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<document id="1249D605150EDB3705494F4F82AAA5C4" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia &amp; Ahrens, Dirk" docDate="2017" docId="03CB6967FF9738403876FBBD934AFCB3" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla charlyi AHRENS, BECKETT, EBERLE &amp; FABRIZI 2017, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="341" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="340" updateTime="1738712560284" updateUser="ExternalLinkService">
<mods:mods id="86B547D3C36418638786312A707CC3C6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="BEBFF4A26CB4D891C82D89AF3CA7410F">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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<mods:namePart id="21BBC279C954C139E03E6C2C9ACCF3E8">Eberle, Jonas</mods:namePart>
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<mods:namePart id="50ABA3CC01E5E292B62411B29D87099F">Beckett, Marc</mods:namePart>
</mods:name>
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<mods:namePart id="242B9044FFE336D7E6F9522F975E6197">Ozgul-Siemund, Alev</mods:namePart>
</mods:name>
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<mods:namePart id="968DAFFD09BC8C791C1CC883050DB3B0">Frings, Jujina</mods:namePart>
</mods:name>
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<mods:namePart id="D600D368DA0785D5D5F7A6829E59991E">Fabrizi, Silvia</mods:namePart>
</mods:name>
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<mods:namePart id="B3BC842ADBC159E563CCB1F1DBEF663B">Ahrens, Dirk</mods:namePart>
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<mods:typeOfResource id="FDC2E1319655B9BF1C56B4EA94EC0978">text</mods:typeOfResource>
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<mods:date id="1996FF8ECF287A1843D6FBDB46B2A39E">2017</mods:date>
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<mods:title id="E411E6AAFDD1203E4C60590D414B828B">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="16A424517902C6503EEDEFF069847021">180</mods:number>
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<treatment id="03CB6967FF9738403876FBBD934AFCB3" LSID="urn:lsid:plazi:treatment:03CB6967FF9738403876FBBD934AFCB3" httpUri="http://treatment.plazi.org/id/03CB6967FF9738403876FBBD934AFCB3" lastPageId="20" lastPageNumber="341" pageId="19" pageNumber="340">
<subSubSection id="C3788BFAFF9738473876FBBD928EFB3A" pageId="19" pageNumber="340" type="nomenclature">
<paragraph id="8BDDD871FF9738473876FBBD9422FB88" blockId="19.[842,1431,1030,1087]" pageId="19" pageNumber="340">
<heading id="D0956F1DFF9738473876FBBD9422FB88" centered="true" fontSize="9" level="2" pageId="19" pageNumber="340" reason="2">
<taxonomicName id="4C62A3F2FF9738473876FBBD9573FB8B" ID-CoL="782N7" authority="AHRENS, BECKETT, EBERLE &amp; FABRIZI" authorityName="AHRENS, BECKETT, EBERLE &amp; FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
<emphasis id="B9160463FF9738473876FBBD9537FBA8" bold="true" box="[842,1105,1034,1055]" italics="true" pageId="19" pageNumber="340">
<smallCapsWord id="8D3B4EADFF9738473876FBBD9285FBA8" baselines="1050,1050" box="[842,995,1034,1055]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="19" pageNumber="340">PLEOPHYLLA</smallCapsWord>
<smallCapsWord id="8D3B4EADFF97384738D1FBBA9537FBA8" baselines="1050" box="[1005,1105,1037,1054]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="charlyi" pageId="19" pageNumber="340">CHARLYI</smallCapsWord>
</emphasis>
<smallCapsWord id="8D3B4EADFF9738473F66FBB095D3FBA8" baselines="1049,1050" box="[1114,1205,1031,1055]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="19" pageNumber="340">AHRENS</smallCapsWord>
,
<smallCapsWord id="8D3B4EADFF9738473FFBFBB0944BFBA8" baselines="1049,1050" box="[1223,1325,1031,1055]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="19" pageNumber="340">BECKETT</smallCapsWord>
,
<smallCapsWord id="8D3B4EADFF9738473E03FBB094F1FBAB" baselines="1049,1049" box="[1343,1431,1031,1055]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="19" pageNumber="340">EBERLE</smallCapsWord>
&amp;
<smallCapsWord id="8D3B4EADFF9738473881FB919573FB8B" baselines="1080,1081" box="[957,1045,1062,1086]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabrizi" pageId="19" pageNumber="340">FABRIZI</smallCapsWord>
</taxonomicName>
<emphasis id="B9160463FF9738473F22FB9C95E5FB89" bold="true" box="[1054,1155,1060,1087]" pageId="19" pageNumber="340">
<taxonomicNameLabel id="A225B918FF9738473F22FB9C95E5FB89" box="[1054,1155,1060,1087]" pageId="19" pageNumber="340" rank="species">
<smallCapsWord id="8D3B4EADFF9738473F22FB9C955CFB8A" baselines="1080" box="[1054,1082,1067,1084]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="19" pageNumber="340">SP</smallCapsWord>
.
<smallCapsWord id="8D3B4EADFF9738473F70FB9C951AFB8A" baselines="1080" box="[1100,1148,1067,1084]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="19" pageNumber="340">NOV</smallCapsWord>
.
</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1359C4F4FF9738473FAAFB91947CFB88" box="[1174,1306,1060,1086]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="19" pageNumber="340">
<smallCapsWord id="8D3B4EADFF9738473FAAFB9195ADFB8B" baselines="1080,1081" box="[1174,1227,1062,1086]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="19" pageNumber="340">FIGS</smallCapsWord>
4E
<emphasis id="B9160463FF9738473FCBFB9F9463FB88" box="[1271,1285,1064,1086]" italics="true" pageId="19" pageNumber="340">–</emphasis>
H
</figureCitation>
,
<figureCitation id="1359C4F4FF9738473E10FB93945AFB88" box="[1324,1340,1060,1086]" captionStart="Figure 7" captionStartId="14.[144,223,925,944]" captionTargetBox="[144,1428,192,893]" captionTargetId="figure-25@14.[144,1428,192,893]" captionTargetPageId="14" captionText="Figure 7. Distribution of Pleophylla species (left side) and absence records (right side), represented by sampling locali- ties of all other sampled Sericini species without records of Pleophylla (white dots)." figureDoi="http://doi.org/10.5281/zenodo.14807097" httpUri="https://zenodo.org/record/14807097/files/figure.png" pageId="19" pageNumber="340">7</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BDDD871FF9738473807FBE0928EFB3A" blockId="19.[827,1442,1111,1164]" pageId="19" pageNumber="340">
<uri id="FFF3D473FF9738473807FBE0928EFB3A" pageId="19" pageNumber="340">
urn:lsid:zoobank.org:act:
<uuid id="FFC4E2A4FF9738473F7FFBE0928EFB3A" pageId="19" pageNumber="340">00BFEBA3-5CCA-4189-B934- 3B4587703F0A</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3788BFAFF9738403801FB1F934AFCB3" lastPageId="20" lastPageNumber="341" pageId="19" pageNumber="340" type="materials_examined">
<paragraph id="8BDDD871FF9738473801FB1F9542FB4C" blockId="19.[829,1444,1191,1275]" pageId="19" pageNumber="340">
<emphasis id="B9160463FF9738473801FB1F9510FB0A" box="[829,1142,1191,1213]" italics="true" pageId="19" pageNumber="340">Type material examined:</emphasis>
<materialsCitation id="3B0AD22CFF9738473FB9FB109542FB4C" collectingDate="1998-12-14" collectionCode="ZFMK" collectorName="Lizler" country="Tanzania" location="Southern Highland Werner" pageId="19" pageNumber="340" specimenCount="1" specimenCount-male="1" stateProvince="Njombe" typeStatus="holotype">
<typeStatus id="54D966D3FF9738473FB9FB109588FB0B" box="[1157,1262,1191,1213]" pageId="19" pageNumber="340" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D6413F8FF9738473E34FB1D947EFB0A" box="[1288,1304,1194,1212]" count="1" pageId="19" pageNumber="340" type="male">♂</specimenCount>
“
<collectingCountry id="F37598E1FF9738473E05FB1094C5FB0B" box="[1337,1443,1191,1213]" name="Tanzania" pageId="19" pageNumber="340">Tanzania</collectingCountry>
<date id="FFDCFEB1FF9738473801FB7192A4FB6D" box="[829,962,1222,1243]" pageId="19" pageNumber="340" value="1998-12-14">
<collectingDate id="EF980759FF9738473801FB7192A4FB6D" box="[829,962,1222,1243]" pageId="19" pageNumber="340" value="1998-12-14">14.XII.1998</collectingDate>
</date>
<collectingRegion id="49A61693FF97384738EDFB71954EFB6A" box="[977,1064,1222,1244]" country="Tanzania" name="Njombe" pageId="19" pageNumber="340">Njombe</collectingRegion>
<location id="8EBD8EAAFF9738473F0BFB7194E7FB6D" LSID="urn:lsid:plazi:treatment:03CB6967FF9738403876FBBD934AFCB3:8EBD8EAAFF9738473F0BFB7194E7FB6D" box="[1079,1409,1222,1244]" country="Tanzania" name="Southern Highland Werner" pageId="19" pageNumber="340" stateProvince="Njombe">Southern Highland Werner</location>
&amp;
<collectorName id="2697BDA7FF9738473801FB5292E6FB4D" box="[829,896,1253,1275]" pageId="19" pageNumber="340">Lizler</collectorName>
leg.” (
<collectionCode id="ED7340B4FF97384738F5FB52957EFB4C" box="[969,1048,1253,1274]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="19" pageNumber="340" type="Museum">ZFMK</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BDDD871FF9738403801FAA493A1FE81" blockId="19.[829,1444,1298,1901]" lastBlockId="20.[144,758,199,312]" lastPageId="20" lastPageNumber="341" pageId="19" pageNumber="340">
<emphasis id="B9160463FF9738473801FAA492A0FA9E" box="[829,966,1299,1320]" italics="true" pageId="19" pageNumber="340">Description:</emphasis>
Body length:
<quantity id="4C9A7594FF9738473F42FAA49582FA9E" box="[1150,1252,1299,1320]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="19" pageNumber="340" unit="mm" value="12.1">12.1 mm</quantity>
, elytral length:
<quantity id="4C9A7594FF9738473801FA8692F0FAF0" box="[829,918,1329,1350]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="19" pageNumber="340" unit="mm" value="8.4">8.4 mm</quantity>
, maximum width: 6.0 mm. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus strongly reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.7. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/ metacoxa: 1/1.39. Metatibia, ratio of width/length: 1/ 4.28. Metatarsi dorsally smooth; first metatarsomere slightly longer than subsequent one. Female unknown. Aedeagus:
<figureCitation id="1359C4F4FF9038403ABEFE959090FE8E" box="[386,502,290,312]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">Fig. 4E, F</figureCitation>
. Habitus:
<figureCitation id="1359C4F4FF9038403955FE9593A5FE81" box="[617,707,290,312]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">Fig. 4H</figureCitation>
.
</paragraph>
<paragraph id="8BDDD871FF9038403BACFEFA91B6FD22" blockId="20.[144,759,332,660]" pageId="20" pageNumber="341">
<emphasis id="B9160463FF9038403BACFEFA9090FED7" box="[144,502,332,354]" italics="true" pageId="20" pageNumber="341">
Diagnosis:
<taxonomicName id="4C62A3F2FF9038403A29FEFB9090FED7" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[277,502,332,353]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">Pleophylla charlyi</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A225B918FF9038403933FEFA931EFED4" box="[527,632,333,354]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
is quite similar to
<taxonomicName id="4C62A3F2FF9038403A30FEDB90CBFE36" box="[268,429,363,385]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="navicularis">
<emphasis id="B9160463FF9038403A30FEDB90CBFE36" box="[268,429,363,385]" italics="true" pageId="20" pageNumber="341">P. navicularis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C62A3F2FF9038403AD2FEDB9308FE36" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[494,622,363,385]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="silvatica" status="sp. nov.">
<emphasis id="B9160463FF9038403AD2FEDB9308FE36" box="[494,622,363,385]" italics="true" pageId="20" pageNumber="341">P. silvatica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF9038403945FEDB93B3FE37" box="[633,725,364,385]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
in external morphology.
<taxonomicName id="4C62A3F2FF9038403A9EFE3D9318FE29" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[418,638,394,415]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
<emphasis id="B9160463FF9038403A9EFE3D9318FE29" box="[418,638,394,415]" italics="true" pageId="20" pageNumber="341">Pleophylla charlyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF90384039AEFE3D9391FE29" box="[658,759,394,415]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
differs from both in the strongly asymmetrical ventral lobes of the parameres being distinctly produced dorsally. From
<taxonomicName id="4C62A3F2FF9038403AFBFE519319FE4C" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[455,639,485,507]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="mpumalanga" status="sp. nov.">
<emphasis id="B9160463FF9038403AFBFE519319FE4C" box="[455,639,485,507]" italics="true" pageId="20" pageNumber="341">P. mpumalanga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF90384039B3FE519394FE4D" box="[655,754,486,507]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
, which is similar in the shape of the parameres,
<taxonomicName id="4C62A3F2FF9038403BACFD949066FD81" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[144,256,546,568]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
<emphasis id="B9160463FF9038403BACFD949066FD81" box="[144,256,546,568]" italics="true" pageId="20" pageNumber="341">P. charlyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF9038403A31FD94900CFD8E" box="[269,362,547,568]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
differs in the evenly distributed white pilosity on the elytra and the large lobes produced dorsally by the extensions of the ventral lobes.
</paragraph>
<paragraph id="8BDDD871FF9038403BACFD05934AFCB3" blockId="20.[144,758,690,773]" pageId="20" pageNumber="341">
<emphasis id="B9160463FF9038403BACFD05906BFD71" box="[144,269,690,711]" italics="true" pageId="20" pageNumber="341">Etymology:</emphasis>
The new species is named after one of its collectors, Karl Werner (his nickname was ‘Charly’). He passed away too early.
</paragraph>
</subSubSection>
</treatment>
</document>
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