312 lines
36 KiB
XML
312 lines
36 KiB
XML
<document id="370A50B5A396DA143107915F37DB7F65" ID-CLB-Dataset="21808" ID-DOI="10.5852/ejt.2014.100" ID-GBIF-Dataset="5329bfc4-61b2-48f5-a96a-caf4ba31039b" ID-ISSN="2118-9773" ID-Zenodo-Dep="3860450" ID-ZooBank="B3E6C489-6D96-4AF5-A33D-EE8329A9321B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1590595554426" checkinUser="carolina" docAuthor="Enghoff, Henrik" docDate="2014" docId="334F2769C153FFC81C3EF2BBFC03FAFF" docLanguage="en" docName="ejt-2014-100.pdf.imf" docOrigin="European Journal of Taxonomy 100" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chaleponcus howelli Enghoff 2014, sp. nov." docType="treatment" docUuid="F7D97CB8-4791-4584-AD89-F2BCC6589590" docUuidSource="ZooBank" docVersion="7" lastPageNumber="65" masterDocId="CF765F11C16DFF881E57F137FFA0FFA3" masterDocTitle="A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="63" updateTime="1698841637397" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="48070A39CD8C052D9AA64D184CA01826">A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="D8BC3CF82D03EC3231DF2D612215DF01">Enghoff, Henrik</mods:namePart>
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<mods:typeOfResource id="C0DE070F2E6F68B65AC56EF45EB2AD36">text</mods:typeOfResource>
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<mods:title id="EFCDF7F375B29D8A352F0CEE417060E1">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="503B86B6D88CD6BF75A27A92FAA98D88">
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<mods:date id="68D0EB2F30AE0B3722FEDD1607267271">2014</mods:date>
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<mods:number id="66FCDB8CD4D49E8C619889D97F3A6F2C">2014-10-24</mods:number>
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<mods:number id="C47C0F293EFF16D49874357E06A011E9">100</mods:number>
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<mods:classification id="D7EA6D792697E0B4B9FEA9CE801BDF0D">journal article</mods:classification>
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<mods:identifier id="BA9869FF8B5DE1EE64F6A80D63F3C3FE" type="DOI">10.5852/ejt.2014.100</mods:identifier>
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<treatment id="334F2769C153FFC81C3EF2BBFC03FAFF" ID-DOI="http://doi.org/10.5281/zenodo.3861205" ID-GBIF-Taxon="164362779" ID-Zenodo-Dep="3861205" LSID="urn:lsid:zoobank.org:act:F7D97CB8-4791-4584-AD89-F2BCC6589590" httpUri="http://treatment.plazi.org/id/334F2769C153FFC81C3EF2BBFC03FAFF" lastPageId="64" lastPageNumber="65" pageId="62" pageNumber="63">
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<subSubSection id="F3FCC5F4C153FFB61C3EF2BBFC6AFC04" box="[617,970,908,935]" pageId="62" pageNumber="63" type="nomenclature">
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<paragraph id="BB59967FC153FFB61C3EF2BBFC6AFC04" blockId="62.[369,1218,908,1005]" box="[617,970,908,935]" pageId="62" pageNumber="63">
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<heading id="E0112113C153FFB61C3EF2BBFC6AFC04" box="[617,970,908,935]" centered="true" fontSize="11" level="2" pageId="62" pageNumber="63" reason="2">
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<taxonomicName id="7CE6EDFCC153FFB61C3EF2BBFCC7FC04" ID-CoL="TH6N" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[617,871,908,935]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="howelli" status="sp. nov.">
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<emphasis id="89924A6DC153FFB61C3EF2BBFCC7FC04" bold="true" box="[617,871,908,935]" italics="true" pageId="62" pageNumber="63">Chaleponcus howelli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="92A1F716C153FFB61D39F2BAFC6AFC04" box="[878,970,909,935]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F3FCC5F4C153FFB61F26F287FCC2FC4E" pageId="62" pageNumber="63" type="description">
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<paragraph id="BB59967FC153FFB61F26F287FB62FC68" blockId="62.[369,1218,908,1005]" box="[369,1218,944,971]" pageId="62" pageNumber="63">
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<uri id="CF779A7DC153FFB61F26F287FB62FC68" box="[369,1218,944,971]" pageId="62" pageNumber="63">
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urn:lsid:zoobank.org:act:
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<uuid id="CF40ACAAC153FFB61CC0F286FB62FC68" box="[663,1218,944,971]" pageId="62" pageNumber="63">F7D97CB8-4791-4584-AD89-F2BCC6589590</uuid>
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</uri>
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</paragraph>
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<paragraph id="BB59967FC153FFB61C86F2E4FCC2FC4E" blockId="62.[369,1218,908,1005]" box="[721,866,979,1005]" pageId="62" pageNumber="63">
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<figureCitation id="23DD8AFAC153FFB61C86F2E4FCBCFC4E" box="[721,796,979,1005]" captionStart="Fig" captionStartId="11.[189,231,1754,1780]" captionTargetBox="[189,1398,416,1713]" captionTargetId="figure@11.[189,1398,416,1713]" captionTargetPageId="11" captionText="Fig. 4. Limbus types in Chaleponcus species, SEM micrographs, external view, except I: internal view. — A. Lobes pointed, more than twice as long as broad (C. scopus sp. nov.). B. Lobes pointed, slightly less than twice as long as broad (C. howelli sp. nov.). C. Lobes pointed, almost equilateral triangles (C. netus sp. nov.), D. Lobes rounded, slightly longer than broad (C. dabagaensis Kraus, 1958). E. Lobes broadly rounded, slightly shorter than broad (C. hamerae sp. nov.). F. Lobes spatulate (C. gracilior sp. nov.). G. Lobes rounded, broader than long, spinose, ridged (C. krai sp. nov.). H. Lobes ca. 3 times as broad as high, rounded, spinose (C. teres sp. nov.). I. Lobes rounded, much broader than high, smooth (C. malleolus sp. nov.). J. No lobes (C. circumvallatus sp. nov.). Scales: 0.01 mm (A– F, J), 0.001 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.3860460" httpUri="https://zenodo.org/record/3860460/files/figure.png" pageId="62" pageNumber="63">Figs 4</figureCitation>
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,
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<figureCitation id="23DD8AFAC153FFB61D7FF2E4FC99FC4E" box="[808,825,979,1005]" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The ’transverse’ plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="62" pageNumber="63">9</figureCitation>
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,
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<figureCitation id="23DD8AFAC153FFB61D12F2E4FCC2FC4E" box="[837,866,979,1005]" captionStart="Fig" captionStartId="63.[189,232,1860,1886]" captionTargetBox="[206,1385,265,1818]" captionTargetId="figure@63.[206,1385,265,1818]" captionTargetPageId="63" captionText="Fig. 30. Chaleponcus howelli sp. nov., left gonopod. A. Mesal-ventral (-anterior) view. B. Posterior view. C. Anterior view. D. Ventral (-anterior) view. al = anterior telomeral lamella, als = spine on al, cu = cucullus, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior telomeral lamella, ps = proximal spine on solenomere, sts = spine on telomeral main stem. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860514" httpUri="https://zenodo.org/record/3860514/files/figure.png" pageId="62" pageNumber="63">30</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F3FCC5F4C153FFB61EEAF523FCDAFB00" pageId="62" pageNumber="63" type="diagnosis">
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<paragraph id="BB59967FC153FFB61EEAF523FE96FB8D" blockId="62.[189,310,1044,1070]" box="[189,310,1044,1070]" pageId="62" pageNumber="63">
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||
<heading id="E0112113C153FFB61EEAF523FE96FB8D" bold="true" box="[189,310,1044,1070]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
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||
<emphasis id="89924A6DC153FFB61EEAF523FE96FB8D" bold="true" box="[189,310,1044,1070]" pageId="62" pageNumber="63">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="BB59967FC153FFB61EEAF574FCDAFB00" blockId="62.[189,1398,1091,1188]" pageId="62" pageNumber="63">
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||
Medium-sized. Gonopod coxa (
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<figureCitation id="23DD8AFAC153FFB61C6CF574FD24FBFE" box="[571,644,1091,1117]" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The ’transverse’ plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="62" pageNumber="63">Fig. 9</figureCitation>
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) without a lateral process; cucullus small. Distinguished from other species, i.a., by the presence of a spine (
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<emphasis id="89924A6DC153FFB61C83F550FD53FB23" box="[724,755,1127,1152]" italics="true" pageId="62" pageNumber="63">sts</emphasis>
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) on the main stem of the telomere in combination with the large distal spine (
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<emphasis id="89924A6DC153FFB61FE8F5BDFE41FB07" box="[447,481,1162,1188]" italics="true" pageId="62" pageNumber="63">als</emphasis>
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) on the anterior telomeral lamella.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F3FCC5F4C153FFB61EEAF5FCFE9FFA94" pageId="62" pageNumber="63" type="etymology">
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||
<paragraph id="BB59967FC153FFB61EEAF5FCFEE4FB46" blockId="62.[189,324,1227,1253]" box="[189,324,1227,1253]" pageId="62" pageNumber="63">
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||
<heading id="E0112113C153FFB61EEAF5FCFEE4FB46" bold="true" box="[189,324,1227,1253]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
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||
<emphasis id="89924A6DC153FFB61EEAF5FCFEE4FB46" bold="true" box="[189,324,1227,1253]" pageId="62" pageNumber="63">Etymology</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="BB59967FC153FFB61EEAF5CEFE9FFA94" blockId="62.[189,1398,1273,1335]" pageId="62" pageNumber="63">The name honours Kim Howell in recognition of his huge efforts to increase the knowledge of Tanzanian millipedes.</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F3FCC5F4C153FFB61EEAF469FE8DF93B" pageId="62" pageNumber="63" type="materials_examined">
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||
<paragraph id="BB59967FC153FFB61EEAF469FDEBFADA" blockId="62.[189,587,1374,1401]" box="[189,587,1374,1401]" pageId="62" pageNumber="63">
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||
<emphasis id="89924A6DC153FFB61EEAF469FE2CFADB" bold="true" box="[189,396,1374,1400]" pageId="62" pageNumber="63">Material studied</emphasis>
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||
(total:
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||
<specimenCount id="ADE05DF6C153FFB61FB6F469FDE2FADA" box="[481,578,1374,1401]" pageId="62" pageNumber="63" type="male">100 ♂♂</specimenCount>
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||
)
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||
</paragraph>
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<materialsCitation id="0B8E9C22C153FFB61EBDF4BAFC69FA4E" ID-GBIF-Occurrence="2626747317" collectingDate="1984-10-07" collectingDateMax="1984-10-15" collectingDateMin="1984-10-07" collectingMethod="pitfall traps" collectionCode="ZMUC" collectorName="N. Scharff" country="Tanzania" county="Udzungwa Mts" elevation="1900" location="Kigogo Forest Reserve" municipality="Mufindi" pageId="62" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="Iringa Region" typeStatus="holotype">
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<paragraph id="BB59967FC153FFB61EBDF4BAFEFCFA04" blockId="62.[189,1399,1421,1518]" box="[234,348,1421,1447]" pageId="62" pageNumber="63">
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<emphasis id="89924A6DC153FFB61EBDF4BAFEFCFA04" bold="true" box="[234,348,1421,1447]" pageId="62" pageNumber="63">
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<typeStatus id="645D28DDC153FFB61EBDF4BAFEFCFA04" box="[234,348,1421,1447]" pageId="62" pageNumber="63" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
</paragraph>
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||
<paragraph id="BB59967FC153FFB61EEAF487FC69FA4E" blockId="62.[189,1399,1421,1518]" pageId="62" pageNumber="63">
|
||
<collectingCountry id="C3F1D6EFC153FFB61EEAF487FEF7FA69" box="[189,343,1456,1482]" name="Tanzania" pageId="62" pageNumber="63">TANZANIA</collectingCountry>
|
||
:
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||
<specimenCount id="ADE05DF6C153FFB61F34F487FEDBFA69" box="[355,379,1456,1482]" pageId="62" pageNumber="63" type="male">♂</specimenCount>
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||
,
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||
<collectingRegion id="7922589DC153FFB61FDDF487FD91FA69" box="[394,561,1456,1482]" country="Tanzania" name="Iringa" pageId="62" pageNumber="63">Iringa Region</collectingRegion>
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||
,
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||
<collectingCounty id="5238EEF3C153FFB61C69F487FD56FA69" box="[574,758,1456,1482]" pageId="62" pageNumber="63">Udzungwa Mts</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="5B3D0C05C153FFB61D52F487FCC7FA69" box="[773,871,1456,1482]" pageId="62" pageNumber="63">Mufindi</collectingMunicipality>
|
||
,
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||
<location id="BE39C0A4C153FFB61D23F487FB28FA69" LSID="urn:lsid:plazi:treatment:334F2769C153FFC81C3EF2BBFC03FAFF:BE39C0A4C153FFB61D23F487FB28FA69" box="[884,1160,1456,1482]" country="Tanzania" county="Udzungwa Mts" municipality="Mufindi" name="Kigogo Forest Reserve" pageId="62" pageNumber="63" stateProvince="Iringa Region">Kigogo Forest Reserve</location>
|
||
,
|
||
<quantity id="7C1E3B9AC153FFB61AC1F487FB53FA69" box="[1174,1267,1456,1482]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="62" pageNumber="63" unit="m" value="1900.0">
|
||
<elevation id="30CB714CC153FFB61AC1F487FB53FA69" box="[1174,1267,1456,1482]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="62" pageNumber="63" unit="m" value="1900.0">1900 m</elevation>
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||
</quantity>
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||
,
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||
<date id="CF58B0BFC153FFB61B56F487FF5AFA4E" pageId="62" pageNumber="63" value="1984-10-07" valueMax="1984-10-15" valueMin="1984-10-07">
|
||
<collectingDate id="DF1C4957C153FFB61B56F487FF5AFA4E" pageId="62" pageNumber="63" value="1984-10-07" valueMax="1984-10-15" valueMin="1984-10-07">7–15 Oct. 1984</collectingDate>
|
||
</date>
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||
,
|
||
<collectingMethod id="62A7EE68C153FFB61F51F4E4FE2DFA4D" box="[262,397,1491,1518]" pageId="62" pageNumber="63">pitfall traps</collectingMethod>
|
||
in montane rain forest,
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||
<collectorName id="1613F3A9C153FFB61CF2F4E3FCBFFA4E" box="[677,799,1491,1518]" pageId="62" pageNumber="63">N. Scharff</collectorName>
|
||
leg. (
|
||
<collectionCode id="DDF70EBAC153FFB61D36F4E4FC1DFA4E" box="[865,957,1491,1517]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="62" pageNumber="63">ZMUC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="BB59967FC153FFB61EBDF722FEC7F98C" blockId="62.[189,1399,1557,1688]" box="[234,359,1557,1583]" pageId="62" pageNumber="63">
|
||
<emphasis id="89924A6DC153FFB61EBDF722FEC7F98C" bold="true" box="[234,359,1557,1583]" pageId="62" pageNumber="63">
|
||
<typeStatus id="645D28DDC153FFB61EBDF722FEC7F98C" box="[234,359,1557,1583]" pageId="62" pageNumber="63" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="BB59967FC153FFB61EEAF70FFE8DF93B" blockId="62.[189,1399,1557,1688]" pageId="62" pageNumber="63">
|
||
<collectingCountry id="C3F1D6EFC153FFB61EEAF70FFEF7F9F1" box="[189,343,1592,1618]" name="Tanzania" pageId="62" pageNumber="63">TANZANIA</collectingCountry>
|
||
:
|
||
<specimenCount id="ADE05DF6C153FFB61F34F70FFE18F9F1" box="[355,440,1592,1618]" pageId="62" pageNumber="63" type="male">61 ♂♂</specimenCount>
|
||
,
|
||
<collectingRegion id="7922589DC153FFB61F91F70FFDCDF9F1" box="[454,621,1592,1618]" country="Tanzania" name="Iringa" pageId="62" pageNumber="63">Iringa Region</collectingRegion>
|
||
, Udzungwa Mts, Mufindi, Kigogo Forest Reserve,
|
||
<quantity id="7C1E3B9AC153FFB61A86F70FFA8EF9F1" box="[1233,1326,1592,1618]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="62" pageNumber="63" unit="m" value="1700.0">1700 m</quantity>
|
||
,
|
||
<date id="CF58B0BFC153FFB61B6BF70FFE94F9D6" pageId="62" pageNumber="63" value="1984-10-08" valueMax="1984-10-10" valueMin="1984-10-08">8–10 Oct. 1984</date>
|
||
, pitfall traps in montane rain forest, N. Scharff leg. (ZMUC);
|
||
<specimenCount id="ADE05DF6C153FFB61A46F76CFBC5F9D5" box="[1041,1125,1627,1654]" pageId="62" pageNumber="63" type="male">38 ♂♂</specimenCount>
|
||
, same data as
|
||
<typeStatus id="645D28DDC153FFB61B47F76CFAD7F9D6" box="[1296,1399,1627,1653]" pageId="62" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
(ZMUC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3FCC5F4C153FFB61EEAF7F7FEA8F888" pageId="62" pageNumber="63" type="distribution">
|
||
<paragraph id="BB59967FC153FFB61EEAF7F7FEFDF97A" blockId="62.[189,349,1727,1754]" box="[189,349,1727,1754]" pageId="62" pageNumber="63">
|
||
<heading id="E0112113C153FFB61EEAF7F7FEFDF97A" bold="true" box="[189,349,1727,1754]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
|
||
<emphasis id="89924A6DC153FFB61EEAF7F7FEFDF97A" bold="true" box="[189,349,1727,1754]" pageId="62" pageNumber="63">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BB59967FC153FFB61EEAF7D9FEA8F888" blockId="62.[189,1399,1774,1835]" pageId="62" pageNumber="63">
|
||
<collectingCountry id="C3F1D6EFC153FFB61EEAF7D9FEF7F8AB" box="[189,343,1774,1800]" name="Tanzania" pageId="62" pageNumber="63">TANZANIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="7922589DC153FFB61F34F7D9FDABF8AB" box="[355,523,1774,1800]" country="Tanzania" name="Iringa" pageId="62" pageNumber="63">Iringa Region</collectingRegion>
|
||
, Udzungwa Mts, Mufindi, Kigogo Forest Reserve,
|
||
<quantity id="7C1E3B9AC153FFB61A27F7D9FB6CF8AA" box="[1136,1228,1774,1801]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="62" pageNumber="63" unit="m" value="1900.0">1900 m</quantity>
|
||
, montane rain forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3FCC5F4C153FFC81EEAF664FD15FB94" lastPageId="64" lastPageNumber="65" pageId="62" pageNumber="63" type="description">
|
||
<paragraph id="BB59967FC153FFB61EEAF664FE02F8CD" blockId="62.[189,418,1875,1902]" box="[189,418,1875,1902]" pageId="62" pageNumber="63">
|
||
<emphasis id="89924A6DC153FFB61EEAF664FEEEF8CE" bold="true" box="[189,334,1875,1901]" pageId="62" pageNumber="63">Description</emphasis>
|
||
(male)
|
||
</paragraph>
|
||
<paragraph id="BB59967FC153FFB61EEAF6B5FD67F838" blockId="62.[189,711,1921,1948]" box="[189,711,1921,1948]" pageId="62" pageNumber="63">
|
||
DIAMETER. 2.0–
|
||
<quantity id="7C1E3B9AC153FFB61F27F6B6FE6CF83F" box="[368,460,1921,1948]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="62" pageNumber="63" unit="mm" value="2.2">2.2 mm</quantity>
|
||
, 44–47 podous rings.
|
||
</paragraph>
|
||
<caption id="EF99C6F7C152FFB71EEAF673FD34F84F" ID-DOI="http://doi.org/10.5281/zenodo.3860514" ID-Zenodo-Dep="3860514" httpUri="https://zenodo.org/record/3860514/files/figure.png" pageId="63" pageNumber="64" startId="63.[189,232,1860,1886]" targetBox="[206,1385,265,1818]" targetPageId="63">
|
||
<paragraph id="BB59967FC152FFB71EEAF673FD34F84F" blockId="63.[189,1399,1860,2028]" pageId="63" pageNumber="64">
|
||
<emphasis id="89924A6DC152FFB71EEAF673FEBCF8FD" bold="true" box="[189,284,1860,1886]" pageId="63" pageNumber="64">Fig. 30.</emphasis>
|
||
<taxonomicName id="7CE6EDFCC152FFB71F70F672FD81F8FC" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[295,545,1861,1887]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="howelli" status="sp. nov.">
|
||
<emphasis id="89924A6DC152FFB71F70F672FD81F8FC" box="[295,545,1861,1887]" italics="true" pageId="63" pageNumber="64">Chaleponcus howelli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="92A1F716C152FFB71C7BF672FD2CF8FC" box="[556,652,1861,1887]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, left gonopod.
|
||
<emphasis id="89924A6DC152FFB71D11F673FCFBF8FD" bold="true" box="[838,859,1860,1886]" pageId="63" pageNumber="64">A</emphasis>
|
||
. Mesal-ventral (-anterior) view.
|
||
<emphasis id="89924A6DC152FFB71AB0F672FB5BF8FC" bold="true" box="[1255,1275,1861,1887]" pageId="63" pageNumber="64">B</emphasis>
|
||
. Posterior view.
|
||
<emphasis id="89924A6DC152FFB71F52F65FFEBAF821" bold="true" box="[261,282,1896,1922]" pageId="63" pageNumber="64">C</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="89924A6DC152FFB71F8AF65FFE52F821" bold="true" box="[477,498,1896,1922]" pageId="63" pageNumber="64">D</emphasis>
|
||
. Ventral (-anterior) view.
|
||
<emphasis id="89924A6DC152FFB71D7EF65FFCE0F821" box="[809,832,1896,1922]" italics="true" pageId="63" pageNumber="64">al</emphasis>
|
||
= anterior telomeral lamella,
|
||
<emphasis id="89924A6DC152FFB71AF1F65FFB68F821" box="[1190,1224,1896,1922]" italics="true" pageId="63" pageNumber="64">als</emphasis>
|
||
= spine on
|
||
<emphasis id="89924A6DC152FFB71B0FF65FFACFF821" box="[1368,1391,1896,1922]" italics="true" pageId="63" pageNumber="64">al</emphasis>
|
||
,
|
||
<emphasis id="89924A6DC152FFB71EEAF6BBFF79F806" box="[189,217,1932,1957]" italics="true" pageId="63" pageNumber="64">cu</emphasis>
|
||
= cucullus,
|
||
<emphasis id="89924A6DC152FFB71F25F6BCFE2FF806" box="[370,399,1931,1957]" italics="true" pageId="63" pageNumber="64">mf</emphasis>
|
||
= metaplical flange,
|
||
<emphasis id="89924A6DC152FFB71CC4F6BBFD17F806" box="[659,695,1932,1957]" italics="true" pageId="63" pageNumber="64">mp</emphasis>
|
||
= mesal metaplical process,
|
||
<emphasis id="89924A6DC152FFB71A4DF6BBFB9BF806" box="[1050,1083,1932,1957]" italics="true" pageId="63" pageNumber="64">ms</emphasis>
|
||
= metaplical shelf,
|
||
<emphasis id="89924A6DC152FFB71B78F6BBFAFBF806" box="[1327,1371,1932,1957]" italics="true" pageId="63" pageNumber="64">mss</emphasis>
|
||
= metaplical shelf-spine,
|
||
<emphasis id="89924A6DC152FFB71F99F699FE45F86B" box="[462,485,1966,1992]" italics="true" pageId="63" pageNumber="64">pl</emphasis>
|
||
= posterior telomeral lamella,
|
||
<emphasis id="89924A6DC152FFB71D18F698FCC9F86B" box="[847,873,1967,1992]" italics="true" pageId="63" pageNumber="64">ps</emphasis>
|
||
= proximal spine on solenomere,
|
||
<emphasis id="89924A6DC152FFB71AADF698FAB9F86B" box="[1274,1305,1967,1992]" italics="true" pageId="63" pageNumber="64">sts</emphasis>
|
||
= spine on telomeral main stem. Scales 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BB59967FC12DFFC81EEAF03CFD15FEEB" blockId="64.[189,1398,267,329]" pageId="64" pageNumber="65">COLOUR. After 30 years in alcohol almost uniform pale brown, with faint traces of a light dorsal stripe, sometimes with a thin dark line in midline.</paragraph>
|
||
<paragraph id="BB59967FC12DFFC81EEAF047FC59FE0E" blockId="64.[189,1399,368,430]" pageId="64" pageNumber="65">ANAL VALVES. Each with a long, dorsal spine and a well-developed, but small, triangular ventral one; marginal rim raised, setiferous tubercles well developed, on ‘ravelins’.</paragraph>
|
||
<paragraph id="BB59967FC12DFFC81EEAF0E2FEDFFDB1" blockId="64.[189,1398,469,530]" pageId="64" pageNumber="65">
|
||
LIMBUS (
|
||
<figureCitation id="23DD8AFAC12DFFC81F73F0E2FE22FE4C" box="[292,386,469,495]" captionStart="Fig" captionStartId="11.[189,231,1754,1780]" captionTargetBox="[189,1398,416,1713]" captionTargetId="figure@11.[189,1398,416,1713]" captionTargetPageId="11" captionText="Fig. 4. Limbus types in Chaleponcus species, SEM micrographs, external view, except I: internal view. — A. Lobes pointed, more than twice as long as broad (C. scopus sp. nov.). B. Lobes pointed, slightly less than twice as long as broad (C. howelli sp. nov.). C. Lobes pointed, almost equilateral triangles (C. netus sp. nov.), D. Lobes rounded, slightly longer than broad (C. dabagaensis Kraus, 1958). E. Lobes broadly rounded, slightly shorter than broad (C. hamerae sp. nov.). F. Lobes spatulate (C. gracilior sp. nov.). G. Lobes rounded, broader than long, spinose, ridged (C. krai sp. nov.). H. Lobes ca. 3 times as broad as high, rounded, spinose (C. teres sp. nov.). I. Lobes rounded, much broader than high, smooth (C. malleolus sp. nov.). J. No lobes (C. circumvallatus sp. nov.). Scales: 0.01 mm (A– F, J), 0.001 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.3860460" httpUri="https://zenodo.org/record/3860460/files/figure.png" pageId="64" pageNumber="65">Fig. 4B</figureCitation>
|
||
). With long, slender triangular lobes, separated by rounded sinuses; lobes striate on external surface.
|
||
</paragraph>
|
||
<paragraph id="BB59967FC12DFFC81EEAF30DFE4AFDF0" blockId="64.[189,490,569,596]" box="[189,490,569,596]" pageId="64" pageNumber="65">TARSAL SETATION. Normal.</paragraph>
|
||
<paragraph id="BB59967FC12DFFC81EEAF34CFD24FCCB" blockId="64.[189,1399,633,872]" pageId="64" pageNumber="65">
|
||
GONOPOD COXA (
|
||
<figureCitation id="23DD8AFAC12DFFC81FD3F34CFE73FD36" box="[388,467,635,661]" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The ’transverse’ plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="64" pageNumber="65">Figs 9</figureCitation>
|
||
,
|
||
<figureCitation id="23DD8AFAC12DFFC81FB2F34CFDA6FD36" box="[485,518,635,661]" captionStart="Fig" captionStartId="63.[189,232,1860,1886]" captionTargetBox="[206,1385,265,1818]" captionTargetId="figure@63.[206,1385,265,1818]" captionTargetPageId="63" captionText="Fig. 30. Chaleponcus howelli sp. nov., left gonopod. A. Mesal-ventral (-anterior) view. B. Posterior view. C. Anterior view. D. Ventral (-anterior) view. al = anterior telomeral lamella, als = spine on al, cu = cucullus, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior telomeral lamella, ps = proximal spine on solenomere, sts = spine on telomeral main stem. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860514" httpUri="https://zenodo.org/record/3860514/files/figure.png" pageId="64" pageNumber="65">30</figureCitation>
|
||
). About 3½ × as long as wide. Lateral margin almost straight, towards apex gently curving mesad and continuous with apical margin; cucullus (
|
||
<emphasis id="89924A6DC12DFFC81A75F3A8FB9EFD1B" box="[1058,1086,671,696]" italics="true" pageId="64" pageNumber="65">cu</emphasis>
|
||
) an almost perfect hollow hemisphere. Metaplical flange (
|
||
<emphasis id="89924A6DC12DFFC81C6AF3F6FDFAFD78" box="[573,602,705,731]" italics="true" pageId="64" pageNumber="65">mf</emphasis>
|
||
) ending in blunt, equilateral triangular process; metaplical mesal margin with large rounded process (
|
||
<emphasis id="89924A6DC12DFFC81C3FF3D2FD2CFD5D" box="[616,652,741,766]" italics="true" pageId="64" pageNumber="65">mp</emphasis>
|
||
) delimiting semicircular sinus together with base of metaplical shelf. Metaplical shelf (
|
||
<emphasis id="89924A6DC12DFFC81F85F23FFE53FC82" box="[466,499,776,801]" italics="true" pageId="64" pageNumber="65">ms</emphasis>
|
||
) simple, regularly rounded, mostly projecting laterad. Metaplical shelf-spine (
|
||
<emphasis id="89924A6DC12DFFC81E90F21BFF53FCE6" box="[199,243,812,837]" italics="true" pageId="64" pageNumber="65">mss</emphasis>
|
||
) of medium length, originating from anterior-mesal part of
|
||
<emphasis id="89924A6DC12DFFC81DFCF21BFC6CFCE6" box="[939,972,812,837]" italics="true" pageId="64" pageNumber="65">ms</emphasis>
|
||
, projecting disto-mesad, then basad, reaching beyond telopodital basomere.
|
||
</paragraph>
|
||
<paragraph id="BB59967FC12DFFC81EEAF2B8FE8EFC53" blockId="64.[189,1398,911,1079]" pageId="64" pageNumber="65">
|
||
GONOPOD TELOPODITE (
|
||
<figureCitation id="23DD8AFAC12DFFC81FE9F2B8FDB5FC0A" box="[446,533,911,938]" captionStart="Fig" captionStartId="63.[189,232,1860,1886]" captionTargetBox="[206,1385,265,1818]" captionTargetId="figure@63.[206,1385,265,1818]" captionTargetPageId="63" captionText="Fig. 30. Chaleponcus howelli sp. nov., left gonopod. A. Mesal-ventral (-anterior) view. B. Posterior view. C. Anterior view. D. Ventral (-anterior) view. al = anterior telomeral lamella, als = spine on al, cu = cucullus, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior telomeral lamella, ps = proximal spine on solenomere, sts = spine on telomeral main stem. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860514" httpUri="https://zenodo.org/record/3860514/files/figure.png" pageId="64" pageNumber="65">Fig. 30</figureCitation>
|
||
). Solenomere with a very long, straight, meso-basad, proximal spine (
|
||
<emphasis id="89924A6DC12DFFC81B1CF2A7FAC5FC0A" box="[1355,1381,912,937]" italics="true" pageId="64" pageNumber="65">ps</emphasis>
|
||
). Main stem of telomere with a small spine (
|
||
<emphasis id="89924A6DC12DFFC81CFAF284FD6CFC6F" box="[685,716,947,972]" italics="true" pageId="64" pageNumber="65">sts</emphasis>
|
||
)
|
||
<emphasis id="89924A6DC12DFFC81C8BF284FD5FFC6F" box="[732,767,947,972]" italics="true" pageId="64" pageNumber="65">ca.</emphasis>
|
||
in middle of basal surface. Telomere distally with two branches:
|
||
</paragraph>
|
||
<paragraph id="BB59967FC12DFFC81EEAF2CEFB0DFBB0" blockId="64.[189,1398,911,1079]" box="[189,1197,1017,1043]" pageId="64" pageNumber="65">
|
||
• a rounded anterior lamella (
|
||
<emphasis id="89924A6DC12DFFC81C64F2CEFDEAFBB0" box="[563,586,1017,1043]" italics="true" pageId="64" pageNumber="65">al</emphasis>
|
||
) ending in a long, gently sinuous mesad spine (
|
||
<emphasis id="89924A6DC12DFFC81A2DF2CEFB3CFBB0" box="[1146,1180,1017,1043]" italics="true" pageId="64" pageNumber="65">als</emphasis>
|
||
),
|
||
</paragraph>
|
||
<paragraph id="BB59967FC12DFFC81EEAF52AFD15FB94" blockId="64.[189,1398,911,1079]" box="[189,693,1052,1079]" pageId="64" pageNumber="65">
|
||
• a large, irregular posterior lamella (
|
||
<emphasis id="89924A6DC12DFFC81CDAF52BFD04FB95" box="[653,676,1052,1078]" italics="true" pageId="64" pageNumber="65">pl</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3FCC5F4C12DFFC81EEAF56AFEA8FB6A" pageId="64" pageNumber="65" type="multiple">
|
||
<paragraph id="BB59967FC12DFFC81EEAF56AFE4EFBD4" blockId="64.[189,494,1117,1144]" box="[189,494,1117,1144]" pageId="64" pageNumber="65">
|
||
<heading id="E0112113C12DFFC81EEAF56AFE4EFBD4" bold="true" box="[189,494,1117,1144]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
|
||
<emphasis id="89924A6DC12DFFC81EEAF56AFE4EFBD4" bold="true" box="[189,494,1117,1144]" pageId="64" pageNumber="65">Distribution and habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BB59967FC12DFFC81EEAF5BAFEA8FB6A" blockId="64.[189,1399,1164,1225]" pageId="64" pageNumber="65">
|
||
Known only from Udzungwa Mts, Kigogo FR. Altitudinal range:
|
||
<quantity id="7C1E3B9AC12DFFC81D93F5BBFBCAFB04" box="[964,1130,1164,1191]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="64" pageNumber="65" unit="m" value="1800.0" valueMax="1900.0" valueMin="1700.0">1700–1900 m</quantity>
|
||
. Habitat: montane rain forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3FCC5F4C12DFFC81EEAF5C6FC03FAFF" pageId="64" pageNumber="65" type="discussion">
|
||
<paragraph id="BB59967FC12DFFC81EEAF5C6FE3EFAA8" blockId="64.[189,414,1265,1291]" box="[189,414,1265,1291]" pageId="64" pageNumber="65">
|
||
<heading id="E0112113C12DFFC81EEAF5C6FE3EFAA8" bold="true" box="[189,414,1265,1291]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
|
||
<emphasis id="89924A6DC12DFFC81EEAF5C6FE3EFAA8" bold="true" box="[189,414,1265,1291]" pageId="64" pageNumber="65">Coexisting species</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BB59967FC12DFFC81EEAF428FC03FAFF" blockId="64.[189,1398,1311,1373]" pageId="64" pageNumber="65">
|
||
All four species known from Kigogo FR, i.e.,
|
||
<taxonomicName id="7CE6EDFCC12DFFC81C8BF417FC20FA9A" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[732,896,1311,1338]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="quasimodo" status="sp. nov.">
|
||
<emphasis id="89924A6DC12DFFC81C8BF417FC20FA9A" box="[732,896,1311,1338]" italics="true" pageId="64" pageNumber="65">C. quasimodo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="92A1F716C12DFFC81DD0F417FC42FA99" box="[903,994,1312,1338]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="7CE6EDFCC12DFFC81DA6F417FBC3FA9A" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1009,1123,1312,1338]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="scopus" status="sp. nov.">
|
||
<emphasis id="89924A6DC12DFFC81DA6F417FBC3FA9A" box="[1009,1123,1312,1338]" italics="true" pageId="64" pageNumber="65">C. scopus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="92A1F716C12DFFC81A3CF417FB66FA99" box="[1131,1222,1312,1338]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="7CE6EDFCC12DFFC81A83F417FAEDFA9A" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1236,1357,1311,1338]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="howelli" status="sp. nov.">
|
||
<emphasis id="89924A6DC12DFFC81A83F417FAEDFA9A" box="[1236,1357,1311,1338]" italics="true" pageId="64" pageNumber="65">C. howelli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="92A1F716C12DFFC81B02F417FF4FFAFE" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="7CE6EDFCC12DFFC81F70F474FE14FAFF" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[295,436,1346,1373]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="mwabvui" status="sp. nov.">
|
||
<emphasis id="89924A6DC12DFFC81F70F474FE14FAFF" box="[295,436,1346,1373]" italics="true" pageId="64" pageNumber="65">C. mwabvui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="92A1F716C12DFFC81FECF474FDB8FAFE" box="[443,536,1347,1373]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, were found in the same samples.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |