURE 1" captionStartId="3.[151,250,1888,1910]" captionTargetBox="[329,1288,217,1862]" captionTargetId="figure@3.[208,1381,195,1871]" captionTargetPageId="3" captionText="FIGURE 1. Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (A) carapace, dorsal view. (B) frontal-orbital region of carapace, frontal view. (C) carapace, lateral view, emphasizing the branchiostegite. (D) right antennule, ventral view. (E) right antennal peduncle, dorsal view (flagellum omitted). (F) left Mxp 3, ventral view (exopod flagellum not shown). (G) left minor P1, dorsal view. (H) right major P1, dorsal view. (I) right P2, lateral view. (J) telson, external view. (K) thoracic sternum, ventral view. Setae all omitted. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2599189/files/figure.png" pageId="2" pageNumber="187">Figs. 1K</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB04391FA67FD08FA60" box="[705,739,1448,1472]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="2" pageNumber="187">2B</figureCitation>
|
||
) rather than distinctly subtriangular and separated from the lateral lobes by straight, laterally V-shaped margins in
|
||
<taxonomicName id="4DAC4D32FFDBFFB043ABFA02FC65FA44" authorityName="A. Milne-Edwards" authorityYear="1880" box="[763,910,1485,1508]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDBFFB043ABFA02FC65FA44" box="[763,910,1485,1508]" italics="true" pageId="2" pageNumber="187">P. rugimanus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="12972A34FFDBFFB042CDFA03FC1EFA44" box="[925,1013,1484,1509]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="2" pageNumber="187">Fig. 2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8A1336B1FFDBFFB44197FA3EFBC2FEBD" blockId="2.[151,1437,151,1977]" lastBlockId="6.[151,1437,151,608]" lastPageId="6" lastPageNumber="191" pageId="2" pageNumber="187">
|
||
In addition to the above mentioned distinctions,
|
||
<taxonomicName id="4DAC4D32FFDBFFB043ADFA3EFC84F9A8" authority="Ferreira & Tavares, 2019" authorityName="Ferreira & Tavares" authorityYear="2019" box="[765,879,1520,1544]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="coelhoi" status="sp. nov.">
|
||
<emphasis id="B8D8EAA3FFDBFFB043ADFA3EFC84F9A8" box="[765,879,1520,1544]" italics="true" pageId="2" pageNumber="187">P. coelhoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B8D8EAA3FFDBFFB0422AFA20FC3CF9A8" bold="true" box="[890,983,1519,1544]" pageId="2" pageNumber="187">
|
||
<taxonomicNameLabel id="A3EB57D8FFDBFFB0422AFA20FC3CF9A8" box="[890,983,1519,1544]" pageId="2" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4DAC4D32FFDBFFB04525FA3EFB0CF9A8" authorityName="Haig" authorityYear="1960" box="[1141,1255,1520,1544]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDBFFB04525FA3EFB0CF9A8" box="[1141,1255,1520,1544]" italics="true" pageId="2" pageNumber="187">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
in that: (1) the frontal, protogastric and hepatic regions of the carapace are strongly striated transversally, and the epibranchial region is coarsely squamose (
|
||
<figureCitation id="12972A34FFDBFFB040B8F9F8FDA6F9F0" box="[488,589,1591,1616]" captionStart="FIGURE 1" captionStartId="3.[151,250,1888,1910]" captionTargetBox="[329,1288,217,1862]" captionTargetId="figure@3.[208,1381,195,1871]" captionTargetPageId="3" captionText="FIGURE 1. Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (A) carapace, dorsal view. (B) frontal-orbital region of carapace, frontal view. (C) carapace, lateral view, emphasizing the branchiostegite. (D) right antennule, ventral view. (E) right antennal peduncle, dorsal view (flagellum omitted). (F) left Mxp 3, ventral view (exopod flagellum not shown). (G) left minor P1, dorsal view. (H) right major P1, dorsal view. (I) right P2, lateral view. (J) telson, external view. (K) thoracic sternum, ventral view. Setae all omitted. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2599189/files/figure.png" pageId="2" pageNumber="187">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB0430CF9F7FD6BF9F0" box="[604,640,1592,1616]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="2" pageNumber="187">2A</figureCitation>
|
||
) (in
|
||
<taxonomicName id="4DAC4D32FFDBFFB043E8F9F6FCC3F9F0" authorityName="Haig" authorityYear="1960" box="[696,808,1592,1616]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDBFFB043E8F9F6FCC3F9F0" box="[696,808,1592,1616]" italics="true" pageId="2" pageNumber="187">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
the frontal, protogastric and hepatic regions are nearly smooth and the epibranchial region have few minute squamae (
|
||
<figureCitation id="12972A34FFDBFFB04233F993FC55F9D4" box="[867,958,1628,1653]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1931]" captionTargetBox="[201,1388,193,1893]" captionTargetId="figure@5.[201,1388,193,1893]" captionTargetPageId="5" captionText="FIGURE 3. (A–C) Pachycheles ackleianus (A. Milne-Edwards, 1880), male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D)." httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="2" pageNumber="187">Fig. 3D</figureCitation>
