346 lines
50 KiB
XML
346 lines
50 KiB
XML
<document id="B8A1D1E522C5AFE712323E49657EF46A" ID-DOI="10.5070/P9401462457" ID-ISSN="0031-0298" ID-Zenodo-Dep="10913330" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1712167746961" checkinUser="felipe" docAuthor="Wheeler, Elisabeth A., Manchester, Steven R. & Baas, Pieter" docDate="2023" docId="038AF505A316960D57D7F8A2FE899A9C" docLanguage="en" docName="PaleoBios.40.14.1-55.pdf" docOrigin="PaleoBios 40 (14)" docSource="http://dx.doi.org/10.5070/p9401462457" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.3:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="3" docTitle="Plerandreoxylon oskolskii Wheeler & Manchester & Baas 2023, SP. NOV." docType="treatment" docVersion="1" lastPageNumber="42" masterDocId="FFB38D7DA33F9621547BFFC0FFD49B4F" masterDocTitle="A late Eocene wood assemblage from the Crooked River Basin, Oregon, USA" masterLastPageNumber="55" masterPageNumber="1" pageNumber="40" updateTime="1728052592909" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="B6EC5CB35C290E7E4C9C523D0119496C">A late Eocene wood assemblage from the Crooked River Basin, Oregon, USA</mods:title>
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<mods:namePart id="567F5F4828613271A1486BCFD3E460DA">Wheeler, Elisabeth A.</mods:namePart>
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<mods:namePart id="0CAE3BFFB64154729BC13FC10E0A23C1">Manchester, Steven R.</mods:namePart>
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<mods:namePart id="5848F7EF77740249ECD41DF2B8391114">Baas, Pieter</mods:namePart>
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<mods:typeOfResource id="E04FFC8EF841EF60792B82C67169426A">text</mods:typeOfResource>
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<mods:title id="18E09A588DD9B5262F70830D917D5BD5">PaleoBios</mods:title>
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<mods:part id="A99B5C4172BB0CAA72D51ECF9702A7E3">
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<mods:date id="712BA627931383969B567C125C65EE1D">2023</mods:date>
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<mods:number id="45A916DBA237276FEBA12C5D5F14937C">2023-11-01</mods:number>
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<mods:number id="BD5ADCB4FBF35783429720AA8D3FC960">14</mods:number>
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<mods:url id="E10C4B6D631C40EB7B4E8DDB41C859F6">http://dx.doi.org/10.5070/p9401462457</mods:url>
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<mods:classification id="5A47E446C847A89E02DAB2CA53F7DF46">journal article</mods:classification>
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<mods:identifier id="6381A23F2D6AC58F5F0BEF648792D461" type="DOI">10.5070/P9401462457</mods:identifier>
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<treatment id="038AF505A316960D57D7F8A2FE899A9C" LSID="urn:lsid:plazi:treatment:038AF505A316960D57D7F8A2FE899A9C" httpUri="http://treatment.plazi.org/id/038AF505A316960D57D7F8A2FE899A9C" lastPageId="44" lastPageNumber="42" pageId="41" pageNumber="40">
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<subSubSection id="C3391798A316960857D7F8A2FA519C31" box="[940,1413,1890,1918]" pageId="41" pageNumber="40" type="nomenclature">
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<paragraph id="8B9C4413A316960857D7F8A2FA519C31" blockId="41.[940,1413,1890,1953]" box="[940,1413,1890,1918]" pageId="41" pageNumber="40">
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<heading id="D0D4F37FA316960857D7F8A2FA519C31" allCaps="true" box="[940,1413,1890,1918]" centered="true" fontSize="10" level="2" pageId="41" pageNumber="40" reason="2">
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<taxonomicName id="4C233F90A316960857D7F8A2FAFF9C31" authority="Wheeler & Manchester & Baas, 2023" authorityName="Wheeler & Manchester & Baas" authorityYear="2023" box="[940,1323,1890,1918]" class="Magnoliopsida" family="Araliaceae" genus="Plerandreoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="41" pageNumber="1" phylum="Tracheophyta" rank="species" species="oskolskii" status="sp. nov.">
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<emphasis id="B9579801A316960857D7F8A2FAFF9C31" box="[940,1323,1890,1918]" italics="true" pageId="41" pageNumber="40">PLERANDREOXYLON OSKOLSKII</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A264257AA31696085149F8A6FA519C31" box="[1330,1413,1894,1918]" pageId="41" pageNumber="40" rank="species">
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<collectionCode id="ED32DCD6A31696085149F8A6FA989C31" box="[1330,1356,1894,1918]" country="Brazil" name="Instituto de Botânica" pageId="41" pageNumber="40" type="Herbarium">SP</collectionCode>
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. NOV.
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</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3391798A31696085020F849FB019CEE" box="[1115,1237,1929,1953]" pageId="41" pageNumber="40" type="description">
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<paragraph id="8B9C4413A31696085020F849FB019CEE" blockId="41.[940,1413,1890,1953]" box="[1115,1237,1929,1953]" pageId="41" pageNumber="40">
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<figureCitation id="13185896A31696085020F849FB019CEE" box="[1115,1237,1929,1953]" captionStart="Figure 19" captionStartId="43.[112,187,1396,1420]" captionTargetBox="[147,1488,194,1368]" captionTargetId="figure-318@43.[147,1488,194,1371]" captionTargetPageId="43" captionText="Figure 19. Araliaceae. Plerandreoxylon oskolski sp. nov., UF 278-84906. A. Semi-ring-porous wood with latewood vessels arranged in wavy tangential bands/diagonal arrangement, vessel clusters in latest latewood, axial parenchyma rare, T.S. B. Semi-ring-porous wood, vessels solitary and in short radial multiples, axial parenchyma rare, TS. C. Simple perforation plates, rays with procumbent body cells, RLS. D. Crowded alternate intervessel pitting, TLS. E. Vessel-ray parenchyma pits with reduced borders, oval to slightly horizontally elongate in outline, tyloses, RLS. F. Rays 5-6 cells wide, septate fibers, TLS. G. Rays predominantly multiseriate, TLS. Scale bars: 200 µm in A, B, G; 100 µm in C, F. 50 µm in E; 20 µm in D." figureDoi="http://doi.org/10.5281/zenodo.10913368" httpUri="https://zenodo.org/record/10913368/files/figure.png" pageId="41" pageNumber="40">FIG. 19A–G</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3391798A315960B54EBFF03FDE49835" pageId="42" pageNumber="41" type="diagnosis">
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<paragraph id="8B9C4413A315960B54EBFF03FED19A09" blockId="42.[112,801,195,1947]" pageId="42" pageNumber="41">
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<emphasis id="B9579801A315960B54EBFF03FEE59B92" bold="true" box="[144,305,195,221]" pageId="42" pageNumber="41">Diagnosis—</emphasis>
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Growth rings distinct, marked by radially flattened latewood fibers, and differences in vessel diameter between latewood and earlywood of subsequent rings.
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</paragraph>
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<paragraph id="8B9C4413A315960B54EBFE90FE66992E" blockId="42.[112,801,195,1947]" pageId="42" pageNumber="41">Semi-ring-porous to ring-porous. Vessels solitary and in radial multiples, small clusters in the latewood. Earlywood with multiple rows of vessels, latewood vessels arranged in a tangential to diagonal pattern in narrower rings. Perforation plates simple, intervessel pits alternate, medium to large; vessel-ray parenchyma pits with reduced borders to simple, oval to slightly horizontally elongate. Tyloses present.</paragraph>
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<paragraph id="8B9C4413A315960B54EBFDA9FEAE99E8" blockId="42.[112,801,195,1947]" pageId="42" pageNumber="41">Fibers with simple to minutely bordered pits; thin to thick-walled; septate.</paragraph>
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<paragraph id="8B9C4413A315960B54EBFD70FDE09985" blockId="42.[112,801,195,1947]" box="[144,564,688,714]" pageId="42" pageNumber="41">Axial parenchyma extremely rare.</paragraph>
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<paragraph id="8B9C4413A315960B54EBFD13FDE49835" blockId="42.[112,801,195,1947]" pageId="42" pageNumber="41">
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Rays predominantly multiseriate, sheath cells occasional, average ray height <
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<quantity id="4CDBE9F6A315960B559EFD36FDFF985E" box="[485,555,758,785]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="41" unit="mm" value="1.0">1 mm</quantity>
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. Rays homocellular, composed of procumbent body cells, or heterocellular with a single marginal row of square cells and weakly procumbent cells. Rays per mm few.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3391798A315960B54EBFC43FDFC9F48" pageId="42" pageNumber="41" type="materials_examined">
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<paragraph id="8B9C4413A315960B54EBFC43FEE5988E" blockId="42.[112,801,195,1947]" pageId="42" pageNumber="41">
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<materialsCitation id="3B4B4E4EA315960B54EBFC43FEE5988E" collectionCode="UF" pageId="42" pageNumber="41" specimenCode="UF 278-84872" specimenCount="1" typeStatus="holotype">
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<emphasis id="B9579801A315960B54EBFC43FEFD98D2" bold="true" box="[144,297,899,925]" pageId="42" pageNumber="41">
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<typeStatus id="5498FAB1A315960B54EBFC43FEDD98D2" box="[144,265,899,925]" pageId="42" pageNumber="41" type="holotype">Holotype</typeStatus>
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—
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</emphasis>
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<specimenCode id="DB85EC68A315960B5553FC43FE3698D2" box="[296,482,899,925]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="42" pageNumber="41" type="Museum">UF 278-84872</specimenCode>
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, estimated maximum diameter
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<quantity id="4CDBE9F6A315960B54ABFC66FEF9988E" box="[208,301,934,961]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="42" pageNumber="41" unit="cm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 cm</quantity>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B9C4413A315960B54EBFC09FE3598AB" blockId="42.[112,801,195,1947]" box="[144,481,969,996]" pageId="42" pageNumber="41">
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<materialsCitation id="3B4B4E4EA315960B54EBFC09FE0A98AB" box="[144,478,969,996]" collectionCode="UF" pageId="42" pageNumber="41" specimenCode="UF 278-84905" specimenCount="1" typeStatus="paratype">
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<emphasis id="B9579801A315960B54EBFC09FEF298AC" bold="true" box="[144,294,969,995]" pageId="42" pageNumber="41">
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<typeStatus id="5498FAB1A315960B54EBFC09FED398AC" box="[144,263,969,995]" pageId="42" pageNumber="41" type="paratype">Paratype</typeStatus>
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||
—
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</emphasis>
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||
<specimenCode id="DB85EC68A315960B555DFC0AFE0A98AB" box="[294,478,969,996]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="42" pageNumber="41" type="Museum">UF 278-84905</specimenCode>
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</materialsCitation>
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||
.
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||
</paragraph>
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<paragraph id="8B9C4413A315960B54EBFC2CFDFC9F48" blockId="42.[112,801,195,1947]" box="[144,552,1004,1031]" pageId="42" pageNumber="41">
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<emphasis id="B9579801A315960B54EBFC2CFE939F49" bold="true" box="[144,327,1004,1030]" pageId="42" pageNumber="41">Occurrence—</emphasis>
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Dietz Hill,
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<collectionCode id="ED32DCD6A315960B55B3FC2DFE3F9F48" box="[456,491,1005,1031]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="42" pageNumber="41" type="Museum">UF</collectionCode>
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278.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3391798A315960B54EBFBD0FF009FDC" pageId="42" pageNumber="41" type="etymology">
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<paragraph id="8B9C4413A315960B54EBFBD0FF009FDC" blockId="42.[112,801,195,1947]" pageId="42" pageNumber="41">
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<emphasis id="B9579801A315960B54EBFBD0FE909F65" bold="true" box="[144,324,1040,1066]" pageId="42" pageNumber="41">Etymology—</emphasis>
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Named for our good colleague and friend Alexei Oskolski who has studied
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<taxonomicName id="4C233F90A315960B5629FBF3FD019F02" box="[594,725,1075,1101]" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="family">Araliaceae</taxonomicName>
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wood and done much to further the field of wood anatomy in general.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3391798A315960D54EBFB5CFE899A9C" lastPageId="44" lastPageNumber="43" pageId="42" pageNumber="41" type="description">
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<paragraph id="8B9C4413A315960B54EBFB5CFDAA9DCF" blockId="42.[112,801,195,1947]" pageId="42" pageNumber="41">
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<emphasis id="B9579801A315960B54EBFB5CFE989FF9" bold="true" box="[144,332,1180,1206]" pageId="42" pageNumber="41">Description—</emphasis>
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Growth rings distinct, marked by radially flattened latewood fibers. Ring-porous to semi-ring-porous. Vessels solitary and in short radial multiples, small clusters in the latewood. Earlywood with multiple rows of vessels; latewood vessels arranged in a tangential to diagonal pattern in narrower rings, mostly in radial multiples, solitary vessels not common (
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<figureCitation id="13185896A315960B5601FAB0FCD99EC5" box="[634,781,1392,1418]" captionStart="Figure 19" captionStartId="43.[112,187,1396,1420]" captionTargetBox="[147,1488,194,1368]" captionTargetId="figure-318@43.[147,1488,194,1371]" captionTargetPageId="43" captionText="Figure 19. Araliaceae. Plerandreoxylon oskolski sp. nov., UF 278-84906. A. Semi-ring-porous wood with latewood vessels arranged in wavy tangential bands/diagonal arrangement, vessel clusters in latest latewood, axial parenchyma rare, T.S. B. Semi-ring-porous wood, vessels solitary and in short radial multiples, axial parenchyma rare, TS. C. Simple perforation plates, rays with procumbent body cells, RLS. D. Crowded alternate intervessel pitting, TLS. E. Vessel-ray parenchyma pits with reduced borders, oval to slightly horizontally elongate in outline, tyloses, RLS. F. Rays 5-6 cells wide, septate fibers, TLS. G. Rays predominantly multiseriate, TLS. Scale bars: 200 µm in A, B, G; 100 µm in C, F. 50 µm in E; 20 µm in D." figureDoi="http://doi.org/10.5281/zenodo.10913368" httpUri="https://zenodo.org/record/10913368/files/figure.png" pageId="42" pageNumber="41">Fig. 19A, B</figureCitation>
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||
.). Earlywood vessels 97 (
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<collectionCode id="ED32DCD6A315960B55EEFA53FE699EE2" box="[405,445,1427,1453]" country="USA" lsid="urn:lsid:biocol.org:col:15659" name="San Diego Natural History Museum" pageId="42" pageNumber="41" type="Herbarium">SD</collectionCode>
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=22) µm, range 48–137 µm. Vessel element lengths commonly 350–500 µm. Perforation plates simple (
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<figureCitation id="13185896A315960B551BFA19FE189EBC" box="[352,460,1497,1523]" captionStart="Figure 19" captionStartId="43.[112,187,1396,1420]" captionTargetBox="[147,1488,194,1368]" captionTargetId="figure-318@43.[147,1488,194,1371]" captionTargetPageId="43" captionText="Figure 19. Araliaceae. Plerandreoxylon oskolski sp. nov., UF 278-84906. A. Semi-ring-porous wood with latewood vessels arranged in wavy tangential bands/diagonal arrangement, vessel clusters in latest latewood, axial parenchyma rare, T.S. B. Semi-ring-porous wood, vessels solitary and in short radial multiples, axial parenchyma rare, TS. C. Simple perforation plates, rays with procumbent body cells, RLS. D. Crowded alternate intervessel pitting, TLS. E. Vessel-ray parenchyma pits with reduced borders, oval to slightly horizontally elongate in outline, tyloses, RLS. F. Rays 5-6 cells wide, septate fibers, TLS. G. Rays predominantly multiseriate, TLS. Scale bars: 200 µm in A, B, G; 100 µm in C, F. 50 µm in E; 20 µm in D." figureDoi="http://doi.org/10.5281/zenodo.10913368" httpUri="https://zenodo.org/record/10913368/files/figure.png" pageId="42" pageNumber="41">Fig. 19C</figureCitation>
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||
). Intervessel pits alternate, 8–10 µm in horizontal diameter, tending to be angular in outline (
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<figureCitation id="13185896A315960B5487F9E0FEB89D75" box="[252,364,1568,1594]" captionStart="Figure 19" captionStartId="43.[112,187,1396,1420]" captionTargetBox="[147,1488,194,1368]" captionTargetId="figure-318@43.[147,1488,194,1371]" captionTargetPageId="43" captionText="Figure 19. Araliaceae. Plerandreoxylon oskolski sp. nov., UF 278-84906. A. Semi-ring-porous wood with latewood vessels arranged in wavy tangential bands/diagonal arrangement, vessel clusters in latest latewood, axial parenchyma rare, T.S. B. Semi-ring-porous wood, vessels solitary and in short radial multiples, axial parenchyma rare, TS. C. Simple perforation plates, rays with procumbent body cells, RLS. D. Crowded alternate intervessel pitting, TLS. E. Vessel-ray parenchyma pits with reduced borders, oval to slightly horizontally elongate in outline, tyloses, RLS. F. Rays 5-6 cells wide, septate fibers, TLS. G. Rays predominantly multiseriate, TLS. Scale bars: 200 µm in A, B, G; 100 µm in C, F. 50 µm in E; 20 µm in D." figureDoi="http://doi.org/10.5281/zenodo.10913368" httpUri="https://zenodo.org/record/10913368/files/figure.png" pageId="42" pageNumber="41">Fig. 19D</figureCitation>
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); vessel-ray parenchyma pits with reduced borders and oval to slightly horizontally elongate (
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<figureCitation id="13185896A315960B5402F9A6FF379DCF" box="[121,227,1638,1664]" captionStart="Figure 19" captionStartId="43.[112,187,1396,1420]" captionTargetBox="[147,1488,194,1368]" captionTargetId="figure-318@43.[147,1488,194,1371]" captionTargetPageId="43" captionText="Figure 19. Araliaceae. Plerandreoxylon oskolski sp. nov., UF 278-84906. A. Semi-ring-porous wood with latewood vessels arranged in wavy tangential bands/diagonal arrangement, vessel clusters in latest latewood, axial parenchyma rare, T.S. B. Semi-ring-porous wood, vessels solitary and in short radial multiples, axial parenchyma rare, TS. C. Simple perforation plates, rays with procumbent body cells, RLS. D. Crowded alternate intervessel pitting, TLS. E. Vessel-ray parenchyma pits with reduced borders, oval to slightly horizontally elongate in outline, tyloses, RLS. F. Rays 5-6 cells wide, septate fibers, TLS. G. Rays predominantly multiseriate, TLS. Scale bars: 200 µm in A, B, G; 100 µm in C, F. 50 µm in E; 20 µm in D." figureDoi="http://doi.org/10.5281/zenodo.10913368" httpUri="https://zenodo.org/record/10913368/files/figure.png" pageId="42" pageNumber="41">Fig. 19E</figureCitation>
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). Tyloses present (
|
||
<figureCitation id="13185896A315960B55B1F9A6FDA69DCF" box="[458,626,1638,1665]" captionStart="Figure 19" captionStartId="43.[112,187,1396,1420]" captionTargetBox="[147,1488,194,1368]" captionTargetId="figure-318@43.[147,1488,194,1371]" captionTargetPageId="43" captionText="Figure 19. Araliaceae. Plerandreoxylon oskolski sp. nov., UF 278-84906. A. Semi-ring-porous wood with latewood vessels arranged in wavy tangential bands/diagonal arrangement, vessel clusters in latest latewood, axial parenchyma rare, T.S. B. Semi-ring-porous wood, vessels solitary and in short radial multiples, axial parenchyma rare, TS. C. Simple perforation plates, rays with procumbent body cells, RLS. D. Crowded alternate intervessel pitting, TLS. E. Vessel-ray parenchyma pits with reduced borders, oval to slightly horizontally elongate in outline, tyloses, RLS. F. Rays 5-6 cells wide, septate fibers, TLS. G. Rays predominantly multiseriate, TLS. Scale bars: 200 µm in A, B, G; 100 µm in C, F. 50 µm in E; 20 µm in D." figureDoi="http://doi.org/10.5281/zenodo.10913368" httpUri="https://zenodo.org/record/10913368/files/figure.png" pageId="42" pageNumber="41">Fig. 19C, E, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A315960B54EBF949FE249D89" blockId="42.[112,801,195,1947]" pageId="42" pageNumber="41">
|
||
Fibers with simple to minutely bordered pits, septate (
|
||
<figureCitation id="13185896A315960B5402F96CFF359D88" box="[121,225,1708,1735]" captionStart="Figure 19" captionStartId="43.[112,187,1396,1420]" captionTargetBox="[147,1488,194,1368]" captionTargetId="figure-318@43.[147,1488,194,1371]" captionTargetPageId="43" captionText="Figure 19. Araliaceae. Plerandreoxylon oskolski sp. nov., UF 278-84906. A. Semi-ring-porous wood with latewood vessels arranged in wavy tangential bands/diagonal arrangement, vessel clusters in latest latewood, axial parenchyma rare, T.S. B. Semi-ring-porous wood, vessels solitary and in short radial multiples, axial parenchyma rare, TS. C. Simple perforation plates, rays with procumbent body cells, RLS. D. Crowded alternate intervessel pitting, TLS. E. Vessel-ray parenchyma pits with reduced borders, oval to slightly horizontally elongate in outline, tyloses, RLS. F. Rays 5-6 cells wide, septate fibers, TLS. G. Rays predominantly multiseriate, TLS. Scale bars: 200 µm in A, B, G; 100 µm in C, F. 50 µm in E; 20 µm in D." figureDoi="http://doi.org/10.5281/zenodo.10913368" httpUri="https://zenodo.org/record/10913368/files/figure.png" pageId="42" pageNumber="41">Fig. 19F</figureCitation>
|
||
), thin to thick-walled.
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A315960B54EBF910FDE09DA5" blockId="42.[112,801,195,1947]" box="[144,564,1744,1770]" pageId="42" pageNumber="41">Axial parenchyma extremely rare.</paragraph>
|
||
<paragraph id="8B9C4413A315960B54EBF933FBCD9B92" blockId="42.[112,801,195,1947]" lastBlockId="42.[832,1521,195,1946]" pageId="42" pageNumber="41">
|
||
Rays homocellular, composed of procumbent body cells, or heterocellular with a single marginal row of square cells and barely procumbent cells; sheath cells occasional; 3–7 cells wide, mostly 5–6 cells (
|
||
<figureCitation id="13185896A315960B56EBF89CFCC09C38" box="[656,788,1884,1911]" captionStart="Figure 19" captionStartId="43.[112,187,1396,1420]" captionTargetBox="[147,1488,194,1368]" captionTargetId="figure-318@43.[147,1488,194,1371]" captionTargetPageId="43" captionText="Figure 19. Araliaceae. Plerandreoxylon oskolski sp. nov., UF 278-84906. A. Semi-ring-porous wood with latewood vessels arranged in wavy tangential bands/diagonal arrangement, vessel clusters in latest latewood, axial parenchyma rare, T.S. B. Semi-ring-porous wood, vessels solitary and in short radial multiples, axial parenchyma rare, TS. C. Simple perforation plates, rays with procumbent body cells, RLS. D. Crowded alternate intervessel pitting, TLS. E. Vessel-ray parenchyma pits with reduced borders, oval to slightly horizontally elongate in outline, tyloses, RLS. F. Rays 5-6 cells wide, septate fibers, TLS. G. Rays predominantly multiseriate, TLS. Scale bars: 200 µm in A, B, G; 100 µm in C, F. 50 µm in E; 20 µm in D." figureDoi="http://doi.org/10.5281/zenodo.10913368" httpUri="https://zenodo.org/record/10913368/files/figure.png" pageId="42" pageNumber="41">Fig. 19F, G</figureCitation>
|
||
). Ray height averages 449 (126) µm, range 126–728 µm; 3–4 rays per mm.
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A315960B571BFF26FBEA9A6C" blockId="42.[832,1521,195,1946]" pageId="42" pageNumber="41">Crystals, oil/mucilage cells, storied structure, and radial canals absent.</paragraph>
|
||
<paragraph id="8B9C4413A315960B571BFEECFB1C9E6F" blockId="42.[832,1521,195,1946]" pageId="42" pageNumber="41">
|
||
<emphasis id="B9579801A315960B571BFEECFA139A08" bold="true" box="[864,1479,300,327]" pageId="42" pageNumber="41">Comparisons to extant and fossil woods—</emphasis>
|
||
We searched the InsideWood database multiple times. One search used: distinct growth ring boundaries (1p); wood ring-porous to semi-ring porous (5a, not diffuse-porous); vessels in wavy tangential bands (6p);simple perforation plates (13p); intervessel pits alternate and not minute (22p 24a); vessel-ray parenchyma pits rounded with much reduced borders (30a, 31p); septate fibers (65p); axial parenchyma rare (75p); rays more than 4-seriate, <
|
||
<quantity id="4CDBE9F6A315960B572EFDA9FC4D99CB" box="[853,921,617,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="41" unit="mm" value="1.0">1 mm</quantity>
|
||
high and with one row of square marginal cells (98p 102a 106p); storied structure, oil/mucilage cells, and radial canals absent (118a 120a 124a 125a 126a 130a). The IW matches only included
|
||
<taxonomicName id="4C233F90A315960B516DFD13FA4D99A2" box="[1302,1433,723,749]" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="family">Araliaceae</taxonomicName>
|
||
(order
|
||
<taxonomicName id="4C233F90A315960B573BFD36FC7C985E" authorityName="NAKAI" authorityYear="1930" box="[832,936,758,785]" class="Magnoliopsida" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="order">Apiales</taxonomicName>
|
||
): two modern (
|
||
<taxonomicName id="4C233F90A315960B5001FD35FC26987C" authority="J. Wen (1994)" authorityName="J. Wen" authorityYear="1994" class="Magnoliopsida" family="Araliaceae" genus="Aralia" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="variety" species="elata" variety="ryukyuensis">
|
||
<emphasis id="B9579801A315960B5001FD35FAC0985E" box="[1146,1300,757,785]" italics="true" pageId="42" pageNumber="41">Aralia elata</emphasis>
|
||
var.
|
||
<emphasis id="B9579801A315960B5123FD36FA3B985D" box="[1368,1519,758,786]" italics="true" pageId="42" pageNumber="41">ryukyuensis</emphasis>
|
||
J.Wen (1994)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C233F90A315960B5041FCD9FC249818" authority="(Thunb.) Koidz. (1925)" authorityName="Koidz." authorityYear="1925" baseAuthorityName="Thunb." class="Magnoliopsida" family="Araliaceae" genus="Kalopanax" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="species" species="septemlobus">
|
||
<emphasis id="B9579801A315960B5041FCD9FAA4987C" box="[1082,1392,793,819]" italics="true" pageId="42" pageNumber="41">Kalopanax septemlobus</emphasis>
|
||
(Thunb.) Koidz. (1925)
|
||
</taxonomicName>
|
||
, both East Asian) and two fossil woods (
|
||
<taxonomicName id="4C233F90A315960B5730FCA0FC5698D2" authority="Wheeler and Manchester, 2002" authorityName="Wheeler" authorityYear="2002" class="Magnoliopsida" family="Araliaceae" genus="Plerandreoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="species" species="oregonensis">
|
||
<emphasis id="B9579801A315960B5730FCA0FB7F9833" box="[843,1195,864,892]" italics="true" pageId="42" pageNumber="41">Plerandreoxylon oregonensis</emphasis>
|
||
<bibRefCitation id="EFB239E2A315960B50C8FCA0FC5698D2" author="Wheeler, E. A. & S. R. Manchester" pageId="42" pageNumber="41" pagination="1 - 188" refId="ref41110" refString="Wheeler, E. A. and S. R. Manchester. 2002. Woods of the Eocene Nut Beds flora, Clarno Formation, Oregon, USA. IAWA Journal Supplement 3: 1 - 188." type="journal article" year="2002">Wheeler and Manchester, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C233F90A315960B57BEFC43FA8D98D2" authority="Platen, 1908" authorityName="Platen" authorityYear="1908" box="[965,1369,899,925]" class="Magnoliopsida" family="Araliaceae" genus="Aralinium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="species" species="excellens">
|
||
<emphasis id="B9579801A315960B57BEFC43FB6098D2" box="[965,1204,899,925]" italics="true" pageId="42" pageNumber="41">Aralinium excellens</emphasis>
|
||
<bibRefCitation id="EFB239E2A315960B50C0FC43FA8D98D2" author="Platen, P." box="[1211,1369,899,925]" pageId="42" pageNumber="41" pagination="161 - 164" refId="ref38303" refString="Platen, P. 1908. Untersuchungen fossiler Holzer aus dem Westen der vereinigten Staaten von Nordamerika. Naturforschende Gesellschaft zu Leipzig Sitzungsberichte 34: 1 - 155, 161 - 164." type="journal article" year="1908">Platen, 1908</bibRefCitation>
|
||
</taxonomicName>
|
||
), both from the Eocene of the
|
||
<collectingCountry id="F3340483A315960B505AFC66FB83988F" box="[1057,1111,934,960]" name="United States of America" pageId="42" pageNumber="41">USA</collectingCountry>
|
||
. These two Dietz Hill woods' absence of uniseriate rays and the occasional occurrence of sheath cells are additional features that are consistent with the
|
||
<taxonomicName id="4C233F90A315960B57CDFBD0FBE89F65" box="[950,1084,1040,1066]" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="family">Araliaceae</taxonomicName>
|
||
. Unfortunately, Platen’s
|
||
<taxonomicName id="4C233F90A315960B510FFBD0FC789F02" authorityName="Platen" authorityYear="1908" class="Magnoliopsida" family="Araliaceae" genus="Aralinium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="species" species="excellens">
|
||
<emphasis id="B9579801A315960B510FFBD0FC789F02" italics="true" pageId="42" pageNumber="41">Aralinium excellens</emphasis>
|
||
</taxonomicName>
|
||
cannot be placed with certainty in the
|
||
<taxonomicName id="4C233F90A315960B51E7FBF3FCAC9F3E" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="family">Araliaceae</taxonomicName>
|
||
and reasonably compared with this Dietz Hill (
|
||
<collectionCode id="ED32DCD6A315960B51B1FB96FA249F3F" box="[1482,1520,1110,1136]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="42" pageNumber="41" type="Museum">UF</collectionCode>
|
||
278) wood because its description lacks details on ray cellular composition and height and whether fibers are septate or non-septate (
|
||
<bibRefCitation id="EFB239E2A315960B5014FB00FADC9F95" author="Platen, P." box="[1135,1288,1216,1242]" pageId="42" pageNumber="41" pagination="161 - 164" refId="ref38303" refString="Platen, P. 1908. Untersuchungen fossiler Holzer aus dem Westen der vereinigten Staaten von Nordamerika. Naturforschende Gesellschaft zu Leipzig Sitzungsberichte 34: 1 - 155, 161 - 164." type="journal article" year="1908">Platen 1908</bibRefCitation>
|
||
). Moreover, it was described as having crystals in enlarged cells, a feature not reported for the
|
||
<taxonomicName id="4C233F90A315960B5044FAC6FB109E6F" box="[1087,1220,1286,1312]" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="family">Araliaceae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A315960B571BFAE9FB8F9C39" blockId="42.[832,1521,195,1946]" pageId="42" pageNumber="41">
|
||
Another search used absence of vessel-ray parenchyma pits similar to intervessel pits (30a) rather than 31p and absence of obvious axial parenchyma features (77a, 79–86a) rather than
|
||
<specimenCount id="9D258F9AA315960B50F1FA53FB149EE1" box="[1162,1216,1427,1454]" count="75" pageId="42" pageNumber="41" type="generic">75p.</specimenCount>
|
||
This search returned an additional two modern woods (
|
||
<taxonomicName id="4C233F90A315960B50B5FA75FAA99E9E" box="[1230,1405,1461,1489]" class="Magnoliopsida" family="Araliaceae" genus="Aralia" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="species" species="spinosa">
|
||
<emphasis id="B9579801A315960B50B5FA75FAA99E9E" box="[1230,1405,1461,1489]" italics="true" pageId="42" pageNumber="41">Aralia spinosa</emphasis>
|
||
</taxonomicName>
|
||
<collectionCode id="ED32DCD6A315960B51FFFA77FA409E9E" box="[1412,1428,1463,1489]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="42" pageNumber="41" type="Herbarium">L</collectionCode>
|
||
., 1753;
|
||
<taxonomicName id="4C233F90A315960B573BFA19FB389EBC" authority="C. B. Clarke, 1879" authorityName="C. B. Clarke" authorityYear="1879" box="[832,1260,1497,1523]" class="Magnoliopsida" family="Araliaceae" genus="Gamblea" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="species" species="ciliata">
|
||
<emphasis id="B9579801A315960B573BFA19FBD19EBC" box="[832,1029,1497,1523]" italics="true" pageId="42" pageNumber="41">Gamblea ciliata</emphasis>
|
||
C.B. Clarke, 1879
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C233F90A315960B5085FA19FA529EBB" box="[1278,1414,1497,1524]" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="family">Araliaceae</taxonomicName>
|
||
) and an additional fossil wood (
|
||
<taxonomicName id="4C233F90A315960B501CFA3BFA3E9D59" authority="Gregory, 1976" authorityName="Gregory" authorityYear="1976" box="[1127,1514,1531,1561]" class="Magnoliopsida" family="Rutaceae" genus="Evodia" kingdom="Plantae" order="Sapindales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="species" species="gadjirian">
|
||
<emphasis id="B9579801A315960B501CFA3BFAE79D56" box="[1127,1331,1531,1561]" italics="true" pageId="42" pageNumber="41">Evodia gadjirian</emphasis>
|
||
<bibRefCitation id="EFB239E2A315960B5142FA3DFA3E9D59" author="Gregory, I." box="[1337,1514,1532,1559]" pageId="42" pageNumber="41" pagination="135 - 139" refId="ref35240" refString="Gregory, I. 1976. An extinct Evodia wood from Oregon. The Ore Bin 38 (9): 135 - 139." type="journal article" year="1976">Gregory, 1976</bibRefCitation>
|
||
</taxonomicName>
|
||
, from an unspecified Eocene site in the vicinity of Post,
|
||
<collectingRegion id="49E78AF1A315960B573BF983FC499D12" box="[832,925,1603,1629]" country="United States of America" name="Oregon" pageId="42" pageNumber="41">Oregon</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3340483A315960B57DDF983FC349D12" box="[934,992,1603,1629]" name="United States of America" pageId="42" pageNumber="41">USA</collectingCountry>
|
||
). Unfortunately, the description of the putative
|
||
<taxonomicName id="4C233F90A315960B570EF9A5FB289DCE" authority="Forst. and Forst. (1775)" authorityName="Forst. and Forst." authorityYear="1775" box="[885,1276,1637,1665]" class="Magnoliopsida" family="Rutaceae" genus="Evodia" kingdom="Plantae" order="Sapindales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A315960B570EF9A5FC1D9DCE" box="[885,969,1637,1665]" italics="true" pageId="42" pageNumber="41">Evodia</emphasis>
|
||
Forst. and Forst. (1775)
|
||
</taxonomicName>
|
||
lacked information on intervessel and vessel-ray parenchyma pitting and ray cellular composition; the accompanying images were at too low of a magnification to determine those details. Whether or not this wood truly is
|
||
<taxonomicName id="4C233F90A315960B5091F931FAEA9C42" box="[1258,1342,1777,1805]" class="Magnoliopsida" family="Rutaceae" genus="Evodia" kingdom="Plantae" order="Sapindales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A315960B5091F931FAEA9C42" box="[1258,1342,1777,1805]" italics="true" pageId="42" pageNumber="41">Evodia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C233F90A315960B5136F933FA109C42" box="[1357,1476,1779,1805]" class="Magnoliopsida" family="Rutaceae" kingdom="Plantae" order="Sapindales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="family">Rutaceae</taxonomicName>
|
||
) or might be
|
||
<taxonomicName id="4C233F90A315960B57C8F8D6FBE29C7F" box="[947,1078,1814,1840]" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="42" pageNumber="41" phylum="Tracheophyta" rank="family">Araliaceae</taxonomicName>
|
||
is impossible to determine because the location of Gregory’s material is unknown and so it cannot be reexamined.
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A315960A571BF840FBC59DC7" blockId="42.[832,1521,195,1946]" lastBlockId="43.[833,1522,1611,1954]" lastPageId="43" lastPageNumber="42" pageId="42" pageNumber="41">
|
||
Alternative searches, e.g., 1p, 4p, 9a, 10a, 13p, 26p, 30a, 31p, 56p, 61p, 65p, 98p, 106p, in addition to
|
||
<taxonomicName id="4C233F90A314960A5691F98BFF1D9DC7" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="43" pageNumber="42" phylum="Tracheophyta" rank="family">Araliaceae</taxonomicName>
|
||
, also retrieved two extant species of
|
||
<taxonomicName id="4C233F90A314960A56D6F9ADFF259DE3" authority="DC (1830)" authorityName="DC" authorityYear="1830" class="Magnoliopsida" family="Cornaceae" genus="Toricellia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cornales" pageId="43" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A314960A56D6F9ADFCF49DC6" box="[685,800,1645,1673]" italics="true" pageId="43" pageNumber="42">Toricellia</emphasis>
|
||
DC (1830)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C233F90A314960A557BF951FDF59DE4" authority="Hu 1934" authorityName="Hu" authorityYear="1934" box="[256,545,1681,1708]" class="Magnoliopsida" family="Toricelliaceae" higherTaxonomySource="IPNI" kingdom="Plantae" order="Apiales" pageId="43" pageNumber="42" phylum="Magnoliophyta" rank="family">Toricelliaceae Hu 1934</taxonomicName>
|
||
), also in the
|
||
<taxonomicName id="4C233F90A314960A56C5F951FCC89DE4" authorityName="NAKAI" authorityYear="1930" box="[702,796,1681,1707]" class="Magnoliopsida" kingdom="Plantae" order="Apiales" pageId="43" pageNumber="42" phylum="Tracheophyta" rank="order">Apiales</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFB239E2A314960A540BF974FECC9D80" author="Adams, J. E." box="[112,280,1716,1743]" pageId="43" pageNumber="42" pagination="218 - 244" refId="ref32161" refString="Adams, J. E. 1949. Studies in the comparative anatomy of the Cornaceae. Journal of the Elisha Mitchell Scientific Society 65 (2): 218 - 244. [https: // www. jstor. org / stable / 24333908]." type="journal article" year="1949">Adams (1949</bibRefCitation>
|
||
, p. 239) remarked “[the] overall character of the wood is suggestive of araliaceaeous affinities.”
|
||
<taxonomicName id="4C233F90A314960A540BF939FF369C5A" authorityName="DC" authorityYear="1830" box="[112,226,1785,1813]" class="Magnoliopsida" family="Cornaceae" genus="Toricellia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cornales" pageId="43" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A314960A540BF939FF369C5A" box="[112,226,1785,1813]" italics="true" pageId="43" pageNumber="42">Toricellia</emphasis>
|
||
</taxonomicName>
|
||
differs from the fossil in having helical vessel wall thickenings, taller rays (to
|
||
<quantity id="4CDBE9F6A314960A567AF8DEFDB19C76" box="[513,613,1822,1849]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="43" pageNumber="42" unit="mm" value="1.8">1.8 mm</quantity>
|
||
), slightly more rays per mm, scanty paratracheal parenchyma more obvious, and conspicuous tyloses (
|
||
<bibRefCitation id="EFB239E2A314960A5642F8A4FF659CED" author="Noshiro, S. & P. Baas" pageId="43" pageNumber="42" pagination="43 - 97" refId="ref37877" refString="Noshiro, S. and P. Baas. 1998. Systematic wood anatomy of Cornaceae and allies. IAWA Journal 19 (1): 43 - 97. [https: // doi. org / 10.1163 / 22941932 - 90000652]." type="journal article" year="1998">Noshiro and Baas 1998</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C233F90A314960A54B2F846FEBC9CED" authority="DC" authorityName="DC" authorityYear="1830" box="[201,360,1926,1954]" class="Magnoliopsida" family="Cornaceae" genus="Toricellia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cornales" pageId="43" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A314960A54B2F846FEEF9CED" box="[201,315,1926,1954]" italics="true" pageId="43" pageNumber="42">Toricellia</emphasis>
|
||
DC
|
||
</taxonomicName>
|
||
, is known by fruits from the Clarno Nut Beds (
|
||
<bibRefCitation id="EFB239E2A314960A57BBF98BFB019D2A" author="Manchester, S. R. & M. E. Collinson & C Soriano and & D. Sykes" box="[960,1237,1611,1637]" pageId="43" pageNumber="42" pagination="567 - 579" refId="ref37208" refString="Manchester, S. R., M. E. Collinson, C Soriano and D. Sykes. 2017. Homologous fruit characters in geographically separated genera of extant and fossil Torricelliaceae. International Journal of Plant Sciences 178 (7): 567 - 579. [https: // doi. org / 10.1086 / 692988]." type="journal article" year="2017">
|
||
<collectingRegion id="49E78AF1A314960A57BBF98BFB869D2A" box="[960,1106,1611,1637]" country="United Kingdom" name="Manchester" pageId="43" pageNumber="42">Manchester</collectingRegion>
|
||
et al. 2017
|
||
</bibRefCitation>
|
||
) but is not known from any Post locality.
|
||
</paragraph>
|
||
<caption id="DF5C149BA314960A540BFAB4FC9D9D5E" ID-DOI="http://doi.org/10.5281/zenodo.10913368" ID-Zenodo-Dep="10913368" httpUri="https://zenodo.org/record/10913368/files/figure.png" pageId="43" pageNumber="42" startId="43.[112,187,1396,1420]" targetBox="[147,1488,194,1368]" targetPageId="43" targetType="figure">
|
||
<paragraph id="8B9C4413A314960A540BFAB4FC9D9D5E" blockId="43.[112,1517,1396,1554]" pageId="43" pageNumber="42">
|
||
<emphasis id="B9579801A314960A540BFAB4FF319EC3" bold="true" box="[112,229,1396,1420]" pageId="43" pageNumber="42">Figure 19.</emphasis>
|
||
<taxonomicName id="4C233F90A314960A5491FAB4FE899EC3" box="[234,349,1396,1420]" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="43" pageNumber="42" phylum="Tracheophyta" rank="family">Araliaceae</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C233F90A314960A551DFAB5FDB89EC2" authority="Wheeler & Manchester & Baas, 2023" authorityName="Wheeler & Manchester & Baas" authorityYear="2023" box="[358,620,1397,1421]" class="Magnoliopsida" family="Araliaceae" genus="Plerandreoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="43" pageNumber="42" phylum="Tracheophyta" rank="species" species="oskolski" status="sp. nov.">
|
||
<emphasis id="B9579801A314960A551DFAB5FDB89EC2" box="[358,620,1397,1421]" italics="true" pageId="43" pageNumber="42">Plerandreoxylon oskolski</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A264257AA314960A560AFAB5FD169EC3" box="[625,706,1397,1420]" pageId="43" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, UF 278-84906.
|
||
<emphasis id="B9579801A314960A5709FAB4FC579EC3" bold="true" box="[882,899,1396,1420]" pageId="43" pageNumber="42">A</emphasis>
|
||
. Semi-ring-porous wood with latewood vessels arranged in wavy tangential bands/diagonal arrangement, vessel clusters in latest latewood, axial parenchyma rare, T.S.
|
||
<emphasis id="B9579801A314960A516FFA4FFAF09EE8" bold="true" box="[1300,1316,1423,1447]" pageId="43" pageNumber="42">B</emphasis>
|
||
. Semi-ring-porous wood, vessels solitary and in short radial multiples, axial parenchyma rare, TS.
|
||
<emphasis id="B9579801A314960A57BBFA69FC1B9E8E" bold="true" box="[960,975,1449,1473]" pageId="43" pageNumber="42">C</emphasis>
|
||
. Simple perforation plates, rays with procumbent body cells, RLS.
|
||
<emphasis id="B9579801A314960A5560FA04FEF99E93" bold="true" box="[283,301,1476,1500]" pageId="43" pageNumber="42">D</emphasis>
|
||
. Crowded alternate intervessel pitting, TLS.
|
||
<emphasis id="B9579801A314960A5773FA04FCC39E93" bold="true" box="[776,791,1476,1500]" pageId="43" pageNumber="42">E</emphasis>
|
||
. Vessel-ray parenchyma pits with reduced borders, oval to slightly horizontally elongate in outline, tyloses, RLS.
|
||
<emphasis id="B9579801A314960A562CFA1FFDBC9EB8" bold="true" box="[599,616,1503,1527]" pageId="43" pageNumber="42">F.</emphasis>
|
||
Rays 5-6 cells wide, septate fibers, TLS.
|
||
<emphasis id="B9579801A314960A506CFA1FFBF39EB8" bold="true" box="[1047,1063,1503,1527]" pageId="43" pageNumber="42">G</emphasis>
|
||
. Rays predominantly multiseriate, TLS. Scale bars: 200 µm in A, B, G; 100 µm in C, F. 50 µm in E; 20 µm in D.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9C4413A314960D571AF951FE899A9C" blockId="43.[833,1522,1611,1954]" lastBlockId="44.[112,800,193,469]" lastPageId="44" lastPageNumber="43" pageId="43" pageNumber="42">
|
||
There are araliaceous fruits (
|
||
<taxonomicName id="4C233F90A314960A50A0F950FC8B9D80" authorityName="Wheeler and Manchester" authorityYear="2002" class="Magnoliopsida" family="Araliaceae" genus="Paleopanax" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="43" pageNumber="42" phylum="Tracheophyta" rank="species" species="oregonensis">
|
||
<emphasis id="B9579801A314960A50A0F950FC8B9D80" italics="true" pageId="43" pageNumber="42">Paleopanax oregonensis</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="EFB239E2A314960A5711F974FB9F9D80" author="Manchester, S. R." box="[874,1099,1716,1743]" pageId="43" pageNumber="42" pagination="1 - 205" refId="ref37109" refString="Manchester, S. R. 1994. Fruits and seeds of the middle Eocene Nut Beds Flora, Clarno Formation, Oregon. Palaeontographica Americana 58: 1 - 205." type="journal article" year="1994">
|
||
<collectingRegion id="49E78AF1A314960A5711F974FC289D81" box="[874,1020,1716,1742]" country="United Kingdom" name="Manchester" pageId="43" pageNumber="42">Manchester</collectingRegion>
|
||
, 1994
|
||
</bibRefCitation>
|
||
) and two species of fossil wood,
|
||
<taxonomicName id="4C233F90A314960A573AF918FC4F9C5A" authority="Wheeler and Manchester (2002)" authorityName="Wheeler and Manchester" authorityYear="2002" class="Magnoliopsida" family="Araliaceae" genus="Plerandreoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="43" pageNumber="42" phylum="Tracheophyta" rank="species" species="nutbedensis">
|
||
<emphasis id="B9579801A314960A573AF918FB729DBD" box="[833,1190,1750,1780]" italics="true" pageId="43" pageNumber="42">Plerandreoxylon nutbedensis</emphasis>
|
||
<bibRefCitation id="EFB239E2A314960A50CBF918FC4F9C5A" author="Wheeler, E. A. & S. R. Manchester" pageId="43" pageNumber="42" pagination="1 - 188" refId="ref41110" refString="Wheeler, E. A. and S. R. Manchester. 2002. Woods of the Eocene Nut Beds flora, Clarno Formation, Oregon, USA. IAWA Journal Supplement 3: 1 - 188." type="journal article" year="2002">Wheeler and Manchester (2002)</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C233F90A314960A579BF939FC4D9C76" authority="Wheeler and Manchester (2002)" authorityName="Wheeler and Manchester" authorityYear="2002" class="Magnoliopsida" family="Araliaceae" genus="Paleopanax" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="43" pageNumber="42" phylum="Tracheophyta" rank="species" species="oregonensis">
|
||
<emphasis id="B9579801A314960A579BF939FB4A9C58" box="[992,1182,1785,1815]" italics="true" pageId="43" pageNumber="42">Pl. oregonensis</emphasis>
|
||
<bibRefCitation id="EFB239E2A314960A50D2F93BFC4D9C76" author="Wheeler, E. A. & S. R. Manchester" pageId="43" pageNumber="42" pagination="1 - 188" refId="ref41110" refString="Wheeler, E. A. and S. R. Manchester. 2002. Woods of the Eocene Nut Beds flora, Clarno Formation, Oregon, USA. IAWA Journal Supplement 3: 1 - 188." type="journal article" year="2002">Wheeler and Manchester (2002)</bibRefCitation>
|
||
</taxonomicName>
|
||
, known from the middle Eocene Clarno Formation. This Dietz Hill wood agrees with the diagnosis of
|
||
<taxonomicName id="4C233F90A314960A573AF8A5FBD09CCE" authorityName="WHEELER & MANCHESTER" authorityYear="2002" box="[833,1028,1893,1921]" class="Magnoliopsida" family="Araliaceae" genus="Plerandreoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="43" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A314960A573AF8A5FBD09CCE" box="[833,1028,1893,1921]" italics="true" pageId="43" pageNumber="42">Plerandreoxylon</emphasis>
|
||
</taxonomicName>
|
||
, aside from “rays usually less than 4-seriate” and so we have emended the diagnosis to rays less than 10-seriate.
|
||
<taxonomicName id="4C233F90A313960D553AFF03FD709B92" authorityName="Wheeler and Manchester" authorityYear="2002" box="[321,676,193,223]" class="Magnoliopsida" family="Araliaceae" genus="Plerandreoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="44" pageNumber="43" phylum="Tracheophyta" rank="species" species="nutbedensis">
|
||
<emphasis id="B9579801A313960D553AFF03FD709B92" box="[321,676,193,223]" italics="true" pageId="44" pageNumber="43">Plerandreoxylon nutbedensis</emphasis>
|
||
</taxonomicName>
|
||
differs in having some scalariform perforation plates, some scalariform intervessel pitting, with uniseriate rays more common;
|
||
<taxonomicName id="4C233F90A313960D5496FEEAFE779A07" authorityName="Wheeler and Manchester" authorityYear="2002" box="[237,419,298,328]" class="Magnoliopsida" family="Araliaceae" genus="Paleopanax" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="44" pageNumber="43" phylum="Tracheophyta" rank="species" species="oregonensis">
|
||
<emphasis id="B9579801A313960D5496FEEAFE779A07" box="[237,419,298,328]" italics="true" pageId="44" pageNumber="43">Pl. oregonensis</emphasis>
|
||
</taxonomicName>
|
||
has mostly non-septate fibers and some scalariform intervessel pitting (
|
||
<bibRefCitation id="EFB239E2A313960D5604FE8FFE999AC3" author="Wheeler, E. A. & S. R. Manchester" pageId="44" pageNumber="43" pagination="1 - 188" refId="ref41110" refString="Wheeler, E. A. and S. R. Manchester. 2002. Woods of the Eocene Nut Beds flora, Clarno Formation, Oregon, USA. IAWA Journal Supplement 3: 1 - 188." type="journal article" year="2002">
|
||
Wheeler and
|
||
<collectingRegion id="49E78AF1A313960D540BFEB2FED79AC3" box="[112,259,370,396]" country="United Kingdom" name="Manchester" pageId="44" pageNumber="43">Manchester</collectingRegion>
|
||
2002
|
||
</bibRefCitation>
|
||
). We consider these differences sufficient to warrant assigning this wood to a new species of
|
||
<taxonomicName id="4C233F90A313960D54F4FE79FE839A9A" authorityName="WHEELER & MANCHESTER" authorityYear="2002" box="[143,343,441,469]" class="Magnoliopsida" family="Araliaceae" genus="Plerandreoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="44" pageNumber="43" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A313960D54F4FE79FE839A9A" box="[143,343,441,469]" italics="true" pageId="44" pageNumber="43">Plerandreoxylon</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |