A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae) Author Enghoff, Henrik text European Journal of Taxonomy 2014 2014-10-24 100 1 75 journal article 21808 10.5852/ejt.2014.100 5329bfc4-61b2-48f5-a96a-caf4ba31039b 2118-9773 3860450 B3E6C489-6D96-4AF5-A33D-EE8329A9321B Chaleponcus howelli sp. nov. urn:lsid:zoobank.org:act: F7D97CB8-4791-4584-AD89-F2BCC6589590 Figs 4 , 9 , 30 Diagnosis Medium-sized. Gonopod coxa ( Fig. 9 ) without a lateral process; cucullus small. Distinguished from other species, i.a., by the presence of a spine ( sts ) on the main stem of the telomere in combination with the large distal spine ( als ) on the anterior telomeral lamella. Etymology The name honours Kim Howell in recognition of his huge efforts to increase the knowledge of Tanzanian millipedes. Material studied (total: 100 ♂♂ ) Holotype TANZANIA : ♂ , Iringa Region , Udzungwa Mts , Mufindi , Kigogo Forest Reserve , 1900 m , 7–15 Oct. 1984 , pitfall traps in montane rain forest, N. Scharff leg. ( ZMUC ). Paratypes TANZANIA : 61 ♂♂ , Iringa Region , Udzungwa Mts, Mufindi, Kigogo Forest Reserve, 1700 m , 8–10 Oct. 1984 , pitfall traps in montane rain forest, N. Scharff leg. (ZMUC); 38 ♂♂ , same data as holotype (ZMUC). Type locality TANZANIA : Iringa Region , Udzungwa Mts, Mufindi, Kigogo Forest Reserve, 1900 m , montane rain forest. Description (male) DIAMETER. 2.0– 2.2 mm , 44–47 podous rings. Fig. 30. Chaleponcus howelli sp. nov. , left gonopod. A . Mesal-ventral (-anterior) view. B . Posterior view. C . Anterior view. D . Ventral (-anterior) view. al = anterior telomeral lamella, als = spine on al , cu = cucullus, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior telomeral lamella, ps = proximal spine on solenomere, sts = spine on telomeral main stem. Scales 0.1 mm. COLOUR. After 30 years in alcohol almost uniform pale brown, with faint traces of a light dorsal stripe, sometimes with a thin dark line in midline. ANAL VALVES. Each with a long, dorsal spine and a well-developed, but small, triangular ventral one; marginal rim raised, setiferous tubercles well developed, on ‘ravelins’. LIMBUS ( Fig. 4B ). With long, slender triangular lobes, separated by rounded sinuses; lobes striate on external surface. TARSAL SETATION. Normal. GONOPOD COXA ( Figs 9 , 30 ). About 3½ × as long as wide. Lateral margin almost straight, towards apex gently curving mesad and continuous with apical margin; cucullus ( cu ) an almost perfect hollow hemisphere. Metaplical flange ( mf ) ending in blunt, equilateral triangular process; metaplical mesal margin with large rounded process ( mp ) delimiting semicircular sinus together with base of metaplical shelf. Metaplical shelf ( ms ) simple, regularly rounded, mostly projecting laterad. Metaplical shelf-spine ( mss ) of medium length, originating from anterior-mesal part of ms , projecting disto-mesad, then basad, reaching beyond telopodital basomere. GONOPOD TELOPODITE ( Fig. 30 ). Solenomere with a very long, straight, meso-basad, proximal spine ( ps ). Main stem of telomere with a small spine ( sts ) ca. in middle of basal surface. Telomere distally with two branches: • a rounded anterior lamella ( al ) ending in a long, gently sinuous mesad spine ( als ), • a large, irregular posterior lamella ( pl ). Distribution and habitat Known only from Udzungwa Mts, Kigogo FR. Altitudinal range: 1700–1900 m . Habitat: montane rain forest. Coexisting species All four species known from Kigogo FR, i.e., C. quasimodo sp. nov. , C. scopus sp. nov. , C. howelli sp. nov. and C. mwabvui sp. nov. , were found in the same samples.