|
||
); (2) the major P1 fingers cutting surfaces are markedly hollowed along the longitudinal line, with incisor teeth along hollows, and the dactylus cutting edge has no molar tooth (
|
||
<figureCitation id="12972A34FFDBFFB04026F96BFE31F91C" box="[374,474,1700,1725]" captionStart="FIGURE 1" captionStartId="3.[151,250,1888,1910]" captionTargetBox="[329,1288,217,1862]" captionTargetId="figure@3.[208,1381,195,1871]" captionTargetPageId="3" captionText="FIGURE 1. Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (A) carapace, dorsal view. (B) frontal-orbital region of carapace, frontal view. (C) carapace, lateral view, emphasizing the branchiostegite. (D) right antennule, ventral view. (E) right antennal peduncle, dorsal view (flagellum omitted). (F) left Mxp 3, ventral view (exopod flagellum not shown). (G) left minor P1, dorsal view. (H) right major P1, dorsal view. (I) right P2, lateral view. (J) telson, external view. (K) thoracic sternum, ventral view. Setae all omitted. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2599189/files/figure.png" pageId="2" pageNumber="187">Figs. 1H</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB040B6F96BFDE3F91C" box="[486,520,1700,1724]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="2" pageNumber="187">2C</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB04344F96BFDB2F91C" box="[532,601,1700,1724]" captionStart="FIGURE 4" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[151,1436,661,1843]" captionTargetId="figure@6.[151,1436,661,1843]" captionTargetPageId="6" captionText="FIGURE 4. Major chelae. (A, B) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E)." httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="2" pageNumber="187">4A, B</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="4DAC4D32FFDBFFB043BDF96AFCB0F91C" authorityName="Haig" authorityYear="1960" box="[749,859,1700,1724]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDBFFB043BDF96AFCB0F91C" box="[749,859,1700,1724]" italics="true" pageId="2" pageNumber="187">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
the dactylus and fixed finger are hollowed out only along the distal half and only slightly hollowed distally, respectively. The dactylus and fixed finger each have two longitudinal, subparallel rows of molariform flat teeth along the hollow axes and the dactylus has one strong molar tooth subproximally (considered an autapomorphy of
|
||
<taxonomicName id="4DAC4D32FFDBFFB043BEF8DEFCB0F888" authorityName="Haig" authorityYear="1960" box="[750,859,1808,1832]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDBFFB043BEF8DEFCB0F888" box="[750,859,1808,1832]" italics="true" pageId="2" pageNumber="187">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EE3D4B40FFDBFFB042D4F8DEFBFCF888" author="Harvey, A." box="[900,1047,1808,1833]" pageId="2" pageNumber="187" pagination="746 - 752" refId="ref3973" refString="Harvey, A. (1998) Rediscovery and range extension of the Galapagos " endemic " Pachycheles velerae (Decapoda: Anomura: Porcellanidae). Journal of Crustacean Biology, 18 (4), 746 - 752. https: // doi. org / 10.2307 / 1549151" type="journal article" year="1998">Harvey 1998</bibRefCitation>
|
||
) (
|
||
<figureCitation id="12972A34FFDBFFB0457DF8C0FB62F888" box="[1069,1161,1807,1832]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1931]" captionTargetBox="[201,1388,193,1893]" captionTargetId="figure@5.[201,1388,193,1893]" captionTargetPageId="5" captionText="FIGURE 3. (A–C) Pachycheles ackleianus (A. Milne-Edwards, 1880), male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D)." httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="2" pageNumber="187">Figs. 3F</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDBFFB045C4F8DFFB38F888" box="[1172,1235,1808,1832]" captionStart="FIGURE 4" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[151,1436,661,1843]" captionTargetId="figure@6.[151,1436,661,1843]" captionTargetPageId="6" captionText="FIGURE 4. Major chelae. (A, B) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E)." httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="2" pageNumber="187">4E, F</figureCitation>
|
||
); (3) the major P1 propodus dorsal surface has a wide, high, heavily tuberculated, elongated upper bulge (
|
||
<figureCitation id="12972A34FFDBFFB04524F8FBFB24F8EC" box="[1140,1231,1844,1869]" captionStart="FIGURE 4" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[151,1436,661,1843]" captionTargetId="figure@6.[151,1436,661,1843]" captionTargetPageId="6" captionText="FIGURE 4. Major chelae. (A, B) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E)." httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="2" pageNumber="187">Fig. 4A</figureCitation>
|
||
), versus a narrow, low, rather smooth, elongated upper ridge in
|
||
<taxonomicName id="4DAC4D32FFDBFFB043C3F896FCEAF8D0" authorityName="Haig" authorityYear="1960" box="[659,769,1880,1904]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDBFFB043C3F896FCEAF8D0" box="[659,769,1880,1904]" italics="true" pageId="2" pageNumber="187">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="12972A34FFDBFFB04242F898FC81F8D0" box="[786,874,1879,1904]" captionStart="FIGURE 4" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[151,1436,661,1843]" captionTargetId="figure@6.[151,1436,661,1843]" captionTargetPageId="6" captionText="FIGURE 4. Major chelae. (A, B) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E)." httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="2" pageNumber="187">Fig. 4E</figureCitation>
|
||
); (4) the major P1 propodus dorsal surface has an elongated, large, high, heavily tuberculated axial bulge and is provided with discrete, transverse ridges between its upper margin and the upper bulge (
|
||
<figureCitation id="12972A34FFDBFFB0434DF86FFD9CF818" box="[541,631,1952,1977]" captionStart="FIGURE 4" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[151,1436,661,1843]" captionTargetId="figure@6.[151,1436,661,1843]" captionTargetPageId="6" captionText="FIGURE 4. Major chelae. (A, B) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E)." httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="2" pageNumber="187">Fig. 4A</figureCitation>
|
||
), whereas in
|
||
<emphasis id="B8D8EAA3FFDBFFB04258F86EFC90F818" box="[776,891,1952,1976]" italics="true" pageId="2" pageNumber="187">
|
||
<taxonomicName id="4DAC4D32FFDBFFB04258F86EFC93F818" authorityName="Haig" authorityYear="1960" box="[776,888,1952,1976]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="velerae">P. velerae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the axial bulge is rounded, far smaller and lower (
|
||
<figureCitation id="12972A34FFDFFFB441CFFF58FF1FFF10" box="[159,244,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[151,1436,661,1843]" captionTargetId="figure@6.[151,1436,661,1843]" captionTargetPageId="6" captionText="FIGURE 4. Major chelae. (A, B) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E)." httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="6" pageNumber="191">Fig. 4E</figureCitation>
|
||
) and there is a net of ridges of different sizes forming a reticulated, wrinkled pattern instead of transverse ridges (
|
||
<figureCitation id="12972A34FFDFFFB441A0FF73FEA5FF74" box="[240,334,188,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[151,1436,661,1843]" captionTargetId="figure@6.[151,1436,661,1843]" captionTargetPageId="6" captionText="FIGURE 4. Major chelae. (A, B) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E)." httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="6" pageNumber="191">Fig. 4E</figureCitation>
|
||
). The more elongate carapace and projecting front, the sparse setation on the chelipeds and carapace, and the dense plumose setae on pereopods 2–4 were also regarded as additional autapomorphies of
|
||
<taxonomicName id="4DAC4D32FFDFFFB444D7FF2EFF01FEBC" authorityName="Haig" authorityYear="1960" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDFFFB444D7FF2EFF01FEBC" italics="true" pageId="6" pageNumber="191">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EE3D4B40FFDFFFB44043FECAFE52FEBC" author="Harvey, A." box="[275,441,260,285]" pageId="6" pageNumber="191" pagination="746 - 752" refId="ref3973" refString="Harvey, A. (1998) Rediscovery and range extension of the Galapagos " endemic " Pachycheles velerae (Decapoda: Anomura: Porcellanidae). Journal of Crustacean Biology, 18 (4), 746 - 752. https: // doi. org / 10.2307 / 1549151" type="journal article" year="1998">Harvey (1998)</bibRefCitation>
|
||
, but these characters are shared with
|
||
<taxonomicName id="4DAC4D32FFDFFFB4420BFECAFC22FEBC" authority="Ferreira & Tavares, 2019" authorityName="Ferreira & Tavares" authorityYear="2019" box="[859,969,260,284]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="coelhoi" status="sp. nov.">
|
||
<emphasis id="B8D8EAA3FFDFFFB4420BFECAFC22FEBC" box="[859,969,260,284]" italics="true" pageId="6" pageNumber="191">P. coelhoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B8D8EAA3FFDFFFB44280FECBFBC2FEBD" bold="true" box="[976,1065,260,285]" pageId="6" pageNumber="191">
|
||
<taxonomicNameLabel id="A3EB57D8FFDFFFB44280FECBFBC2FEBD" box="[976,1065,260,285]" pageId="6" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<caption id="DED36639FFDAFFB141C7F8AFFBEBF851" httpUri="https://zenodo.org/record/2599189/files/figure.png" pageId="3" pageNumber="188" startId="3.[151,250,1888,1910]" targetBox="[329,1288,217,1862]" targetPageId="3">
|
||
<paragraph id="8A1336B1FFDAFFB141C7F8AFFBEBF851" blockId="3.[151,1436,1888,2033]" pageId="3" pageNumber="188">
|
||
<emphasis id="B8D8EAA3FFDAFFB141C7F8AFFEF8F8D5" bold="true" box="[151,275,1888,1910]" pageId="3" pageNumber="188">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4DAC4D32FFDAFFB14049F8AFFE02F8D6" authority="Ferreira & Tavares, 2019" authorityName="Ferreira & Tavares" authorityYear="2019" box="[281,489,1888,1910]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="188" phylum="Arthropoda" rank="species" species="coelhoi" status="sp. nov.">
|
||
<emphasis id="B8D8EAA3FFDAFFB14049F8AFFE02F8D6" box="[281,489,1888,1910]" italics="true" pageId="3" pageNumber="188">Pachycheles coelhoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B8D8EAA3FFDAFFB140BFF8AFFDABF8D5" bold="true" box="[495,576,1888,1909]" pageId="3" pageNumber="188">
|
||
<taxonomicNameLabel id="A3EB57D8FFDAFFB140BFF8AFFDABF8D5" box="[495,576,1888,1909]" pageId="3" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (A) carapace, dorsal view. (B) frontal-orbital region of carapace, frontal view. (C) carapace, lateral view, emphasizing the branchiostegite. (D) right antennule, ventral view. (E) right antennal peduncle, dorsal view (flagellum omitted). (F) left Mxp 3, ventral view (exopod flagellum not shown). (G) left minor P1, dorsal view. (H) right major P1, dorsal view. (I) right P2, lateral view. (J) telson, external view. (K) thoracic sternum, ventral view. Setae all omitted. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DED36639FFDDFFB641C7F8D2FD6CF84A" httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="4" pageNumber="189" startId="4.[151,250,1821,1843]" targetBox="[302,1288,193,1808]" targetPageId="4">
|
||
<paragraph id="8A1336B1FFDDFFB641C7F8D2FD6CF84A" blockId="4.[151,1436,1820,2026]" pageId="4" pageNumber="189">
|
||
<emphasis id="B8D8EAA3FFDDFFB641C7F8D2FEF9F892" bold="true" box="[151,274,1821,1843]" pageId="4" pageNumber="189">FIGURE 2.</emphasis>
|
||
(A–C)
|
||
<taxonomicName id="4DAC4D32FFDDFFB64030F8D3FDDBF892" authority="Ferreira & Tavares, 2019" authorityName="Ferreira & Tavares" authorityYear="2019" box="[352,560,1820,1842]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="coelhoi" status="sp. nov.">
|
||
<emphasis id="B8D8EAA3FFDDFFB64030F8D3FDDBF892" box="[352,560,1820,1842]" italics="true" pageId="4" pageNumber="189">Pachycheles coelhoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B8D8EAA3FFDDFFB64366F8D2FD6CF892" bold="true" box="[566,647,1821,1842]" pageId="4" pageNumber="189">
|
||
<taxonomicNameLabel id="A3EB57D8FFDDFFB64366F8D2FD6CF892" box="[566,647,1821,1842]" pageId="4" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F)
|
||
<taxonomicName id="4DAC4D32FFDDFFB6444DF8D3FDF4F8F1" authority="(A. Milne-Edwards, 1880)" authorityName="A. Milne-Edwards" authorityYear="1880" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDDFFB6444DF8D3FEEFF8F1" italics="true" pageId="4" pageNumber="189">Pachycheles rugimanus</emphasis>
|
||
(A. Milne-Edwards, 1880)
|
||
</taxonomicName>
|
||
, male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DED36639FFDCFFB741C7F8BAFB20F847" httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="5" pageNumber="190" startId="5.[151,250,1909,1931]" targetBox="[201,1388,193,1893]" targetPageId="5">
|
||
<paragraph id="8A1336B1FFDCFFB741C7F8BAFB20F847" blockId="5.[151,1436,1909,2023]" pageId="5" pageNumber="190">
|
||
<emphasis id="B8D8EAA3FFDCFFB741C7F8BAFEF8F82A" bold="true" box="[151,275,1909,1931]" pageId="5" pageNumber="190">FIGURE 3.</emphasis>
|
||
(A–C)
|
||
<taxonomicName id="4DAC4D32FFDCFFB74032F8BAFC80F82B" authority="(A. Milne-Edwards, 1880)" authorityName="A. Milne-Edwards" authorityYear="1880" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" box="[354,875,1909,1931]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="ackleianus">
|
||
<emphasis id="B8D8EAA3FFDCFFB74032F8BAFDBFF82B" box="[354,596,1909,1931]" italics="true" pageId="5" pageNumber="190">Pachycheles ackleianus</emphasis>
|
||
(A. Milne-Edwards, 1880)
|
||
</taxonomicName>
|
||
, male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F)
|
||
<taxonomicName id="4DAC4D32FFDCFFB741C7F85BFE04F80A" authority="Haig, 1960" authorityName="Haig" authorityYear="1960" box="[151,495,1940,1962]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDCFFB741C7F85BFE87F80A" box="[151,364,1940,1962]" italics="true" pageId="5" pageNumber="190">Pachycheles velerae</emphasis>
|
||
Haig, 1960
|
||
</taxonomicName>
|
||
, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8A1336B1FFDFFFB44197FEE7FE00FDFF" blockId="6.[151,1437,151,608]" pageId="6" pageNumber="191">
|
||
In the past,
|
||
<taxonomicName id="4DAC4D32FFDFFFB4401DFEE7FDD8FEE0" authority="Ferreira & Tavares, 2019" authorityName="Ferreira & Tavares" authorityYear="2019" box="[333,563,296,320]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="coelhoi" status="sp. nov.">
|
||
<emphasis id="B8D8EAA3FFDFFFB4401DFEE7FDD8FEE0" box="[333,563,296,320]" italics="true" pageId="6" pageNumber="191">Pachycheles coelhoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B8D8EAA3FFDFFFB4436BFEE8FD7DFEE0" bold="true" box="[571,662,295,320]" pageId="6" pageNumber="191">
|
||
<taxonomicNameLabel id="A3EB57D8FFDFFFB4436BFEE8FD7DFEE0" box="[571,662,295,320]" pageId="6" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has been confused with
|
||
<taxonomicName id="4DAC4D32FFDFFFB442E0FEE6FBAFFEE0" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" box="[944,1092,296,320]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="ackleianus">
|
||
<emphasis id="B8D8EAA3FFDFFFB442E0FEE6FBAFFEE0" box="[944,1092,296,320]" italics="true" pageId="6" pageNumber="191">P. ackleianus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="12972A34FFDFFFB44505FEE8FB7DFEE0" box="[1109,1174,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1931]" captionTargetBox="[201,1388,193,1893]" captionTargetId="figure@5.[201,1388,193,1893]" captionTargetPageId="5" captionText="FIGURE 3. (A–C) Pachycheles ackleianus (A. Milne-Edwards, 1880), male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D)." httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="6" pageNumber="191">Fig. 3</figureCitation>
|
||
A–C). The two species are somewhat similar, but can be easily separated in that: (1) the carapace is longer than broad in males and females (
|
||
<figureCitation id="12972A34FFDFFFB441CFFEA0FEEFFE28" box="[159,260,367,392]" captionStart="FIGURE 1" captionStartId="3.[151,250,1888,1910]" captionTargetBox="[329,1288,217,1862]" captionTargetId="figure@3.[208,1381,195,1871]" captionTargetPageId="3" captionText="FIGURE 1. Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (A) carapace, dorsal view. (B) frontal-orbital region of carapace, frontal view. (C) carapace, lateral view, emphasizing the branchiostegite. (D) right antennule, ventral view. (E) right antennal peduncle, dorsal view (flagellum omitted). (F) left Mxp 3, ventral view (exopod flagellum not shown). (G) left minor P1, dorsal view. (H) right major P1, dorsal view. (I) right P2, lateral view. (J) telson, external view. (K) thoracic sternum, ventral view. Setae all omitted. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2599189/files/figure.png" pageId="6" pageNumber="191">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDFFFB44041FEBFFEDDFE28" box="[273,310,368,392]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="6" pageNumber="191">2A</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="4DAC4D32FFDFFFB4409CFEBEFD8AFE28" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" box="[460,609,368,392]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="ackleianus">
|
||
<emphasis id="B8D8EAA3FFDFFFB4409CFEBEFD8AFE28" box="[460,609,368,392]" italics="true" pageId="6" pageNumber="191">P. ackleianus</emphasis>
|
||
</taxonomicName>
|
||
, it is distinctly broader than long in males and females (
|
||
<figureCitation id="12972A34FFDFFFB445B0FEA0FAD0FE28" box="[1248,1339,367,392]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1931]" captionTargetBox="[201,1388,193,1893]" captionTargetId="figure@5.[201,1388,193,1893]" captionTargetPageId="5" captionText="FIGURE 3. (A–C) Pachycheles ackleianus (A. Milne-Edwards, 1880), male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D)." httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="6" pageNumber="191">Fig. 3A</figureCitation>
|
||
); (2) the front is slightly deflexed and distinctly trilobate in dorsal view (
|
||
<figureCitation id="12972A34FFDFFFB4420CFE5BFC54FE0C" box="[860,959,404,429]" captionStart="FIGURE 1" captionStartId="3.[151,250,1888,1910]" captionTargetBox="[329,1288,217,1862]" captionTargetId="figure@3.[208,1381,195,1871]" captionTargetPageId="3" captionText="FIGURE 1. Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (A) carapace, dorsal view. (B) frontal-orbital region of carapace, frontal view. (C) carapace, lateral view, emphasizing the branchiostegite. (D) right antennule, ventral view. (E) right antennal peduncle, dorsal view (flagellum omitted). (F) left Mxp 3, ventral view (exopod flagellum not shown). (G) left minor P1, dorsal view. (H) right major P1, dorsal view. (I) right P2, lateral view. (J) telson, external view. (K) thoracic sternum, ventral view. Setae all omitted. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2599189/files/figure.png" pageId="6" pageNumber="191">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDFFFB4429BFE5BFC1BFE0C" box="[971,1008,404,428]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="6" pageNumber="191">2A</figureCitation>
|
||
), versus distinctly deflexed and nearly straight in dorsal view in
|
||
<taxonomicName id="4DAC4D32FFDFFFB440EBFE76FDBBFE70" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" box="[443,592,440,464]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="ackleianus">
|
||
<emphasis id="B8D8EAA3FFDFFFB440EBFE76FDBBFE70" box="[443,592,440,464]" italics="true" pageId="6" pageNumber="191">P. ackleianus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="12972A34FFDFFFB44330FE78FD50FE70" box="[608,699,439,464]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1931]" captionTargetBox="[201,1388,193,1893]" captionTargetId="figure@5.[201,1388,193,1893]" captionTargetPageId="5" captionText="FIGURE 3. (A–C) Pachycheles ackleianus (A. Milne-Edwards, 1880), male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D)." httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="6" pageNumber="191">Fig. 3A</figureCitation>
|
||
); (3) the P1 carpus and propodus are distinctly granulate and the carpus has 3 pronounced, longitudinal carinae and a series of transverse, discrete ridges between the longitudinal carinae (
|
||
<figureCitation id="12972A34FFDFFFB441AAFE30FE69FDB7" box="[250,386,511,536]" captionStart="FIGURE 1" captionStartId="3.[151,250,1888,1910]" captionTargetBox="[329,1288,217,1862]" captionTargetId="figure@3.[208,1381,195,1871]" captionTargetPageId="3" captionText="FIGURE 1. Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (A) carapace, dorsal view. (B) frontal-orbital region of carapace, frontal view. (C) carapace, lateral view, emphasizing the branchiostegite. (D) right antennule, ventral view. (E) right antennal peduncle, dorsal view (flagellum omitted). (F) left Mxp 3, ventral view (exopod flagellum not shown). (G) left minor P1, dorsal view. (H) right major P1, dorsal view. (I) right P2, lateral view. (J) telson, external view. (K) thoracic sternum, ventral view. Setae all omitted. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2599189/files/figure.png" pageId="6" pageNumber="191">Figs. 1G, H</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDFFFB440C0FDCFFE58FDB8" box="[400,435,512,536]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="6" pageNumber="191">2C</figureCitation>
|
||
,
|
||
<figureCitation id="12972A34FFDFFFB44092FDCFFDE1FDB8" box="[450,522,512,536]" captionStart="FIGURE 4" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[151,1436,661,1843]" captionTargetId="figure@6.[151,1436,661,1843]" captionTargetPageId="6" captionText="FIGURE 4. Major chelae. (A, B) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E)." httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="6" pageNumber="191">4A, B</figureCitation>
|
||
), whereas in
|
||
<emphasis id="B8D8EAA3FFDFFFB443F6FDCEFCA8FDB8" box="[678,835,512,536]" italics="true" pageId="6" pageNumber="191">
|
||
<taxonomicName id="4DAC4D32FFDFFFB443F6FDCEFCABFDB8" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" box="[678,832,512,536]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="ackleianus">P. ackleianus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the carpus and propodus are covered with flattened tubercles grouped together in longitudinal rows (
|
||
<figureCitation id="12972A34FFDFFFB44380FDEBFCC7FD9C" box="[720,812,548,573]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1931]" captionTargetBox="[201,1388,193,1893]" captionTargetId="figure@5.[201,1388,193,1893]" captionTargetPageId="5" captionText="FIGURE 3. (A–C) Pachycheles ackleianus (A. Milne-Edwards, 1880), male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D)." httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="6" pageNumber="191">Fig. 3C</figureCitation>
|
||
); and (4) the male gonopods are present on somite 2 versus absent in
|
||
<taxonomicName id="4DAC4D32FFDFFFB44001FD86FE0FFDC0" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" box="[337,484,584,608]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="ackleianus">
|
||
<emphasis id="B8D8EAA3FFDFFFB44001FD86FE0FFDC0" box="[337,484,584,608]" italics="true" pageId="6" pageNumber="191">P. ackleianus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DED36639FFDFFFB441C7F8F5FDDBF849" httpUri="https://zenodo.org/record/2599195/files/figure.png" pageId="6" pageNumber="191" startId="6.[151,250,1850,1872]" targetBox="[151,1436,661,1843]" targetPageId="6">
|
||
<paragraph id="8A1336B1FFDFFFB441C7F8F5FDDBF849" blockId="6.[151,1436,1850,2025]" pageId="6" pageNumber="191">
|
||
<emphasis id="B8D8EAA3FFDFFFB441C7F8F5FEF8F8EF" bold="true" box="[151,275,1850,1872]" pageId="6" pageNumber="191">FIGURE 4.</emphasis>
|
||
Major chelae. (A, B)
|
||
<taxonomicName id="4DAC4D32FFDFFFB440A7F8F5FD23F8F0" authority="Ferreira & Tavares, 2019" authorityName="Ferreira & Tavares" authorityYear="2019" box="[503,712,1850,1872]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="coelhoi" status="sp. nov.">
|
||
<emphasis id="B8D8EAA3FFDFFFB440A7F8F5FD23F8F0" box="[503,712,1850,1872]" italics="true" pageId="6" pageNumber="191">Pachycheles coelhoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B8D8EAA3FFDFFFB4439FF8F5FCCBF8EF" bold="true" box="[719,800,1850,1871]" pageId="6" pageNumber="191">
|
||
<taxonomicNameLabel id="A3EB57D8FFDFFFB4439FF8F5FCCBF8EF" box="[719,800,1850,1871]" pageId="6" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (C, D)
|
||
<taxonomicName id="4DAC4D32FFDFFFB441E7F897FD50F8CE" authority="(A. Milne-Edwards, 1880)" authorityName="A. Milne-Edwards" authorityYear="1880" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" box="[183,699,1880,1902]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDFFFB441E7F897FE4CF8CE" box="[183,423,1880,1902]" italics="true" pageId="6" pageNumber="191">Pachycheles rugimanus</emphasis>
|
||
(A. Milne-Edwards, 1880)
|
||
</taxonomicName>
|
||
, male, cl 8.0 mm, cw 7.8 mm (USNM 188445), dorsal and ventral views, respectively. (E, F)
|
||
<taxonomicName id="4DAC4D32FFDFFFB4400FF8B8FD43F82D" authority="Haig, 1960" authorityName="Haig" authorityYear="1960" box="[351,680,1911,1933]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDFFFB4400FF8B8FE36F82D" box="[351,477,1911,1933]" italics="true" pageId="6" pageNumber="191">Pachycheles</emphasis>
|
||
<emphasis id="B8D8EAA3FFDFFFB440B4F8B8FDC4F82D" box="[484,559,1911,1933]" italics="true" pageId="6" pageNumber="191">velerae</emphasis>
|
||
Haig, 1960
|
||
</taxonomicName>
|
||
, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, C, E) dorsal view. (B, D, F) ventral view. Scale bars: 1 mm. White arrows: high, heavily tuberculated, elongate upper bulge (A); fixed finger hollowed, interlocking with dactylus (B); narrow, low, rather smooth, elongated upper ridge (E). Black arrows: elongate axial bulge (A); small, rounded bulges (C, E).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8A1336B1FFDEFFB54197FF58FB07FEC4" blockId="7.[151,1437,151,464]" pageId="7" pageNumber="192">
|
||
<bibRefCitation id="EE3D4B40FFDEFFB54197FF58FEBBFF10" author="Haig, J." box="[199,336,151,176]" pageId="7" pageNumber="192" pagination="1 - 440" refId="ref3943" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig (1960)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EE3D4B40FFDEFFB540DAFF57FDC4FF10" author="Harvey, A." box="[394,559,152,176]" pageId="7" pageNumber="192" pagination="746 - 752" refId="ref3973" refString="Harvey, A. (1998) Rediscovery and range extension of the Galapagos " endemic " Pachycheles velerae (Decapoda: Anomura: Porcellanidae). Journal of Crustacean Biology, 18 (4), 746 - 752. https: // doi. org / 10.2307 / 1549151" type="journal article" year="1998">Harvey (1998)</bibRefCitation>
|
||
put forward a number of distinguishing characters between
|
||
<taxonomicName id="4DAC4D32FFDEFFB5458FFF56FAA4FF10" authorityName="Haig" authorityYear="1960" box="[1247,1359,152,176]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDEFFB5458FFF56FAA4FF10" box="[1247,1359,152,176]" italics="true" pageId="7" pageNumber="192">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4DAC4D32FFDEFFB544D7FF56FEE4FF74" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDEFFB544D7FF56FEE4FF74" italics="true" pageId="7" pageNumber="192">P. rugimanus</emphasis>
|
||
</taxonomicName>
|
||
. Additional characters differentiating
|
||
<taxonomicName id="4DAC4D32FFDEFFB543ECFF72FCC0FF74" authorityName="Haig" authorityYear="1960" box="[700,811,188,212]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDEFFB543ECFF72FCC0FF74" box="[700,811,188,212]" italics="true" pageId="7" pageNumber="192">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4DAC4D32FFDEFFB54220FF72FBE8FF74" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" box="[880,1027,189,212]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDEFFB54220FF72FBE8FF74" box="[880,1027,189,212]" italics="true" pageId="7" pageNumber="192">P. rugimanus</emphasis>
|
||
</taxonomicName>
|
||
include the poorly defined carapace regions (
|
||
<figureCitation id="12972A34FFDEFFB541AAFF10FEBFFF58" box="[250,340,223,248]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1931]" captionTargetBox="[201,1388,193,1893]" captionTargetId="figure@5.[201,1388,193,1893]" captionTargetPageId="5" captionText="FIGURE 3. (A–C) Pachycheles ackleianus (A. Milne-Edwards, 1880), male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D)." httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="7" pageNumber="192">Fig. 3D</figureCitation>
|
||
), compared to well defined in
|
||
<taxonomicName id="4DAC4D32FFDEFFB543E2FF2EFCADFF57" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" box="[690,838,224,248]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDEFFB543E2FF2EFCADFF57" box="[690,838,224,248]" italics="true" pageId="7" pageNumber="192">P. rugimanus</emphasis>
|
||
</taxonomicName>
|
||
, especially the protogastric and epibranchial regions (
|
||
<figureCitation id="12972A34FFDEFFB541CFFECBFF17FEBC" box="[159,252,260,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="7" pageNumber="192">Fig. 2D</figureCitation>
|
||
), and the anteriorly trilobate thoracic sternite 3 with the median lobe rounded and separated from the lateral lobes by shallow, gently concave margins (
|
||
<figureCitation id="12972A34FFDEFFB5439BFEE8FCC9FEE0" box="[715,802,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1931]" captionTargetBox="[201,1388,193,1893]" captionTargetId="figure@5.[201,1388,193,1893]" captionTargetPageId="5" captionText="FIGURE 3. (A–C) Pachycheles ackleianus (A. Milne-Edwards, 1880), male, cl 3.6 mm, cw 4.2 mm (MZUSP 12892). (D–F) Pachycheles velerae Haig, 1960, male, cl 5.4 mm, cw 4.9 mm (USNM 110551). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front distinctly deflexed, nearly straight on dorsal view (A); front slightly deflexed, distinctly trilobate in dorsal view (D)." httpUri="https://zenodo.org/record/2599193/files/figure.png" pageId="7" pageNumber="192">Fig. 3E</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="4DAC4D32FFDEFFB542E9FEE6FBA6FE9F" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" box="[953,1101,296,320]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDEFFB542E9FEE6FBA6FE9F" box="[953,1101,296,320]" italics="true" pageId="7" pageNumber="192">P. rugimanus</emphasis>
|
||
</taxonomicName>
|
||
, the median lobe is distinctly subtriangular and separated from the lateral lobes by straight, laterally V-shaped margins (
|
||
<figureCitation id="12972A34FFDEFFB545D8FE83FB34FEC4" box="[1160,1247,332,357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[302,1288,193,1808]" captionTargetId="figure@4.[302,1288,193,1808]" captionTargetPageId="4" captionText="FIGURE 2. (A–C) Pachycheles coelhoi sp. nov., male holotype, cl 6.4 mm, cw 5.9 mm (MOUFPE 7491). (D–F) Pachycheles rugimanus (A. Milne-Edwards, 1880), male, cl 8.0 mm, cw 7.8 mm (USNM 188445). (A, D) carapace, dorsal view. (B, E) cephalothorax and pleon, ventral view. (C, F) minor and major P1, dorsal view. Scale bars: 1 mm. Arrows: front median lobe strongly projected, separated from lateral lobes by broad concave margins (A); front median lobe very low, poorly projected, separated from lateral lobes by nearly straight margins (D); thoracic sternite 3 median lobe rounded, separated from lateral lobes by shallow, gently concave margins (B); thoracic sternite 3 median lobe distinctly subtriangular, separated from lateral lobes by straight, laterally V-shaped margins (E)." httpUri="https://zenodo.org/record/2599191/files/figure.png" pageId="7" pageNumber="192">Fig. 2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8A1336B1FFDEFFB54197FEBFFBE4FE70" blockId="7.[151,1437,151,464]" pageId="7" pageNumber="192">
|
||
<bibRefCitation id="EE3D4B40FFDEFFB54197FEBFFE87FE28" author="Harvey, A." box="[199,364,368,392]" pageId="7" pageNumber="192" pagination="746 - 752" refId="ref3973" refString="Harvey, A. (1998) Rediscovery and range extension of the Galapagos " endemic " Pachycheles velerae (Decapoda: Anomura: Porcellanidae). Journal of Crustacean Biology, 18 (4), 746 - 752. https: // doi. org / 10.2307 / 1549151" type="journal article" year="1998">Harvey (1998)</bibRefCitation>
|
||
considered that
|
||
<taxonomicName id="4DAC4D32FFDEFFB54377FEBEFD7CFE28" authorityName="Haig" authorityYear="1960" box="[551,663,368,392]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDEFFB54377FEBEFD7CFE28" box="[551,663,368,392]" italics="true" pageId="7" pageNumber="192">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4DAC4D32FFDEFFB5439FFEBEFC88FE27" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1880" box="[719,867,368,392]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="rugimanus">
|
||
<emphasis id="B8D8EAA3FFDEFFB5439FFEBEFC88FE27" box="[719,867,368,392]" italics="true" pageId="7" pageNumber="192">P. rugimanus</emphasis>
|
||
</taxonomicName>
|
||
are sister taxa on opposite sides of the Isthmus of
|
||
<collectingCountry id="F2BB7621FFDEFFB541C7FE5AFF1BFE0C" box="[151,240,405,428]" name="Panama" pageId="7" pageNumber="192">Panama</collectingCountry>
|
||
based on an unpublished phylogenetic analysis of
|
||
<taxonomicName id="4DAC4D32FFDEFFB54215FE5BFC3AFE0C" authorityName="Stimpson" authorityYear="1858" box="[837,977,404,428]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B8D8EAA3FFDEFFB54215FE5BFC3AFE0C" box="[837,977,404,428]" italics="true" pageId="7" pageNumber="192">Pachycheles</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4DAC4D32FFDEFFB5455FFE5AFB4DFE0C" authorityName="Haig" authorityYear="1960" box="[1039,1190,405,428]" class="Malacostraca" family="Porcellanidae" genus="Neopisosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B8D8EAA3FFDEFFB5455FFE5AFB4DFE0C" box="[1039,1190,405,428]" italics="true" pageId="7" pageNumber="192">Neopisosoma</emphasis>
|
||
</taxonomicName>
|
||
. Future phylogenetic analysis might well reveal that
|
||
<taxonomicName id="4DAC4D32FFDEFFB540A4FE76FD89FE70" authorityName="Haig" authorityYear="1960" box="[500,610,440,464]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="velerae">
|
||
<emphasis id="B8D8EAA3FFDEFFB540A4FE76FD89FE70" box="[500,610,440,464]" italics="true" pageId="7" pageNumber="192">P. velerae</emphasis>
|
||
</taxonomicName>
|
||
is actually sister to
|
||
<taxonomicName id="4DAC4D32FFDEFFB54211FE76FC40FE70" authority="Ferreira & Tavares, 2019" authorityName="Ferreira & Tavares" authorityYear="2019" box="[833,939,440,464]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="coelhoi" status="sp. nov.">
|
||
<emphasis id="B8D8EAA3FFDEFFB54211FE76FC40FE70" box="[833,939,440,464]" italics="true" pageId="7" pageNumber="192">P. coelhoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B8D8EAA3FFDEFFB542E6FE78FBE4FE70" bold="true" box="[950,1039,439,464]" pageId="7" pageNumber="192">
|
||
<taxonomicNameLabel id="A3EB57D8FFDEFFB542E6FE78FBE4FE70" box="[950,1039,439,464]" pageId="7" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |