Add updates up until 2025-02-04 23:45:25
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03C4526D0325A056029626BCFD0CFDC4.xml03C4526D0331A042029626BCFA57F836.xml03C4526D0339A04A029626BCFA57F834.xml03C4526D033FA04C029626BCFA57F834.xml
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03CB6967FF82385C387FFB619019FB26.xml03CB6967FF8838583870FCE594EBF8DB.xml03CB6967FF8838583B94FCAC920BFC9F.xml03CB6967FF8A385B3BA9F9C993BAFD73.xml03CB6967FF8B38443876FDA091B0FDDC.xml03CB6967FF8B385B3BEEFD4A92F7FE52.xml03CB6967FF8C385C3BD6FB7095DBFC41.xml03CB6967FF8C385E3803FB9990E4FBCF.xml03CB6967FF8E38583805FB2E906BFD5A.xml03CB6967FF8E385E3BACFB289438FBD5.xml03CB6967FF9038403BADFCFC9449FD2C.xml03CB6967FF903841386FFD699300FEDE.xml03CB6967FF913841387DFEE7944AF95F.xml03CB6967FF9138413B91FE2C9538FEAC.xml03CB6967FF913842385BF8A69194FB1D.xml03CB6967FF9238433BAFFB69908CFCE8.xml03CB6967FF9438443B85FD16953DFE13.xml03CB6967FF9438453816FE6193D8FD80.xml03CB6967FF9538453B87FD3794F4FD22.xml03CB6967FF9538463871FD6591B0FC26.xml03CB6967FF9638463B84FC729570FBBF.xml03CB6967FF963847386AFBF9939FFB20.xml03CB6967FF9738403876FBBD934AFCB3.xml03CB6967FF9738473BFAFB6792B3FC79.xml
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<mods:title id="D2C1F5C6F8A83CA3CE06C8C2DB25C0FA">Small survivors: unexpected endemic diversity of Hyposmocoma (Lepidoptera: Cosmopterigidae) moths on Kaho olawe, a degraded Hawaiian island</mods:title>
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<mods:namePart id="AA87ED804294B315478D64395F4613B2">Doorenweerd, Camiel</mods:namePart>
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<mods:namePart id="F9D402EBF0BA64DF33EC5295F30DAB0F">Austin, Kyhl A.</mods:namePart>
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<mods:namePart id="78268FBD744F409B55AF9C53007D260A">Rubinoff, Daniel</mods:namePart>
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<mods:title id="5BD06274A9FF67B68CF5047B8C4A29D2">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="8253AD0B13370160F10E6CF165D3D397">
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<mods:date id="675F61B494ADB24563B843147A62AF4F">2023</mods:date>
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<mods:title id="5D5F5A15D244ABA52275A5B76F85F07B">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="3E8646B0F10FF2CB9F38739A0AB46AE7">180</mods:number>
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<mods:number id="B50D7B5158F051D5013FB7CBEB43957F">3</mods:number>
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<mods:identifier id="D332DA8B44565C5092D1065BAA1B1CF2" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03C4526D0325A056029626BCFD0CFDC4" LSID="urn:lsid:plazi:treatment:03C4526D0325A056029626BCFD0CFDC4" httpUri="http://treatment.plazi.org/id/03C4526D0325A056029626BCFD0CFDC4" lastPageNumber="587" pageId="17" pageNumber="587">
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NEW
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<emphasis id="B9193F690325A056035326BCFB95FF61" box="[815,1025,189,210]" italics="true" pageId="17" pageNumber="587">HYPOSMOCOMA</emphasis>
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FROM
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</taxonomicName>
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KAHOʻOLAWE
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587
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<emphasis id="B9193F690325A05603C727D3FBFBFE58" bold="true" box="[955,1135,466,491]" pageId="17" pageNumber="587">APPENDIX 2</emphasis>
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</heading>
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</paragraph>
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.
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Sort Rearing order
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<taxonomicName id="4C6D98F80325A056014B2463FE32FDC4" box="[311,422,610,631]" order="Extraction" pageId="17" pageNumber="587" rank="order">Extraction</taxonomicName>
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code Species
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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<mods:title id="C03B1EECB6DBC5B0434D4736BAF9364C">Small survivors: unexpected endemic diversity of Hyposmocoma (Lepidoptera: Cosmopterigidae) moths on Kaho olawe, a degraded Hawaiian island</mods:title>
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<paragraph id="8BD2E37B0331A042029626BCFA5DFF60" blockId="5.[746,1481,184,211]" box="[746,1481,184,211]" pageId="5" pageNumber="575">
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<heading id="D09A54170331A042029626BCFA5DFF60" box="[746,1481,184,211]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="575" reason="2">
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NEW
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<taxonomicName id="4C6D98F80331A042035326BCFBC1FF62" authority="FROM" authorityName="FROM" box="[815,1109,188,210]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="genus">
|
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<emphasis id="B9193F690331A042035326BCFB95FF61" box="[815,1025,189,210]" italics="true" pageId="5" pageNumber="575">HYPOSMOCOMA</emphasis>
|
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FROM
|
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</taxonomicName>
|
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KAHOʻOLAWE
|
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<collectingCountry id="F37AA3EB0331A042056A26BDFAEDFF62" box="[1302,1401,188,209]" name="Iceland" pageId="5" pageNumber="575">ISLAND</collectingCountry>
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575
|
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<emphasis id="B9193F690331A04200BA237CFEA4FA23" bold="true" box="[198,304,1405,1424]" pageId="5" pageNumber="575">Figure 3.</emphasis>
|
||||
Adult
|
||||
<taxonomicName id="4C6D98F80331A0420103237FFD81FA23" box="[383,533,1406,1424]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="genus">
|
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<emphasis id="B9193F690331A0420103237FFD81FA23" box="[383,533,1406,1424]" italics="true" pageId="5" pageNumber="575">Hyposmocoma</emphasis>
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</taxonomicName>
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. A,
|
||||
<emphasis id="B9193F690331A0420238237FFC0DFA22" bold="true" box="[580,921,1405,1425]" pageId="5" pageNumber="575">
|
||||
<taxonomicName id="4C6D98F80331A0420238237FFCA1FA23" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[580,821,1405,1425]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="anoai" status="sp. nov.">
|
||||
<emphasis id="B9193F690331A0420238237FFCA1FA23" bold="true" box="[580,821,1405,1425]" italics="true" pageId="5" pageNumber="575">Hyposmocoma anoai</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22A82120331A0420342237FFC0DFA22" box="[830,921,1406,1425]" pageId="5" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
|
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</emphasis>
|
||||
, holotype ♂, forewing
|
||||
<emphasis id="B9193F690331A04204F7237DFB0CFA23" box="[1163,1176,1404,1424]" italics="true" pageId="5" pageNumber="575">=</emphasis>
|
||||
4.5 mm; B,
|
||||
<emphasis id="B9193F690331A0420561237FFEFBFA1D" bold="true" pageId="5" pageNumber="575">
|
||||
<taxonomicName id="4C6D98F80331A0420561237FFE9AFA1E" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="hooilo" status="sp. nov.">
|
||||
<emphasis id="B9193F690331A0420561237FFE9AFA1E" bold="true" italics="true" pageId="5" pageNumber="575">Hyposmocoma hooilo</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22A82120331A0420169239AFEFBFA1D" box="[277,367,1435,1454]" pageId="5" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype ♀, forewing
|
||||
<emphasis id="B9193F690331A0420223239BFDF8FA1D" box="[607,620,1434,1454]" italics="true" pageId="5" pageNumber="575">=</emphasis>
|
||||
5.8 mm; C,
|
||||
<emphasis id="B9193F690331A042028C239AFBFCFA1D" bold="true" box="[752,1128,1434,1454]" pageId="5" pageNumber="575">
|
||||
<taxonomicName id="4C6D98F80331A042028C239AFB92FA1E" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[752,1030,1434,1454]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="kamaula" status="sp. nov.">
|
||||
<emphasis id="B9193F690331A042028C239AFB92FA1E" bold="true" box="[752,1030,1434,1454]" italics="true" pageId="5" pageNumber="575">Hyposmocoma kamaula</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22A82120331A0420472239AFBFCFA1D" box="[1038,1128,1435,1454]" pageId="5" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype ♀, forewing
|
||||
<emphasis id="B9193F690331A0420524239BFAF1FA1D" box="[1368,1381,1434,1454]" italics="true" pageId="5" pageNumber="575">=</emphasis>
|
||||
4.5 mm; D,
|
||||
<emphasis id="B9193F690331A042009A23B9FDCDFA78" bold="true" box="[230,601,1464,1483]" pageId="5" pageNumber="575">
|
||||
<taxonomicName id="4C6D98F80331A042009A23B9FE62FA78" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[230,502,1464,1483]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="kanaloa" status="sp. nov.">
|
||||
<emphasis id="B9193F690331A042009A23B9FE62FA78" bold="true" box="[230,502,1464,1483]" italics="true" pageId="5" pageNumber="575">Hyposmocoma kanaloa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22A82120331A042018223B9FDCDFA78" box="[510,601,1464,1483]" pageId="5" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype ♂, forewing
|
||||
<emphasis id="B9193F690331A042033723B6FCCCFA78" box="[843,856,1463,1483]" italics="true" pageId="5" pageNumber="575">=</emphasis>
|
||||
3.2 mm; E,
|
||||
<emphasis id="B9193F690331A04203A123B9FAC3FA78" bold="true" box="[989,1367,1464,1483]" pageId="5" pageNumber="575">
|
||||
<taxonomicName id="4C6D98F80331A04203A123B9FB67FA78" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[989,1267,1464,1483]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="mahoepo" status="sp. nov.">
|
||||
<emphasis id="B9193F690331A04203A123B9FB67FA78" bold="true" box="[989,1267,1464,1483]" italics="true" pageId="5" pageNumber="575">Hyposmocoma mahoepo</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22A82120331A042048023B9FAC3FA78" box="[1276,1367,1464,1483]" pageId="5" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype ♂, forewing
|
||||
<emphasis id="B9193F690331A042013023D5FECDFA5B" box="[332,345,1492,1512]" italics="true" pageId="5" pageNumber="575">=</emphasis>
|
||||
5.6 mm; F,
|
||||
<emphasis id="B9193F690331A042019B23D4FC14FA5B" bold="true" box="[487,896,1493,1512]" pageId="5" pageNumber="575">
|
||||
<taxonomicName id="4C6D98F80331A042019B23D4FC87FA5B" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[487,787,1493,1512]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="nohomeha" status="sp. nov.">
|
||||
<emphasis id="B9193F690331A042019B23D4FC87FA5B" bold="true" box="[487,787,1493,1512]" italics="true" pageId="5" pageNumber="575">Hyposmocoma nohomeha</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22A82120331A042035C23D4FC14FA5B" box="[800,896,1493,1512]" pageId="5" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype ♂, forewing
|
||||
<emphasis id="B9193F690331A04204FC23D5FB19FA5B" box="[1152,1165,1492,1512]" italics="true" pageId="5" pageNumber="575">=</emphasis>
|
||||
4.3 mm; G,
|
||||
<emphasis id="B9193F690331A042056123D4FEF1F9B5" bold="true" pageId="5" pageNumber="575">
|
||||
<taxonomicName id="4C6D98F80331A042056123D4FE97F9B6" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="oolea" status="sp. nov.">
|
||||
<emphasis id="B9193F690331A042056123D4FE97F9B6" bold="true" italics="true" pageId="5" pageNumber="575">Hyposmocoma oolea</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22A82120331A042017723F2FEF1F9B5" box="[267,357,1523,1542]" pageId="5" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype ♂, forewing
|
||||
<emphasis id="B9193F690331A042022923F0FDF6F9B6" box="[597,610,1521,1541]" italics="true" pageId="5" pageNumber="575">=</emphasis>
|
||||
5.0 mm; H,
|
||||
<emphasis id="B9193F690331A042029523F2FBFDF9B5" bold="true" box="[745,1129,1522,1542]" pageId="5" pageNumber="575">
|
||||
<taxonomicName id="4C6D98F80331A042029523F2FB92F9B6" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[745,1030,1522,1542]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="pahanalo" status="sp. nov.">
|
||||
<emphasis id="B9193F690331A042029523F2FB92F9B6" bold="true" box="[745,1030,1522,1542]" italics="true" pageId="5" pageNumber="575">Hyposmocoma pahanalo</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22A82120331A042047323F2FBFDF9B5" box="[1039,1129,1523,1542]" pageId="5" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype ♂, forewing
|
||||
<emphasis id="B9193F690331A042052423F0FAF1F9B6" box="[1368,1381,1521,1541]" italics="true" pageId="5" pageNumber="575">=</emphasis>
|
||||
5.8 mm; I,
|
||||
<emphasis id="B9193F690331A04200A12011FDD0F990" bold="true" box="[221,580,1551,1571]" pageId="5" pageNumber="575">
|
||||
<taxonomicName id="4C6D98F80331A04200A12011FE77F991" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[221,483,1551,1571]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="waauhi" status="sp. nov.">
|
||||
<emphasis id="B9193F690331A04200A12011FE77F991" bold="true" box="[221,483,1551,1571]" italics="true" pageId="5" pageNumber="575">Hyposmocoma waauhi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22A82120331A04201962011FDD0F990" box="[490,580,1552,1571]" pageId="5" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, paratype ♂, forewing
|
||||
<emphasis id="B9193F690331A042034B200EFCD0F990" box="[823,836,1551,1571]" italics="true" pageId="5" pageNumber="575">=</emphasis>
|
||||
4.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BD2E37B0331A04200BA205CFACDF95E" blockId="5.[198,1476,1629,1926]" pageId="5" pageNumber="575">
|
||||
flagellomeres very pale brown; scape dark brown. scales clustered near tip. Subcostal brush absent. Thorax, tegula, and metascutellum pale brown Sclerotized hook (pseuduncus
|
||||
<emphasis id="B9193F690331A04204A0207DFA8FF922" box="[1244,1307,1660,1681]" italics="true" pageId="5" pageNumber="575">sensu</emphasis>
|
||||
Zimmerman, dorsally, dark brown along sides; abdomen nearly 1978) elongate, slightly curved, slender; distinct white. Foreleg brown with rings of lighter scales at sclerotized ring on segment VII. Females similar to distal end of each segment. Midleg and hindleg males except frenulum with three acanthae.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C377B0F00331A04200BA20F4FA57F836" pageId="5" pageNumber="575" type="description">
|
||||
<paragraph id="8BD2E37B0331A04200BA20F4FA57F836" blockId="5.[198,1476,1629,1926]" pageId="5" pageNumber="575">
|
||||
almost entirely nearly white, with some dark scales
|
||||
<emphasis id="B9193F690331A042030920F4FBB3F8B9" box="[885,1063,1781,1802]" italics="true" pageId="5" pageNumber="575">Male genitalia:</emphasis>
|
||||
(
|
||||
<emphasis id="B9193F690331A042044220F7FBFDF8B9" box="[1086,1129,1782,1803]" italics="true" pageId="5" pageNumber="575">N =</emphasis>
|
||||
1) (
|
||||
<figureCitation id="1356FFFE0331A04204E220F4FB61F8B8" box="[1182,1269,1781,1803]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." pageId="5" pageNumber="575">Fig. 4B</figureCitation>
|
||||
). Valvae roughly present on femur. Forewing ground colour pale symmetrical, both with long and somewhat robust brown, with very dark brown band extending from arms, although left valva slightly larger than right. base of wing to termen; additional dark brown scales Both valvae bent nearly 90
|
||||
<emphasis id="B9193F690331A04204DF2153FB39F8D5" box="[1187,1197,1874,1894]" italics="true" pageId="5" pageNumber="575">°</emphasis>
|
||||
upward in the middle, in thin band along costal margin; fringe pale brown. with four heavy, long, sclerotized spur-like setae
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
325
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325
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@ -0,0 +1,325 @@
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<heading id="D09A54170339A04A029626BCFA5DFF60" box="[746,1481,184,211]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="583" reason="2">
|
||||
NEW
|
||||
<taxonomicName id="4C6D98F80339A04A035326BCFBC1FF62" authority="FROM" authorityName="FROM" box="[815,1109,188,210]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="583" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B9193F690339A04A035326BCFB95FF61" box="[815,1025,189,210]" italics="true" pageId="13" pageNumber="583">HYPOSMOCOMA</emphasis>
|
||||
FROM
|
||||
</taxonomicName>
|
||||
KAHOʻOLAWE
|
||||
<collectingCountry id="F37AA3EB0339A04A056A26BDFAEDFF62" box="[1302,1401,188,209]" name="Iceland" pageId="13" pageNumber="583">ISLAND</collectingCountry>
|
||||
583
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C377B0F00339A04A00A226FEFA57F834" pageId="13" pageNumber="583" type="description">
|
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<paragraph id="8BD2E37B0339A04A00A226FEFA57FDF5" blockId="13.[198,1476,253,1927]" pageId="13" pageNumber="583">
|
||||
<emphasis id="B9193F690339A04A00A226FEFE32FEA7" box="[222,422,255,276]" italics="true" pageId="13" pageNumber="583">Female genitalia:</emphasis>
|
||||
(
|
||||
<emphasis id="B9193F690339A04A01C426FEFE58FEA7" box="[440,460,255,276]" italics="true" pageId="13" pageNumber="583">N</emphasis>
|
||||
= 2) (
|
||||
<figureCitation id="1356FFFE0339A04A026D26FEFDFEFEA7" box="[529,618,254,277]" captionStart="Figure 5" captionStartId="7.[198,277,1241,1260]" captionTargetBox="[428,1254,253,1205]" captionTargetId="figure-325@7.[427,1255,251,1205]" captionTargetPageId="7" captionText="Figure 5. Female genitalia and larval cases of Hyposmocoma. A, Hyposmocoma hooilo sp. nov., slide WPH042- 13; B, Hyposmocoma kamaula sp. nov., slide WPH050-13; C, Hyposmocoma kanaloa sp. nov., slide WPH123- 13; D, Hyposmocoma mahoepo sp. nov., slide WPH053-13; E, Hyposmocoma nohomeha sp. nov., slide WPH076-13; F, Hyposmocoma oolea sp. nov., slide WPH048-13; G, Hyposmocoma pahanalo sp. nov., slide WPH044-13; H, Hyposmocoma kanaloa sp. nov., larval case, dorsal view, length = 4 mm; I, Hyposmocoma oolea sp. nov., larval case, dorsal view, length = 6 mm. Abbreviations: a, anterior apophysis; cb, corpus bursa; db, ductus bursa; o, ostium-bearing process; p, posterior apophysis; pa, papillae anales." pageId="13" pageNumber="583">Fig. 5G</figureCitation>
|
||||
). Papillae anales Antennal flagellomeres with alternating rings of short. Apophyses thin and straight; posterior apophy- dark and pale brown scales; scape dark brown; ses very long, about two times length of anterior pecten absent. Thorax, tegula, and metascutellum apophyses. Ostium-bearing process heavily sclero- mottled pale brown to brown. Foreleg and midleg tized and very large, externally protruding, snail-shell coxa, femur, tibia, and tarsomeres mostly dark shaped and curled, with broad base. Ductus bursae brown; fringe brown. Hindleg entirely nearly white. long, about same length as anterior apophysis. Corpus Forewing ground colour pale brown, with three or bursae oval, about same length as anterior apophysis; four wide bands of brown scales extending more or signum absent. Inception of ductus seminalis situated less vertically from costa to anal margin; several at posterior end of corpus bursae. brown spots near cell; some scattered brown scales near anal margin. Hindwing and fringe uniformly
|
||||
</paragraph>
|
||||
<paragraph id="8BD2E37B0339A04A00BA244EFB37FCAF" blockId="13.[198,1476,253,1927]" pageId="13" pageNumber="583">
|
||||
<emphasis id="B9193F690339A04A00BA244EFDEAFDD7" box="[198,638,591,612]" italics="true" pageId="13" pageNumber="583">
|
||||
Etymology:
|
||||
<taxonomicName id="4C6D98F80339A04A012E244EFDEAFDD7" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[338,638,591,612]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="583" phylum="Arthropoda" rank="species" species="pahanalo">Hyposmocoma pahanalo</taxonomicName>
|
||||
</emphasis>
|
||||
, or ‘hidden pale brown. Subcostal brush absent. Abdomen surfboard’, from the Hawaiian paha (a kind of brown. Sclerotized hook (pseuduncus
|
||||
<emphasis id="B9193F690339A04A05F8246FFA57FD30" box="[1412,1475,622,643]" italics="true" pageId="13" pageNumber="583">sensu</emphasis>
|
||||
surfboard) and nalo (missing); this refers to the Zimmerman, 1978) elongate, slender, with pointed presumed larval case of this species, as the ‘flat apex; distinct sclerotized ring on segment VII. purse’ cases in this clade are similar in shape to Female similar to males except frenulum with Hawaiian surfboards. apparently two acanthae and antennae slightly thinner than those of males.
|
||||
</paragraph>
|
||||
<paragraph id="8BD2E37B0339A04A00BA2524FA50FC06" blockId="13.[198,1476,253,1927]" pageId="13" pageNumber="583">
|
||||
<emphasis id="B9193F690339A04A00BA2524FEB5FC89" box="[198,289,805,826]" italics="true" pageId="13" pageNumber="583">Biology:</emphasis>
|
||||
Based on its position in the phylogenetic
|
||||
<emphasis id="B9193F690339A04A03092524FBB0FC89" box="[885,1060,805,826]" italics="true" pageId="13" pageNumber="583">Male genitalia:</emphasis>
|
||||
(
|
||||
<emphasis id="B9193F690339A04A044B2527FBDFFC88" box="[1079,1099,806,827]" italics="true" pageId="13" pageNumber="583">N</emphasis>
|
||||
= 2) (
|
||||
<figureCitation id="1356FFFE0339A04A04E82524FB77FC89" box="[1172,1251,805,827]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." pageId="13" pageNumber="583">Fig. 4I</figureCitation>
|
||||
). Valvae asymmettree, we expect larvae of
|
||||
<taxonomicName id="4C6D98F80339A04A027E2545FD05FCEB" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[514,657,835,857]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="583" phylum="Arthropoda" rank="species" species="pahanalo">
|
||||
<emphasis id="B9193F690339A04A027E2545FD05FCEB" box="[514,657,835,857]" italics="true" pageId="13" pageNumber="583">H. pahanalo</emphasis>
|
||||
</taxonomicName>
|
||||
to construct rical, left arm long and robust, enlarged apically, ‘flat purse’ case
|
||||
<typeStatus id="54D65DD90339A04A01F22563FE59FCC4" box="[398,461,866,887]" pageId="13" pageNumber="583">types</typeStatus>
|
||||
, based on its being sister to with four heavy, long, sclerotized spur-like setae proanother clade that makes this case
|
||||
<typeStatus id="54D65DD90339A04A02232580FD07FC25" box="[607,659,897,918]" pageId="13" pageNumber="583">type</typeStatus>
|
||||
. jecting dorsally near apex (lost in
|
||||
<figureCitation id="1356FFFE0339A04A04912581FAA8FC25" box="[1261,1340,896,918]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." pageId="13" pageNumber="583">Fig. 4I</figureCitation>
|
||||
); right arm long and slender with no heavy setae, apex deeply
|
||||
</paragraph>
|
||||
<paragraph id="8BD2E37B0339A04A00BA25BCFA50FBA2" blockId="13.[198,1476,253,1927]" pageId="13" pageNumber="583">
|
||||
<emphasis id="B9193F690339A04A00BA25BCFEC3FC61" box="[198,343,957,978]" italics="true" pageId="13" pageNumber="583">Distribution:</emphasis>
|
||||
Known only from Kahoʻolawe
|
||||
<collectingCountry id="F37AA3EB0339A04A02A225BCFCBCFC60" box="[734,808,957,979]" name="Iceland" pageId="13" pageNumber="583">Island</collectingCountry>
|
||||
, notched resulting in the appearance of one large and Hawai i, where it is presumed to be endemic. one small apical lobe. Right valva with a small lateral lobe projecting from near base; left valva with-
|
||||
</paragraph>
|
||||
<paragraph id="8BD2E37B0339A04A00942228FBE5FB39" blockId="13.[198,1476,253,1927]" pageId="13" pageNumber="583">
|
||||
<taxonomicName id="4C6D98F80339A04A00942228FD97FB8E" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[232,515,1065,1086]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="591" phylum="Arthropoda" rank="species" species="waauhi">
|
||||
<emphasis id="B9193F690339A04A00942228FD97FB8E" bold="true" box="[232,515,1065,1086]" italics="true" pageId="13" pageNumber="583">
|
||||
<smallCapsWord id="8D3475A70339A04A00942228FE0AFB8E" baselines="1081,1081" box="[232,414,1065,1086]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Hyposmocoma" pageId="13" pageNumber="583">HYPOSMOCOMA</smallCapsWord>
|
||||
<smallCapsWord id="8D3475A70339A04A01D4222DFD97FB8E" baselines="1081" box="[424,515,1068,1085]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="waauhi" pageId="13" pageNumber="583">WAAUHI</smallCapsWord>
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
<smallCapsWord id="8D3475A70339A04A02702227FD16FB8E" baselines="1080,1081" box="[524,642,1062,1086]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Medeiros" pageId="13" pageNumber="583">MEDEIROS</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3475A70339A04A02EF2227FD7DFB8E" baselines="1080,1081" box="[659,745,1062,1086]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Haines" pageId="13" pageNumber="583">HAINES</smallCapsWord>
|
||||
& out such a lobe. Anellus with two roughly symmetrical lobes, both slender and slightly apically
|
||||
<smallCapsWord id="8D3475A70339A04A01442244FE3EFBE8" baselines="1111,1111" box="[312,426,1093,1117]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Rubinoff" pageId="13" pageNumber="583">RUBINOFF</smallCapsWord>
|
||||
<emphasis id="B9193F690339A04A01C8224BFD8EFBED" bold="true" box="[436,538,1091,1118]" pageId="13" pageNumber="583">
|
||||
<taxonomicNameLabel id="A22A82120339A04A01C8224BFD8EFBED" box="[436,538,1091,1118]" pageId="13" pageNumber="583" rank="species">
|
||||
<smallCapsWord id="8D3475A70339A04A01C8224BFE44FBE8" baselines="1111" box="[436,464,1098,1115]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="13" pageNumber="583">SP</smallCapsWord>
|
||||
.
|
||||
<smallCapsWord id="8D3475A70339A04A019E224BFD86FBE8" baselines="1111" box="[482,530,1098,1115]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="13" pageNumber="583">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1356FFFE0339A04A02502244FD11FBEE" box="[556,645,1091,1117]" captionStart="Figure 3" captionStartId="5.[198,277,1405,1424]" captionTargetBox="[420,1247,251,1369]" captionTargetId="figure-28@5.[420,1248,251,1369]" captionTargetPageId="5" captionText="Figure 3. Adult Hyposmocoma. A, Hyposmocoma anoai sp. nov., holotype ♂, forewing = 4.5 mm; B, Hyposmocoma hooilo sp. nov., paratype ♀, forewing = 5.8 mm; C, Hyposmocoma kamaula sp. nov., paratype ♀, forewing = 4.5 mm; D, Hyposmocoma kanaloa sp. nov., holotype ♂, forewing = 3.2 mm; E, Hyposmocoma mahoepo sp. nov., paratype ♂, forewing = 5.6 mm; F, Hyposmocoma nohomeha sp. nov., holotype ♂, forewing = 4.3 mm; G, Hyposmocoma oolea sp. nov., holotype ♂, forewing = 5.0 mm; H, Hyposmocoma pahanalo sp. nov., holotype ♂, forewing = 5.8 mm; I, Hyposmocoma waauhi sp. nov., paratype ♂, forewing = 4.5 mm." pageId="13" pageNumber="583">
|
||||
<smallCapsWord id="8D3475A70339A04A02502244FDF5FBE8" baselines="1111,1111" box="[556,609,1093,1117]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="13" pageNumber="583">FIGS</smallCapsWord>
|
||||
3I
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1356FFFE0339A04A02EB2242FD24FBEE" box="[663,688,1091,1117]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." pageId="13" pageNumber="583">4I</figureCitation>
|
||||
) enlarged. Phallus large, cylindrical, heavily sclerotized, curved near apex.
|
||||
</paragraph>
|
||||
<paragraph id="8BD2E37B0339A04A00BA2292FC76FACC" blockId="13.[198,1476,253,1927]" pageId="13" pageNumber="583">
|
||||
<emphasis id="B9193F690339A04A00BA2292FE3BFB1B" box="[198,431,1171,1192]" italics="true" pageId="13" pageNumber="583">Material examined:</emphasis>
|
||||
<typeStatus id="54D65DD90339A04A01C22292FDC5FB1B" box="[446,593,1171,1192]" pageId="13" pageNumber="583" type="holotype">HOLOTYPE</typeStatus>
|
||||
: HI: Kahoʻolawe:
|
||||
<emphasis id="B9193F690339A04A03092295FBAAFB1A" box="[885,1086,1172,1193]" italics="true" pageId="13" pageNumber="583">Female genitalia:</emphasis>
|
||||
(
|
||||
<emphasis id="B9193F690339A04A042E2295FBF2FB1A" box="[1106,1126,1172,1193]" italics="true" pageId="13" pageNumber="583">N</emphasis>
|
||||
= 1). Specimen is in unsatis- Honokanaiʻa camp, 20 30
|
||||
<geoCoordinate id="EE5985BC0339A04A027B22AFFDDEFB74" box="[519,586,1198,1224]" degrees="0" direction="north" minutes="54" orientation="latitude" pageId="13" pageNumber="583" precision="15" seconds="00" value="0.9">
|
||||
<emphasis id="B9193F690339A04A027B22AFFD98FB0E" bold="true" box="[519,524,1198,1213]" italics="true" pageId="13" pageNumber="583">
|
||||
<superScript id="7C184E330339A04A027B22AFFD98FB0E" attach="none" box="[519,524,1198,1213]" fontSize="6" pageId="13" pageNumber="583">0</superScript>
|
||||
</emphasis>
|
||||
54
|
||||
<emphasis id="B9193F690339A04A025422AFFDA7FB0E" bold="true" box="[552,563,1198,1213]" italics="true" pageId="13" pageNumber="583">
|
||||
<superScript id="7C184E330339A04A025422AFFDA7FB0E" attach="right" box="[552,563,1198,1213]" fontSize="6" pageId="13" pageNumber="583">00</superScript>
|
||||
</emphasis>
|
||||
N
|
||||
</geoCoordinate>
|
||||
, 156 40
|
||||
<geoCoordinate id="EE5985BC0339A04A02CF22AFFD68FB74" box="[691,764,1198,1224]" degrees="0" direction="west" minutes="56" orientation="longitude" pageId="13" pageNumber="583" precision="15" seconds="00" value="-0.93333334">
|
||||
<emphasis id="B9193F690339A04A02CF22AFFD2CFB0E" bold="true" box="[691,696,1198,1213]" italics="true" pageId="13" pageNumber="583">
|
||||
<superScript id="7C184E330339A04A02CF22AFFD2CFB0E" attach="none" box="[691,696,1198,1213]" fontSize="6" pageId="13" pageNumber="583">0</superScript>
|
||||
</emphasis>
|
||||
56
|
||||
<emphasis id="B9193F690339A04A02A822AFFD4BFB0E" bold="true" box="[724,735,1198,1213]" italics="true" pageId="13" pageNumber="583">
|
||||
<superScript id="7C184E330339A04A02A822AFFD4BFB0E" attach="right" box="[724,735,1198,1213]" fontSize="6" pageId="13" pageNumber="583">00</superScript>
|
||||
</emphasis>
|
||||
W
|
||||
</geoCoordinate>
|
||||
.
|
||||
<specimenCount id="9D6B28F20339A04A036922AEFCB1FB74" box="[789,805,1199,1223]" count="1" pageId="13" pageNumber="583" type="male">♂</specimenCount>
|
||||
, factory condition to illustrate, as the genitalia are
|
||||
<date id="FFD3C5BB0339A04A00BA22D1FEA5FB55" box="[198,305,1232,1254]" pageId="13" pageNumber="583" value="2013-03-05">5.iii.2013</date>
|
||||
, W. Haines & D. Rubinoff, WPH074-13, damaged. Ductus bursa, corpus bursa, and ostium DNA extraction DN2180, slide WPH074-
|
||||
<specimenCount id="9D6B28F20339A04A02E122EEFD51FAB7" box="[669,709,1260,1284]" count="13" pageId="13" pageNumber="583" type="male">13♂</specimenCount>
|
||||
. unavailable for study. Papillae anales short. Apophy-
|
||||
<materialsCitation id="3B05E9260339A04A00A2230FFC49FACC" collectingDate="2008-10-20" collectingDateMax="2013-03-05" collectingDateMin="2008-10-20" collectorName="Hakioawa, D & M. J. Medeiros & W. Haines" latitude="0.9" location="Planting Area" longLatPrecision="21" longitude="-0.93333334" municipality="Inception" pageId="13" pageNumber="583" specimenCount="51" specimenCount-female="1" specimenCount-male="50" typeStatus="paratype">
|
||||
<typeStatus id="54D65DD90339A04A00A2230FFEEEFA90" box="[222,378,1294,1315]" pageId="13" pageNumber="583" type="paratype">PARATYPES</typeStatus>
|
||||
: HI: Kahoʻolawe:
|
||||
<collectorName id="269886AD0339A04A0205230CFC8AFA90" box="[633,798,1293,1315]" pageId="13" pageNumber="583">Hakioawa, D</collectorName>
|
||||
4 ses thin and straight; posterior apophyses long,
|
||||
<location id="8EB2B5A00339A04A00BA232DFEECFAF2" LSID="urn:lsid:plazi:treatment:03C4526D0339A04A029626BCFA57F834:8EB2B5A00339A04A00BA232DFEECFAF2" box="[198,376,1324,1346]" latitude="0.9" longLatPrecision="21" longitude="-0.93333334" municipality="Inception" name="Planting Area" pageId="13" pageNumber="583">Planting Area</location>
|
||||
.
|
||||
<specimenCount id="9D6B28F20339A04A01ED232DFE51FAF2" box="[401,453,1321,1345]" count="1" pageId="13" pageNumber="583" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<date id="FFD3C5BB0339A04A019E232DFDC5FAF2" box="[482,593,1324,1345]" pageId="13" pageNumber="583" value="2008-10-20">
|
||||
<collectingDate id="EF973C530339A04A019E232DFDC5FAF2" box="[482,593,1324,1345]" pageId="13" pageNumber="583" value="2008-10-20">20.x.2008</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="269886AD0339A04A0217232DFCBCFAF1" box="[619,808,1324,1346]" pageId="13" pageNumber="583">M. J. Medeiros</collectorName>
|
||||
, about 1.5
|
||||
<emphasis id="B9193F690339A04A03A8232CFC73FAF2" bold="true" box="[980,999,1325,1345]" italics="true" pageId="13" pageNumber="583">9</emphasis>
|
||||
length of anterior apophyses.
|
||||
<collectingMunicipality id="6BB679010339A04A0524232DFA57FAF1" box="[1368,1475,1324,1346]" pageId="13" pageNumber="583">Inception</collectingMunicipality>
|
||||
slide 08A
|
||||
<specimenCount id="9D6B28F20339A04A0139234AFEE4FAD3" box="[325,368,1352,1376]" count="48" pageId="13" pageNumber="583" type="male">48♂</specimenCount>
|
||||
. Honokanaiʻa camp, 20 30
|
||||
<geoCoordinate id="EE5985BC0339A04A02982347FCB3FAD3" box="[740,807,1350,1376]" degrees="0" direction="north" minutes="54" orientation="latitude" pageId="13" pageNumber="583" precision="15" seconds="00" value="0.9">
|
||||
<emphasis id="B9193F690339A04A02982347FD7DFAE6" bold="true" box="[740,745,1350,1365]" italics="true" pageId="13" pageNumber="583">
|
||||
<superScript id="7C184E330339A04A02982347FD7DFAE6" attach="none" box="[740,745,1350,1365]" fontSize="6" pageId="13" pageNumber="583">0</superScript>
|
||||
</emphasis>
|
||||
54
|
||||
<emphasis id="B9193F690339A04A03792347FC84FAE6" bold="true" box="[773,784,1350,1365]" italics="true" pageId="13" pageNumber="583">
|
||||
<superScript id="7C184E330339A04A03792347FC84FAE6" attach="right" box="[773,784,1350,1365]" fontSize="6" pageId="13" pageNumber="583">00</superScript>
|
||||
</emphasis>
|
||||
N
|
||||
</geoCoordinate>
|
||||
, of ductus seminalis situated at posterior end of cor- 156 40
|
||||
<geoCoordinate id="EE5985BC0339A04A016F2364FEC8FACD" box="[275,348,1381,1407]" degrees="0" direction="west" minutes="56" orientation="longitude" pageId="13" pageNumber="583" precision="15" seconds="00" value="-0.93333334">
|
||||
<emphasis id="B9193F690339A04A016F2364FE8CFAC7" bold="true" box="[275,280,1381,1396]" italics="true" pageId="13" pageNumber="583">
|
||||
<superScript id="7C184E330339A04A016F2364FE8CFAC7" attach="none" box="[275,280,1381,1396]" fontSize="6" pageId="13" pageNumber="583">0</superScript>
|
||||
</emphasis>
|
||||
56
|
||||
<emphasis id="B9193F690339A04A01482364FEABFAC7" bold="true" box="[308,319,1381,1396]" italics="true" pageId="13" pageNumber="583">
|
||||
<superScript id="7C184E330339A04A01482364FEABFAC7" attach="right" box="[308,319,1381,1396]" fontSize="6" pageId="13" pageNumber="583">00</superScript>
|
||||
</emphasis>
|
||||
W
|
||||
</geoCoordinate>
|
||||
.
|
||||
<specimenCount id="9D6B28F20339A04A01112368FE0CFACD" box="[365,408,1382,1406]" count="1" pageId="13" pageNumber="583" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D6B28F20339A04A01D12368FE41FACD" box="[429,469,1382,1406]" count="1" pageId="13" pageNumber="583" type="female">1 ♀</specimenCount>
|
||||
,
|
||||
<date id="FFD3C5BB0339A04A01952368FDC0FACC" box="[489,596,1385,1407]" pageId="13" pageNumber="583" value="2013-03-05">
|
||||
<collectingDate id="EF973C530339A04A01952368FDC0FACC" box="[489,596,1385,1407]" pageId="13" pageNumber="583" value="2013-03-05">5.iii.2013</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="269886AD0339A04A021A2368FD77FACC" box="[614,739,1385,1407]" pageId="13" pageNumber="583">W. Haines</collectorName>
|
||||
& D. pus bursae
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BD2E37B0339A04A00BA2389FCB8FA2D" blockId="13.[198,1476,253,1927]" box="[198,812,1415,1438]" pageId="13" pageNumber="583">Rubinoff, WPH057-13, DNA extraction DN2174, slide</paragraph>
|
||||
<paragraph id="8BD2E37B0339A04A00BA23A7FA57FA4A" blockId="13.[198,1476,253,1927]" pageId="13" pageNumber="583">
|
||||
WPH057-
|
||||
<specimenCount id="9D6B28F20339A04A014B23A7FECBFA08" box="[311,351,1443,1468]" count="13" pageId="13" pageNumber="583" type="male">13♂</specimenCount>
|
||||
; WPH075-13, DNA extraction DN2181,
|
||||
<emphasis id="B9193F690339A04A032123A6FB7FFA0F" box="[861,1259,1447,1468]" italics="true" pageId="13" pageNumber="583">
|
||||
Etymology:
|
||||
<taxonomicName id="4C6D98F80339A04A039523A6FB7FFA0F" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[1001,1259,1447,1468]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="583" phylum="Arthropoda" rank="species" species="waauhi">Hyposmocoma waauhi</taxonomicName>
|
||||
</emphasis>
|
||||
, or ‘hidden canoe’, slide WPH075-
|
||||
<specimenCount id="9D6B28F20339A04A010823C5FE0DFA69" box="[372,409,1474,1498]" count="13" pageId="13" pageNumber="583" type="female">13♀</specimenCount>
|
||||
. from the Hawaiian waʻa (canoe) and uhi (concealed or hidden). This species is so named because we
|
||||
</paragraph>
|
||||
<paragraph id="8BD2E37B0339A04A00BA2002FBD6F9E6" blockId="13.[198,1476,253,1927]" pageId="13" pageNumber="583">
|
||||
<emphasis id="B9193F690339A04A00BA2002FEA9F9AB" box="[198,317,1539,1560]" italics="true" pageId="13" pageNumber="583">Diagnosis:</emphasis>
|
||||
The forewing pattern is very similar to hypothesize that it builds ‘tube purse’ cases, their
|
||||
<taxonomicName id="4C6D98F80339A04A00BA2020FEDBF986" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[198,335,1568,1590]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="583" phylum="Arthropoda" rank="species" species="kamaula">
|
||||
<emphasis id="B9193F690339A04A00BA2020FEDBF986" box="[198,335,1568,1590]" italics="true" pageId="13" pageNumber="583">H. kamaula</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C6D98F80339A04A01E02020FE6EF986" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[412,506,1568,1590]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="583" phylum="Arthropoda" rank="species" species="oolea">
|
||||
<emphasis id="B9193F690339A04A01E02020FE6EF986" box="[412,506,1568,1590]" italics="true" pageId="13" pageNumber="583">H. oolea</emphasis>
|
||||
</taxonomicName>
|
||||
, in being mostly brown shapes resembling canoes, but we still have not with irregular darker spots and irregular dark bands collected the larvae.
|
||||
</paragraph>
|
||||
<paragraph id="8BD2E37B0339A04A00BA205CFA57F8BE" blockId="13.[198,1476,253,1927]" pageId="13" pageNumber="583">
|
||||
of scales extending from costal to anal margin of the forewing. The male genitalia are therefore required
|
||||
<emphasis id="B9193F690339A04A0321207DFC2CF922" box="[861,952,1660,1681]" italics="true" pageId="13" pageNumber="583">Biology:</emphasis>
|
||||
Based on its position in the phylogenetic tree, to diagnose this species. This is the only species of we expect larvae of
|
||||
<taxonomicName id="4C6D98F80339A04A042F209DFB58F903" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[1107,1228,1691,1713]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="583" phylum="Arthropoda" rank="species" species="waauhi">
|
||||
<emphasis id="B9193F690339A04A042F209DFB58F903" box="[1107,1228,1691,1713]" italics="true" pageId="13" pageNumber="583">H. waauhi</emphasis>
|
||||
</taxonomicName>
|
||||
to construct a ‘tube Kahoʻolawe
|
||||
<taxonomicName id="4C6D98F80339A04A011D20BBFD90F97C" box="[353,516,1722,1743]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="583" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B9193F690339A04A011D20BBFD90F97C" box="[353,516,1722,1743]" italics="true" pageId="13" pageNumber="583">Hyposmocoma</emphasis>
|
||||
</taxonomicName>
|
||||
with the right valva purse’ case, based on its being nested within a clade deeply notched. that makes this case
|
||||
<typeStatus id="54D65DD90339A04A042A20D9FB1EF95E" box="[1110,1162,1752,1773]" pageId="13" pageNumber="583">type</typeStatus>
|
||||
. Larvae of other species in the ‘tube purse’ clade can be found in a diversity of
|
||||
</paragraph>
|
||||
<paragraph id="8BD2E37B0339A04A00BA2114FAADF8FA" blockId="13.[198,1476,253,1927]" pageId="13" pageNumber="583">
|
||||
<emphasis id="B9193F690339A04A00BA2114FEDAF899" box="[198,334,1813,1834]" italics="true" pageId="13" pageNumber="583">Description:</emphasis>
|
||||
(
|
||||
<emphasis id="B9193F690339A04A01192114FEEDF899" box="[357,377,1813,1834]" italics="true" pageId="13" pageNumber="583">N</emphasis>
|
||||
= 4) (
|
||||
<figureCitation id="1356FFFE0339A04A01A32114FDBAF899" box="[479,558,1813,1835]" captionStart="Figure 3" captionStartId="5.[198,277,1405,1424]" captionTargetBox="[420,1247,251,1369]" captionTargetId="figure-28@5.[420,1248,251,1369]" captionTargetPageId="5" captionText="Figure 3. Adult Hyposmocoma. A, Hyposmocoma anoai sp. nov., holotype ♂, forewing = 4.5 mm; B, Hyposmocoma hooilo sp. nov., paratype ♀, forewing = 5.8 mm; C, Hyposmocoma kamaula sp. nov., paratype ♀, forewing = 4.5 mm; D, Hyposmocoma kanaloa sp. nov., holotype ♂, forewing = 3.2 mm; E, Hyposmocoma mahoepo sp. nov., paratype ♂, forewing = 5.6 mm; F, Hyposmocoma nohomeha sp. nov., holotype ♂, forewing = 4.3 mm; G, Hyposmocoma oolea sp. nov., holotype ♂, forewing = 5.0 mm; H, Hyposmocoma pahanalo sp. nov., holotype ♂, forewing = 5.8 mm; I, Hyposmocoma waauhi sp. nov., paratype ♂, forewing = 4.5 mm." pageId="13" pageNumber="583">Fig. 3I</figureCitation>
|
||||
). Wingspan 9.2
|
||||
<emphasis id="B9193F690339A04A035C2119FCB8F899" box="[800,812,1816,1834]" italics="true" pageId="13" pageNumber="583">–</emphasis>
|
||||
habitats, including decaying logs and leaf litter. They
|
||||
<quantity id="4C954E9E0339A04A00BA2135FEB9F8FA" box="[198,301,1844,1865]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="13" pageNumber="583" unit="mm" value="10.1">10.1 mm</quantity>
|
||||
. Head pale to medium brown. Haustellum presumably feed on decaying plant matter.
|
||||
</paragraph>
|
||||
<paragraph id="8BD2E37B0339A04A00BA2153FA57F834" blockId="13.[198,1476,253,1927]" pageId="13" pageNumber="583">
|
||||
with beige scales. Maxillary palpus reduced. Second segment of recurved labial palpus dark brown; third
|
||||
<emphasis id="B9193F690339A04A03212170FC7AF835" box="[861,1006,1905,1926]" italics="true" pageId="13" pageNumber="583">Distribution:</emphasis>
|
||||
Known only from Kahoʻolawe
|
||||
<collectingCountry id="F37AA3EB0339A04A05092170FA2BF834" box="[1397,1471,1905,1927]" name="Iceland" pageId="13" pageNumber="583">Island</collectingCountry>
|
||||
,
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
435
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435
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@ -0,0 +1,435 @@
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<mods:title id="D057B459576083B38EEEFD8949564E21">Small survivors: unexpected endemic diversity of Hyposmocoma (Lepidoptera: Cosmopterigidae) moths on Kaho olawe, a degraded Hawaiian island</mods:title>
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|
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|
||||
NEW
|
||||
<taxonomicName id="4C6D98F8033FA04C035326BCFBC1FF62" authority="FROM" authorityName="FROM" box="[815,1109,188,210]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B9193F69033FA04C035326BCFB95FF61" box="[815,1025,189,210]" italics="true" pageId="11" pageNumber="581">HYPOSMOCOMA</emphasis>
|
||||
FROM
|
||||
</taxonomicName>
|
||||
KAHOʻOLAWE
|
||||
<collectingCountry id="F37AA3EB033FA04C056A26BDFAEDFF62" box="[1302,1401,188,209]" name="Iceland" pageId="11" pageNumber="581">ISLAND</collectingCountry>
|
||||
581
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C377B0F0033FA04C00BA26FEFA57F834" pageId="11" pageNumber="581" type="materials_examined">
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<paragraph id="8BD2E37B033FA04C00BA26FEFA57F834" blockId="11.[198,1476,255,1927]" pageId="11" pageNumber="581">
|
||||
<materialsCitation id="3B05E926033FA04C00BA26FEFA57F834" collectingDate="2013-03-05" collectingDateMax="2013-04-26" collectingDateMin="2013-03-05" collectionCode="NOFF" collectorName="Thorax & W. Haines & D. Rubinoff & Foreleg & Base & Iki, N & D Rubinoff" country="Iceland" county="Material" latitude="0.38333333" location="Anellar" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" pageId="11" pageNumber="581" specimenCount="45" specimenCount-female="30" specimenCount-male="15" typeStatus="paratype">
|
||||
pecten present.
|
||||
<collectorName id="269886AD033FA04C01F426FEFE48FEA6" box="[392,476,255,277]" pageId="11" pageNumber="581">Thorax</collectorName>
|
||||
, tegula, and metascutellum 2013,
|
||||
<collectorName id="269886AD033FA04C03DB26FEFBB7FEA6" box="[935,1059,255,277]" pageId="11" pageNumber="581">W. Haines</collectorName>
|
||||
&
|
||||
<collectorName id="269886AD033FA04C042826FEFB4AFEA6" box="[1108,1246,255,277]" pageId="11" pageNumber="581">D. Rubinoff</collectorName>
|
||||
, WPH047-13, DNA mostly white, some scales tipped with brown; extraction DN2164, slide WPH047-
|
||||
<specimenCount id="9D6B28F2033FA04C049F271CFA9EFE81" box="[1251,1290,282,306]" count="13" pageId="11" pageNumber="581" type="male">13♂</specimenCount>
|
||||
. abdomen pale grey.
|
||||
<collectorName id="269886AD033FA04C01CF273DFD9CFEE1" box="[435,520,316,338]" pageId="11" pageNumber="581">Foreleg</collectorName>
|
||||
and midleg coxa, femur,
|
||||
<typeStatus id="54D65DD9033FA04C0309273DFB85FEE2" box="[885,1041,316,337]" pageId="11" pageNumber="581" type="paratype">PARATYPES</typeStatus>
|
||||
: HI: Kahoʻolawe:
|
||||
<collectorName id="269886AD033FA04C0482273DFAA1FEE2" box="[1278,1333,316,337]" pageId="11" pageNumber="581">Base</collectorName>
|
||||
of Puʻu o tibia, and tarsomeres mostly dark brown with some Moaʻula
|
||||
<collectorName id="269886AD033FA04C03B5275BFB85FEC3" box="[969,1041,346,368]" pageId="11" pageNumber="581">Iki, N</collectorName>
|
||||
<geoCoordinate id="EE5985BC033FA04C046E275BFB0FFEC3" box="[1042,1179,346,368]" degrees="20.56457" direction="west" orientation="longitude" pageId="11" pageNumber="581" precision="1" value="-20.56457">20.56457 W</geoCoordinate>
|
||||
156.58519.
|
||||
<specimenCount id="9D6B28F2033FA04C055E275BFADFFEDC" box="[1314,1355,343,367]" count="1" pageId="11" pageNumber="581" type="female">1 ♀</specimenCount>
|
||||
, on boullighter scales scattered throughout, especially as ders, ‘burrito’ case,
|
||||
<date id="FFD3C5BB033FA04C04272778FB52FE3C" box="[1115,1222,377,399]" pageId="11" pageNumber="581" value="2013-03-07">
|
||||
<collectingDate id="EF973C53033FA04C04272778FB52FE3C" box="[1115,1222,377,399]" pageId="11" pageNumber="581" value="2013-03-07">7.iii.2013</collectingDate>
|
||||
</date>
|
||||
, em.
|
||||
<date id="FFD3C5BB033FA04C056B2778FA1AFE3C" box="[1303,1422,377,399]" pageId="11" pageNumber="581" value="2013-04-26">
|
||||
<collectingDate id="EF973C53033FA04C056B2778FA1AFE3C" box="[1303,1422,377,399]" pageId="11" pageNumber="581" value="2013-04-26">26.iv.2013</collectingDate>
|
||||
</date>
|
||||
, W. rings around tarsomeres. Hindleg entirely nearly Haines &
|
||||
<collectorName id="269886AD033FA04C03AB2799FBF5FE1E" box="[983,1121,407,429]" pageId="11" pageNumber="581">D. Rubinoff</collectorName>
|
||||
, WPH147-13, DR13C7C, slide white. Forewing ground colour white, with dark WPH147-
|
||||
<specimenCount id="9D6B28F2033FA04C03B127B7FC67FE78" box="[973,1011,435,459]" count="13" pageId="11" pageNumber="581" type="female">13♀</specimenCount>
|
||||
. Honokanaiʻa camp. 20 30
|
||||
<geoCoordinate id="EE5985BC033FA04C050727B3FA2AFE78" box="[1403,1470,434,460]" degrees="0" direction="north" minutes="54" orientation="latitude" pageId="11" pageNumber="581" precision="15" seconds="00" value="0.9">
|
||||
<emphasis id="B9193F69033FA04C050727B3FA14FE72" bold="true" box="[1403,1408,434,449]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C050727B3FA14FE72" attach="none" box="[1403,1408,434,449]" fontSize="6" pageId="11" pageNumber="581">0</superScript>
|
||||
</emphasis>
|
||||
54
|
||||
<emphasis id="B9193F69033FA04C05E027B3FA33FE72" bold="true" box="[1436,1447,434,449]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C05E027B3FA33FE72" attach="right" box="[1436,1447,434,449]" fontSize="6" pageId="11" pageNumber="581">00</superScript>
|
||||
</emphasis>
|
||||
N
|
||||
</geoCoordinate>
|
||||
, brown scales scattered throughout, especially near 156 40
|
||||
<geoCoordinate id="EE5985BC033FA04C03D627D1FC67FE59" box="[938,1011,464,490]" degrees="0" direction="west" minutes="56" orientation="longitude" pageId="11" pageNumber="581" precision="15" seconds="00" value="-0.93333334">
|
||||
<emphasis id="B9193F69033FA04C03D627D1FC3BFE6C" bold="true" box="[938,943,464,479]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C03D627D1FC3BFE6C" attach="none" box="[938,943,464,479]" fontSize="6" pageId="11" pageNumber="581">0</superScript>
|
||||
</emphasis>
|
||||
56
|
||||
<emphasis id="B9193F69033FA04C03B727D1FC42FE6C" bold="true" box="[971,982,464,479]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C03B727D1FC42FE6C" attach="right" box="[971,982,464,479]" fontSize="6" pageId="11" pageNumber="581">00</superScript>
|
||||
</emphasis>
|
||||
W
|
||||
</geoCoordinate>
|
||||
.
|
||||
<specimenCount id="9D6B28F2033FA04C047827D4FBBBFE59" box="[1028,1071,466,490]" count="1" pageId="11" pageNumber="581" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D6B28F2033FA04C043827D4FBF8FE59" box="[1092,1132,466,490]" count="1" pageId="11" pageNumber="581" type="female">1 ♀</specimenCount>
|
||||
,
|
||||
<date id="FFD3C5BB033FA04C04FC27D5FB78FE59" box="[1152,1260,468,490]" pageId="11" pageNumber="581" value="2013-03-05">
|
||||
<collectingDate id="EF973C53033FA04C04FC27D5FB78FE59" box="[1152,1260,468,490]" pageId="11" pageNumber="581" value="2013-03-05">5.iii.2013</collectingDate>
|
||||
</date>
|
||||
.
|
||||
<collectorName id="269886AD033FA04C048127D4FAEEFE58" box="[1277,1402,469,491]" pageId="11" pageNumber="581">W. Haines</collectorName>
|
||||
& D. costal margin; fringe minimal, nearly white. Rubinoff, WHP051-13. DNA extraction DN2168; Hindwing and fringe uniformly brown. Subcostal WPH073-13, DNA extraction 2179. Kaukamoku brush apparently absent. No sclerotized hook present gulch, 20 33
|
||||
<geoCoordinate id="EE5985BC033FA04C0388242DFBACFDF5" box="[1012,1080,556,582]" degrees="0" direction="north" minutes="33" orientation="latitude" pageId="11" pageNumber="581" precision="15" seconds="00" value="0.55">
|
||||
<emphasis id="B9193F69033FA04C0388242DFC6DFD88" bold="true" box="[1012,1017,556,571]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C0388242DFC6DFD88" attach="none" box="[1012,1017,556,571]" fontSize="6" pageId="11" pageNumber="581">0</superScript>
|
||||
</emphasis>
|
||||
33
|
||||
<emphasis id="B9193F69033FA04C046A242DFBB5FD88" bold="true" box="[1046,1057,556,571]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C046A242DFBB5FD88" attach="right" box="[1046,1057,556,571]" fontSize="6" pageId="11" pageNumber="581">00</superScript>
|
||||
</emphasis>
|
||||
N
|
||||
</geoCoordinate>
|
||||
, 156 35
|
||||
<geoCoordinate id="EE5985BC033FA04C04E1242DFB72FDF5" box="[1181,1254,556,582]" degrees="0" direction="west" minutes="13" orientation="longitude" pageId="11" pageNumber="581" precision="15" seconds="00" value="-0.21666667">
|
||||
<emphasis id="B9193F69033FA04C04E1242DFB36FD88" bold="true" box="[1181,1186,556,571]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C04E1242DFB36FD88" attach="none" box="[1181,1186,556,571]" fontSize="6" pageId="11" pageNumber="581">0</superScript>
|
||||
</emphasis>
|
||||
13
|
||||
<emphasis id="B9193F69033FA04C04C2242DFB5DFD88" bold="true" box="[1214,1225,556,571]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C04C2242DFB5DFD88" attach="right" box="[1214,1225,556,571]" fontSize="6" pageId="11" pageNumber="581">00</superScript>
|
||||
</emphasis>
|
||||
W
|
||||
</geoCoordinate>
|
||||
.
|
||||
<specimenCount id="9D6B28F2033FA04C0480242CFA9DFDF6" box="[1276,1289,557,581]" count="1" pageId="11" pageNumber="581" type="female">♀</specimenCount>
|
||||
,
|
||||
<date id="FFD3C5BB033FA04C055E2431FA19FDF5" box="[1314,1421,560,582]" pageId="11" pageNumber="581" value="2013-03-07">
|
||||
<collectingDate id="EF973C53033FA04C055E2431FA19FDF5" box="[1314,1421,560,582]" pageId="11" pageNumber="581" value="2013-03-07">7.iii.2013</collectingDate>
|
||||
</date>
|
||||
, W. on segment VII. Females similar to males except Haines &
|
||||
<collectorName id="269886AD033FA04C03A4244EFBF6FDD6" box="[984,1122,591,613]" pageId="11" pageNumber="581">D. Rubinoff</collectorName>
|
||||
, WPH048-13, DNA extraction frenulum with three acanthae and antennae slightly DN2165, slide WPH048-
|
||||
<specimenCount id="9D6B28F2033FA04C0402246CFB37FD31" box="[1150,1187,618,642]" count="13" pageId="11" pageNumber="581" type="female">13♀</specimenCount>
|
||||
. Kealialalo, 20 32
|
||||
<geoCoordinate id="EE5985BC033FA04C05072468FA2AFD30" box="[1403,1470,617,643]" degrees="0" direction="north" minutes="23" orientation="latitude" pageId="11" pageNumber="581" precision="15" seconds="00" value="0.38333333">
|
||||
<emphasis id="B9193F69033FA04C05072468FA14FDCB" bold="true" box="[1403,1408,617,632]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C05072468FA14FDCB" attach="none" box="[1403,1408,617,632]" fontSize="6" pageId="11" pageNumber="581">0</superScript>
|
||||
</emphasis>
|
||||
23
|
||||
<emphasis id="B9193F69033FA04C05E02468FA33FDCB" bold="true" box="[1436,1447,617,632]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C05E02468FA33FDCB" attach="right" box="[1436,1447,617,632]" fontSize="6" pageId="11" pageNumber="581">00</superScript>
|
||||
</emphasis>
|
||||
N
|
||||
</geoCoordinate>
|
||||
, thinner than those of males. 156 38
|
||||
<geoCoordinate id="EE5985BC033FA04C03D62489FC67FD12" box="[938,1011,648,674]" degrees="0" direction="west" minutes="15" orientation="longitude" pageId="11" pageNumber="581" precision="15" seconds="00" value="-0.25">
|
||||
<emphasis id="B9193F69033FA04C03D62489FC3BFD24" bold="true" box="[938,943,648,663]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C03D62489FC3BFD24" attach="none" box="[938,943,648,663]" fontSize="6" pageId="11" pageNumber="581">0</superScript>
|
||||
</emphasis>
|
||||
15
|
||||
<emphasis id="B9193F69033FA04C03B72489FC42FD24" bold="true" box="[971,982,648,663]" italics="true" pageId="11" pageNumber="581">
|
||||
<superScript id="7C184E33033FA04C03B72489FC42FD24" attach="right" box="[971,982,648,663]" fontSize="6" pageId="11" pageNumber="581">00</superScript>
|
||||
</emphasis>
|
||||
W
|
||||
</geoCoordinate>
|
||||
.
|
||||
<specimenCount id="9D6B28F2033FA04C047E248DFBBDFD12" box="[1026,1065,649,673]" count="1" pageId="11" pageNumber="581" type="female">1 ♀</specimenCount>
|
||||
,
|
||||
<date id="FFD3C5BB033FA04C0440248DFB32FD11" box="[1084,1190,652,674]" pageId="11" pageNumber="581" value="2013-03-05">
|
||||
<collectingDate id="EF973C53033FA04C0440248DFB32FD11" box="[1084,1190,652,674]" pageId="11" pageNumber="581" value="2013-03-05">5.iii.2013</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="269886AD033FA04C04CB248DFAA6FD11" box="[1207,1330,652,674]" pageId="11" pageNumber="581">W. Haines</collectorName>
|
||||
& D. Rubi-
|
||||
<emphasis id="B9193F69033FA04C00A224ABFE1FFD0C" box="[222,395,682,703]" italics="true" pageId="11" pageNumber="581">Male genitalia:</emphasis>
|
||||
(
|
||||
<emphasis id="B9193F69033FA04C01E024AAFE24FD73" box="[412,432,683,704]" italics="true" pageId="11" pageNumber="581">N</emphasis>
|
||||
= 1) (
|
||||
<figureCitation id="1356FFFE033FA04C018A24ABFDD9FD73" box="[502,589,682,704]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." pageId="11" pageNumber="581">Fig. 4F</figureCitation>
|
||||
). Valvae symmetri- noff, WPH081-13, DNA extraction DN2187. Near cal, simple, slightly upcurved, enlarged and rounded Honokanaiʻa camp, N
|
||||
<geoCoordinate id="EE5985BC033FA04C042124C8FB73FD6C" box="[1117,1255,713,735]" degrees="20.52256" direction="west" orientation="longitude" pageId="11" pageNumber="581" precision="1" value="-20.52256">20.52256 W</geoCoordinate>
|
||||
156.68657.
|
||||
<specimenCount id="9D6B28F2033FA04C051724C8FA01FD6D" box="[1387,1429,710,734]" count="1" pageId="11" pageNumber="581" type="male">1 ♂</specimenCount>
|
||||
, on apically.
|
||||
<location id="8EB2B5A0033FA04C014B24E6FE18FD4E" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C014B24E6FE18FD4E" box="[311,396,743,765]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Anellar" pageId="11" pageNumber="581">Anellar</location>
|
||||
lobes not visible.
|
||||
<location id="8EB2B5A0033FA04C02FF24E6FD4EFD4E" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C02FF24E6FD4EFD4E" box="[643,730,743,765]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Phallus" pageId="11" pageNumber="581">Phallus</location>
|
||||
large, boulder, ‘burrito’ case,
|
||||
<date id="FFD3C5BB033FA04C040424E6FB77FD4E" box="[1144,1251,743,765]" pageId="11" pageNumber="581" value="2013-03-06">
|
||||
<collectingDate id="EF973C53033FA04C040424E6FB77FD4E" box="[1144,1251,743,765]" pageId="11" pageNumber="581" value="2013-03-06">6.iii.2013</collectingDate>
|
||||
</date>
|
||||
, em.
|
||||
<date id="FFD3C5BB033FA04C055324E6FA0CFD4E" box="[1327,1432,743,765]" pageId="11" pageNumber="581" value="2013-04-09">
|
||||
<collectingDate id="EF973C53033FA04C055324E6FA0CFD4E" box="[1327,1432,743,765]" pageId="11" pageNumber="581" value="2013-04-09">9.iv.2013</collectingDate>
|
||||
</date>
|
||||
, W cylindrical, heavily sclerotized, straight.
|
||||
<location id="8EB2B5A0033FA04C03212507FC39FCAF" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C03212507FC39FCAF" box="[861,941,774,796]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Haines" pageId="11" pageNumber="581">Haines</location>
|
||||
&
|
||||
<collectorName id="269886AD033FA04C03A62506FBCAFCAF" box="[986,1118,774,796]" pageId="11" pageNumber="581">D Rubinoff</collectorName>
|
||||
, WPH121-13, DNA extraction
|
||||
<emphasis id="B9193F69033FA04C00A22524FE31FC89" box="[222,421,805,826]" italics="true" pageId="11" pageNumber="581">
|
||||
<location id="8EB2B5A0033FA04C00A22524FEA4FC89" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C00A22524FEA4FC89" box="[222,304,805,826]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Female" pageId="11" pageNumber="581">Female</location>
|
||||
genitalia:
|
||||
</emphasis>
|
||||
(
|
||||
<emphasis id="B9193F69033FA04C01CA2524FE5EFC89" box="[438,458,805,826]" italics="true" pageId="11" pageNumber="581">N</emphasis>
|
||||
= 1) (
|
||||
<figureCitation id="1356FFFE033FA04C026D2524FDFCFC88" box="[529,616,805,827]" captionStart="Figure 5" captionStartId="7.[198,277,1241,1260]" captionTargetBox="[428,1254,253,1205]" captionTargetId="figure-325@7.[427,1255,251,1205]" captionTargetPageId="7" captionText="Figure 5. Female genitalia and larval cases of Hyposmocoma. A, Hyposmocoma hooilo sp. nov., slide WPH042- 13; B, Hyposmocoma kamaula sp. nov., slide WPH050-13; C, Hyposmocoma kanaloa sp. nov., slide WPH123- 13; D, Hyposmocoma mahoepo sp. nov., slide WPH053-13; E, Hyposmocoma nohomeha sp. nov., slide WPH076-13; F, Hyposmocoma oolea sp. nov., slide WPH048-13; G, Hyposmocoma pahanalo sp. nov., slide WPH044-13; H, Hyposmocoma kanaloa sp. nov., larval case, dorsal view, length = 4 mm; I, Hyposmocoma oolea sp. nov., larval case, dorsal view, length = 6 mm. Abbreviations: a, anterior apophysis; cb, corpus bursa; db, ductus bursa; o, ostium-bearing process; p, posterior apophysis; pa, papillae anales." pageId="11" pageNumber="581">Fig. 5E</figureCitation>
|
||||
).
|
||||
<location id="8EB2B5A0033FA04C02012524FD4EFC88" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C02012524FD4EFC88" box="[637,730,805,827]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Papillae" pageId="11" pageNumber="581">Papillae</location>
|
||||
anales DN2219. short.
|
||||
<location id="8EB2B5A0033FA04C015F2542FE08FCEA" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C015F2542FE08FCEA" box="[291,412,835,857]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Apophyses" pageId="11" pageNumber="581">Apophyses</location>
|
||||
thin and straight; posterior apophyses very long, about two times length of ante-
|
||||
<emphasis id="B9193F69033FA04C03212563FC40FCC4" box="[861,980,866,887]" italics="true" pageId="11" pageNumber="581">
|
||||
<location id="8EB2B5A0033FA04C03212563FC5AFCC4" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C03212563FC5AFCC4" box="[861,974,866,887]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Diagnosis" pageId="11" pageNumber="581">Diagnosis</location>
|
||||
:
|
||||
</emphasis>
|
||||
<location id="8EB2B5A0033FA04C039E2563FC69FCCB" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C039E2563FC69FCCB" box="[994,1021,866,888]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Of" pageId="11" pageNumber="581">Of</location>
|
||||
the
|
||||
<emphasis id="B9193F69033FA04C04372563FA17FCC4" box="[1099,1411,866,887]" italics="true" pageId="11" pageNumber="581">
|
||||
Kahoʻolawe
|
||||
<taxonomicName id="4C6D98F8033FA04C049C2563FA17FCC4" box="[1248,1411,866,887]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="genus">Hyposmocoma</taxonomicName>
|
||||
</emphasis>
|
||||
, the rior apophyses.
|
||||
<location id="8EB2B5A0033FA04C01ED2580FE76FC25" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C01ED2580FE76FC25" box="[401,482,897,918]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Ductus" pageId="11" pageNumber="581">Ductus</location>
|
||||
bursae long, about same forewing pattern of
|
||||
<taxonomicName id="4C6D98F8033FA04C042F2580FB23FC26" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[1107,1207,896,918]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="oolea">
|
||||
<emphasis id="B9193F69033FA04C042F2580FB23FC26" box="[1107,1207,896,918]" italics="true" pageId="11" pageNumber="581">H. oolea</emphasis>
|
||||
</taxonomicName>
|
||||
is very similar to
|
||||
<emphasis id="B9193F69033FA04C05D42580FA50FC25" box="[1448,1476,897,918]" italics="true" pageId="11" pageNumber="581">H.</emphasis>
|
||||
length as anterior apophysis.
|
||||
<location id="8EB2B5A0033FA04C0245259EFD1EFC07" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C0245259EFD1EFC07" box="[569,650,927,948]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Corpus" pageId="11" pageNumber="581">Corpus</location>
|
||||
bursae oval,
|
||||
<taxonomicName id="4C6D98F8033FA04C0321259EFC56FC07" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[861,962,927,948]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="kamaula">
|
||||
<emphasis id="B9193F69033FA04C0321259EFC56FC07" box="[861,962,927,948]" italics="true" pageId="11" pageNumber="581">kamaula</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C6D98F8033FA04C038325A1FBEDFC07" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[1023,1145,927,949]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="waauhi">
|
||||
<emphasis id="B9193F69033FA04C038325A1FBEDFC07" box="[1023,1145,927,949]" italics="true" pageId="11" pageNumber="581">H. waauhi</emphasis>
|
||||
</taxonomicName>
|
||||
in being mostly brown with about same length as anterior apophysis; two small irregular darker bands that appear as spots, and so circular signa present near centre of corpus bursa. the male genitalia are required to diagnose this
|
||||
<location id="8EB2B5A0033FA04C00BA25FAFEA5FBA2" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C00BA25FAFEA5FBA2" box="[198,305,1019,1041]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Inception" pageId="11" pageNumber="581">Inception</location>
|
||||
of ductus seminalis situated at posterior species.
|
||||
<location id="8EB2B5A0033FA04C03BD25FBFC79FBA3" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C03BD25FBFC79FBA3" box="[961,1005,1018,1040]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="The" pageId="11" pageNumber="581">The</location>
|
||||
male genitalia have a crescent-shaped end of corpus bursae. left lobe of the anellus, as well as a fifth seta on the lobe of the left valva; no other Kahoʻolawe
|
||||
<emphasis id="B9193F69033FA04C00BA2257FD15FBD8" box="[198,641,1110,1132]" italics="true" pageId="11" pageNumber="581">
|
||||
<location id="8EB2B5A0033FA04C00BA2257FEAAFBD8" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C00BA2257FEAAFBD8" box="[198,318,1110,1131]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Etymology" pageId="11" pageNumber="581">Etymology</location>
|
||||
:
|
||||
<taxonomicName id="4C6D98F8033FA04C012E2256FD15FBD8" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[338,641,1110,1132]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="nohomeha">Hyposmocoma nohomeha</taxonomicName>
|
||||
</emphasis>
|
||||
, or ‘hermit’,
|
||||
<taxonomicName id="4C6D98F8033FA04C03212256FB94FBDF" box="[861,1024,1111,1132]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B9193F69033FA04C03212256FB94FBDF" box="[861,1024,1111,1132]" italics="true" pageId="11" pageNumber="581">Hyposmocoma</emphasis>
|
||||
</taxonomicName>
|
||||
has this combination of characters. from the
|
||||
<location id="8EB2B5A0033FA04C01242274FE53FB38" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C01242274FE53FB38" box="[344,455,1141,1163]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Hawaiian" pageId="11" pageNumber="581">Hawaiian</location>
|
||||
noho (reside) and meha (isolated/alone), so named because the phylogeny
|
||||
<emphasis id="B9193F69033FA04C03212295FC71FB1A" box="[861,997,1172,1193]" italics="true" pageId="11" pageNumber="581">
|
||||
<location id="8EB2B5A0033FA04C03212295FC4BFB1A" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C03212295FC4BFB1A" box="[861,991,1172,1193]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Description" pageId="11" pageNumber="581">Description</location>
|
||||
:
|
||||
</emphasis>
|
||||
(
|
||||
<emphasis id="B9193F69033FA04C03802295FB84FB1A" box="[1020,1040,1172,1193]" italics="true" pageId="11" pageNumber="581">N</emphasis>
|
||||
= 7) (
|
||||
<figureCitation id="1356FFFE033FA04C04132292FB5CFB1B" box="[1135,1224,1171,1193]" captionStart="Figure 3" captionStartId="5.[198,277,1405,1424]" captionTargetBox="[420,1247,251,1369]" captionTargetId="figure-28@5.[420,1248,251,1369]" captionTargetPageId="5" captionText="Figure 3. Adult Hyposmocoma. A, Hyposmocoma anoai sp. nov., holotype ♂, forewing = 4.5 mm; B, Hyposmocoma hooilo sp. nov., paratype ♀, forewing = 5.8 mm; C, Hyposmocoma kamaula sp. nov., paratype ♀, forewing = 4.5 mm; D, Hyposmocoma kanaloa sp. nov., holotype ♂, forewing = 3.2 mm; E, Hyposmocoma mahoepo sp. nov., paratype ♂, forewing = 5.6 mm; F, Hyposmocoma nohomeha sp. nov., holotype ♂, forewing = 4.3 mm; G, Hyposmocoma oolea sp. nov., holotype ♂, forewing = 5.0 mm; H, Hyposmocoma pahanalo sp. nov., holotype ♂, forewing = 5.8 mm; I, Hyposmocoma waauhi sp. nov., paratype ♂, forewing = 4.5 mm." pageId="11" pageNumber="581">Fig. 3G</figureCitation>
|
||||
).
|
||||
<location id="8EB2B5A0033FA04C04882292FAF2FB1A" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C04882292FAF2FB1A" box="[1268,1382,1171,1193]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Wingspan" pageId="11" pageNumber="581">Wingspan</location>
|
||||
10.0
|
||||
<emphasis id="B9193F69033FA04C05CB2297FA57FB1B" box="[1463,1475,1174,1192]" italics="true" pageId="11" pageNumber="581">–</emphasis>
|
||||
suggests that this species has been isolated from
|
||||
<quantity id="4C954E9E033FA04C032122B3FC50FB74" box="[861,964,1202,1224]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="11" pageNumber="581" unit="mm" value="11.6">11.6 mm</quantity>
|
||||
.
|
||||
<location id="8EB2B5A0033FA04C03AE22B3FB9AFB7B" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C03AE22B3FB9AFB7B" box="[978,1038,1202,1224]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Head" pageId="11" pageNumber="581">Head</location>
|
||||
dark brown.
|
||||
<location id="8EB2B5A0033FA04C04D322B3FAA1FB7B" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C04D322B3FAA1FB7B" box="[1199,1333,1202,1224]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Haustellum" pageId="11" pageNumber="581">Haustellum</location>
|
||||
with brown its nearest relatives for a long time on a small scales.
|
||||
<location id="8EB2B5A0033FA04C03C422D1FBB3FB55" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C03C422D1FBB3FB55" box="[952,1063,1232,1254]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Maxillary" pageId="11" pageNumber="581">Maxillary</location>
|
||||
palpus reduced.
|
||||
<location id="8EB2B5A0033FA04C057B22D1FAE6FB55" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C057B22D1FAE6FB55" box="[1287,1394,1232,1254]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Recurved" pageId="11" pageNumber="581">Recurved</location>
|
||||
labial island. palpus with second segment nearly white with darker scales scattered throughout; third segment
|
||||
<emphasis id="B9193F69033FA04C00BA232DFEB5FAF2" box="[198,289,1324,1345]" italics="true" pageId="11" pageNumber="581">
|
||||
<location id="8EB2B5A0033FA04C00BA232DFE8FFAF2" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C00BA232DFE8FFAF2" box="[198,283,1324,1345]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Biology" pageId="11" pageNumber="581">Biology</location>
|
||||
:
|
||||
</emphasis>
|
||||
<location id="8EB2B5A0033FA04C0153232DFEE0FAF1" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C0153232DFEE0FAF1" box="[303,372,1324,1346]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Based" pageId="11" pageNumber="581">Based</location>
|
||||
on its position in the phylogenetic dark brown.
|
||||
<location id="8EB2B5A0033FA04C0475232DFBE7FAF1" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C0475232DFBE7FAF1" box="[1033,1139,1324,1346]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Antennal" pageId="11" pageNumber="581">Antennal</location>
|
||||
flagellomeres dark brown tree, we expect larvae of
|
||||
<taxonomicName id="4C6D98F8033FA04C0187234AFD00FAD3" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[507,660,1355,1376]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="nohomeha">
|
||||
<emphasis id="B9193F69033FA04C0187234AFD00FAD3" box="[507,660,1355,1376]" italics="true" pageId="11" pageNumber="581">H. nohomeha</emphasis>
|
||||
</taxonomicName>
|
||||
to construct scales; scape dark brown; pecten present.
|
||||
<location id="8EB2B5A0033FA04C0517234AFA2BFAD2" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C0517234AFA2BFAD2" box="[1387,1471,1355,1377]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Thorax" pageId="11" pageNumber="581">Thorax</location>
|
||||
, ‘burrito’ case types, as it is sister to another clade tegula, metascutellum, and abdomen dark brown. that makes this case type. Fore-, mid-, and hindleg coxa dark brown; femur, tibia, and tarsomeres dark brown with each segment
|
||||
<emphasis id="B9193F69033FA04C00BA23C4FEC3FA69" box="[198,343,1477,1498]" italics="true" pageId="11" pageNumber="581">
|
||||
<location id="8EB2B5A0033FA04C00BA23C4FEC6FA69" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C00BA23C4FEC6FA69" box="[198,338,1477,1498]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Distribution" pageId="11" pageNumber="581">Distribution</location>
|
||||
:
|
||||
</emphasis>
|
||||
<location id="8EB2B5A0033FA04C011A23C4FE23FA69" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C011A23C4FE23FA69" box="[358,439,1477,1498]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Known" pageId="11" pageNumber="581">Known</location>
|
||||
only from Kahoʻolawe
|
||||
<collectingCountry id="F37AA3EB033FA04C02A223C4FCBCFA68" box="[734,808,1477,1499]" name="Iceland" pageId="11" pageNumber="581">Island</collectingCountry>
|
||||
, terminating with a light brown ring.
|
||||
<location id="8EB2B5A0033FA04C052423C4FE84FA49" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C052423C4FE84FA49" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Forewing Hawai" pageId="11" pageNumber="581">Forewing Hawai</location>
|
||||
i, where it is presumed to be endemic. ground colour brown, with several bands of darker scales extending from costal margin to anal margin,
|
||||
<emphasis id="B9193F69033FA04C00BA2020FEA5F985" box="[198,305,1569,1590]" italics="true" pageId="11" pageNumber="581">
|
||||
<location id="8EB2B5A0033FA04C00BA2020FEBFF985" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C00BA2020FEBFF985" box="[198,299,1569,1590]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Remarks" pageId="11" pageNumber="581">Remarks</location>
|
||||
:
|
||||
</emphasis>
|
||||
<location id="8EB2B5A0033FA04C013C2020FEE6F984" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C013C2020FEE6F984" box="[320,370,1569,1591]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="This" pageId="11" pageNumber="581">This</location>
|
||||
species is one of several independent alternating with bands of lighter scales; fringe lineages with the ‘burrito’ case type that inhabit brown.
|
||||
<location id="8EB2B5A0033FA04C03BF203EFBA0F9E6" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C03BF203EFBA0F9E6" box="[963,1076,1599,1621]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Hindwing" pageId="11" pageNumber="581">Hindwing</location>
|
||||
and fringe uniformly brown. Kahoʻolawe.
|
||||
<location id="8EB2B5A0033FA04C0115205FFE16F9C0" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C0115205FFE16F9C0" box="[361,386,1630,1651]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="In" pageId="11" pageNumber="581">In</location>
|
||||
the phylogeny,
|
||||
<taxonomicName id="4C6D98F8033FA04C0215205FFC96F9C0" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[617,770,1630,1651]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="581" phylum="Arthropoda" rank="species" species="nohomeha">
|
||||
<emphasis id="B9193F69033FA04C0215205FFC96F9C0" box="[617,770,1630,1651]" italics="true" pageId="11" pageNumber="581">H. nohomeha</emphasis>
|
||||
</taxonomicName>
|
||||
is
|
||||
<location id="8EB2B5A0033FA04C0321205FFC5EF9C7" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C0321205FFC5EF9C7" box="[861,970,1630,1652]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Subcostal" pageId="11" pageNumber="581">Subcostal</location>
|
||||
brush present.
|
||||
<location id="8EB2B5A0033FA04C049A205FFAF7F9C7" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C049A205FFAF7F9C7" box="[1254,1379,1630,1652]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Sclerotized" pageId="11" pageNumber="581">Sclerotized</location>
|
||||
hook separated from its closest known relatives by a long (pseuduncus
|
||||
<emphasis id="B9193F69033FA04C047F207CFBD6F921" box="[1027,1090,1661,1682]" italics="true" pageId="11" pageNumber="581">sensu</emphasis>
|
||||
Zimmerman, 1978) elongate, branch (
|
||||
<figureCitation id="1356FFFE033FA04C0152209AFEE6F903" box="[302,370,1691,1713]" captionStart="Figure 2" captionStartId="4.[222,301,257,276]" captionText="Figure 2. Bayesian phylogeny of Kahoʻolawe Hyposmocoma, with selected Hyposmocoma from other islands, and of various case types, included. See Material and methods for details about tree construction. Branches coloured using the same case type colour scheme as in Haines et al. (2014). Within the ingroup, posterior probabilities are only labelled when values were <1, and are not labelled for relationships amongst individuals within species. Scale bar indicates substitutions per site. New taxa from Kahoʻolawe have extraction codes in red." pageId="11" pageNumber="581">Fig. 2</figureCitation>
|
||||
), implying either a long history of slender, straight; distinct sclerotized ring on segment isolation, or extinction of related taxa. VII.
|
||||
<location id="8EB2B5A0033FA04C03E820B8FC67F97C" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C03E820B8FC67F97C" box="[916,1011,1721,1743]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Females" pageId="11" pageNumber="581">Females</location>
|
||||
similar to males except frenulum with three acanthae.
|
||||
<taxonomicName id="4C6D98F8033FA04C00B72107FE43F8A9" authorityName="Doorenweerd & Austin & Rubinoff & n & n & n & n & n" authorityYear="2023" box="[203,471,1798,1819]" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="582" phylum="Arthropoda" rank="species" species="oolea">
|
||||
<emphasis id="B9193F69033FA04C00B72107FE43F8A9" bold="true" box="[203,471,1798,1819]" italics="true" pageId="11" pageNumber="581">
|
||||
<smallCapsWord id="8D3475A7033FA04C00B72107FE15F8A9" baselines="1814,1814" box="[203,385,1798,1819]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Hyposmocoma" pageId="11" pageNumber="581">HYPOSMOCOMA</smallCapsWord>
|
||||
<smallCapsWord id="8D3475A7033FA04C01F72108FE43F8A9" baselines="1814" box="[395,471,1801,1818]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="oolea" pageId="11" pageNumber="581">OOLEA</smallCapsWord>
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
<smallCapsWord id="8D3475A7033FA04C019C2102FDC2F8A9" baselines="1813,1814" box="[480,598,1795,1819]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Medeiros" pageId="11" pageNumber="581">MEDEIROS</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3475A7033FA04C02142102FD2AF8A9" baselines="1813,1814" box="[616,702,1795,1819]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Haines" pageId="11" pageNumber="581">HAINES</smallCapsWord>
|
||||
& RUBI-
|
||||
<emphasis id="B9193F69033FA04C030920F7FBB6F8B8" box="[885,1058,1782,1803]" italics="true" pageId="11" pageNumber="581">
|
||||
<location id="8EB2B5A0033FA04C030920F7FC39F8B8" LSID="urn:lsid:plazi:treatment:03C4526D033FA04C029626BCFA57F834:8EB2B5A0033FA04C030920F7FC39F8B8" box="[885,941,1782,1803]" country="Iceland" county="Material" latitude="0.38333333" longLatPrecision="15" longitude="-20.52256" municipality="Valvae" name="Male" pageId="11" pageNumber="581">Male</location>
|
||||
genitalia:
|
||||
</emphasis>
|
||||
(
|
||||
<emphasis id="B9193F69033FA04C044820F6FBDCF8BF" box="[1076,1096,1783,1804]" italics="true" pageId="11" pageNumber="581">N</emphasis>
|
||||
= 1) (
|
||||
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asymmetrical, left valva robust and with four heavy, long,
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(
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3G
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,
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<figureCitation id="1356FFFE033FA04C02282121FDE3F889" box="[596,631,1824,1850]" captionStart="Figure 4" captionStartId="6.[222,301,1472,1491]" captionTargetBox="[446,1270,251,1436]" captionTargetId="figure-263@6.[446,1271,251,1436]" captionTargetPageId="6" captionText="Figure 4. Male genitalia of Hyposmocoma. A, Hyposmocoma anoai sp. nov., slide WPH055-13; B, Hyposmocoma hooilo sp. nov., slide WPH040-13; C, Hyposmocoma kamaula sp. nov., slide WPH095-13; D, Hyposmocoma kanaloa sp. nov., slide WPH122-13; E, Hyposmocoma mahoepo sp. nov., slide 11A02; F, Hyposmocoma nohomeha sp. nov., slide WPH084-13; G, Hyposmocoma oolea sp. nov., slide WPH047-13; H, Hyposmocoma pahanalo sp. nov., slide 08A42; I, Hyposmocoma waauhi sp. nov., slide WPH074-13. Abbreviations: la, left anellus lobe; lv, left valva; ra, right anellus lobe; rv, right valva; s, setae." pageId="11" pageNumber="581">4G</figureCitation>
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,
|
||||
<figureCitation id="1356FFFE033FA04C02F42121FD50F889" box="[648,708,1824,1851]" captionStart="Figure 5" captionStartId="7.[198,277,1241,1260]" captionTargetBox="[428,1254,253,1205]" captionTargetId="figure-325@7.[427,1255,251,1205]" captionTargetPageId="7" captionText="Figure 5. Female genitalia and larval cases of Hyposmocoma. A, Hyposmocoma hooilo sp. nov., slide WPH042- 13; B, Hyposmocoma kamaula sp. nov., slide WPH050-13; C, Hyposmocoma kanaloa sp. nov., slide WPH123- 13; D, Hyposmocoma mahoepo sp. nov., slide WPH053-13; E, Hyposmocoma nohomeha sp. nov., slide WPH076-13; F, Hyposmocoma oolea sp. nov., slide WPH048-13; G, Hyposmocoma pahanalo sp. nov., slide WPH044-13; H, Hyposmocoma kanaloa sp. nov., larval case, dorsal view, length = 4 mm; I, Hyposmocoma oolea sp. nov., larval case, dorsal view, length = 6 mm. Abbreviations: a, anterior apophysis; cb, corpus bursa; db, ductus bursa; o, ostium-bearing process; p, posterior apophysis; pa, papillae anales." pageId="11" pageNumber="581">5F, I</figureCitation>
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) sclerotized spur-like setae projecting dorsally near apex of both valvae; one additional heavy seta pro-
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examined:
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: HI: Kahoʻolawe: jecting dorsally from an acute dorsal lobe located
|
||||
</materialsCitation>
|
||||
</paragraph>
|
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</subSubSection>
|
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</treatment>
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|
||||
371
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<taxonomicName id="4C62A3F2FF823852387FFB619417FB5D" ID-CoL="4K83W" authority="BLANCHARD, 1850" authorityName="Blanchard" authorityYear="1850" box="[835,1393,1233,1260]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="fasciatipennis">
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, 1850
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(
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1A
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<emphasis id="B9160463FF8238523F1AFB449552FABF" box="[1062,1076,1267,1289]" italics="true" pageId="6" pageNumber="327">–</emphasis>
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L
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</figureCitation>
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,
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<figureCitation id="1359C4F4FF8238523F6BFB5895FDFABC" box="[1111,1179,1263,1290]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="6" pageNumber="327">
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2A
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<emphasis id="B9160463FF8238523F44FB4495E0FABF" box="[1144,1158,1267,1289]" italics="true" pageId="6" pageNumber="327">–</emphasis>
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D
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,
|
||||
<figureCitation id="1359C4F4FF8238523F90FB5895ABFABC" box="[1196,1229,1263,1290]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="6" pageNumber="327">5A</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF823852381BFA9695D3FA58" pageId="6" pageNumber="327" type="reference_group">
|
||||
<paragraph id="8BDDD871FF823852381BFA969512FAE3" blockId="6.[807,1421,1313,1519]" pageId="6" pageNumber="327">
|
||||
<treatmentCitationGroup id="AB72FF5FFF823852381BFA969512FAE3" pageId="6" pageNumber="327">
|
||||
<treatmentCitation id="0AC3FE60FF823852381BFA969421FA80" author="Blanchard ME" box="[807,1351,1313,1335]" page="83" pageId="6" pageNumber="327" year="1850">
|
||||
<taxonomicName id="4C62A3F2FF823852381BFA969421FA80" authority="Blanchard, 1850: 83" authorityName="Blanchard" authorityPageNumber="83" authorityYear="1850" box="[807,1351,1313,1335]" class="Insecta" family="Scarabaeidae" genus="Omaloplia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="fasciatipennis">
|
||||
<emphasis id="B9160463FF823852381BFA96952CFA80" box="[807,1098,1313,1334]" italics="true" pageId="6" pageNumber="327">Omaloplia fasciatipennis</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF8238523F6BFA969421FA80" box="[1111,1351,1313,1335]" location="Good Hope" pageId="6" pageNumber="327" specimenCount="1" stateProvince="Western Cape">
|
||||
<bibRefCitation id="EFF3A580FF8238523F6BFA969421FA80" author="Blanchard ME" box="[1111,1351,1313,1335]" pageId="6" pageNumber="327" pagination="1 - 128" refId="ref23143" refString="Blanchard ME. 1850. Catalogue de la collection Entomologique. Classes des Insectes. Ordre des Coleopteres. part.: Melolonthidae. Paris: Tom I. Museum National d'Histoire Naturelle, 1-128." type="book chapter" year="1850">Blanchard, 1850: 83</bibRefCitation>
|
||||
</materialsCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
[
|
||||
<typeStatus id="54D966D3FF8238523E66FA9594EBFA81" box="[1370,1421,1314,1335]" pageId="6" pageNumber="327">type</typeStatus>
|
||||
locality:
|
||||
<collectingRegion id="49A61693FF82385238B4FAF792A4FAE3" box="[904,962,1344,1365]" country="South Africa" name="Western Cape" pageId="6" pageNumber="327">Cape</collectingRegion>
|
||||
of
|
||||
<location id="8EBD8EAAFF82385238DBFAF7950FFAE3" LSID="urn:lsid:plazi:treatment:03CB6967FF82385C387FFB619019FB26:8EBD8EAAFF82385238DBFAF7950FFAE3" box="[999,1129,1344,1366]" name="Good Hope" pageId="6" pageNumber="327" stateProvince="Western Cape">Good Hope</location>
|
||||
].
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF823852381BFAE995BCFA24" blockId="6.[807,1421,1313,1519]" pageId="6" pageNumber="327">
|
||||
<treatmentCitationGroup id="AB72FF5FFF823852381BFAE995BCFA24" pageId="6" pageNumber="327">
|
||||
<taxonomicName id="4C62A3F2FF823852381BFAE99533FAC5" ID-CoL="4K83W" authorityName="Blanchard" authorityYear="1850" box="[807,1109,1374,1395]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="fasciatipennis">
|
||||
<emphasis id="B9160463FF823852381BFAE99533FAC5" box="[807,1109,1374,1395]" italics="true" pageId="6" pageNumber="327">Pleophylla fasciatipennis</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<treatmentCitation id="0AC3FE60FF8238523F49FAE994EFFAC2" author="Schaufuss LW" box="[1141,1417,1374,1396]" page="231" pageId="6" pageNumber="327" year="1871">
|
||||
<bibRefCitation id="EFF3A580FF8238523F49FAE994EFFAC2" author="Schaufuss LW" box="[1141,1417,1374,1396]" pageId="6" pageNumber="327" pagination="231 - 233" refId="ref24095" refString="Schaufuss LW. 1871. Die Arten der Gattung Pleophylla Er. Nunquam Otiosus I 6: 231-233." type="journal article" year="1871">Schaufuss, 1871: 231</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
; Brenske 1900: 84; Peringuey, 1904: 8.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF823852381BFA2C95D3FA58" blockId="6.[807,1421,1313,1519]" pageId="6" pageNumber="327">
|
||||
<treatmentCitationGroup id="AB72FF5FFF823852381BFA2C95D3FA58" pageId="6" pageNumber="327">
|
||||
<treatmentCitation id="0AC3FE60FF823852381BFA2C9421FA07" author="Boheman CH" box="[807,1351,1435,1457]" page="124" pageId="6" pageNumber="327" year="1857">
|
||||
<taxonomicName id="4C62A3F2FF823852381BFA2C9421FA07" ID-CoL="4K844" authority="Boheman, 1857: 124" authorityName="Boheman" authorityPageNumber="124" authorityYear="1857" box="[807,1351,1435,1457]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="maculipennis">
|
||||
<emphasis id="B9160463FF823852381BFA2C9523FA06" box="[807,1093,1435,1456]" italics="true" pageId="6" pageNumber="327">Pleophylla maculipennis</emphasis>
|
||||
<bibRefCitation id="EFF3A580FF8238523F6FFA2C9421FA07" author="Boheman CH" box="[1107,1351,1435,1457]" pageId="6" pageNumber="327" refId="ref23181" refString="Boheman CH. 1857. Insecta Caffrariae. Annis 1838 - 1845. Part II. Holmiae, 1 - 395." type="book" year="1857">Boheman, 1857: 124</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
[
|
||||
<typeStatus id="54D966D3FF8238523E66FA2B94EBFA07" box="[1370,1421,1436,1457]" pageId="6" pageNumber="327">type</typeStatus>
|
||||
locality: Gariep]; syn. by Peringuey, 1904: 8;
|
||||
<treatmentCitation id="0AC3FE60FF8238523E03FA0D92B2FA58" author="Schaufuss LW" page="232" pageId="6" pageNumber="327" year="1871">
|
||||
<bibRefCitation id="EFF3A580FF8238523E03FA0D92B2FA58" author="Schaufuss LW" pageId="6" pageNumber="327" pagination="231 - 233" refId="ref24095" refString="Schaufuss LW. 1871. Die Arten der Gattung Pleophylla Er. Nunquam Otiosus I 6: 231-233." type="journal article" year="1871">Schaufuss, 1871: 232</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
; Brenske, 1900: 83.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF82385C381BF9919019FB26" lastPageId="8" lastPageNumber="329" pageId="6" pageNumber="327" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF823853381BF99194C2F8DF" blockId="6.[807,1422,1573,1870]" lastBlockId="7.[829,1444,1814,1897]" lastPageId="7" lastPageNumber="328" pageId="6" pageNumber="327">
|
||||
<emphasis id="B9160463FF823852381BF9919558F98C" box="[807,1086,1573,1595]" italics="true" pageId="6" pageNumber="327">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF8238523F71F99294ECF902" collectionCode="MNHN" pageId="6" pageNumber="327" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
|
||||
<typeStatus id="54D966D3FF8238523F71F99295DDF98C" box="[1101,1211,1573,1594]" pageId="6" pageNumber="327" type="lectotype">Lectotype</typeStatus>
|
||||
(here designated):
|
||||
<specimenCount id="9D6413F8FF823852381BF9F19251F9EE" box="[807,823,1606,1624]" count="1" pageId="6" pageNumber="327" type="male">♂</specimenCount>
|
||||
“
|
||||
<taxonomicName id="4C62A3F2FF8238523875F9F39446F9EF" authority="Cat. Mus. Cap de B. Esp." box="[841,1312,1603,1625]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="fasciatipennis">P. fasciatipennis Cat. Mus. Cap de B. Esp.</taxonomicName>
|
||||
/Museum Paris Afrique Hottentot Delalande/
|
||||
<taxonomicName id="4C62A3F2FF8238523E29F9D592A3F920" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="fasciatipennis">
|
||||
<emphasis id="B9160463FF8238523E29F9D592A3F920" italics="true" pageId="6" pageNumber="327">Pleophylla fasciatipennis</emphasis>
|
||||
</taxonomicName>
|
||||
Type Blanch. [unknown handwriting]/ Afriq. Hottentot Delalande/IMG0032
|
||||
<emphasis id="B9160463FF8238523FC5F9289465F902" bold="true" box="[1273,1283,1695,1716]" italics="true" pageId="6" pageNumber="327">″</emphasis>
|
||||
(
|
||||
<collectionCode id="ED7340B4FF8238523E16F92894E5F902" box="[1322,1411,1695,1716]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="327">MNHN</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B0AD22CFF823852381BF90992EAF8A6" collectionCode="MNHN" location="Museum Paris Cap de Bonne Esperance" pageId="6" pageNumber="327" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
|
||||
<typeStatus id="54D966D3FF823852381BF90992A9F962" box="[807,975,1726,1748]" pageId="6" pageNumber="327" type="paralectotype">Paralectotypes</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF82385238DDF909956DF964" box="[993,1035,1726,1747]" count="1" pageId="6" pageNumber="327" type="female">1 ♀</specimenCount>
|
||||
“
|
||||
<location id="8EBD8EAAFF8238523F18F90992FFF947" LSID="urn:lsid:plazi:treatment:03CB6967FF82385C387FFB619019FB26:8EBD8EAAFF8238523F18F90992FFF947" name="Museum Paris Cap de Bonne Esperance" pageId="6" pageNumber="327">Museum Paris Cap de Bonne Esperance</location>
|
||||
/
|
||||
<taxonomicName id="4C62A3F2FF823852389DF96B95AFF947" box="[929,1225,1756,1777]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="fasciatipennis">
|
||||
<emphasis id="B9160463FF823852389DF96B95AFF947" box="[929,1225,1756,1777]" italics="true" pageId="6" pageNumber="327">Pleophylla fasciatipennis</emphasis>
|
||||
</taxonomicName>
|
||||
Type Blanch.” (
|
||||
<collectionCode id="ED7340B4FF8238523812F94C92E3F8A6" box="[814,901,1787,1808]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="327">MNHN</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="3B0AD22CFF8238523899F94C92EAF8FB" collectionCode="MNHN" location="Museum Paris Afrique Hottentot Delalande" pageId="6" pageNumber="327" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
|
||||
<specimenCount id="9D6413F8FF8238523899F94C92B2F8A6" box="[933,980,1787,1808]" count="1" pageId="6" pageNumber="327" type="female">1 ♀</specimenCount>
|
||||
“
|
||||
<location id="8EBD8EAAFF82385238CEF94C92F1F899" LSID="urn:lsid:plazi:treatment:03CB6967FF82385C387FFB619019FB26:8EBD8EAAFF82385238CEF94C92F1F899" name="Museum Paris Afrique Hottentot Delalande" pageId="6" pageNumber="327">Museum Paris Afrique Hottentot Delalande</location>
|
||||
/
|
||||
<taxonomicName id="4C62A3F2FF82385238A2F8AE95A1F898" box="[926,1223,1817,1838]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="fasciatipennis">
|
||||
<emphasis id="B9160463FF82385238A2F8AE95A1F898" box="[926,1223,1817,1838]" italics="true" pageId="6" pageNumber="327">Pleophylla fasciatipennis</emphasis>
|
||||
</taxonomicName>
|
||||
Type Blanch.” (
|
||||
<collectionCode id="ED7340B4FF8238523812F88F92E3F8FB" box="[814,901,1848,1869]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="327">MNHN</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B0AD22CFF82385338ABF88F90B8F8DF" collectionCode="NHRS" collectorName="J. Wahlb." lastPageId="7" lastPageNumber="328" location="Stockholm" pageId="6" pageNumber="327" specimenCount="1" specimenCount-male="1" stateProvince="Stockholm" typeStatus="lectotype">
|
||||
<typeStatus id="54D966D3FF82385238ABF88F9562F8FB" box="[919,1028,1848,1869]" pageId="6" pageNumber="327" type="lectotype">Lectotype</typeStatus>
|
||||
(
|
||||
<taxonomicName id="4C62A3F2FF8238523F2EF88F95CEF8FB" authorityName="Boheman" authorityYear="1857" box="[1042,1192,1848,1869]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="maculipennis">
|
||||
<emphasis id="B9160463FF8238523F2EF88F95CEF8FB" box="[1042,1192,1848,1869]" italics="true" pageId="6" pageNumber="327">maculipennis</emphasis>
|
||||
</taxonomicName>
|
||||
, here designated):
|
||||
<specimenCount id="9D6413F8FF8238523E41F88C94EBF8FB" box="[1405,1421,1851,1869]" count="1" pageId="6" pageNumber="327" type="male">♂</specimenCount>
|
||||
“Caffraria./
|
||||
<collectorName id="2697BDA7FF8338533A15F8A190F5F89A" box="[297,403,1814,1836]" pageId="7" pageNumber="328">J. Wahlb.</collectorName>
|
||||
/Type./Typus [red printed label]/ 3674E91/Naturhistoriska Riksmuseet
|
||||
<collectingRegion id="49A61693FF83385339ABF882926BF8FD" box="[663,781,1845,1867]" country="Sweden" name="Stockholm" pageId="7" pageNumber="328">Stockholm</collectingRegion>
|
||||
Loan no. 1743/07” (
|
||||
<collectionCode id="ED7340B4FF8338533AB5F8E390B0F8DF" box="[393,470,1876,1897]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="7" pageNumber="328">NHRS</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B0AD22CFF8338533AD0F8E394C2F8DF" collectionCode="NHRS" collectorName="Cap. B. Spei." location="Loan" municipality="Naturhistoriska Riksmuseet" pageId="7" pageNumber="328" specimenCount="2" specimenCount-male="2" stateProvince="Stockholm" typeStatus="paralectotype">
|
||||
<typeStatus id="54D966D3FF8338533AD0F8E393F7F8DC" box="[492,657,1876,1898]" pageId="7" pageNumber="328" type="paralectotype">Paralectotypes</typeStatus>
|
||||
(
|
||||
<taxonomicName id="4C62A3F2FF833853399FF8E39383F8DF" authorityName="Boheman" authorityYear="1857" box="[675,741,1876,1897]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B9160463FF833853399FF8E39383F8DF" box="[675,741,1876,1897]" italics="true" pageId="7" pageNumber="328">pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
):
|
||||
<specimenCount id="9D6413F8FF83385339C3F8E3922BF89D" box="[767,845,1817,1897]" count="1" pageId="7" pageNumber="328" type="male">1 ♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF83385338B1F8A09510F89A" box="[909,1142,1814,1836]" pageId="7" pageNumber="328">Cap. B. Spei.</collectorName>
|
||||
/Victorin./3676 E91 + /
|
||||
<collectingMunicipality id="6BB9420BFF8338533801F88295EAF8FD" box="[829,1164,1845,1867]" pageId="7" pageNumber="328">Naturhistoriska Riksmuseet</collectingMunicipality>
|
||||
<collectingRegion id="49A61693FF8338533F98F882947CF8FD" box="[1188,1306,1845,1867]" country="Sweden" name="Stockholm" pageId="7" pageNumber="328">Stockholm</collectingRegion>
|
||||
<location id="8EBD8EAAFF8338533E0DF882940CF8FC" LSID="urn:lsid:plazi:treatment:03CB6967FF82385C387FFB619019FB26:8EBD8EAAFF8338533E0DF882940CF8FC" box="[1329,1386,1845,1866]" municipality="Naturhistoriska Riksmuseet" name="Loan" pageId="7" pageNumber="328" stateProvince="Stockholm">Loan</location>
|
||||
no. 1745/07 “(
|
||||
<collectionCode id="ED7340B4FF8338533888F8E39566F8DF" box="[948,1024,1876,1897]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="7" pageNumber="328">NHRS</collectionCode>
|
||||
),
|
||||
<specimenCount id="9D6413F8FF8338533F26F8E39521F8DE" box="[1050,1095,1876,1897]" count="1" pageId="7" pageNumber="328" type="male">1 ♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF8338533F5CF8E3959BF8DF" box="[1120,1277,1875,1897]" pageId="7" pageNumber="328">Cap. B. Spei.</collectorName>
|
||||
/Victorin./3675
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<caption id="DF1D88F9FF8338533B9AF9819569F96D" ID-DOI="http://doi.org/10.5281/zenodo.14807091" ID-Zenodo-Dep="14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="7" pageNumber="328" startId="7.[166,245,1590,1609]" targetBox="[178,1431,206,1549]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="8BDDD871FF8338533B9AF9819569F96D" blockId="7.[166,1444,1590,1756]" pageId="7" pageNumber="328">
|
||||
<emphasis id="B9160463FF8338533B9AF9819076F9FF" bold="true" box="[166,272,1590,1609]" pageId="7" pageNumber="328">Figure 3.</emphasis>
|
||||
A
|
||||
<emphasis id="B9160463FF8338533A11F98E905EF9FF" box="[301,312,1593,1609]" italics="true" pageId="7" pageNumber="328">–</emphasis>
|
||||
D,
|
||||
<taxonomicName id="4C62A3F2FF8338533A64F98193DDF9FF" authority="Burmeister" authorityName="Burmeister" authorityYear="1855" box="[344,699,1590,1609]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="ferruginea">
|
||||
<emphasis id="B9160463FF8338533A64F981935DF9FF" box="[344,571,1590,1609]" italics="true" pageId="7" pageNumber="328">Pleophylla ferruginea</emphasis>
|
||||
Burmeister
|
||||
</taxonomicName>
|
||||
(RSA, Tygerkloof); E
|
||||
<emphasis id="B9160463FF83385338A2F98E92CFF9FF" box="[926,937,1593,1609]" italics="true" pageId="7" pageNumber="328">–</emphasis>
|
||||
H,
|
||||
<emphasis id="B9160463FF83385338F7F9819476F9FF" bold="true" box="[971,1296,1590,1609]" pageId="7" pageNumber="328">
|
||||
<taxonomicName id="4C62A3F2FF83385338F7F98195CBF9FF" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[971,1197,1590,1609]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="murzini" status="sp. nov.">
|
||||
<emphasis id="B9160463FF83385338F7F98195CBF9FF" bold="true" box="[971,1197,1590,1609]" italics="true" pageId="7" pageNumber="328">Pleophylla murzini</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8338533F8AF9819476F9FF" box="[1206,1296,1590,1609]" pageId="7" pageNumber="328" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype); I
|
||||
<emphasis id="B9160463FF8338533EA4F98E94C5F9FF" box="[1432,1443,1593,1609]" italics="true" pageId="7" pageNumber="328">–</emphasis>
|
||||
L,
|
||||
<taxonomicName id="4C62A3F2FF8338533BFAF9E49353F9D0" authority="Burmeister" authorityName="Burmeister" authorityYear="1855" box="[198,565,1619,1638]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="navicularis">
|
||||
<emphasis id="B9160463FF8338533BFAF9E490D2F9D0" box="[198,436,1619,1638]" italics="true" pageId="7" pageNumber="328">Pleophylla navicularis</emphasis>
|
||||
Burmeister
|
||||
</taxonomicName>
|
||||
(RSA, Weza); M
|
||||
<emphasis id="B9160463FF83385339D0F9E19391F9D0" box="[748,759,1622,1638]" italics="true" pageId="7" pageNumber="328">–</emphasis>
|
||||
P,
|
||||
<emphasis id="B9160463FF833853382BF9E4950FF9D0" bold="true" box="[791,1129,1619,1638]" pageId="7" pageNumber="328">
|
||||
<taxonomicName id="4C62A3F2FF833853382BF9E49564F9D0" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[791,1026,1619,1638]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="silvatica" status="sp. nov.">
|
||||
<emphasis id="B9160463FF833853382BF9E49564F9D0" bold="true" box="[791,1026,1619,1638]" italics="true" pageId="7" pageNumber="328">Pleophylla silvatica</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8338533F30F9E4950FF9D0" box="[1036,1129,1619,1638]" pageId="7" pageNumber="328" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype); Q
|
||||
<emphasis id="B9160463FF8338533FC1F9E1946EF9D0" box="[1277,1288,1622,1638]" italics="true" pageId="7" pageNumber="328">–</emphasis>
|
||||
T,
|
||||
<emphasis id="B9160463FF8338533E1BF9E490CEF932" bold="true" pageId="7" pageNumber="328">
|
||||
<taxonomicName id="4C62A3F2FF8338533E1BF9E49027F935" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="transkeiensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8338533E1BF9E49027F935" bold="true" italics="true" pageId="7" pageNumber="328">Pleophylla transkeiensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8338533A70F9C690CEF932" box="[332,424,1649,1668]" pageId="7" pageNumber="328" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype); U
|
||||
<emphasis id="B9160463FF8338533902F9C4932FF935" box="[574,585,1651,1667]" italics="true" pageId="7" pageNumber="328">–</emphasis>
|
||||
X,
|
||||
<emphasis id="B9160463FF8338533956F9C792C0F932" bold="true" box="[618,934,1648,1668]" pageId="7" pageNumber="328">
|
||||
<taxonomicName id="4C62A3F2FF8338533956F9C79226F935" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[618,832,1648,1667]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="ruthae" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8338533956F9C79226F935" bold="true" box="[618,832,1648,1667]" italics="true" pageId="7" pageNumber="328">Pleophylla ruthae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8338533876F9C692C0F932" box="[842,934,1649,1668]" pageId="7" pageNumber="328" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype); Y
|
||||
<emphasis id="B9160463FF8338533F05F9C49522F935" box="[1081,1092,1651,1667]" italics="true" pageId="7" pageNumber="328">–</emphasis>
|
||||
Ab,
|
||||
<emphasis id="B9160463FF8338533F4EF9C791BEF917" bold="true" pageId="7" pageNumber="328">
|
||||
<taxonomicName id="4C62A3F2FF8338533F4EF9C7941FF935" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1138,1401,1648,1667]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="warnockae" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8338533F4EF9C7941FF935" bold="true" box="[1138,1401,1648,1667]" italics="true" pageId="7" pageNumber="328">Pleophylla warnockae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8338533EBFF9C691BEF917" pageId="7" pageNumber="328" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype); Ac
|
||||
<emphasis id="B9160463FF8338533A48F9279019F916" box="[372,383,1680,1696]" italics="true" pageId="7" pageNumber="328">–</emphasis>
|
||||
Af,
|
||||
<emphasis id="B9160463FF8338533A9BF93A9265F917" bold="true" box="[423,771,1677,1697]" pageId="7" pageNumber="328">
|
||||
<taxonomicName id="4C62A3F2FF8338533A9BF93A93FBF916" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[423,669,1677,1696]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="harrisoni" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8338533A9BF93A93FBF916" bold="true" box="[423,669,1677,1696]" italics="true" pageId="7" pageNumber="328">Pleophylla harrisoni</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF833853399BF9399265F917" box="[679,771,1678,1697]" pageId="7" pageNumber="328" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BDDD871FF8C385C3BACFF7090E3FF4C" blockId="8.[144,758,199,251]" pageId="8" pageNumber="329">
|
||||
E91 + /Naturhistoriska Riksmuseet
|
||||
<collectingRegion id="49A61693FF8C385C3904FF7093C8FF6B" box="[568,686,199,221]" country="Sweden" name="Stockholm" pageId="8" pageNumber="329">Stockholm</collectingRegion>
|
||||
Loan no. 1744/07 “(
|
||||
<collectionCode id="ED7340B4FF8C385C3A11FF52901FFF4C" box="[301,377,229,250]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="8" pageNumber="329">NHRS</collectionCode>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8C385C3BACFE9593A0FE81" blockId="8.[144,710,290,312]" box="[144,710,290,312]" pageId="8" pageNumber="329">
|
||||
<emphasis id="B9160463FF8C385C3BACFE959089FE81" box="[144,495,290,311]" italics="true" pageId="8" pageNumber="329">Additional material examined:</emphasis>
|
||||
See Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8C385C3BACFEE89390FC41" blockId="8.[144,759,351,1015]" pageId="8" pageNumber="329">
|
||||
<emphasis id="B9160463FF8C385C3BACFEE89057FEC2" box="[144,305,351,372]" italics="true" pageId="8" pageNumber="329">Redescription:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8C385C3AD9FEE8935BFEC3" box="[485,573,351,373]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="8" pageNumber="329" unit="mm" value="7.9">7.9 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF8C385C3BACFEC9918EFE25" box="[144,232,382,404]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="8" pageNumber="329" unit="mm" value="5.9">5.9 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF8C385C3AE5FEC99354FE25" box="[473,562,382,403]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="8" pageNumber="329" unit="mm" value="4.2">4.2 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes 1.5 times as wide as long; ratio of eye diameter/interocular width: 0.5. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.21. Metatibia, ratio of width/length: 1/3.59. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF8C385C3BACFC569062FC40" box="[144,260,993,1015]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="8" pageNumber="329">
|
||||
Fig. 2A
|
||||
<emphasis id="B9160463FF8C385C3BDFFC539189FC40" box="[227,239,996,1014]" italics="true" pageId="8" pageNumber="329">–</emphasis>
|
||||
C
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF8C385C3A4FFC5690ADFC40" box="[371,459,993,1015]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="8" pageNumber="329">Fig. 2D</figureCitation>
|
||||
. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF8C385C39A0FC569395FC41" box="[668,755,993,1015]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="8" pageNumber="329">Fig. 5A</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8C385C3BACFBA99019FB26" blockId="8.[144,759,1054,1168]" pageId="8" pageNumber="329">
|
||||
<emphasis id="B9160463FF8C385C3BACFBA9919DFB85" box="[144,251,1054,1075]" italics="true" pageId="8" pageNumber="329">Remarks:</emphasis>
|
||||
In previous publications we referred to this species as ‘sp2’ (for
|
||||
<taxonomicName id="4C62A3F2FF8C385C3AF6FB899348FBE4" authorityName="Boheman" authorityYear="1857" box="[458,558,1085,1107]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B9160463FF8C385C3AF6FB899348FBE4" box="[458,558,1085,1107]" italics="true" pageId="8" pageNumber="329">P. pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
) and ‘sp12’ (for
|
||||
<taxonomicName id="4C62A3F2FF8C385C3BACFBEB902FFBC6" authorityName="Boheman" authorityYear="1857" box="[144,329,1115,1137]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="maculipennis">
|
||||
<emphasis id="B9160463FF8C385C3BACFBEB902FFBC6" box="[144,329,1115,1137]" italics="true" pageId="8" pageNumber="329">P. maculipennis</emphasis>
|
||||
</taxonomicName>
|
||||
) (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF8C385C3973FBEC9394FBC7" author="Ahrens D" box="[591,754,1115,1137]" pageId="8" pageNumber="329" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFF3A580FF8C385C3BACFBCD9015FB26" author="Eberle J & Warnock RCM & Ahrens D" box="[144,371,1146,1168]" pageId="8" pageNumber="329" refId="ref23428" refString="Eberle J, Warnock RCM, Ahrens D. 2016 b. Bayesian species delimitation in Pleophylla chafers (Coleoptera) - the importance of prior choice and morphology. BMC Evolutionary Biology 16: 94." type="journal volume" year="2016">
|
||||
Eberle
|
||||
<emphasis id="B9160463FF8C385C3BDEFBCC907AFB39" box="[226,284,1146,1168]" italics="true" pageId="8" pageNumber="329">et al.</emphasis>
|
||||
, 2016b
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
204
data/03/CB/69/03CB6967FF8838583870FCE594EBF8DB.xml
Normal file
204
data/03/CB/69/03CB6967FF8838583870FCE594EBF8DB.xml
Normal file
|
|
@ -0,0 +1,204 @@
|
|||
<document id="593941E03F793D43D396E784C540639C" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia & Ahrens, Dirk" docDate="2017" docId="03CB6967FF8838583870FCE594EBF8DB" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla congoensis AHRENS, BECKETT, EBERLE & FABRIZI 2017, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="333" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="333" updateTime="1738712560284" updateUser="ExternalLinkService">
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<mods:mods id="4F7BA1A75C26AE49969382D4FF2AC031" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F63F073A73577FE89D941D640DA2881F">
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<mods:title id="AA5329CDB89ED797D22EBCBD72151AD9">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
|
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|
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<mods:role id="FA491B977C52FC03179723F38A0F7A54">
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<mods:roleTerm id="430A7E6D8C7B8B0F54BBF77DDF094204">Author</mods:roleTerm>
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|
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<mods:namePart id="A826B4373AA8A03FCFE387947C811C7A">Eberle, Jonas</mods:namePart>
|
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</mods:name>
|
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</mods:role>
|
||||
<mods:namePart id="374B4C9B824FFF2196718E7E6FF40E6E">Beckett, Marc</mods:namePart>
|
||||
</mods:name>
|
||||
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<mods:namePart id="20A76799787EEB7B952EC4DFD2D5B255">Ozgul-Siemund, Alev</mods:namePart>
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<mods:namePart id="99962480B044BAFFCB2FD19F1663269D">Frings, Jujina</mods:namePart>
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<mods:namePart id="7DD831F14BC93C4175D35BEF9153F61E">Fabrizi, Silvia</mods:namePart>
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||||
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|
||||
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<mods:namePart id="C676AB82E64AF22CCC4AE108CF6758A1">Ahrens, Dirk</mods:namePart>
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||||
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|
||||
<mods:typeOfResource id="50EFAD4A4462601399981EE38B2DA886">text</mods:typeOfResource>
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|
||||
<mods:title id="BDB1705C0313F075482292F1B609247D">Zoological Journal of the Linnean Society</mods:title>
|
||||
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|
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<mods:part id="F216CBF3103EA3E84C599B0A9BCC0940">
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<mods:date id="136B08EBF2AFBBDC33ED5AEFEEA2CF2F">2017</mods:date>
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<mods:title id="6FE772E28179D908FAD0C03FFDE4F11E">Zool. J. Linn. Soc.</mods:title>
|
||||
</mods:detail>
|
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<mods:detail id="107E03681ACBA26F1E79C2C08CD61E7D" type="volume">
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<mods:number id="B8B03A1A3FB7370CEC01F94981EC3EB9">180</mods:number>
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||||
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|
||||
<mods:detail id="7487331A87692D40A820A7EC910B7E6A" type="issue">
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<mods:number id="D984814F95D47A7F703A44D58183EAA2">2</mods:number>
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<mods:start id="951BEC081A6ECA37DF87750E033CD2F6">321</mods:start>
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<mods:url id="11FA54261450EC0DC530CB180D88C692">https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/</mods:url>
|
||||
</mods:location>
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||||
<mods:classification id="C7AAB298FB731341B641C0359438C9E7">journal article</mods:classification>
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<mods:identifier id="75C6574324CA9829494FBD1A6B81BDA2" type="DOI">10.1111/zoj.12489</mods:identifier>
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<mods:identifier id="B34048DA73B21F8D614A6986FA11C885" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="D838E3882B58AA672AFD10A3825345E4" type="Zenodo-Dep">14807083</mods:identifier>
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<mods:identifier id="F50D8C7AACBB1133399CC34C48BED584" type="ZooBank">3663E0A-2E17-4346-A388-6BEE53081814</mods:identifier>
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<treatment id="03CB6967FF8838583870FCE594EBF8DB" LSID="urn:lsid:plazi:treatment:03CB6967FF8838583870FCE594EBF8DB" httpUri="http://treatment.plazi.org/id/03CB6967FF8838583870FCE594EBF8DB" lastPageNumber="333" pageId="12" pageNumber="333">
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<subSubSection id="C3788BFAFF8838583870FCE59284FC62" pageId="12" pageNumber="333" type="nomenclature">
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<paragraph id="8BDDD871FF8838583870FCE59402FC31" blockId="12.[844,1384,847,904]" pageId="12" pageNumber="333">
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<heading id="D0956F1DFF8838583870FCE59402FC31" centered="true" fontSize="9" level="2" pageId="12" pageNumber="333" reason="2">
|
||||
<taxonomicName id="4C62A3F2FF8838583870FCE5954FFC33" ID-CoL="4K83V" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="333" phylum="Arthropoda" rank="species" species="congoensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8838583870FCE595E5FCD1" bold="true" box="[844,1155,850,871]" italics="true" pageId="12" pageNumber="333">
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<smallCapsWord id="8D3B4EADFF88385838D2FCE195E5FCD1" baselines="867" box="[1006,1155,854,871]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="congoensis" pageId="12" pageNumber="333">CONGOENSIS</smallCapsWord>
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</emphasis>
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<smallCapsWord id="8D3B4EADFF8838583FB0FCE7958EFCD1" baselines="866,867" box="[1164,1256,848,872]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="12" pageNumber="333">AHRENS</smallCapsWord>
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,
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<smallCapsWord id="8D3B4EADFF8838583FC5FCE79407FCD1" baselines="866,867" box="[1273,1377,848,872]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="12" pageNumber="333">BECKETT</smallCapsWord>
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,
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<smallCapsWord id="8D3B4EADFF8838583873FCD892CEFC33" baselines="897,897" box="[847,936,879,903]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="12" pageNumber="333">EBERLE</smallCapsWord>
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&
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</taxonomicName>
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<emphasis id="B9160463FF8838583F0FFCC395FEFC3E" bold="true" box="[1075,1176,877,904]" pageId="12" pageNumber="333">
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<taxonomicNameLabel id="A225B918FF8838583F0FFCC395FEFC3E" box="[1075,1176,877,904]" pageId="12" pageNumber="333" rank="species">
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<smallCapsWord id="8D3B4EADFF8838583F0FFCC39529FC33" baselines="897" box="[1075,1103,884,901]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="12" pageNumber="333">SP</smallCapsWord>
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.
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<smallCapsWord id="8D3B4EADFF8838583F5CFCC395F6FC33" baselines="897" box="[1120,1168,884,901]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="12" pageNumber="333">NOV</smallCapsWord>
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.
|
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</taxonomicNameLabel>
|
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</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF8838583F97FCD8945CFC33" box="[1195,1338,877,903]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="12" pageNumber="333">
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|
||||
2Y
|
||||
<emphasis id="B9160463FF8838583E30FCC6947CFC31" box="[1292,1306,881,903]" italics="true" pageId="12" pageNumber="333">–</emphasis>
|
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<smallCapsWord id="8D3B4EADFF8838583E26FCD8945CFC33" baselines="897,897" box="[1306,1338,879,903]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ab" pageId="12" pageNumber="333">AB</smallCapsWord>
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||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF8838583E77FCDA943DFC31" box="[1355,1371,877,903]" captionStart="Figure 7" captionStartId="14.[144,223,925,944]" captionTargetBox="[144,1428,192,893]" captionTargetId="figure-25@14.[144,1428,192,893]" captionTargetPageId="14" captionText="Figure 7. Distribution of Pleophylla species (left side) and absence records (right side), represented by sampling locali- ties of all other sampled Sericini species without records of Pleophylla (white dots)." figureDoi="http://doi.org/10.5281/zenodo.14807097" httpUri="https://zenodo.org/record/14807097/files/figure.png" pageId="12" pageNumber="333">7</figureCitation>
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||||
)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8838583815FC289284FC62" blockId="12.[809,1419,927,980]" pageId="12" pageNumber="333">
|
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<uri id="FFF3D473FF8838583815FC289284FC62" pageId="12" pageNumber="333">
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urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF8838583F06FC289284FC62" pageId="12" pageNumber="333">566E6542-C1A3-4176-A8D9- 1F62CD9CE164</uuid>
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</uri>
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|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF883858381BFC4694EBF8DB" pageId="12" pageNumber="333" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF883858381BFC469513FB6D" blockId="12.[807,1422,1008,1244]" pageId="12" pageNumber="333">
|
||||
<emphasis id="B9160463FF883858381BFC46953EFBB3" box="[807,1112,1008,1030]" italics="true" pageId="12" pageNumber="333">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF8838583F5BFC4794ECFB92" collectingDate="1948-04" collectionCode="RMCA" collectorName="Coll. Mus. & J. V. Leroy" country="Democratic Republic of the Congo" location="Bukima" pageId="12" pageNumber="333" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF8838583F5BFC4795ABFBB0" box="[1127,1229,1008,1030]" pageId="12" pageNumber="333" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D6413F8FF8838583FDCFC449596FBB3" box="[1248,1264,1011,1029]" count="1" pageId="12" pageNumber="333" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF8838583E31FC4794E8FBB0" box="[1293,1422,1008,1030]" pageId="12" pageNumber="333">Coll. Mus.</collectorName>
|
||||
<collectingCountry id="F37598E1FF883858381BFBB89208FB92" box="[807,878,1039,1060]" name="Democratic Republic of the Congo" pageId="12" pageNumber="333">Congo</collectingCountry>
|
||||
<location id="8EBD8EAAFF88385838B3FBB8928FFB93" LSID="urn:lsid:plazi:treatment:03CB6967FF8838583870FCE594EBF8DB:8EBD8EAAFF88385838B3FBB8928FFB93" box="[911,1001,1039,1061]" country="Democratic Republic of the Congo" name="Bukima" pageId="12" pageNumber="333">Bukima</location>
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||||
<date id="FFDCFEB1FF8838583F37FBB89503FB92" box="[1035,1125,1039,1060]" pageId="12" pageNumber="333" value="1948-04">
|
||||
<collectingDate id="EF980759FF8838583F37FBB89503FB92" box="[1035,1125,1039,1060]" pageId="12" pageNumber="333" value="1948-04">IV-1948</collectingDate>
|
||||
</date>
|
||||
<collectorName id="2697BDA7FF8838583F52FBB89598FB92" box="[1134,1278,1039,1060]" pageId="12" pageNumber="333">J.V. Leroy</collectorName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF8838583E0DFBB894E5FB92" box="[1329,1411,1039,1060]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="12" pageNumber="333" type="Museum">RMCA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B0AD22CFF883858381BFB9992E0FB36" collectingDate="1935-02-19" collectionCode="RMCA" collectorName="S. Bishoke & G. F. de Witte" country="Democratic Republic of the Congo" pageId="12" pageNumber="333" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||||
<typeStatus id="54D966D3FF883858381BFB9992FBFBF5" box="[807,925,1070,1091]" pageId="12" pageNumber="333" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF8838583890FB9A92B3FBF4" box="[940,981,1069,1090]" count="1" pageId="12" pageNumber="333" type="male">1 ♂</specimenCount>
|
||||
“1083,
|
||||
<collectorName id="2697BDA7FF8838583F0EFB9A95C9FBF5" box="[1074,1199,1069,1091]" pageId="12" pageNumber="333">S. Bishoke</collectorName>
|
||||
(2400), (Kibga), 8/
|
||||
<date id="FFDCFEB1FF883858381BFBFB92F9FBD4" box="[807,927,1100,1122]" pageId="12" pageNumber="333" value="1935-02-19">
|
||||
<collectingDate id="EF980759FF883858381BFBFB92F9FBD4" box="[807,927,1100,1122]" pageId="12" pageNumber="333" value="1935-02-19">19-II-1935</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="2697BDA7FF883858388EFBFB950DFBD4" box="[946,1131,1100,1122]" pageId="12" pageNumber="333">G. F. de Witte</collectorName>
|
||||
, PARC NAT. ALBERT” (
|
||||
<collectionCode id="ED7340B4FF8838583812FBDC9219FB36" box="[814,895,1131,1152]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="12" pageNumber="333" type="Museum">RMCA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="3B0AD22CFF88385838ABFBDC942DFB28" collectionCode="ZFMK" collectorName="Th. Wagner" country="Rwanda" elevation="2950" latitude="-1.46" location="Bisoke" longLatPrecision="785" longitude="29.48" pageId="12" pageNumber="333" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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||||
<specimenCount id="9D6413F8FF88385838ABFBDC92D8FBC9" box="[919,958,1131,1152]" count="1" pageId="12" pageNumber="333" type="female">1 ♀</specimenCount>
|
||||
“
|
||||
<collectingCountry id="F37598E1FF88385838E8FBDD9555FB36" box="[980,1075,1130,1152]" name="Rwanda" pageId="12" pageNumber="333">Rwanda</collectingCountry>
|
||||
,
|
||||
<location id="8EBD8EAAFF8838583F78FBDD95F5FB36" LSID="urn:lsid:plazi:treatment:03CB6967FF8838583870FCE594EBF8DB:8EBD8EAAFF8838583F78FBDD95F5FB36" box="[1092,1171,1130,1152]" country="Rwanda" latitude="-1.46" longLatPrecision="785" longitude="29.48" name="Bisoke" pageId="12" pageNumber="333">Bisoke</location>
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||||
,
|
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<geoCoordinate id="EE56BEB6FF8838583F9FFBDD95B9FB36" box="[1187,1247,1130,1152]" degrees="1,46" direction="south" orientation="latitude" pageId="12" pageNumber="333" precision="555" value="-1.46">1,46S</geoCoordinate>
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/
|
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<geoCoordinate id="EE56BEB6FF8838583FD4FBDD945CFB36" box="[1256,1338,1130,1152]" degrees="29,48" direction="east" orientation="longitude" pageId="12" pageNumber="333" precision="555" value="29.48">29,48E</geoCoordinate>
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,
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||||
<quantity id="4C9A7594FF8838583E77FBDD921AFB28" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.3" metricValueMin="2.6" pageId="12" pageNumber="333" unit="m" value="2950.0" valueMax="3300.0" valueMin="2600.0">
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<elevation id="004F3F42FF8838583E77FBDD921AFB28" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.3" metricValueMin="2.6" pageId="12" pageNumber="333" unit="m" value="2950.0" valueMax="3300.0" valueMin="2600.0">
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||||
2600
|
||||
<emphasis id="B9160463FF8838583EBDFBDA94EBFBC9" box="[1409,1421,1133,1151]" italics="true" pageId="12" pageNumber="333">–</emphasis>
|
||||
3300 m
|
||||
</elevation>
|
||||
</quantity>
|
||||
19.IX.02,
|
||||
<collectorName id="2697BDA7FF8838583F3AFB3E95FEFB28" box="[1030,1176,1161,1183]" pageId="12" pageNumber="333">Th. Wagner</collectorName>
|
||||
leg.” (
|
||||
<collectionCode id="ED7340B4FF8838583FC9FB3E9422FB28" box="[1269,1348,1161,1182]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="12" pageNumber="333" type="Museum">ZFMK</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="3B0AD22CFF8838583E5DFB3E9517FB6D" collectingDate="2007-09" collectionCode="ISNB" collectorName="Coll. I. R. Sc. N. B. & Coll. I. R. Sc. & P. N. Nyungwe & Leg. E. Vingerhoedt" country="Rwanda" location="Rwanda" pageId="12" pageNumber="333" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||||
<specimenCount id="9D6413F8FF8838583E5DFB3E94E8FB28" box="[1377,1422,1161,1182]" count="1" pageId="12" pageNumber="333" type="female">1 ♀</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF883858380CFB109292FB0B" box="[816,1012,1191,1213]" pageId="12" pageNumber="333">Coll. I.R.Sc.N.B.</collectorName>
|
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<collectingCountry id="F37598E1FF8838583F39FB109504FB0B" box="[1029,1122,1191,1213]" name="Rwanda" pageId="12" pageNumber="333">Rwanda</collectingCountry>
|
||||
<collectorName id="2697BDA7FF8838583F4FFB1F9444FB0B" box="[1139,1314,1192,1213]" pageId="12" pageNumber="333">P.N. Nyungwe</collectorName>
|
||||
<date id="FFDCFEB1FF8838583E0FFB1F94EBFB0B" box="[1331,1421,1192,1213]" pageId="12" pageNumber="333" value="2007-09">
|
||||
<collectingDate id="EF980759FF8838583E0FFB1F94EBFB0B" box="[1331,1421,1192,1213]" pageId="12" pageNumber="333" value="2007-09">IX-2007</collectingDate>
|
||||
</date>
|
||||
<collectorName id="2697BDA7FF883858381BFB71956AFB6A" box="[807,1036,1222,1244]" pageId="12" pageNumber="333">Leg. E. Vingerhoedt</collectorName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF8838583F1AFB71950FFB6D" box="[1062,1129,1222,1243]" pageId="12" pageNumber="333">ISNB</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF883858381BFB4294EBF8DB" blockId="12.[807,1422,1268,1901]" pageId="12" pageNumber="333">
|
||||
<emphasis id="B9160463FF883858381BFB4292C9FABC" box="[807,943,1269,1290]" italics="true" pageId="12" pageNumber="333">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8838583F56FB4395ADFABF" box="[1130,1227,1268,1289]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="12" pageNumber="333" unit="mm" value="9.1">9.1 mm</quantity>
|
||||
, elytral length: 7.0 mm, maximum width:
|
||||
<quantity id="4C9A7594FF8838583F53FAA495AFFA9E" box="[1135,1225,1299,1320]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="12" pageNumber="333" unit="mm" value="4.7">4.7 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially weakly sinuate; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.73. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra distinctly elevated but convex; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.27. Metatibia, ratio of width/length: 1/4.31. Metatarsi dorsally smooth; first
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
206
data/03/CB/69/03CB6967FF8838583B94FCAC920BFC9F.xml
Normal file
206
data/03/CB/69/03CB6967FF8838583B94FCAC920BFC9F.xml
Normal file
|
|
@ -0,0 +1,206 @@
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<taxonomicName id="4C62A3F2FF8838583B94FCAC9019FCF8" ID-CoL="4K83T" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="333" phylum="Arthropoda" rank="species" species="burundiensis" status="sp. nov.">
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,
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,
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&
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.
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<smallCapsWord id="8D3B4EADFF8838583A8AFC8A9080FCF8" baselines="842" box="[438,486,829,846]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="12" pageNumber="333">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF883858393CFC8093E2FCF9" box="[512,644,821,847]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="12" pageNumber="333">
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<smallCapsWord id="8D3B4EADFF883858393CFC809353FCF8" baselines="841,842" box="[512,565,823,847]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="12" pageNumber="333">FIGS</smallCapsWord>
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||||
2U
|
||||
<emphasis id="B9160463FF8838583958FC8E9314FCF9" box="[612,626,825,847]" italics="true" pageId="12" pageNumber="333">–</emphasis>
|
||||
X
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF88385839A9FC8293D1FCF9" box="[661,695,821,847]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="12" pageNumber="333">5E</figureCitation>
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,
|
||||
<figureCitation id="1359C4F4FF88385839F4FC8293BEFCF9" box="[712,728,821,847]" captionStart="Figure 7" captionStartId="14.[144,223,925,944]" captionTargetBox="[144,1428,192,893]" captionTargetId="figure-25@14.[144,1428,192,893]" captionTargetPageId="14" captionText="Figure 7. Distribution of Pleophylla species (left side) and absence records (right side), represented by sampling locali- ties of all other sampled Sericini species without records of Pleophylla (white dots)." figureDoi="http://doi.org/10.5281/zenodo.14807097" httpUri="https://zenodo.org/record/14807097/files/figure.png" pageId="12" pageNumber="333">7</figureCitation>
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)
|
||||
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|
||||
</paragraph>
|
||||
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<uri id="FFF3D473FF8838583BADFCDC905CFC29" pageId="12" pageNumber="333">
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urn:lsid:zoobank.org:act:
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</subSubSection>
|
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<subSubSection id="C3788BFAFF8838583BACFC0C920BFC9F" pageId="12" pageNumber="333" type="materials_examined">
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|
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<emphasis id="B9160463FF8838583BACFC0C90A8FC66" box="[144,462,955,976]" italics="true" pageId="12" pageNumber="333">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF8838583AE0FC0C9026FC58" collectionCode="ZFMK" country="Burundi" location="Ngozi" pageId="12" pageNumber="333" specimenCount="1" specimenCount-male="1" stateProvince="Ngozi" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF8838583AE0FC0C9323FC67" box="[476,581,955,977]" pageId="12" pageNumber="333" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF883858395FFC0A9315FC79" box="[611,627,957,975]" count="1" pageId="12" pageNumber="333" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectingCountry id="F37598E1FF88385839ABFC0C9390FC67" box="[663,758,955,977]" name="Burundi" pageId="12" pageNumber="333">Burundi</collectingCountry>
|
||||
<collectingRegion id="49A61693FF8838583BACFC6E91B7FC59" box="[144,209,985,1007]" country="Burundi" name="Ngozi" pageId="12" pageNumber="333">Ngozi</collectingRegion>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF8838583BD6FC6E905FFC58" box="[234,313,985,1006]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="12" pageNumber="333" type="Museum">ZFMK</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<typeStatus id="54D966D3FF8838583A70FC6D90A4FC59" box="[332,450,986,1007]" pageId="12" pageNumber="333" type="paratype">Paratypes</typeStatus>
|
||||
: see Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8838583BACFBB89372F973" blockId="12.[144,759,1039,1733]" pageId="12" pageNumber="333">
|
||||
<emphasis id="B9160463FF8838583BACFBB8907EFB92" box="[144,280,1039,1060]" italics="true" pageId="12" pageNumber="333">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8838583AECFBB89351FB92" box="[464,567,1039,1060]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="12" pageNumber="333" unit="mm" value="10.3">10.3 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF8838583BACFB9A918FFBF5" box="[144,233,1069,1091]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="12" pageNumber="333" unit="mm" value="7.1">7.1 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF8838583AE4FB9A9355FBF5" box="[472,563,1069,1091]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="12" pageNumber="333" unit="mm" value="4.6">4.6 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially weakly sinuate; margins of labroclypeus strongly reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.7. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity condensed to transverse patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/ metacoxa: 1/1.33. Metatibia, ratio of width/length: 1/4. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF8838583938F92793E5F910" box="[516,643,1680,1702]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="12" pageNumber="333">
|
||||
Fig. 2U
|
||||
<emphasis id="B9160463FF8838583967F9249301F913" box="[603,615,1683,1701]" italics="true" pageId="12" pageNumber="333">–</emphasis>
|
||||
W
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF8838583BACF9189181F972" box="[144,231,1711,1733]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="12" pageNumber="333">Fig. 2X</figureCitation>
|
||||
. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF8838583A85F9189376F973" box="[441,528,1711,1733]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="12" pageNumber="333">Fig. 5E</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8838583BACF96A943AFE8E" blockId="12.[144,759,1757,1901]" lastBlockId="12.[807,1421,199,312]" pageId="12" pageNumber="333">
|
||||
<emphasis id="B9160463FF8838583BACF96A9328F944" box="[144,590,1757,1778]" italics="true" pageId="12" pageNumber="333">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF8838583A29F96A9328F944" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[277,590,1757,1778]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="333" phylum="Arthropoda" rank="species" species="burundiensis" status="sp. nov.">Pleophylla burundiensis</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF8838583946F96A9391F944" box="[634,759,1757,1778]" pageId="12" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
resembles
|
||||
<taxonomicName id="4C62A3F2FF8838583A31F94B9017F8A7" box="[269,369,1788,1809]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="333" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B9160463FF8838583A31F94B9017F8A7" box="[269,369,1788,1809]" italics="true" pageId="12" pageNumber="333">P. pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
in external shape; however, the punctures on the dorsal surface are denser and coarser, and the sutural interval is flat and not limited by a sharp carina in the apical declivity of the elytra. This new species is in genital shape similar to
|
||||
<taxonomicName id="4C62A3F2FF8838583875FF519563FF4C" authorityName="Boheman" authorityYear="1857" box="[841,1029,229,251]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="333" phylum="Arthropoda" rank="species" species="maculipennis">
|
||||
<emphasis id="B9160463FF8838583875FF519563FF4C" box="[841,1029,229,251]" italics="true" pageId="12" pageNumber="333">P. maculipennis</emphasis>
|
||||
</taxonomicName>
|
||||
; however, the right paramere is basally tooth-like and extended, and the basal dorsal portion of the left paramere bears two small teeth.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF883858381BFEED9553FDD2" blockId="12.[807,1422,346,612]" pageId="12" pageNumber="333">
|
||||
<emphasis id="B9160463FF883858381BFEED92FCFED9" box="[807,922,346,367]" italics="true" pageId="12" pageNumber="333">Variation:</emphasis>
|
||||
Body length: 8.0
|
||||
<emphasis id="B9160463FF8838583F54FEEA9512FED9" box="[1128,1140,349,367]" italics="true" pageId="12" pageNumber="333">–</emphasis>
|
||||
<quantity id="4C9A7594FF8838583F48FEED95BCFED9" box="[1140,1242,346,367]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="12" pageNumber="333" unit="mm" value="10.3">10.3 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF883858381BFECF92C8FE38" box="[807,942,376,398]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.1" metricValueMin="6.5" pageId="12" pageNumber="333" unit="mm" value="6.8" valueMax="7.1" valueMin="6.5">
|
||||
6.5
|
||||
<emphasis id="B9160463FF8838583875FECC9233FE3B" box="[841,853,379,397]" italics="true" pageId="12" pageNumber="333">–</emphasis>
|
||||
7.1 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF8838583FA6FECE9444FE38" box="[1178,1314,377,398]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.8" metricValueMin="4.2" pageId="12" pageNumber="333" unit="mm" value="4.5" valueMax="4.8" valueMin="4.2">
|
||||
4.2
|
||||
<emphasis id="B9160463FF8838583F80FECC95AEFE3B" box="[1212,1224,379,397]" italics="true" pageId="12" pageNumber="333">–</emphasis>
|
||||
4.8 mm
|
||||
</quantity>
|
||||
. Density of dark spot may vary slightly, but never completely without spots. Female: antennal club of female straight, composed of five antennomeres, as long as the remaining antennomeres combined; sixth antennomere very short, less than 1/5 of club length. Eyes smaller than in male: ratio of eye diameter/ interocular width: 0.52.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF883858381BFD379549FD03" blockId="12.[807,1421,640,693]" pageId="12" pageNumber="333">
|
||||
<emphasis id="B9160463FF883858381BFD3792C2FD23" box="[807,932,640,661]" italics="true" pageId="12" pageNumber="333">Etymology:</emphasis>
|
||||
The new species is named after its occurrence in
|
||||
<collectingCountry id="F37598E1FF88385838F4FD28954DFD03" box="[968,1067,671,693]" name="Burundi" pageId="12" pageNumber="333">Burundi</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF883858381BFD61920BFC9F" blockId="12.[807,1421,726,809]" pageId="12" pageNumber="333">
|
||||
<emphasis id="B9160463FF883858381BFD6192F4FD5D" box="[807,914,726,747]" italics="true" pageId="12" pageNumber="333">Remarks:</emphasis>
|
||||
In a previous publication we referred to this species as ‘SpM-21’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF8838583E08FD429207FC9F" author="Ahrens D" pageId="12" pageNumber="333" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
213
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213
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<mods:title id="6560E179918CD6D812C5AA84C163F4B0">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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|
||||
<heading id="D0956F1DFF8A385A3BA9F9C9938BF904" centered="true" fontSize="9" level="2" pageId="14" pageNumber="335" reason="2">
|
||||
<taxonomicName id="4C62A3F2FF8A385A3BA9F9C99015F907" ID-CoL="6W2MY" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="species" species="settentrionalis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8A385A3BA9F9C9936DF924" bold="true" box="[149,523,1662,1683]" italics="true" pageId="14" pageNumber="335">
|
||||
<smallCapsWord id="8D3B4EADFF8A385A3BA9F9C99049F924" baselines="1678,1678" box="[149,303,1662,1683]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="14" pageNumber="335">PLEOPHYLLA</smallCapsWord>
|
||||
<smallCapsWord id="8D3B4EADFF8A385A3A04F936936DF924" baselines="1678" box="[312,523,1665,1682]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="settentrionalis" pageId="14" pageNumber="335">SETTENTRIONALIS</smallCapsWord>
|
||||
</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF8A385A3929F9CB9316F924" baselines="1678,1678" box="[533,624,1660,1684]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="14" pageNumber="335">AHRENS</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF8A385A39BEF9CB938FF924" baselines="1678,1678" box="[642,745,1660,1684]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="14" pageNumber="335">BECKETT</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF8A385A3BA5F92D9197F906" baselines="1708,1708" box="[153,241,1690,1714]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="14" pageNumber="335">EBERLE</smallCapsWord>
|
||||
&
|
||||
<smallCapsWord id="8D3B4EADFF8A385A3A27F92D9015F907" baselines="1708,1709" box="[283,371,1690,1714]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabrizi" pageId="14" pageNumber="335">FABRIZI</smallCapsWord>
|
||||
</taxonomicName>
|
||||
<emphasis id="B9160463FF8A385A3A40F9179084F905" bold="true" box="[380,482,1688,1715]" pageId="14" pageNumber="335">
|
||||
<taxonomicNameLabel id="A225B918FF8A385A3A40F9179084F905" box="[380,482,1688,1715]" pageId="14" pageNumber="335" rank="species">
|
||||
<smallCapsWord id="8D3B4EADFF8A385A3A40F91790FEF907" baselines="1709" box="[380,408,1696,1713]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="14" pageNumber="335">SP</smallCapsWord>
|
||||
.
|
||||
<smallCapsWord id="8D3B4EADFF8A385A3A96F91790BCF907" baselines="1709" box="[426,474,1696,1713]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="14" pageNumber="335">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF8A385A3AC8F92D93F7F906" box="[500,657,1688,1714]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="14" pageNumber="335">
|
||||
<smallCapsWord id="8D3B4EADFF8A385A3AC8F92D934FF907" baselines="1708,1709" box="[500,553,1690,1714]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="14" pageNumber="335">FIGS</smallCapsWord>
|
||||
2
|
||||
<smallCapsWord id="8D3B4EADFF8A385A397FF92D9305F907" baselines="1708,1709" box="[579,611,1690,1714]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ac" pageId="14" pageNumber="335">AC</smallCapsWord>
|
||||
<emphasis id="B9160463FF8A385A3958F92B9314F904" box="[612,626,1692,1714]" italics="true" pageId="14" pageNumber="335">–</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF8A385A394EF92D93F7F906" baselines="1708,1708" box="[626,657,1690,1714]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Af" pageId="14" pageNumber="335">AF</smallCapsWord>
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF8A385A399EF92F93A5F904" box="[674,707,1688,1714]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="14" pageNumber="335">5F</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF8A385A39E8F92F9382F904" box="[724,740,1688,1714]" captionStart="Figure 7" captionStartId="14.[144,223,925,944]" captionTargetBox="[144,1428,192,893]" captionTargetId="figure-25@14.[144,1428,192,893]" captionTargetPageId="14" captionText="Figure 7. Distribution of Pleophylla species (left side) and absence records (right side), represented by sampling locali- ties of all other sampled Sericini species without records of Pleophylla (white dots)." figureDoi="http://doi.org/10.5281/zenodo.14807097" httpUri="https://zenodo.org/record/14807097/files/figure.png" pageId="14" pageNumber="335">7</figureCitation>
|
||||
)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8A385A3BADF978905EF8B5" blockId="14.[145,761,1743,1795]" pageId="14" pageNumber="335">
|
||||
<uri id="FFF3D473FF8A385A3BADF978905EF8B5" pageId="14" pageNumber="335">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF8A385A3AA5F978905EF8B5" pageId="14" pageNumber="335">FAA40DFF-EB25-413D-8573- 6993258C3090</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF8A385B3BACF8AD93BAFD73" lastPageId="15" lastPageNumber="336" pageId="14" pageNumber="335" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF8A385A3BACF8AD9324F8DA" blockId="14.[144,759,1818,1901]" pageId="14" pageNumber="335">
|
||||
<emphasis id="B9160463FF8A385A3BACF8AD90A7F899" box="[144,449,1818,1839]" italics="true" pageId="14" pageNumber="335">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF8A385A3AECF8AD9196F8DA" collectingDate="1951-11" collectionCode="RMCA" collectorName="Coll. Mus. & N. Lac & J. V. Leroy" country="Democratic Republic of the Congo" location="Rwankwi" pageId="14" pageNumber="335" specimenCount="1" specimenCount-male="1" stateProvince="Sud-Kivu" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF8A385A3AECF8AD9350F886" box="[464,566,1818,1840]" pageId="14" pageNumber="335" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D6413F8FF8A385A3975F8AA933FF899" box="[585,601,1821,1839]" count="1" pageId="14" pageNumber="335" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF8A385A394BF8AD9391F899" box="[631,759,1818,1840]" pageId="14" pageNumber="335">Coll. Mus.</collectorName>
|
||||
<collectingCountry id="F37598E1FF8A385A3BACF88F91B1F8FB" box="[144,215,1848,1869]" name="Democratic Republic of the Congo" pageId="14" pageNumber="335">Congo</collectingCountry>
|
||||
<collectorName id="2697BDA7FF8A385A3BDAF88E905CF8F8" box="[230,314,1849,1870]" pageId="14" pageNumber="335">N. Lac</collectorName>
|
||||
<collectingRegion id="49A61693FF8A385A3A76F88F90E7F8F8" box="[330,385,1848,1870]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="14" pageNumber="335">Kivu</collectingRegion>
|
||||
<location id="8EBD8EAAFF8A385A3AADF88F909AF8F8" LSID="urn:lsid:plazi:treatment:03CB6967FF8A385B3BA9F9C993BAFD73:8EBD8EAAFF8A385A3AADF88F909AF8F8" box="[401,508,1848,1870]" country="Democratic Republic of the Congo" name="Rwankwi" pageId="14" pageNumber="335" stateProvince="Sud-Kivu">Rwankwi</location>
|
||||
<date id="FFDCFEB1FF8A385A3930F88F9300F8F8" box="[524,614,1848,1870]" pageId="14" pageNumber="335" value="1951-11">
|
||||
<collectingDate id="EF980759FF8A385A3930F88F9300F8F8" box="[524,614,1848,1870]" pageId="14" pageNumber="335" value="1951-11">XI-1951</collectingDate>
|
||||
</date>
|
||||
<collectorName id="2697BDA7FF8A385A3953F88E938AF8FB" box="[623,748,1848,1870]" pageId="14" pageNumber="335">J.V. Leroy</collectorName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF8A385A3BABF8E0918FF8DA" box="[151,233,1879,1900]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="14" pageNumber="335" type="Museum">RMCA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<typeStatus id="54D966D3FF8A385A3BC7F8E09017F8DA" box="[251,369,1879,1900]" pageId="14" pageNumber="335" type="paratype">Paratypes</typeStatus>
|
||||
: see Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8A385A381BFBB7928BF900" blockId="14.[807,1422,1023,1718]" pageId="14" pageNumber="335">
|
||||
<emphasis id="B9160463FF8A385A381BFBB792C9FBA3" box="[807,943,1024,1045]" italics="true" pageId="14" pageNumber="335">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8A385A3F53FBB795A1FBA3" box="[1135,1223,1024,1045]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="14" pageNumber="335" unit="mm" value="7.7">7.7 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF8A385A381BFBA99219FB82" box="[807,895,1054,1076]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="14" pageNumber="335" unit="mm" value="5.6">5.6 mm</quantity>
|
||||
, maximum width: 4.0 mm. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially weakly sinuate; margins of labroclypeus strongly reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.77. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.35. Metatibia, ratio of width/length: 1/4. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF8A385A3891F9369288F920" box="[941,1006,1665,1687]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="14" pageNumber="335">Fig. 2</figureCitation>
|
||||
Ac
|
||||
<emphasis id="B9160463FF8A385A3F37F9339571F920" box="[1035,1047,1668,1686]" italics="true" pageId="14" pageNumber="335">–</emphasis>
|
||||
Ae. Habitus:
|
||||
<figureCitation id="1359C4F4FF8A385A3FF8F9369463F921" box="[1220,1285,1665,1687]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="14" pageNumber="335">Fig. 2</figureCitation>
|
||||
Af. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF8A385A38A8F917928FF900" box="[916,1001,1696,1718]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="14" pageNumber="335">
|
||||
Fig.
|
||||
<specimenCount id="9D6413F8FF8A385A38F5F917928FF900" box="[969,1001,1696,1718]" count="5" pageId="14" pageNumber="335" type="generic">5F</specimenCount>
|
||||
</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8A385A381BF96992F4F8DB" blockId="14.[807,1422,1757,1901]" pageId="14" pageNumber="335">
|
||||
<emphasis id="B9160463FF8A385A381BF96995B4F944" box="[807,1234,1757,1779]" italics="true" pageId="14" pageNumber="335">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF8A385A3890F96A95B4F944" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[940,1234,1757,1778]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="species" species="settentrionalis" status="sp. nov.">Pleophylla settentrionalis</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF8A385A3FE1F96A945FF944" box="[1245,1337,1757,1778]" pageId="14" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from the previous two species,
|
||||
<taxonomicName id="4C62A3F2FF8A385A3FADF94B944CF8A7" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1169,1322,1788,1809]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="species" species="congoensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8A385A3FADF94B944CF8A7" box="[1169,1322,1788,1809]" italics="true" pageId="14" pageNumber="335">P. congoensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8A385A3E08F94B94E8F8A7" box="[1332,1422,1788,1809]" pageId="14" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C62A3F2FF8A385A3866F8AC9569F899" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[858,1039,1818,1840]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="species" species="burundiensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8A385A3866F8AC9569F899" box="[858,1039,1818,1840]" italics="true" pageId="14" pageNumber="335">P. burundiensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8A385A3F24F8AC9512F886" box="[1048,1140,1819,1840]" pageId="14" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, by having the centre of the elytral intervals impunctate rather than densely punctate.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8B385B3B9AFF7093CCFE67" blockId="15.[166,781,199,465]" pageId="15" pageNumber="336">
|
||||
<emphasis id="B9160463FF8B385B3B9AFF70907CFF6A" box="[166,282,199,220]" italics="true" pageId="15" pageNumber="336">Variation:</emphasis>
|
||||
Body length: 7.6
|
||||
<emphasis id="B9160463FF8B385B3AD2FF7E909CFF6D" box="[494,506,201,219]" italics="true" pageId="15" pageNumber="336">–</emphasis>
|
||||
9.0 mm, elytral length: 5.6
|
||||
<emphasis id="B9160463FF8B385B3BF4FF5F91B2FF4C" box="[200,212,232,250]" italics="true" pageId="15" pageNumber="336">–</emphasis>
|
||||
7.0 mm, maximum width: 4.0
|
||||
<emphasis id="B9160463FF8B385B3900FF5F932EFF4C" box="[572,584,232,250]" italics="true" pageId="15" pageNumber="336">–</emphasis>
|
||||
<quantity id="4C9A7594FF8B385B3974FF5293C4FF4D" box="[584,674,229,251]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="15" pageNumber="336" unit="mm" value="4.8">4.8 mm</quantity>
|
||||
. Density of dark spots may vary, sometimes completely without spots. Female: antennal club of female straight, composed of five antennomeres, slightly longer than the remaining antennomeres combined; sixth antennomere short, slightly shorter than a quarter of club length. Eyes smaller than in male: ratio of eye diameter/interocular width: 0.53.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8B385B3B9AFE4F9198FDDC" blockId="15.[166,781,504,618]" pageId="15" pageNumber="336">
|
||||
<emphasis id="B9160463FF8B385B3B9AFE4F9042FDBB" box="[166,292,504,525]" italics="true" pageId="15" pageNumber="336">Etymology:</emphasis>
|
||||
This new species is named
|
||||
<taxonomicName id="4C62A3F2FF8B385B3B9AFDA0902BFD9A" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[166,333,535,556]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="settentrionalis">
|
||||
‘
|
||||
<emphasis id="B9160463FF8B385B3B97FDA0902BFD9A" box="[171,333,535,556]" italics="true" pageId="15" pageNumber="336">settentrionalis</emphasis>
|
||||
</taxonomicName>
|
||||
’ in reference to its occurrence in the northernmost part of the known range of
|
||||
<taxonomicName id="4C62A3F2FF8B385B39A9FD829198FDDC" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="undetermined">
|
||||
<emphasis id="B9160463FF8B385B39A9FD82926BFDFC" box="[661,781,565,586]" italics="true" pageId="15" pageNumber="336">Pleophylla</emphasis>
|
||||
species.
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8B385B3B9AFD2693BAFD73" blockId="15.[166,780,657,709]" pageId="15" pageNumber="336">
|
||||
<emphasis id="B9160463FF8B385B3B9AFD269074FD10" box="[166,274,657,678]" italics="true" pageId="15" pageNumber="336">Remarks:</emphasis>
|
||||
In a previous publication we referred to this species as ‘sp27’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF8B385B390BFD1893B6FD73" author="Ahrens D" box="[567,720,687,709]" pageId="15" pageNumber="336" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
209
data/03/CB/69/03CB6967FF8B38443876FDA091B0FDDC.xml
Normal file
209
data/03/CB/69/03CB6967FF8B38443876FDA091B0FDDC.xml
Normal file
|
|
@ -0,0 +1,209 @@
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<mods:title id="93FEF849CD1E799A07FA99C7C45F5F3A">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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<mods:namePart id="73F408866AED6C7A837E28BA3755E404">Eberle, Jonas</mods:namePart>
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|
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|
||||
<taxonomicName id="4C62A3F2FF8B385B3876FDA0929CFDFC" ID-CoL="4K847" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="murzini" status="sp. nov.">
|
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,
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||||
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|
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,
|
||||
<smallCapsWord id="8D3B4EADFF8B385B3E03FDA294F1FD9D" baselines="551,551" box="[1343,1431,533,557]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="15" pageNumber="336">EBERLE</smallCapsWord>
|
||||
&
|
||||
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|
||||
</taxonomicName>
|
||||
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|
||||
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|
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.
|
||||
<smallCapsWord id="8D3B4EADFF8B385B3F0CFD8E9506FDFC" baselines="582" box="[1072,1120,569,586]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="15" pageNumber="336">NOV</smallCapsWord>
|
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.
|
||||
</taxonomicNameLabel>
|
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</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF8B385B3F47FD849599FDFD" box="[1147,1279,561,587]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="15" pageNumber="336">
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<smallCapsWord id="8D3B4EADFF8B385B3F47FD8495D6FDFC" baselines="581,582" box="[1147,1200,563,587]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="15" pageNumber="336">FIGS</smallCapsWord>
|
||||
3E
|
||||
<emphasis id="B9160463FF8B385B3FE0FD82958CFDFD" box="[1244,1258,565,587]" italics="true" pageId="15" pageNumber="336">–</emphasis>
|
||||
H
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF8B385B3E2DFD869453FDFD" box="[1297,1333,561,587]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="15" pageNumber="336">5H</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
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<uri id="FFF3D473FF8B385B3807FDDD928BFD29" pageId="15" pageNumber="336">
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urn:lsid:zoobank.org:act:
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|
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|
||||
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|
||||
<emphasis id="B9160463FF8B385B3801FD0E9515FD78" box="[829,1139,697,718]" italics="true" pageId="15" pageNumber="336">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF8B385B3FBDFD0E9586FCBD" collectionCode="ZFMK" collectorName="S. Africa & S. Murzin" elevation="1250" location="Katberg" pageId="15" pageNumber="336" specimenCount="1" specimenCount-male="1" stateProvince="Eastern Cape" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF8B385B3FBDFD0E958CFD79" box="[1153,1258,697,719]" pageId="15" pageNumber="336" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF8B385B3E38FD0C9472FD7B" box="[1284,1300,699,717]" count="1" pageId="15" pageNumber="336" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF8B385B3E09FD0E94C2FD79" box="[1333,1444,697,719]" pageId="15" pageNumber="336">S. Africa</collectorName>
|
||||
<collectingRegion id="49A61693FF8B385B3801FD6F9296FD5A" box="[829,1008,727,749]" country="South Africa" name="Eastern Cape" pageId="15" pageNumber="336">Eastern Cape</collectingRegion>
|
||||
pr
|
||||
<location id="8EBD8EAAFF8B385B3F7AFD6095C4FD5B" LSID="urn:lsid:plazi:treatment:03CB6967FF8B38443876FDA091B0FDDC:8EBD8EAAFF8B385B3F7AFD6095C4FD5B" box="[1094,1186,727,749]" name="Katberg" pageId="15" pageNumber="336" stateProvince="Eastern Cape">Katberg</location>
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pass, h =
|
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<quantity id="4C9A7594FF8B385B3E74FD6094F9FD5B" box="[1352,1439,727,749]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="336" unit="m" value="1250.0">
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||||
<elevation id="004F3F42FF8B385B3E74FD6094F9FD5B" box="[1352,1439,727,749]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="336" unit="m" value="1250.0">1250 m</elevation>
|
||||
</quantity>
|
||||
; 09.1.2002
|
||||
<collectorName id="2697BDA7FF8B385B3887FD419550FCBA" box="[955,1078,758,780]" pageId="15" pageNumber="336">S. Murzin</collectorName>
|
||||
leg” (
|
||||
<collectionCode id="ED7340B4FF8B385B3FB6FD4195BFFCBD" box="[1162,1241,758,779]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="15" pageNumber="336" type="Museum">ZFMK</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<typeStatus id="54D966D3FF8B385B3FC9FD41940DFCBD" box="[1269,1387,758,779]" pageId="15" pageNumber="336" type="paratype">Paratypes</typeStatus>
|
||||
: see Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8B385B3801FCF094C4FA68" blockId="15.[829,1445,838,1502]" pageId="15" pageNumber="336">
|
||||
<emphasis id="B9160463FF8B385B3801FCF092A0FCEA" box="[829,966,839,860]" italics="true" pageId="15" pageNumber="336">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8B385B3FB5FCF195BCFCEA" box="[1161,1242,838,860]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="336" unit="mm" value="11.0">11 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF8B385B3801FCD292C4FCCC" box="[829,930,869,890]" metricMagnitude="-3" metricUnit="m" metricValue="8.14" pageId="15" pageNumber="336" unit="mm" value="8.14">8.14 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF8B385B3FB8FCD2958DFCCC" box="[1156,1259,869,891]" metricMagnitude="-3" metricUnit="m" metricValue="6.14" pageId="15" pageNumber="336" unit="mm" value="6.14">6.14 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.63. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra distinctly elevated but convex; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.44. Metatibia, ratio of width/length: 1/4.11. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF8B385B3801FA7F92C9FA6B" box="[829,943,1480,1502]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="15" pageNumber="336">Fig. 3E, F</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF8B385B3F22FA7F9510FA6B" box="[1054,1142,1480,1502]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="15" pageNumber="336">Fig. 3H</figureCitation>
|
||||
. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF8B385B3E79FA7F94F8FA68" box="[1349,1438,1480,1502]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="15" pageNumber="336">Fig. 5H</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8B385B3801FA4D95C1F894" blockId="15.[829,1444,1530,1827]" pageId="15" pageNumber="336">
|
||||
<emphasis id="B9160463FF8B385B3801FA4D92D2F9B9" box="[829,948,1530,1551]" italics="true" pageId="15" pageNumber="336">Diagnosis:</emphasis>
|
||||
The new species is rather similar to
|
||||
<taxonomicName id="4C62A3F2FF8B385B3801F9AE92B2F99B" box="[829,980,1560,1582]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="ferruginea">
|
||||
<emphasis id="B9160463FF8B385B3801F9AE92B2F99B" box="[829,980,1560,1582]" italics="true" pageId="15" pageNumber="336">P. ferruginea</emphasis>
|
||||
</taxonomicName>
|
||||
in the shape of the male genitalia.
|
||||
<taxonomicName id="4C62A3F2FF8B385B3801F9809546F9FA" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" box="[829,1056,1591,1612]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="murzini">
|
||||
<emphasis id="B9160463FF8B385B3801F9809546F9FA" box="[829,1056,1591,1612]" italics="true" pageId="15" pageNumber="336">Pleophylla murzini</emphasis>
|
||||
</taxonomicName>
|
||||
differs from the latter by the slightly larger body size and the shape of parameres: the basal basomedian branches (tooth) of the left and right paramere are much larger than the distal basomedian branches (whereas in
|
||||
<taxonomicName id="4C62A3F2FF8B385B3FEDF905940EF970" box="[1233,1384,1713,1735]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="ferruginea">
|
||||
<emphasis id="B9160463FF8B385B3FEDF905940EF970" box="[1233,1384,1713,1735]" italics="true" pageId="15" pageNumber="336">P. ferruginea</emphasis>
|
||||
</taxonomicName>
|
||||
they are subequal in size). From the other large species (following below) it differs by the complete basal marginal line on the pronotum.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8B38443801F88E90F8FE22" blockId="15.[829,1444,1849,1901]" lastBlockId="16.[144,759,199,404]" lastPageId="16" lastPageNumber="337" pageId="15" pageNumber="336">
|
||||
<emphasis id="B9160463FF8B385B3801F88E92D7F8F8" box="[829,945,1849,1870]" italics="true" pageId="15" pageNumber="336">Variation:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8B385B3F5AF88E9581F8F8" box="[1126,1255,1849,1870]" metricMagnitude="-3" metricUnit="m" metricValue="9.95" metricValueMax="11.0" metricValueMin="8.9" pageId="15" pageNumber="336" unit="mm" value="9.95" valueMax="11.0" valueMin="8.9">
|
||||
8.9
|
||||
<emphasis id="B9160463FF8B385B3FB4F88B95F2F8F8" box="[1160,1172,1852,1870]" italics="true" pageId="15" pageNumber="336">–</emphasis>
|
||||
11 mm
|
||||
</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF8B385B3801F8E092A3F8DB" box="[829,965,1879,1901]" metricMagnitude="-3" metricUnit="m" metricValue="7.65" metricValueMax="8.5" metricValueMin="6.8" pageId="15" pageNumber="336" unit="mm" value="7.65" valueMax="8.5" valueMin="6.8">
|
||||
6.8
|
||||
<emphasis id="B9160463FF8B385B3863F8ED920DF8DA" box="[863,875,1882,1900]" italics="true" pageId="15" pageNumber="336">–</emphasis>
|
||||
8.5 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF8B385B3F8DF8E0945FF8DB" box="[1201,1337,1879,1901]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="6.5" metricValueMin="4.8" pageId="15" pageNumber="336" unit="mm" value="5.65" valueMax="6.5" valueMin="4.8">
|
||||
4.8
|
||||
<emphasis id="B9160463FF8B385B3FEFF8ED95B9F8DA" box="[1235,1247,1882,1900]" italics="true" pageId="15" pageNumber="336">–</emphasis>
|
||||
6.5 mm
|
||||
</quantity>
|
||||
. Density of dark spot may vary, sometimes completely without spots. Female: antennal club of female straight, composed of five antennomeres, slightly longer than the remaining antennomeres combined; sixth antennomere short, half as long as club length. Eyes smaller than in male: ratio of eye diameter/ interocular width: 0.53.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9438443BACFE0C937DFE59" blockId="16.[144,758,443,496]" pageId="16" pageNumber="337">
|
||||
<emphasis id="B9160463FF9438443BACFE0C906BFE66" box="[144,269,443,464]" italics="true" pageId="16" pageNumber="337">Etymology:</emphasis>
|
||||
The new species is named after its collector, Sergej Murzin (Moscow).
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9438443BACFDA091B0FDDC" blockId="16.[144,758,535,618]" pageId="16" pageNumber="337">
|
||||
<emphasis id="B9160463FF9438443BACFDA0919DFD9A" box="[144,251,535,556]" italics="true" pageId="16" pageNumber="337">Remarks:</emphasis>
|
||||
In a previous publication we referred to this species as ‘SpM-16’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF94384439A1FD8291ACFDDC" author="Ahrens D" pageId="16" pageNumber="337" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
175
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Normal file
175
data/03/CB/69/03CB6967FF8B385B3BEEFD4A92F7FE52.xml
Normal file
|
|
@ -0,0 +1,175 @@
|
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<mods:namePart id="CAD935F9A10D71EBC5AAEB517715237E">Beckett, Marc</mods:namePart>
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<mods:namePart id="5559BDFA6E2F24E59055C629A287E1E0">Ozgul-Siemund, Alev</mods:namePart>
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<mods:namePart id="671CB812511A68BB8A9CA2BDFAFC13E3">Frings, Jujina</mods:namePart>
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<mods:namePart id="4A3C39D4FE04802D29994370B87B2D4B">Fabrizi, Silvia</mods:namePart>
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<mods:namePart id="5B0F9665831214B98F375E1DBD81A285">Ahrens, Dirk</mods:namePart>
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||||
</mods:mods>
|
||||
<treatment id="03CB6967FF8B385B3BEEFD4A92F7FE52" LSID="urn:lsid:plazi:treatment:03CB6967FF8B385B3BEEFD4A92F7FE52" httpUri="http://treatment.plazi.org/id/03CB6967FF8B385B3BEEFD4A92F7FE52" lastPageNumber="336" pageId="15" pageNumber="336">
|
||||
<subSubSection id="C3788BFAFF8B385B3BEEFD4A9329FC86" pageId="15" pageNumber="336" type="nomenclature">
|
||||
<paragraph id="8BDDD871FF8B385B3BEEFD4A9329FC86" blockId="15.[210,737,760,817]" pageId="15" pageNumber="336">
|
||||
<heading id="D0956F1DFF8B385B3BEEFD4A9329FC86" centered="true" fontSize="9" level="2" pageId="15" pageNumber="336" reason="2">
|
||||
<taxonomicName id="4C62A3F2FF8B385B3BEEFD4A9387FCA4" ID-CoL="4K83X" authority="BURMEISTER, 1855" authorityName="Burmeister" authorityYear="1855" box="[210,737,760,787]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="ferruginea">
|
||||
<emphasis id="B9160463FF8B385B3BEEFD4A909EFCA6" box="[210,504,765,786]" italics="true" pageId="15" pageNumber="336">
|
||||
<smallCapsWord id="8D3B4EADFF8B385B3BEEFD4A9005FCA6" baselines="781,780" box="[210,355,765,786]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="15" pageNumber="336">PLEOPHYLLA</smallCapsWord>
|
||||
<smallCapsWord id="8D3B4EADFF8B385B3A50FCB7909EFCA6" baselines="780" box="[364,504,768,784]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="ferruginea" pageId="15" pageNumber="336">FERRUGINEA</smallCapsWord>
|
||||
</emphasis>
|
||||
<bibRefCitation id="EFF3A580FF8B385B393EFD4D9387FCA4" author="Burmeister H" box="[514,737,760,787]" pageId="15" pageNumber="336" pagination="1 - 467" refId="ref23262" refString="Burmeister H. 1855. Handbuch der Entomologie. 4. Band. Besondere Entomologie, Fortsetzung. 2. Abteilung, Coleoptera Lamellicornia Phyllophaga chaenochela. Berlin: Theodor Johann Christian Friedrich Enslin, 1-467." type="book chapter" year="1855">
|
||||
<smallCapsWord id="8D3B4EADFF8B385B393EFD4D93F7FCA7" baselines="780,781" box="[514,657,762,786]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Burmeister" pageId="15" pageNumber="336">BURMEISTER</smallCapsWord>
|
||||
, 1855
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF8B385B3A51FCAF9096FC86" box="[365,496,790,816]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="15" pageNumber="336">
|
||||
<smallCapsWord id="8D3B4EADFF8B385B3A51FCAF90C4FC99" baselines="810,811" box="[365,418,792,816]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="15" pageNumber="336">FIGS</smallCapsWord>
|
||||
3A
|
||||
<emphasis id="B9160463FF8B385B3AF2FCAD90BAFC86" box="[462,476,794,816]" italics="true" pageId="15" pageNumber="336">–</emphasis>
|
||||
D
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF8B385B393EFCA19343FC86" box="[514,549,790,816]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="15" pageNumber="336">5G</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF8B385B3B9AFCFF9067FC0F" pageId="15" pageNumber="336" type="reference_group">
|
||||
<paragraph id="8BDDD871FF8B385B3B9AFCFF9067FC0F" blockId="15.[166,781,840,953]" pageId="15" pageNumber="336">
|
||||
<treatmentCitationGroup id="AB72FF5FFF8B385B3B9AFCFF9067FC0F" pageId="15" pageNumber="336">
|
||||
<treatmentCitation id="0AC3FE60FF8B385B3B9AFCFF93A2FCEB" author="Burmeister H" box="[166,708,840,862]" page="181" pageId="15" pageNumber="336" year="1855">
|
||||
<taxonomicName id="4C62A3F2FF8B385B3B9AFCFF93A2FCEB" ID-CoL="4K83X" authority="Burmeister, 1855: 181" authorityName="Burmeister" authorityPageNumber="181" authorityYear="1855" box="[166,708,840,862]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="ferruginea">
|
||||
<emphasis id="B9160463FF8B385B3B9AFCFF90C2FCEB" box="[166,420,840,861]" italics="true" pageId="15" pageNumber="336">Pleophylla ferruginea</emphasis>
|
||||
<bibRefCitation id="EFF3A580FF8B385B3A88FCFF93A2FCEB" author="Burmeister H" box="[436,708,840,862]" pageId="15" pageNumber="336" pagination="1 - 467" refId="ref23262" refString="Burmeister H. 1855. Handbuch der Entomologie. 4. Band. Besondere Entomologie, Fortsetzung. 2. Abteilung, Coleoptera Lamellicornia Phyllophaga chaenochela. Berlin: Theodor Johann Christian Friedrich Enslin, 1-467." type="book chapter" year="1855">Burmeister, 1855: 181</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
[
|
||||
<typeStatus id="54D966D3FF8B385B39E6FCFE926BFCE8" box="[730,781,841,862]" pageId="15" pageNumber="336">type</typeStatus>
|
||||
locality: Weihnachtsbai [i.e. Durban](Ecklon)];
|
||||
<treatmentCitation id="0AC3FE60FF8B385B3B9AFC3290F8FC2C" author="Schaufuss LW" box="[166,414,901,923]" page="231" pageId="15" pageNumber="336" year="1871">
|
||||
<bibRefCitation id="EFF3A580FF8B385B3B9AFC3290F8FC2C" author="Schaufuss LW" box="[166,414,901,923]" pageId="15" pageNumber="336" pagination="231 - 233" refId="ref24095" refString="Schaufuss LW. 1871. Die Arten der Gattung Pleophylla Er. Nunquam Otiosus I 6: 231-233." type="journal article" year="1871">Schaufuss, 1871: 231</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
; Brenske, 1900: 84; Peringuey, 1904: 8.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF8B385B3B9AFC5592F7FE52" pageId="15" pageNumber="336" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF8B385B3B9AFC559068FBC6" blockId="15.[166,781,993,1136]" pageId="15" pageNumber="336">
|
||||
<emphasis id="B9160463FF8B385B3B9AFC559378FC40" box="[166,542,993,1015]" italics="true" pageId="15" pageNumber="336">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF8B385B3911FC569068FBC6" collectingDate="2014-07-03" collectionCode="MLUH" pageId="15" pageNumber="336" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
|
||||
<typeStatus id="54D966D3FF8B385B3911FC5693FDFC40" box="[557,667,993,1014]" pageId="15" pageNumber="336" type="lectotype">Lectotype</typeStatus>
|
||||
(here designated):
|
||||
<specimenCount id="9D6413F8FF8B385B3A05FBB5902FFBA2" box="[313,329,1026,1044]" count="1" pageId="15" pageNumber="336" type="male">♂</specimenCount>
|
||||
<taxonomicName id="4C62A3F2FF8B385B3A6EFC489358FBA3" authority="Eckl. Pt." authorityName="Eckl. Pt." box="[338,574,1023,1045]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="ferruginea">“ferruginea Eckl. Pt.</taxonomicName>
|
||||
nat. Eck./Martin- Luther Universitat Halle-Wittenberg Zentralmagazin Naturw. Sammlungen 08/03/12
|
||||
<date id="FFDCFEB1FF8B385B3963FB8A9265FBE4" box="[607,771,1085,1107]" pageId="15" pageNumber="336" value="2014-07-03">
|
||||
<collectingDate id="EF980759FF8B385B3963FB8A9265FBE4" box="[607,771,1085,1107]" pageId="15" pageNumber="336" value="2014-07-03">03.Jul. 2014</collectingDate>
|
||||
</date>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF8B385B3B92FBEC9065FBC6" box="[174,259,1115,1136]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="15" pageNumber="336">MLUH</collectionCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8B385B3B9AFB2F93BBFB1B" blockId="15.[166,733,1176,1198]" box="[166,733,1176,1198]" pageId="15" pageNumber="336">
|
||||
<emphasis id="B9160463FF8B385B3B9AFB2F9363FB1B" box="[166,517,1176,1197]" italics="true" pageId="15" pageNumber="336">Additional material examined:</emphasis>
|
||||
See Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8B385B3B9AFB619452FF4D" blockId="15.[166,781,1237,1901]" lastBlockId="15.[829,1444,199,251]" pageId="15" pageNumber="336">
|
||||
<emphasis id="B9160463FF8B385B3B9AFB619049FB5D" box="[166,303,1238,1259]" italics="true" pageId="15" pageNumber="336">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8B385B3ADBFB61932BFB5D" box="[487,589,1238,1259]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="15" pageNumber="336" unit="mm" value="10.3">10.3 mm</quantity>
|
||||
, elytral length: 7.0 mm, maximum width: 5.0 mm. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.76. Pronotum with dense and thick erect setae; unicoloured. Hypomeron not carinate. Basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/ metacoxa: 1/1.43. Metatibia, ratio of width/length: 1/ 4.4. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF8B385B3E16FF7094C6FF6A" box="[1322,1440,199,221]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="15" pageNumber="336">
|
||||
Fig. 3A
|
||||
<emphasis id="B9160463FF8B385B3E43FF7D94EDFF6A" box="[1407,1419,202,220]" italics="true" pageId="15" pageNumber="336">–</emphasis>
|
||||
C
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF8B385B389AFF529566FF4C" box="[934,1024,229,251]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="15" pageNumber="336">Fig. 3D</figureCitation>
|
||||
. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF8B385B3FEBFF529456FF4D" box="[1239,1328,229,251]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="15" pageNumber="336">Fig. 5G</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8B385B3801FEA092F7FE52" blockId="15.[829,1444,279,484]" pageId="15" pageNumber="336">
|
||||
<emphasis id="B9160463FF8B385B3801FEA095AAFE9A" box="[829,1228,279,300]" italics="true" pageId="15" pageNumber="336">
|
||||
Remarks:
|
||||
<taxonomicName id="4C62A3F2FF8B385B388BFEA095AAFE9A" box="[951,1228,279,300]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="transvaalica">Pleophylla transvaalica</taxonomicName>
|
||||
</emphasis>
|
||||
is an invalid ‘in litteris’ name of a specimen without a locality label that is housed in ZMHB (ex coll. Brenske) but originally came from Peringuey (
|
||||
<collectionCode id="ED7340B4FF8B385B3F81FEC4946AFE3E" box="[1213,1292,371,392]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="15" pageNumber="336" type="Museum">SAMC</collectionCode>
|
||||
). In previous publications we referred to this species as ‘sp10’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF8B385B3F06FE079582FE70" author="Ahrens D" box="[1082,1252,432,454]" pageId="15" pageNumber="336" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFF3A580FF8B385B3E3DFE0792E3FE52" author="Eberle J & Warnock RCM & Ahrens D" pageId="15" pageNumber="336" refId="ref23428" refString="Eberle J, Warnock RCM, Ahrens D. 2016 b. Bayesian species delimitation in Pleophylla chafers (Coleoptera) - the importance of prior choice and morphology. BMC Evolutionary Biology 16: 94." type="journal volume" year="2016">
|
||||
Eberle
|
||||
<emphasis id="B9160463FF8B385B3E5FFE0694FBFE73" box="[1379,1437,432,454]" italics="true" pageId="15" pageNumber="336">et al.</emphasis>
|
||||
, 2016b
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
226
data/03/CB/69/03CB6967FF8C385C3BD6FB7095DBFC41.xml
Normal file
226
data/03/CB/69/03CB6967FF8C385C3BD6FB7095DBFC41.xml
Normal file
|
|
@ -0,0 +1,226 @@
|
|||
<document id="ACDFE587110F9E46D9A466049DE17914" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia & Ahrens, Dirk" docDate="2017" docId="03CB6967FF8C385C3BD6FB7095DBFC41" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla pilosa Boheman 1857" docType="treatment" docVersion="2" lastPageNumber="329" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="329" updateTime="1738712560284" updateUser="ExternalLinkService">
|
||||
<mods:mods id="751303A4860C874895D58F50FCA9D1E6" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="F939013B99F318A5CF91AF0028894EB1">
|
||||
<mods:title id="3AE2A5AF413CF63BC8DAD8CCCDCEDEEB">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="8AE72077E18B4528F66880F8586721FC" type="personal">
|
||||
<mods:role id="A906C6F40AE805B35DEBE42EE8765997">
|
||||
<mods:roleTerm id="C29DBAB837B85BAF55C21E92EECD1288">Author</mods:roleTerm>
|
||||
</mods:role>
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<treatment id="03CB6967FF8C385C3BD6FB7095DBFC41" ID-DOI="http://doi.org/10.5281/zenodo.14807089" ID-Zenodo-Dep="14807089" LSID="urn:lsid:plazi:treatment:03CB6967FF8C385C3BD6FB7095DBFC41" httpUri="http://treatment.plazi.org/id/03CB6967FF8C385C3BD6FB7095DBFC41" lastPageNumber="329" pageId="8" pageNumber="329">
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<taxonomicName id="4C62A3F2FF8C385C3BD6FB7093FDFB6A" ID-CoL="4K84B" authority="BOHEMAN, 1857" authorityName="Boheman" authorityYear="1857" box="[234,667,1218,1245]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pilosa">
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<smallCapsWord id="8D3B4EADFF8C385C3AB8FB7D90B7FB6C" baselines="1238" box="[388,465,1226,1242]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="pilosa" pageId="8" pageNumber="329">PILOSA</smallCapsWord>
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<bibRefCitation id="EFF3A580FF8C385C3AE7FB7393FDFB6A" author="Boheman CH" box="[475,667,1218,1245]" pageId="8" pageNumber="329" refId="ref23181" refString="Boheman CH. 1857. Insecta Caffrariae. Annis 1838 - 1845. Part II. Holmiae, 1 - 395." type="book" year="1857">
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<smallCapsWord id="8D3B4EADFF8C385C3AE7FB73932AFB6D" baselines="1238,1239" box="[475,588,1220,1244]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Boheman" pageId="8" pageNumber="329">BOHEMAN</smallCapsWord>
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, 1857
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(
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<figureCitation id="1359C4F4FF8C385C3A6BFB5490BDFB4D" box="[343,475,1249,1275]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="8" pageNumber="329">
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2E
|
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<emphasis id="B9160463FF8C385C3A84FB5290A0FB4D" box="[440,454,1253,1275]" italics="true" pageId="8" pageNumber="329">–</emphasis>
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H
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</figureCitation>
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,
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<figureCitation id="1359C4F4FF8C385C3AD0FB569368FB4D" box="[492,526,1249,1275]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="8" pageNumber="329">5B</figureCitation>
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, 6)
|
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<taxonomicName id="4C62A3F2FF8C385C3BACFAA59328FA9E" ID-CoL="4K84B" authority="Boheman, 1857: 125" authorityName="Boheman" authorityPageNumber="125" authorityYear="1857" box="[144,590,1298,1320]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pilosa">
|
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<emphasis id="B9160463FF8C385C3BACFAA59030FA91" box="[144,342,1298,1319]" italics="true" pageId="8" pageNumber="329">Pleophylla pilosa</emphasis>
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[
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|
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locality:
|
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<collectingRegion id="49A61693FF8C385C3BACFA869194FAF1" box="[144,242,1329,1351]" country="South Africa" name="Limpopo" pageId="8" pageNumber="329">Limpopo</collectingRegion>
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river]
|
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<treatmentCitation id="0AC3FE60FF8C385C3A79FA86935AFAF0" author="Schaufuss LW" box="[325,572,1329,1351]" page="232" pageId="8" pageNumber="329" year="1871">
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<bibRefCitation id="EFF3A580FF8C385C3A79FA86935AFAF0" author="Schaufuss LW" box="[325,572,1329,1351]" pageId="8" pageNumber="329" pagination="231 - 233" refId="ref24095" refString="Schaufuss LW. 1871. Die Arten der Gattung Pleophylla Er. Nunquam Otiosus I 6: 231-233." type="journal article" year="1871">Schaufuss, 1871: 232</bibRefCitation>
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|
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; Brenske, 1900: 84; Peringuey, 1904: 8.
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</paragraph>
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<treatmentCitation id="0AC3FE60FF8C385C3BADFAD9931EFA35" author="Schaufuss LW" box="[145,632,1390,1412]" page="232" pageId="8" pageNumber="329" year="1871">
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<taxonomicName id="4C62A3F2FF8C385C3BADFAD9931EFA35" ID-CoL="6VQMY" authority="Schaufuss, 1871: 232" authorityName="Schaufuss" authorityPageNumber="232" authorityYear="1871" box="[145,632,1390,1412]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="opalina">
|
||||
<emphasis id="B9160463FF8C385C3BADFAD9900BFA35" box="[145,365,1390,1411]" italics="true" pageId="8" pageNumber="329">Pleophylla opalina</emphasis>
|
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<bibRefCitation id="EFF3A580FF8C385C3A47FAD9931EFA35" author="Schaufuss LW" box="[379,632,1390,1412]" pageId="8" pageNumber="329" pagination="231 - 233" refId="ref24095" refString="Schaufuss LW. 1871. Die Arten der Gattung Pleophylla Er. Nunquam Otiosus I 6: 231-233." type="journal article" year="1871">Schaufuss, 1871: 232</bibRefCitation>
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|
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|
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[Caffraria interia]; Brenske, 1900: 84, Peringuey, 1904: 8;
|
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<emphasis id="B9160463FF8C385C39FFFA3A91A3FA76" bold="true" pageId="8" pageNumber="329">syn. nov.</emphasis>
|
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|
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|
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<paragraph id="8BDDD871FF8C385C3BACFA5E9084F965" blockId="8.[144,759,1512,1748]" pageId="8" pageNumber="329">
|
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<emphasis id="B9160463FF8C385C3BACFA5E90CAFA4B" box="[144,428,1512,1534]" italics="true" pageId="8" pageNumber="329">Type material examined:</emphasis>
|
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<materialsCitation id="3B0AD22CFF8C385C3A87FA5E9395F9EF" collectionCode="NHRS" collectorName="J. Wahlb." location="Stockholm" pageId="8" pageNumber="329" specimenCount="1" specimenCount-male="1" stateProvince="Stockholm" typeStatus="lectotype">
|
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<typeStatus id="54D966D3FF8C385C3A87FA5E934EFA48" box="[443,552,1513,1534]" pageId="8" pageNumber="329" type="lectotype">Lectotype</typeStatus>
|
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(here designated):
|
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<specimenCount id="9D6413F8FF8C385C3BACF9BD91C6F9AA" box="[144,160,1546,1564]" count="1" pageId="8" pageNumber="329" type="male">♂</specimenCount>
|
||||
“Caffraria./
|
||||
<collectorName id="2697BDA7FF8C385C3A12F9B090F0F9AB" box="[302,406,1543,1565]" pageId="8" pageNumber="329">J. Wahlb.</collectorName>
|
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/Type./
|
||||
<emphasis id="B9160463FF8C385C3ADEF9B093D5F9AA" box="[482,691,1543,1564]" italics="true" pageId="8" pageNumber="329">
|
||||
<taxonomicName id="4C62A3F2FF8C385C3ADEF9B093C0F9AA" authorityName="Boheman" authorityYear="1857" box="[482,678,1543,1564]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pilosa">Pleophylla pilosa</taxonomicName>
|
||||
/
|
||||
</emphasis>
|
||||
<taxonomicName id="4C62A3F2FF8C385C398FF9B090D0F98D" authority="Bhn. Ent. Kaffr." class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B9160463FF8C385C398FF9B09390F9AA" box="[691,758,1543,1564]" italics="true" pageId="8" pageNumber="329">pilosa</emphasis>
|
||||
Bhn. Ent. Kaffr.
|
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</taxonomicName>
|
||||
/3680E91 + /Naturhistoriska Riksmuseet
|
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<collectingRegion id="49A61693FF8C385C3A18F9F390FCF9EC" box="[292,410,1604,1626]" country="Sweden" name="Stockholm" pageId="8" pageNumber="329">Stockholm</collectingRegion>
|
||||
Loan no. 1746/07” (
|
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<collectionCode id="ED7340B4FF8C385C39A3F9F3938AF9EF" box="[671,748,1604,1625]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="8" pageNumber="329">NHRS</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B0AD22CFF8C385C3BACF9D490B8F965" collectionCode="ZMHB" collectorName="Coll. L. W. Schaufuss" pageId="8" pageNumber="329" specimenCount="1" specimenCount-male="1" stateProvince="Stockholm" typeStatus="lectotype">
|
||||
<typeStatus id="54D966D3FF8C385C3BACF9D4919BF9CE" box="[144,253,1635,1656]" pageId="8" pageNumber="329" type="lectotype">Lectotype</typeStatus>
|
||||
(
|
||||
<taxonomicName id="4C62A3F2FF8C385C3A2FF9D49001F9CE" authorityName="Schaufuss" authorityYear="1871" box="[275,359,1635,1656]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="opalina">
|
||||
<emphasis id="B9160463FF8C385C3A2FF9D49001F9CE" box="[275,359,1635,1656]" italics="true" pageId="8" pageNumber="329">opalina</emphasis>
|
||||
</taxonomicName>
|
||||
, here designated):
|
||||
<specimenCount id="9D6413F8FF8C385C396AF9D29300F9C1" box="[598,614,1637,1655]" count="1" pageId="8" pageNumber="329" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF8C385C3943F9D49067F921" pageId="8" pageNumber="329">Coll. L.W. Schaufuss</collectorName>
|
||||
/Nunqn at pag. 232/
|
||||
<taxonomicName id="4C62A3F2FF8C385C3ACAF9369391F921" authority="Schauf. Caffr." authorityName="Schauf. Caffr." box="[502,759,1665,1687]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="opalina">opalina Schauf. Caffr.</taxonomicName>
|
||||
inter./Type./
|
||||
<taxonomicName id="4C62A3F2FF8C385C3A24F9179089F903" box="[280,495,1696,1717]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="opalina">
|
||||
<emphasis id="B9160463FF8C385C3A24F9179089F903" box="[280,495,1696,1717]" italics="true" pageId="8" pageNumber="329">Pleophylla opalina</emphasis>
|
||||
</taxonomicName>
|
||||
m.
|
||||
<taxonomicName id="4C62A3F2FF8C385C3919F9179391F903" authority="Burm." authorityName="Burmeister" authorityYear="1855" box="[549,759,1696,1718]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="navicularis">navicularis Burm.</taxonomicName>
|
||||
[sic! canceled] Natal” (
|
||||
<collectionCode id="ED7340B4FF8C385C3ABAF90990B1F965" box="[390,471,1726,1747]" pageId="8" pageNumber="329">ZMHB</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8C385C3BACF94C93A0F8A7" blockId="8.[144,710,1787,1809]" box="[144,710,1787,1809]" pageId="8" pageNumber="329">
|
||||
<emphasis id="B9160463FF8C385C3BACF94C9089F8A6" box="[144,495,1787,1808]" italics="true" pageId="8" pageNumber="329">Additional material examined:</emphasis>
|
||||
See Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8C385C3BACF88F9514FC89" blockId="8.[144,759,1848,1901]" lastBlockId="8.[807,1422,199,832]" pageId="8" pageNumber="329">
|
||||
<emphasis id="B9160463FF8C385C3BACF88F9057F8FB" box="[144,305,1848,1869]" italics="true" pageId="8" pageNumber="329">Redescription:</emphasis>
|
||||
Body length: 9.0 mm, elytral length:
|
||||
<quantity id="4C9A7594FF8C385C3BACF8E0918FF8DB" box="[144,233,1879,1901]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="8" pageNumber="329" unit="mm" value="6.1">6.1 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF8C385C3AE4F8E09355F8DB" box="[472,563,1879,1901]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="8" pageNumber="329" unit="mm" value="4.7">4.7 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.68. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum moderately large. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.33. Metatibia, ratio of width/length: 1/3.87. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF8C385C381BFC9D92FBFC89" box="[807,925,810,832]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="8" pageNumber="329">
|
||||
Fig. 2E
|
||||
<emphasis id="B9160463FF8C385C3847FC9A92E1FC89" box="[891,903,813,831]" italics="true" pageId="8" pageNumber="329">–</emphasis>
|
||||
G
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF8C385C3F28FC9D9509FC89" box="[1044,1135,810,832]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="8" pageNumber="329">Fig. 2H</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8C385C381BFCD095DBFC41" blockId="8.[807,1421,871,1015]" pageId="8" pageNumber="329">
|
||||
<emphasis id="B9160463FF8C385C381BFCD092F4FCCA" box="[807,914,871,892]" italics="true" pageId="8" pageNumber="329">Remarks:</emphasis>
|
||||
The shape of the parameres of
|
||||
<taxonomicName id="4C62A3F2FF8C385C3E2BFCDF94EBFCCA" box="[1303,1421,871,893]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="opalina">
|
||||
<emphasis id="B9160463FF8C385C3E2BFCDF94EBFCCA" box="[1303,1421,871,893]" italics="true" pageId="8" pageNumber="329">P. opalina</emphasis>
|
||||
</taxonomicName>
|
||||
are virtually identical in shape to those of the
|
||||
<typeStatus id="54D966D3FF8C385C381BFC1392EBFC0C" box="[807,909,932,954]" pageId="8" pageNumber="329" type="lectotype">lectotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C62A3F2FF8C385C38FCFC129542FC0F" box="[960,1060,932,954]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B9160463FF8C385C38FCFC129542FC0F" box="[960,1060,932,954]" italics="true" pageId="8" pageNumber="329">P. pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
. In previous publications we referred to this species as ‘sp6’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF8C385C381BFC5692A7FC41" author="Ahrens D" box="[807,961,993,1015]" pageId="8" pageNumber="329" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFF3A580FF8C385C38F1FC5695D7FC41" author="Eberle J & Warnock RCM & Ahrens D" box="[973,1201,993,1015]" pageId="8" pageNumber="329" refId="ref23428" refString="Eberle J, Warnock RCM, Ahrens D. 2016 b. Bayesian species delimitation in Pleophylla chafers (Coleoptera) - the importance of prior choice and morphology. BMC Evolutionary Biology 16: 94." type="journal volume" year="2016">
|
||||
Eberle
|
||||
<emphasis id="B9160463FF8C385C3F1CFC55953CFC40" box="[1056,1114,993,1015]" italics="true" pageId="8" pageNumber="329">et al.</emphasis>
|
||||
, 2016b
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
346
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346
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<taxonomicName id="4C62A3F2FF8C385C3803FB99957DFBD7" ID-CoL="4K84C" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pseudopilosa" status="sp. nov.">
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,
|
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<smallCapsWord id="8D3B4EADFF8C385C3E3AFB9C9408FBF4" baselines="1085,1086" box="[1286,1390,1067,1091]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="8" pageNumber="329">BECKETT</smallCapsWord>
|
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,
|
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|
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&
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</taxonomicName>
|
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|
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|
||||
.
|
||||
<smallCapsWord id="8D3B4EADFF8C385C3F6EFBF895E4FBD6" baselines="1116" box="[1106,1154,1103,1120]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="8" pageNumber="329">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF8C385C3FA1FBFD9473FBD4" box="[1181,1301,1096,1122]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="8" pageNumber="329">
|
||||
<smallCapsWord id="8D3B4EADFF8C385C3FA1FBFD95B4FBD7" baselines="1116,1117" box="[1181,1234,1098,1122]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="8" pageNumber="329">FIGS</smallCapsWord>
|
||||
2I
|
||||
<emphasis id="B9160463FF8C385C3FCAFBFB9462FBD4" box="[1270,1284,1100,1122]" italics="true" pageId="8" pageNumber="329">–</emphasis>
|
||||
L
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF8C385C3E1AFBFF942EFBD4" box="[1318,1352,1096,1122]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="8" pageNumber="329">5C</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8C385C3816FB399286FB77" blockId="8.[810,1428,1165,1217]" pageId="8" pageNumber="329">
|
||||
<uri id="FFF3D473FF8C385C3816FB399286FB77" pageId="8" pageNumber="329">
|
||||
<emphasis id="B9160463FF8C385C3816FB39923CFB15" box="[810,858,1166,1187]" italics="true" pageId="8" pageNumber="329">urn:</emphasis>
|
||||
lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF8C385C3F02FB3A9286FB77" pageId="8" pageNumber="329">877B006F-29D1-44E7-AB0D- FB3F8189AE48</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF8C385E381BFB0090E4FBCF" lastPageId="10" lastPageNumber="331" pageId="8" pageNumber="329" type="description">
|
||||
<paragraph id="8BDDD871FF8C385C381BFB0094EBFB7A" box="[807,1421,1207,1229]" pageId="8" pageNumber="329">
|
||||
X-DA0394
|
||||
<collectingCountry id="F37598E1FF8C385C38A1FB009541FB7B" box="[925,1063,1207,1229]" name="South Africa" pageId="8" pageNumber="329">South Africa</collectingCountry>
|
||||
Cheerio Farm, Haenertsburg, ca
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8C385C381BFB6E940CFADE" blockId="8.[807,1421,1241,1385]" pageId="8" pageNumber="329">
|
||||
<emphasis id="B9160463FF8C385C381BFB6E9509FB58" box="[807,1135,1241,1262]" italics="true" pageId="8" pageNumber="329">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF8C385C3F42FB6E9548FADE" collectingDate="2006-11-20" collectingDateMax="2006-11-22" collectingDateMin="2006-11-20" collectionCode="TMSA" collectorName="D. Ahrens & S. Fabrizi" country="South Africa" elevation="1492" latitude="-53.01186" location="Cheerio Farm" longLatPrecision="1" longitude="57.002693" municipality="Haenertsburg" pageId="8" pageNumber="329" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF8C385C3F42FB6E9580FB59" box="[1150,1254,1241,1263]" pageId="8" pageNumber="329" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF8C385C3E3AFB6C9470FB5B" box="[1286,1302,1243,1261]" count="1" pageId="8" pageNumber="329" type="male">♂</specimenCount>
|
||||
“834699 X-DA0394
|
||||
<collectingCountry id="F37598E1FF8C385C38A1FB409541FABB" box="[925,1063,1271,1293]" name="South Africa" pageId="8" pageNumber="329">South Africa</collectingCountry>
|
||||
<location id="8EBD8EAAFF8C385C3F10FB4095A0FABB" LSID="urn:lsid:plazi:treatment:03CB6967FF8C385E3803FB9990E4FBCF:8EBD8EAAFF8C385C3F10FB4095A0FABB" box="[1068,1222,1271,1293]" country="South Africa" latitude="-53.01186" longLatPrecision="1" longitude="57.002693" municipality="Haenertsburg" name="Cheerio Farm" pageId="8" pageNumber="329">Cheerio Farm</location>
|
||||
,
|
||||
<collectingMunicipality id="6BB9420BFF8C385C3FF3FB40940AFABB" box="[1231,1388,1271,1293]" pageId="8" pageNumber="329">Haenertsburg</collectingMunicipality>
|
||||
, ca
|
||||
<locationDeviation id="9A6FC153FF8C385C381BFAA19569FA9D" box="[807,1039,1302,1324]" location="Tzaneen" pageId="8" pageNumber="329">
|
||||
<quantity id="4C9A7594FF8C385C381BFAA19208FA9A" box="[807,878,1302,1324]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="329" unit="km" value="20.0">20 km</quantity>
|
||||
W of Tzaneen
|
||||
</locationDeviation>
|
||||
,
|
||||
<quantity id="4C9A7594FF8C385C3F26FAA19508FA9D" box="[1050,1134,1302,1323]" metricMagnitude="3" metricUnit="m" metricValue="1.4919999999999998" pageId="8" pageNumber="329" unit="m" value="1492.0">
|
||||
<elevation id="004F3F42FF8C385C3F26FAA19508FA9D" box="[1050,1134,1302,1323]" metricMagnitude="3" metricUnit="m" metricValue="1.4919999999999998" pageId="8" pageNumber="329" unit="m" value="1492.0">1492 m</elevation>
|
||||
</quantity>
|
||||
023°
|
||||
<geoCoordinate id="EE56BEB6FF8C385C3F99FAA19475FA9D" box="[1189,1299,1297,1324]" degrees="53" direction="south" minutes="0" orientation="latitude" pageId="8" pageNumber="329" precision="1" seconds="42, 7" value="-53.01186">
|
||||
53
|
||||
<emphasis id="B9160463FF8C385C3F83FAA695A2FA96" bold="true" box="[1215,1220,1297,1312]" italics="true" pageId="8" pageNumber="329">
|
||||
<superScript id="7C177539FF8C385C3F83FAA695A2FA96" attach="none" box="[1215,1220,1297,1312]" fontSize="6" pageId="8" pageNumber="329">0</superScript>
|
||||
</emphasis>
|
||||
42, 7
|
||||
<emphasis id="B9160463FF8C385C3FC7FAA19463FA9D" bold="true" box="[1275,1285,1302,1323]" italics="true" pageId="8" pageNumber="329">″</emphasis>
|
||||
S
|
||||
</geoCoordinate>
|
||||
029°
|
||||
<geoCoordinate id="EE56BEB6FF8C385C3E77FAA29229FAFC" degrees="57" direction="east" minutes="0" orientation="longitude" pageId="8" pageNumber="329" precision="1" seconds="09, 7" value="57.002693">
|
||||
57
|
||||
<emphasis id="B9160463FF8C385C3E5AFAA6940DFA96" bold="true" box="[1382,1387,1297,1312]" italics="true" pageId="8" pageNumber="329">
|
||||
<superScript id="7C177539FF8C385C3E5AFAA6940DFA96" attach="none" box="[1382,1387,1297,1312]" fontSize="6" pageId="8" pageNumber="329">0</superScript>
|
||||
</emphasis>
|
||||
09, 7
|
||||
<emphasis id="B9160463FF8C385C3808FA839258FAFF" bold="true" box="[820,830,1332,1353]" italics="true" pageId="8" pageNumber="329">″</emphasis>
|
||||
E
|
||||
</geoCoordinate>
|
||||
<date id="FFDCFEB1FF8C385C3858FA83929BFAFC" box="[868,1021,1332,1354]" pageId="8" pageNumber="329" value="2006-11-20" valueMax="2006-11-22" valueMin="2006-11-20">
|
||||
<collectingDate id="EF980759FF8C385C3858FA83929BFAFC" box="[868,1021,1332,1354]" pageId="8" pageNumber="329" value="2006-11-20" valueMax="2006-11-22" valueMin="2006-11-20">20-22.XI.2006</collectingDate>
|
||||
</date>
|
||||
leg.
|
||||
<collectorName id="2697BDA7FF8C385C3F73FA8295A9FAFC" box="[1103,1231,1332,1354]" pageId="8" pageNumber="329">D. Ahrens</collectorName>
|
||||
&
|
||||
<collectorName id="2697BDA7FF8C385C3E30FA8394EFFAFC" box="[1292,1417,1332,1354]" pageId="8" pageNumber="329">S. Fabrizi</collectorName>
|
||||
,
|
||||
<taxonomicName id="4C62A3F2FF8C385C381BFAE492A0FADE" box="[807,966,1363,1384]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="undefined-1">
|
||||
<emphasis id="B9160463FF8C385C381BFAE492FDFADE" box="[807,923,1363,1384]" italics="true" pageId="8" pageNumber="329">Pleophylla</emphasis>
|
||||
sp1
|
||||
</taxonomicName>
|
||||
<emphasis id="B9160463FF8C385C38F9FAE492A9FADE" bold="true" box="[965,975,1363,1384]" italics="true" pageId="8" pageNumber="329">″</emphasis>
|
||||
(
|
||||
<collectionCode id="ED7340B4FF8C385C38E7FAE49541FADE" box="[987,1063,1363,1384]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="8" pageNumber="329">TMSA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<typeStatus id="54D966D3FF8C385C3F0AFAE495CFFADE" box="[1078,1193,1363,1384]" pageId="8" pageNumber="329" type="paratype">Paratypes</typeStatus>
|
||||
: see Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8C385E381BFA2793E6FE2F" blockId="8.[807,1422,1424,1904]" lastBlockId="10.[144,762,203,409]" lastPageId="10" lastPageNumber="331" pageId="8" pageNumber="329">
|
||||
<emphasis id="B9160463FF8C385C381BFA2792C9FA13" box="[807,943,1424,1445]" italics="true" pageId="8" pageNumber="329">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8C385C3F52FA2795A1FA13" box="[1134,1223,1424,1445]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="8" pageNumber="329" unit="mm" value="8.3">8.3 mm</quantity>
|
||||
, elytral length: 6.0 mm, maximum width:
|
||||
<quantity id="4C9A7594FF8C385C3F53FA1995AFFA72" box="[1135,1225,1454,1476]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="329" unit="mm" value="4.6">4.6 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.65. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/ metacoxa: 1/1.34. Metatibia, ratio of width/length: 1/4. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF8E385E39BFFED39394FECC" box="[643,754,356,378]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="10" pageNumber="331">
|
||||
Fig. 2I
|
||||
<emphasis id="B9160463FF8E385E39ECFED093BAFECF" box="[720,732,359,377]" italics="true" pageId="10" pageNumber="331">–</emphasis>
|
||||
K
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF8E385E3BC4FE349029FE2E" box="[248,335,387,409]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="10" pageNumber="331">
|
||||
Fig.
|
||||
<specimenCount id="9D6413F8FF8E385E3A11FE349029FE2E" box="[301,335,387,408]" count="2" pageId="10" pageNumber="331" type="generic">2L</specimenCount>
|
||||
</figureCitation>
|
||||
. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF8E385E3918FE34931AFE2F" box="[548,636,387,409]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="10" pageNumber="331">Fig. 5C</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF1D88F9FF8D385D3B9AF93B9569F884" ID-DOI="http://doi.org/10.5281/zenodo.14807093" ID-Zenodo-Dep="14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="9" pageNumber="330" startId="9.[166,245,1676,1695]" targetBox="[189,1421,218,1624]" targetPageId="9" targetType="figure">
|
||||
<paragraph id="8BDDD871FF8D385D3B9AF93B9569F884" blockId="9.[166,1444,1676,1842]" pageId="9" pageNumber="330">
|
||||
<emphasis id="B9160463FF8D385D3B9AF93B9076F929" bold="true" box="[166,272,1676,1695]" pageId="9" pageNumber="330">Figure 4.</emphasis>
|
||||
A
|
||||
<emphasis id="B9160463FF8D385D3A11F938905EF929" box="[301,312,1679,1695]" italics="true" pageId="9" pageNumber="330">–</emphasis>
|
||||
D,
|
||||
<emphasis id="B9160463FF8D385D3A60F93B938FF916" bold="true" box="[348,745,1676,1696]" pageId="9" pageNumber="330">
|
||||
<taxonomicName id="4C62A3F2FF8D385D3A60F93B93E6F929" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[348,640,1676,1695]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="mpumalanga" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8D385D3A60F93B93E6F929" bold="true" box="[348,640,1676,1695]" italics="true" pageId="9" pageNumber="330">Pleophylla mpumalanga</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8D385D39B7F93A938FF916" box="[651,745,1677,1696]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype); E
|
||||
<emphasis id="B9160463FF8D385D3843F93892ECF929" box="[895,906,1679,1695]" italics="true" pageId="9" pageNumber="330">–</emphasis>
|
||||
H,
|
||||
<emphasis id="B9160463FF8D385D3893F93B9592F916" bold="true" box="[943,1268,1676,1696]" pageId="9" pageNumber="330">
|
||||
<taxonomicName id="4C62A3F2FF8D385D3893F93B95EDF929" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[943,1163,1676,1695]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8D385D3893F93B95EDF929" bold="true" box="[943,1163,1676,1695]" italics="true" pageId="9" pageNumber="330">Pleophylla charlyi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8D385D3FABF93A9592F916" box="[1175,1268,1677,1696]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype); I
|
||||
<emphasis id="B9160463FF8D385D3EBFF93894E8F929" box="[1411,1422,1679,1695]" italics="true" pageId="9" pageNumber="330">–</emphasis>
|
||||
L,
|
||||
<emphasis id="B9160463FF8D385D3B9AF91E90BEF90B" bold="true" box="[166,472,1705,1725]" pageId="9" pageNumber="330">
|
||||
<taxonomicName id="4C62A3F2FF8D385D3B9AF91E9014F90A" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[166,370,1705,1724]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="lizleri" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8D385D3B9AF91E9014F90A" bold="true" box="[166,370,1705,1724]" italics="true" pageId="9" pageNumber="330">Pleophylla lizleri</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8D385D3A40F91D90BEF90B" box="[380,472,1706,1725]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype); M
|
||||
<emphasis id="B9160463FF8D385D3953F91B931CF90A" box="[623,634,1708,1724]" italics="true" pageId="9" pageNumber="330">–</emphasis>
|
||||
P,
|
||||
<emphasis id="B9160463FF8D385D39A5F91E9291F90B" bold="true" box="[665,1015,1705,1725]" pageId="9" pageNumber="330">
|
||||
<taxonomicName id="4C62A3F2FF8D385D39A5F91E92F4F90B" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[665,914,1705,1725]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="taitaensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8D385D39A5F91E92F4F90B" bold="true" box="[665,914,1705,1725]" italics="true" pageId="9" pageNumber="330">Pleophylla taitaensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8D385D38A7F91D9291F90B" box="[923,1015,1706,1725]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype); Q
|
||||
<emphasis id="B9160463FF8D385D3FB6F91B95F3F90A" box="[1162,1173,1708,1724]" italics="true" pageId="9" pageNumber="330">–</emphasis>
|
||||
T,
|
||||
<emphasis id="B9160463FF8D385D3F88F91E91BEF96C" bold="true" pageId="9" pageNumber="330">
|
||||
<taxonomicName id="4C62A3F2FF8D385D3F88F91E941FF90A" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1204,1401,1705,1724]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="stalsi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8D385D3F88F91E941FF90A" bold="true" box="[1204,1401,1705,1724]" italics="true" pageId="9" pageNumber="330">Pleophylla stalsi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8D385D3EBFF91D91BEF96C" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype); U
|
||||
<emphasis id="B9160463FF8D385D3A52F97E901FF96F" box="[366,377,1737,1753]" italics="true" pageId="9" pageNumber="330">–</emphasis>
|
||||
X,
|
||||
<emphasis id="B9160463FF8D385D3AA6F970920EF96C" bold="true" box="[410,872,1735,1754]" pageId="9" pageNumber="330">
|
||||
<taxonomicName id="4C62A3F2FF8D385D3AA6F9709266F96C" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[410,768,1735,1754]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="wakkerstroomensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8D385D3AA6F9709266F96C" bold="true" box="[410,768,1735,1754]" italics="true" pageId="9" pageNumber="330">Pleophylla wakkerstroomensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8D385D3837F970920EF96C" box="[779,872,1735,1754]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype); Y
|
||||
<emphasis id="B9160463FF8D385D38C7F97E9560F96F" box="[1019,1030,1737,1753]" italics="true" pageId="9" pageNumber="330">–</emphasis>
|
||||
Ab,
|
||||
<taxonomicName id="4C62A3F2FF8D385D3F08F97094C2F96C" authority="Peringuey" authorityName="Peringuey" authorityYear="1904" box="[1076,1444,1735,1754]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="tongaatsana">
|
||||
<emphasis id="B9160463FF8D385D3F08F9709448F96C" box="[1076,1326,1735,1754]" italics="true" pageId="9" pageNumber="330">Pleophylla tongaatsana</emphasis>
|
||||
Peringuey
|
||||
</taxonomicName>
|
||||
(RSA, Sjonajona); Ac
|
||||
<emphasis id="B9160463FF8D385D3AB9F95090F6F941" box="[389,400,1767,1783]" italics="true" pageId="9" pageNumber="330">–</emphasis>
|
||||
Af,
|
||||
<emphasis id="B9160463FF8D385D3A84F9539261F941" bold="true" box="[440,775,1764,1783]" pageId="9" pageNumber="330">
|
||||
<taxonomicName id="4C62A3F2FF8D385D3A84F95393C4F941" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[440,674,1764,1783]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="kruegeri" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8D385D3A84F95393C4F941" bold="true" box="[440,674,1764,1783]" italics="true" pageId="9" pageNumber="330">Pleophylla kruegeri</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8D385D3997F9539261F941" box="[683,775,1764,1783]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BDDD871FF8E385E3BADFE1993F8FD8B" blockId="10.[145,760,430,574]" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E3BADFE199356FE75" box="[145,560,430,451]" italics="true" pageId="10" pageNumber="331">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF8E385E3A2BFE199356FE75" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[279,560,430,451]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="pseudopilosa" status="sp. nov.">Pleophylla pseudopilosa</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF8E385E397CFE1993C6FE75" box="[576,672,430,451]" pageId="10" pageNumber="331" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from
|
||||
<taxonomicName id="4C62A3F2FF8E385E3BE6FE7A9059FE57" box="[218,319,460,482]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B9160463FF8E385E3BE6FE7A9059FE57" box="[218,319,460,482]" italics="true" pageId="10" pageNumber="331">P. pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
in the parameres by having the ventral lobe in lateral view wider and shorter; the base of the dorsal lobe of left paramere has a single narrow medial tooth (not two as in
|
||||
<taxonomicName id="4C62A3F2FF8E385E3916FD9F93E8FD8B" box="[554,654,552,573]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B9160463FF8E385E3916FD9F93E8FD8B" box="[554,654,552,573]" italics="true" pageId="10" pageNumber="331">P. pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8E385E3BADFDEF938BFCD4" blockId="10.[145,760,599,866]" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E3BADFDEF9063FDDB" box="[145,261,600,621]" italics="true" pageId="10" pageNumber="331">Variation:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8E385E3A8BFDE09359FDDB" box="[439,575,599,621]" metricMagnitude="-3" metricUnit="m" metricValue="7.65" metricValueMax="8.5" metricValueMin="6.8" pageId="10" pageNumber="331" unit="mm" value="7.65" valueMax="8.5" valueMin="6.8">
|
||||
6.8
|
||||
<emphasis id="B9160463FF8E385E3AE5FDED9083FDDA" box="[473,485,602,620]" italics="true" pageId="10" pageNumber="331">–</emphasis>
|
||||
8.5 mm
|
||||
</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF8E385E3BADFDC1907FFD3A" box="[145,281,630,652]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="6.1" metricValueMin="4.2" pageId="10" pageNumber="331" unit="mm" value="5.15" valueMax="6.1" valueMin="4.2">
|
||||
4.2
|
||||
<emphasis id="B9160463FF8E385E3B8FFDCE91D9FD3D" box="[179,191,633,651]" italics="true" pageId="10" pageNumber="331">–</emphasis>
|
||||
6.1 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF8E385E3932FDC193F0FD3A" box="[526,662,630,652]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.8" metricValueMin="3.8" pageId="10" pageNumber="331" unit="mm" value="4.3" valueMax="4.8" valueMin="3.8">
|
||||
3.8
|
||||
<emphasis id="B9160463FF8E385E390CFDCE935AFD3D" box="[560,572,633,651]" italics="true" pageId="10" pageNumber="331">–</emphasis>
|
||||
4.8 mm
|
||||
</quantity>
|
||||
. Colour varies from reddish brown with dark spot to entirely yellowish brown without spots. Female: antennal club of female straight, composed of five antennomeres, slightly longer than the remaining antennomeres combined; sixth antennomere subequal a quarter of club length. Eyes smaller than in male: ratio of eye diameter/interocular width: 0.5.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8E385E3BAEFCCD9086FBBC" blockId="10.[146,761,890,1034]" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E3BAEFCCD9076FC39" box="[146,272,890,911]" italics="true" pageId="10" pageNumber="331">Etymology:</emphasis>
|
||||
The name of the new species is composed of the Latinized Greek prefix pseudo- (false) and pilosa (species name of
|
||||
<taxonomicName id="4C62A3F2FF8E385E3910FC009394FC7A" box="[556,754,951,972]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B9160463FF8E385E3910FC009394FC7A" box="[556,754,951,972]" italics="true" pageId="10" pageNumber="331">Pleophylla pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
, with reference to its general similarity with this species) (noun in apposition).
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8E385E3BAEFB9190E4FBCF" blockId="10.[146,761,1062,1145]" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E3BAEFB919198FB8D" box="[146,254,1062,1083]" italics="true" pageId="10" pageNumber="331">Remarks:</emphasis>
|
||||
In previous publications we referred to this species as ‘sp1
|
||||
<emphasis id="B9160463FF8E385E3A51FBF79014FBF9" bold="true" box="[365,370,1088,1103]" italics="true" pageId="10" pageNumber="331">
|
||||
<superScript id="7C177539FF8E385E3A51FBF79014FBF9" attach="left" box="[365,370,1088,1103]" fontSize="6" pageId="10" pageNumber="331">0</superScript>
|
||||
</emphasis>
|
||||
(Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF8E385E3966FBF39393FBEC" author="Ahrens D" box="[602,757,1092,1114]" pageId="10" pageNumber="331" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFF3A580FF8E385E3BAEFBD49010FBCF" author="Eberle J & Warnock RCM & Ahrens D" box="[146,374,1123,1145]" pageId="10" pageNumber="331" refId="ref23428" refString="Eberle J, Warnock RCM, Ahrens D. 2016 b. Bayesian species delimitation in Pleophylla chafers (Coleoptera) - the importance of prior choice and morphology. BMC Evolutionary Biology 16: 94." type="journal volume" year="2016">
|
||||
Eberle
|
||||
<emphasis id="B9160463FF8E385E3BD9FBD39079FBCE" box="[229,287,1123,1145]" italics="true" pageId="10" pageNumber="331">et al.</emphasis>
|
||||
, 2016b
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
296
data/03/CB/69/03CB6967FF8E38583805FB2E906BFD5A.xml
Normal file
296
data/03/CB/69/03CB6967FF8E38583805FB2E906BFD5A.xml
Normal file
|
|
@ -0,0 +1,296 @@
|
|||
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<mods:titleInfo id="71DE468F5EFD427E6D7B111821A31B2A">
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<mods:title id="0887F557ACB5BDDF72C8A90AFE460AE4">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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<mods:namePart id="72CCBA9C4BE30A47F9A8393E4D1CF5C9">Eberle, Jonas</mods:namePart>
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<mods:namePart id="B0FD7627746D9C98E0BD0D46ECD0CFCD">Beckett, Marc</mods:namePart>
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<mods:namePart id="F7B6111B6107556DF2B761126D09A537">Ozgul-Siemund, Alev</mods:namePart>
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<mods:namePart id="1286D95E6A12AADB85BAAFDDFEF45C72">Fabrizi, Silvia</mods:namePart>
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<mods:number id="ED9897804425B387EF24592869C574E4">180</mods:number>
|
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</mods:detail>
|
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<mods:detail id="DFB8AC98C90D4F8CB937544E79E1EA06" type="issue">
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<mods:number id="0569B8E221710B8CF5942D06E105F233">2</mods:number>
|
||||
</mods:detail>
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<mods:start id="BD8AEE91874B78057AADA22FDE18D89B">321</mods:start>
|
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<mods:end id="4C30788A75234C3A0EF4D7CC96A1A3AC">353</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location id="60AF0DF833E427F6162012F5273E8ACC">
|
||||
<mods:url id="DBCF067C23BB838C3CF3FE36493DD00B">https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="FE0C3CF4C4AFC07FA2050E2F66D49032">journal article</mods:classification>
|
||||
<mods:identifier id="0BCBBB9D3EB835ECEB3C439E27B8B734" type="DOI">10.1111/zoj.12489</mods:identifier>
|
||||
<mods:identifier id="258A20BAD3C34CFCC6CEFAF832723F5F" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="8EF10D6D89A67CCF974953501622CFD6" type="Zenodo-Dep">14807083</mods:identifier>
|
||||
<mods:identifier id="52E8C2975F3FB1A5B1746C9B696D0F35" type="ZooBank">3663E0A-2E17-4346-A388-6BEE53081814</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03CB6967FF8E38583805FB2E906BFD5A" LSID="urn:lsid:plazi:treatment:03CB6967FF8E38583805FB2E906BFD5A" httpUri="http://treatment.plazi.org/id/03CB6967FF8E38583805FB2E906BFD5A" lastPageId="12" lastPageNumber="333" pageId="10" pageNumber="331">
|
||||
<subSubSection id="C3788BFAFF8E385E3805FB2E92B6FAA9" pageId="10" pageNumber="331" type="nomenclature">
|
||||
<paragraph id="8BDDD871FF8E385E3805FB2E9439FB78" blockId="10.[825,1405,1174,1231]" pageId="10" pageNumber="331">
|
||||
<heading id="D0956F1DFF8E385E3805FB2E9439FB78" centered="true" fontSize="9" level="2" pageId="10" pageNumber="331" reason="2">
|
||||
<taxonomicName id="4C62A3F2FF8E385E3805FB2E9554FB7A" ID-CoL="4K845" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="mlilwaneensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8E385E3805FB2E95FEFB18" bold="true" box="[825,1176,1177,1198]" italics="true" pageId="10" pageNumber="331">
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3805FB2E92B5FB18" baselines="1193,1194" box="[825,979,1177,1198]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="10" pageNumber="331">PLEOPHYLLA</smallCapsWord>
|
||||
<smallCapsWord id="8D3B4EADFF8E385E38E0FB2A95FEFB18" baselines="1194" box="[988,1176,1181,1198]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="mlilwaneensis" pageId="10" pageNumber="331">MLILWANEENSIS</smallCapsWord>
|
||||
</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3F9EFB20959BFB18" baselines="1193,1194" box="[1186,1277,1175,1199]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="10" pageNumber="331">AHRENS</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3E33FB209410FB18" baselines="1193,1194" box="[1295,1398,1175,1199]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="10" pageNumber="331">BECKETT</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3864FB0192D6FB7A" baselines="1224,1224" box="[856,944,1206,1230]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="10" pageNumber="331">EBERLE</smallCapsWord>
|
||||
&
|
||||
<smallCapsWord id="8D3B4EADFF8E385E38E6FB019554FB7A" baselines="1224,1224" box="[986,1074,1206,1230]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabrizi" pageId="10" pageNumber="331">FABRIZI</smallCapsWord>
|
||||
</taxonomicName>
|
||||
<emphasis id="B9160463FF8E385E3F07FB0C95C7FB79" bold="true" box="[1083,1185,1204,1231]" pageId="10" pageNumber="331">
|
||||
<taxonomicNameLabel id="A225B918FF8E385E3F07FB0C95C7FB79" box="[1083,1185,1204,1231]" pageId="10" pageNumber="331" rank="species">
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3F07FB0C9531FB7A" baselines="1224" box="[1083,1111,1211,1228]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="10" pageNumber="331">SP</smallCapsWord>
|
||||
.
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3F54FB0C95FEFB7A" baselines="1224" box="[1128,1176,1211,1228]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="10" pageNumber="331">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF8E385E3F8FFB019453FB78" box="[1203,1333,1204,1230]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="10" pageNumber="331">
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3F8FFB01958EFB7A" baselines="1224,1224" box="[1203,1256,1206,1230]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="10" pageNumber="331">FIGS</smallCapsWord>
|
||||
2Q
|
||||
<emphasis id="B9160463FF8E385E3E2AFB0F9442FB78" box="[1302,1316,1208,1230]" italics="true" pageId="10" pageNumber="331">–</emphasis>
|
||||
T
|
||||
</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8E385E381BFB5C92B6FAA9" blockId="10.[807,1428,1259,1311]" pageId="10" pageNumber="331">
|
||||
<uri id="FFF3D473FF8E385E381BFB5C92B6FAA9" pageId="10" pageNumber="331">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF8E385E3F04FB5C92B6FAA9" pageId="10" pageNumber="331">3E523A53-33D3-43CD-B4D7- 1103C2602228</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF8E38583815FAF0906BFD5A" lastPageId="12" lastPageNumber="333" pageId="10" pageNumber="331" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF8E385E3815FAF094E1FA2E" blockId="10.[809,1424,1350,1433]" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E3815FAF0953AFAED" box="[809,1116,1350,1372]" italics="true" pageId="10" pageNumber="331">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF8E385E3F56FAF194E1FA2E" collectingDate="1996-11-17" collectingDateMax="1996-11-18" collectingDateMin="1996-11-17" collectionCode="NME" collectorName="M. Hartmann" country="Swaziland" latitude="-22.0" location="Mlilwane Wildlife Sanctuary" longLatPrecision="925" longitude="31.183332" pageId="10" pageNumber="331" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF8E385E3F56FAF195B6FAEA" box="[1130,1232,1350,1372]" pageId="10" pageNumber="331" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D6413F8FF8E385E3FD8FAFE9592FAED" box="[1252,1268,1353,1371]" count="1" pageId="10" pageNumber="331" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectingCountry id="F37598E1FF8E385E3E2FFAF194EDFAEA" box="[1299,1419,1350,1372]" name="Swaziland" pageId="10" pageNumber="331">Swaziland</collectingCountry>
|
||||
,
|
||||
<location id="8EBD8EAAFF8E385E3815FAD295EBFACC" LSID="urn:lsid:plazi:treatment:03CB6967FF8E38583805FB2E906BFD5A:8EBD8EAAFF8E385E3815FAD295EBFACC" box="[809,1165,1381,1403]" country="Swaziland" latitude="-22.0" longLatPrecision="925" longitude="31.183332" name="Mlilwane Wildlife Sanctuary" pageId="10" pageNumber="331">Mlilwane Wildlife Sanctuary</location>
|
||||
26
|
||||
<emphasis id="B9160463FF8E385E3F83FAD195AFFACC" box="[1215,1225,1382,1402]" italics="true" pageId="10" pageNumber="331">°</emphasis>
|
||||
29,
|
||||
<geoCoordinate id="EE56BEB6FF8E385E3FD5FAD2947BFACC" box="[1257,1309,1376,1402]" degrees="22" direction="south" minutes="0" orientation="latitude" pageId="10" pageNumber="331" precision="925" value="-22.0">
|
||||
22
|
||||
<emphasis id="B9160463FF8E385E3E3AFAD7946DFAD9" bold="true" box="[1286,1291,1376,1391]" italics="true" pageId="10" pageNumber="331">
|
||||
<superScript id="7C177539FF8E385E3E3AFAD7946DFAD9" attach="right" box="[1286,1291,1376,1391]" fontSize="6" pageId="10" pageNumber="331">0</superScript>
|
||||
</emphasis>
|
||||
S
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE56BEB6FF8E385E3E04FAD294F6FACC" box="[1336,1424,1376,1402]" degrees="31" direction="east" minutes="11" orientation="longitude" pageId="10" pageNumber="331" precision="15" seconds="0" value="31.183332">
|
||||
31
|
||||
<emphasis id="B9160463FF8E385E3E6FFAD1943BFACC" box="[1363,1373,1382,1402]" italics="true" pageId="10" pageNumber="331">°</emphasis>
|
||||
11
|
||||
<emphasis id="B9160463FF8E385E3E44FAD7941BFAD9" bold="true" box="[1400,1405,1376,1391]" italics="true" pageId="10" pageNumber="331">
|
||||
<superScript id="7C177539FF8E385E3E44FAD7941BFAD9" attach="right" box="[1400,1405,1376,1391]" fontSize="6" pageId="10" pageNumber="331">0</superScript>
|
||||
</emphasis>
|
||||
E
|
||||
</geoCoordinate>
|
||||
800mNN,
|
||||
<date id="FFDCFEB1FF8E385E38A2FA349520FA2F" box="[926,1094,1411,1433]" pageId="10" pageNumber="331" value="1996-11-17" valueMax="1996-11-18" valueMin="1996-11-17">
|
||||
<collectingDate id="EF980759FF8E385E38A2FA349520FA2F" box="[926,1094,1411,1433]" pageId="10" pageNumber="331" value="1996-11-17" valueMax="1996-11-18" valueMin="1996-11-17">17.-18.XI.1996</collectingDate>
|
||||
</date>
|
||||
, leg.
|
||||
<collectorName id="2697BDA7FF8E385E3FBFFA339446FA2E" box="[1155,1312,1411,1433]" pageId="10" pageNumber="331">M. Hartmann</collectorName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF8E385E3E06FA34941AFA2E" box="[1338,1404,1411,1432]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="10" pageNumber="331">NME</collectionCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8E38583815FA769329FE04" blockId="10.[809,1424,1472,1922]" lastBlockId="12.[144,759,199,435]" lastPageId="12" lastPageNumber="333" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E3815FA7692D7FA60" box="[809,945,1473,1494]" italics="true" pageId="10" pageNumber="331">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8E385E3F4CFA7695AFFA60" box="[1136,1225,1473,1494]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="10" pageNumber="331" unit="mm" value="7.7">7.7 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF8E385E3815FA6892E4FA42" box="[809,898,1503,1525]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="10" pageNumber="331" unit="mm" value="6.8">6.8 mm</quantity>
|
||||
, maximum width: 4.0 mm. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus strongly reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.74. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum moderately large. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.44. Metatibia, ratio of width/length: 1/3.71. Metatarsi dorsally smooth; first metatarsomere slightly shorter than subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF8838583A37FE2A90E6FE04" box="[267,384,413,435]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="12" pageNumber="333">
|
||||
Fig. 2Q
|
||||
<emphasis id="B9160463FF8838583A5DFE28900BFE07" box="[353,365,415,433]" italics="true" pageId="12" pageNumber="333">–</emphasis>
|
||||
S
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF8838583AC9FE2A932AFE04" box="[501,588,413,435]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="12" pageNumber="333">Fig. 2T</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF1D88F9FF8F385F3B9AF9029473F888" ID-DOI="http://doi.org/10.5281/zenodo.14807095" ID-Zenodo-Dep="14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="11" pageNumber="332" startId="11.[166,245,1717,1736]" targetBox="[237,1338,218,1653]" targetPageId="11" targetType="figure">
|
||||
<paragraph id="8BDDD871FF8F385F3B9AF9029473F888" blockId="11.[166,1444,1717,1854]" pageId="11" pageNumber="332">
|
||||
<emphasis id="B9160463FF8F385F3B9AF9029076F97F" bold="true" box="[166,272,1717,1737]" pageId="11" pageNumber="332">Figure 5.</emphasis>
|
||||
Female genitalia of
|
||||
<taxonomicName id="4C62A3F2FF8F385F3AC2F9019381F97F" box="[510,743,1718,1737]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="undefined-A">
|
||||
<emphasis id="B9160463FF8F385F3AC2F901930AF97F" box="[510,620,1718,1737]" italics="true" pageId="11" pageNumber="332">Pleophylla</emphasis>
|
||||
species. A
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C62A3F2FF8F385F39CBF9019565F97F" box="[759,1027,1718,1737]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="fasciatipennis">
|
||||
<emphasis id="B9160463FF8F385F39CBF9019565F97F" box="[759,1027,1718,1737]" italics="true" pageId="11" pageNumber="332">Pleophylla fasciatipennis</emphasis>
|
||||
</taxonomicName>
|
||||
; B,
|
||||
<taxonomicName id="4C62A3F2FF8F385F3F05F9019597F97F" box="[1081,1265,1718,1737]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B9160463FF8F385F3F05F9019597F97F" box="[1081,1265,1718,1737]" italics="true" pageId="11" pageNumber="332">Pleophylla pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
; C,
|
||||
<emphasis id="B9160463FF8F385F3E1BF90290C0F950" bold="true" pageId="11" pageNumber="332">
|
||||
<taxonomicName id="4C62A3F2FF8F385F3E1BF902905BF953" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="pseudopilosa" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8F385F3E1BF902905BF953" bold="true" italics="true" pageId="11" pageNumber="332">Pleophylla pseudopilosa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8F385F3A75F96490C0F950" box="[329,422,1747,1766]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
; D,
|
||||
<emphasis id="B9160463FF8F385F3AE6F965921BF950" bold="true" box="[474,893,1746,1766]" pageId="11" pageNumber="332">
|
||||
<taxonomicName id="4C62A3F2FF8F385F3AE6F9659273F953" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[474,789,1746,1765]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="nelshoogteensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8F385F3AE6F9659273F953" bold="true" box="[474,789,1746,1765]" italics="true" pageId="11" pageNumber="332">Pleophylla nelshoogteensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8F385F381CF964921BF950" box="[800,893,1747,1766]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
; E,
|
||||
<emphasis id="B9160463FF8F385F388CF965945FF950" bold="true" box="[944,1337,1746,1766]" pageId="11" pageNumber="332">
|
||||
<taxonomicName id="4C62A3F2FF8F385F388CF96595B6F953" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[944,1232,1746,1765]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="burundiensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8F385F388CF96595B6F953" bold="true" box="[944,1232,1746,1765]" italics="true" pageId="11" pageNumber="332">Pleophylla burundiensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8F385F3FE0F964945FF950" box="[1244,1337,1747,1766]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
; F,
|
||||
<emphasis id="B9160463FF8F385F3E56F9659363F8B5" bold="true" pageId="11" pageNumber="332">
|
||||
<taxonomicName id="4C62A3F2FF8F385F3E56F96590C4F8B5" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="settentrionalis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8F385F3E56F96590C4F8B5" bold="true" italics="true" pageId="11" pageNumber="332">Pleophylla settentrionalis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8F385F3A96F9479363F8B5" box="[426,517,1776,1795]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
; G,
|
||||
<taxonomicName id="4C62A3F2FF8F385F3908F9479270F8B5" authorityName="Burmeister" authorityYear="1855" box="[564,790,1776,1795]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="ferruginea">
|
||||
<emphasis id="B9160463FF8F385F3908F9479270F8B5" box="[564,790,1776,1795]" italics="true" pageId="11" pageNumber="332">Pleophylla ferruginea</emphasis>
|
||||
</taxonomicName>
|
||||
; H,
|
||||
<emphasis id="B9160463FF8F385F387AF94795EDF8B5" bold="true" box="[838,1163,1776,1795]" pageId="11" pageNumber="332">
|
||||
<taxonomicName id="4C62A3F2FF8F385F387AF947954EF8B5" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[838,1064,1776,1795]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="murzini" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8F385F387AF947954EF8B5" bold="true" box="[838,1064,1776,1795]" italics="true" pageId="11" pageNumber="332">Pleophylla murzini</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8F385F3F0CF94795EDF8B5" box="[1072,1163,1776,1795]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
; I,
|
||||
<taxonomicName id="4C62A3F2FF8F385F3F8DF94794FBF8B5" authorityName="Burmeister" authorityYear="1855" box="[1201,1437,1776,1795]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="navicularis">
|
||||
<emphasis id="B9160463FF8F385F3F8DF94794FBF8B5" box="[1201,1437,1776,1795]" italics="true" pageId="11" pageNumber="332">Pleophylla navicularis</emphasis>
|
||||
</taxonomicName>
|
||||
; J,
|
||||
<taxonomicName id="4C62A3F2FF8F385F3BFFF8BA90F4F896" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[195,402,1805,1824]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="silvatica">
|
||||
<emphasis id="B9160463FF8F385F3BFFF8BA90F4F896" box="[195,402,1805,1824]" italics="true" pageId="11" pageNumber="332">Pleophylla silvatica</emphasis>
|
||||
</taxonomicName>
|
||||
; K,
|
||||
<emphasis id="B9160463FF8F385F3AF8F8BA922CF897" bold="true" box="[452,842,1805,1825]" pageId="11" pageNumber="332">
|
||||
<taxonomicName id="4C62A3F2FF8F385F3AF8F8BA9383F896" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[452,741,1805,1824]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="transkeiensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8F385F3AF8F8BA9383F896" bold="true" box="[452,741,1805,1824]" italics="true" pageId="11" pageNumber="332">Pleophylla transkeiensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8F385F39D3F8B9922CF897" box="[751,842,1806,1825]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
; L,
|
||||
<emphasis id="B9160463FF8F385F3845F8BA95D5F897" bold="true" box="[889,1203,1805,1825]" pageId="11" pageNumber="332">
|
||||
<taxonomicName id="4C62A3F2FF8F385F3845F8BA9528F896" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[889,1102,1805,1824]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="ruthae" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8F385F3845F8BA9528F896" bold="true" box="[889,1102,1805,1824]" italics="true" pageId="11" pageNumber="332">Pleophylla ruthae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8F385F3F64F8B995D5F897" box="[1112,1203,1806,1825]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
; M,
|
||||
<emphasis id="B9160463FF8F385F3FD5F8BA903CF888" bold="true" pageId="11" pageNumber="332">
|
||||
<taxonomicName id="4C62A3F2FF8F385F3FD5F8BA919EF88B" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="warnockae" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8F385F3FD5F8BA919EF88B" bold="true" italics="true" pageId="11" pageNumber="332">Pleophylla warnockae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8F385F3A3CF89C903CF888" box="[256,346,1835,1854]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
; N,
|
||||
<emphasis id="B9160463FF8F385F3AB4F89D93BAF888" bold="true" box="[392,732,1834,1854]" pageId="11" pageNumber="332">
|
||||
<taxonomicName id="4C62A3F2FF8F385F3AB4F89D931DF88B" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[392,635,1834,1853]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="harrisoni" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8F385F3AB4F89D931DF88B" bold="true" box="[392,635,1834,1853]" italics="true" pageId="11" pageNumber="332">Pleophylla harrisoni</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8F385F39BEF89C93BAF888" box="[642,732,1835,1854]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
; O,
|
||||
<emphasis id="B9160463FF8F385F3835F89D9548F888" bold="true" box="[777,1070,1834,1854]" pageId="11" pageNumber="332">
|
||||
<taxonomicName id="4C62A3F2FF8F385F3835F89D92AAF88B" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[777,972,1834,1853]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="stalsi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8F385F3835F89D92AAF88B" bold="true" box="[777,972,1834,1853]" italics="true" pageId="11" pageNumber="332">Pleophylla stalsi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF8F385F38E8F89C9548F888" box="[980,1070,1835,1854]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
Scale bars = 0.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BDDD871FF8838583BACFE669048FDC9" blockId="12.[144,759,464,639]" pageId="12" pageNumber="333">
|
||||
<emphasis id="B9160463FF8838583BACFE66932EFE53" box="[144,584,464,486]" italics="true" pageId="12" pageNumber="333">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF8838583A29FE67932EFE53" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[277,584,464,485]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="333" phylum="Arthropoda" rank="species" species="mlilwaneensis" status="sp. nov.">Pleophylla mlilwaneensis</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF883858395DFE6693ADFE50" box="[609,715,465,486]" pageId="12" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is rather similar to
|
||||
<taxonomicName id="4C62A3F2FF8838583A5BFE589353FDB2" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[359,565,495,516]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="333" phylum="Arthropoda" rank="species" species="nelshoogteensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8838583A5BFE589353FDB2" box="[359,565,495,516]" italics="true" pageId="12" pageNumber="333">P. nelshoogteensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF883858397FFE5893C6FDB2" box="[579,672,495,516]" pageId="12" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
in the shape of parameres. It differs from the latter in having the distal basomedial tooth in the apical third of the left paramere (rather than having both in basal half).
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8838583BACFD2E906BFD5A" blockId="12.[144,758,665,748]" pageId="12" pageNumber="333">
|
||||
<emphasis id="B9160463FF8838583BACFD2E906BFD18" box="[144,269,665,686]" italics="true" pageId="12" pageNumber="333">Etymology:</emphasis>
|
||||
The new species is named after its
|
||||
<typeStatus id="54D966D3FF88385839F9FD2E9390FD18" box="[709,758,665,686]" pageId="12" pageNumber="333">type</typeStatus>
|
||||
locality, the Mlilwane Wildlife Sanctuary in
|
||||
<collectingCountry id="F37598E1FF8838583BACFD61906FFD5A" box="[144,265,726,748]" name="Swaziland" pageId="12" pageNumber="333">Swaziland</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
220
data/03/CB/69/03CB6967FF8E385E3BACFB289438FBD5.xml
Normal file
220
data/03/CB/69/03CB6967FF8E385E3BACFB289438FBD5.xml
Normal file
|
|
@ -0,0 +1,220 @@
|
|||
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<mods:titleInfo id="4891B45F9C02A3482D63C4D843F92DEB">
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<mods:title id="C3D9D2B6D050A73EFD66752630D0F93B">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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|
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|
||||
<mods:extent id="746F8B91071AF3388E06D3D6B775D74F" unit="page">
|
||||
<mods:start id="C20DE4A9D4AF7984B1633ACC71EFD5EE">321</mods:start>
|
||||
<mods:end id="AA15CBDCBFE37C85CA9EA9133ED86188">353</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location id="B520897C2CEF6D75D4E7099D897AF3FB">
|
||||
<mods:url id="EE36D73B8D9B6E2C45F9FBC964523043">https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="AFEDD0B6D69ECB903CFC0D33BC41D8E8">journal article</mods:classification>
|
||||
<mods:identifier id="CCA99EE454B0C484125854A54E7A20CB" type="DOI">10.1111/zoj.12489</mods:identifier>
|
||||
<mods:identifier id="8D5CA687A3CA0749D071567991692D87" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="457523F4A058DB8154A120DFE9C8B298" type="Zenodo-Dep">14807083</mods:identifier>
|
||||
<mods:identifier id="A5E448E9A746C34CB6F61CD2969864F1" type="ZooBank">3663E0A-2E17-4346-A388-6BEE53081814</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03CB6967FF8E385E3BACFB289438FBD5" LSID="urn:lsid:plazi:treatment:03CB6967FF8E385E3BACFB289438FBD5" httpUri="http://treatment.plazi.org/id/03CB6967FF8E385E3BACFB289438FBD5" lastPageNumber="331" pageId="10" pageNumber="331">
|
||||
<subSubSection id="C3788BFAFF8E385E3BACFB28902EFA90" pageId="10" pageNumber="331" type="nomenclature">
|
||||
<paragraph id="8BDDD871FF8E385E3BACFB289385FB65" blockId="10.[144,759,1179,1236]" pageId="10" pageNumber="331">
|
||||
<heading id="D0956F1DFF8E385E3BACFB289385FB65" centered="true" fontSize="9" level="2" pageId="10" pageNumber="331" reason="2">
|
||||
<taxonomicName id="4C62A3F2FF8E385E3BACFB28901BFB64" ID-CoL="4K849" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="nelshoogteensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF8E385E3BACFB289377FB05" bold="true" box="[144,529,1183,1204]" italics="true" pageId="10" pageNumber="331">
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3BACFB28904CFB05" baselines="1199,1199" box="[144,298,1183,1204]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="10" pageNumber="331">PLEOPHYLLA</smallCapsWord>
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3A0FFB159377FB05" baselines="1199" box="[307,529,1186,1203]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nelshoogteensis" pageId="10" pageNumber="331">NELSHOOGTEENSIS</smallCapsWord>
|
||||
</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3927FB2B9310FB05" baselines="1198,1199" box="[539,630,1180,1204]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="10" pageNumber="331">AHRENS</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF8E385E39B4FB2B9389FB05" baselines="1198,1199" box="[648,751,1180,1204]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="10" pageNumber="331">BECKETT</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3B9FFB0C919AFB67" baselines="1229,1229" box="[163,252,1211,1235]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="10" pageNumber="331">EBERLE</smallCapsWord>
|
||||
&
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3A19FB0C901BFB64" baselines="1229,1230" box="[293,381,1211,1235]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabrizi" pageId="10" pageNumber="331">FABRIZI</smallCapsWord>
|
||||
</taxonomicName>
|
||||
<emphasis id="B9160463FF8E385E3ABBFB77908AFB62" bold="true" box="[391,492,1209,1236]" pageId="10" pageNumber="331">
|
||||
<taxonomicNameLabel id="A225B918FF8E385E3ABBFB77908AFB62" box="[391,492,1209,1236]" pageId="10" pageNumber="331" rank="species">
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3ABBFB7790C5FB67" baselines="1229" box="[391,419,1216,1233]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="10" pageNumber="331">SP</smallCapsWord>
|
||||
.
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3A88FB779082FB67" baselines="1229" box="[436,484,1216,1233]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="10" pageNumber="331">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF8E385E3AC3FB0C93E3FB65" box="[511,645,1209,1235]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="10" pageNumber="331">
|
||||
<smallCapsWord id="8D3B4EADFF8E385E3AC3FB0C9352FB64" baselines="1229,1230" box="[511,564,1211,1235]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="10" pageNumber="331">FIGS</smallCapsWord>
|
||||
2M
|
||||
<emphasis id="B9160463FF8E385E395AFB0A9312FB65" box="[614,628,1213,1235]" italics="true" pageId="10" pageNumber="331">–</emphasis>
|
||||
P
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF8E385E39AAFB0E93DCFB65" box="[662,698,1209,1235]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="10" pageNumber="331">5D</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8E385E3BB3FB46902EFA90" blockId="10.[143,757,1265,1318]" pageId="10" pageNumber="331">
|
||||
<uri id="FFF3D473FF8E385E3BB3FB46902EFA90" pageId="10" pageNumber="331">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF8E385E3AA2FB46902EFA90" pageId="10" pageNumber="331">9FB34217-3242-4B0D-BFF7- 21960F3BC6DA</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF8E385E3BAEFA809438FBD5" pageId="10" pageNumber="331" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF8E385E3BAEFA8093CFFA11" blockId="10.[146,761,1335,1448]" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E3BAEFA8090DBFAFA" box="[146,445,1335,1356]" italics="true" pageId="10" pageNumber="331">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF8E385E3AF7FA809031FA11" collectingDate="1986-02-12" collectionCode="TMSA" collectorName="S. Afr & Endrody-Younga" latitude="-25.5" location="Tv. Nelshoogte Forest Station" longLatPrecision="748" longitude="30.5" pageId="10" pageNumber="331" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF8E385E3AF7FA809352FAFB" box="[459,564,1335,1357]" pageId="10" pageNumber="331" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF8E385E3974FA8D933EFAFA" box="[584,600,1338,1356]" count="1" pageId="10" pageNumber="331" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF8E385E3948FA8093A6FAFB" box="[628,704,1335,1357]" pageId="10" pageNumber="331">S. Afr</collectorName>
|
||||
;
|
||||
<location id="8EBD8EAAFF8E385E39E8FA8090BCFADD" LSID="urn:lsid:plazi:treatment:03CB6967FF8E385E3BACFB289438FBD5:8EBD8EAAFF8E385E39E8FA8090BCFADD" latitude="-25.5" longLatPrecision="748" longitude="30.5" name="Tv. Nelshoogte Forest Station" pageId="10" pageNumber="331">Tv. Nelshoogte Forest Station</location>
|
||||
<geoCoordinate id="EE56BEB6FF8E385E3ACDFAE29333FADC" box="[497,597,1365,1387]" degrees="25.50" direction="south" orientation="latitude" pageId="10" pageNumber="331" precision="555" value="-25.5">25.50 S</geoCoordinate>
|
||||
-
|
||||
<geoCoordinate id="EE56BEB6FF8E385E39B0FAE29396FADC" box="[652,752,1365,1387]" degrees="30.50" direction="east" orientation="longitude" pageId="10" pageNumber="331" precision="555" value="30.5">30.50 E</geoCoordinate>
|
||||
/
|
||||
<date id="FFDCFEB1FF8E385E3BAEFAC39199FA3C" box="[146,255,1396,1418]" pageId="10" pageNumber="331" value="1986-02-12">
|
||||
<collectingDate id="EF980759FF8E385E3BAEFAC39199FA3C" box="[146,255,1396,1418]" pageId="10" pageNumber="331" value="1986-02-12">2.12.1986</collectingDate>
|
||||
</date>
|
||||
E-Y: 2346 UV-light collection leg.
|
||||
<collectorName id="2697BDA7FF8E385E39B3FAC3918FFA11" pageId="10" pageNumber="331">Endrody-Younga</collectorName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF8E385E3A3EFA259036FA11" box="[258,336,1426,1447]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="10" pageNumber="331">TMSA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<typeStatus id="54D966D3FF8E385E3A5FFA2490BFFA1E" box="[355,473,1427,1448]" pageId="10" pageNumber="331" type="paratype">Paratypes</typeStatus>
|
||||
: see Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8E385E3BAEFA77947EFE67" blockId="10.[146,761,1471,1921]" lastBlockId="10.[807,1421,199,465]" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E3BAEFA77907CFA63" box="[146,282,1472,1493]" italics="true" pageId="10" pageNumber="331">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8E385E3AE5FA089354FA63" box="[473,562,1471,1493]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="10" pageNumber="331" unit="mm" value="9.6">9.6 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF8E385E3BAEFA69918DFA45" box="[146,235,1502,1524]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="10" pageNumber="331" unit="mm" value="6.3">6.3 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF8E385E3AE7FA699353FA45" box="[475,565,1502,1524]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="10" pageNumber="331" unit="mm" value="4.6">4.6 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.77. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/ metacoxa: 1/1.5. Metatibia, ratio of width/length: 1/ 4.14. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF8E385E3E31FE2A94EFFE04" box="[1293,1417,413,435]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="10" pageNumber="331">
|
||||
Fig. 2M
|
||||
<emphasis id="B9160463FF8E385E3E5BFE179415FE04" box="[1383,1395,416,434]" italics="true" pageId="10" pageNumber="331">–</emphasis>
|
||||
O
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF8E385E38B3FE0C9283FE66" box="[911,997,443,465]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="10" pageNumber="331">
|
||||
Fig.
|
||||
<specimenCount id="9D6413F8FF8E385E38F8FE0C9283FE66" box="[964,997,443,464]" count="2" pageId="10" pageNumber="331" type="generic">2P</specimenCount>
|
||||
</figureCitation>
|
||||
. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF8E385E3F86FE0C9475FE67" box="[1210,1299,443,465]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="10" pageNumber="331">Fig. 5D</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8E385E381BFE5D95BEFD2E" blockId="10.[807,1421,489,664]" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E381BFE5D95BDFE48" box="[807,1243,489,511]" italics="true" pageId="10" pageNumber="331">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF8E385E3890FE5E95BDFE48" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[940,1243,489,510]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="nelshoogteensis" status="sp. nov.">Pleophylla nelshoogteensis</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF8E385E3FDFFE5D945AFE49" box="[1251,1340,490,511]" pageId="10" pageNumber="331" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from
|
||||
<taxonomicName id="4C62A3F2FF8E385E384CFDBF92B2FDAB" box="[880,980,520,541]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B9160463FF8E385E384CFDBF92B2FDAB" box="[880,980,520,541]" italics="true" pageId="10" pageNumber="331">P. pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
in the parameres by having the ventral lobe in lateral view wider and shorter, and from
|
||||
<taxonomicName id="4C62A3F2FF8E385E3856FDF1957AFDEC" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[874,1052,581,603]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="pseudopilosa">
|
||||
<emphasis id="B9160463FF8E385E3856FDF1957AFDEC" box="[874,1052,581,603]" italics="true" pageId="10" pageNumber="331">P. pseudopilosa</emphasis>
|
||||
</taxonomicName>
|
||||
by the base of left paramere being widened, the base of dorsal lobe of left paramere with a double medial tooth.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8E385E3816FD0795F1FC0C" blockId="10.[810,1425,687,954]" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E3816FD0792FBFD73" box="[810,925,688,709]" italics="true" pageId="10" pageNumber="331">Variation:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF8E385E3F73FD1895B1FD73" box="[1103,1239,687,709]" metricMagnitude="-3" metricUnit="m" metricValue="8.35" metricValueMax="9.6" metricValueMin="7.1" pageId="10" pageNumber="331" unit="mm" value="8.35" valueMax="9.6" valueMin="7.1">
|
||||
7.1
|
||||
<emphasis id="B9160463FF8E385E3F4EFD059518FD72" box="[1138,1150,690,708]" italics="true" pageId="10" pageNumber="331">–</emphasis>
|
||||
9.6 mm
|
||||
</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF8E385E3816FD7992D7FD55" box="[810,945,718,740]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.5" metricValueMin="5.6" pageId="10" pageNumber="331" unit="mm" value="6.05" valueMax="6.5" valueMin="5.6">
|
||||
5.6
|
||||
<emphasis id="B9160463FF8E385E3870FD66923EFD55" box="[844,856,721,739]" italics="true" pageId="10" pageNumber="331">–</emphasis>
|
||||
6.5 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF8E385E3FA1FD799440FD55" box="[1181,1318,718,739]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.8" metricValueMin="3.9" pageId="10" pageNumber="331" unit="mm" value="4.35" valueMax="4.8" valueMin="3.9">
|
||||
3.9
|
||||
<emphasis id="B9160463FF8E385E3F83FD6695ADFD55" box="[1215,1227,721,739]" italics="true" pageId="10" pageNumber="331">–</emphasis>
|
||||
4.8 mm
|
||||
</quantity>
|
||||
. Density of dark spot may vary slightly, but never completely without spots. Female: antennal club of female straight, composed of five antennomeres, slightly longer than the remaining antennomeres combined; sixth antennomere subequal one to three quarters of club length. Eyes smaller than in male: ratio of eye diameter/interocular width: 0.5.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8E385E3814FC6295DDFBBC" blockId="10.[808,1423,981,1034]" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E3814FC6292C0FC5C" box="[808,934,981,1002]" italics="true" pageId="10" pageNumber="331">Etymology:</emphasis>
|
||||
The new species is named after its
|
||||
<typeStatus id="54D966D3FF8E385E3E62FC6194E9FC5D" box="[1374,1423,982,1003]" pageId="10" pageNumber="331">type</typeStatus>
|
||||
locality, Nelshoogte Forest Station.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF8E385E3816FB999438FBD5" blockId="10.[810,1425,1070,1123]" pageId="10" pageNumber="331">
|
||||
<emphasis id="B9160463FF8E385E3816FB9992F3FBF5" box="[810,917,1070,1091]" italics="true" pageId="10" pageNumber="331">Remarks:</emphasis>
|
||||
In a previous publication we referred to this species as ‘sp1A’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF8E385E3F87FBFA9434FBD5" author="Ahrens D" box="[1211,1362,1101,1123]" pageId="10" pageNumber="331" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
241
data/03/CB/69/03CB6967FF9038403BADFCFC9449FD2C.xml
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241
data/03/CB/69/03CB6967FF9038403BADFCFC9449FD2C.xml
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|
|
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<taxonomicName id="4C62A3F2FF9038403BADFCFC903BFCC8" ID-CoL="4K843" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="lizleri" status="sp. nov.">
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,
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,
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&
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.
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|
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.
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</taxonomicNameLabel>
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</emphasis>
|
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(
|
||||
<figureCitation id="1359C4F4FF9038403AE3FCD09331FCC9" box="[479,599,869,895]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">
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4I
|
||||
<emphasis id="B9160463FF9038403904FCDE9320FCC9" box="[568,582,873,895]" italics="true" pageId="20" pageNumber="341">–</emphasis>
|
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L
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF9038403954FCD2931EFCC9" box="[616,632,869,895]" captionStart="Figure 7" captionStartId="14.[144,223,925,944]" captionTargetBox="[144,1428,192,893]" captionTargetId="figure-25@14.[144,1428,192,893]" captionTargetPageId="14" captionText="Figure 7. Distribution of Pleophylla species (left side) and absence records (right side), represented by sampling locali- ties of all other sampled Sericini species without records of Pleophylla (white dots)." figureDoi="http://doi.org/10.5281/zenodo.14807097" httpUri="https://zenodo.org/record/14807097/files/figure.png" pageId="20" pageNumber="341">7</figureCitation>
|
||||
)
|
||||
</heading>
|
||||
</paragraph>
|
||||
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urn:lsid:zoobank.org:act:
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|
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</subSubSection>
|
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|
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<emphasis id="B9160463FF9038403BACFC5C90AEFC49" box="[144,456,1002,1024]" italics="true" pageId="20" pageNumber="341">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9038403AEBFC5D904AFB8A" collectingDate="1999-12-07" collectionCode="ZFMK" collectorName="Lizler" country="Tanzania" location="Porote Mts." pageId="20" pageNumber="341" specimenCount="1" specimenCount-male="1" stateProvince="Mbeya" typeStatus="holotype">
|
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<typeStatus id="54D966D3FF9038403AEBFC5D9326FBB6" box="[471,576,1002,1024]" pageId="20" pageNumber="341" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF9038403967FC5A930DFC49" box="[603,619,1005,1023]" count="1" pageId="20" pageNumber="341" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectingCountry id="F37598E1FF90384039B0FC5D9390FBB6" box="[652,758,1002,1024]" name="Tanzania" pageId="20" pageNumber="341">Tanzania</collectingCountry>
|
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<date id="FFDCFEB1FF9038403BACFBBE9060FBA8" box="[144,262,1033,1054]" pageId="20" pageNumber="341" value="1999-12-07">
|
||||
<collectingDate id="EF980759FF9038403BACFBBE9060FBA8" box="[144,262,1033,1054]" pageId="20" pageNumber="341" value="1999-12-07">7.XII.1999</collectingDate>
|
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</date>
|
||||
<location id="8EBD8EAAFF9038403A2CFBBE90F0FBA8" LSID="urn:lsid:plazi:treatment:03CB6967FF9038403BADFCFC9449FD2C:8EBD8EAAFF9038403A2CFBBE90F0FBA8" box="[272,406,1033,1054]" country="Tanzania" name="Porote Mts." pageId="20" pageNumber="341" stateProvince="Mbeya">Porote Mts.</location>
|
||||
<collectingRegion id="49A61693FF9038403AA3FBBE9348FBA8" box="[415,558,1033,1055]" country="Tanzania" name="Mbeya" pageId="20" pageNumber="341">Mbeya prov.</collectingRegion>
|
||||
Werner &
|
||||
<collectorName id="2697BDA7FF903840398FFBBE9390FBA9" box="[691,758,1033,1055]" pageId="20" pageNumber="341">Lizler</collectorName>
|
||||
leg.” (
|
||||
<collectionCode id="ED7340B4FF9038403BEAFB909043FB8A" box="[214,293,1063,1084]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="20" pageNumber="341" type="Museum">ZFMK</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B0AD22CFF9038403A01FB9F918BFBCF" collectingDate="1999-12-07" collectionCode="ZFMK" collectorName="Lizler" country="Tanzania" location="Porote Mts." pageId="20" pageNumber="341" specimenCount="2" specimenCount-male="2" stateProvince="Mbeya" typeStatus="paratype">
|
||||
<typeStatus id="54D966D3FF9038403A01FB9F90D5FB8B" box="[317,435,1064,1085]" pageId="20" pageNumber="341" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF9038403AF9FB909096FB8A" box="[453,496,1063,1084]" count="2" pageId="20" pageNumber="341" type="male">2 ♂</specimenCount>
|
||||
“
|
||||
<collectingCountry id="F37598E1FF903840393BFB909317FB8B" box="[519,625,1063,1085]" name="Tanzania" pageId="20" pageNumber="341">Tanzania</collectingCountry>
|
||||
<date id="FFDCFEB1FF9038403943FB909390FB8A" box="[639,758,1063,1084]" pageId="20" pageNumber="341" value="1999-12-07">
|
||||
<collectingDate id="EF980759FF9038403943FB909390FB8A" box="[639,758,1063,1084]" pageId="20" pageNumber="341" value="1999-12-07">7.XII.1999</collectingDate>
|
||||
</date>
|
||||
<location id="8EBD8EAAFF9038403BACFBF19046FBED" LSID="urn:lsid:plazi:treatment:03CB6967FF9038403BADFCFC9449FD2C:8EBD8EAAFF9038403BACFBF19046FBED" box="[144,288,1094,1115]" country="Tanzania" name="Porote Mts." pageId="20" pageNumber="341" stateProvince="Mbeya">Porote Mts.</location>
|
||||
<collectingRegion id="49A61693FF9038403A0FFBF190AAFBED" box="[307,460,1094,1116]" country="Tanzania" name="Mbeya" pageId="20" pageNumber="341">Mbeya prov.</collectingRegion>
|
||||
Werner &
|
||||
<collectorName id="2697BDA7FF9038403953FBF193D4FBEA" box="[623,690,1094,1116]" pageId="20" pageNumber="341">Lizler</collectorName>
|
||||
leg.” (
|
||||
<collectionCode id="ED7340B4FF9038403BABFBD39180FBCF" box="[151,230,1124,1145]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="20" pageNumber="341" type="Museum">ZFMK</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9038403BACFB2090A8F8DC" blockId="20.[144,759,1174,1899]" pageId="20" pageNumber="341">
|
||||
<emphasis id="B9160463FF9038403BACFB20907EFB1A" box="[144,280,1175,1196]" italics="true" pageId="20" pageNumber="341">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9038403AECFB219351FB1A" box="[464,567,1174,1196]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="20" pageNumber="341" unit="mm" value="11.4">11.4 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9038403BACFB02918FFB7C" box="[144,233,1205,1227]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="20" pageNumber="341" unit="mm" value="7.6">7.6 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9038403AE4FB029355FB7C" box="[472,563,1205,1227]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="20" pageNumber="341" unit="mm" value="5.6">5.6 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus strongly reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.78. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.18. Metatibia, ratio of width/length: 1/3.84. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF9038403BACF8E29199F8DC" box="[144,255,1877,1899]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">
|
||||
Fig. 4I
|
||||
<emphasis id="B9160463FF9038403BE0F8EF918EF8DC" box="[220,232,1880,1898]" italics="true" pageId="20" pageNumber="341">–</emphasis>
|
||||
K
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF9038403A48F8E290ACF8DC" box="[372,458,1877,1899]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">
|
||||
Fig.
|
||||
<specimenCount id="9D6413F8FF9038403A95F8E290ACF8DC" box="[425,458,1877,1898]" count="4" pageId="20" pageNumber="341" type="generic">4L</specimenCount>
|
||||
</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF903840381BFF7095F6FE67" blockId="20.[806,1421,199,465]" pageId="20" pageNumber="341">
|
||||
<emphasis id="B9160463FF903840381BFF709509FF6A" box="[807,1135,199,220]" italics="true" pageId="20" pageNumber="341">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF9038403890FF709509FF6A" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[940,1135,199,220]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="lizleri" status="sp. nov.">Pleophylla lizleri</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF9038403F44FF7095B7FF6A" box="[1144,1233,199,220]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from the quite similar
|
||||
<taxonomicName id="4C62A3F2FF90384038F8FF519553FF4C" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[964,1077,229,251]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF90384038F8FF519553FF4C" box="[964,1077,229,251]" italics="true" pageId="20" pageNumber="341">P. charlyi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9038403F02FF5195FEFF4D" box="[1086,1176,230,251]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C62A3F2FF9038403FE9FF5194EBFF4C" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1237,1421,229,251]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="mpumalanga" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9038403FE9FF5194EBFF4C" box="[1237,1421,229,251]" italics="true" pageId="20" pageNumber="341">P. mpumalanga</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF903840381BFEB392E2FEAF" box="[807,900,260,281]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
in the less extended lobe produced by the dorsolateral extension of the ventral lobe of the right paramere. Furthermore,
|
||||
<taxonomicName id="4C62A3F2FF9038403F67FEF595DBFEE0" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1115,1213,321,343]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="lizleri" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9038403F67FEF595DBFEE0" box="[1115,1213,321,343]" italics="true" pageId="20" pageNumber="341">P. lizleri</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9038403FF3FEF69454FEE0" box="[1231,1330,321,342]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from
|
||||
<taxonomicName id="4C62A3F2FF9038403857FED792BDFEC3" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[875,987,352,373]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9038403857FED792BDFEC3" box="[875,987,352,373]" italics="true" pageId="20" pageNumber="341">P. charlyi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF90384038D6FED7952FFEC3" box="[1002,1097,352,373]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
by the robust basomedial teeth of the left paramere, which is basally more widened, and from
|
||||
<taxonomicName id="4C62A3F2FF9038403F2FFE2A95ACFE04" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1043,1226,413,434]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="mpumalanga" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9038403F2FFE2A95ACFE04" box="[1043,1226,413,434]" italics="true" pageId="20" pageNumber="341">P. mpumalanga</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9038403FEBFE2A9452FE04" box="[1239,1332,413,434]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
by the wider parameres (lateral view).
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF903840381AFE4692FEFDF2" blockId="20.[806,1421,497,580]" pageId="20" pageNumber="341">
|
||||
<emphasis id="B9160463FF903840381AFE4692FCFDB0" box="[806,922,497,518]" italics="true" pageId="20" pageNumber="341">Variation:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9038403F7AFE4695BCFDB0" box="[1094,1242,497,518]" metricMagnitude="-2" metricUnit="m" metricValue="1.065" metricValueMax="1.14" metricValueMin="0.99" pageId="20" pageNumber="341" unit="mm" value="10.65" valueMax="11.4" valueMin="9.9">
|
||||
9.9
|
||||
<emphasis id="B9160463FF9038403F54FE439512FDB0" box="[1128,1140,500,518]" italics="true" pageId="20" pageNumber="341">–</emphasis>
|
||||
11.4 mm
|
||||
</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF903840381AFDA792C8FD93" box="[806,942,527,549]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="7.6" metricValueMin="7.1" pageId="20" pageNumber="341" unit="mm" value="7.35" valueMax="7.6" valueMin="7.1">
|
||||
7.1
|
||||
<emphasis id="B9160463FF9038403875FDA59233FD92" box="[841,853,530,548]" italics="true" pageId="20" pageNumber="341">–</emphasis>
|
||||
7.6 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9038403FA1FDA79440FD93" box="[1181,1318,527,549]" metricMagnitude="-3" metricUnit="m" metricValue="5.199999999999999" metricValueMax="5.6" metricValueMin="4.8" pageId="20" pageNumber="341" unit="mm" value="5.199999999999999" valueMax="5.6" valueMin="4.8">
|
||||
4.8
|
||||
<emphasis id="B9160463FF9038403F83FDA595ADFD92" box="[1215,1227,530,548]" italics="true" pageId="20" pageNumber="341">–</emphasis>
|
||||
5.6 mm
|
||||
</quantity>
|
||||
. Female unknown.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF903840381AFDD29449FD2C" blockId="20.[806,1420,613,666]" pageId="20" pageNumber="341">
|
||||
<emphasis id="B9160463FF903840381AFDD292C2FDCC" box="[806,932,613,634]" italics="true" pageId="20" pageNumber="341">Etymology:</emphasis>
|
||||
The new species is named after one of its collectors, Robert Lızler (
|
||||
<collectingRegion id="49A61693FF9038403F57FD339445FD2C" box="[1131,1315,644,666]" country="Czech Republic" name="Kralovehradecky" pageId="20" pageNumber="341">Hradec Kralove</collectingRegion>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
230
data/03/CB/69/03CB6967FF903841386FFD699300FEDE.xml
Normal file
230
data/03/CB/69/03CB6967FF903841386FFD699300FEDE.xml
Normal file
|
|
@ -0,0 +1,230 @@
|
|||
<document id="EC91F089875905CDE26FE6D845396116" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia & Ahrens, Dirk" docDate="2017" docId="03CB6967FF903841386FFD699300FEDE" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla taitaensis AHRENS, BECKETT, EBERLE & FABRIZI 2017, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="342" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="341" updateTime="1738712560284" updateUser="ExternalLinkService">
|
||||
<mods:mods id="884485681F0633B1DE3CDFD5DCAD9266" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="4BF56717C22D106BF5E6C6C9EB53256A">
|
||||
<mods:title id="FF1BF5D13FA470CB74FFC3B55F4B0DF1">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="94C921B0AD298A2D8658B7F31F3ACE47" type="personal">
|
||||
<mods:role id="427FD640CA582E9EC97E31F44A4216DF">
|
||||
<mods:roleTerm id="EDBD045BA0126CBA156E39A8E924D8C6">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="77025C9E3C07F1686FBF8C10F1DA9A2D">Eberle, Jonas</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="D425F725E56F5DF31ED36ABF9052B496" type="personal">
|
||||
<mods:role id="7F3157710D18E12144D23A42A017C47C">
|
||||
<mods:roleTerm id="7A29758C8853C651A08BC8C2240BFA87">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="0B65B7DF5B7059B9E8633C5552852E98">Beckett, Marc</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="6170135175C57549B30856C07D718D4D" type="personal">
|
||||
<mods:role id="808BD98F467F45DB0886F2B3B74E78BF">
|
||||
<mods:roleTerm id="374617ACFEB35CA885ED5CC19720BD92">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="5ADBC96DAA65ED2E74B77088E6AD8DB3">Ozgul-Siemund, Alev</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="37298FE707852BFF568630E357ACCA42" type="personal">
|
||||
<mods:role id="F1870C5A27211F252D07E80F95AE068D">
|
||||
<mods:roleTerm id="98D17A5A7236D51034F3BB172BEB5622">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="C05AA6BAA0E4E346E28A0202BF446850">Frings, Jujina</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="CEAFEACC90094239AD0BA4820E17ABA7" type="personal">
|
||||
<mods:role id="9C5165E3A66618019DEC313000B93446">
|
||||
<mods:roleTerm id="99D84936A7D7C7EAA7ED6386D6C1E814">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="7820AC7B6A5DA902813E533126660421">Fabrizi, Silvia</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="386C0BF1DDE21A1F1B4C9567F49D66DC" type="personal">
|
||||
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|
||||
<mods:roleTerm id="336B8E3E0A4ACED75B6A569EE81C0642">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="819CEB44B2AAC35AF84AA45E01F29351">Ahrens, Dirk</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="9100E43B4B11B69624C68AC91FDF1E45">text</mods:typeOfResource>
|
||||
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|
||||
<mods:title id="BE588D2F780AD2A82ACBF1678B26BD0C">Zoological Journal of the Linnean Society</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="536525651443FAEB767FC355B6062A16">
|
||||
<mods:date id="53ECC7F15B795FE12A1C436BDE39B457">2017</mods:date>
|
||||
<mods:detail id="9D6678055E4E61D54821A925A8245070" type="series">
|
||||
<mods:title id="CDE6B47A621A3722A808D889DB37D71B">Zool. J. Linn. Soc.</mods:title>
|
||||
</mods:detail>
|
||||
<mods:detail id="082192DCD50BAB881ED7722AB4A5322A" type="volume">
|
||||
<mods:number id="841FE8D72F8D6B6F12194FDDE2689B77">180</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="17E23C1BB552B526AD4C465486AB34D9" type="issue">
|
||||
<mods:number id="26C5532B1DD0D64729020A1FCCEE5070">2</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="D16B6BA9A91AAFA7724FA4DDAB8F98D3" unit="page">
|
||||
<mods:start id="D3754D090B61969C0841FD9E6576E8FA">321</mods:start>
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<mods:end id="3766A8D0E6834E72147195B0411F0BF4">353</mods:end>
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<mods:location id="EC507BB33E7781519AF8C163B33FCCB9">
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||||
<mods:url id="54556D4D727C14011D53527EECB0B6AA">https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="15FD7E3ACA54EC3C32F7E0715477A792">journal article</mods:classification>
|
||||
<mods:identifier id="D5F0AEC693FD458369AD9874F7FEDC3D" type="DOI">10.1111/zoj.12489</mods:identifier>
|
||||
<mods:identifier id="01A5AA8C23A47CAA36F6C63064E470F3" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="7EF9EFA5C4CF4E84E3041396B0EA6765" type="Zenodo-Dep">14807083</mods:identifier>
|
||||
<mods:identifier id="E7DE5A57D0F2E26B1D0FD4FAADF023EA" type="ZooBank">3663E0A-2E17-4346-A388-6BEE53081814</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03CB6967FF903841386FFD699300FEDE" LSID="urn:lsid:plazi:treatment:03CB6967FF903841386FFD699300FEDE" httpUri="http://treatment.plazi.org/id/03CB6967FF903841386FFD699300FEDE" lastPageId="21" lastPageNumber="342" pageId="20" pageNumber="341">
|
||||
<subSubSection id="C3788BFAFF903840386FFD6992B0FCD1" pageId="20" pageNumber="341" type="nomenclature">
|
||||
<paragraph id="8BDDD871FF903840386FFD699439FCA4" blockId="20.[851,1377,730,787]" pageId="20" pageNumber="341">
|
||||
<heading id="D0956F1DFF903840386FFD699439FCA4" centered="true" fontSize="9" level="2" pageId="20" pageNumber="341" reason="2">
|
||||
<taxonomicName id="4C62A3F2FF903840386FFD699548FCA7" ID-CoL="4K84H" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="taitaensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF903840386FFD69951AFD44" bold="true" box="[851,1148,734,755]" italics="true" pageId="20" pageNumber="341">
|
||||
<smallCapsWord id="8D3B4EADFF903840386FFD69928AFD44" baselines="750,750" box="[851,1004,734,755]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="20" pageNumber="341">PLEOPHYLLA</smallCapsWord>
|
||||
<smallCapsWord id="8D3B4EADFF90384038CAFD56951AFD44" baselines="750" box="[1014,1148,737,754]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="taitaensis" pageId="20" pageNumber="341">TAITAENSIS</smallCapsWord>
|
||||
</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF9038403FB9FD6C9586FD44" baselines="749,750" box="[1157,1248,731,755]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="20" pageNumber="341">AHRENS</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF9038403FCEFD6C943EFD44" baselines="749,750" box="[1266,1368,731,755]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="20" pageNumber="341">BECKETT</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF9038403868FD4D92CAFCA6" baselines="780,780" box="[852,940,762,786]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="20" pageNumber="341">EBERLE</smallCapsWord>
|
||||
&
|
||||
<smallCapsWord id="8D3B4EADFF90384038EAFD4D9548FCA7" baselines="780,781" box="[982,1070,762,786]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabrizi" pageId="20" pageNumber="341">FABRIZI</smallCapsWord>
|
||||
</taxonomicName>
|
||||
<emphasis id="B9160463FF9038403F0BFD4895FBFCA5" bold="true" box="[1079,1181,760,787]" pageId="20" pageNumber="341">
|
||||
<taxonomicNameLabel id="A225B918FF9038403F0BFD4895FBFCA5" box="[1079,1181,760,787]" pageId="20" pageNumber="341" rank="species">
|
||||
<smallCapsWord id="8D3B4EADFF9038403F0BFD489535FCA6" baselines="780" box="[1079,1107,767,784]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="20" pageNumber="341">SP</smallCapsWord>
|
||||
.
|
||||
<smallCapsWord id="8D3B4EADFF9038403F59FD4895F3FCA6" baselines="780" box="[1125,1173,767,784]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="20" pageNumber="341">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF9038403F8CFD4D9453FCA4" box="[1200,1333,760,786]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">
|
||||
<smallCapsWord id="8D3B4EADFF9038403F8CFD4D9583FCA7" baselines="780,781" box="[1200,1253,762,786]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="20" pageNumber="341">FIGS</smallCapsWord>
|
||||
4M
|
||||
<emphasis id="B9160463FF9038403E2AFD4B9442FCA4" box="[1302,1316,764,786]" italics="true" pageId="20" pageNumber="341">–</emphasis>
|
||||
P
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF9038403E7BFD4F9431FCA4" box="[1351,1367,760,786]" captionStart="Figure 7" captionStartId="14.[144,223,925,944]" captionTargetBox="[144,1428,192,893]" captionTargetId="figure-25@14.[144,1428,192,893]" captionTargetPageId="14" captionText="Figure 7. Distribution of Pleophylla species (left side) and absence records (right side), represented by sampling locali- ties of all other sampled Sericini species without records of Pleophylla (white dots)." figureDoi="http://doi.org/10.5281/zenodo.14807097" httpUri="https://zenodo.org/record/14807097/files/figure.png" pageId="20" pageNumber="341">7</figureCitation>
|
||||
)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9038403815FC8592B0FCD1" blockId="20.[809,1424,818,871]" pageId="20" pageNumber="341">
|
||||
<uri id="FFF3D473FF9038403815FC8592B0FCD1" pageId="20" pageNumber="341">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF9038403F04FC8592B0FCD1" pageId="20" pageNumber="341">8E31225B-A3BB-4BB3-91B0- 8A9295AC7295</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF903841381BFCCB9300FEDE" lastPageId="21" lastPageNumber="342" pageId="20" pageNumber="341" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF903840381BFCCB92BCFBBD" blockId="20.[807,1422,891,1035]" pageId="20" pageNumber="341">
|
||||
<emphasis id="B9160463FF903840381BFCCB9528FC26" box="[807,1102,891,913]" italics="true" pageId="20" pageNumber="341">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9038403F60FCCC92E2FC7B" collectingDate="1997-03-22" collectionCode="ZFMK" collectorName="Lgt. M. Snizek" country="Kenya" location="Hills Wundanyi" pageId="20" pageNumber="341" specimenCount="1" specimenCount-male="1" stateProvince="Taita/Taveta" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF9038403F60FCCC95A4FC27" box="[1116,1218,891,913]" pageId="20" pageNumber="341" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D6413F8FF9038403FEDFCC99587FC26" box="[1233,1249,894,912]" count="1" pageId="20" pageNumber="341" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectingCountry id="F37598E1FF9038403FC5FCCC9424FC26" box="[1273,1346,891,912]" name="Kenya" pageId="20" pageNumber="341">Kenya</collectingCountry>
|
||||
<collectingRegion id="49A61693FF9038403E6DFCCC94E8FC27" box="[1361,1422,891,913]" country="Kenya" name="Taita/Taveta" pageId="20" pageNumber="341">Taita</collectingRegion>
|
||||
<location id="8EBD8EAAFF903840381BFC2D92B8FC06" LSID="urn:lsid:plazi:treatment:03CB6967FF903841386FFD699300FEDE:8EBD8EAAFF903840381BFC2D92B8FC06" box="[807,990,922,944]" country="Kenya" name="Hills Wundanyi" pageId="20" pageNumber="341" stateProvince="Taita/Taveta">Hills Wundanyi</location>
|
||||
18.3.-
|
||||
<date id="FFDCFEB1FF9038403F1CFC2D95F5FC19" box="[1056,1171,922,943]" pageId="20" pageNumber="341" value="1997-03-22">
|
||||
<collectingDate id="EF980759FF9038403F1CFC2D95F5FC19" box="[1056,1171,922,943]" pageId="20" pageNumber="341" value="1997-03-22">22.3.1997</collectingDate>
|
||||
</date>
|
||||
<collectorName id="2697BDA7FF9038403FA2FC2D942DFC06" box="[1182,1355,922,944]" pageId="20" pageNumber="341">Lgt. M. Snizek</collectorName>
|
||||
[sic]” (
|
||||
<collectionCode id="ED7340B4FF9038403812FC0F921BFC7B" box="[814,893,952,973]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="20" pageNumber="341" type="Museum">ZFMK</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B0AD22CFF90384038A9FC0E92B0FBBD" collectingDate="2001-01" collectionCode="ISNB" collectorName="Coll. I. R. Sc. N. B. & Coll. I. R. Sc. & Leg. S. Sabari" country="Kenya" location="Hills Mbololo" pageId="20" pageNumber="341" specimenCount="2" specimenCount-male="2" stateProvince="Taita/Taveta" typeStatus="paratype">
|
||||
<typeStatus id="54D966D3FF90384038A9FC0E956DFC78" box="[917,1035,953,974]" pageId="20" pageNumber="341" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF9038403F21FC0F953FFC7B" box="[1053,1113,952,973]" count="2" pageId="20" pageNumber="341" type="male">2 ♂♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF9038403F4CFC0F9449FC7B" box="[1136,1327,952,974]" pageId="20" pageNumber="341">Coll. I.R.Sc.N.B.</collectorName>
|
||||
<collectingCountry id="F37598E1FF9038403E01FC0E94EFFC78" box="[1341,1417,953,974]" name="Kenya" pageId="20" pageNumber="341">Kenya</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="49A61693FF903840381BFC609202FC5B" box="[807,868,983,1005]" country="Kenya" name="Taita/Taveta" pageId="20" pageNumber="341">Taita</collectingRegion>
|
||||
<location id="8EBD8EAAFF9038403849FC60957FFC5B" LSID="urn:lsid:plazi:treatment:03CB6967FF903841386FFD699300FEDE:8EBD8EAAFF9038403849FC60957FFC5B" box="[885,1049,983,1005]" country="Kenya" name="Hills Mbololo" pageId="20" pageNumber="341" stateProvince="Taita/Taveta">Hills Mbololo</location>
|
||||
, III.
|
||||
<date id="FFDCFEB1FF9038403F6CFC6095CCFC5A" box="[1104,1194,983,1004]" pageId="20" pageNumber="341" value="2001-01">
|
||||
<collectingDate id="EF980759FF9038403F6CFC6095CCFC5A" box="[1104,1194,983,1004]" pageId="20" pageNumber="341" value="2001-01">2001 I.</collectingDate>
|
||||
</date>
|
||||
G. 31.839
|
||||
<collectorName id="2697BDA7FF9038403E08FC609217FBBD" pageId="20" pageNumber="341">Leg. S. Sabari</collectorName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF90384038B7FC4192A8FBBD" box="[907,974,1014,1035]" pageId="20" pageNumber="341">ISNB</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF903840381BFB979588F963" blockId="20.[807,1422,1055,1750]" pageId="20" pageNumber="341">
|
||||
<emphasis id="B9160463FF903840381BFB9792C9FB83" box="[807,943,1056,1077]" italics="true" pageId="20" pageNumber="341">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9038403F5BFBA895A8FB83" box="[1127,1230,1055,1077]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="20" pageNumber="341" unit="mm" value="11.5">11.5 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF903840381BFB8992E6FBE2" box="[807,896,1086,1108]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="20" pageNumber="341" unit="mm" value="7.4">7.4 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9038403F53FB8995AFFBE2" box="[1135,1225,1086,1108]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="20" pageNumber="341" unit="mm" value="5.4">5.4 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus strongly reflexed; smooth area in front of eyes 1.5 times as wide as long; ratio of eye diameter/interocular width: 0.83. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.4. Metatibia, ratio of width/length: 1/4.13. Metatarsi dorsally smooth; first metatarsomere slightly longer than subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF903840389EF9779579F963" box="[930,1055,1728,1750]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">
|
||||
Fig. 4M
|
||||
<emphasis id="B9160463FF90384038C0F974956EF963" box="[1020,1032,1731,1749]" italics="true" pageId="20" pageNumber="341">–</emphasis>
|
||||
O
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF9038403FA8F977958CF963" box="[1172,1258,1728,1750]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">
|
||||
Fig.
|
||||
<specimenCount id="9D6413F8FF9038403FF5F977958CF963" box="[1225,1258,1728,1749]" count="4" pageId="20" pageNumber="341" type="generic">4P</specimenCount>
|
||||
</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF903840381BF94F958BF8DF" blockId="20.[807,1421,1783,1897]" pageId="20" pageNumber="341">
|
||||
<emphasis id="B9160463FF903840381BF94F95FDF8BB" box="[807,1179,1783,1805]" italics="true" pageId="20" pageNumber="341">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF9038403890F94095FDF8BB" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[940,1179,1783,1805]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="taitaensis" status="sp. nov.">Pleophylla taitaensis</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF9038403F98F94F9598F8BB" box="[1188,1278,1784,1805]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from the very similar
|
||||
<taxonomicName id="4C62A3F2FF90384038D1F8A09529F89D" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1005,1103,1814,1836]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="lizleri" status="sp. nov.">
|
||||
<emphasis id="B9160463FF90384038D1F8A09529F89D" box="[1005,1103,1814,1836]" italics="true" pageId="20" pageNumber="341">P. lizleri</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9038403F65F8A195D2F89D" box="[1113,1204,1814,1835]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C62A3F2FF9038403FCEF8A09405F89D" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1266,1379,1814,1836]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9038403FCEF8A09405F89D" box="[1266,1379,1814,1836]" italics="true" pageId="20" pageNumber="341">P. charlyi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9038403E51F8A19231F8FC" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
by the ventral lobes of parameres being symmetrical and not extended dorsally.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9138413B9AFF709012FEAC" blockId="21.[166,781,198,282]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413B9AFF70907CFF6A" box="[166,282,199,220]" italics="true" pageId="21" pageNumber="342">Variation:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9138413AC4FF7093FCFF6A" box="[504,666,198,220]" metricMagnitude="-2" metricUnit="m" metricValue="1.095" metricValueMax="1.15" metricValueMin="1.04" pageId="21" pageNumber="342" unit="mm" value="10.95" valueMax="11.5" valueMin="10.4">
|
||||
10.4
|
||||
<emphasis id="B9160463FF9138413914FF7E9352FF6D" box="[552,564,201,219]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||||
11.5 mm
|
||||
</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9138413A3BFF5290E8FF4D" box="[263,398,229,251]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="7.8" metricValueMin="7.4" pageId="21" pageNumber="342" unit="mm" value="7.6" valueMax="7.8" valueMin="7.4">
|
||||
7.4
|
||||
<emphasis id="B9160463FF9138413A15FF5F9053FF4C" box="[297,309,232,250]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||||
7.8 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF91384139BDFF52926FFF4D" box="[641,777,229,251]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.6" metricValueMin="5.4" pageId="21" pageNumber="342" unit="mm" value="5.5" valueMax="5.6" valueMin="5.4">
|
||||
5.4
|
||||
<emphasis id="B9160463FF913841399FFF5F93C9FF4C" box="[675,687,232,250]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||||
5.6 mm
|
||||
</quantity>
|
||||
. Female unknown.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9138413B9AFE839300FEDE" blockId="21.[166,780,308,360]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413B9AFE839042FEFF" box="[166,292,308,329]" italics="true" pageId="21" pageNumber="342">Etymology:</emphasis>
|
||||
The new species is named after its occurrence in the
|
||||
<collectingRegion id="49A61693FF9138413A48FEE590D7FEDE" box="[372,433,338,360]" country="Kenya" name="Taita/Taveta" pageId="21" pageNumber="342">Taita</collectingRegion>
|
||||
Hills of
|
||||
<collectingCountry id="F37598E1FF9138413929FEE49304FEDE" box="[533,610,339,360]" name="Kenya" pageId="21" pageNumber="342">Kenya</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
227
data/03/CB/69/03CB6967FF913841387DFEE7944AF95F.xml
Normal file
227
data/03/CB/69/03CB6967FF913841387DFEE7944AF95F.xml
Normal file
|
|
@ -0,0 +1,227 @@
|
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<mods:title id="3E5E6C287AF71D3D7FC9770302F01EF7">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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<mods:namePart id="F8B5E4FFCD1647439F0655B57FA743C6">Eberle, Jonas</mods:namePart>
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<mods:namePart id="5AE3315BC97E06ECF76E1241771FAC81">Beckett, Marc</mods:namePart>
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<mods:date id="ECD05A846B0283A347D43256D533D879">2017</mods:date>
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<taxonomicName id="4C62A3F2FF913841387DFEE79505FE35" ID-CoL="4K84M" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="342" phylum="Arthropoda" rank="species" species="wakkerstroomensis" status="sp. nov.">
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|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF9138413E60FEF99211FE35" baselines="383" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="ett" pageId="21" pageNumber="342">BECKETT</smallCapsWord>
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,
|
||||
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|
||||
&
|
||||
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|
||||
</taxonomicName>
|
||||
<emphasis id="B9160463FF9138413F50FEC595B4FE30" bold="true" box="[1132,1234,363,390]" pageId="21" pageNumber="342">
|
||||
<taxonomicNameLabel id="A225B918FF9138413F50FEC595B4FE30" box="[1132,1234,363,390]" pageId="21" pageNumber="342" rank="species">
|
||||
<smallCapsWord id="8D3B4EADFF9138413F50FEC595EEFE35" baselines="383" box="[1132,1160,370,387]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="21" pageNumber="342">SP</smallCapsWord>
|
||||
.
|
||||
<smallCapsWord id="8D3B4EADFF9138413FA6FEC595ACFE35" baselines="383" box="[1178,1226,370,387]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="21" pageNumber="342">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF9138413FD9FEDA940EFE33" box="[1253,1384,363,389]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="21" pageNumber="342">
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|
||||
4U
|
||||
<emphasis id="B9160463FF9138413E74FED99430FE32" box="[1352,1366,366,388]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||||
X
|
||||
</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
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<uri id="FFF3D473FF9138413807FE1D9292FE69" pageId="21" pageNumber="342">
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urn:lsid:zoobank.org:act:
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</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF9138413801FE4D944AF95F" pageId="21" pageNumber="342" type="materials_examined">
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|
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<emphasis id="B9160463FF9138413801FE4D950AFDB8" box="[829,1132,505,527]" italics="true" pageId="21" pageNumber="342">Type material examined:</emphasis>
|
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:
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||||
“835047 X-DA1676
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:
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<collectingMunicipality id="6BB9420BFF9138413FD7FDAF9412FD98" box="[1259,1396,536,558]" pageId="21" pageNumber="342">Wakkerstro</collectingMunicipality>
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[o]m (
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env.);
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</quantity>
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27
|
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<emphasis id="B9160463FF9138413E2EFD8F947AFDFA" box="[1298,1308,568,588]" italics="true" pageId="21" pageNumber="342">°</emphasis>
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20
|
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<emphasis id="B9160463FF9138413E0BFD85945AFDF7" bold="true" box="[1335,1340,562,577]" italics="true" pageId="21" pageNumber="342">
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</emphasis>
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19, 6
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<emphasis id="B9160463FF9138413EB6FD8194F2FDFD" bold="true" box="[1418,1428,566,587]" italics="true" pageId="21" pageNumber="342">″</emphasis>
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S
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</geoCoordinate>
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30
|
||||
<emphasis id="B9160463FF9138413864FDE19204FDDC" box="[856,866,598,618]" italics="true" pageId="21" pageNumber="342">°</emphasis>
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<geoCoordinate id="EE56BEB6FF913841385EFDE292B8FDDD" box="[866,990,593,619]" degrees="09" direction="east" minutes="0" orientation="longitude" pageId="21" pageNumber="342" precision="1" seconds="11, 1" value="9.003083">
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09
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</emphasis>
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11, 1
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<emphasis id="B9160463FF91384138FEFDE292AAFDDC" bold="true" box="[962,972,597,618]" italics="true" pageId="21" pageNumber="342">″</emphasis>
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E
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</geoCoordinate>
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<date id="FFDCFEB1FF91384138D0FDE295FDFDDD" box="[1004,1179,597,619]" pageId="21" pageNumber="342" value="2007-12-14" valueMax="2007-12-15" valueMin="2007-12-14">
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<collectingDate id="EF980759FF91384138D0FDE295FDFDDD" box="[1004,1179,597,619]" pageId="21" pageNumber="342" value="2007-12-14" valueMax="2007-12-15" valueMin="2007-12-14">14.-15.XII.2007</collectingDate>
|
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</date>
|
||||
leg.
|
||||
<collectorName id="2697BDA7FF9138413FDDFDE1943AFDDD" box="[1249,1372,597,619]" pageId="21" pageNumber="342">D. Ahrens</collectorName>
|
||||
&
|
||||
<collectorName id="2697BDA7FF9138413EB2FDE292F7FD3C" pageId="21" pageNumber="342">S. Fabrizi</collectorName>
|
||||
,
|
||||
<taxonomicName id="4C62A3F2FF91384138A2FDC39570FD3F" authorityName="Erichson" authorityYear="1847" box="[926,1046,628,649]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="342" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B9160463FF91384138A2FDC39570FD3F" box="[926,1046,628,649]" italics="true" pageId="21" pageNumber="342">Pleophylla</emphasis>
|
||||
</taxonomicName>
|
||||
spSAX2” (
|
||||
<collectionCode id="ED7340B4FF9138413FACFDC395B9FD3F" box="[1168,1247,628,649]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="21" pageNumber="342" type="Museum">ZFMK</collectionCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9138413801FD159441FAE1" blockId="21.[829,1444,674,1368]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413801FD1592A0FD01" box="[829,966,674,695]" italics="true" pageId="21" pageNumber="342">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9138413FBCFD159582FD01" box="[1152,1252,674,695]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="21" pageNumber="342" unit="mm" value="10.4">10.4 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9138413801FD7792F2FD60" box="[829,916,704,726]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="21" pageNumber="342" unit="mm" value="7.6">7.6 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9138413FBBFD7795B9FD60" box="[1159,1247,704,726]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="21" pageNumber="342" unit="mm" value="5.4">5.4 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus strongly reflexed; smooth area in front of eyes 1.5 times as wide as long; ratio of eye diameter/interocular width: 0.55. Pronotum apparently glabrous (with only a few erect setae anteriorly, otherwise with only minute setae); unicoloured. Hypomeron not carinate. Basal margin of pronotum complete; larger punctures absent from pronotum. Elytra without dark spots; erect dorsal pilosity on elytra sparse (fewer than ten setae per interval); sutural interval in apical declivity of elytra delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity short, less than 1/4 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.39. Metatibia, ratio of width/length: 1/4.06. Metatarsi dorsally smooth; first metatarsomere slightly longer than subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF91384138DAFAF59507FAE1" box="[998,1121,1346,1368]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="21" pageNumber="342">
|
||||
Fig. 4U
|
||||
<emphasis id="B9160463FF9138413F06FAF29520FAE1" box="[1082,1094,1349,1367]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||||
W
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF9138413FF2FAF59445FAE1" box="[1230,1315,1346,1368]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="21" pageNumber="342">Fig. 4X</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9138413801FADA95B5F98F" blockId="21.[829,1444,1388,1593]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413801FADA9475FA37" box="[829,1299,1388,1410]" italics="true" pageId="21" pageNumber="342">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF91384138FFFADB9475FA37" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[963,1299,1388,1409]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="342" phylum="Arthropoda" rank="species" species="wakkerstroomensis" status="sp. nov.">Pleophylla wakkerstroomensis</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF9138413E25FADA9409FA34" box="[1305,1391,1389,1410]" pageId="21" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is in external appearance rather similar to
|
||||
<taxonomicName id="4C62A3F2FF9138413FD4FA3C9427FA16" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1256,1345,1419,1440]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="342" phylum="Arthropoda" rank="species" species="stalsi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9138413FD4FA3C9427FA16" box="[1256,1345,1419,1440]" italics="true" pageId="21" pageNumber="342">P. stalsi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9138413E74FA3C94C6FA16" box="[1352,1440,1419,1440]" pageId="21" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
; however, the dorsal pilosity is nearly completely absent in the first and the parameres differ significantly in shape: the ventral lobes are widened in lateral view and convexly rounded at the apex (whereas they are sharply pointed in
|
||||
<taxonomicName id="4C62A3F2FF9138413F31F993950FF98E" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1037,1129,1571,1593]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="342" phylum="Arthropoda" rank="species" species="stalsi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9138413F31F993950FF98E" box="[1037,1129,1571,1593]" italics="true" pageId="21" pageNumber="342">P. stalsi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9138413F53F99395A1F98F" box="[1135,1223,1572,1593]" pageId="21" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9138413801F9EE9468F93B" blockId="21.[829,1444,1625,1677]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413801F9EE92DDF9D8" box="[829,955,1625,1646]" italics="true" pageId="21" pageNumber="342">Etymology:</emphasis>
|
||||
The new species is named after its
|
||||
<typeStatus id="54D966D3FF9138413E4FF9EE94C2F9D8" box="[1395,1444,1625,1646]" pageId="21" pageNumber="342">type</typeStatus>
|
||||
locality, Wakkerstroom, in
|
||||
<collectingCountry id="F37598E1FF9138413F49F9C0946CF93B" box="[1141,1290,1655,1677]" name="South Africa" pageId="21" pageNumber="342">South Africa</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9138413801F902944AF95F" blockId="21.[829,1444,1717,1769]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413801F90292CFF97C" box="[829,937,1717,1738]" italics="true" pageId="21" pageNumber="342">Remarks:</emphasis>
|
||||
In a previous publication we referred to this species as ‘spX2’ (
|
||||
<bibRefCitation id="EFF3A580FF9138413F00F9649446F95F" author="Eberle J & Warnock RCM & Ahrens D" box="[1084,1312,1747,1769]" pageId="21" pageNumber="342" refId="ref23428" refString="Eberle J, Warnock RCM, Ahrens D. 2016 b. Bayesian species delimitation in Pleophylla chafers (Coleoptera) - the importance of prior choice and morphology. BMC Evolutionary Biology 16: 94." type="journal volume" year="2016">
|
||||
Eberle
|
||||
<emphasis id="B9160463FF9138413FACF96395ACF95E" box="[1168,1226,1747,1769]" italics="true" pageId="21" pageNumber="342">et al.</emphasis>
|
||||
, 2016b
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
228
data/03/CB/69/03CB6967FF9138413B91FE2C9538FEAC.xml
Normal file
228
data/03/CB/69/03CB6967FF9138413B91FE2C9538FEAC.xml
Normal file
|
|
@ -0,0 +1,228 @@
|
|||
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<mods:mods id="F2C7B524409584A1F220976C8287DFFE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1FA170C4BD2578A9CAA1324CE589AB4A">
|
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<mods:title id="8E7D6CBF62E405013496F038E219DD73">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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|
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<mods:roleTerm id="C81C028A0DA81DE726FE64547E60B21B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D9B26BC1B9ED8443974A73A464A92D8C">Eberle, Jonas</mods:namePart>
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<mods:namePart id="D7B2B631DEE7B164A4B338E1286AD6AD">Beckett, Marc</mods:namePart>
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<mods:namePart id="182CEB7AA1927EBD4E9CAE513F208048">Ozgul-Siemund, Alev</mods:namePart>
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</mods:name>
|
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<mods:roleTerm id="9BCFAD572509212F7695ABD2683832E1">Author</mods:roleTerm>
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<mods:namePart id="B46DE2E5EC69F0F5201467E9C95F825E">Frings, Jujina</mods:namePart>
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<mods:roleTerm id="FEA318C8AFC385B9508CCBA1017D4A67">Author</mods:roleTerm>
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</mods:role>
|
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<mods:namePart id="1449BF3AAC94807726F98AF42426AC13">Fabrizi, Silvia</mods:namePart>
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</mods:name>
|
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<mods:name id="199DF5153806DCB2CA76E2E4912F7985" type="personal">
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<mods:roleTerm id="27F9DD76C5371EA2E784DAA7DDD04D60">Author</mods:roleTerm>
|
||||
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|
||||
<mods:namePart id="70047B633EE81BE26A8242D6EB76BB42">Ahrens, Dirk</mods:namePart>
|
||||
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|
||||
<mods:typeOfResource id="6DC6C67AB37FA4A96B76C193CD4F3956">text</mods:typeOfResource>
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|
||||
<mods:title id="8F41700DE1DF4B796ECABECB51EF0507">Zoological Journal of the Linnean Society</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="2B59782D35BBC328C3338B827B718C32">
|
||||
<mods:date id="9DD4E64A8A0E8F69F181F60883E971F4">2017</mods:date>
|
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|
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<mods:title id="44E7524BAC91F59A455738AD4D6BCF5D">Zool. J. Linn. Soc.</mods:title>
|
||||
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|
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<mods:detail id="B28336814E98EF52FB98367F779AAFB3" type="volume">
|
||||
<mods:number id="C7109F60949B37D03F8E00D2DC154CFF">180</mods:number>
|
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</mods:detail>
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<mods:number id="280D2B378AAF442DAAA884B21893DBA5">2</mods:number>
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<mods:classification id="49C9A1DAA07E08131BA52E73AA70D2F6">journal article</mods:classification>
|
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<mods:identifier id="26ABCF0461E3F99CB92BD1969950E99F" type="DOI">10.1111/zoj.12489</mods:identifier>
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<mods:identifier id="A89D267A61C90FA4769A0562BD48BF15" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="B1A91A058432FAAE8EE5326B40FE4B20" type="ZooBank">3663E0A-2E17-4346-A388-6BEE53081814</mods:identifier>
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</mods:mods>
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<treatment id="03CB6967FF9138413B91FE2C9538FEAC" LSID="urn:lsid:plazi:treatment:03CB6967FF9138413B91FE2C9538FEAC" httpUri="http://treatment.plazi.org/id/03CB6967FF9138413B91FE2C9538FEAC" lastPageNumber="342" pageId="21" pageNumber="342">
|
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<subSubSection id="C3788BFAFF9138413B91FE2C902EFD9E" pageId="21" pageNumber="342" type="nomenclature">
|
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<paragraph id="8BDDD871FF9138413B91FE2C93D0FE79" blockId="21.[173,774,407,464]" pageId="21" pageNumber="342">
|
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<heading id="D0956F1DFF9138413B91FE2C93D0FE79" centered="true" fontSize="9" level="2" pageId="21" pageNumber="342" reason="2">
|
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<taxonomicName id="4C62A3F2FF9138413B91FE2C9032FE78" ID-CoL="4K84G" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="342" phylum="Arthropoda" rank="species" species="stalsi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9138413B91FE2C90F9FE19" bold="true" box="[173,415,411,432]" italics="true" pageId="21" pageNumber="342">
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<smallCapsWord id="8D3B4EADFF9138413A6CFE2990F9FE19" baselines="427" box="[336,415,414,431]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="stalsi" pageId="21" pageNumber="342">STALSI</smallCapsWord>
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</emphasis>
|
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<smallCapsWord id="8D3B4EADFF9138413A94FE2E9362FE06" baselines="427,428" box="[424,516,409,433]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="21" pageNumber="342">AHRENS</smallCapsWord>
|
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,
|
||||
<smallCapsWord id="8D3B4EADFF9138413929FE2E931AFE06" baselines="427,428" box="[533,636,409,433]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="21" pageNumber="342">BECKETT</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF91384139B1FE2E9380FE19" baselines="427,427" box="[653,742,409,433]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="21" pageNumber="342">EBERLE</smallCapsWord>
|
||||
&
|
||||
<smallCapsWord id="8D3B4EADFF9138413BC0FE009032FE78" baselines="457,458" box="[252,340,439,463]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabrizi" pageId="21" pageNumber="342">FABRIZI</smallCapsWord>
|
||||
</taxonomicName>
|
||||
<emphasis id="B9160463FF9138413A62FE0A90A5FE66" bold="true" box="[350,451,437,464]" pageId="21" pageNumber="342">
|
||||
<taxonomicNameLabel id="A225B918FF9138413A62FE0A90A5FE66" box="[350,451,437,464]" pageId="21" pageNumber="342" rank="species">
|
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<smallCapsWord id="8D3B4EADFF9138413A62FE0A901CFE78" baselines="458" box="[350,378,445,462]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="21" pageNumber="342">SP</smallCapsWord>
|
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.
|
||||
<smallCapsWord id="8D3B4EADFF9138413AB7FE0A90DDFE78" baselines="458" box="[395,443,445,462]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="21" pageNumber="342">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
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</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF9138413AEAFE00933EFE79" box="[470,600,437,463]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="21" pageNumber="342">
|
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<smallCapsWord id="8D3B4EADFF9138413AEAFE00936DFE78" baselines="457,458" box="[470,523,439,463]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="21" pageNumber="342">FIGS</smallCapsWord>
|
||||
4Q
|
||||
<emphasis id="B9160463FF9138413905FE0E9321FE79" box="[569,583,441,463]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||||
T
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF9138413955FE0293EBFE79" box="[617,653,437,463]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="21" pageNumber="342">5O</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9138413B9FFE43902EFD9E" blockId="21.[163,778,500,552]" pageId="21" pageNumber="342">
|
||||
<uri id="FFF3D473FF9138413B9FFE43902EFD9E" pageId="21" pageNumber="342">
|
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urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF9138413A97FE43902EFD9E" pageId="21" pageNumber="342">0F7D7614-7CFD-44C3-9AAC-398093991B09</uuid>
|
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</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF9138413B9AFDF29538FEAC" pageId="21" pageNumber="342" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF9138413B9AFDF29382FD65" blockId="21.[166,782,580,724]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413B9AFDF290AFFDEF" box="[166,457,580,602]" italics="true" pageId="21" pageNumber="342">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9138413AE4FDF390FFFD65" collectingDate="2005-11-12" collectionCode="SANC" collectorName="W. Breytenbach" country="South Africa" elevation="1639" latitude="-24.933332" location="Graskop" longLatPrecision="503" longitude="30.47" municipality="Pilgrimsrest" pageId="21" pageNumber="342" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF9138413AE4FDF39327FDEC" box="[472,577,580,602]" pageId="21" pageNumber="342" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF9138413973FDF09339FDEF" box="[591,607,583,601]" count="1" pageId="21" pageNumber="342" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectingCountry id="F37598E1FF9138413948FDF3926CFDEC" box="[628,778,580,602]" name="South Africa" pageId="21" pageNumber="342">South Africa</collectingCountry>
|
||||
: MPU
|
||||
<location id="8EBD8EAAFF9138413BCAFDD49030FDCF" LSID="urn:lsid:plazi:treatment:03CB6967FF9138413B91FE2C9538FEAC:8EBD8EAAFF9138413BCAFDD49030FDCF" box="[246,342,611,633]" country="South Africa" latitude="-24.933332" longLatPrecision="503" longitude="30.47" municipality="Pilgrimsrest" name="Graskop" pageId="21" pageNumber="342">Graskop</location>
|
||||
<quantity id="4C9A7594FF9138413A56FDD590C3FDCF" box="[362,421,610,633]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="342" unit="km" value="5.0">5 km</quantity>
|
||||
on
|
||||
<collectingMunicipality id="6BB9420BFF9138413AD5FDD49310FDCF" box="[489,630,611,633]" pageId="21" pageNumber="342">Pilgrimsrest</collectingMunicipality>
|
||||
rd
|
||||
<geoCoordinate id="EE56BEB6FF9138413984FDD4926AFDCE" box="[696,780,606,632]" degrees="24" direction="south" minutes="56" orientation="latitude" pageId="21" pageNumber="342" precision="15" seconds="0" value="-24.933332">
|
||||
24
|
||||
<emphasis id="B9160463FF91384139EFFDD393BBFDCE" box="[723,733,612,632]" italics="true" pageId="21" pageNumber="342">°</emphasis>
|
||||
56
|
||||
<emphasis id="B9160463FF91384139C4FDE9939BFDDB" bold="true" box="[760,765,606,621]" italics="true" pageId="21" pageNumber="342">
|
||||
<superScript id="7C177539FF91384139C4FDE9939BFDDB" attach="right" box="[760,765,606,621]" fontSize="6" pageId="21" pageNumber="342">0</superScript>
|
||||
</emphasis>
|
||||
S
|
||||
</geoCoordinate>
|
||||
<geoCoordinate id="EE56BEB6FF9138413B9AFD369193FD20" box="[166,245,641,662]" degrees="30.47" direction="east" orientation="longitude" pageId="21" pageNumber="342" precision="555" value="30.47">30.47E</geoCoordinate>
|
||||
<date id="FFDCFEB1FF9138413A26FD3690F2FD21" box="[282,404,641,663]" pageId="21" pageNumber="342" value="2005-11-12">
|
||||
<collectingDate id="EF980759FF9138413A26FD3690F2FD21" box="[282,404,641,663]" pageId="21" pageNumber="342" value="2005-11-12">12.XI.2005</collectingDate>
|
||||
</date>
|
||||
<quantity id="4C9A7594FF9138413A85FD369368FD21" box="[441,526,641,663]" metricMagnitude="3" metricUnit="m" metricValue="1.639" pageId="21" pageNumber="342" unit="m" value="1639.0">
|
||||
<elevation id="004F3F42FF9138413A85FD369368FD21" box="[441,526,641,663]" metricMagnitude="3" metricUnit="m" metricValue="1.639" pageId="21" pageNumber="342" unit="m" value="1639.0">1639 m</elevation>
|
||||
</quantity>
|
||||
<collectorName id="2697BDA7FF913841390EFD359263FD21" box="[562,773,641,663]" pageId="21" pageNumber="342">W. Breytenbach</collectorName>
|
||||
/ Collected from
|
||||
<taxonomicName id="4C62A3F2FF9138413A6EFD179083FD03" box="[338,485,672,693]" class="Magnoliopsida" family="Proteaceae" genus="Protea" kingdom="Plantae" order="Proteales" pageId="21" pageNumber="342" phylum="Tracheophyta" rank="species" species="caffra">
|
||||
<emphasis id="B9160463FF9138413A6EFD179083FD03" box="[338,485,672,693]" italics="true" pageId="21" pageNumber="342">Protea caffra</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C62A3F2FF9138413AC8FD179314FD03" box="[500,626,672,693]" class="Magnoliopsida" family="Proteaceae" kingdom="Plantae" order="Proteales" pageId="21" pageNumber="342" phylum="Tracheophyta" rank="family">Proteaceae</taxonomicName>
|
||||
) many leaves were beaten” (
|
||||
<collectionCode id="ED7340B4FF9138413A7BFD0990F5FD65" box="[327,403,702,723]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="21" pageNumber="342" type="Museum">SANC</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<typeStatus id="54D966D3FF9138413A9FFD08937FFD62" box="[419,537,703,724]" pageId="21" pageNumber="342" type="paratype">Paratypes</typeStatus>
|
||||
: see Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9138413B9AFD5893FAFA75" blockId="21.[166,781,750,1475]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413B9AFD589049FCB2" box="[166,303,751,772]" italics="true" pageId="21" pageNumber="342">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9138413AD2FD599321FCB2" box="[494,583,750,772]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="21" pageNumber="342" unit="mm" value="9.9">9.9 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9138413B9AFCBA9199FC94" box="[166,255,781,803]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="21" pageNumber="342" unit="mm" value="6.7">6.7 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9138413AD3FCBB932FFC94" box="[495,585,780,802]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="21" pageNumber="342" unit="mm" value="5.1">5.1 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.64. Pronotum apparently glabrous (with only a few erect setae anteriorly, otherwise with only minute setae); unicoloured; basal margin of pronotum complete; larger punctures absent from pronotum. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra sparse (fewer than ten setae per interval); sutural interval in apical declivity of elytra distinctly elevated but convex; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity short, less than 1/4 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/ 1.38. Metatibia, ratio of width/length: 1/3.94. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF91384139A8FA39926FFA12" box="[660,777,1422,1444]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="21" pageNumber="342">
|
||||
Fig. 4Q
|
||||
<emphasis id="B9160463FF91384139D6FA269390FA15" box="[746,758,1425,1443]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||||
S
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF9138413A2CFA1A9000FA74" box="[272,358,1453,1475]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="21" pageNumber="342">Fig. 4T</figureCitation>
|
||||
. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF9138413903FA1A93FEFA75" box="[575,664,1453,1475]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="21" pageNumber="342">Fig. 5O</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9138413B9AFA6B90AFF9FA" blockId="21.[166,781,1499,1613]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413B9AFA6B908AFA46" box="[166,492,1499,1521]" italics="true" pageId="21" pageNumber="342">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF9138413A10FA6C908AFA46" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[300,492,1499,1520]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="342" phylum="Arthropoda" rank="species" species="stalsi" status="sp. nov.">Pleophylla stalsi</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF9138413ACBFA6C9334FA46" box="[503,594,1499,1520]" pageId="21" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from all other
|
||||
<taxonomicName id="4C62A3F2FF9138413BD3FA4D90A4F9A6" box="[239,450,1530,1552]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413BD3FA4D9001F9B9" box="[239,359,1530,1551]" italics="true" pageId="21" pageNumber="342">Pleophylla</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
by the short white pilosity on the dorsal surface, and the almost absent erect pilosity on the pronotum.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9138413B9AF9D49372F8DB" blockId="21.[166,781,1635,1901]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413B9AF9D4907CF9CE" box="[166,282,1635,1656]" italics="true" pageId="21" pageNumber="342">Variation:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9138413AF9F9D4933CF9CE" box="[453,602,1635,1656]" metricMagnitude="-2" metricUnit="m" metricValue="1.015" metricValueMax="1.04" metricValueMin="0.99" pageId="21" pageNumber="342" unit="mm" value="10.15" valueMax="10.4" valueMin="9.9">
|
||||
9.9
|
||||
<emphasis id="B9160463FF9138413ADBF9D19095F9CE" box="[487,499,1638,1656]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||||
10.4 mm
|
||||
</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9138413B9AF9369048F921" box="[166,302,1665,1687]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="6.7" metricValueMin="6.6" pageId="21" pageNumber="342" unit="mm" value="6.65" valueMax="6.7" valueMin="6.6">
|
||||
6.6
|
||||
<emphasis id="B9160463FF9138413BF4F93391B2F920" box="[200,212,1668,1686]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||||
6.7 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9138413926F93693C4F921" box="[538,674,1665,1687]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="21" pageNumber="342" unit="mm" value="4.85" valueMax="5.1" valueMin="4.6">
|
||||
4.6
|
||||
<emphasis id="B9160463FF9138413900F933932EF920" box="[572,584,1668,1686]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||||
5.1 mm
|
||||
</quantity>
|
||||
. Density of dark spots may vary, often body reddish brown and completely without spots. Female: antennal club of female straight, composed of five antennomeres, as long as the remaining antennomeres combined; sixth antennomere shorter than a quarter of club length. Eyes smaller than in male: ratio of eye diameter/interocular width: 0.5.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9138413801FF709538FEAC" blockId="21.[829,1444,199,282]" pageId="21" pageNumber="342">
|
||||
<emphasis id="B9160463FF9138413801FF7092DDFF6A" box="[829,955,199,220]" italics="true" pageId="21" pageNumber="342">Etymology:</emphasis>
|
||||
This new species is dedicated to Riaan Stals (
|
||||
<collectionCode id="ED7340B4FF91384138B2FF5192B1FF4D" box="[910,983,230,251]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="21" pageNumber="342" type="Museum">SANC</collectionCode>
|
||||
) in gratitude for his support of our research in
|
||||
<collectingCountry id="F37598E1FF91384138FAFEB3953CFEAC" box="[966,1114,260,282]" name="South Africa" pageId="21" pageNumber="342">South Africa</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
158
data/03/CB/69/03CB6967FF913842385BF8A69194FB1D.xml
Normal file
158
data/03/CB/69/03CB6967FF913842385BF8A69194FB1D.xml
Normal file
|
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@ -0,0 +1,158 @@
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<mods:namePart id="E2D4598A7DDC2746E9A78D178DBE4375">Eberle, Jonas</mods:namePart>
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<treatment id="03CB6967FF913842385BF8A69194FB1D" LSID="urn:lsid:plazi:treatment:03CB6967FF913842385BF8A69194FB1D" httpUri="http://treatment.plazi.org/id/03CB6967FF913842385BF8A69194FB1D" lastPageId="22" lastPageNumber="343" pageId="21" pageNumber="342">
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|
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<paragraph id="8BDDD871FF913841385BF8A695B4F8F3" blockId="21.[871,1402,1804,1862]" pageId="21" pageNumber="342">
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<taxonomicName id="4C62A3F2FF913841385BF8A6941CF890" ID-CoL="4K84J" authority="PERINGUEY, 1904" authorityName="Peringuey" authorityYear="1904" box="[871,1402,1804,1831]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="342" phylum="Arthropoda" rank="species" species="tongaatsana">
|
||||
<emphasis id="B9160463FF913841385BF8A695C6F892" box="[871,1184,1809,1830]" italics="true" pageId="21" pageNumber="342">
|
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<smallCapsWord id="8D3B4EADFF913841385BF8A69290F892" baselines="1825,1824" box="[871,1014,1809,1830]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="21" pageNumber="342">PLEOPHYLLA</smallCapsWord>
|
||||
<smallCapsWord id="8D3B4EADFF9138413F3CF8A395C6F892" baselines="1824" box="[1024,1184,1812,1828]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="tongaatsana" pageId="21" pageNumber="342">TONGAATSANA</smallCapsWord>
|
||||
</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF9138413F95F8B9944CF893" baselines="1824,1825" box="[1193,1322,1806,1830]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Peringuey" pageId="21" pageNumber="342">PERINGUEY</smallCapsWord>
|
||||
, 1904
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF9138413F24F89A95C1F8F5" box="[1048,1191,1835,1861]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="21" pageNumber="342">
|
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<smallCapsWord id="8D3B4EADFF9138413F24F89A952BF8F2" baselines="1855,1856" box="[1048,1101,1837,1861]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="21" pageNumber="342">FIGS</smallCapsWord>
|
||||
4Y
|
||||
<emphasis id="B9160463FF9138413F45F89895E1F8F3" box="[1145,1159,1839,1861]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF9138413FBBF89A95C1F8F5" baselines="1855,1855" box="[1159,1191,1837,1861]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ab" pageId="21" pageNumber="342">AB</smallCapsWord>
|
||||
</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF9138423801F8E09194FB1D" lastPageId="22" lastPageNumber="343" pageId="21" pageNumber="342" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF9138423801F8E093B2FEAF" blockId="21.[829,1327,1879,1901]" lastBlockId="22.[144,759,199,281]" lastPageId="22" lastPageNumber="343" pageId="21" pageNumber="342">
|
||||
<taxonomicName id="4C62A3F2FF9138413801F8E0944DF8DA" authority="Peringuey, 1904: 9" authorityName="Peringuey" authorityPageNumber="9" authorityYear="1904" box="[829,1323,1879,1901]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="342" phylum="Arthropoda" rank="species" species="tongaatsana">
|
||||
<emphasis id="B9160463FF9138413801F8E0952FF8DB" box="[829,1097,1879,1901]" italics="true" pageId="21" pageNumber="342">Pleophylla tongaatsana</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9138413F6EF8E0944DF8DA" box="[1106,1323,1879,1901]" collectionCode="SAMC" location="Upper Tongaat" pageId="21" pageNumber="342" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">Peringuey, 1904: 9</materialsCitation>
|
||||
</taxonomicName>
|
||||
.
|
||||
<emphasis id="B9160463FF9238423BACFF7090C1FF6A" box="[144,423,199,220]" italics="true" pageId="22" pageNumber="343">Type material examined:</emphasis>
|
||||
<typeStatus id="54D966D3FF9238423A8AFF709342FF6A" box="[438,548,199,220]" pageId="22" pageNumber="343" type="lectotype">Lectotype</typeStatus>
|
||||
(here designated):
|
||||
<specimenCount id="9D6413F8FF9238423BACFF5F91C6FF4C" box="[144,160,232,250]" count="1" pageId="22" pageNumber="343" type="male">♂</specimenCount>
|
||||
“
|
||||
<location id="8EBD8EAAFF9238423BF6FF5290F6FF4D" LSID="urn:lsid:plazi:treatment:03CB6967FF913842385BF8A69194FB1D:8EBD8EAAFF9238423BF6FF5290F6FF4D" box="[202,400,229,251]" name="Upper Tongaat" pageId="22" pageNumber="343">Upper Tongaat</location>
|
||||
N. [sic] 11/01/
|
||||
<taxonomicName id="4C62A3F2FF9238423942FF52907AFEAF" authorityName="Peringuey" authorityYear="1904" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="343" phylum="Arthropoda" rank="species" species="tongaatsana">
|
||||
<emphasis id="B9160463FF9238423942FF52907AFEAF" italics="true" pageId="22" pageNumber="343">Pleophylla tongaatsana</emphasis>
|
||||
</taxonomicName>
|
||||
Type Py/Type SAM/Ent 31/4” (
|
||||
<collectionCode id="ED7340B4FF9238423945FEB393AEFEAF" box="[633,712,260,281]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="22" pageNumber="343" type="Museum">SAMC</collectionCode>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9238423BACFE829196FE10" blockId="22.[144,759,309,422]" pageId="22" pageNumber="343">
|
||||
<emphasis id="B9160463FF9238423BACFE82937DFEFC" box="[144,539,309,330]" italics="true" pageId="22" pageNumber="343">Additional material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9238423916FE81918AFE10" collectingDate="2002-11-11" collectionCode="TMSA" collectorName="S. Afr." elevation="1410" latitude="-25.44" longLatPrecision="748" longitude="30.4" pageId="22" pageNumber="343" specimenCount="1" specimenCount-male="1">
|
||||
<specimenCount id="9D6413F8FF9238423916FE819300FEFC" box="[554,614,310,331]" count="1" pageId="22" pageNumber="343" type="male">1 ♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF92384239ADFE819395FEFD" box="[657,755,309,331]" pageId="22" pageNumber="343">S. Afr.</collectorName>
|
||||
:
|
||||
<taxonomicName id="4C62A3F2FF9238423BACFEE390C4FEDC" authority=", Badplaas" authorityName="Badplaas" box="[144,418,340,362]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="343" phylum="Arthropoda" rank="species" species="mpumalanga">Mpumalanga, Badplaas</taxonomicName>
|
||||
<geoCoordinate id="EE56BEB6FF9238423A92FEE39369FEDF" box="[430,527,340,362]" degrees="25.44" direction="south" orientation="latitude" pageId="22" pageNumber="343" precision="555" value="-25.44">25.44 S-</geoCoordinate>
|
||||
<geoCoordinate id="EE56BEB6FF9238423926FEE39315FEDF" box="[538,627,340,361]" degrees="30.40" direction="east" orientation="longitude" pageId="22" pageNumber="343" precision="555" value="30.4">30.40 E</geoCoordinate>
|
||||
/
|
||||
<date id="FFDCFEB1FF9238423940FEE39390FEDF" box="[636,758,340,361]" pageId="22" pageNumber="343" value="2002-11-11">
|
||||
<collectingDate id="EF980759FF9238423940FEE39390FEDF" box="[636,758,340,361]" pageId="22" pageNumber="343" value="2002-11-11">11.11.2002</collectingDate>
|
||||
</date>
|
||||
E-Y: 3566 general collect
|
||||
<quantity id="4C9A7594FF9238423AF7FEC49346FE3E" box="[459,544,371,392]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="22" pageNumber="343" unit="m" value="1410.0">
|
||||
<elevation id="004F3F42FF9238423AF7FEC49346FE3E" box="[459,544,371,392]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="22" pageNumber="343" unit="m" value="1410.0">1410 m</elevation>
|
||||
</quantity>
|
||||
leg. TMSA staff” (
|
||||
<collectionCode id="ED7340B4FF9238423BABFE269183FE10" box="[151,229,401,422]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="343">TMSA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9238423BACFE0F9194FB1D" blockId="22.[144,759,440,1195]" pageId="22" pageNumber="343">
|
||||
<emphasis id="B9160463FF9238423BACFE0F907EFE7B" box="[144,280,440,461]" italics="true" pageId="22" pageNumber="343">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9238423ADDFE0F9340FE7B" box="[481,550,440,461]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="343" unit="mm" value="9.0">9 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9238423BACFE61918FFE5A" box="[144,233,470,492]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="22" pageNumber="343" unit="mm" value="6.3">6.3 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9238423AE4FE619355FE5A" box="[472,563,470,492]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="22" pageNumber="343" unit="mm" value="4.8">4.8 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus distinct; anterior margin of labroclypeus medially distinctly sinuate; margins of labroclypeus moderately reflexed; smooth area in front of eyes 1.5 times as wide as long; ratio of eye diameter/ interocular width: 0.69. Pronotum with dense and thick erect setae; bicoloured; basal margin widely incomplete medially (equal to or more than scutellum width); larger punctures on pronotum moderately large. Hypomeron carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process long. Ratio of length of metepisternum/metacoxa: 1/1.45. Metatibia, ratio of width/length: 1/4.07. Metatarsi dorsally punctate; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF9238423ACBFBC1932DFB3A" box="[503,587,1142,1164]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="22" pageNumber="343">Fig. 4Y</figureCitation>
|
||||
<emphasis id="B9160463FF9238423977FBCE9331FB3D" box="[587,599,1145,1163]" italics="true" pageId="22" pageNumber="343">–</emphasis>
|
||||
Aa. Habitus:
|
||||
<figureCitation id="1359C4F4FF9238423BACFB2291AAFB1D" box="[144,204,1173,1195]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="22" pageNumber="343">Fig 4</figureCitation>
|
||||
Ab.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
326
data/03/CB/69/03CB6967FF9238433BAFFB69908CFCE8.xml
Normal file
326
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@ -0,0 +1,326 @@
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<taxonomicName id="4C62A3F2FF9238423BAFFB69903DFAA7" ID-CoL="4K842" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="343" phylum="Arthropoda" rank="species" species="kruegeri" status="sp. nov.">
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,
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,
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&
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</taxonomicName>
|
||||
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.
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.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF9238423AE0FB4D931FFAA6" box="[476,633,1272,1298]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="22" pageNumber="343">
|
||||
<smallCapsWord id="8D3B4EADFF9238423AE0FB4D9377FAA7" baselines="1292,1293" box="[476,529,1274,1298]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="22" pageNumber="343">FIGS</smallCapsWord>
|
||||
4
|
||||
<smallCapsWord id="8D3B4EADFF9238423917FB4D932DFAA7" baselines="1292,1293" box="[555,587,1274,1298]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ac" pageId="22" pageNumber="343">AC</smallCapsWord>
|
||||
<emphasis id="B9160463FF9238423977FB4B933FFAA4" box="[587,601,1276,1298]" italics="true" pageId="22" pageNumber="343">–</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF9238423966FB4D931FFAA6" baselines="1292,1292" box="[602,633,1274,1298]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Af" pageId="22" pageNumber="343">AF</smallCapsWord>
|
||||
</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9238423BADFA879026FAD3" blockId="22.[145,760,1328,1381]" pageId="22" pageNumber="343">
|
||||
<uri id="FFF3D473FF9238423BADFA879026FAD3" pageId="22" pageNumber="343">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF9238423AA3FA879026FAD3" pageId="22" pageNumber="343">E6A56F8B-F12A-4D52-901E-01AF81C762F4</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF9238433BACFA33908CFCE8" lastPageId="23" lastPageNumber="344" pageId="22" pageNumber="343" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF9238423BACFA3390F7F93B" blockId="22.[144,759,1411,1677]" pageId="22" pageNumber="343">
|
||||
<emphasis id="B9160463FF9238423BACFA3390AAFA2E" box="[144,460,1411,1433]" italics="true" pageId="22" pageNumber="343">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9238423AE7FA349395FA63" collectingDate="1998-12-12" collectionCode="TMSA" collectorName="S. Afr." elevation="1700" pageId="22" pageNumber="343" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF9238423AE7FA349322FA2F" box="[475,580,1411,1433]" pageId="22" pageNumber="343" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF923842395DFA319317FA2E" box="[609,625,1414,1432]" count="1" pageId="22" pageNumber="343" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF92384239AAFA349395FA2F" box="[662,755,1411,1433]" pageId="22" pageNumber="343">S. Afr.</collectorName>
|
||||
:
|
||||
<taxonomicName id="4C62A3F2FF9238423BACFA1690EAFA01" authority="Marie" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[144,396,1441,1463]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="343" phylum="Arthropoda" rank="species" species="mpumalanga">Mpumalanga Marie</taxonomicName>
|
||||
[p]skop,
|
||||
<quantity id="4C9A7594FF923842393DFA159331FA01" box="[513,599,1442,1463]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="22" pageNumber="343" unit="m" value="1700.0">
|
||||
<elevation id="004F3F42FF923842393DFA159331FA01" box="[513,599,1442,1463]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="22" pageNumber="343" unit="m" value="1700.0">1700 m</elevation>
|
||||
</quantity>
|
||||
<date id="FFDCFEB1FF9238423949FA159394FA01" box="[629,754,1442,1463]" pageId="22" pageNumber="343" value="1998-12-12">
|
||||
<collectingDate id="EF980759FF9238423949FA159394FA01" box="[629,754,1442,1463]" pageId="22" pageNumber="343" value="1998-12-12">12.12.1998</collectingDate>
|
||||
</date>
|
||||
; Kruger/DNA voucher BMNH 837896” (
|
||||
<collectionCode id="ED7340B4FF92384239A2FA77938AFA63" box="[670,748,1472,1493]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="343">TMSA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B0AD22CFF9238423BACFA689304F987" collectingDate="2008-11-21" collectingDateMax="2008-11-26" collectingDateMin="2008-11-21" collectionCode="SMNS" collectorName="W. Schawaller" country="South Africa" elevation="1450" location="RSA" pageId="22" pageNumber="343" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||||
<typeStatus id="54D966D3FF9238423BACFA689060FA42" box="[144,262,1503,1524]" pageId="22" pageNumber="343" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF9238423A22FA689036FA42" box="[286,336,1503,1524]" count="1" pageId="22" pageNumber="343" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D6413F8FF9238423A50FA6890FAFA42" box="[364,412,1503,1524]" count="1" pageId="22" pageNumber="343" type="female">1 ♀</specimenCount>
|
||||
“
|
||||
<collectingCountry id="F37598E1FF9238423A86FA689096FA42" box="[442,496,1503,1524]" name="South Africa" pageId="22" pageNumber="343">RSA</collectingCountry>
|
||||
:
|
||||
<taxonomicName id="4C62A3F2FF9238423934FA68932AF9A5" authority="Prov., Mariepskop Forest Reserve, 1300" authorityName="Prov., Mariepskop Forest Reserve" authorityYear="1300" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="343" phylum="Arthropoda" rank="species" species="mpumalanga">
|
||||
Mpumalanga Prov., Mariepskop Forest Reserve,
|
||||
<quantity id="4C9A7594FF9238423929FA4A932AF9A5" box="[533,588,1533,1555]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="22" pageNumber="343" unit="m" value="1450.0" valueMax="1600.0" valueMin="1300.0">
|
||||
<elevation id="004F3F42FF9238423929FA4A932AF9A5" box="[533,588,1533,1555]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="22" pageNumber="343" unit="m" value="1450.0" valueMax="1600.0" valueMin="1300.0">1300</elevation>
|
||||
</quantity>
|
||||
</taxonomicName>
|
||||
-1600 m,
|
||||
<date id="FFDCFEB1FF92384239F0FA4A906BF984" pageId="22" pageNumber="343" value="2008-11-21" valueMax="2008-11-26" valueMin="2008-11-21">
|
||||
<collectingDate id="EF980759FF92384239F0FA4A906BF984" pageId="22" pageNumber="343" value="2008-11-21" valueMax="2008-11-26" valueMin="2008-11-21">21.- 26.XI.2008</collectingDate>
|
||||
</date>
|
||||
, leg.
|
||||
<collectorName id="2697BDA7FF9238423A77F9AB9097F984" box="[331,497,1564,1586]" pageId="22" pageNumber="343">W. Schawaller</collectorName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF9238423930F9AB933CF987" box="[524,602,1564,1585]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="22" pageNumber="343">SMNS</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="3B0AD22CFF9238423953F9AB90EAF93B" collectingDate="2008-11-23" collectionCode="TMSA" collectorName="S. Afr. & Ruth M" country="South Africa" elevation="1613" latitude="-24.34" longLatPrecision="751" longitude="30.52" pageId="22" pageNumber="343" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
|
||||
<specimenCount id="9D6413F8FF9238423953F9AB93F4F987" box="[623,658,1564,1585]" count="2" pageId="22" pageNumber="343" type="female">2 ♀</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF923842399BF9AB9395F984" box="[679,755,1564,1586]" pageId="22" pageNumber="343">S. Afr.</collectorName>
|
||||
;
|
||||
<taxonomicName id="4C62A3F2FF9238423BACF98D90DFF9E6" authority="Mariepskop" authorityName="Prov., Mariepskop Forest Reserve" authorityYear="1300" box="[144,441,1594,1616]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="343" phylum="Arthropoda" rank="species" species="mpumalanga">Mpumalanga Mariepskop</taxonomicName>
|
||||
forest,
|
||||
<geoCoordinate id="EE56BEB6FF9238423924F98D9316F9F9" box="[536,624,1594,1615]" degrees="24.34" direction="south" orientation="latitude" pageId="22" pageNumber="343" precision="555" value="-24.34">24.34 S</geoCoordinate>
|
||||
<emphasis id="B9160463FF9238423940F98A93EEF9F9" box="[636,648,1597,1615]" italics="true" pageId="22" pageNumber="343">–</emphasis>
|
||||
<geoCoordinate id="EE56BEB6FF92384239A8F98D9388F9F9" box="[660,750,1594,1616]" degrees="30.52" direction="east" orientation="longitude" pageId="22" pageNumber="343" precision="555" value="30.52">30.52 E</geoCoordinate>
|
||||
/
|
||||
<date id="FFDCFEB1FF9238423BACF9EE906BF9D8" box="[144,269,1625,1646]" pageId="22" pageNumber="343" value="2008-11-23">
|
||||
<collectingDate id="EF980759FF9238423BACF9EE906BF9D8" box="[144,269,1625,1646]" pageId="22" pageNumber="343" value="2008-11-23">23.11.2008</collectingDate>
|
||||
</date>
|
||||
, E-Y:3802 night with torch,
|
||||
<quantity id="4C9A7594FF9238423950F9EE93A7F9D8" box="[620,705,1625,1647]" metricMagnitude="3" metricUnit="m" metricValue="1.613" pageId="22" pageNumber="343" unit="m" value="1613.0">
|
||||
<elevation id="004F3F42FF9238423950F9EE93A7F9D8" box="[620,705,1625,1647]" metricMagnitude="3" metricUnit="m" metricValue="1.613" pageId="22" pageNumber="343" unit="m" value="1613.0">1613 m</elevation>
|
||||
</quantity>
|
||||
leg.
|
||||
<collectorName id="2697BDA7FF9238423BACF9C0918EF93B" box="[144,232,1655,1677]" pageId="22" pageNumber="343">Ruth M</collectorName>
|
||||
uller € ” (
|
||||
<collectionCode id="ED7340B4FF9238423A0BF9CF90E3F93B" box="[311,389,1656,1677]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="343">TMSA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9238423BACF917959BFD52" blockId="22.[144,759,1696,1901]" lastBlockId="22.[807,1422,199,740]" pageId="22" pageNumber="343">
|
||||
<emphasis id="B9160463FF9238423BACF917907EF903" box="[144,280,1696,1717]" italics="true" pageId="22" pageNumber="343">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9238423AECF9179351F903" box="[464,567,1696,1718]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="22" pageNumber="343" unit="mm" value="10.6">10.6 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9238423BACF909918FF962" box="[144,233,1726,1748]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="22" pageNumber="343" unit="mm" value="7.1">7.1 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9238423AE4F9099355F962" box="[472,563,1726,1748]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="22" pageNumber="343" unit="mm" value="5.1">5.1 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus distinct; anterior margin of labroclypeus medially distinctly sinuate; margins of labroclypeus strongly reflexed; smooth area in front of eyes 1.5 times as wide as long; ratio of eye diameter/interocular width: 0.63. Pronotum apparently glabrous (with only a few erect setae anteriorly, otherwise with only minute setae); unicoloured; basal margin widely incomplete medially (equal to or more than scutellum width); larger punctures on pronotum very coarse. Hypomeron carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity condensed to longitudinal patches; impunctate (only striae punctate); adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle blunt; Mesosternal process long. Ratio of length of metepisternum/metacoxa: 1/1.33. Metatibia, ratio of width/length: 1/4.33. Metatarsi dorsally smooth; first metatarsomere slightly longer than subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF923842389FFD7992B8FD55" box="[931,990,718,740]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="22" pageNumber="343">Fig 4</figureCitation>
|
||||
Ac
|
||||
<emphasis id="B9160463FF92384238C0FD66956EFD55" box="[1020,1032,721,739]" italics="true" pageId="22" pageNumber="343">–</emphasis>
|
||||
Ae. Habitus:
|
||||
<figureCitation id="1359C4F4FF9238423FA3FD7995BDFD52" box="[1183,1243,718,740]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="22" pageNumber="343">Fig 4</figureCitation>
|
||||
Af.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF923842381BFCBB947FFCCA" blockId="22.[807,1421,779,893]" pageId="22" pageNumber="343">
|
||||
<emphasis id="B9160463FF923842381BFCBB92F8FC97" box="[807,926,780,801]" italics="true" pageId="22" pageNumber="343">Diagnosis:</emphasis>
|
||||
This species differs from all other known
|
||||
<taxonomicName id="4C62A3F2FF923842381BFC9D929FFCF6" box="[807,1017,810,832]" pageId="22" pageNumber="343">
|
||||
<emphasis id="B9160463FF923842381BFC9D92F9FC89" box="[807,927,810,831]" italics="true" pageId="22" pageNumber="343">Pleophylla</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
by having the ventral and dorsal lobes of the parameres completely fused with each other, and without setae on the parameres.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF923842381BFC129541FB18" blockId="22.[806,1422,932,1199]" pageId="22" pageNumber="343">
|
||||
<emphasis id="B9160463FF923842381BFC1292FCFC0C" box="[807,922,933,954]" italics="true" pageId="22" pageNumber="343">Variation:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9238423F44FC13947DFC0C" box="[1144,1307,932,954]" metricMagnitude="-2" metricUnit="m" metricValue="1.105" metricValueMax="1.15" metricValueMin="1.06" pageId="22" pageNumber="343" unit="mm" value="11.05" valueMax="11.5" valueMin="10.6">
|
||||
10.6
|
||||
<emphasis id="B9160463FF9238423F94FC1095D2FC0F" box="[1192,1204,935,953]" italics="true" pageId="22" pageNumber="343">–</emphasis>
|
||||
11.5 mm
|
||||
</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF92384238BBFC749569FC6E" box="[903,1039,963,984]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="8.1" metricValueMin="7.1" pageId="22" pageNumber="343" unit="mm" value="7.6" valueMax="8.1" valueMin="7.1">
|
||||
7.1
|
||||
<emphasis id="B9160463FF9238423895FC7292D3FC61" box="[937,949,965,983]" italics="true" pageId="22" pageNumber="343">–</emphasis>
|
||||
8.1 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9238423E3DFC7594EFFC6E" box="[1281,1417,962,984]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.6" metricValueMin="5.1" pageId="22" pageNumber="343" unit="mm" value="5.35" valueMax="5.6" valueMin="5.1">
|
||||
5.1
|
||||
<emphasis id="B9160463FF9238423E1FFC729449FC61" box="[1315,1327,965,983]" italics="true" pageId="22" pageNumber="343">–</emphasis>
|
||||
5.6 mm
|
||||
</quantity>
|
||||
. Density of dark spots may vary, but never completely without spots. Female: antennal club of female straight, composed of five antennomeres, as long as the remaining antennomeres combined; sixth antennomere shorter than a quarter of club length. Eyes smaller than in male: ratio of eye diameter/ interocular width: 0.5.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF923842381AFB6195B1FABF" blockId="22.[806,1420,1238,1290]" pageId="22" pageNumber="343">
|
||||
<emphasis id="B9160463FF923842381AFB6192C2FB5D" box="[806,932,1238,1259]" italics="true" pageId="22" pageNumber="343">Etymology:</emphasis>
|
||||
The new species is named after one of its collectors, Martin Kr uger € (
|
||||
<collectionCode id="ED7340B4FF9238423F42FB4395AAFABF" box="[1150,1228,1268,1289]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="343">TMSA</collectionCode>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF923842381AFA8695EBF921" blockId="22.[806,1421,1329,1749]" pageId="22" pageNumber="343">
|
||||
<emphasis id="B9160463FF923842381AFA8692F4FAF0" box="[806,914,1329,1350]" italics="true" pageId="22" pageNumber="343">Remarks:</emphasis>
|
||||
The systematic placement of this species is still uncertain. The species differs clearly from all other
|
||||
<taxonomicName id="4C62A3F2FF9238423844FAD9953EFA32" box="[888,1112,1390,1412]" pageId="22" pageNumber="343">
|
||||
<emphasis id="B9160463FF9238423844FAD99296FA35" box="[888,1008,1390,1411]" italics="true" pageId="22" pageNumber="343">Pleophylla</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
, which might justify a separate genus. Many symplesiomorphies of
|
||||
<taxonomicName id="4C62A3F2FF923842381AFA1B92F8FA77" authorityName="Erichson" authorityYear="1847" box="[806,926,1452,1473]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="343" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B9160463FF923842381AFA1B92F8FA77" box="[806,926,1452,1473]" italics="true" pageId="22" pageNumber="343">Pleophylla</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C62A3F2FF92384238CEFA1B950CFA77" baseAuthorityName="Ahrens" baseAuthorityYear="2006" box="[1010,1130,1452,1473]" class="Insecta" family="Scarabaeidae" genus="Omaloplia" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="343" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B9160463FF92384238CEFA1B950CFA77" box="[1010,1130,1452,1473]" italics="true" pageId="22" pageNumber="343">Omaloplia</emphasis>
|
||||
</taxonomicName>
|
||||
are absent from this species, such as the separate dorsal and ventral lobes on the parameres or the pilosity of parameres. However,
|
||||
<taxonomicName id="4C62A3F2FF923842389FF9BF9544F9AA" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" box="[931,1058,1543,1565]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="343" phylum="Arthropoda" rank="species" species="kruegeri">
|
||||
<emphasis id="B9160463FF923842389FF9BF9544F9AA" box="[931,1058,1543,1565]" italics="true" pageId="22" pageNumber="343">P. kruegeri</emphasis>
|
||||
</taxonomicName>
|
||||
shares with
|
||||
<taxonomicName id="4C62A3F2FF9238423FEFF9B0942DF9AA" baseAuthorityName="Ahrens" baseAuthorityYear="2006" box="[1235,1355,1543,1564]" class="Insecta" family="Scarabaeidae" genus="Omaloplia" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="343" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B9160463FF9238423FEFF9B0942DF9AA" box="[1235,1355,1543,1564]" italics="true" pageId="22" pageNumber="343">Omaloplia</emphasis>
|
||||
</taxonomicName>
|
||||
only two plesiomorphies and therefore it does not occupy a basal position amongst the
|
||||
<taxonomicName id="4C62A3F2FF9238423FAFF9F3940FF9EC" box="[1171,1385,1604,1626]" pageId="22" pageNumber="343">
|
||||
<emphasis id="B9160463FF9238423FAFF9F3946DF9EF" box="[1171,1291,1604,1625]" italics="true" pageId="22" pageNumber="343">Pleophylla</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
in the cladistic analysis (
|
||||
<figureCitation id="1359C4F4FF9238423F75F9D495E8F9CE" box="[1097,1166,1635,1657]" captionStart="Figure 9" captionStartId="28.[144,223,1791,1810]" captionTargetBox="[151,1413,580,1756]" captionTargetId="graphics-206@28.[151,1269,580,1747]" captionTargetPageId="28" captionText="Figure 9. Strict consensus tree of the WINCLADA/NONA tree search (parsimony ratchet). Black squares, unambiguous apomorphies; white squares, unambiguous but to some degree homoplastic apomorphies. Small numbers above squares are characters; small numbers below squares are character states. Bootstrap values> 50 are shown in large and bold font under branches." figureDoi="http://doi.org/10.5281/zenodo.14807101" httpUri="https://zenodo.org/record/14807101/files/figure.png" pageId="22" pageNumber="343">Fig. 9</figureCitation>
|
||||
), a prerequisite for placement in a separate genus.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9238423803F917940FF963" blockId="22.[806,1421,1329,1749]" pageId="22" pageNumber="343">
|
||||
In a previous publication we referred to this species as ‘SpM-24’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF9238423FFDF908943BF963" author="Ahrens D" box="[1217,1373,1727,1749]" pageId="22" pageNumber="343" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9238423864F8BC943DF8A9" blockId="22.[856,1371,1803,1824]" box="[856,1371,1803,1824]" pageId="22" pageNumber="343">
|
||||
<smallCapsWord id="8D3B4EADFF9238423864F8BC92ADF8A9" baselines="1819,1819" box="[856,971,1803,1824]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Patterns" pageId="22" pageNumber="343">PATTERNS</smallCapsWord>
|
||||
<smallCapsWord id="8D3B4EADFF92384238E9F8B99294F8A9" baselines="1819" box="[981,1010,1806,1823]" lowerCaseFontSize="7" mainFontSize="9" normCase="lower" normString="of" pageId="22" pageNumber="343">OF</smallCapsWord>
|
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<smallCapsWord id="8D3B4EADFF92384238C7F8B995F0F8A9" baselines="1819" box="[1019,1174,1806,1823]" lowerCaseFontSize="7" mainFontSize="9" normCase="lower" normString="distribution" pageId="22" pageNumber="343">DISTRIBUTION</smallCapsWord>
|
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<smallCapsWord id="8D3B4EADFF9238423FA3F8B995ABF8A9" baselines="1819" box="[1183,1229,1806,1823]" lowerCaseFontSize="7" mainFontSize="9" normCase="lower" normString="and" pageId="22" pageNumber="343">AND</smallCapsWord>
|
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<smallCapsWord id="8D3B4EADFF9238423FEAF8B9943DF8A9" baselines="1819" box="[1238,1371,1806,1823]" lowerCaseFontSize="7" mainFontSize="9" normCase="lower" normString="phenology" pageId="22" pageNumber="343">PHENOLOGY</smallCapsWord>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF923843381BF88F9181FD10" blockId="22.[807,1421,1848,1901]" lastBlockId="23.[166,781,199,862]" lastPageId="23" lastPageNumber="344" pageId="22" pageNumber="343">
|
||||
Although the distribution of
|
||||
<taxonomicName id="4C62A3F2FF9238423FABF88F9469F8FB" authorityName="Erichson" authorityYear="1847" box="[1175,1295,1848,1869]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="343" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B9160463FF9238423FABF88F9469F8FB" box="[1175,1295,1848,1869]" italics="true" pageId="22" pageNumber="343">Pleophylla</emphasis>
|
||||
</taxonomicName>
|
||||
in
|
||||
<collectingCountry id="F37598E1FF9238423E76F88F920AF8DB" name="South Africa" pageId="22" pageNumber="343">South Africa</collectingCountry>
|
||||
is focussed to the eastern and southern parts of the country, its northern occurrences extend patchily along the African Eastern Arc Mountains to the
|
||||
<collectingCountry id="F37598E1FF9338433B9AFEB39099FEAF" box="[166,511,260,282]" name="Democratic Republic of the Congo" pageId="23" pageNumber="344">Democratic Republic of Congo</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="F37598E1FF9338433906FEB393F2FEAC" box="[570,660,260,282]" name="Uganda" pageId="23" pageNumber="344">Uganda</collectingCountry>
|
||||
as well as
|
||||
<collectingCountry id="F37598E1FF9338433B9AFE949195FE8E" box="[166,243,291,312]" name="Kenya" pageId="23" pageNumber="344">Kenya</collectingCountry>
|
||||
. Between these areas records are lacking over long distances. Current distribution data suggest that the species of
|
||||
<taxonomicName id="4C62A3F2FF9338433AB1FEE89363FEC2" authorityName="Erichson" authorityYear="1847" box="[397,517,351,372]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B9160463FF9338433AB1FEE89363FEC2" box="[397,517,351,372]" italics="true" pageId="23" pageNumber="344">Pleophylla</emphasis>
|
||||
</taxonomicName>
|
||||
are closely associated with the Afrotropical forest, but as far as known, they are absent from the tropical lowland forests of western Central Africa and western Africa. The ranges of many species overlap, at least partly, and numerous taxa co-occur syntopically (
|
||||
<tableCitation id="C6E0EDCAFF9338433960FE4F93D7FDBB" box="[604,689,504,526]" captionStart="Table 1" captionStartId="24.[87,153,504,525]" captionText="Table 1. Number of pairwise co-occurrences of the Pleophylla species compared to the number of sites with only one recorded species (i.e. spatially unique samples)." pageId="23" pageNumber="344">Table 1</tableCitation>
|
||||
); so far we have found up to five species in the same locality (in
|
||||
<collectingCountry id="F37598E1FF9338433BF3FD829002FDFD" box="[207,356,565,587]" name="South Africa" pageId="23" pageNumber="344">South Africa</collectingCountry>
|
||||
, Natal, Karkloof forest). The species with the widest range is
|
||||
<taxonomicName id="4C62A3F2FF9338433AE5FDE293F1FDDF" box="[473,663,596,618]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="344" phylum="Arthropoda" rank="species" species="fasciatipennis">
|
||||
<emphasis id="B9160463FF9338433AE5FDE293F1FDDF" box="[473,663,596,618]" italics="true" pageId="23" pageNumber="344">P. fasciatipennis</emphasis>
|
||||
</taxonomicName>
|
||||
, which is found from north-eastern
|
||||
<collectingCountry id="F37598E1FF9338433AE0FDC59316FD3E" box="[476,624,626,648]" name="South Africa" pageId="23" pageNumber="344">South Africa</collectingCountry>
|
||||
south to the
|
||||
<collectingRegion id="49A61693FF9338433B9AFD269185FD10" box="[166,227,657,678]" country="South Africa" name="Western Cape" pageId="23" pageNumber="344">Cape</collectingRegion>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9338433B83FD18908CFCE8" blockId="23.[166,781,199,862]" pageId="23" pageNumber="344">
|
||||
The main temporal occurrence in terms of numbers of species and individuals is during November and December; after February species richness and abundance drop considerably (
|
||||
<figureCitation id="1359C4F4FF933843393BFCBC932DFC96" box="[519,587,779,801]" captionStart="Figure 8" captionStartId="25.[166,245,1510,1529]" captionTargetBox="[396,1216,201,1477]" captionTargetPageId="25" captionText="Figure 8. Phenology of Pleophylla in terms of species richness (below) and numbers of recorded adult specimens (above)." figureDoi="http://doi.org/10.5281/zenodo.14807099" httpUri="https://zenodo.org/record/14807099/files/figure.png" pageId="23" pageNumber="344">Fig. 8</figureCitation>
|
||||
). During May to August records are very rare, except in June when records are entirely lacking.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
223
data/03/CB/69/03CB6967FF9438443B85FD16953DFE13.xml
Normal file
223
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Normal file
|
|
@ -0,0 +1,223 @@
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|
||||
<heading id="D0956F1DFF9438443B85FD169348FD63" centered="true" fontSize="9" level="2" pageId="16" pageNumber="337" reason="2">
|
||||
<taxonomicName id="4C62A3F2FF9438443B85FD1693ABFD00" ID-CoL="4K848" authority="BURMEISTER, 1855" authorityName="Burmeister" authorityYear="1855" box="[185,717,668,695]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="species" species="navicularis">
|
||||
<emphasis id="B9160463FF9438443B85FD169083FD02" box="[185,485,673,694]" italics="true" pageId="16" pageNumber="337">
|
||||
<smallCapsWord id="8D3B4EADFF9438443B85FD16902FFD02" baselines="689,688" box="[185,329,673,694]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="16" pageNumber="337">PLEOPHYLLA</smallCapsWord>
|
||||
<smallCapsWord id="8D3B4EADFF9438443A6FFD139083FD02" baselines="688" box="[339,485,676,692]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="navicularis" pageId="16" pageNumber="337">NAVICULARIS</smallCapsWord>
|
||||
</emphasis>
|
||||
<bibRefCitation id="EFF3A580FF9438443AD2FD2993ABFD00" author="Burmeister H" box="[494,717,668,695]" pageId="16" pageNumber="337" pagination="1 - 467" refId="ref23262" refString="Burmeister H. 1855. Handbuch der Entomologie. 4. Band. Besondere Entomologie, Fortsetzung. 2. Abteilung, Coleoptera Lamellicornia Phyllophaga chaenochela. Berlin: Theodor Johann Christian Friedrich Enslin, 1-467." type="book chapter" year="1855">
|
||||
<smallCapsWord id="8D3B4EADFF9438443AD2FD29931BFD03" baselines="688,689" box="[494,637,670,694]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Burmeister" pageId="16" pageNumber="337">BURMEISTER</smallCapsWord>
|
||||
, 1855
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF9438443A5DFD0A90BFFD63" box="[353,473,699,725]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="16" pageNumber="337">
|
||||
<smallCapsWord id="8D3B4EADFF9438443A5DFD0A90F0FD65" baselines="719,719" box="[353,406,701,725]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="16" pageNumber="337">FIGS</smallCapsWord>
|
||||
3I
|
||||
<emphasis id="B9160463FF9438443A86FD0890AEFD63" box="[442,456,703,725]" italics="true" pageId="16" pageNumber="337">–</emphasis>
|
||||
L
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF9438443AD6FD0C9362FD63" box="[490,516,699,725]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="16" pageNumber="337">5I</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF9438443BACFD5B9067FC2C" pageId="16" pageNumber="337" type="reference_group">
|
||||
<paragraph id="8BDDD871FF9438443BACFD5B93E7FC89" blockId="16.[144,759,748,922]" pageId="16" pageNumber="337">
|
||||
<treatmentCitationGroup id="AB72FF5FFF9438443BACFD5B93E7FC89" pageId="16" pageNumber="337">
|
||||
<treatmentCitation id="0AC3FE60FF9438443BACFD5B93C9FCB4" author="Burmeister H" box="[144,687,748,770]" page="181" pageId="16" pageNumber="337" year="1855">
|
||||
<taxonomicName id="4C62A3F2FF9438443BACFD5B93C9FCB4" ID-CoL="4K848" authority="Burmeister, 1855: 181" authorityName="Burmeister" authorityPageNumber="181" authorityYear="1855" box="[144,687,748,770]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="species" species="navicularis">
|
||||
<emphasis id="B9160463FF9438443BACFD5B90F0FCB7" box="[144,406,748,769]" italics="true" pageId="16" pageNumber="337">Pleophylla navicularis</emphasis>
|
||||
<bibRefCitation id="EFF3A580FF9438443A98FD5B93C9FCB4" author="Burmeister H" box="[420,687,748,770]" pageId="16" pageNumber="337" pagination="1 - 467" refId="ref23262" refString="Burmeister H. 1855. Handbuch der Entomologie. 4. Band. Besondere Entomologie, Fortsetzung. 2. Abteilung, Coleoptera Lamellicornia Phyllophaga chaenochela. Berlin: Theodor Johann Christian Friedrich Enslin, 1-467." type="book chapter" year="1855">Burmeister, 1855: 181</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
[
|
||||
<typeStatus id="54D966D3FF94384439FFFD5A9390FCB4" box="[707,758,749,770]" pageId="16" pageNumber="337">type</typeStatus>
|
||||
locality: Weihnachtsbai (Hoppig)];
|
||||
<treatmentCitation id="0AC3FE60FF943844390CFCBC91DDFC89" author="Schaufuss LW" page="231" pageId="16" pageNumber="337" year="1871">
|
||||
<bibRefCitation id="EFF3A580FF943844390CFCBC91DDFC89" author="Schaufuss LW" pageId="16" pageNumber="337" pagination="231 - 233" refId="ref24095" refString="Schaufuss LW. 1871. Die Arten der Gattung Pleophylla Er. Nunquam Otiosus I 6: 231-233." type="journal article" year="1871">Schaufuss, 1871: 231</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
; Brenske, 1900: 83; Peringuey, 1904: 7.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9438443BADFCFF9067FC2C" blockId="16.[144,759,748,922]" pageId="16" pageNumber="337">
|
||||
<treatmentCitationGroup id="AB72FF5FFF9438443BADFCFF9067FC2C" pageId="16" pageNumber="337">
|
||||
<treatmentCitation id="0AC3FE60FF9438443BADFCFF93CDFCEB" author="Schaufuss LW" box="[145,683,840,862]" page="232" pageId="16" pageNumber="337" year="1871">
|
||||
<taxonomicName id="4C62A3F2FF9438443BADFCFF93CDFCEB" ID-CoL="4K83Y" authority="Schaufuss, 1871: 232" authorityName="Schaufuss" authorityPageNumber="232" authorityYear="1871" box="[145,683,840,862]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="species" species="flavicornis">
|
||||
<emphasis id="B9160463FF9438443BADFCFF90F4FCEB" box="[145,402,840,861]" italics="true" pageId="16" pageNumber="337">Pleophylla flavicornis</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9438443A98FCFF93CDFCEB" box="[420,683,840,862]" country="Germany" location="Brenske" pageId="16" pageNumber="337" specimenCount="1">
|
||||
<bibRefCitation id="EFF3A580FF9438443A98FCFF93CDFCEB" author="Schaufuss LW" box="[420,683,840,862]" pageId="16" pageNumber="337" pagination="231 - 233" refId="ref24095" refString="Schaufuss LW. 1871. Die Arten der Gattung Pleophylla Er. Nunquam Otiosus I 6: 231-233." type="journal article" year="1871">Schaufuss, 1871: 232</bibRefCitation>
|
||||
</materialsCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
[
|
||||
<typeStatus id="54D966D3FF94384439FFFCFE9390FCE8" box="[707,758,841,862]" pageId="16" pageNumber="337">type</typeStatus>
|
||||
locality: Caffr., New
|
||||
<collectingCountry id="F37598E1FF9438443AB3FCD09098FCCA" box="[399,510,871,892]" name="Germany" pageId="16" pageNumber="337">Germany</collectingCountry>
|
||||
];
|
||||
<location id="8EBD8EAAFF943844392AFCD09311FCCB" LSID="urn:lsid:plazi:treatment:03CB6967FF9438443B85FD16953DFE13:8EBD8EAAFF943844392AFCD09311FCCB" box="[534,631,871,893]" country="Germany" name="Brenske" pageId="16" pageNumber="337">Brenske</location>
|
||||
, 1900: 83,
|
||||
<emphasis id="B9160463FF9438443BACFC329067FC2C" bold="true" box="[144,257,901,922]" pageId="16" pageNumber="337">syn. nov.</emphasis>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF9438443BACFC74953DFE13" pageId="16" pageNumber="337" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF9438443BACFC74908BFB18" blockId="16.[144,759,962,1198]" pageId="16" pageNumber="337">
|
||||
<emphasis id="B9160463FF9438443BACFC7490A9FC61" box="[144,463,962,984]" italics="true" pageId="16" pageNumber="337">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9438443AE1FC74903FFBE4" collectionCode="MLUH" collectorName="Pt. Nat. Gm." pageId="16" pageNumber="337" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="lectotype">
|
||||
<typeStatus id="54D966D3FF9438443AE1FC74932DFC6E" box="[477,587,963,984]" pageId="16" pageNumber="337" type="lectotype">Lectotype</typeStatus>
|
||||
(
|
||||
<taxonomicName id="4C62A3F2FF9438443953FC759396FC61" authorityName="Burmeister" authorityYear="1855" box="[623,752,962,983]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="species" species="navicularis">
|
||||
<emphasis id="B9160463FF9438443953FC759396FC61" box="[623,752,962,983]" italics="true" pageId="16" pageNumber="337">navicularis</emphasis>
|
||||
</taxonomicName>
|
||||
, here designated):
|
||||
<specimenCount id="9D6413F8FF9438443A5CFC539016FC40" box="[352,368,996,1014]" count="1" pageId="16" pageNumber="337" type="male">♂</specimenCount>
|
||||
<taxonomicName id="4C62A3F2FF9438443A47FC569325FC41" authority="Nab." authorityName="Nab." box="[379,579,993,1015]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="species" species="navicularis">“navicularis Nab.</taxonomicName>
|
||||
subcylindr.
|
||||
<collectorName id="2697BDA7FF94384439E9FC569065FBA3" pageId="16" pageNumber="337">Pt. Nat. Gm.</collectorName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF9438443A19FBB7901CFBA3" box="[293,378,1024,1045]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="16" pageNumber="337">MLUH</collectionCode>
|
||||
). Paralectoptype:
|
||||
<specimenCount id="9D6413F8FF9438443965FBB793E3FBA2" box="[601,645,1024,1045]" count="1" pageId="16" pageNumber="337" type="female">1 ♀</specimenCount>
|
||||
[without labels, provided with a copy of the original collection label] (
|
||||
<collectionCode id="ED7340B4FF9438443BC0FB8A9037FBE4" box="[252,337,1085,1106]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="16" pageNumber="337">MLUH</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B0AD22CFF9438443ABFFB8A908FFB18" collectionCode="ZMHB" collectorName="N. Germ. & Coll. L. W. Schaufuss" pageId="16" pageNumber="337" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
|
||||
<typeStatus id="54D966D3FF9438443ABFFB8A9097FBE4" box="[387,497,1085,1106]" pageId="16" pageNumber="337" type="lectotype">Lectotype</typeStatus>
|
||||
(
|
||||
<taxonomicName id="4C62A3F2FF943844391CFB8A93F1FBE4" authorityName="Schaufuss" authorityYear="1871" box="[544,663,1085,1106]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="species" species="flavicornis">
|
||||
<emphasis id="B9160463FF943844391CFB8A93F1FBE4" box="[544,663,1085,1106]" italics="true" pageId="16" pageNumber="337">flavicornis</emphasis>
|
||||
</taxonomicName>
|
||||
, here designated):
|
||||
<specimenCount id="9D6413F8FF9438443A19FBE99054FBC6" box="[293,306,1118,1136]" count="1" pageId="16" pageNumber="337" type="female">♀</specimenCount>
|
||||
“Type./flavicornis m.
|
||||
<collectorName id="2697BDA7FF9438443908FBEB93F9FBC6" box="[564,671,1115,1137]" pageId="16" pageNumber="337">N. Germ.</collectorName>
|
||||
/Nunqu at. pag. 232/
|
||||
<collectorName id="2697BDA7FF9438443A1AFBCD9373FB26" box="[294,533,1146,1168]" pageId="16" pageNumber="337">Coll. L.W. Schaufuss</collectorName>
|
||||
/
|
||||
<taxonomicName id="4C62A3F2FF9438443922FBCD919EFB18" authority="Schauf. Caffraria" authorityName="Schauf. Caffraria" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="species" species="flavicornis">flavicornis Schauf. Caffraria</taxonomicName>
|
||||
/
|
||||
<taxonomicName id="4C62A3F2FF9438443A3CFB2F9011FB18" authority="(ZMHB)" authorityName="Erichson" authorityYear="1847" baseAuthorityName="ZMHB" box="[256,375,1176,1198]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="genus">Pleophylla</taxonomicName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF9438443AACFB2E9084FB18" box="[400,482,1177,1198]" pageId="16" pageNumber="337">ZMHB</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9438443BACFB6293A0FB5D" blockId="16.[144,710,1237,1259]" box="[144,710,1237,1259]" pageId="16" pageNumber="337">
|
||||
<emphasis id="B9160463FF9438443BACFB629089FB5C" box="[144,495,1237,1258]" italics="true" pageId="16" pageNumber="337">Additional material examined:</emphasis>
|
||||
See Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9438443BACFAA594E1FE8E" blockId="16.[144,759,1298,1901]" lastBlockId="16.[807,1422,199,312]" pageId="16" pageNumber="337">
|
||||
<emphasis id="B9160463FF9438443BACFAA59057FA91" box="[144,305,1298,1319]" italics="true" pageId="16" pageNumber="337">Redescription:</emphasis>
|
||||
Body length: 13.0 mm, elytral length:
|
||||
<quantity id="4C9A7594FF9438443BACFA86918FFAF1" box="[144,233,1329,1351]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="16" pageNumber="337" unit="mm" value="9.4">9.4 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9438443AE4FA869355FAF1" box="[472,563,1329,1351]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="16" pageNumber="337" unit="mm" value="6.6">6.6 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes three times as wide as long; ratio of eye diameter/interocular width: 0.68. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra distinctly elevated but convex; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.47. Metatibia, ratio of width/length: 1/4.3. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF943844381BFE9592F0FE8E" box="[807,918,290,312]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="16" pageNumber="337">
|
||||
Fig. 3I
|
||||
<emphasis id="B9160463FF943844384FFE929219FE81" box="[883,895,293,311]" italics="true" pageId="16" pageNumber="337">–</emphasis>
|
||||
K
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF9438443F34FE959539FE8E" box="[1032,1119,290,312]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="16" pageNumber="337">
|
||||
Fig.
|
||||
<specimenCount id="9D6413F8FF9438443F01FE949539FE8E" box="[1085,1119,291,312]" count="3" pageId="16" pageNumber="337" type="generic">3L</specimenCount>
|
||||
</figureCitation>
|
||||
. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF9438443E08FE9594E5FE8E" box="[1332,1411,290,312]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="16" pageNumber="337">Fig. 5I</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF943844381BFEE5953DFE13" blockId="16.[807,1421,338,422]" pageId="16" pageNumber="337">
|
||||
<emphasis id="B9160463FF943844381BFEE592F4FED1" box="[807,914,338,359]" italics="true" pageId="16" pageNumber="337">Remarks:</emphasis>
|
||||
In previous publications we referred to this species as ‘sp11’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF9438443E08FEC69206FE13" author="Ahrens D" pageId="16" pageNumber="337" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFF3A580FF9438443850FE279536FE13" author="Eberle J & Warnock RCM & Ahrens D" box="[876,1104,399,422]" pageId="16" pageNumber="337" refId="ref23428" refString="Eberle J, Warnock RCM, Ahrens D. 2016 b. Bayesian species delimitation in Pleophylla chafers (Coleoptera) - the importance of prior choice and morphology. BMC Evolutionary Biology 16: 94." type="journal volume" year="2016">
|
||||
Eberle
|
||||
<emphasis id="B9160463FF9438443882FE27929FFE13" box="[958,1017,400,421]" italics="true" pageId="16" pageNumber="337">et al.</emphasis>
|
||||
, 2016b
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
226
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226
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<taxonomicName id="4C62A3F2FF9438443816FE619280FDBF" ID-CoL="4K84F" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="species" species="silvatica" status="sp. nov.">
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,
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,
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&
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.
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.
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(
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<figureCitation id="1359C4F4FF9438443F5BFE44958BFDBD" box="[1127,1261,497,523]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="16" pageNumber="337">
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|
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3M
|
||||
<emphasis id="B9160463FF9438443FF2FE4395BAFDBC" box="[1230,1244,500,522]" italics="true" pageId="16" pageNumber="337">–</emphasis>
|
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P
|
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</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF9438443FC2FE46947AFDBD" box="[1278,1308,497,523]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="16" pageNumber="337">5J</figureCitation>
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, 6)
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urn:lsid:zoobank.org:act:
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<emphasis id="B9160463FF943844381BFDCD9535FD39" box="[807,1107,634,655]" italics="true" pageId="16" pageNumber="337">Type material examined:</emphasis>
|
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<materialsCitation id="3B0AD22CFF9438443F5DFDCD95E4FD5C" collectingDate="1986-08-12" collectionCode="TMSA" collectorName="S. Afr. & E. Transvaal & Endrody-Younga" latitude="-25.31" location="Karst" longLatPrecision="748" longitude="30.46" pageId="16" pageNumber="337" specimenCount="1" specimenCount-male="1" stateProvince="Berlin" typeStatus="holotype">
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|
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:
|
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<specimenCount id="9D6413F8FF9438443FE3FDCB9589FD38" box="[1247,1263,636,654]" count="1" pageId="16" pageNumber="337" type="male">♂</specimenCount>
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“
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,
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plat.
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-
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||||
/
|
||||
<date id="FFDCFEB1FF943844381BFD0092C6FD7B" box="[807,928,695,717]" pageId="16" pageNumber="337" value="1986-08-12">
|
||||
<collectingDate id="EF980759FF943844381BFD0092C6FD7B" box="[807,928,695,717]" pageId="16" pageNumber="337" value="1986-08-12">08.12.1986</collectingDate>
|
||||
</date>
|
||||
E-Y: 2363 fungous Pinus logs leg.
|
||||
<collectorName id="2697BDA7FF943844381BFD629281FD5D" box="[807,999,725,747]" pageId="16" pageNumber="337">Endrody-Younga</collectorName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF9438443F11FD62951DFD5C" box="[1069,1147,725,746]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="16" pageNumber="337">TMSA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<typeStatus id="54D966D3FF9438443F87FD619457FD5D" box="[1211,1329,726,747]" pageId="16" pageNumber="337" type="paratype">Paratypes</typeStatus>
|
||||
: see Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF943844381BFC9F95C0FA4A" blockId="16.[807,1422,808,1532]" pageId="16" pageNumber="337">
|
||||
<emphasis id="B9160463FF943844381BFC9F92C9FC8B" box="[807,943,808,829]" italics="true" pageId="16" pageNumber="337">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9438443F5BFC9F95A8FC8B" box="[1127,1230,808,829]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="16" pageNumber="337" unit="mm" value="13.4">13.4 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF943844381BFCF192E6FCEA" box="[807,896,838,860]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="16" pageNumber="337" unit="mm" value="9.3">9.3 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9438443F53FCF195AFFCEA" box="[1135,1225,838,860]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="16" pageNumber="337" unit="mm" value="7.1">7.1 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes three times as wide as long; ratio of eye diameter/interocular width: 0.67. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra distinctly elevated but convex; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.54. Metatibia, ratio of width/length: 1/4.14. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF9438443FAFFA7F9469FA6B" box="[1171,1295,1480,1502]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="16" pageNumber="337">
|
||||
Fig. 3M
|
||||
<emphasis id="B9160463FF9438443FD1FA7C959FFA6B" box="[1261,1273,1483,1501]" italics="true" pageId="16" pageNumber="337">–</emphasis>
|
||||
O
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF943844381BFA51921BFA4A" box="[807,893,1510,1532]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="16" pageNumber="337">
|
||||
Fig.
|
||||
<specimenCount id="9D6413F8FF9438443860FA50921BFA4A" box="[860,893,1511,1532]" count="3" pageId="16" pageNumber="337" type="generic">3P</specimenCount>
|
||||
</figureCitation>
|
||||
. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF9438443F73FA5195C4FA4A" box="[1103,1186,1510,1532]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="16" pageNumber="337">Fig. 5J</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF943844381BF9AE9566F953" blockId="16.[807,1421,1560,1765]" pageId="16" pageNumber="337">
|
||||
<emphasis id="B9160463FF943844381BF9AE95C7F99B" box="[807,1185,1560,1582]" italics="true" pageId="16" pageNumber="337">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF9438443890F9AF95C7F99B" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[940,1185,1560,1581]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="species" species="silvatica" status="sp. nov.">Pleophylla silvatica</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF9438443F82F9AE944DF998" box="[1214,1323,1561,1582]" pageId="16" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is in external and genital shape similar to
|
||||
<taxonomicName id="4C62A3F2FF9438443FD9F98094E0F9FA" box="[1253,1414,1591,1612]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="species" species="navicularis">
|
||||
<emphasis id="B9160463FF9438443FD9F98094E0F9FA" box="[1253,1414,1591,1612]" italics="true" pageId="16" pageNumber="337">P. navicularis</emphasis>
|
||||
</taxonomicName>
|
||||
. It differs from the latter in having the two narrow basal branches of both parameres (dorsal lobe) subequal in length, whereas in
|
||||
<taxonomicName id="4C62A3F2FF9438443F8BF924943FF911" box="[1207,1369,1682,1704]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="337" phylum="Arthropoda" rank="species" species="navicularis">
|
||||
<emphasis id="B9160463FF9438443F8BF924943FF911" box="[1207,1369,1682,1704]" italics="true" pageId="16" pageNumber="337">P. navicularis</emphasis>
|
||||
</taxonomicName>
|
||||
the basal branches of both parameres (dorsal lobe) are different in length.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF943845381BF94B90C1FEE1" blockId="16.[807,1422,1788,1901]" lastBlockId="17.[166,781,199,343]" lastPageId="17" lastPageNumber="338" pageId="16" pageNumber="337">
|
||||
<emphasis id="B9160463FF943844381BF94B92FCF8A7" box="[807,922,1788,1809]" italics="true" pageId="16" pageNumber="337">Variation:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9438443F7AF94B95BCF8A7" box="[1094,1242,1788,1810]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.34" metricValueMin="0.86" pageId="16" pageNumber="337" unit="mm" value="11.0" valueMax="13.4" valueMin="8.6">
|
||||
8.6
|
||||
<emphasis id="B9160463FF9438443F54F9499512F8A6" box="[1128,1140,1790,1808]" italics="true" pageId="16" pageNumber="337">–</emphasis>
|
||||
13.4 mm
|
||||
</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF943844381BF8AD92C8F886" box="[807,942,1818,1840]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" metricValueMax="9.4" metricValueMin="6.4" pageId="16" pageNumber="337" unit="mm" value="7.9" valueMax="9.4" valueMin="6.4">
|
||||
6.4
|
||||
<emphasis id="B9160463FF9438443875F8AA9233F899" box="[841,853,1821,1839]" italics="true" pageId="16" pageNumber="337">–</emphasis>
|
||||
9.4 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9438443FA6F8AD9444F886" box="[1178,1314,1818,1840]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="7.1" metricValueMin="4.6" pageId="16" pageNumber="337" unit="mm" value="5.85" valueMax="7.1" valueMin="4.6">
|
||||
4.6
|
||||
<emphasis id="B9160463FF9438443F80F8AA95AEF899" box="[1212,1224,1821,1839]" italics="true" pageId="16" pageNumber="337">–</emphasis>
|
||||
7.1 mm
|
||||
</quantity>
|
||||
. Density of dark spot may vary, but never completely without spots. Female: antennal club of female straight, composed of five antennomeres, slightly longer than the remaining antennomeres combined; sixth antennomere short, half as long as club length. Eyes smaller than in male: ratio of eye diameter/ interocular width: 0.5.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9538453B9AFED8939DFE15" blockId="17.[166,781,367,419]" pageId="17" pageNumber="338">
|
||||
<emphasis id="B9160463FF9538453B9AFED89042FE32" box="[166,292,367,388]" italics="true" pageId="17" pageNumber="338">Etymology:</emphasis>
|
||||
The new species is named
|
||||
<taxonomicName id="4C62A3F2FF953845399EFED8926EFE32" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[674,776,367,388]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="silvatica">
|
||||
‘
|
||||
<emphasis id="B9160463FF9538453994FED8926EFE32" box="[680,776,367,388]" italics="true" pageId="17" pageNumber="338">silvatica</emphasis>
|
||||
</taxonomicName>
|
||||
’ following its strict occurrence only in native forests.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9538453B9AFE7293D8FD80" blockId="17.[166,781,453,566]" pageId="17" pageNumber="338">
|
||||
<emphasis id="B9160463FF9538453B9AFE729074FE6C" box="[166,274,453,474]" italics="true" pageId="17" pageNumber="338">Remarks:</emphasis>
|
||||
The northern and southern populations are slightly differentiated in shape of parameres. In a previous publication we referred to this species as ‘SpeciesM-1’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF953845392BFD9793D4FD80" author="Ahrens D" box="[535,690,544,566]" pageId="17" pageNumber="338" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
221
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Normal file
221
data/03/CB/69/03CB6967FF9538453B87FD3794F4FD22.xml
Normal file
|
|
@ -0,0 +1,221 @@
|
|||
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<mods:mods id="626B7E57CA79C7B7AF0060C1601A03AA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="ADED020781C58673F341265384708DB1">
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<mods:title id="751A9309A0CA7EE8F6995B93831AA7E1">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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</mods:titleInfo>
|
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<mods:role id="9AB671204249D691014B7A09AC27C90E">
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<mods:roleTerm id="3028CE7093D97D266E250F28717912AF">Author</mods:roleTerm>
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||||
<mods:namePart id="3D06F9753FC139507AF49630C8A5645E">Eberle, Jonas</mods:namePart>
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<mods:namePart id="F3B616083B8F94734A797B96E1486614">Beckett, Marc</mods:namePart>
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<mods:namePart id="FD2622AB5A121D1743E37F415631E5F5">Ozgul-Siemund, Alev</mods:namePart>
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<mods:namePart id="7B7A5F1A71AEA23EC1E41F44B399BFB6">Frings, Jujina</mods:namePart>
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</mods:role>
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<mods:namePart id="AB290ACBDA8586DA4FF05A469397ECA7">Fabrizi, Silvia</mods:namePart>
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</mods:name>
|
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</mods:role>
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<mods:namePart id="9C24F7CE04B5D47C8C084E891DFECF7D">Ahrens, Dirk</mods:namePart>
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<mods:title id="C80336B0B38D8F0BB57F81E87D35D12A">Zoological Journal of the Linnean Society</mods:title>
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|
||||
<mods:location id="1B2402CF44E317A5520B476EC850BF59">
|
||||
<mods:url id="17E7E047C8622C07FC2267F6D14203CC">https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="484C5F2B4C20E9E46852D1FF1641B691">journal article</mods:classification>
|
||||
<mods:identifier id="2D0F43805C99A0B389A8343A354E2F7E" type="DOI">10.1111/zoj.12489</mods:identifier>
|
||||
<mods:identifier id="134608395868E4A4A2A09A20E832FB24" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="11EC1DABD624F139386AAAAB9358FC6A" type="Zenodo-Dep">14807083</mods:identifier>
|
||||
<mods:identifier id="31079C3A314DE2535BC2FD3041D1E72D" type="ZooBank">3663E0A-2E17-4346-A388-6BEE53081814</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03CB6967FF9538453B87FD3794F4FD22" LSID="urn:lsid:plazi:treatment:03CB6967FF9538453B87FD3794F4FD22" httpUri="http://treatment.plazi.org/id/03CB6967FF9538453B87FD3794F4FD22" lastPageNumber="338" pageId="17" pageNumber="338">
|
||||
<subSubSection id="C3788BFAFF9538453B87FD37903AFCAE" pageId="17" pageNumber="338" type="nomenclature">
|
||||
<paragraph id="8BDDD871FF9538453B87FD379391FD02" blockId="17.[187,760,637,693]" pageId="17" pageNumber="338">
|
||||
<heading id="D0956F1DFF9538453B87FD379391FD02" centered="true" fontSize="9" level="2" pageId="17" pageNumber="338" reason="2">
|
||||
<taxonomicName id="4C62A3F2FF9538453B87FD3790F3FD05" ID-CoL="4K84K" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="transkeiensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9538453B87FD379372FD22" bold="true" box="[187,532,640,661]" italics="true" pageId="17" pageNumber="338">
|
||||
<smallCapsWord id="8D3B4EADFF9538453B87FD379032FD22" baselines="656,656" box="[187,340,640,661]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="17" pageNumber="338">PLEOPHYLLA</smallCapsWord>
|
||||
<smallCapsWord id="8D3B4EADFF9538453A62FD349372FD22" baselines="656" box="[350,532,643,660]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="transkeiensis" pageId="17" pageNumber="338">TRANSKEIENSIS</smallCapsWord>
|
||||
</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF9538453921FDC9931EFD22" baselines="656,656" box="[541,632,638,662]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="17" pageNumber="338">AHRENS</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF95384539B6FDC99397FD22" baselines="656,656" box="[650,753,638,662]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="17" pageNumber="338">BECKETT</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF9538453B87FD2B9072FD04" baselines="686,686" box="[187,276,668,692]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="17" pageNumber="338">EBERLE</smallCapsWord>
|
||||
&
|
||||
<smallCapsWord id="8D3B4EADFF9538453A02FD2B90F3FD05" baselines="686,687" box="[318,405,668,692]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabrizi" pageId="17" pageNumber="338">FABRIZI</smallCapsWord>
|
||||
</taxonomicName>
|
||||
<emphasis id="B9160463FF9538453AA3FD159362FD03" bold="true" box="[415,516,666,693]" pageId="17" pageNumber="338">
|
||||
<taxonomicNameLabel id="A225B918FF9538453AA3FD159362FD03" box="[415,516,666,693]" pageId="17" pageNumber="338" rank="species">
|
||||
<smallCapsWord id="8D3B4EADFF9538453AA3FD1590DDFD05" baselines="687" box="[415,443,674,691]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="17" pageNumber="338">SP</smallCapsWord>
|
||||
.
|
||||
<smallCapsWord id="8D3B4EADFF9538453AF0FD15909AFD05" baselines="687" box="[460,508,674,691]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="17" pageNumber="338">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF953845392BFD2B93FFFD02" box="[535,665,666,692]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="17" pageNumber="338">
|
||||
<smallCapsWord id="8D3B4EADFF953845392BFD2B932AFD05" baselines="686,687" box="[535,588,668,692]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="17" pageNumber="338">FIGS</smallCapsWord>
|
||||
3Q
|
||||
<emphasis id="B9160463FF9538453946FD2993EEFD02" box="[634,648,670,692]" italics="true" pageId="17" pageNumber="338">–</emphasis>
|
||||
T
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF9538453996FD2D93A8FD02" box="[682,718,666,692]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="17" pageNumber="338">5K</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9538453B9FFD54903AFCAE" blockId="17.[163,779,739,792]" pageId="17" pageNumber="338">
|
||||
<uri id="FFF3D473FF9538453B9FFD54903AFCAE" pageId="17" pageNumber="338">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF9538453A8AFD54903AFCAE" pageId="17" pageNumber="338">74086E27-186C-4C5A-80F7- 868A2FAAD37E</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF9538453B9AFC8E94F4FD22" pageId="17" pageNumber="338" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF9538453B9AFC8E9059FC7E" blockId="17.[166,781,824,968]" pageId="17" pageNumber="338">
|
||||
<emphasis id="B9160463FF9538453B9AFC8E9085FCFB" box="[166,483,824,846]" italics="true" pageId="17" pageNumber="338">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9538453ACEFC8F9364FC1F" collectingDate="1987-11-25" collectionCode="TMSA" collectorName="S. Afr." latitude="-31.27" location="Transkei Ntsubane forest" longLatPrecision="730" longitude="29.44" pageId="17" pageNumber="338" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF9538453ACEFC8F933DFCF8" box="[498,603,824,846]" pageId="17" pageNumber="338" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF9538453944FC8C93EEFCFB" box="[632,648,827,845]" count="1" pageId="17" pageNumber="338" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF9538453991FC8F926FFCF8" box="[685,777,824,846]" pageId="17" pageNumber="338">S. Afr.</collectorName>
|
||||
,
|
||||
<location id="8EBD8EAAFF9538453B9AFCE0908CFCDB" LSID="urn:lsid:plazi:treatment:03CB6967FF9538453B87FD3794F4FD22:8EBD8EAAFF9538453B9AFCE0908CFCDB" box="[166,490,855,877]" latitude="-31.27" longLatPrecision="730" longitude="29.44" name="Transkei Ntsubane forest" pageId="17" pageNumber="338">Transkei Ntsubane forest</location>
|
||||
<geoCoordinate id="EE56BEB6FF953845393FFCE09301FCDA" box="[515,615,855,876]" degrees="31.27" direction="south" orientation="latitude" pageId="17" pageNumber="338" precision="555" value="-31.27">31.27 S</geoCoordinate>
|
||||
-
|
||||
<geoCoordinate id="EE56BEB6FF95384539A3FCE09262FCDA" box="[671,772,855,876]" degrees="29.44" direction="east" orientation="longitude" pageId="17" pageNumber="338" precision="555" value="29.44">29.44 E</geoCoordinate>
|
||||
/
|
||||
<date id="FFDCFEB1FF9538453B9AFCC29046FC3D" box="[166,288,885,907]" pageId="17" pageNumber="338" value="1987-11-25">
|
||||
<collectingDate id="EF980759FF9538453B9AFCC29046FC3D" box="[166,288,885,907]" pageId="17" pageNumber="338" value="1987-11-25">25.11.1987</collectingDate>
|
||||
</date>
|
||||
E-Y: 2537 Fungi & for. litter leg Endrody-Younga” (
|
||||
<collectionCode id="ED7340B4FF9538453A91FC23909DFC1F" box="[429,507,916,937]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="17" pageNumber="338">TMSA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<typeStatus id="54D966D3FF9538453906FC2393D6FC1F" box="[570,688,916,937]" pageId="17" pageNumber="338" type="paratype">Paratypes</typeStatus>
|
||||
: see Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9538453B9AFC529357F90F" blockId="17.[166,781,996,1721]" pageId="17" pageNumber="338">
|
||||
<emphasis id="B9160463FF9538453B9AFC529049FC4C" box="[166,303,997,1018]" italics="true" pageId="17" pageNumber="338">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9538453ADBFC53932BFC4C" box="[487,589,996,1018]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="17" pageNumber="338" unit="mm" value="10.6">10.6 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9538453B9AFBB49199FBAE" box="[166,255,1027,1048]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="17" pageNumber="338" unit="mm" value="7.4">7.4 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9538453AD3FBB4932FFBAE" box="[495,585,1027,1049]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="17" pageNumber="338" unit="mm" value="5.7">5.7 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.6. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.53. Metatibia, ratio of width/length: 1/4.38. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF9538453924F93393EAF92C" box="[536,652,1668,1690]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="17" pageNumber="338">
|
||||
Fig. 3Q
|
||||
<emphasis id="B9160463FF9538453952F930931CF92F" box="[622,634,1671,1689]" italics="true" pageId="17" pageNumber="338">–</emphasis>
|
||||
S
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF9538453B9AF914919BF90E" box="[166,253,1699,1721]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="17" pageNumber="338">Fig. 3T</figureCitation>
|
||||
. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF9538453AEFF914934BF90F" box="[467,557,1699,1721]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="17" pageNumber="338">Fig. 5K</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9538453B9AF96290D2F8D3" blockId="17.[166,782,1749,1893]" pageId="17" pageNumber="338">
|
||||
<emphasis id="B9160463FF9538453B9AF9629333F95C" box="[166,597,1749,1770]" italics="true" pageId="17" pageNumber="338">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF9538453A10F9629333F95C" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[300,597,1749,1770]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="transkeiensis" status="sp. nov.">Pleophylla transkeiensis</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF953845394DF96293B8F95C" box="[625,734,1749,1770]" pageId="17" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is similar to
|
||||
<taxonomicName id="4C62A3F2FF9538453A0AF94390ABF8BE" box="[310,461,1779,1801]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="ferruginea">
|
||||
<emphasis id="B9160463FF9538453A0AF94390ABF8BE" box="[310,461,1779,1801]" italics="true" pageId="17" pageNumber="338">P. ferruginea</emphasis>
|
||||
</taxonomicName>
|
||||
in external morphology.
|
||||
<taxonomicName id="4C62A3F2FF9538453B9AF8A590A2F891" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[166,452,1810,1831]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="transkeiensis" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9538453B9AF8A590A2F891" box="[166,452,1810,1831]" italics="true" pageId="17" pageNumber="338">Pleophylla transkeiensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9538453AE9F8A59351F891" box="[469,567,1810,1831]" pageId="17" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from the latter by the wider labroclypeus and the distinctly asymmetric parameres.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9538453801FF70952DFE67" blockId="17.[829,1444,199,465]" pageId="17" pageNumber="338">
|
||||
<emphasis id="B9160463FF9538453801FF7092D7FF6A" box="[829,945,199,220]" italics="true" pageId="17" pageNumber="338">Variation:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9538453F60FF709597FF6A" box="[1116,1265,199,221]" metricMagnitude="-2" metricUnit="m" metricValue="1.015" metricValueMax="1.06" metricValueMin="0.97" pageId="17" pageNumber="338" unit="mm" value="10.149999999999999" valueMax="10.6" valueMin="9.7">
|
||||
9.7
|
||||
<emphasis id="B9160463FF9538453F42FF7D95ECFF6A" box="[1150,1162,202,220]" italics="true" pageId="17" pageNumber="338">–</emphasis>
|
||||
10.6 mm
|
||||
</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9538453801FF5192A3FF4D" box="[829,965,230,251]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" metricValueMax="7.4" metricValueMin="7.2" pageId="17" pageNumber="338" unit="mm" value="7.3" valueMax="7.4" valueMin="7.2">
|
||||
7.2
|
||||
<emphasis id="B9160463FF9538453863FF5F920DFF4C" box="[863,875,232,250]" italics="true" pageId="17" pageNumber="338">–</emphasis>
|
||||
7.4 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9538453F8DFF52945FFF4D" box="[1201,1337,229,251]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="5.7" metricValueMin="5.1" pageId="17" pageNumber="338" unit="mm" value="5.4" valueMax="5.7" valueMin="5.1">
|
||||
5.1
|
||||
<emphasis id="B9160463FF9538453FEFFF5F95B9FF4C" box="[1235,1247,232,250]" italics="true" pageId="17" pageNumber="338">–</emphasis>
|
||||
5.7 mm
|
||||
</quantity>
|
||||
. Density of dark spot may vary, sometimes completely without spots. Female: antennal club of female straight, composed of five antennomeres, as long as the remaining antennomeres combined; sixth antennomere short, a quarter as long as club length. Eyes smaller than in male: ratio of eye diameter/ interocular width: 0.51.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9538453801FE4692ECFDF2" blockId="17.[829,1444,497,580]" pageId="17" pageNumber="338">
|
||||
<emphasis id="B9160463FF9538453801FE4692DDFDB0" box="[829,955,497,518]" italics="true" pageId="17" pageNumber="338">Etymology:</emphasis>
|
||||
The new species is named
|
||||
<taxonomicName id="4C62A3F2FF9538453E39FE4694F9FDB0" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1285,1439,497,518]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="transkeiensis">
|
||||
‘
|
||||
<emphasis id="B9160463FF9538453E36FE4694F9FDB0" box="[1290,1439,497,518]" italics="true" pageId="17" pageNumber="338">transkeiensis</emphasis>
|
||||
</taxonomicName>
|
||||
’ after its occurrence in the Transkei Region of
|
||||
<collectingCountry id="F37598E1FF9538453E5CFDB892E0FDF2" name="South Africa" pageId="17" pageNumber="338">South Africa</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9538453801FDD794F4FD22" blockId="17.[829,1443,608,660]" pageId="17" pageNumber="338">
|
||||
<emphasis id="B9160463FF9538453801FDD792CFFDC3" box="[829,937,608,629]" italics="true" pageId="17" pageNumber="338">Remarks:</emphasis>
|
||||
In a previous publication we referred to this species as ‘SpM-16’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF9538453FD1FDC994E0FD22" author="Ahrens D" box="[1261,1414,638,660]" pageId="17" pageNumber="338" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
232
data/03/CB/69/03CB6967FF9538463871FD6591B0FC26.xml
Normal file
232
data/03/CB/69/03CB6967FF9538463871FD6591B0FC26.xml
Normal file
|
|
@ -0,0 +1,232 @@
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<mods:namePart id="5D245B67D2D1BC4B2CEBE56405D55EDB">Beckett, Marc</mods:namePart>
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<mods:namePart id="9E8E5912AACF5925165065C288A8A15D">Fabrizi, Silvia</mods:namePart>
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<mods:date id="95266F212AB5E56DE67C73429C1A270A">2017</mods:date>
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|
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<taxonomicName id="4C62A3F2FF9538453871FD65929AFCB3" ID-CoL="4K84D" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="ruthae" status="sp. nov.">
|
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,
|
||||
<smallCapsWord id="8D3B4EADFF9538453FFFFD67944CFD51" baselines="738,739" box="[1219,1322,720,744]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="17" pageNumber="338">BECKETT</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF9538453E07FD6794F2FD50" baselines="738,738" box="[1339,1428,720,744]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="17" pageNumber="338">EBERLE</smallCapsWord>
|
||||
&
|
||||
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|
||||
</taxonomicName>
|
||||
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|
||||
<taxonomicNameLabel id="A225B918FF9538453F39FD43950DFCBE" box="[1029,1131,749,776]" pageId="17" pageNumber="338" rank="species">
|
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|
||||
.
|
||||
<smallCapsWord id="8D3B4EADFF9538453F0FFD439505FCB3" baselines="769" box="[1075,1123,756,773]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="17" pageNumber="338">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF9538453F42FD589467FCB1" box="[1150,1281,749,775]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="17" pageNumber="338">
|
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<smallCapsWord id="8D3B4EADFF9538453F42FD5895D5FCB3" baselines="769,769" box="[1150,1203,751,775]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="17" pageNumber="338">FIGS</smallCapsWord>
|
||||
3U
|
||||
<emphasis id="B9160463FF9538453FDDFD479589FCB0" box="[1249,1263,752,774]" italics="true" pageId="17" pageNumber="338">–</emphasis>
|
||||
X
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF9538453E2EFD5A9455FCB1" box="[1298,1331,749,775]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="17" pageNumber="338">5L</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
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</paragraph>
|
||||
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|
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<uri id="FFF3D473FF9538453807FC929556FCEF" pageId="17" pageNumber="338">
|
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urn:lsid:zoobank.org:act:
|
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|
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|
||||
<subSubSection id="C3788BFAFF9538463801FCCF91B0FC26" lastPageId="18" lastPageNumber="339" pageId="17" pageNumber="338" type="materials_examined">
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|
||||
<emphasis id="B9160463FF9538453801FCCF9538FC3A" box="[829,1118,887,909]" italics="true" pageId="17" pageNumber="338">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9538453F51FCC094C6FC7C" collectingDate="1978-11" collectionCode="SANC" collectorName="G. L. Prinsloo & L. van Luik" country="South Africa" latitude="-23.0" location="Entabeni Forest" longLatPrecision="754" longitude="30.16" pageId="17" pageNumber="338" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF9538453F51FCC095B0FC3B" box="[1133,1238,887,909]" pageId="17" pageNumber="338" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF9538453FD9FCCD9593FC3A" box="[1253,1269,890,908]" count="1" pageId="17" pageNumber="338" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectingCountry id="F37598E1FF9538453E36FCC094C6FC3B" box="[1290,1440,887,909]" name="South Africa" pageId="17" pageNumber="338">South Africa</collectingCountry>
|
||||
, TVL
|
||||
<location id="8EBD8EAAFF9538453843FC219526FC1A" LSID="urn:lsid:plazi:treatment:03CB6967FF9538463871FD6591B0FC26:8EBD8EAAFF9538453843FC219526FC1A" box="[895,1088,918,940]" country="South Africa" latitude="-23.0" longLatPrecision="754" longitude="30.16" name="Entabeni Forest" pageId="17" pageNumber="338">Entabeni Forest</location>
|
||||
Res.
|
||||
<geoCoordinate id="EE56BEB6FF9538453FB3FC21958DFC1D" box="[1167,1259,918,939]" degrees="23.00" direction="south" orientation="latitude" pageId="17" pageNumber="338" precision="555" value="-23.0">23.00 S</geoCoordinate>
|
||||
<geoCoordinate id="EE56BEB6FF9538453FC6FC21943AFC1A" box="[1274,1372,918,940]" degrees="30.16" direction="east" orientation="longitude" pageId="17" pageNumber="338" precision="555" value="30.16">30.16 E</geoCoordinate>
|
||||
.
|
||||
<date id="FFDCFEB1FF9538453E53FC209211FC7C" pageId="17" pageNumber="338" value="1978-11">
|
||||
<collectingDate id="EF980759FF9538453E53FC209211FC7C" pageId="17" pageNumber="338" value="1978-11">Nov. 1978</collectingDate>
|
||||
</date>
|
||||
.
|
||||
<collectorName id="2697BDA7FF95384538AEFC02955CFC7D" box="[914,1082,949,971]" pageId="17" pageNumber="338">G.L. Prinsloo</collectorName>
|
||||
&
|
||||
<collectorName id="2697BDA7FF9538453F41FC029443FC7D" box="[1149,1317,949,971]" pageId="17" pageNumber="338">L. van Luik</collectorName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF9538453E71FC0294FFFC7C" box="[1357,1433,949,970]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="17" pageNumber="338" type="Museum">SANC</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<typeStatus id="54D966D3FF9538453801FC6392D5FC5F" box="[829,947,980,1001]" pageId="17" pageNumber="338" type="paratype">Paratypes</typeStatus>
|
||||
: see Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9538453801FBB0944BF96D" blockId="17.[829,1444,1031,1756]" pageId="17" pageNumber="338">
|
||||
<emphasis id="B9160463FF9538453801FBB092A0FBAA" box="[829,966,1031,1052]" italics="true" pageId="17" pageNumber="338">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9538453F42FBB09582FBAA" box="[1150,1252,1031,1052]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="17" pageNumber="338" unit="mm" value="10.7">10.7 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9538453801FB9192F0FB8D" box="[829,918,1062,1083]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="17" pageNumber="338" unit="mm" value="7.7">7.7 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9538453FBAFB929586FB8D" box="[1158,1248,1061,1083]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="17" pageNumber="338" unit="mm" value="5.7">5.7 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially weakly sinuate; margins of labroclypeus strongly reflexed; smooth area in front of eyes three times as wide as long; ratio of eye diameter/interocular width: 0.7. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra distinctly elevated but convex; white, fine pilosity condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/ metacoxa: 1/1.44. Metatibia, ratio of width/length: 1/ 3.84. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF9538453E1EF91094C7F90B" box="[1314,1441,1703,1725]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="17" pageNumber="338">
|
||||
Fig. 3U
|
||||
<emphasis id="B9160463FF9538453E45F91D94E3F90A" box="[1401,1413,1706,1724]" italics="true" pageId="17" pageNumber="338">–</emphasis>
|
||||
W
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF9538453899F971929BF96D" box="[933,1021,1734,1756]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="17" pageNumber="338">Fig. 3X</figureCitation>
|
||||
. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF9538453FEEF971944FF96D" box="[1234,1321,1733,1756]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="17" pageNumber="338">
|
||||
Fig.
|
||||
<specimenCount id="9D6413F8FF9538453E3BF972944FF96D" box="[1287,1321,1733,1755]" count="5" pageId="17" pageNumber="338" type="generic">5L</specimenCount>
|
||||
</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9538463801F9439056FEC3" blockId="17.[829,1444,1780,1893]" lastBlockId="18.[144,759,199,373]" lastPageId="18" lastPageNumber="339" pageId="17" pageNumber="338">
|
||||
<emphasis id="B9160463FF9538453801F94395C6F8BF" box="[829,1184,1780,1801]" italics="true" pageId="17" pageNumber="338">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF95384538FFF94395C6F8BF" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[963,1184,1780,1801]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="ruthae" status="sp. nov.">Pleophylla ruthae</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF9538453F85F9439442F8BF" box="[1209,1316,1780,1801]" pageId="17" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is quite similar to
|
||||
<taxonomicName id="4C62A3F2FF9538453881F8A4951AF891" box="[957,1148,1810,1832]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="fasciatipennis">
|
||||
<emphasis id="B9160463FF9538453881F8A4951AF891" box="[957,1148,1810,1832]" italics="true" pageId="17" pageNumber="338">P. fasciatipennis</emphasis>
|
||||
</taxonomicName>
|
||||
in external morphology.
|
||||
<taxonomicName id="4C62A3F2FF9538453801F8869574F8F0" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[829,1042,1841,1862]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="ruthae" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9538453801F8869574F8F0" box="[829,1042,1841,1862]" italics="true" pageId="17" pageNumber="338">Pleophylla ruthae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9538453F1FF88695E0F8F0" box="[1059,1158,1841,1862]" pageId="17" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs in the sutural interval being convex in the apical declivity of the elytra and not delimitated by a sharp carina.
|
||||
<taxonomicName id="4C62A3F2FF9638463BACFF529011FF4C" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[144,375,229,250]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="339" phylum="Arthropoda" rank="species" species="ruthae" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9638463BACFF529011FF4C" box="[144,375,229,250]" italics="true" pageId="18" pageNumber="339">Pleophylla ruthae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9638463AA7FF519376FF4D" box="[411,528,230,251]" pageId="18" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is similar to
|
||||
<taxonomicName id="4C62A3F2FF9638463BACFEB39023FEAF" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[144,325,260,281]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="339" phylum="Arthropoda" rank="species" species="transkeiensis">
|
||||
<emphasis id="B9160463FF9638463BACFEB39023FEAF" box="[144,325,260,281]" italics="true" pageId="18" pageNumber="339">P. transkeiensis</emphasis>
|
||||
</taxonomicName>
|
||||
in the shape of the parameres, but the left paramere is basally less produced medially and the right paramere lacks a blunt median tooth.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9638463BACFE3B90F6FD20" blockId="18.[144,759,396,662]" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF9638463BACFE3B9065FE17" box="[144,259,396,417]" italics="true" pageId="18" pageNumber="339">Variation:</emphasis>
|
||||
Body length: 9.0
|
||||
<emphasis id="B9160463FF9638463AEDFE3990BBFE16" box="[465,477,398,416]" italics="true" pageId="18" pageNumber="339">–</emphasis>
|
||||
<quantity id="4C9A7594FF9638463AE1FE3B9322FE17" box="[477,580,396,417]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="18" pageNumber="339" unit="mm" value="10.7">10.7 mm</quantity>
|
||||
, elytral length: 6.6
|
||||
<emphasis id="B9160463FF9638463B8EFE1A91D8FE09" box="[178,190,429,447]" italics="true" pageId="18" pageNumber="339">–</emphasis>
|
||||
8.0 mm, maximum width:
|
||||
<quantity id="4C9A7594FF963846393FFE1D93EAFE76" box="[515,652,426,448]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.7" metricValueMin="4.4" pageId="18" pageNumber="339" unit="mm" value="5.05" valueMax="5.7" valueMin="4.4">
|
||||
4.4
|
||||
<emphasis id="B9160463FF9638463919FE1A9357FE09" box="[549,561,429,447]" italics="true" pageId="18" pageNumber="339">–</emphasis>
|
||||
5.7 mm
|
||||
</quantity>
|
||||
. Density of dark spots may vary, never completely without spots. Female: antennal club of female straight, composed of five antennomeres, as long as the remaining antennomeres combined; sixth antennomere shorter than a quarter of club length. Eyes smaller than in male: ratio of eye diameter/ interocular width: 0.5.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9638463BACFD07933DFC94" blockId="18.[144,758,688,802]" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF9638463BACFD07906BFD73" box="[144,269,688,709]" italics="true" pageId="18" pageNumber="339">Etymology:</emphasis>
|
||||
This new species is dedicated to Ruth M uller € (
|
||||
<collectionCode id="ED7340B4FF9638463BD1FD78905CFD52" box="[237,314,719,740]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="18" pageNumber="339">TMSA</collectionCode>
|
||||
), collector of many
|
||||
<taxonomicName id="4C62A3F2FF9638463922FD799394FD52" box="[542,754,718,740]" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF9638463922FD7993F0FD55" box="[542,662,718,739]" italics="true" pageId="18" pageNumber="339">Pleophylla</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
, in gratitude for her hospitality and long-lasting support of our research in
|
||||
<collectingCountry id="F37598E1FF9638463AFFFCBB9331FC94" box="[451,599,780,802]" name="South Africa" pageId="18" pageNumber="339">South Africa</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9638463BACFC8A91B0FC26" blockId="18.[144,758,829,912]" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF9638463BACFC8A919DFCE4" box="[144,251,829,850]" italics="true" pageId="18" pageNumber="339">Remarks:</emphasis>
|
||||
In a previous publication we referred to this species as ‘SpM-23’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF96384639A1FCEB91ACFC26" author="Ahrens D" pageId="18" pageNumber="339" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
235
data/03/CB/69/03CB6967FF9638463B84FC729570FBBF.xml
Normal file
235
data/03/CB/69/03CB6967FF9638463B84FC729570FBBF.xml
Normal file
|
|
@ -0,0 +1,235 @@
|
|||
<document id="84D1A2AAEB2901B6B6F5A2AAA083C6DA" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia & Ahrens, Dirk" docDate="2017" docId="03CB6967FF9638463B84FC729570FBBF" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla warnockae AHRENS, BECKETT, EBERLE & FABRIZI 2017, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="339" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="339" updateTime="1738712560284" updateUser="ExternalLinkService">
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<mods:mods id="CCE265E736C496612F42E8B3192B0123" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5F05AD0E2761AB6B75AF92DAA43F1A71">
|
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<mods:title id="EA7756386DC2EEC48CC898B26593AE02">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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</mods:titleInfo>
|
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<mods:role id="7896489351B5EAD96FCDB5400CB8365D">
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<mods:roleTerm id="72AFB5EF12363FD8422F9AB4CCEDB1C7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1651FBFF5BAD0CE869AF61DF9A550449">Eberle, Jonas</mods:namePart>
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</mods:role>
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<mods:namePart id="50CFD419A0F662B3F67477B0F6C58664">Beckett, Marc</mods:namePart>
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<mods:roleTerm id="2D0BE633A2B3230EAA6A9A3B25B52B23">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="513E84E611F96B31BAA48B413A33B10A">Ozgul-Siemund, Alev</mods:namePart>
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</mods:name>
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<mods:namePart id="945B6AB06A8BEE494C9DD4ABDBF69B26">Frings, Jujina</mods:namePart>
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</mods:name>
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<mods:roleTerm id="A696F15681CB5691CF89EB6F5B5B27C2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="10FE400E0BB0E4EC66D89F7DAAC4AB27">Fabrizi, Silvia</mods:namePart>
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</mods:name>
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</mods:role>
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<mods:namePart id="89C59F491E565E4B6A32404921125D53">Ahrens, Dirk</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="205BC60336DC2599596D196B8205C923">text</mods:typeOfResource>
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||||
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||||
<mods:titleInfo id="2D8BAB0191455C0D527058B13F90FDEF">
|
||||
<mods:title id="FBBCDCB79B0FE606CD36F31DEE689B15">Zoological Journal of the Linnean Society</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="8F96DDCA0C1A68F12E6D944771884180">
|
||||
<mods:date id="E18E2BA04F08ACFE64FB5C7D87279D5D">2017</mods:date>
|
||||
<mods:detail id="832AAC2E08095FC4172A7BE99E474AA7" type="series">
|
||||
<mods:title id="AA09BA25B6CDEB5DF868230935BF2355">Zool. J. Linn. Soc.</mods:title>
|
||||
</mods:detail>
|
||||
<mods:detail id="C50288A728CFE7D31B29083454EBBF04" type="volume">
|
||||
<mods:number id="735009416EF6F7556B827510C6A749BE">180</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="08F170E5AEC7B9E3C640A4CF787637AE" type="issue">
|
||||
<mods:number id="46967331349AA9660F8D9AA000DB83C6">2</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="8EE84D3EA1971B60435F7DBAD94D17B6" unit="page">
|
||||
<mods:start id="ADF3C3B6ED6959AEB94EF6C84961B3C6">321</mods:start>
|
||||
<mods:end id="3AEA4A0038EDC00ABC8700C659FA2D58">353</mods:end>
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</mods:extent>
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<mods:location id="B7B8A2B51C50103C27FA1D7C689F2339">
|
||||
<mods:url id="D4F46BA93054BB31EDF9E42B1C5256FC">https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="AC3232470FBAB16951E46F228E7FB302">journal article</mods:classification>
|
||||
<mods:identifier id="55DADF02F312696A219C434473E2A4FB" type="DOI">10.1111/zoj.12489</mods:identifier>
|
||||
<mods:identifier id="FADE812056592F316BD6EABDF271DCEE" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="6619C61DCD2C7BCE078EA26E9AB9B124" type="Zenodo-Dep">14807083</mods:identifier>
|
||||
<mods:identifier id="2418B265478E0897C02427C3AC355F7C" type="ZooBank">3663E0A-2E17-4346-A388-6BEE53081814</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03CB6967FF9638463B84FC729570FBBF" LSID="urn:lsid:plazi:treatment:03CB6967FF9638463B84FC729570FBBF" httpUri="http://treatment.plazi.org/id/03CB6967FF9638463B84FC729570FBBF" lastPageNumber="339" pageId="18" pageNumber="339">
|
||||
<subSubSection id="C3788BFAFF9638463B84FC72905CFBE7" pageId="18" pageNumber="339" type="nomenclature">
|
||||
<paragraph id="8BDDD871FF9638463B84FC72938CFC4F" blockId="18.[156,746,962,1018]" pageId="18" pageNumber="339">
|
||||
<heading id="D0956F1DFF9638463B84FC72938CFC4F" centered="true" fontSize="9" level="2" pageId="18" pageNumber="339" reason="2">
|
||||
<taxonomicName id="4C62A3F2FF9638463B84FC729010FC4E" ID-CoL="4K84N" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="339" phylum="Arthropoda" rank="species" species="warnockae" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9638463B84FC72908FFC6F" bold="true" box="[184,489,965,986]" italics="true" pageId="18" pageNumber="339">
|
||||
<smallCapsWord id="8D3B4EADFF9638463B84FC729037FC6F" baselines="981,981" box="[184,337,965,986]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="18" pageNumber="339">PLEOPHYLLA</smallCapsWord>
|
||||
<smallCapsWord id="8D3B4EADFF9638463A67FC7F908FFC6F" baselines="981" box="[347,489,968,985]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="warnockae" pageId="18" pageNumber="339">WARNOCKAE</smallCapsWord>
|
||||
</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF9638463ACEFC749328FC6F" baselines="981,981" box="[498,590,963,987]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="18" pageNumber="339">AHRENS</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF9638463963FC7493A1FC6F" baselines="981,981" box="[607,711,963,987]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="18" pageNumber="339">BECKETT</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF9638463BA0FC569192FC41" baselines="1011,1011" box="[156,244,993,1017]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="18" pageNumber="339">EBERLE</smallCapsWord>
|
||||
&
|
||||
<smallCapsWord id="8D3B4EADFF9638463A22FC569010FC4E" baselines="1011,1012" box="[286,374,993,1017]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabrizi" pageId="18" pageNumber="339">FABRIZI</smallCapsWord>
|
||||
</taxonomicName>
|
||||
<emphasis id="B9160463FF9638463A43FC509083FC4C" bold="true" box="[383,485,991,1018]" pageId="18" pageNumber="339">
|
||||
<taxonomicNameLabel id="A225B918FF9638463A43FC509083FC4C" box="[383,485,991,1018]" pageId="18" pageNumber="339" rank="species">
|
||||
<smallCapsWord id="8D3B4EADFF9638463A43FC5090FDFC4E" baselines="1012" box="[383,411,999,1016]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="18" pageNumber="339">SP</smallCapsWord>
|
||||
.
|
||||
<smallCapsWord id="8D3B4EADFF9638463A91FC5090BBFC4E" baselines="1012" box="[429,477,999,1016]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="18" pageNumber="339">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF9638463AC4FC5693E1FC41" box="[504,647,991,1017]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="18" pageNumber="339">
|
||||
<smallCapsWord id="8D3B4EADFF9638463AC4FC56934BFC4E" baselines="1011,1012" box="[504,557,993,1017]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="18" pageNumber="339">FIGS</smallCapsWord>
|
||||
3Y
|
||||
<emphasis id="B9160463FF9638463964FC549300FC4F" box="[600,614,995,1017]" italics="true" pageId="18" pageNumber="339">–</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF963846395AFC5693E1FC41" baselines="1011,1011" box="[614,647,993,1017]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ab" pageId="18" pageNumber="339">AB</smallCapsWord>
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF96384639A4FC6893A6FC4F" box="[664,704,991,1017]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="18" pageNumber="339">5M</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9638463BADFBAB905CFBE7" blockId="18.[145,761,1052,1105]" pageId="18" pageNumber="339">
|
||||
<uri id="FFF3D473FF9638463BADFBAB905CFBE7" pageId="18" pageNumber="339">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF9638463A9AFBAB905CFBE7" pageId="18" pageNumber="339">12F9482F-5443-442E-970F- C0113C594151</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF9638463BACFBDA9570FBBF" pageId="18" pageNumber="339" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF9638463BACFBDA93FDFB08" blockId="18.[144,758,1132,1215]" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF9638463BACFBDA90B4FB37" box="[144,466,1132,1154]" italics="true" pageId="18" pageNumber="339">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9638463ADDFBDB902FFB08" collectingDate="1980-11-06" collectingDateMax="1980-11-09" collectingDateMin="1980-11-06" collectionCode="TMSA" collectorName="D. H. Jacobs" latitude="-24.183332" location="Balloon Forest Tzaneen" longLatPrecision="21" longitude="30.333334" municipality="Balloon Forest" pageId="18" pageNumber="339" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF9638463ADDFBDB932CFB34" box="[481,586,1132,1154]" pageId="18" pageNumber="339" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF9638463956FBD8931CFB37" box="[618,634,1135,1153]" count="1" pageId="18" pageNumber="339" type="male">♂</specimenCount>
|
||||
“
|
||||
<location id="8EBD8EAAFF963846399CFBDB9021FB16" LSID="urn:lsid:plazi:treatment:03CB6967FF9638463B84FC729570FBBF:8EBD8EAAFF963846399CFBDB9021FB16" latitude="-24.183332" longLatPrecision="21" longitude="30.333334" municipality="Balloon Forest" name="Balloon Forest Tzaneen" pageId="18" pageNumber="339">
|
||||
<collectingMunicipality id="6BB9420BFF963846399CFBDB91BFFB16" pageId="18" pageNumber="339">Balloon Forest</collectingMunicipality>
|
||||
Tzaneen
|
||||
</location>
|
||||
<geoCoordinate id="EE56BEB6FF9638463A6AFB3C90CDFB16" box="[342,427,1159,1184]" degrees="24" direction="south" minutes="11" orientation="latitude" pageId="18" pageNumber="339" precision="15" seconds="0" value="-24.183332">
|
||||
24
|
||||
<emphasis id="B9160463FF9638463A4DFB3B901DFB16" box="[369,379,1164,1184]" italics="true" pageId="18" pageNumber="339">°</emphasis>
|
||||
11
|
||||
<emphasis id="B9160463FF9638463AAAFB3090FDFB20" bold="true" box="[406,411,1159,1174]" italics="true" pageId="18" pageNumber="339">
|
||||
<superScript id="7C177539FF9638463AAAFB3090FDFB20" attach="right" box="[406,411,1159,1174]" fontSize="6" pageId="18" pageNumber="339">0</superScript>
|
||||
</emphasis>
|
||||
S
|
||||
</geoCoordinate>
|
||||
<geoCoordinate id="EE56BEB6FF9638463A86FB3C9374FB16" box="[442,530,1159,1184]" degrees="30" direction="east" minutes="20" orientation="longitude" pageId="18" pageNumber="339" precision="15" seconds="0" value="30.333334">
|
||||
30
|
||||
<emphasis id="B9160463FF9638463AE9FB3B90B9FB16" box="[469,479,1164,1184]" italics="true" pageId="18" pageNumber="339">°</emphasis>
|
||||
20
|
||||
<emphasis id="B9160463FF9638463AC6FB309099FB20" bold="true" box="[506,511,1159,1174]" italics="true" pageId="18" pageNumber="339">
|
||||
<superScript id="7C177539FF9638463AC6FB309099FB20" attach="right" box="[506,511,1159,1174]" fontSize="6" pageId="18" pageNumber="339">0</superScript>
|
||||
</emphasis>
|
||||
E
|
||||
</geoCoordinate>
|
||||
<date id="FFDCFEB1FF963846391CFB3C93D4FB16" box="[544,690,1163,1185]" pageId="18" pageNumber="339" value="1980-11-06" valueMax="1980-11-09" valueMin="1980-11-06">
|
||||
<collectingDate id="EF980759FF963846391CFB3C93D4FB16" box="[544,690,1163,1185]" pageId="18" pageNumber="339" value="1980-11-06" valueMax="1980-11-09" valueMin="1980-11-06">6-9 XI 1980</collectingDate>
|
||||
</date>
|
||||
<collectorName id="2697BDA7FF96384639FDFB3C91BDFB09" pageId="18" pageNumber="339">D.H. Jacobs</collectorName>
|
||||
” (
|
||||
<collectionCode id="ED7340B4FF9638463BC8FB1E9024FB08" box="[244,322,1193,1214]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="18" pageNumber="339">TMSA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<typeStatus id="54D966D3FF9638463A69FB1D90ADFB09" box="[341,459,1194,1215]" pageId="18" pageNumber="339" type="paratype">Paratypes</typeStatus>
|
||||
: see Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9638463BACFB6192EEFEAC" blockId="18.[144,759,1237,1901]" lastBlockId="18.[807,1422,199,282]" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF9638463BACFB61907EFB5D" box="[144,280,1238,1259]" italics="true" pageId="18" pageNumber="339">Description:</emphasis>
|
||||
Body length: 10.0 mm, elytral length:
|
||||
<quantity id="4C9A7594FF9638463BACFB43918FFABF" box="[144,233,1268,1290]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="18" pageNumber="339" unit="mm" value="6.9">6.9 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9638463AE4FB439355FABF" box="[472,563,1268,1289]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="18" pageNumber="339" unit="mm" value="4.9">4.9 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus strongly reflexed; smooth area in front of eyes three times as wide as long; ratio of eye diameter/interocular width: 0.73. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra distinctly elevated but convex; white, fine pilosity condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.6. Metatibia, ratio of width/length: 1/3.62. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF963846381BFF52921DFF4C" box="[807,891,229,251]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="18" pageNumber="339">Fig. 3Y</figureCitation>
|
||||
<emphasis id="B9160463FF9638463847FF5F92E1FF4C" box="[891,903,232,250]" italics="true" pageId="18" pageNumber="339">–</emphasis>
|
||||
Aa. Habitus:
|
||||
<figureCitation id="1359C4F4FF9638463F06FF52951DFF4D" box="[1082,1147,229,251]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="18" pageNumber="339">Fig. 3</figureCitation>
|
||||
Ab. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF963846381BFEB392E2FEAC" box="[807,900,260,282]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="18" pageNumber="339">
|
||||
Fig.
|
||||
<specimenCount id="9D6413F8FF9638463860FEB392E2FEAC" box="[860,900,260,282]" count="5" pageId="18" pageNumber="339" type="generic">5M</specimenCount>
|
||||
</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF963846381BFE83958EFDB7" blockId="18.[807,1421,308,513]" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF963846381BFE8395C8FEFF" box="[807,1198,308,329]" italics="true" pageId="18" pageNumber="339">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF9638463890FE8395C8FEFF" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[940,1198,308,329]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="339" phylum="Arthropoda" rank="species" species="warnockae" status="sp. nov.">Pleophylla warnockae</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF9638463F83FE839446FEFF" box="[1215,1312,308,329]" pageId="18" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is quite similar to
|
||||
<taxonomicName id="4C62A3F2FF963846389EFEE49568FED1" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[930,1038,338,360]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="339" phylum="Arthropoda" rank="species" species="ruthae" status="sp. nov.">
|
||||
<emphasis id="B9160463FF963846389EFEE49568FED1" box="[930,1038,338,360]" italics="true" pageId="18" pageNumber="339">P. ruthae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9638463F24FEE49515FEDE" box="[1048,1139,339,360]" pageId="18" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
in external morphology but it differs in the slightly shorter metatibia (ratio length/maximum width: 3.6) and the strongly reflexed margins of the labroclypeus; the basomedial teeth are widely separated from each other, whereas in
|
||||
<taxonomicName id="4C62A3F2FF963846387AFE5B92D7FDB6" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[838,945,491,513]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="339" phylum="Arthropoda" rank="species" species="ruthae" status="sp. nov.">
|
||||
<emphasis id="B9160463FF963846387AFE5B92D7FDB6" box="[838,945,491,513]" italics="true" pageId="18" pageNumber="339">P. ruthae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9638463885FE5B9574FDB7" box="[953,1042,492,513]" pageId="18" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
they are adjacent.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF963846381BFDA89438FCBD" blockId="18.[806,1422,543,779]" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF963846381BFDA892FCFD82" box="[807,922,543,564]" italics="true" pageId="18" pageNumber="339">Variation:</emphasis>
|
||||
Body length: 8.1
|
||||
<emphasis id="B9160463FF9638463F54FD959512FD82" box="[1128,1140,546,564]" italics="true" pageId="18" pageNumber="339">–</emphasis>
|
||||
10.0 mm, elytral length:
|
||||
<quantity id="4C9A7594FF963846381BFD8A92C8FDE5" box="[807,942,573,595]" metricMagnitude="-3" metricUnit="m" metricValue="6.699999999999999" metricValueMax="7.1" metricValueMin="6.3" pageId="18" pageNumber="339" unit="mm" value="6.699999999999999" valueMax="7.1" valueMin="6.3">
|
||||
6.3
|
||||
<emphasis id="B9160463FF9638463875FDF79233FDE4" box="[841,853,576,594]" italics="true" pageId="18" pageNumber="339">–</emphasis>
|
||||
7.1 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9638463FB9FD8A946BFDE5" box="[1157,1293,573,595]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.8" metricValueMin="4.4" pageId="18" pageNumber="339" unit="mm" value="5.1" valueMax="5.8" valueMin="4.4">
|
||||
4.4
|
||||
<emphasis id="B9160463FF9638463F9BFDF795D5FDE4" box="[1191,1203,576,594]" italics="true" pageId="18" pageNumber="339">–</emphasis>
|
||||
5.8 mm
|
||||
</quantity>
|
||||
. Density of dark spots may vary, never completely without spots. Female: antennal club of female straight, composed of five antennomeres, as long as the remaining antennomeres combined; sixth antennomere shorter than a quarter of club length. Eyes smaller than in male: ratio of eye diameter/interocular width: 0.5.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF963846381AFC9D92C3FC2D" blockId="18.[806,1421,810,923]" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF963846381AFC9D92C2FC89" box="[806,932,810,831]" italics="true" pageId="18" pageNumber="339">Etymology:</emphasis>
|
||||
The new species is dedicated to Rachel Warnock; her Master’s thesis contributed to a better understanding of the integrative taxonomy of
|
||||
<taxonomicName id="4C62A3F2FF963846381AFC3192F8FC2D" authorityName="Erichson" authorityYear="1847" box="[806,926,902,923]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B9160463FF963846381AFC3192F8FC2D" box="[806,926,902,923]" italics="true" pageId="18" pageNumber="339">Pleophylla</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF963846381AFC019570FBBF" blockId="18.[806,1421,950,1033]" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF963846381AFC0192F4FC7D" box="[806,914,950,971]" italics="true" pageId="18" pageNumber="339">Remarks:</emphasis>
|
||||
In previous publications we referred to this species as ‘sp9’ (Ozg € ul-Siemund € &
|
||||
<bibRefCitation id="EFF3A580FF9638463FD2FC6394EFFC5C" author="Ahrens D" box="[1262,1417,980,1002]" pageId="18" pageNumber="339" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFF3A580FF963846381BFC44956CFBBF" author="Eberle J & Warnock RCM & Ahrens D" box="[807,1034,1011,1033]" pageId="18" pageNumber="339" refId="ref23428" refString="Eberle J, Warnock RCM, Ahrens D. 2016 b. Bayesian species delimitation in Pleophylla chafers (Coleoptera) - the importance of prior choice and morphology. BMC Evolutionary Biology 16: 94." type="journal volume" year="2016">
|
||||
Eberle
|
||||
<emphasis id="B9160463FF9638463845FC4392D5FBBE" box="[889,947,1011,1033]" italics="true" pageId="18" pageNumber="339">et al.</emphasis>
|
||||
, 2016b
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
219
data/03/CB/69/03CB6967FF963847386AFBF9939FFB20.xml
Normal file
219
data/03/CB/69/03CB6967FF963847386AFBF9939FFB20.xml
Normal file
|
|
@ -0,0 +1,219 @@
|
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|
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<taxonomicName id="4C62A3F2FF963846386AFBF9956EFB37" ID-CoL="4K83Z" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="339" phylum="Arthropoda" rank="species" species="harrisoni" status="sp. nov.">
|
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|
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,
|
||||
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|
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,
|
||||
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&
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</taxonomicName>
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|
||||
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|
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|
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.
|
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<smallCapsWord id="8D3B4EADFF9638463F03FBC79509FB37" baselines="1149" box="[1087,1135,1136,1153]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="18" pageNumber="339">NOV</smallCapsWord>
|
||||
.
|
||||
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|
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|
||||
(
|
||||
<figureCitation id="1359C4F4FF9638463FB5FBDC9440FB37" box="[1161,1318,1129,1155]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="18" pageNumber="339">
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|
||||
3
|
||||
<smallCapsWord id="8D3B4EADFF9638463FE4FBDC959EFB37" baselines="1149,1149" box="[1240,1272,1131,1155]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ac" pageId="18" pageNumber="339">AC</smallCapsWord>
|
||||
<emphasis id="B9160463FF9638463FC5FBDA9461FB35" box="[1273,1287,1133,1155]" italics="true" pageId="18" pageNumber="339">–</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF9638463E3BFBDC9440FB37" baselines="1149,1149" box="[1287,1318,1131,1155]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Af" pageId="18" pageNumber="339">AF</smallCapsWord>
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF9638463E0BFBDE943AFB35" box="[1335,1372,1129,1155]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="18" pageNumber="339">5N</figureCitation>
|
||||
, 6)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9638463815FB2A92B1FB67" blockId="18.[809,1425,1181,1233]" pageId="18" pageNumber="339">
|
||||
<uri id="FFF3D473FF9638463815FB2A92B1FB67" pageId="18" pageNumber="339">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF9638463F0AFB2A92B1FB67" pageId="18" pageNumber="339">FA467F76-B3E5-4C3C-9DA0- 8102E00BC156</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF963847381BFB52939FFB20" lastPageId="19" lastPageNumber="340" pageId="18" pageNumber="339" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF963846381BFB5295B3FAE3" blockId="18.[807,1422,1252,1366]" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF963846381BFB529537FB4F" box="[807,1105,1252,1274]" italics="true" pageId="18" pageNumber="339">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9638463F5CFB5392E5FAE3" collectingDate="1987-02-09" collectionCode="TMSA" collectorName="S. Afr" latitude="-25.29" longLatPrecision="748" longitude="30.55" pageId="18" pageNumber="339" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF9638463F5CFB5395A0FB4C" box="[1120,1222,1252,1274]" pageId="18" pageNumber="339" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="9D6413F8FF9638463FE9FB509583FB4F" box="[1237,1253,1255,1273]" count="1" pageId="18" pageNumber="339" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF9638463E3DFB52942BFB4C" box="[1281,1357,1252,1274]" pageId="18" pageNumber="339">S. Afr</collectorName>
|
||||
, Tvl. Nelspruit Nat. Res, rivulet val.
|
||||
<geoCoordinate id="EE56BEB6FF9638463F92FAB49461FAAE" box="[1198,1287,1283,1305]" degrees="25.29" direction="south" orientation="latitude" pageId="18" pageNumber="339" precision="555" value="-25.29">25.29 S</geoCoordinate>
|
||||
-
|
||||
<geoCoordinate id="EE56BEB6FF9638463E16FAB494E3FAAE" box="[1322,1413,1283,1305]" degrees="30.55" direction="east" orientation="longitude" pageId="18" pageNumber="339" precision="555" value="30.55">30.55 E</geoCoordinate>
|
||||
/
|
||||
<date id="FFDCFEB1FF963846381BFA9592E0FA81" box="[807,902,1314,1335]" pageId="18" pageNumber="339" value="1987-02-09">
|
||||
<collectingDate id="EF980759FF963846381BFA9592E0FA81" box="[807,902,1314,1335]" pageId="18" pageNumber="339" value="1987-02-09">9.2.1987</collectingDate>
|
||||
</date>
|
||||
E-Y: 2433 beating leg Endrody-Younga” (
|
||||
<collectionCode id="ED7340B4FF9638463812FAF7921AFAE3" box="[814,892,1344,1365]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="18" pageNumber="339">TMSA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<typeStatus id="54D966D3FF96384638B3FAF69563FAE0" box="[911,1029,1345,1366]" pageId="18" pageNumber="339" type="paratype">Paratypes</typeStatus>
|
||||
: see Appendix S1.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF963847381BFAD893C5FE22" blockId="18.[807,1422,1390,1901]" lastBlockId="19.[166,781,199,404]" lastPageId="19" lastPageNumber="340" pageId="18" pageNumber="339">
|
||||
<emphasis id="B9160463FF963846381BFAD892C9FA32" box="[807,943,1391,1412]" italics="true" pageId="18" pageNumber="339">Description:</emphasis>
|
||||
Body length: 9.0 mm, elytral length:
|
||||
<quantity id="4C9A7594FF963846381BFA3B92E6FA14" box="[807,896,1420,1442]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="18" pageNumber="339" unit="mm" value="5.7">5.7 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9638463F53FA3A95AFFA14" box="[1135,1225,1421,1443]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="18" pageNumber="339" unit="mm" value="4.6">4.6 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus strongly reflexed; smooth area in front of eyes three times as wide as long; ratio of eye diameter/interocular width: 0.74. Pronotum with dense and thick erect setae; unicoloured. Hypomeron not carinate. Basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse; Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/ metacoxa: 1/1.41. Metatibia, ratio of width/length: 1/ 3.87. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF9738473940FEE893DBFEC3" box="[636,701,351,373]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="19" pageNumber="340">Fig. 3</figureCitation>
|
||||
Ac
|
||||
<emphasis id="B9160463FF97384739E0FED5938EFEC2" box="[732,744,354,372]" italics="true" pageId="19" pageNumber="340">–</emphasis>
|
||||
Ae. Habitus:
|
||||
<figureCitation id="1359C4F4FF9738473A33FEC99029FE22" box="[271,335,382,404]" captionStart="Figure 3" captionStartId="7.[166,245,1590,1609]" captionTargetBox="[178,1431,206,1549]" captionTargetId="figure-186@7.[162,1441,183,1568]" captionTargetPageId="7" captionText="Figure 3. A–D, Pleophylla ferruginea Burmeister (RSA, Tygerkloof); E–H, Pleophylla murzini sp. nov. (holotype); I– L, Pleophylla navicularis Burmeister (RSA, Weza); M–P, Pleophylla silvatica sp. nov. (holotype); Q–T, Pleophylla transkeiensis sp. nov. (holotype); U–X, Pleophylla ruthae sp. nov. (holotype); Y–Ab, Pleophylla warnockae sp. nov. (holotype); Ac–Af, Pleophylla harrisoni sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807091" httpUri="https://zenodo.org/record/14807091/files/figure.png" pageId="19" pageNumber="340">Fig. 3</figureCitation>
|
||||
Af. Female genitalia:
|
||||
<figureCitation id="1359C4F4FF9738473979FEC993F8FE22" box="[581,670,382,404]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="19" pageNumber="340">
|
||||
Fig.
|
||||
<specimenCount id="9D6413F8FF9738473946FEC993F8FE22" box="[634,670,382,404]" count="5" pageId="19" pageNumber="340" type="generic">5N</specimenCount>
|
||||
</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9738473B9AFE1C90F7FD23" blockId="19.[166,781,426,662]" pageId="19" pageNumber="340">
|
||||
<emphasis id="B9160463FF9738473B9AFE1C9379FE09" box="[166,543,426,448]" italics="true" pageId="19" pageNumber="340">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF9738473A10FE1D9379FE09" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[300,543,426,447]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="harrisoni" status="sp. nov.">Pleophylla harrisoni</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF9738473913FE1C93F6FE76" box="[559,656,427,448]" pageId="19" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is rather similar to
|
||||
<taxonomicName id="4C62A3F2FF9738473A1AFE7E9083FE68" box="[294,485,457,478]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="fasciatipennis">
|
||||
<emphasis id="B9160463FF9738473A1AFE7E9083FE68" box="[294,485,457,478]" italics="true" pageId="19" pageNumber="340">P. fasciatipennis</emphasis>
|
||||
</taxonomicName>
|
||||
in external morphology, but it differs from the latter by the smooth area in front of the eyes being three times as wide as long. The parameres are quite similar to
|
||||
<taxonomicName id="4C62A3F2FF9738473975FD929385FD8F" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[585,739,548,570]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="warnockae" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9738473975FD929385FD8F" box="[585,739,548,570]" italics="true" pageId="19" pageNumber="340">P. warnockae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF97384739D1FD9291B0FDEE" pageId="19" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C62A3F2FF9738473A2DFDF3901AFDEE" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[273,380,579,601]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="ruthae" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9738473A2DFDF3901AFDEE" box="[273,380,579,601]" italics="true" pageId="19" pageNumber="340">P. ruthae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9738473AB9FDF490B9FDEE" box="[389,479,579,600]" pageId="19" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, but the dorsal lobe of the left paramere has only a single basal tooth in
|
||||
<taxonomicName id="4C62A3F2FF9738473B9AFD369057FD23" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[166,305,640,662]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="harrisoni" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9738473B9AFD369057FD23" box="[166,305,640,662]" italics="true" pageId="19" pageNumber="340">P. harrisoni</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9738473A05FD3790F7FD23" box="[313,401,640,661]" pageId="19" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9738473B9AFD1C90C1FC02" blockId="19.[166,781,682,949]" pageId="19" pageNumber="340">
|
||||
<emphasis id="B9160463FF9738473B9AFD1C907CFD76" box="[166,282,683,704]" italics="true" pageId="19" pageNumber="340">Variation:</emphasis>
|
||||
Body length: 6.9
|
||||
<emphasis id="B9160463FF9738473AD2FD1A909CFD09" box="[494,506,685,703]" italics="true" pageId="19" pageNumber="340">–</emphasis>
|
||||
9.0 mm, elytral length:
|
||||
<quantity id="4C9A7594FF9738473B9AFD7E9048FD68" box="[166,302,713,735]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" metricValueMax="5.7" metricValueMin="5.2" pageId="19" pageNumber="340" unit="mm" value="5.45" valueMax="5.7" valueMin="5.2">
|
||||
5.2
|
||||
<emphasis id="B9160463FF9738473BF4FD7B91B2FD68" box="[200,212,716,734]" italics="true" pageId="19" pageNumber="340">–</emphasis>
|
||||
5.7 mm
|
||||
</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9738473926FD7E93C4FD68" box="[538,674,713,735]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.6" metricValueMin="3.6" pageId="19" pageNumber="340" unit="mm" value="4.1" valueMax="4.6" valueMin="3.6">
|
||||
3.6
|
||||
<emphasis id="B9160463FF9738473900FD7B932EFD68" box="[572,584,716,734]" italics="true" pageId="19" pageNumber="340">–</emphasis>
|
||||
4.6 mm
|
||||
</quantity>
|
||||
. Density of dark spots may vary, never completely without spots. Female: antennal club of female straight, composed of five antennomeres, as long as the remaining antennomeres combined; sixth antennomere shorter than a quarter of club length. Eyes smaller than in male: ratio of eye diameter/ interocular width: 0.5.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9738473B9AFC7A90B8FB88" blockId="19.[166,781,973,1086]" pageId="19" pageNumber="340">
|
||||
<emphasis id="B9160463FF9738473B9AFC7A9042FC54" box="[166,292,973,994]" italics="true" pageId="19" pageNumber="340">Etymology:</emphasis>
|
||||
This new species is dedicated to James du Guesclin Harrison (Pretoria). James supports our research in
|
||||
<collectingCountry id="F37598E1FF9738473A0FFBBD90A3FB96" box="[307,453,1034,1056]" name="South Africa" pageId="19" pageNumber="340">South Africa</collectingCountry>
|
||||
and recollected this species for our molecular analyses.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9738473B9AFBD5939FFB20" blockId="19.[166,781,1122,1174]" pageId="19" pageNumber="340">
|
||||
<emphasis id="B9160463FF9738473B9AFBD59074FBC1" box="[166,274,1122,1143]" italics="true" pageId="19" pageNumber="340">Remarks:</emphasis>
|
||||
In a previous publication we referred to this species as ‘SpM-17’ (Ozgul-Siemund € € &
|
||||
<bibRefCitation id="EFF3A580FF973847396BFB37938BFB20" author="Ahrens D" box="[599,749,1152,1174]" pageId="19" pageNumber="340" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
188
data/03/CB/69/03CB6967FF9738403876FBBD934AFCB3.xml
Normal file
188
data/03/CB/69/03CB6967FF9738403876FBBD934AFCB3.xml
Normal file
|
|
@ -0,0 +1,188 @@
|
|||
<document id="1249D605150EDB3705494F4F82AAA5C4" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia & Ahrens, Dirk" docDate="2017" docId="03CB6967FF9738403876FBBD934AFCB3" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla charlyi AHRENS, BECKETT, EBERLE & FABRIZI 2017, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="341" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="340" updateTime="1738712560284" updateUser="ExternalLinkService">
|
||||
<mods:mods id="86B547D3C36418638786312A707CC3C6" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="30C504D92908049834CC97CD67ECB8B1">
|
||||
<mods:title id="BEBFF4A26CB4D891C82D89AF3CA7410F">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="64053AB61C9DD8B6BE5E9161D3252063" type="personal">
|
||||
<mods:role id="44DE74A9332C6B8133817CF32249BA25">
|
||||
<mods:roleTerm id="E0004AD798C00A54CAC03F942558FE73">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="21BBC279C954C139E03E6C2C9ACCF3E8">Eberle, Jonas</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="9FFE31E56A604E350D2115F7A5F60889" type="personal">
|
||||
<mods:role id="7992322F2A5F952874CCC25C8A75FBC8">
|
||||
<mods:roleTerm id="763A6B8138B2096A69877D90C1EEA3F2">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="50ABA3CC01E5E292B62411B29D87099F">Beckett, Marc</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="867509BAA18B6E24416BD6747E1C53F7" type="personal">
|
||||
<mods:role id="A5514FFDA8259408CF5C8D8E6F31C8CD">
|
||||
<mods:roleTerm id="D082CF6168D7B00B5B8446779A05B401">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="242B9044FFE336D7E6F9522F975E6197">Ozgul-Siemund, Alev</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="AFC4CE4AA81D39129BCBE6111FEC7677" type="personal">
|
||||
<mods:role id="FC1E4387C2DABB8FD6E42AFFFC50DBCC">
|
||||
<mods:roleTerm id="749C4A642B1607DB00B21C565F372168">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="968DAFFD09BC8C791C1CC883050DB3B0">Frings, Jujina</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="7CCFE2C1665FA0979A8D3B218E2D81F2" type="personal">
|
||||
<mods:role id="392303881CCB8320531E96F889A8BA25">
|
||||
<mods:roleTerm id="6FE4417AB53839687230AC558FFE0F74">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="D600D368DA0785D5D5F7A6829E59991E">Fabrizi, Silvia</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="846AD37CCB5A0B4D93C441519C30B77F" type="personal">
|
||||
<mods:role id="8F613C5E3DEAD63F5117561596689554">
|
||||
<mods:roleTerm id="F33F493787E4F093B2C97577C44BCB8C">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="B3BC842ADBC159E563CCB1F1DBEF663B">Ahrens, Dirk</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="FDC2E1319655B9BF1C56B4EA94EC0978">text</mods:typeOfResource>
|
||||
<mods:relatedItem id="37515D71528E65C6A17AB6C979713D43" type="host">
|
||||
<mods:titleInfo id="2C00BD7A8A9EE20AEC80F85E4F20F7C1">
|
||||
<mods:title id="F02A2F030B88D5EF887FEA8D723D5557">Zoological Journal of the Linnean Society</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="A7CCC50D654B8C59BC04471777160E89">
|
||||
<mods:date id="1996FF8ECF287A1843D6FBDB46B2A39E">2017</mods:date>
|
||||
<mods:detail id="D9D4D209ACB9F1B93B18F00AB52B665F" type="series">
|
||||
<mods:title id="E411E6AAFDD1203E4C60590D414B828B">Zool. J. Linn. Soc.</mods:title>
|
||||
</mods:detail>
|
||||
<mods:detail id="ADA72DD98867AA606E788B21463EE5C6" type="volume">
|
||||
<mods:number id="16A424517902C6503EEDEFF069847021">180</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="21AB9C378B577A3683A1139F78642684" type="issue">
|
||||
<mods:number id="06BC3FB58A26BBCE7BF2871F3995388C">2</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="080F76191A8089DD7147ACB4F0E68BB8" unit="page">
|
||||
<mods:start id="6B6BCE8153F3BC7865F9620373FE25ED">321</mods:start>
|
||||
<mods:end id="1DA595839C5E353CD8FCE124B89D41AB">353</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location id="D129C5769B5E4CFE53BB2071B5E70E7B">
|
||||
<mods:url id="DBC01333DE8DD894DF046F224B9B2072">https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="B2FF7374F520E2B48AFB0AF1A35559D8">journal article</mods:classification>
|
||||
<mods:identifier id="A9F7173E5E794514ED6CF1452928AA44" type="DOI">10.1111/zoj.12489</mods:identifier>
|
||||
<mods:identifier id="7E2D87E1A4D69F46D9083CACCA3456B8" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="54E53D896227C5F39745F5A5A1159EA3" type="Zenodo-Dep">14807083</mods:identifier>
|
||||
<mods:identifier id="0542C573BAAFA9799E005BF5993379C1" type="ZooBank">3663E0A-2E17-4346-A388-6BEE53081814</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03CB6967FF9738403876FBBD934AFCB3" LSID="urn:lsid:plazi:treatment:03CB6967FF9738403876FBBD934AFCB3" httpUri="http://treatment.plazi.org/id/03CB6967FF9738403876FBBD934AFCB3" lastPageId="20" lastPageNumber="341" pageId="19" pageNumber="340">
|
||||
<subSubSection id="C3788BFAFF9738473876FBBD928EFB3A" pageId="19" pageNumber="340" type="nomenclature">
|
||||
<paragraph id="8BDDD871FF9738473876FBBD9422FB88" blockId="19.[842,1431,1030,1087]" pageId="19" pageNumber="340">
|
||||
<heading id="D0956F1DFF9738473876FBBD9422FB88" centered="true" fontSize="9" level="2" pageId="19" pageNumber="340" reason="2">
|
||||
<taxonomicName id="4C62A3F2FF9738473876FBBD9573FB8B" ID-CoL="782N7" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9738473876FBBD9537FBA8" bold="true" box="[842,1105,1034,1055]" italics="true" pageId="19" pageNumber="340">
|
||||
<smallCapsWord id="8D3B4EADFF9738473876FBBD9285FBA8" baselines="1050,1050" box="[842,995,1034,1055]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="19" pageNumber="340">PLEOPHYLLA</smallCapsWord>
|
||||
<smallCapsWord id="8D3B4EADFF97384738D1FBBA9537FBA8" baselines="1050" box="[1005,1105,1037,1054]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="charlyi" pageId="19" pageNumber="340">CHARLYI</smallCapsWord>
|
||||
</emphasis>
|
||||
<smallCapsWord id="8D3B4EADFF9738473F66FBB095D3FBA8" baselines="1049,1050" box="[1114,1205,1031,1055]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="19" pageNumber="340">AHRENS</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF9738473FFBFBB0944BFBA8" baselines="1049,1050" box="[1223,1325,1031,1055]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="19" pageNumber="340">BECKETT</smallCapsWord>
|
||||
,
|
||||
<smallCapsWord id="8D3B4EADFF9738473E03FBB094F1FBAB" baselines="1049,1049" box="[1343,1431,1031,1055]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="19" pageNumber="340">EBERLE</smallCapsWord>
|
||||
&
|
||||
<smallCapsWord id="8D3B4EADFF9738473881FB919573FB8B" baselines="1080,1081" box="[957,1045,1062,1086]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabrizi" pageId="19" pageNumber="340">FABRIZI</smallCapsWord>
|
||||
</taxonomicName>
|
||||
<emphasis id="B9160463FF9738473F22FB9C95E5FB89" bold="true" box="[1054,1155,1060,1087]" pageId="19" pageNumber="340">
|
||||
<taxonomicNameLabel id="A225B918FF9738473F22FB9C95E5FB89" box="[1054,1155,1060,1087]" pageId="19" pageNumber="340" rank="species">
|
||||
<smallCapsWord id="8D3B4EADFF9738473F22FB9C955CFB8A" baselines="1080" box="[1054,1082,1067,1084]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="19" pageNumber="340">SP</smallCapsWord>
|
||||
.
|
||||
<smallCapsWord id="8D3B4EADFF9738473F70FB9C951AFB8A" baselines="1080" box="[1100,1148,1067,1084]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="19" pageNumber="340">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<figureCitation id="1359C4F4FF9738473FAAFB91947CFB88" box="[1174,1306,1060,1086]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="19" pageNumber="340">
|
||||
<smallCapsWord id="8D3B4EADFF9738473FAAFB9195ADFB8B" baselines="1080,1081" box="[1174,1227,1062,1086]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="19" pageNumber="340">FIGS</smallCapsWord>
|
||||
4E
|
||||
<emphasis id="B9160463FF9738473FCBFB9F9463FB88" box="[1271,1285,1064,1086]" italics="true" pageId="19" pageNumber="340">–</emphasis>
|
||||
H
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="1359C4F4FF9738473E10FB93945AFB88" box="[1324,1340,1060,1086]" captionStart="Figure 7" captionStartId="14.[144,223,925,944]" captionTargetBox="[144,1428,192,893]" captionTargetId="figure-25@14.[144,1428,192,893]" captionTargetPageId="14" captionText="Figure 7. Distribution of Pleophylla species (left side) and absence records (right side), represented by sampling locali- ties of all other sampled Sericini species without records of Pleophylla (white dots)." figureDoi="http://doi.org/10.5281/zenodo.14807097" httpUri="https://zenodo.org/record/14807097/files/figure.png" pageId="19" pageNumber="340">7</figureCitation>
|
||||
)
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9738473807FBE0928EFB3A" blockId="19.[827,1442,1111,1164]" pageId="19" pageNumber="340">
|
||||
<uri id="FFF3D473FF9738473807FBE0928EFB3A" pageId="19" pageNumber="340">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFC4E2A4FF9738473F7FFBE0928EFB3A" pageId="19" pageNumber="340">00BFEBA3-5CCA-4189-B934- 3B4587703F0A</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3788BFAFF9738403801FB1F934AFCB3" lastPageId="20" lastPageNumber="341" pageId="19" pageNumber="340" type="materials_examined">
|
||||
<paragraph id="8BDDD871FF9738473801FB1F9542FB4C" blockId="19.[829,1444,1191,1275]" pageId="19" pageNumber="340">
|
||||
<emphasis id="B9160463FF9738473801FB1F9510FB0A" box="[829,1142,1191,1213]" italics="true" pageId="19" pageNumber="340">Type material examined:</emphasis>
|
||||
<materialsCitation id="3B0AD22CFF9738473FB9FB109542FB4C" collectingDate="1998-12-14" collectionCode="ZFMK" collectorName="Lizler" country="Tanzania" location="Southern Highland Werner" pageId="19" pageNumber="340" specimenCount="1" specimenCount-male="1" stateProvince="Njombe" typeStatus="holotype">
|
||||
<typeStatus id="54D966D3FF9738473FB9FB109588FB0B" box="[1157,1262,1191,1213]" pageId="19" pageNumber="340" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF9738473E34FB1D947EFB0A" box="[1288,1304,1194,1212]" count="1" pageId="19" pageNumber="340" type="male">♂</specimenCount>
|
||||
“
|
||||
<collectingCountry id="F37598E1FF9738473E05FB1094C5FB0B" box="[1337,1443,1191,1213]" name="Tanzania" pageId="19" pageNumber="340">Tanzania</collectingCountry>
|
||||
<date id="FFDCFEB1FF9738473801FB7192A4FB6D" box="[829,962,1222,1243]" pageId="19" pageNumber="340" value="1998-12-14">
|
||||
<collectingDate id="EF980759FF9738473801FB7192A4FB6D" box="[829,962,1222,1243]" pageId="19" pageNumber="340" value="1998-12-14">14.XII.1998</collectingDate>
|
||||
</date>
|
||||
<collectingRegion id="49A61693FF97384738EDFB71954EFB6A" box="[977,1064,1222,1244]" country="Tanzania" name="Njombe" pageId="19" pageNumber="340">Njombe</collectingRegion>
|
||||
<location id="8EBD8EAAFF9738473F0BFB7194E7FB6D" LSID="urn:lsid:plazi:treatment:03CB6967FF9738403876FBBD934AFCB3:8EBD8EAAFF9738473F0BFB7194E7FB6D" box="[1079,1409,1222,1244]" country="Tanzania" name="Southern Highland Werner" pageId="19" pageNumber="340" stateProvince="Njombe">Southern Highland Werner</location>
|
||||
&
|
||||
<collectorName id="2697BDA7FF9738473801FB5292E6FB4D" box="[829,896,1253,1275]" pageId="19" pageNumber="340">Lizler</collectorName>
|
||||
leg.” (
|
||||
<collectionCode id="ED7340B4FF97384738F5FB52957EFB4C" box="[969,1048,1253,1274]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="19" pageNumber="340" type="Museum">ZFMK</collectionCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9738403801FAA493A1FE81" blockId="19.[829,1444,1298,1901]" lastBlockId="20.[144,758,199,312]" lastPageId="20" lastPageNumber="341" pageId="19" pageNumber="340">
|
||||
<emphasis id="B9160463FF9738473801FAA492A0FA9E" box="[829,966,1299,1320]" italics="true" pageId="19" pageNumber="340">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9738473F42FAA49582FA9E" box="[1150,1252,1299,1320]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="19" pageNumber="340" unit="mm" value="12.1">12.1 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9738473801FA8692F0FAF0" box="[829,918,1329,1350]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="19" pageNumber="340" unit="mm" value="8.4">8.4 mm</quantity>
|
||||
, maximum width: 6.0 mm. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus strongly reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.7. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/ metacoxa: 1/1.39. Metatibia, ratio of width/length: 1/ 4.28. Metatarsi dorsally smooth; first metatarsomere slightly longer than subsequent one. Female unknown. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF9038403ABEFE959090FE8E" box="[386,502,290,312]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">Fig. 4E, F</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF9038403955FE9593A5FE81" box="[617,707,290,312]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">Fig. 4H</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9038403BACFEFA91B6FD22" blockId="20.[144,759,332,660]" pageId="20" pageNumber="341">
|
||||
<emphasis id="B9160463FF9038403BACFEFA9090FED7" box="[144,502,332,354]" italics="true" pageId="20" pageNumber="341">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF9038403A29FEFB9090FED7" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[277,502,332,353]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">Pleophylla charlyi</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF9038403933FEFA931EFED4" box="[527,632,333,354]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is quite similar to
|
||||
<taxonomicName id="4C62A3F2FF9038403A30FEDB90CBFE36" box="[268,429,363,385]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="navicularis">
|
||||
<emphasis id="B9160463FF9038403A30FEDB90CBFE36" box="[268,429,363,385]" italics="true" pageId="20" pageNumber="341">P. navicularis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C62A3F2FF9038403AD2FEDB9308FE36" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[494,622,363,385]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="silvatica" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9038403AD2FEDB9308FE36" box="[494,622,363,385]" italics="true" pageId="20" pageNumber="341">P. silvatica</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9038403945FEDB93B3FE37" box="[633,725,364,385]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
in external morphology.
|
||||
<taxonomicName id="4C62A3F2FF9038403A9EFE3D9318FE29" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[418,638,394,415]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9038403A9EFE3D9318FE29" box="[418,638,394,415]" italics="true" pageId="20" pageNumber="341">Pleophylla charlyi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF90384039AEFE3D9391FE29" box="[658,759,394,415]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from both in the strongly asymmetrical ventral lobes of the parameres being distinctly produced dorsally. From
|
||||
<taxonomicName id="4C62A3F2FF9038403AFBFE519319FE4C" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[455,639,485,507]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="mpumalanga" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9038403AFBFE519319FE4C" box="[455,639,485,507]" italics="true" pageId="20" pageNumber="341">P. mpumalanga</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF90384039B3FE519394FE4D" box="[655,754,486,507]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, which is similar in the shape of the parameres,
|
||||
<taxonomicName id="4C62A3F2FF9038403BACFD949066FD81" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[144,256,546,568]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9038403BACFD949066FD81" box="[144,256,546,568]" italics="true" pageId="20" pageNumber="341">P. charlyi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9038403A31FD94900CFD8E" box="[269,362,547,568]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs in the evenly distributed white pilosity on the elytra and the large lobes produced dorsally by the extensions of the ventral lobes.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9038403BACFD05934AFCB3" blockId="20.[144,758,690,773]" pageId="20" pageNumber="341">
|
||||
<emphasis id="B9160463FF9038403BACFD05906BFD71" box="[144,269,690,711]" italics="true" pageId="20" pageNumber="341">Etymology:</emphasis>
|
||||
The new species is named after one of its collectors, Karl Werner (his nickname was ‘Charly’). He passed away too early.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
224
data/03/CB/69/03CB6967FF9738473BFAFB6792B3FC79.xml
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224
data/03/CB/69/03CB6967FF9738473BFAFB6792B3FC79.xml
Normal file
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<taxonomicName id="4C62A3F2FF9738473BFAFB6790C9FAB5" ID-CoL="4K846" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="mpumalanga" status="sp. nov.">
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,
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,
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&
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</taxonomicName>
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.
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.
|
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|
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|
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(
|
||||
<figureCitation id="1359C4F4FF973847390DFB5A93D2FAB3" box="[561,692,1259,1285]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="19" pageNumber="340">
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4A
|
||||
<emphasis id="B9160463FF97384739AEFB5893C6FAB3" box="[658,672,1263,1285]" italics="true" pageId="19" pageNumber="340">–</emphasis>
|
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D
|
||||
</figureCitation>
|
||||
, 6)
|
||||
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||||
</paragraph>
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urn:lsid:zoobank.org:act:
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<emphasis id="B9160463FF9738473B9AFAD89085FA35" box="[166,483,1390,1412]" italics="true" pageId="19" pageNumber="340">Type material examined:</emphasis>
|
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<materialsCitation id="3B0AD22CFF9738473ACEFAD99021FA69" collectingDate="2002-10-28" collectionCode="TMSA" collectorName="S. Afr." latitude="-27.51" longLatPrecision="742" longitude="31.19" pageId="19" pageNumber="340" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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:
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“
|
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<collectorName id="2697BDA7FF9738473991FAD9926CFA32" box="[685,778,1390,1412]" pageId="19" pageNumber="340">S. Afr.</collectorName>
|
||||
: KwaZuluNatal Tygerkloof
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|
||||
<geoCoordinate id="EE56BEB6FF97384739ABFA3A9262FA14" box="[663,772,1421,1442]" degrees="31.19" direction="east" orientation="longitude" pageId="19" pageNumber="340" precision="555" value="31.19">31.19 E</geoCoordinate>
|
||||
/
|
||||
<date id="FFDCFEB1FF9738473B9AFA1C9042FA76" box="[166,292,1451,1472]" pageId="19" pageNumber="340" value="2002-10-28">
|
||||
<collectingDate id="EF980759FF9738473B9AFA1C9042FA76" box="[166,292,1451,1472]" pageId="19" pageNumber="340" value="2002-10-28">28.10.2002</collectingDate>
|
||||
</date>
|
||||
: E-Y: 3563 grassland & forest leg. TMSA staff” (
|
||||
<collectionCode id="ED7340B4FF9738473BCEFA7D9026FA69" box="[242,320,1482,1503]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="19" pageNumber="340">TMSA</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B0AD22CFF9738473A6FFA7D9346F9AA" collectingDate="1993-01-22" collectionCode="ZFMK" collectorName="S. Afr." latitude="-27.48" location="KwaZulu Natal Ngome State forest" longLatPrecision="742" longitude="31.25" pageId="19" pageNumber="340" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||||
<typeStatus id="54D966D3FF9738473A6FFA7D90D8FA69" box="[339,446,1482,1503]" pageId="19" pageNumber="340" type="paratype">Paratype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D6413F8FF9738473AF6FA7D9096FA69" box="[458,496,1482,1503]" count="1" pageId="19" pageNumber="340" type="male">1 ♂</specimenCount>
|
||||
“
|
||||
<collectorName id="2697BDA7FF9738473938FA7D9336FA56" box="[516,592,1482,1504]" pageId="19" pageNumber="340">S. Afr.</collectorName>
|
||||
:
|
||||
<location id="8EBD8EAAFF9738473967FA7D90EDFA48" LSID="urn:lsid:plazi:treatment:03CB6967FF9738473BFAFB6792B3FC79:8EBD8EAAFF9738473967FA7D90EDFA48" latitude="-27.48" longLatPrecision="742" longitude="31.25" name="KwaZulu Natal Ngome State forest" pageId="19" pageNumber="340">KwaZulu Natal Ngome State forest</location>
|
||||
/
|
||||
<geoCoordinate id="EE56BEB6FF9738473AA8FA5F908AFA48" box="[404,492,1512,1534]" degrees="27.48" direction="south" orientation="latitude" pageId="19" pageNumber="340" precision="555" value="-27.48">27.48 S</geoCoordinate>
|
||||
-
|
||||
<geoCoordinate id="EE56BEB6FF9738473932FA5F930DFA48" box="[526,619,1512,1534]" degrees="31.25" direction="east" orientation="longitude" pageId="19" pageNumber="340" precision="555" value="31.25">31.25 E</geoCoordinate>
|
||||
; 18-
|
||||
<date id="FFDCFEB1FF97384739A0FA5E926BFA48" box="[668,781,1513,1534]" pageId="19" pageNumber="340" value="1993-01-22">
|
||||
<collectingDate id="EF980759FF97384739A0FA5E926BFA48" box="[668,781,1513,1534]" pageId="19" pageNumber="340" value="1993-01-22">22.1.1993</collectingDate>
|
||||
</date>
|
||||
Kr uger € & Dombrowsky” (
|
||||
<collectionCode id="ED7340B4FF9738473AF6F9B0937FF9AA" box="[458,537,1543,1564]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="19" pageNumber="340" type="Museum">ZFMK</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9738473B9AF9F2951BFDFD" blockId="19.[166,781,1604,1901]" lastBlockId="19.[829,1445,199,587]" pageId="19" pageNumber="340">
|
||||
<emphasis id="B9160463FF9738473B9AF9F29049F9EC" box="[166,303,1605,1626]" italics="true" pageId="19" pageNumber="340">Description:</emphasis>
|
||||
Body length:
|
||||
<quantity id="4C9A7594FF9738473AD4F9F39328F9EC" box="[488,590,1604,1626]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="19" pageNumber="340" unit="mm" value="12.1">12.1 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF9738473B9AF9D49199F9CE" box="[166,255,1635,1656]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="19" pageNumber="340" unit="mm" value="8.9">8.9 mm</quantity>
|
||||
, maximum width:
|
||||
<quantity id="4C9A7594FF9738473ACCF9D4932FF9CE" box="[496,585,1635,1657]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="19" pageNumber="340" unit="mm" value="6.1">6.1 mm</quantity>
|
||||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.61. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum narrowly interrupted medially (distinctly less than scutellum width); larger punctures on pronotum very coarse. Hypomeron carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra distinctly elevated but convex; white, fine pilosity condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.28. Metatibia, ratio of width/length: 1/4.2. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||||
<figureCitation id="1359C4F4FF9738473801FD8292D7FDFD" box="[829,945,565,587]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="19" pageNumber="340">
|
||||
Fig. 4A
|
||||
<emphasis id="B9160463FF97384738ACFD8F92FAFDFC" box="[912,924,568,586]" italics="true" pageId="19" pageNumber="340">–</emphasis>
|
||||
C
|
||||
</figureCitation>
|
||||
. Habitus:
|
||||
<figureCitation id="1359C4F4FF9738473F1DFD82951FFDFD" box="[1057,1145,565,587]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="19" pageNumber="340">Fig. 4D</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9738473801FDD592A4FD44" blockId="19.[829,1444,609,754]" pageId="19" pageNumber="340">
|
||||
<emphasis id="B9160463FF9738473801FDD59586FDC0" box="[829,1248,609,631]" italics="true" pageId="19" pageNumber="340">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C62A3F2FF97384738FFFDD69586FDC0" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[963,1248,609,630]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="mpumalanga" status="sp. nov.">Pleophylla mpumalanga</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A225B918FF9738473FD3FDD5942BFDC1" box="[1263,1357,610,631]" pageId="19" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from
|
||||
<taxonomicName id="4C62A3F2FF9738473843FD369546FD23" box="[895,1056,640,662]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="navicularis">
|
||||
<emphasis id="B9160463FF9738473843FD369546FD23" box="[895,1056,640,662]" italics="true" pageId="19" pageNumber="340">P. navicularis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C62A3F2FF9738473F59FD369583FD23" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1125,1253,640,662]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="silvatica" status="sp. nov.">
|
||||
<emphasis id="B9160463FF9738473F59FD369583FD23" box="[1125,1253,640,662]" italics="true" pageId="19" pageNumber="340">P. silvatica</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A225B918FF9738473FCEFD379429FD23" box="[1266,1359,640,661]" pageId="19" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
in the thick, white pilosity of the elytra being condensed to longitudinal patches rather than being nearly evenly distributed.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9738473801FCBE9574FCED" blockId="19.[829,1444,776,859]" pageId="19" pageNumber="340">
|
||||
<emphasis id="B9160463FF9738473801FCBE92D7FCA8" box="[829,945,777,798]" italics="true" pageId="19" pageNumber="340">Variation:</emphasis>
|
||||
Body length: 11.0
|
||||
<emphasis id="B9160463FF9738473F82FCBC95ACFCAB" box="[1214,1226,779,797]" italics="true" pageId="19" pageNumber="340">–</emphasis>
|
||||
<quantity id="4C9A7594FF9738473FF7FCBF9457FCAB" box="[1227,1329,776,797]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="19" pageNumber="340" unit="mm" value="12.1">12.1 mm</quantity>
|
||||
, elytral length:
|
||||
<quantity id="4C9A7594FF97384738A2FC909543FC8A" box="[926,1061,807,828]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" metricValueMax="8.9" metricValueMin="8.7" pageId="19" pageNumber="340" unit="mm" value="8.8" valueMax="8.9" valueMin="8.7">
|
||||
8.7
|
||||
<emphasis id="B9160463FF97384738FCFC9E92AAFC8D" box="[960,972,809,827]" italics="true" pageId="19" pageNumber="340">–</emphasis>
|
||||
8.9 mm
|
||||
</quantity>
|
||||
, maximum width: 6.0
|
||||
<emphasis id="B9160463FF9738473E06FC9E9420FC8D" box="[1338,1350,809,827]" italics="true" pageId="19" pageNumber="340">–</emphasis>
|
||||
<quantity id="4C9A7594FF9738473E7AFC9194C6FC8A" box="[1350,1440,806,828]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="19" pageNumber="340" unit="mm" value="6.1">6.1 mm</quantity>
|
||||
. Female: unknown.
|
||||
</paragraph>
|
||||
<paragraph id="8BDDD871FF9738473801FCCB92B3FC79" blockId="19.[829,1444,892,975]" pageId="19" pageNumber="340">
|
||||
<emphasis id="B9160463FF9738473801FCCB92DDFC27" box="[829,955,892,913]" italics="true" pageId="19" pageNumber="340">Etymology:</emphasis>
|
||||
The new species is named
|
||||
<taxonomicName id="4C62A3F2FF9738473E3EFCCA94F8FC27" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[1282,1438,892,914]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="340" phylum="Arthropoda" rank="species" species="mpumalanga">
|
||||
‘
|
||||
<emphasis id="B9160463FF9738473E3BFCCB94F8FC27" box="[1287,1438,892,913]" italics="true" pageId="19" pageNumber="340">mpumalanga</emphasis>
|
||||
</taxonomicName>
|
||||
’ after its occurrence in the
|
||||
<collectingRegion id="49A61693FF9738473F43FC2C94E2FC07" box="[1151,1412,923,945]" country="South Africa" name="Mpumalanga" pageId="19" pageNumber="340">Mpumalanga Province</collectingRegion>
|
||||
of
|
||||
<collectingCountry id="F37598E1FF9738473801FC0E92B4FC79" box="[829,978,953,975]" name="South Africa" pageId="19" pageNumber="340">South Africa</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
282
data/03/EC/87/03EC8792CC20715E7FAA1417083BAEEC.xml
Normal file
282
data/03/EC/87/03EC8792CC20715E7FAA1417083BAEEC.xml
Normal file
|
|
@ -0,0 +1,282 @@
|
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<mods:roleTerm id="5AED522A74AB08CBA8A209EAC6DAB9A2">Author</mods:roleTerm>
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<mods:namePart id="14EC00F4BB8711B5F3C6D8E7D0D34F08">Packer, Laurence</mods:namePart>
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|
||||
<taxonomicName id="4C454D07CC20715E7FAA14170FA7AF82" authority="MITCHELL, 1973" authorityName="Mitchell" authorityYear="1973" box="[197,741,363,390]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="383" phylum="Arthropoda" rank="subGenus" subGenus="Xerocoelioxys">
|
||||
<emphasis id="B931EA96CC20715E7FAA14170F55AF82" box="[197,535,363,389]" italics="true" pageId="52" pageNumber="406">
|
||||
<smallCapsWord id="8D1CA058CC20715E7FAA14170C01AF85" baselines="383,382" box="[197,323,367,388]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Coelioxys" pageId="52" pageNumber="406">COELIOXYS</smallCapsWord>
|
||||
(
|
||||
<smallCapsWord id="8D1CA058CC20715E7E3914160F4CAF85" baselines="383,382" box="[342,526,366,389]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Xerocoelioxys" pageId="52" pageNumber="406">XEROCOELIOXYS</smallCapsWord>
|
||||
)
|
||||
</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC20715E7D4F14150FA7AF82" author="Mitchell TB" box="[544,741,363,390]" pageId="52" pageNumber="406" refId="ref43263" refString="Mitchell TB. 1973. A subgeneric revision of the genus Coelioxys of the Western Hemisphere (Hymenoptera: Megachilidae). Raleigh: Department of Entomology, North Carolina State University." type="book" year="1973">
|
||||
<smallCapsWord id="8D1CA058CC20715E7D4F14150FD4AF84" baselines="383,383" box="[544,662,365,389]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Mitchell" pageId="52" pageNumber="406">MITCHELL</smallCapsWord>
|
||||
, 1973
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F650FCC20715E7FCD14C4083BAEEC" pageId="52" pageNumber="406" type="materials_examined">
|
||||
<paragraph id="8BFA3684CC20715E7FCD14C40FDFAFD7" blockId="52.[162,669,443,465]" box="[162,669,443,465]" pageId="52" pageNumber="406">
|
||||
<emphasis id="B931EA96CC20715E7FCD14C40C76AFD6" box="[162,308,444,465]" italics="true" pageId="52" pageNumber="406">
|
||||
<typeStatus id="54FE8826CC20715E7FCD14C40D9AAFD6" box="[162,216,444,465]" pageId="52" pageNumber="406">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C454D07CC20715E7E2C14C30FDBAFD7" authority="Cresson, 1872" authorityName="Cresson" authorityYear="1872" box="[323,665,443,464]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="edita">
|
||||
<emphasis id="B931EA96CC20715E7E2C14C30CAFAFD7" box="[323,493,443,464]" italics="true" pageId="52" pageNumber="406">Coelioxys edita</emphasis>
|
||||
Cresson, 1872
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC20715E7FCD14810C33AA76" blockId="52.[162,777,504,1137]" pageId="52" pageNumber="406">
|
||||
<emphasis id="B931EA96CC20715E7FCD14810C5BAC09" box="[162,281,505,526]" italics="true" pageId="52" pageNumber="406">Diagnosis:</emphasis>
|
||||
Species in this subgenus are very similar to
|
||||
<emphasis id="B931EA96CC20715E7FD0176F0D93AC2B" box="[191,209,535,556]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC20715E7F86176F0C3CAC2B" box="[233,382,535,556]" italics="true" pageId="52" pageNumber="406">
|
||||
<taxonomicName id="4C454D07CC20715E7F86176F0C10AC2B" box="[233,338,535,556]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
|
||||
<taxonomicNameLabel id="A20257EDCC20715E7E34176F0C3CAC2B" box="[347,382,535,556]" pageId="52" pageNumber="406" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
) spp., but can be differentiated by the pronotal lobe distinctly lamellate (Fig. 31) or carinate throughout, whereas in species of
|
||||
<emphasis id="B931EA96CC20715E7FCD170B0DF6AC8F" box="[162,180,627,648]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC20715E7FA4170A0C23AC8F" box="[203,353,626,648]" italics="true" pageId="52" pageNumber="406">
|
||||
<taxonomicName id="4C454D07CC20715E7FA4170A0C76AC80" box="[203,308,626,647]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
|
||||
<taxonomicNameLabel id="A20257EDCC20715E7E51170B0C23AC8F" box="[318,353,627,648]" pageId="52" pageNumber="406" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
) the pronotal lobe is conspicuously rounded on its outer margin and weakly carinate on its inner margin (
|
||||
<figureCitation id="137E2A01CC20715E7E0517D70CFFACC2" box="[362,445,687,709]" captionStart="Figures 30 and 31" captionStartId="28.[162,252,820,839]" captionTargetBox="[170,1433,198,787]" captionTargetId="figure-156@28.[170,1433,198,787]" captionTargetPageId="28" captionText="Figures 30 and 31. Female, frontal view of pronotal lobe (outer margin indicated by red arrow): 30, Coelioxys sodalis; 31, Coelioxys grindeliae. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807021" httpUri="https://zenodo.org/record/14807021/files/figure.png" pageId="52" pageNumber="406">Fig. 30</figureCitation>
|
||||
). Another difference between species of these two subgenera is in the mesosomal pilosity, which is short, composed of subappressed or appressed hairs, forming marginal lines or spots on the mesoscutum in
|
||||
<emphasis id="B931EA96CC20715E7EFE16520CE1AD38" box="[401,419,810,831]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC20715E7ED516520F12AD38" box="[442,592,810,831]" italics="true" pageId="52" pageNumber="406">Xerocoelioxys</emphasis>
|
||||
) spp. (Fig. 31), contrasting with the long and dense pubescence in species of
|
||||
<emphasis id="B931EA96CC20715E7E70161F0C73AD7B" box="[287,305,871,892]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC20715E7E26161F0C9DAD7A" box="[329,479,871,893]" italics="true" pageId="52" pageNumber="406">
|
||||
<taxonomicName id="4C454D07CC20715E7E26161F0CF0AD7B" box="[329,434,871,892]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
|
||||
<taxonomicNameLabel id="A20257EDCC20715E7ED416100C9DAD7A" box="[443,479,872,893]" pageId="52" pageNumber="406" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
) (
|
||||
<figureCitation id="137E2A01CC20715E7E93161F0F0DAD7B" box="[508,591,871,893]" captionStart="Figures 30 and 31" captionStartId="28.[162,252,820,839]" captionTargetBox="[170,1433,198,787]" captionTargetId="figure-156@28.[170,1433,198,787]" captionTargetPageId="28" captionText="Figures 30 and 31. Female, frontal view of pronotal lobe (outer margin indicated by red arrow): 30, Coelioxys sodalis; 31, Coelioxys grindeliae. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807021" httpUri="https://zenodo.org/record/14807021/files/figure.png" pageId="52" pageNumber="406">Fig. 30</figureCitation>
|
||||
). The following combination of characteristics together with the two preceding ones separate species of
|
||||
<emphasis id="B931EA96CC20715E7D2F16DC0F10ADBE" box="[576,594,932,953]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC20715E7D0516DC0E42ADBE" box="[618,768,932,953]" italics="true" pageId="52" pageNumber="406">Xerocoelioxys</emphasis>
|
||||
) from all other subgenera. Mesoscutum coarsely and densely punctate, no interspaces linear to shorter than puncture diameter (as in
|
||||
<figureCitation id="137E2A01CC20715E7D0811780FB9AA12" box="[615,763,1023,1046]" captionStart="Figures 83–86" captionStartId="45.[148,238,1329,1348]" captionTargetBox="[155,1417,198,1296]" captionTargetId="figure-283@45.[155,1417,198,1296]" captionTargetPageId="45" captionText="Figures 83–86. Female. 83, 84, Coelioxys rufescens: 83, dorsal view of mesoscutellum (for pilosity on anterior margin of mesoscutellum); 84, dorsal view of terga (for apical tergal fasciae). 85, 86, Coelioxys conoidea: 85, dorsal view of mesoscutellum (for pilosity on anterior margin of mesoscutellum); 86, dorsal view of terga (for apical tergal fasciae). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807051" httpUri="https://zenodo.org/record/14807051/files/figure.png" pageId="52" pageNumber="406">Figs 83, 85</figureCitation>
|
||||
); postgradular grooves of
|
||||
<collectionCode id="ED54AE41CC20715E7D4811660F7AAA34" box="[551,568,1054,1075]" name="Tavera, Department of Geology and Geophysics" pageId="52" pageNumber="406">T</collectionCode>
|
||||
2
|
||||
<emphasis id="B931EA96CC20715E7D2A11590F13AA34" box="[581,593,1057,1075]" italics="true" pageId="52" pageNumber="406">–</emphasis>
|
||||
<collectionCode id="ED54AE41CC20715E7D3E11660F21AA34" box="[593,611,1054,1075]" name="Tavera, Department of Geology and Geophysics" pageId="52" pageNumber="406">T</collectionCode>
|
||||
3 complete, uninterrupted medially (
|
||||
<figureCitation id="137E2A01CC20715E7EAF11450F56AA55" box="[448,532,1085,1107]" captionStart="Figures 24 and 25" captionStartId="26.[162,252,1593,1612]" captionText="Figures 24 and 25. Female, dorsal view of T2–T3 (for postgradular grooves): 24, Coelioxys rufitarsis; 25, Coelioxys apacheorum. Scale bars: 1.5 mm." pageId="52" pageNumber="406">Fig. 24</figureCitation>
|
||||
), and gradulus of
|
||||
<collectionCode id="ED54AE41CC20715E7D8511450FB9AA55" box="[746,763,1085,1106]" name="Tavera, Department of Geology and Geophysics" pageId="52" pageNumber="406">T</collectionCode>
|
||||
2 bowed posteriorly.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC20715E7FCD11E10FCBA81A" blockId="52.[162,776,1176,1565]" pageId="52" pageNumber="406">
|
||||
<emphasis id="B931EA96CC20715E7FCD11E10C5DAAA9" box="[162,287,1177,1198]" italics="true" pageId="52" pageNumber="406">Comments:</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC20715E7E4111E00C8DAAA9" author="Baker JR" box="[302,463,1176,1198]" pageId="52" pageNumber="406" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker (1975)</bibRefCitation>
|
||||
revised and keyed ten species in this subgenus. As a result of our phylogenies,
|
||||
<taxonomicName id="4C454D07CC20715E7E2D11AE0CB1AAEC" authorityName="Baker" authorityYear="1975" box="[322,499,1238,1259]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="serricaudata">
|
||||
<emphasis id="B931EA96CC20715E7E2D11AE0C16AAEC" box="[322,340,1238,1259]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC20715E7E0C11AE0CB1AAEC" box="[355,499,1238,1259]" italics="true" pageId="52" pageNumber="406">serricaudata</emphasis>
|
||||
</taxonomicName>
|
||||
, previously placed in
|
||||
<emphasis id="B931EA96CC20715E7FCD118C0DF6AB0E" box="[162,180,1268,1289]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC20715E7FA4118C0C23AB0D" box="[203,353,1268,1290]" italics="true" pageId="52" pageNumber="406">
|
||||
<taxonomicName id="4C454D07CC20715E7FA4118C0C76AB0E" box="[203,308,1268,1289]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
|
||||
<taxonomicNameLabel id="A20257EDCC20715E7E51118D0C23AB0D" box="[318,353,1269,1290]" pageId="52" pageNumber="406" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
), is now regarded as belonging to
|
||||
<emphasis id="B931EA96CC20715E7FCD106B0DF6AB2F" box="[162,180,1299,1320]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC20715E7FA4106B0C23AB2F" box="[203,353,1299,1320]" italics="true" pageId="52" pageNumber="406">Xerocoelioxys</emphasis>
|
||||
). Also, considering the diagnostic characteristics of this subgenus, the other three North American species assigned by
|
||||
<bibRefCitation id="EFD44B75CC20715E7D3C10280FABAB61" author="Baker JR" box="[595,745,1360,1382]" pageId="52" pageNumber="406" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker (1975)</bibRefCitation>
|
||||
to
|
||||
<emphasis id="B931EA96CC20715E7FCD10170DF6AB83" box="[162,180,1391,1412]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC20715E7FA410160C23AB83" box="[203,353,1390,1412]" italics="true" pageId="52" pageNumber="406">
|
||||
<taxonomicName id="4C454D07CC20715E7FA410160C76AB84" box="[203,308,1390,1411]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
|
||||
<taxonomicNameLabel id="A20257EDCC20715E7E5110170C23AB83" box="[318,353,1391,1412]" pageId="52" pageNumber="406" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
),
|
||||
<taxonomicName id="4C454D07CC20715E7EEF10160D99ABA5" authority="Cockerell, 1912" authorityName="Cockerell" authorityYear="1912" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="hirsutissima">
|
||||
<emphasis id="B931EA96CC20715E7EEF10160FC5AB84" box="[384,647,1390,1411]" italics="true" pageId="52" pageNumber="406">Coelioxys hirsutissima</emphasis>
|
||||
Cockerell, 1912
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7F9D10F50F84ABA5" authority="Cockerell, 1912" authorityName="Cockerell" authorityYear="1912" box="[242,710,1421,1443]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="immaculata">
|
||||
<emphasis id="B931EA96CC20715E7F9D10F50CB5ABA5" box="[242,503,1421,1442]" italics="true" pageId="52" pageNumber="406">Coelioxys immaculata</emphasis>
|
||||
Cockerell, 1912
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C454D07CC20715E7FCD10D30F57ABC6" authority="Baker, 1975" authorityName="Baker" authorityYear="1975" box="[162,533,1451,1473]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="mitchelli">
|
||||
<emphasis id="B931EA96CC20715E7FCD10D30C38ABC7" box="[162,378,1451,1472]" italics="true" pageId="52" pageNumber="406">Coelioxys mitchelli</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC20715E7EEA10D30F57ABC6" author="Baker JR" box="[389,533,1451,1473]" pageId="52" pageNumber="406" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker, 1975</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, should be classified as
|
||||
<emphasis id="B931EA96CC20715E7F8110B20C42ABD8" box="[238,256,1482,1503]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC20715E7E7710B20CECABD8" box="[280,430,1482,1503]" italics="true" pageId="52" pageNumber="406">Xerocoelioxys</emphasis>
|
||||
). This subgenus is predominantly Nearctic but with some species reaching the Neotropical region of
|
||||
<collectingCountry id="F3527614CC20715E7D5D137F0FC4A81A" box="[562,646,1543,1565]" name="Mexico" pageId="52" pageNumber="406">Mexico</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC20715E7FCD133D0C1BA949" blockId="52.[162,776,1604,1901]" pageId="52" pageNumber="406">
|
||||
<emphasis id="B931EA96CC20715E7FCD133D0C26A85D" box="[162,356,1604,1626]" italics="true" pageId="52" pageNumber="406">List of species:</emphasis>
|
||||
<taxonomicName id="4C454D07CC20715E7E1C133C0E46A85E" authority="Cresson, 1878" authorityName="Cresson" authorityYear="1878" box="[371,772,1604,1626]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="aperta">
|
||||
<emphasis id="B931EA96CC20715E7E1C133C0F7EA85D" box="[371,572,1604,1626]" italics="true" pageId="52" pageNumber="406">Coelioxys aperta</emphasis>
|
||||
Cresson, 1878
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7FCD131B0F69A87E" authority="Hill, 1936" authorityName="Hill" authorityYear="1936" box="[162,555,1635,1657]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="bisoncornua">
|
||||
<emphasis id="B931EA96CC20715E7FCD131B0CE6A87F" box="[162,420,1635,1656]" italics="true" pageId="52" pageNumber="406">Coelioxys bisoncornua</emphasis>
|
||||
Hill, 1936
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7D51131B0C0BA890" authority="Mitchell, 1962" authorityName="Mitchell" authorityYear="1962" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="boharti">
|
||||
<emphasis id="B931EA96CC20715E7D51131B0E4AA87F" box="[574,776,1635,1656]" italics="true" pageId="52" pageNumber="406">Coelioxys boharti</emphasis>
|
||||
Mitchell, 1962
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7E3913FA0CF3A891" authorityName="Cresson" authorityYear="1872" box="[342,433,1665,1687]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="edita">
|
||||
<emphasis id="B931EA96CC20715E7E3913FA0C2AA890" box="[342,360,1666,1687]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC20715E7E1713F90CF3A891" box="[376,433,1665,1686]" italics="true" pageId="52" pageNumber="406">edita</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7EAE13F90D99A8B1" authority="Mitchell, 1962" authorityName="Mitchell" authorityYear="1962" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="galactiae">
|
||||
<emphasis id="B931EA96CC20715E7EAE13F90FDBA891" box="[449,665,1665,1686]" italics="true" pageId="52" pageNumber="406">Coelioxys galactiae</emphasis>
|
||||
Mitchell, 1962
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7E6913D80E46A8B2" authority="Cockerell, 1900" authorityName="Cockerell" authorityYear="1900" box="[262,772,1696,1718]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="grindeliae">
|
||||
<emphasis id="B931EA96CC20715E7E6913D80F48A8B2" box="[262,522,1696,1717]" italics="true" pageId="52" pageNumber="406">Coelioxys grindeliae</emphasis>
|
||||
Cockerell, 1900
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7FCD13C70C10A8D4" authorityName="Cockerell" authorityYear="1912" box="[162,338,1726,1748]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="hirsutissima">
|
||||
<emphasis id="B931EA96CC20715E7FCD13C70C10A8D4" box="[162,338,1726,1748]" italics="true" pageId="52" pageNumber="406">C. hirsutissima</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7E1B13C70F5CA8D4" authorityName="Cockerell" authorityYear="1912" box="[372,542,1726,1748]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="immaculata">
|
||||
<emphasis id="B931EA96CC20715E7E1B13C70CC4A8D3" box="[372,390,1727,1748]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC20715E7EFA13C60F5CA8D4" box="[405,542,1726,1747]" italics="true" pageId="52" pageNumber="406">immaculata</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7D2F13C60C15A8F5" authority="Cockerell, 1921" authorityName="Cockerell" authorityYear="1921" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="mesae">
|
||||
<emphasis id="B931EA96CC20715E7D2F13C60E4AA8D3" box="[576,776,1726,1748]" italics="true" pageId="52" pageNumber="406">Coelioxys mesae</emphasis>
|
||||
Cockerell, 1921
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7E0813A50CAEA8F5" authorityName="Baker" authorityYear="1975" box="[359,492,1757,1778]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="mitchelli">
|
||||
<emphasis id="B931EA96CC20715E7E0813A50C3BA8F5" box="[359,377,1757,1778]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC20715E7EE713A50CAEA8F5" box="[392,492,1757,1778]" italics="true" pageId="52" pageNumber="406">mitchelli</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7E9013A50D99A916" authority="Baker, 1975" authorityName="Baker" authorityYear="1975" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="nodis">
|
||||
<emphasis id="B931EA96CC20715E7E9013A50FF3A8F5" box="[511,689,1757,1778]" italics="true" pageId="52" pageNumber="406">Coelioxys nodis</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC20715E7DD313A50D99A916" author="Baker JR" pageId="52" pageNumber="406" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker, 1975</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7E7F13840E46A916" authority="Crawford, 1914" authorityName="Crawford" authorityYear="1914" box="[272,772,1788,1810]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="piercei">
|
||||
<emphasis id="B931EA96CC20715E7E7F13840CB7A916" box="[272,501,1788,1809]" italics="true" pageId="52" pageNumber="406">Coelioxys piercei</emphasis>
|
||||
Crawford, 1914
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7FCD12630C11A928" authorityName="Baker" authorityYear="1975" box="[162,339,1818,1840]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="serricaudata">
|
||||
<emphasis id="B931EA96CC20715E7FCD12630DF6A937" box="[162,180,1819,1840]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC20715E7FAC12620C11A928" box="[195,339,1818,1839]" italics="true" pageId="52" pageNumber="406">serricaudata</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7E0A12630C82A937" box="[357,448,1819,1840]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="soror">
|
||||
<emphasis id="B931EA96CC20715E7E0A12630C35A937" box="[357,375,1819,1840]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC20715E7EE912630C82A937" box="[390,448,1819,1840]" italics="true" pageId="52" pageNumber="406">soror</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C454D07CC20715E7D6912620C17A949" authority="Cockerell, 1909" authorityName="Cockerell" authorityYear="1909" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="soledadensis">
|
||||
<emphasis id="B931EA96CC20715E7D6912620E4AA928" box="[518,776,1818,1839]" italics="true" pageId="52" pageNumber="406">Coelioxys soledadensis</emphasis>
|
||||
Cockerell, 1909
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC20715E7FD5122F0FF1A96A" blockId="52.[162,776,1604,1901]" box="[186,691,1879,1901]" pageId="52" pageNumber="406">Six new subgenera are described as follows.</paragraph>
|
||||
<paragraph id="8BFA3684CC20715E7C3115AE083BAEEC" blockId="52.[833,1430,210,328]" box="[862,1401,210,237]" pageId="52" pageNumber="406">
|
||||
<taxonomicName id="4C454D07CC20715E7C3115AE0990AEEA" box="[862,1234,210,237]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="390" phylum="Arthropoda" rank="subGenus" subGenus="Austrocleptria">
|
||||
<smallCapsWord id="8D1CA058CC20715E7C3115AE0EA6AEED" baselines="230" box="[862,996,214,235]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="oelioxys" pageId="52" pageNumber="406">
|
||||
<emphasis id="B931EA96CC20715E7C3115AE0EA6AEED" bold="true" box="[862,996,214,235]" italics="true" pageId="52" pageNumber="406">COELIOXYS</emphasis>
|
||||
</smallCapsWord>
|
||||
<emphasis id="B931EA96CC20715E7C8215AA0EBAAEEA" bold="true" box="[1005,1016,210,237]" pageId="52" pageNumber="406">(</emphasis>
|
||||
<smallCapsWord id="8D1CA058CC20715E7C9715AC0984AEED" baselines="230,230" box="[1016,1222,212,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Austrocleptria" pageId="52" pageNumber="406">
|
||||
<emphasis id="B931EA96CC20715E7C9715AC0984AEED" bold="true" box="[1016,1222,212,236]" italics="true" pageId="52" pageNumber="406">AUSTROCLEPTRIA</emphasis>
|
||||
</smallCapsWord>
|
||||
<emphasis id="B931EA96CC20715E7BA815AA0990AEEA" bold="true" box="[1223,1234,210,237]" pageId="52" pageNumber="406">)</emphasis>
|
||||
</taxonomicName>
|
||||
<smallCapsWord id="8D1CA058CC20715E7BB415AC0871AEEA" baselines="230,231" box="[1243,1331,211,237]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Rocha" pageId="52" pageNumber="406">
|
||||
<collectionCode id="ED54AE41CC20715E7BB415AC0871AEEA" box="[1243,1331,211,237]" pageId="52" pageNumber="406">ROCHA-</collectionCode>
|
||||
</smallCapsWord>
|
||||
<smallCapsWord id="8D1CA058CC20715E7A5C15AC083BAEEC" baselines="230,231" box="[1331,1401,212,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Filho" pageId="52" pageNumber="406">
|
||||
<collectionCode id="ED54AE41CC20715E7A5C15AC083BAEEC" box="[1331,1401,212,236]" pageId="52" pageNumber="406">FILHO</collectionCode>
|
||||
</smallCapsWord>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
203
data/03/EC/87/03EC8792CC2671587C3814F5084FABC6.xml
Normal file
203
data/03/EC/87/03EC8792CC2671587C3814F5084FABC6.xml
Normal file
|
|
@ -0,0 +1,203 @@
|
|||
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||||
<mods:namePart id="85ECA27A7C6E355B147D69A56487A306">Packer, Laurence</mods:namePart>
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|
||||
<subSubSection id="C35F650FCC2671587C3814F508C2AFA4" box="[855,1408,393,420]" pageId="50" pageNumber="404" type="nomenclature">
|
||||
<paragraph id="8BFA3684CC2671587C3814F508C2AFA4" blockId="50.[855,1408,393,420]" box="[855,1408,393,420]" pageId="50" pageNumber="404">
|
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<heading id="D0B281E8CC2671587C3814F508C2AFA4" box="[855,1408,393,420]" centered="true" fontSize="9" level="2" pageId="50" pageNumber="404" reason="2">
|
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<taxonomicName id="4C454D07CC2671587C3814F508C2AFA4" authority="MITCHELL, 1973" authorityName="Mitchell" authorityYear="1973" box="[855,1408,393,420]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="384" phylum="Arthropoda" rank="subGenus" subGenus="Platycoelioxys">
|
||||
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|
||||
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|
||||
(
|
||||
<smallCapsWord id="8D1CA058CC2671587C8714F409EBAFA6" baselines="413,413" box="[1000,1193,396,419]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Platycoelioxys" pageId="50" pageNumber="404">PLATYCOELIOXYS</smallCapsWord>
|
||||
)
|
||||
</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC2671587BD314F308C2AFA4" author="Mitchell TB" box="[1212,1408,393,420]" pageId="50" pageNumber="404" refId="ref43263" refString="Mitchell TB. 1973. A subgeneric revision of the genus Coelioxys of the Western Hemisphere (Hymenoptera: Megachilidae). Raleigh: Department of Entomology, North Carolina State University." type="book" year="1973">
|
||||
<smallCapsWord id="8D1CA058CC2671587BD314F30873AFA5" baselines="413,414" box="[1212,1329,395,419]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Mitchell" pageId="50" pageNumber="404">MITCHELL</smallCapsWord>
|
||||
, 1973
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F650FCC2671587C5714A2084FABC6" pageId="50" pageNumber="404" type="materials_examined">
|
||||
<paragraph id="8BFA3684CC2671587C5714A20826AFE8" blockId="50.[824,1380,474,496]" box="[824,1380,474,496]" pageId="50" pageNumber="404">
|
||||
<emphasis id="B931EA96CC2671587C5714A20E89AFE8" box="[824,971,474,495]" italics="true" pageId="50" pageNumber="404">
|
||||
<typeStatus id="54FE8826CC2671587C5714A20E2CAFE8" box="[824,878,474,495]" pageId="50" pageNumber="404">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C454D07CC2671587CB514A20823AFE8" authority="Friese, 1921" authorityName="Friese" authorityYear="1921" box="[986,1377,474,496]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="alatiformis">
|
||||
<emphasis id="B931EA96CC2671587CB514A2098BAFE8" box="[986,1225,474,495]" italics="true" pageId="50" pageNumber="404">Coelioxys alatiformis</emphasis>
|
||||
Friese, 1921
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2671587C57176F0EFAAB40" blockId="50.[824,1440,535,1351]" pageId="50" pageNumber="404">
|
||||
<emphasis id="B931EA96CC2671587C57176F0EF2AC2B" box="[824,944,535,556]" italics="true" pageId="50" pageNumber="404">Diagnosis:</emphasis>
|
||||
This monotypic subgenus can be distinguished from all other
|
||||
<taxonomicName id="4C454D07CC2671587BE0174D09BAAC4D" box="[1167,1272,565,586]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B931EA96CC2671587BE0174D09BAAC4D" box="[1167,1272,565,586]" italics="true" pageId="50" pageNumber="404">Coelioxys</emphasis>
|
||||
</taxonomicName>
|
||||
subgenera by the shape of S
|
||||
<quantity id="4CBD9B61CC2671587C87172C095BAC6D" box="[1000,1049,596,618]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="50" pageNumber="404" unit="in" value="5.0">5 in</quantity>
|
||||
females, which is spatulate and greatly expanded towards the apex, its margins somewhat swollen and shining, and ventral surface convex (
|
||||
<figureCitation id="137E2A01CC2671587CF317D70EADACC2" box="[924,1007,687,709]" captionStart="Figures 44 and 45" captionStartId="31.[148,238,847,866]" captionTargetBox="[186,1418,198,814]" captionTargetId="figure-121@31.[156,1420,198,814]" captionTargetPageId="31" captionText="Figures 44 and 45. Coelioxys alatiformis: 44, female, dorsal view of T6, S6 and S5 (indicated by black arrow); 45, male, dorsal view of T2–T3 (foveae indicated by red arrows). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807031" httpUri="https://zenodo.org/record/14807031/files/figure.png" pageId="50" pageNumber="404">Fig. 44</figureCitation>
|
||||
), and by the postgradular areas of
|
||||
<collectionCode id="ED54AE41CC2671587C5717B60E0BACE4" box="[824,841,718,739]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="404">T</collectionCode>
|
||||
2
|
||||
<emphasis id="B931EA96CC2671587C3917A90E20ACE4" box="[854,866,721,739]" italics="true" pageId="50" pageNumber="404">–</emphasis>
|
||||
<collectionCode id="ED54AE41CC2671587C0C17B60E37ACE4" box="[867,885,718,739]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="404">T</collectionCode>
|
||||
<quantity id="4CBD9B61CC2671587C1A17B60EE7ACE3" box="[885,933,718,740]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="50" pageNumber="404" unit="in" value="3.0">3 in</quantity>
|
||||
males that are foveate in a manner not found in other subgenera (Fig. 45): on
|
||||
<collectionCode id="ED54AE41CC2671587A6B17950854AD05" box="[1284,1302,749,770]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="404">T</collectionCode>
|
||||
2, the fovea is elliptical and covered with minute whitish hairs, whereas in
|
||||
<collectionCode id="ED54AE41CC2671587CAE16520E91AD38" box="[961,979,810,831]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="404">T</collectionCode>
|
||||
3 it is smaller and circular (Fig. 45). In females from other subgenera the S5 is elongate towards apex, never spatulate nor greatly expanded, except in
|
||||
<taxonomicName id="4C454D07CC2671587CDD16FE0984AD9D" authorityName="Forster" authorityYear="1853" box="[946,1222,901,923]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="alata" subGenus="Paracoelioxys">
|
||||
<emphasis id="B931EA96CC2671587CDD16FE0E86AD9C" box="[946,964,902,923]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587CB316FD0934AD9D" box="[988,1142,901,922]" italics="true" pageId="50" pageNumber="404">Paracoelioxys</emphasis>
|
||||
)
|
||||
<emphasis id="B931EA96CC2671587BE416FD0984AD9D" box="[1163,1222,901,922]" italics="true" pageId="50" pageNumber="404">alata</emphasis>
|
||||
</taxonomicName>
|
||||
, in which the S5 margins are greatly expanded but straight, never swollen, as in
|
||||
<taxonomicName id="4C454D07CC2671587C9C16BB09D0ADD0" authorityName="Friese" authorityYear="1921" box="[1011,1170,962,984]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="alatiformis">
|
||||
<emphasis id="B931EA96CC2671587C9C16BB0947ADDF" box="[1011,1029,963,984]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587B7B16BA09D0ADD0" box="[1044,1170,962,983]" italics="true" pageId="50" pageNumber="404">alatiformis</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="137E2A01CC2671587BC616BA09BFADDF" box="[1193,1277,962,984]" captionStart="Figures 44 and 45" captionStartId="31.[148,238,847,866]" captionTargetBox="[186,1418,198,814]" captionTargetId="figure-121@31.[156,1420,198,814]" captionTargetPageId="31" captionText="Figures 44 and 45. Coelioxys alatiformis: 44, female, dorsal view of T6, S6 and S5 (indicated by black arrow); 45, male, dorsal view of T2–T3 (foveae indicated by red arrows). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807031" httpUri="https://zenodo.org/record/14807031/files/figure.png" pageId="50" pageNumber="404">Fig. 44</figureCitation>
|
||||
); the ventral surface of S5 is concave and the apex is broad and truncate, in contrast to the convex surface and ellipsoid apex in
|
||||
<taxonomicName id="4C454D07CC2671587B7D116609F0AA34" authorityName="Friese" authorityYear="1921" box="[1042,1202,1054,1076]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="alatiformis">
|
||||
<emphasis id="B931EA96CC2671587B7D11660966AA34" box="[1042,1060,1054,1075]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587B5B116609F0AA34" box="[1076,1202,1054,1075]" italics="true" pageId="50" pageNumber="404">alatiformis</emphasis>
|
||||
</taxonomicName>
|
||||
. The fovea on the postgradular area of
|
||||
<collectionCode id="ED54AE41CC2671587B3611450928AA55" box="[1113,1130,1085,1106]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="404">T</collectionCode>
|
||||
2 of males from other subgenera is never elliptical, hairy and deep, but slender as in
|
||||
<taxonomicName id="4C454D07CC2671587C961102093CAA88" baseAuthorityName="Illiger" baseAuthorityYear="1806" box="[1017,1150,1146,1167]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="conoidea">
|
||||
<emphasis id="B931EA96CC2671587C9611020949AA88" box="[1017,1035,1146,1167]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587B741102093CAA88" box="[1051,1150,1146,1167]" italics="true" pageId="50" pageNumber="404">conoidea</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="137E2A01CC2671587BF5110209AFAA88" box="[1178,1261,1146,1168]" captionStart="Figures 87–90" captionStartId="46.[162,252,1545,1564]" captionTargetBox="[177,1417,197,1511]" captionTargetId="figure-170@46.[167,1429,197,1511]" captionTargetPageId="46" captionText="Figures 87–90. 87, 88, Coelioxys rufescens: 87, female, ventral view of S6 (S6 notch indicated by black arrow); 88, male, dorsal view of T2 (fovea indicated by black arrow). 89, 90, Coelioxys conoidea: 89, female, dorsal view of T6 and S6; 90, male, dorsal view of T2 (fovea indicated by black arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807053" httpUri="https://zenodo.org/record/14807053/files/figure.png" pageId="50" pageNumber="404">Fig. 90</figureCitation>
|
||||
),
|
||||
<taxonomicName id="4C454D07CC2671587A61110208DAAA88" authorityName="Lepeletier & Serville" authorityYear="1825" box="[1294,1432,1146,1167]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="rufescens">
|
||||
<emphasis id="B931EA96CC2671587A6111020862AA88" box="[1294,1312,1146,1167]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587A5F110208DAAA88" box="[1328,1432,1146,1167]" italics="true" pageId="50" pageNumber="404">rufescens</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="B931EA96CC2671587C5711E10E08AAA9" box="[824,842,1177,1198]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587C0D11E00EBEAAAA" box="[866,1020,1176,1197]" italics="true" pageId="50" pageNumber="404">Paracoelioxys</emphasis>
|
||||
) (
|
||||
<figureCitation id="137E2A01CC2671587B7111E00932AAA9" box="[1054,1136,1176,1198]" captionStart="Figures 87–90" captionStartId="46.[162,252,1545,1564]" captionTargetBox="[177,1417,197,1511]" captionTargetId="figure-170@46.[167,1429,197,1511]" captionTargetPageId="46" captionText="Figures 87–90. 87, 88, Coelioxys rufescens: 87, female, ventral view of S6 (S6 notch indicated by black arrow); 88, male, dorsal view of T2 (fovea indicated by black arrow). 89, 90, Coelioxys conoidea: 89, female, dorsal view of T6 and S6; 90, male, dorsal view of T2 (fovea indicated by black arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807053" httpUri="https://zenodo.org/record/14807053/files/figure.png" pageId="50" pageNumber="404">Fig. 88</figureCitation>
|
||||
), and
|
||||
<emphasis id="B931EA96CC2671587BA511E1099EAAA9" box="[1226,1244,1177,1198]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587B9B11E008D5AAAA" box="[1268,1431,1176,1197]" italics="true" pageId="50" pageNumber="404">Boreocoelioxys</emphasis>
|
||||
) (
|
||||
<figureCitation id="137E2A01CC2671587C2E11CF0ED6AACB" box="[833,916,1207,1229]" captionStart="Figures 26–29" captionStartId="27.[148,238,1449,1468]" captionTargetBox="[157,1420,199,1417]" captionTargetId="figure-252@27.[156,1420,198,1416]" captionTargetPageId="27" captionText="Figures 26–29. 26, female, Coelioxys porterae: dorsal view of T6 and S6. 27, male, Coelioxys sayi: dorsal view of T2 (fovea indicated by red arrow). 28, female, Coelioxys quadridentata, dorsal view of T6 and S6. 29, male, Coelioxys sodalis: dorsal view of T2 (punctate area indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807017" httpUri="https://zenodo.org/record/14807017/files/figure.png" pageId="50" pageNumber="404">Fig. 27</figureCitation>
|
||||
); minute and circular, as in
|
||||
<emphasis id="B931EA96CC2671587C5611AE0E09AAEC" box="[825,843,1238,1259]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587C0D11AE094DAAEC" box="[866,1039,1238,1259]" italics="true" pageId="50" pageNumber="404">Glyptocoelioxys</emphasis>
|
||||
) (
|
||||
<figureCitation id="137E2A01CC2671587B4511AE093FAAEC" box="[1066,1149,1238,1260]" captionStart="Figures 77–80" captionStartId="43.[148,238,1583,1602]" captionTargetBox="[156,1418,195,1549]" captionTargetId="figure-195@43.[156,1418,195,1549]" captionTargetPageId="43" captionText="Figures 77–80. 77, female, Coelioxys praetextata: dorsal view of T6 and S6 (S6 notch indicated by black arrow). 78, Coelioxys cayennensis: dorsal view of T2–T3 (punctate areas and foveae indicated by red arrows). 79, Coelioxys pergandei: dorsal view of T6 and S6 (S6 notch indicated by black arrow). 80, Coelioxys kuscheli: dorsal view of T2–T3 (fovea indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807047" httpUri="https://zenodo.org/record/14807047/files/figure.png" pageId="50" pageNumber="404">Fig. 78</figureCitation>
|
||||
), except
|
||||
<taxonomicName id="4C454D07CC2671587B8A11AE0828AAEC" authorityName="Reed" authorityYear="1892" box="[1253,1386,1238,1259]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="chilensis">
|
||||
<emphasis id="B931EA96CC2671587B8A11AE09B5AAEC" box="[1253,1271,1238,1259]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587A6811AE0828AAEC" box="[1287,1386,1238,1259]" italics="true" pageId="50" pageNumber="404">chilensis</emphasis>
|
||||
</taxonomicName>
|
||||
[see Discussion of
|
||||
<emphasis id="B931EA96CC2671587CB0118C0EB3AB0E" box="[991,1009,1268,1289]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587B66118C09F4AB0E" box="[1033,1206,1268,1289]" italics="true" pageId="50" pageNumber="404">Glyptocoelioxys</emphasis>
|
||||
)], and circular and bare, as in
|
||||
<taxonomicName id="4C454D07CC2671587B65106B09C2AB2F" authorityName="Cockerell" authorityYear="1905" box="[1034,1152,1299,1320]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="reginae">
|
||||
<emphasis id="B931EA96CC2671587B65106B095EAB2F" box="[1034,1052,1299,1320]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587B43106B09C2AB2F" box="[1068,1152,1299,1320]" italics="true" pageId="50" pageNumber="404">reginae</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2671587BC6106B0808AB2F" authorityName="Cockerell" authorityYear="1905" box="[1193,1354,1299,1320]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="albolineata">
|
||||
<emphasis id="B931EA96CC2671587BC6106B09F9AB2F" box="[1193,1211,1299,1320]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587BA4106B0808AB2F" box="[1227,1354,1299,1320]" italics="true" pageId="50" pageNumber="404">albolineata</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C454D07CC2671587C56104A0EF3AB41" authorityName="Smith" authorityYear="1875" box="[825,945,1329,1351]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="confusa">
|
||||
<emphasis id="B931EA96CC2671587C56104A0E09AB40" box="[825,843,1330,1351]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587C3510490EF3AB41" box="[858,945,1329,1350]" italics="true" pageId="50" pageNumber="404">confusa</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2671587C561017084FABC6" blockId="50.[825,1439,1390,1473]" pageId="50" pageNumber="404">
|
||||
<emphasis id="B931EA96CC2671587C5610170EF4AB83" box="[825,950,1391,1412]" italics="true" pageId="50" pageNumber="404">Comments:</emphasis>
|
||||
This subgenus comprises only one species,
|
||||
<taxonomicName id="4C454D07CC2671587CF510F5097BABA5" authorityName="Friese" authorityYear="1921" box="[922,1081,1421,1442]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="alatiformis">
|
||||
<emphasis id="B931EA96CC2671587CF510F50EEEABA5" box="[922,940,1421,1442]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587CD410F5097BABA5" box="[955,1081,1421,1442]" italics="true" pageId="50" pageNumber="404">alatiformis</emphasis>
|
||||
</taxonomicName>
|
||||
, which ranges from
|
||||
<collectingCountry id="F3527614CC2671587A4010F5083DABA4" box="[1327,1407,1421,1443]" name="Mexico" pageId="50" pageNumber="404">Mexico</collectingCountry>
|
||||
to
|
||||
<collectingCountry id="F3527614CC2671587C5610D30EEEABC6" box="[825,940,1451,1473]" name="Argentina" pageId="50" pageNumber="404">Argentina</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="EFD44B75CC2671587CD410D30843ABC6" author="Rocha-Filho LC & Packer L" box="[955,1281,1451,1473]" pageId="50" pageNumber="404" pagination="151 - 203" refId="ref43662" refString="Rocha-Filho LC, Packer L. 2015. Revision of the Neotropical subgenera Coelioxys (Platycoelioxys) Mitchell and C. (Rhinocoelioxys) Mitchell (Hymenoptera; Megachilidae) with the description of one new species. Zootaxa 3941: 151-203." type="journal article" year="2015">Rocha-Filho & Packer, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
215
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Normal file
215
data/03/EC/87/03EC8792CC2671587FCD170B086AAF50.xml
Normal file
|
|
@ -0,0 +1,215 @@
|
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<mods:title id="C8A56D6B5D219313E5590A84A8894081">Phylogeny of the cleptoparasitic Megachilini genera Coelioxys and Radoszkowskiana, with the description of six new subgenera in Coelioxys (Hymenoptera: Megachilidae)</mods:title>
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|
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<mods:namePart id="4F9310C7B9B5686F100828C9C113CEF2">Packer, Laurence</mods:namePart>
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||||
<mods:part id="0796A18519383535FE077340F5B65445">
|
||||
<mods:date id="460919850A210BC2ACB15A9FACC15812">2017</mods:date>
|
||||
<mods:detail id="72C27326926C6166F31C529BB1651A50" type="pubDate">
|
||||
<mods:number id="CF76D85C6CC5C29A6A144872A7BC6B41">2016-10-14</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="DF4319BF6E09B02DED19B5578C1170CB" type="volume">
|
||||
<mods:number id="463BA3C89170AE8CA4B8BEE236B9AFAB">180</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="BF89378F371BA9B5A725E5B33243229C" type="issue">
|
||||
<mods:number id="57525DF0BF42C45AC4E0854137DEEF2F">2</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="D0C9C633F9730D52467E061A870DB06D" unit="page">
|
||||
<mods:start id="041003A394AF426D37C7CA1416FD7F9A">354</mods:start>
|
||||
<mods:end id="0AE467B0DA47BC235BCC88345121DBAA">413</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location id="4628087882CBD9DC0CAE044F9DEA502D">
|
||||
<mods:url id="A1CB5DEC616ECAB11926CC9D8DB144D7">https://doi.org/10.1111/zoj.12484</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="FFA841D9F7AE5AEB291BEDC8D131AEEC">journal article</mods:classification>
|
||||
<mods:identifier id="5D4F898BC029829C51C75C3E862C761D" type="DOI">10.1111/zoj.12484</mods:identifier>
|
||||
<mods:identifier id="1BE2E1E924E7CE1B494627B996123EB4" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="CC691123A2845D42187E6C4D05C6F31C" type="Zenodo-Dep">14806997</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03EC8792CC2671587FCD170B086AAF50" LSID="urn:lsid:plazi:treatment:03EC8792CC2671587FCD170B086AAF50" httpUri="http://treatment.plazi.org/id/03EC8792CC2671587FCD170B086AAF50" lastPageNumber="404" pageId="50" pageNumber="404">
|
||||
<subSubSection id="C35F650FCC2671587FCD170B0E43AC8F" box="[162,769,626,648]" pageId="50" pageNumber="404" type="nomenclature">
|
||||
<paragraph id="8BFA3684CC2671587FCD170B0E43AC8F" blockId="50.[162,769,626,648]" box="[162,769,626,648]" pageId="50" pageNumber="404">
|
||||
<heading id="D0B281E8CC2671587FCD170B0E43AC8F" box="[162,769,626,648]" centered="true" fontSize="9" level="2" pageId="50" pageNumber="404" reason="2">
|
||||
=
|
||||
<taxonomicName id="4C454D07CC2671587FD7170A0FD3AC80" authority="Mitchell, 1973" authorityName="Mitchell" authorityYear="1973" box="[184,657,626,648]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="subGenus" status="syn. nov." subGenus="Schizocoelioxys">
|
||||
<emphasis id="B931EA96CC2671587FD7170A0C63AC80" box="[184,289,626,647]" italics="true" pageId="50" pageNumber="404">Coelioxys</emphasis>
|
||||
(
|
||||
<emphasis id="B931EA96CC2671587E5E170A0C9CAC80" box="[305,478,626,647]" italics="true" pageId="50" pageNumber="404">Schizocoelioxys</emphasis>
|
||||
)
|
||||
<bibRefCitation id="EFD44B75CC2671587E81170A0FD3AC80" author="Mitchell TB" box="[494,657,626,648]" pageId="50" pageNumber="404" refId="ref43263" refString="Mitchell TB. 1973. A subgeneric revision of the genus Coelioxys of the Western Hemisphere (Hymenoptera: Megachilidae). Raleigh: Department of Entomology, North Carolina State University." type="book" year="1973">Mitchell, 1973</bibRefCitation>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A20257EDCC2671587DF5170B0E43AC8F" box="[666,769,627,648]" pageId="50" pageNumber="404">syn. nov.</taxonomicNameLabel>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F650FCC2671587FCD17C8086AAF50" pageId="50" pageNumber="404" type="materials_examined">
|
||||
<paragraph id="8BFA3684CC2671587FCD17C80FDAACC2" blockId="50.[162,664,687,709]" box="[162,664,687,709]" pageId="50" pageNumber="404">
|
||||
<emphasis id="B931EA96CC2671587FCD17C80C76ACC2" box="[162,308,688,709]" italics="true" pageId="50" pageNumber="404">
|
||||
<typeStatus id="54FE8826CC2671587FCD17C80D9AACC2" box="[162,216,688,709]" pageId="50" pageNumber="404">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C454D07CC2671587E2C17D70FD6ACC2" authority="Forster, 1853" authorityName="Forster" authorityYear="1853" box="[323,660,687,709]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="alata">
|
||||
<emphasis id="B931EA96CC2671587E2C17D70CADACC3" box="[323,495,687,708]" italics="true" pageId="50" pageNumber="404">Coelioxys alata</emphasis>
|
||||
Forster, 1853
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2671587FCD17950F56ABC6" blockId="50.[162,777,748,1473]" pageId="50" pageNumber="404">
|
||||
<emphasis id="B931EA96CC2671587FCD17950C5BAD05" box="[162,281,749,770]" italics="true" pageId="50" pageNumber="404">Diagnosis:</emphasis>
|
||||
This subgenus can be differentiated from all the other subgenera based upon the mandible that is angled dorsally on the outer surface in females (
|
||||
<figureCitation id="137E2A01CC2671587FC516300DB9AD5A" box="[170,251,840,862]" captionStart="Figures 81 and 82" captionStartId="44.[162,252,779,798]" captionTargetBox="[168,1430,198,746]" captionTargetId="figure-571@44.[168,1430,198,746]" captionTargetPageId="44" captionText="Figures 81 and 82. Female, frontal view of head (outer surface of mandible indicated by black arrows): 81, Coelioxys funeraria; 82, Coelioxys conoidea. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807049" httpUri="https://zenodo.org/record/14807049/files/figure.png" pageId="50" pageNumber="404">Fig. 81</figureCitation>
|
||||
).
|
||||
<taxonomicName id="4C454D07CC2671587E7E16300F6BAD5A" authorityName="Gribodo" authorityYear="1884" box="[273,553,840,861]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="subGenus" subGenus="Paracoelioxys">
|
||||
<emphasis id="B931EA96CC2671587E7E16300C3AAD5A" box="[273,376,840,861]" italics="true" pageId="50" pageNumber="404">Coelioxys</emphasis>
|
||||
(
|
||||
<emphasis id="B931EA96CC2671587EE516300F63AD5A" box="[394,545,840,861]" italics="true" pageId="50" pageNumber="404">Paracoelioxys</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
spp. are similar to species of
|
||||
<emphasis id="B931EA96CC2671587E7B161F0C64AD7B" box="[276,294,871,892]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587E52161F0C9EAD7B" box="[317,476,871,892]" italics="true" pageId="50" pageNumber="404">Boreocoelioxys</emphasis>
|
||||
), but can be distinguished by the postgradular grooves of
|
||||
<collectionCode id="ED54AE41CC2671587D5F16FD0F03AD9D" box="[560,577,901,922]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="404">T</collectionCode>
|
||||
2
|
||||
<emphasis id="B931EA96CC2671587D2216F00F1BAD9D" box="[589,601,904,922]" italics="true" pageId="50" pageNumber="404">–</emphasis>
|
||||
<collectionCode id="ED54AE41CC2671587D3616FD0F2EAD9D" box="[601,620,901,922]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="404">T</collectionCode>
|
||||
3, which are broadly interrupted medially in the former (Fig. 25) and complete in the latter (
|
||||
<figureCitation id="137E2A01CC2671587EB016BA0F70ADDF" box="[479,562,962,984]" captionStart="Figures 24 and 25" captionStartId="26.[162,252,1593,1612]" captionText="Figures 24 and 25. Female, dorsal view of T2–T3 (for postgradular grooves): 24, Coelioxys rufitarsis; 25, Coelioxys apacheorum. Scale bars: 1.5 mm." pageId="50" pageNumber="404">Fig. 24</figureCitation>
|
||||
). They can also be differentiated from species of both of the new subgenera described herein,
|
||||
<emphasis id="B931EA96CC2671587E8C11780CB7AA12" box="[483,501,1024,1045]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587D6316870FE1AA13" box="[524,675,1023,1044]" italics="true" pageId="50" pageNumber="404">Melissoctonia</emphasis>
|
||||
) subgen. nov. and
|
||||
<emphasis id="B931EA96CC2671587E6211660C5CAA34" box="[269,286,1054,1075]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587E5911670CE9AA33" box="[310,427,1055,1076]" italics="true" pageId="50" pageNumber="404">Rozeniana</emphasis>
|
||||
) subgen. nov., as well as from all other subgenera, by the following combination of characteristics. Axilla very short, free apical portion much shorter than basal portion, and not produced beyond posterior margin of mesoscutellum (
|
||||
<figureCitation id="137E2A01CC2671587DC511E00FBEAAA9" box="[682,764,1176,1198]" captionStart="Figures 69–72" captionStartId="41.[148,238,1402,1421]" captionTargetBox="[155,1417,195,1366]" captionTargetId="figure-280@41.[155,1417,195,1366]" captionTargetPageId="41" captionText="Figures 69–72. Female. 69, Coelioxys cayennensis: dorsal view of mesoscutellum (for posterior margin). 70, Coelioxys praetextata: lateral view of S1 (basomedial area indicated by black arrow). 71, Coelioxys alata: dorsal view of mesoscutellum (for posterior margin). 72, Coelioxys funeraria: lateral view of S1 (basomedial area indicated by black arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807043" httpUri="https://zenodo.org/record/14807043/files/figure.png" pageId="50" pageNumber="404">Fig. 71</figureCitation>
|
||||
); apical fascia on
|
||||
<collectionCode id="ED54AE41CC2671587E0711CF0C3EAACB" box="[360,380,1207,1228]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="404">T</collectionCode>
|
||||
1 nearly absent, restricted to the lateral margins of tergum; apical fasciae on
|
||||
<collectionCode id="ED54AE41CC2671587DAE11AE0F90AAEC" box="[705,722,1238,1259]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="404">T</collectionCode>
|
||||
2
|
||||
<emphasis id="B931EA96CC2671587DB111A00FA8AAED" box="[734,746,1240,1258]" italics="true" pageId="50" pageNumber="404">–</emphasis>
|
||||
<collectionCode id="ED54AE41CC2671587D8511AD0FBEAAEC" box="[746,764,1237,1259]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="404">T</collectionCode>
|
||||
5 reduced medially or slightly interrupted. Female: mandible angled (
|
||||
<figureCitation id="137E2A01CC2671587E17106B0C8AAB2F" box="[376,456,1299,1321]" captionStart="Figures 81 and 82" captionStartId="44.[162,252,779,798]" captionTargetBox="[168,1430,198,746]" captionTargetId="figure-571@44.[168,1430,198,746]" captionTargetPageId="44" captionText="Figures 81 and 82. Female, frontal view of head (outer surface of mandible indicated by black arrows): 81, Coelioxys funeraria; 82, Coelioxys conoidea. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807049" httpUri="https://zenodo.org/record/14807049/files/figure.png" pageId="50" pageNumber="404">Fig. 81</figureCitation>
|
||||
); S6 apex elongate, longer than broad, notches acute, narrow (
|
||||
<figureCitation id="137E2A01CC2671587D3310490FECAB40" box="[604,686,1329,1351]" captionStart="Figures 26–29" captionStartId="27.[148,238,1449,1468]" captionTargetBox="[157,1420,199,1417]" captionTargetId="figure-252@27.[156,1420,198,1416]" captionTargetPageId="27" captionText="Figures 26–29. 26, female, Coelioxys porterae: dorsal view of T6 and S6. 27, male, Coelioxys sayi: dorsal view of T2 (fovea indicated by red arrow). 28, female, Coelioxys quadridentata, dorsal view of T6 and S6. 29, male, Coelioxys sodalis: dorsal view of T2 (punctate area indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807017" httpUri="https://zenodo.org/record/14807017/files/figure.png" pageId="50" pageNumber="404">Fig. 26</figureCitation>
|
||||
). Male: postgradular area of
|
||||
<collectionCode id="ED54AE41CC2671587EC710280CFBAB62" box="[424,441,1360,1381]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="404">T</collectionCode>
|
||||
2 with sublateral, elliptical, slender fovea on each side (
|
||||
<figureCitation id="137E2A01CC2671587E8D10160F73AB84" box="[482,561,1390,1412]" captionStart="Figures 87–90" captionStartId="46.[162,252,1545,1564]" captionTargetBox="[177,1417,197,1511]" captionTargetId="figure-170@46.[167,1429,197,1511]" captionTargetPageId="46" captionText="Figures 87–90. 87, 88, Coelioxys rufescens: 87, female, ventral view of S6 (S6 notch indicated by black arrow); 88, male, dorsal view of T2 (fovea indicated by black arrow). 89, 90, Coelioxys conoidea: 89, female, dorsal view of T6 and S6; 90, male, dorsal view of T2 (fovea indicated by black arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807053" httpUri="https://zenodo.org/record/14807053/files/figure.png" pageId="50" pageNumber="404">Fig. 88</figureCitation>
|
||||
) (
|
||||
<bibRefCitation id="EFD44B75CC2671587D2410160F9AAB83" author="Baker JR" box="[587,728,1390,1412]" pageId="50" pageNumber="404" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker, 1975</bibRefCitation>
|
||||
: fig. 25B) and gonocoxa subapically not compressed (
|
||||
<figureCitation id="137E2A01CC2671587FC510D30C48ABC6" box="[170,266,1451,1473]" captionStart="Figure 94" captionStartId="47.[148,227,1728,1747]" captionText="Figure 94. A–F, female. A–C, dorsal view of hemitergite 7 (apodemal region indicated by black arrow, lateral process indicated by red arrow): A, Coelioxys ducalis; B, Coelioxys alatiformis; C, Coelioxys caudata. D–F, lateral view of hemitergite 8 (postero expansion indicated by black arrow): D, Coelioxys torrida; E, Coelioxys nasuta; F, Coelioxys porterae (apex of dorsal arm of furcula indicated by red arrow). G–L, male: G, dorsal view of T7 of Coelioxys bisoncornua; H, dorsal view of S7 of Coelioxys apacheorum; I, dorsal view of S8 of Coelioxys alternata (tip producing on apex indicated by black arrow). J–L, genitalia: J, Coelioxys abdominalis; K, Coelioxys apicata; L, Coelioxys sodalis. Scale bars: 1 mm." pageId="50" pageNumber="404">Fig. 94L</figureCitation>
|
||||
) (
|
||||
<bibRefCitation id="EFD44B75CC2671587E7010D30CEAABC6" author="Baker JR" box="[287,424,1451,1473]" pageId="50" pageNumber="404" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker, 1975</bibRefCitation>
|
||||
: fig. 19F).
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2671587FCD10910CC2A916" blockId="50.[162,777,1512,1810]" pageId="50" pageNumber="404">
|
||||
<emphasis id="B931EA96CC2671587FCD10910C62ABF9" box="[162,288,1513,1534]" italics="true" pageId="50" pageNumber="404">Comments:</emphasis>
|
||||
This subgenus had previously been regarded as a synonym of
|
||||
<emphasis id="B931EA96CC2671587D6D137F0F56A81B" box="[514,532,1543,1564]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587D43137F0F80A81A" box="[556,706,1543,1565]" italics="true" pageId="50" pageNumber="404">
|
||||
<taxonomicName id="4C454D07CC2671587D43137F0FD7A81B" box="[556,661,1543,1564]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
|
||||
<taxonomicNameLabel id="A20257EDCC2671587DF113700F80A81A" box="[670,706,1544,1565]" pageId="50" pageNumber="404" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
) (see comments on that subgenus above), whereas
|
||||
<emphasis id="B931EA96CC2671587FCD133D0DF6A85D" box="[162,180,1605,1626]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587FA4133C0C3AA85E" box="[203,376,1604,1625]" italics="true" pageId="50" pageNumber="404">Schizocoelioxys</emphasis>
|
||||
) was synonymized under
|
||||
<emphasis id="B931EA96CC2671587FCD131B0DF6A87F" box="[162,180,1635,1656]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587FA4131B0C2CA87F" box="[203,366,1635,1656]" italics="true" pageId="50" pageNumber="404">Boreocoelioxys</emphasis>
|
||||
) by
|
||||
<bibRefCitation id="EFD44B75CC2671587EB5131B0FF7A87F" author="Michener CD" box="[474,693,1635,1657]" pageId="50" pageNumber="404" refId="ref43240" refString="Michener CD. 2007. The bees of the world, 2 nd edn. Baltimore: The John Hopkins University Press." type="book" year="2007">Michener (2007)</bibRefCitation>
|
||||
. The subgenus
|
||||
<emphasis id="B931EA96CC2671587E4913FA0C7AA890" box="[294,312,1666,1687]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587E3F13F90CA8A891" box="[336,490,1665,1686]" italics="true" pageId="50" pageNumber="404">Paracoelioxys</emphasis>
|
||||
) is resurrected here based on the phylogenetic results (see Results and Discussion). Whereas
|
||||
<emphasis id="B931EA96CC2671587ED713C70C88A8D3" box="[440,458,1727,1748]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<taxonomicName id="4C454D07CC2671587EB513C60F07A8D4" authorityName="Forster" authorityYear="1853" box="[474,581,1726,1748]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="alata" subGenus="Paracoelioxys">
|
||||
(
|
||||
<emphasis id="B931EA96CC2671587E8D13C70CB0A8D3" box="[482,498,1727,1748]" italics="true" pageId="50" pageNumber="404">P</emphasis>
|
||||
.)
|
||||
<emphasis id="B931EA96CC2671587D6513C60F07A8D4" box="[522,581,1726,1747]" italics="true" pageId="50" pageNumber="404">alata</emphasis>
|
||||
</taxonomicName>
|
||||
is Palaearctic,
|
||||
<taxonomicName id="4C454D07CC2671587FCD13A50C6DA8F5" authorityName="Smith" authorityYear="1854" box="[162,303,1757,1778]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="funeraria">
|
||||
<emphasis id="B931EA96CC2671587FCD13A50DF6A8F5" box="[162,180,1757,1778]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587FAC13A50C6DA8F5" box="[195,303,1757,1778]" italics="true" pageId="50" pageNumber="404">funeraria</emphasis>
|
||||
</taxonomicName>
|
||||
is restricted to North America (
|
||||
<bibRefCitation id="EFD44B75CC2671587DD613A50C36A916" author="Ascher JS & Pickering J" pageId="50" pageNumber="404" refId="ref42522" refString="Ascher JS, Pickering J. 2015. Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available via http://www.discoverlife.org/mp/20q?guide=Apoidea_species [Accessed 10 March 2015]" type="url" year="2015">Ascher & Pickering, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2671587FCD1241086AAF50" blockId="50.[162,777,1849,1901]" lastBlockId="50.[825,1439,199,343]" pageId="50" pageNumber="404">
|
||||
<emphasis id="B931EA96CC2671587FCD12410C23A949" box="[162,353,1849,1870]" italics="true" pageId="50" pageNumber="404">List of species:</emphasis>
|
||||
<taxonomicName id="4C454D07CC2671587E0012410C8EA949" authorityName="Forster" authorityYear="1853" box="[367,460,1849,1870]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="alata">
|
||||
<emphasis id="B931EA96CC2671587E0012410CC3A949" box="[367,385,1849,1870]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587EFE12410C8EA949" box="[401,460,1849,1870]" italics="true" pageId="50" pageNumber="404">alata</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C454D07CC2671587D4E12410FEDA949" authorityName="Smith" authorityYear="1854" box="[545,687,1849,1870]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="funeraria">
|
||||
<emphasis id="B931EA96CC2671587D4E12410F71A949" box="[545,563,1849,1870]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2671587D2C12410FEDA949" box="[579,687,1849,1870]" italics="true" pageId="50" pageNumber="404">funeraria</emphasis>
|
||||
</taxonomicName>
|
||||
. Some other Palaearctic species,
|
||||
<taxonomicName id="4C454D07CC2671587EB4122F0E30AEDB" authority="Nagase, 2003" authorityName="Nagase" authorityYear="2003" box="[475,882,199,1901]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="hosoba">
|
||||
<emphasis id="B931EA96CC2671587EB4122F0FE2A96B" box="[475,672,1879,1900]" italics="true" pageId="50" pageNumber="404">Coelioxys hosoba</emphasis>
|
||||
Nagase, 2003
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2671587CE215BF0858AEDB" authority="Nagase, 2003" authorityName="Nagase" authorityYear="2003" box="[909,1306,199,220]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="hiroba">
|
||||
<emphasis id="B931EA96CC2671587CE215BF091AAEDB" box="[909,1112,199,220]" italics="true" pageId="50" pageNumber="404">Coelioxys hiroba</emphasis>
|
||||
Nagase, 2003
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2671587A5915BF0919AEFC" authority="(Kirby, 1802)" baseAuthorityName="Kirby" baseAuthorityYear="1802" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="inermis">
|
||||
<emphasis id="B931EA96CC2671587A5915BF0ECDAEFC" italics="true" pageId="50" pageNumber="404">Coelioxys inermis</emphasis>
|
||||
(Kirby, 1802)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2671587B12159D0EB2AF1E" authority="Nylander, 1848" authorityName="Nylander" authorityYear="1848" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="mandibularis">
|
||||
<emphasis id="B931EA96CC2671587B12159D08DDAEFD" box="[1149,1439,229,250]" italics="true" pageId="50" pageNumber="404">Coelioxys mandibularis</emphasis>
|
||||
Nylander, 1848
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C454D07CC2671587B56147C0E3BAF3F" authority="(Lepeletier, 1841)" baseAuthorityName="Lepeletier" baseAuthorityYear="1841" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="elongata">
|
||||
<emphasis id="B931EA96CC2671587B56147C0852AF1E" box="[1081,1296,260,281]" italics="true" pageId="50" pageNumber="404">Coelioxys elongata</emphasis>
|
||||
(Lepeletier, 1841)
|
||||
</taxonomicName>
|
||||
, might also belong to this subgenus as they share the modified mandible in the female.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
155
data/03/EC/87/03EC8792CC2671597C39134D0C1FADBE.xml
Normal file
155
data/03/EC/87/03EC8792CC2671597C39134D0C1FADBE.xml
Normal file
|
|
@ -0,0 +1,155 @@
|
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<mods:namePart id="D52D0AADE8520A55896E3426791409BB">Packer, Laurence</mods:namePart>
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<subSubSection id="C35F650FCC2671587C39134D08C0A84C" box="[854,1410,1585,1612]" pageId="50" pageNumber="404" type="nomenclature">
|
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<paragraph id="8BFA3684CC2671587C39134D08C0A84C" blockId="50.[854,1410,1585,1612]" box="[854,1410,1585,1612]" pageId="50" pageNumber="404">
|
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<heading id="D0B281E8CC2671587C39134D08C0A84C" box="[854,1410,1585,1612]" centered="true" fontSize="9" level="2" pageId="50" pageNumber="404" reason="2">
|
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<taxonomicName id="4C454D07CC2671587C39134D08C0A84C" authority="MITCHELL, 1973" authorityName="Mitchell" authorityYear="1973" box="[854,1410,1585,1612]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="390" phylum="Arthropoda" rank="subGenus" subGenus="Rhinocoelioxys">
|
||||
<emphasis id="B931EA96CC2671587C39134D09F6A84C" box="[854,1204,1585,1611]" italics="true" pageId="50" pageNumber="404">
|
||||
<smallCapsWord id="8D1CA058CC2671587C39134D0E96A84E" baselines="1605,1605" box="[854,980,1589,1610]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Coelioxys" pageId="50" pageNumber="404">COELIOXYS</smallCapsWord>
|
||||
(
|
||||
<smallCapsWord id="8D1CA058CC2671587C88134C09E8A84E" baselines="1605,1605" box="[999,1194,1588,1611]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Rhinocoelioxys" pageId="50" pageNumber="404">RHINOCOELIOXYS</smallCapsWord>
|
||||
)
|
||||
</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC2671587BD1134B08C0A84C" author="Mitchell TB" box="[1214,1410,1585,1612]" pageId="50" pageNumber="404" refId="ref43263" refString="Mitchell TB. 1973. A subgeneric revision of the genus Coelioxys of the Western Hemisphere (Hymenoptera: Megachilidae). Raleigh: Department of Entomology, North Carolina State University." type="book" year="1973">
|
||||
<smallCapsWord id="8D1CA058CC2671587BD1134B0870A84E" baselines="1605,1605" box="[1214,1330,1587,1611]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Mitchell" pageId="50" pageNumber="404">MITCHELL</smallCapsWord>
|
||||
, 1973
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F650FCC2671597C5713FA0C1FADBE" lastPageId="51" lastPageNumber="405" pageId="50" pageNumber="404" type="materials_examined">
|
||||
<paragraph id="8BFA3684CC2671587C5713FA0819A891" blockId="50.[824,1371,1665,1687]" box="[824,1371,1665,1687]" pageId="50" pageNumber="404">
|
||||
<emphasis id="B931EA96CC2671587C5713FA0E89A890" box="[824,971,1666,1687]" italics="true" pageId="50" pageNumber="404">
|
||||
<typeStatus id="54FE8826CC2671587C5713FA0E2CA890" box="[824,878,1666,1687]" pageId="50" pageNumber="404">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C454D07CC2671587CB513F9081AA891" authority="Cresson, 1878" authorityName="Cresson" authorityYear="1878" box="[986,1368,1665,1687]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="404" phylum="Arthropoda" rank="species" species="zapoteca">
|
||||
<emphasis id="B931EA96CC2671587CB513F909E9A890" box="[986,1195,1665,1687]" italics="true" pageId="50" pageNumber="404">Coelioxys zapoteca</emphasis>
|
||||
Cresson, 1878
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2671597C5713C70CDDAFD6" blockId="50.[824,1440,1726,1901]" lastBlockId="51.[148,763,198,465]" lastPageId="51" lastPageNumber="405" pageId="50" pageNumber="404">
|
||||
<emphasis id="B931EA96CC2671587C5713C70EF2A8D3" box="[824,944,1727,1748]" italics="true" pageId="50" pageNumber="404">Diagnosis:</emphasis>
|
||||
The following characteristics combined (four for either sex) separate species of
|
||||
<emphasis id="B931EA96CC2671587C5713840E08A916" box="[824,842,1788,1809]" italics="true" pageId="50" pageNumber="404">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2671587C0D1383094BA917" box="[866,1033,1787,1808]" italics="true" pageId="50" pageNumber="404">Rhinocoelioxys</emphasis>
|
||||
) from all the other subgenera. Mesoscutellum with a distinct longitudinal median ridge (Fig. 51); posterior margin of mesoscutellum obtusely produced medially into a blunt edge (Fig. 51); axilla long, free apical portion longer or subequal to basal portion (Fig. 51); ocular hairs long (~0.12 mm) (Fig. 5). Female: S6 lateral notches obsolete, a weak impression, not interrupting outline in dorsal view (
|
||||
<figureCitation id="137E2A01CC2771597E2214390CE2AF50" box="[333,416,321,343]" captionStart="Figures 65–68" captionStartId="39.[148,238,1792,1811]" captionText="Figures 65–68. 65, female, Coelioxys agilis: dorsal view of T6 and S6 (S6 notch indicated by black arrow). 66, male, Coelioxys zapoteca. Dorsal view of T2–T3 (punctate areas indicated by red arrows). 67, female, Coelioxys barbata: dorsal view of T6 and S6 (S6 notch indicated by black arrow). 68, male, Coelioxys clypearis: dorsal view of T2–T3 (punctate areas indicated by black arrows). Scale bars: 1.5 mm." pageId="51" pageNumber="405">Fig. 65</figureCitation>
|
||||
). Male: postgradular areas of
|
||||
<collectionCode id="ED54AE41CC2771597FFB14180DE7AF72" box="[148,165,352,373]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
|
||||
2 and
|
||||
<collectionCode id="ED54AE41CC2771597F9914180C4AAF72" box="[246,264,352,373]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
|
||||
3 with a transverse, punctate area either formed by punctures that may be fused or by few smaller punctures basally and larger punctures apicolaterally (
|
||||
<figureCitation id="137E2A01CC2771597E5014C30CD1AFD6" box="[319,403,443,465]" captionStart="Figures 65–68" captionStartId="39.[148,238,1792,1811]" captionText="Figures 65–68. 65, female, Coelioxys agilis: dorsal view of T6 and S6 (S6 notch indicated by black arrow). 66, male, Coelioxys zapoteca. Dorsal view of T2–T3 (punctate areas indicated by red arrows). 67, female, Coelioxys barbata: dorsal view of T6 and S6 (S6 notch indicated by black arrow). 68, male, Coelioxys clypearis: dorsal view of T2–T3 (punctate areas indicated by black arrows). Scale bars: 1.5 mm." pageId="51" pageNumber="405">Fig. 66</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2771597FFB14810C71AD59" blockId="51.[148,763,504,862]" pageId="51" pageNumber="405">
|
||||
<emphasis id="B931EA96CC2771597FFB14810C50AC09" box="[148,274,505,526]" italics="true" pageId="51" pageNumber="405">Comments:</emphasis>
|
||||
The phylogenetic results indicate that
|
||||
<emphasis id="B931EA96CC2771597FFB176F0DE4AC2B" box="[148,166,535,556]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2771597FD1176F0C27AC2B" box="[190,357,535,556]" italics="true" pageId="51" pageNumber="405">Rhinocoelioxys</emphasis>
|
||||
) as previously defined is polyphyletic. The
|
||||
<typeStatus id="54FE8826CC2771597E0A174E0CD4AC4C" box="[357,406,566,587]" pageId="51" pageNumber="405">type</typeStatus>
|
||||
species
|
||||
<taxonomicName id="4C454D07CC2771597E95174E0F3EAC4C" authorityName="Cresson" authorityYear="1878" box="[506,636,566,587]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="zapoteca">
|
||||
<emphasis id="B931EA96CC2771597E95174E0F4EAC4C" box="[506,524,566,587]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2771597D73174E0F3EAC4C" box="[540,636,566,587]" italics="true" pageId="51" pageNumber="405">zapoteca</emphasis>
|
||||
</taxonomicName>
|
||||
as well as
|
||||
<taxonomicName id="4C454D07CC2771597FFB172C0DB7AC6E" authorityName="Smith" authorityYear="1879" box="[148,245,596,617]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="agilis">
|
||||
<emphasis id="B931EA96CC2771597FFB172C0DE4AC6E" box="[148,166,596,617]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2771597FD9172C0DB7AC6E" box="[182,245,596,617]" italics="true" pageId="51" pageNumber="405">agilis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C454D07CC2771597E51172C0C69AC8F" authority="Rocha-Filho & Packer, 2015" authorityName="Rocha-Filho & Packer" authorityYear="2015" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="platygnatha">
|
||||
<emphasis id="B931EA96CC2771597E51172C0F02AC6E" box="[318,576,596,617]" italics="true" pageId="51" pageNumber="405">Coelioxys platygnatha</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC2771597D20172C0C69AC8F" author="Rocha-Filho LC & Packer L" pageId="51" pageNumber="405" pagination="151 - 203" refId="ref43662" refString="Rocha-Filho LC, Packer L. 2015. Revision of the Neotropical subgenera Coelioxys (Platycoelioxys) Mitchell and C. (Rhinocoelioxys) Mitchell (Hymenoptera; Megachilidae) with the description of one new species. Zootaxa 3941: 151-203." type="journal article" year="2015">Rocha-Filho & Packer, 2015</bibRefCitation>
|
||||
</taxonomicName>
|
||||
are in a different clade from the other four species that were included by
|
||||
<bibRefCitation id="EFD44B75CC2771597D2017E90C05ACC2" author="Rocha-Filho LC & Packer L" pageId="51" pageNumber="405" pagination="151 - 203" refId="ref43662" refString="Rocha-Filho LC, Packer L. 2015. Revision of the Neotropical subgenera Coelioxys (Platycoelioxys) Mitchell and C. (Rhinocoelioxys) Mitchell (Hymenoptera; Megachilidae) with the description of one new species. Zootaxa 3941: 151-203." type="journal article" year="2015">Rocha-Filho & Packer (2015)</bibRefCitation>
|
||||
, which are now transferred to
|
||||
<emphasis id="B931EA96CC2771597FFB17B70DE4ACE3" box="[148,166,719,740]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2771597FD117B60C22ACE4" box="[190,352,718,739]" italics="true" pageId="51" pageNumber="405">Austrocleptria</emphasis>
|
||||
) subgen. nov. (see below). The three species are Neotropical.
|
||||
<bibRefCitation id="EFD44B75CC2771597E9517950D99AD26" author="Rocha-Filho LC & Packer L" pageId="51" pageNumber="405" pagination="151 - 203" refId="ref43662" refString="Rocha-Filho LC, Packer L. 2015. Revision of the Neotropical subgenera Coelioxys (Platycoelioxys) Mitchell and C. (Rhinocoelioxys) Mitchell (Hymenoptera; Megachilidae) with the description of one new species. Zootaxa 3941: 151-203." type="journal article" year="2015">Rocha-Filho & Packer (2015)</bibRefCitation>
|
||||
revised and keyed the species of this subgenus [as well as those of
|
||||
<emphasis id="B931EA96CC2771597D4816520F7BAD38" box="[551,569,810,831]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2771597D3E16520FB1AD38" box="[593,755,810,831]" italics="true" pageId="51" pageNumber="405">Austrocleptria</emphasis>
|
||||
) subgen. nov.].
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2771597FFB16FE0C1FADBE" blockId="51.[148,763,901,954]" pageId="51" pageNumber="405">
|
||||
<emphasis id="B931EA96CC2771597FFB16FE0F55AD9D" box="[148,535,901,923]" italics="true" pageId="51" pageNumber="405">
|
||||
List of species:
|
||||
<taxonomicName id="4C454D07CC2771597E3216FD0F55AD9D" authorityName="Smith" authorityYear="1879" box="[349,535,901,922]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="agilis">Coelioxys agilis</taxonomicName>
|
||||
</emphasis>
|
||||
*,
|
||||
<taxonomicName id="4C454D07CC2771597D5316FE0FAAAD9D" authorityName="Rocha-Filho & Packer" authorityYear="2015" box="[572,744,901,923]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="platygnatha">
|
||||
<emphasis id="B931EA96CC2771597D5316FE0F0CAD9C" box="[572,590,902,923]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2771597D3116FD0FAAAD9D" box="[606,744,901,922]" italics="true" pageId="51" pageNumber="405">platygnatha</emphasis>
|
||||
</taxonomicName>
|
||||
*, and
|
||||
<taxonomicName id="4C454D07CC2771597FA816DC0C0BADBD" authorityName="Cresson" authorityYear="1878" box="[199,329,932,954]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="zapoteca">
|
||||
<emphasis id="B931EA96CC2771597FA816DC0D9BADBE" box="[199,217,932,953]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2771597F8616DD0C0BADBD" box="[233,329,933,954]" italics="true" pageId="51" pageNumber="405">zapoteca</emphasis>
|
||||
</taxonomicName>
|
||||
*.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
195
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195
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<mods:title id="3163FA1A701357C6ADAA4A45D1FD1ACB">Phylogeny of the cleptoparasitic Megachilini genera Coelioxys and Radoszkowskiana, with the description of six new subgenera in Coelioxys (Hymenoptera: Megachilidae)</mods:title>
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<taxonomicName id="4C454D07CC2771597FD716890F95AA01" authority="MITCHELL, 1973" authorityName="Mitchell" authorityYear="1973" box="[184,727,1004,1031]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="384" phylum="Arthropoda" rank="subGenus" subGenus="Synocoelioxys">
|
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<emphasis id="B931EA96CC2771597FD716890F4BAA00" box="[184,521,1005,1031]" italics="true" pageId="51" pageNumber="405">
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|
||||
(
|
||||
<smallCapsWord id="8D1CA058CC2771597E2616960F42AA03" baselines="1024,1024" box="[329,512,1006,1030]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Synocoelioxys" pageId="51" pageNumber="405">SYNOCOELIOXYS</smallCapsWord>
|
||||
)
|
||||
</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC2771597D7C16960F95AA01" author="Mitchell TB" box="[531,727,1004,1031]" pageId="51" pageNumber="405" refId="ref43263" refString="Mitchell TB. 1973. A subgeneric revision of the genus Coelioxys of the Western Hemisphere (Hymenoptera: Megachilidae). Raleigh: Department of Entomology, North Carolina State University." type="book" year="1973">
|
||||
<smallCapsWord id="8D1CA058CC2771597D7C16960FC5AA02" baselines="1024,1025" box="[531,647,1006,1030]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Mitchell" pageId="51" pageNumber="405">MITCHELL</smallCapsWord>
|
||||
, 1973
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F650FCC2771597FFB1145081DAC2B" pageId="51" pageNumber="405" type="materials_examined">
|
||||
<paragraph id="8BFA3684CC2771597FFB11450FE0AA55" blockId="51.[148,674,1085,1106]" box="[148,674,1085,1106]" pageId="51" pageNumber="405">
|
||||
<emphasis id="B931EA96CC2771597FFB11450C65AA55" box="[148,295,1085,1106]" italics="true" pageId="51" pageNumber="405">
|
||||
<typeStatus id="54FE8826CC2771597FFB11450D88AA55" box="[148,202,1085,1106]" pageId="51" pageNumber="405">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C454D07CC2771597E5911450FDCAA55" authority="Cresson, 1872" authorityName="Cresson" authorityYear="1872" box="[310,670,1085,1106]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="texana">
|
||||
<emphasis id="B931EA96CC2771597E5911450CB0AA55" box="[310,498,1085,1106]" italics="true" pageId="51" pageNumber="405">Coelioxys texana</emphasis>
|
||||
Cresson, 1872
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2771597FFB11020C09A949" blockId="51.[148,764,1146,1871]" pageId="51" pageNumber="405">
|
||||
<emphasis id="B931EA96CC2771597FFB11020C49AA88" box="[148,267,1146,1167]" italics="true" pageId="51" pageNumber="405">Diagnosis:</emphasis>
|
||||
This is the only
|
||||
<taxonomicName id="4C454D07CC2771597D6211020F3BAA88" box="[525,633,1146,1167]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B931EA96CC2771597D6211020F3BAA88" box="[525,633,1146,1167]" italics="true" pageId="51" pageNumber="405">Coelioxys</emphasis>
|
||||
</taxonomicName>
|
||||
subgenus and perhaps the only members of the
|
||||
<taxonomicName id="4C454D07CC2771597D1D11E00FB9AAA9" box="[626,763,1176,1198]" class="Insecta" family="Megachilidae" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="tribe" tribe="Megachilini">Megachilini</taxonomicName>
|
||||
in which the male S7 is bilobed (
|
||||
<figureCitation id="137E2A01CC2771597D5A11CF0FDFAACB" box="[565,669,1207,1229]" captionStart="Figure 94" captionStartId="47.[148,227,1728,1747]" captionText="Figure 94. A–F, female. A–C, dorsal view of hemitergite 7 (apodemal region indicated by black arrow, lateral process indicated by red arrow): A, Coelioxys ducalis; B, Coelioxys alatiformis; C, Coelioxys caudata. D–F, lateral view of hemitergite 8 (postero expansion indicated by black arrow): D, Coelioxys torrida; E, Coelioxys nasuta; F, Coelioxys porterae (apex of dorsal arm of furcula indicated by red arrow). G–L, male: G, dorsal view of T7 of Coelioxys bisoncornua; H, dorsal view of S7 of Coelioxys apacheorum; I, dorsal view of S8 of Coelioxys alternata (tip producing on apex indicated by black arrow). J–L, genitalia: J, Coelioxys abdominalis; K, Coelioxys apicata; L, Coelioxys sodalis. Scale bars: 1 mm." pageId="51" pageNumber="405">Fig. 94H</figureCitation>
|
||||
) rather than divided into two small sclerites (
|
||||
<bibRefCitation id="EFD44B75CC2771597D0C11AD0FBAAAEC" author="Baker JR" box="[611,760,1237,1259]" pageId="51" pageNumber="405" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker, 1975</bibRefCitation>
|
||||
: fig. 26D). Some characteristics such as the scale-like pubescence in parts of the body (
|
||||
<figureCitation id="137E2A01CC2771597D08106A0FFEAB2F" box="[615,700,1298,1320]" captionStart="Figures 32–34" captionStartId="28.[162,252,1329,1348]" captionTargetBox="[170,1425,933,1296]" captionTargetId="figure-159@28.[170,1433,933,1296]" captionTargetPageId="28" captionText="Figures 32–34. 32, 33, female. 32, Coelioxys texana: dorsal view of mid ocellus. 33, Coelioxys alternata: dorsal view of T6 and S6. 34, male, Coelioxys alternata: dorsal view of T6 (dorsal process indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807023" httpUri="https://zenodo.org/record/14807023/files/figure.png" pageId="51" pageNumber="405">Fig. 32</figureCitation>
|
||||
), the inner ramus of pretarsal claws in females acute (
|
||||
<figureCitation id="137E2A01CC2771597FF210280DB3AB61" box="[157,241,1360,1382]" captionStart="Figures 35–37" captionStartId="28.[162,252,1821,1840]" captionTargetBox="[170,1433,1470,1789]" captionTargetId="figure-106@28.[169,1432,1469,1788]" captionTargetPageId="28" captionText="Figures 35–37. 35, 36, female. 35, Coelioxys albolineata: dorsal view of mesoscutellum (for axilla length). 36, Coelioxys froggatti: lateral view of pretarsal claws (inner rami indicated by black arrows). 37, male, Coelioxys reginae: lateral view of head (for genal excavation). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807025" httpUri="https://zenodo.org/record/14807025/files/figure.png" pageId="51" pageNumber="405">Fig. 36</figureCitation>
|
||||
), and apicodorsal processes of
|
||||
<collectionCode id="ED54AE41CC2771597D0510280F39AB61" box="[618,635,1360,1382]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
|
||||
<quantity id="4CBD9B61CC2771597D1410280FE9AB61" box="[635,683,1360,1382]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="51" pageNumber="405" unit="in" value="6.0">6 in</quantity>
|
||||
males forked (
|
||||
<figureCitation id="137E2A01CC2771597F9710160C08AB84" box="[248,330,1390,1412]" captionStart="Figures 32–34" captionStartId="28.[162,252,1329,1348]" captionTargetBox="[170,1425,933,1296]" captionTargetId="figure-159@28.[170,1433,933,1296]" captionTargetPageId="28" captionText="Figures 32–34. 32, 33, female. 32, Coelioxys texana: dorsal view of mid ocellus. 33, Coelioxys alternata: dorsal view of T6 and S6. 34, male, Coelioxys alternata: dorsal view of T6 (dorsal process indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807023" httpUri="https://zenodo.org/record/14807023/files/figure.png" pageId="51" pageNumber="405">Fig. 34</figureCitation>
|
||||
) are shared only with species of the Old World subgenus
|
||||
<emphasis id="B931EA96CC2771597D4310F50F7CABA5" box="[556,574,1421,1442]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2771597D3810F50FA9ABA5" box="[599,747,1421,1442]" italics="true" pageId="51" pageNumber="405">Allocoelioxys</emphasis>
|
||||
). Species belonging to
|
||||
<emphasis id="B931EA96CC2771597EB010D40CB3ABC6" box="[479,497,1452,1473]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2771597D6510D30FEAABC7" box="[522,680,1451,1472]" italics="true" pageId="51" pageNumber="405">Synocoelioxys</emphasis>
|
||||
) are characterized by the following combination of characteristics that separates them from all other subgenera. Median ocellus surrounded by a swollen and nearly impunctate area (
|
||||
<figureCitation id="137E2A01CC2771597DF6135E0FADA83C" box="[665,751,1574,1596]" captionStart="Figures 32–34" captionStartId="28.[162,252,1329,1348]" captionTargetBox="[170,1425,933,1296]" captionTargetId="figure-159@28.[170,1433,933,1296]" captionTargetPageId="28" captionText="Figures 32–34. 32, 33, female. 32, Coelioxys texana: dorsal view of mid ocellus. 33, Coelioxys alternata: dorsal view of T6 and S6. 34, male, Coelioxys alternata: dorsal view of T6 (dorsal process indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807023" httpUri="https://zenodo.org/record/14807023/files/figure.png" pageId="51" pageNumber="405">Fig. 32</figureCitation>
|
||||
). Female: lateral margins of
|
||||
<collectionCode id="ED54AE41CC2771597D6B133C0F57A85D" box="[516,533,1604,1626]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
|
||||
6 apex with long thick hairs forming an inconspicuous fringe (
|
||||
<figureCitation id="137E2A01CC2771597FF213F90DB3A891" box="[157,241,1665,1687]" captionStart="Figures 32–34" captionStartId="28.[162,252,1329,1348]" captionTargetBox="[170,1425,933,1296]" captionTargetId="figure-159@28.[170,1433,933,1296]" captionTargetPageId="28" captionText="Figures 32–34. 32, 33, female. 32, Coelioxys texana: dorsal view of mid ocellus. 33, Coelioxys alternata: dorsal view of T6 and S6. 34, male, Coelioxys alternata: dorsal view of T6 (dorsal process indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807023" httpUri="https://zenodo.org/record/14807023/files/figure.png" pageId="51" pageNumber="405">Fig. 33</figureCitation>
|
||||
); S6 rounded or elliptical, not notched and straight (
|
||||
<figureCitation id="137E2A01CC2771597E6913D80C19A8B2" box="[262,347,1696,1718]" captionStart="Figures 32–34" captionStartId="28.[162,252,1329,1348]" captionTargetBox="[170,1425,933,1296]" captionTargetId="figure-159@28.[170,1433,933,1296]" captionTargetPageId="28" captionText="Figures 32–34. 32, 33, female. 32, Coelioxys texana: dorsal view of mid ocellus. 33, Coelioxys alternata: dorsal view of T6 and S6. 34, male, Coelioxys alternata: dorsal view of T6 (dorsal process indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807023" httpUri="https://zenodo.org/record/14807023/files/figure.png" pageId="51" pageNumber="405">Fig. 33</figureCitation>
|
||||
), not flexed downwards. Male:
|
||||
<collectionCode id="ED54AE41CC2771597DB313D80FAFA8B1" box="[732,749,1696,1718]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
|
||||
6 apicodorsal processes forked, with processes flattened, pointed, and irregular, almost forming a crenulate plate (
|
||||
<figureCitation id="137E2A01CC2771597E0D13840CF4A916" box="[354,438,1788,1810]" captionStart="Figures 32–34" captionStartId="28.[162,252,1329,1348]" captionTargetBox="[170,1425,933,1296]" captionTargetId="figure-159@28.[170,1433,933,1296]" captionTargetPageId="28" captionText="Figures 32–34. 32, 33, female. 32, Coelioxys texana: dorsal view of mid ocellus. 33, Coelioxys alternata: dorsal view of T6 and S6. 34, male, Coelioxys alternata: dorsal view of T6 (dorsal process indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807023" httpUri="https://zenodo.org/record/14807023/files/figure.png" pageId="51" pageNumber="405">Fig. 34</figureCitation>
|
||||
), and
|
||||
<collectionCode id="ED54AE41CC2771597D6313840F5FA916" box="[524,541,1788,1809]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
|
||||
7 exposed, with a long median tooth-like process medially (
|
||||
<bibRefCitation id="EFD44B75CC2771597DC212620D8DA949" author="Baker JR" pageId="51" pageNumber="405" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker, 1975</bibRefCitation>
|
||||
: fig. 13A).
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2771597C4415BF0E33AF93" blockId="51.[811,1426,199,404]" pageId="51" pageNumber="405">
|
||||
<emphasis id="B931EA96CC2771597C4415BF0EEBAEDB" box="[811,937,199,220]" italics="true" pageId="51" pageNumber="405">Comments:</emphasis>
|
||||
Species of
|
||||
<emphasis id="B931EA96CC2771597B1915BF09CAAEDB" box="[1142,1160,199,220]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2771597BF015BF087BAEDB" box="[1183,1337,199,220]" italics="true" pageId="51" pageNumber="405">Synocoelioxys</emphasis>
|
||||
) are primarily Nearctic; five of them were revised and keyed by
|
||||
<bibRefCitation id="EFD44B75CC2771597CDA147C091BAF1D" author="Baker JR" box="[949,1113,260,282]" pageId="51" pageNumber="405" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker (1975)</bibRefCitation>
|
||||
; another two,
|
||||
<taxonomicName id="4C454D07CC2771597A46147C09C4AF3F" authority="Cockerell, 1923" authorityName="Cockerell" authorityYear="1923" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="toltecoides">
|
||||
<emphasis id="B931EA96CC2771597A46147C0EE2AF30" italics="true" pageId="51" pageNumber="405">Coelioxys toltecoides</emphasis>
|
||||
Cockerell, 1923
|
||||
</taxonomicName>
|
||||
from
|
||||
<collectingCountry id="F3527614CC2771597B96145A080BAF3F" box="[1273,1353,290,312]" name="Mexico" pageId="51" pageNumber="405">Mexico</collectingCountry>
|
||||
and
|
||||
<taxonomicName id="4C454D07CC2771597C4414390981AF50" authority="Cresson, 1869" authorityName="Cresson" authorityYear="1869" box="[811,1219,321,343]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="tegularis">
|
||||
<emphasis id="B931EA96CC2771597C441439094BAF51" box="[811,1033,321,342]" italics="true" pageId="51" pageNumber="405">Coelioxys tegularis</emphasis>
|
||||
Cresson, 1869
|
||||
</taxonomicName>
|
||||
from
|
||||
<collectingCountry id="F3527614CC2771597A761439081AAF50" box="[1305,1368,321,343]" name="Cuba" pageId="51" pageNumber="405">Cuba</collectingCountry>
|
||||
, are also assigned to the subgenus (
|
||||
<bibRefCitation id="EFD44B75CC2771597BCC14270E27AF93" author="Ascher JS & Pickering J" pageId="51" pageNumber="405" refId="ref42522" refString="Ascher JS, Pickering J. 2015. Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available via http://www.discoverlife.org/mp/20q?guide=Apoidea_species [Accessed 10 March 2015]" type="url" year="2015">Ascher & Pickering, 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2771597C4414C4081DAC2B" blockId="51.[811,1426,443,557]" pageId="51" pageNumber="405">
|
||||
<emphasis id="B931EA96CC2771597C4414C40EB6AFD6" box="[811,1012,443,465]" italics="true" pageId="51" pageNumber="405">List of species:</emphasis>
|
||||
<taxonomicName id="4C454D07CC2771597B6C14C308CCAFD7" authority="Say, 1837" authorityName="Say" authorityYear="1837" box="[1027,1422,443,464]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="alternata">
|
||||
<emphasis id="B931EA96CC2771597B6C14C309ADAFD7" box="[1027,1263,443,464]" italics="true" pageId="51" pageNumber="405">Coelioxys alternata</emphasis>
|
||||
Say, 1837
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2771597C4414A2084CAFE8" authority="Cockerell, 1900" authorityName="Cockerell" authorityYear="1900" box="[811,1294,474,496]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="apacheorum">
|
||||
<emphasis id="B931EA96CC2771597C4414A20974AFE8" box="[811,1078,474,495]" italics="true" pageId="51" pageNumber="405">Coelioxys apacheorum</emphasis>
|
||||
Cockerell, 1900
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2771597A4614A2097CAC09" authority="Cockerell, 1912" authorityName="Cockerell" authorityYear="1912" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="erysimi">
|
||||
<emphasis id="B931EA96CC2771597A4614A20E3CAC09" italics="true" pageId="51" pageNumber="405">Coelioxys erysimi</emphasis>
|
||||
Cockerell, 1912
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2771597B2214800E26AC2B" authority="Crawford, 1914" authorityName="Crawford" authorityYear="1914" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="hunteri">
|
||||
<emphasis id="B931EA96CC2771597B2214800857AC0A" box="[1101,1301,504,525]" italics="true" pageId="51" pageNumber="405">Coelioxys hunteri</emphasis>
|
||||
Crawford, 1914
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2771597C1E176F0EB4AC2B" authorityName="Cresson" authorityYear="1869" box="[881,1014,535,556]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="tegularis">
|
||||
<emphasis id="B931EA96CC2771597C1E176F0EC1AC2B" box="[881,899,535,556]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2771597CFD176F0EB4AC2B" box="[914,1014,535,556]" italics="true" pageId="51" pageNumber="405">tegularis</emphasis>
|
||||
</taxonomicName>
|
||||
*,
|
||||
<taxonomicName id="4C454D07CC2771597B7D176F093DAC2A" authorityName="Cresson" authorityYear="1872" box="[1042,1151,535,557]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="texana">
|
||||
<emphasis id="B931EA96CC2771597B7D176F0966AC2B" box="[1042,1060,535,556]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2771597B5B1760093DAC2A" box="[1076,1151,536,557]" italics="true" pageId="51" pageNumber="405">texana</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C454D07CC2771597BAE176F081AAC2B" authorityName="Cockerell" authorityYear="1923" box="[1217,1368,535,556]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="toltecoides">
|
||||
<emphasis id="B931EA96CC2771597BAE176F0991AC2B" box="[1217,1235,535,556]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2771597B8C176F081AAC2B" box="[1251,1368,535,556]" italics="true" pageId="51" pageNumber="405">toltecoides</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
215
data/03/EC/87/03EC8792CC27715E7C05171B0F64AF30.xml
Normal file
215
data/03/EC/87/03EC8792CC27715E7C05171B0F64AF30.xml
Normal file
|
|
@ -0,0 +1,215 @@
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(
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|
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)
|
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</emphasis>
|
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<bibRefCitation id="EFD44B75CC2771597BFB17190811AC7E" author="Pasteels JJ" box="[1172,1363,607,634]" pageId="51" pageNumber="405" pagination="161 - 197" refId="ref43482" refString="Pasteels JJ. 1977. Les Megachilini parasites (Coelioxys s. l.) d'Afrique noire. Revue de Zoologie Africaine 91: 161-197." type="journal article" year="1977">
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<smallCapsWord id="8D1CA058CC2771597BFB17190841AC7F" baselines="627,628" box="[1172,1283,609,633]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Pasteels" pageId="51" pageNumber="405">PASTEELS</smallCapsWord>
|
||||
, 1977
|
||||
</bibRefCitation>
|
||||
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|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F650FCC27715E7C4417C80F64AF30" lastPageId="52" lastPageNumber="406" pageId="51" pageNumber="405" type="materials_examined">
|
||||
<paragraph id="8BFA3684CC2771597C4417C8086BACC2" blockId="51.[811,1321,687,709]" box="[811,1321,687,709]" pageId="51" pageNumber="405">
|
||||
<emphasis id="B931EA96CC2771597C4417C80EFCACC2" box="[811,958,688,709]" italics="true" pageId="51" pageNumber="405">
|
||||
<typeStatus id="54FE8826CC2771597C4417C80E23ACC2" box="[811,865,688,709]" pageId="51" pageNumber="405">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C454D07CC2771597CA217D70867ACC2" authority="Smith, 1854" authorityName="Smith" authorityYear="1854" box="[973,1317,687,709]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="torrida">
|
||||
<emphasis id="B931EA96CC2771597CA217D709CCACC3" box="[973,1166,687,708]" italics="true" pageId="51" pageNumber="405">Coelioxys torrida</emphasis>
|
||||
Smith, 1854
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2771597C4417950909ABC6" blockId="51.[811,1426,748,1473]" pageId="51" pageNumber="405">
|
||||
<emphasis id="B931EA96CC2771597C4417950EE0AD05" box="[811,930,749,770]" italics="true" pageId="51" pageNumber="405">Diagnosis:</emphasis>
|
||||
With
|
||||
<emphasis id="B931EA96CC2771597C9117950952AD05" box="[1022,1040,749,770]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2771597B47179409E9AD06" box="[1064,1195,748,769]" italics="true" pageId="51" pageNumber="405">Liothyrapis</emphasis>
|
||||
) and the species
|
||||
<emphasis id="B931EA96CC2771597C4416740E7FAD26" box="[811,829,780,801]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
.
|
||||
<taxonomicName id="4C454D07CC2771597C2216730909AD27" authorityName="Benoist" authorityYear="1955" box="[845,1099,779,800]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="madagascariensis" subGenus="Allocoelioxys">
|
||||
(
|
||||
<emphasis id="B931EA96CC2771597C3A16730E24AD27" box="[853,870,779,800]" italics="true" pageId="51" pageNumber="405">A</emphasis>
|
||||
.)
|
||||
<emphasis id="B931EA96CC2771597C1016730909AD27" box="[895,1099,779,800]" italics="true" pageId="51" pageNumber="405">madagascariensis</emphasis>
|
||||
</taxonomicName>
|
||||
, this Old World subgenus is the only one within
|
||||
<taxonomicName id="4C454D07CC2771597B7B1652093FAD38" box="[1044,1149,810,831]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B931EA96CC2771597B7B1652093FAD38" box="[1044,1149,810,831]" italics="true" pageId="51" pageNumber="405">Coelioxys</emphasis>
|
||||
</taxonomicName>
|
||||
in which the compound eyes are bare (
|
||||
<figureCitation id="137E2A01CC2771597CB416300962AD5A" box="[987,1056,840,862]" captionStart="Figures 4 and 5" captionStartId="22.[162,252,1021,1040]" captionTargetBox="[172,1435,430,992]" captionTargetId="figure-248@22.[169,1432,426,988]" captionTargetPageId="22" captionText="Figures 4 and 5. Female. Frontal view of head (for compound eye): 4, Coelioxys weinlandi; 5, Coelioxys zapoteca. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807005" httpUri="https://zenodo.org/record/14807005/files/figure.png" pageId="51" pageNumber="405">Fig. 4</figureCitation>
|
||||
). Species of
|
||||
<emphasis id="B931EA96CC2771597BDB16310984AD59" box="[1204,1222,841,862]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2771597BB11630081AAD5A" box="[1246,1368,840,861]" italics="true" pageId="51" pageNumber="405">Torridapis</emphasis>
|
||||
) can be distinguished from
|
||||
<taxonomicName id="4C454D07CC2771597B2A161F0870AD7B" authorityName="Benoist" authorityYear="1955" box="[1093,1330,871,892]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="madagascariensis">
|
||||
<emphasis id="B931EA96CC2771597B2A161F0915AD7B" box="[1093,1111,871,892]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC2771597B09161F0870AD7B" box="[1126,1330,871,892]" italics="true" pageId="51" pageNumber="405">madagascariensis</emphasis>
|
||||
</taxonomicName>
|
||||
by the lateral margins of axilla neither elevated nor carinate (
|
||||
<figureCitation id="137E2A01CC2771597CC016DC0942ADBE" box="[943,1024,932,954]" captionStart="Figures 8–13" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 8–13. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 11</figureCitation>
|
||||
) (conspicuously elevated and carinate in the latter;
|
||||
<figureCitation id="137E2A01CC2771597B5F16BA0934ADDF" box="[1072,1142,962,984]" captionStart="Figures 6 and 7" captionStartId="22.[162,252,1850,1869]" captionTargetBox="[169,1434,1127,1818]" captionTargetId="figure-275@22.[169,1433,1129,1818]" captionTargetPageId="22" captionText="Figures 6 and 7. Female. Dorsal view of mesoscutellum (for lateral margin of axilla): 6, Coelioxys madagascariensis; 7, Coelioxys verticalis. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807009" httpUri="https://zenodo.org/record/14807009/files/figure.png" pageId="51" pageNumber="405">Fig. 6</figureCitation>
|
||||
). They can be separated from those of
|
||||
<emphasis id="B931EA96CC2771597CA116990EA2ADF1" box="[974,992,993,1014]" italics="true" pageId="51" pageNumber="405">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC2771597C9716990939ADF1" box="[1016,1147,993,1014]" italics="true" pageId="51" pageNumber="405">Liothyrapis</emphasis>
|
||||
) by the following list of characteristics. Axilla long, free apical portion produced beyond the posterior margin of mesoscutellum, and curved apically (
|
||||
<figureCitation id="137E2A01CC2771597A5D114508C6AA55" box="[1330,1412,1085,1107]" captionStart="Figures 8–13" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 8–13. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 11</figureCitation>
|
||||
); mesoscutellum with a dorsal median carina at least basally (
|
||||
<figureCitation id="137E2A01CC2771597CF911020EAAAA88" box="[918,1000,1146,1168]" captionStart="Figures 8–13" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 8–13. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 11</figureCitation>
|
||||
). Female:
|
||||
<collectionCode id="ED54AE41CC2771597B1A110209C4AA97" box="[1141,1158,1146,1168]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
|
||||
6 lacking long, erect, thick hairs, with apex ending in an inconspicuous, short, spine-like process (
|
||||
<figureCitation id="137E2A01CC2771597BED11CF0996AACB" box="[1154,1236,1207,1229]" captionStart="Figures 8–13" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 8–13. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 12</figureCitation>
|
||||
);
|
||||
<collectionCode id="ED54AE41CC2771597B9411CF084EAACA" box="[1275,1292,1207,1229]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
|
||||
6 and S6 gradually tapering, greatly elongate, S6 folded, subacute apically, its lateral margins with an inconspicuous fringe composed of short brownish hairs (
|
||||
<figureCitation id="137E2A01CC2771597CE810490E98AB41" box="[903,986,1329,1351]" captionStart="Figures 8–13" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 8–13. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 12</figureCitation>
|
||||
). Male: hypostomal area lacking concavity;
|
||||
<collectionCode id="ED54AE41CC2771597CCA10280EF4AB61" box="[933,950,1360,1382]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
|
||||
6 basally lacking, or with short, flattened lateral expanded areas, never with long, acute teeth (
|
||||
<figureCitation id="137E2A01CC2771597C5C10F50EC4ABA5" box="[819,902,1421,1443]" captionStart="Figures 8–13" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 8–13. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 13</figureCitation>
|
||||
); S4 medially sulcate, hairy, and S6 long, ≥1.2
|
||||
<emphasis id="B931EA96CC2771597C3410D50E2CABC6" bold="true" box="[859,878,1453,1473]" italics="true" pageId="51" pageNumber="405">9</emphasis>
|
||||
longer than broad.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC2771597C4410910E9EA8B1" blockId="51.[811,1426,1513,1718]" pageId="51" pageNumber="405">
|
||||
<emphasis id="B931EA96CC2771597C4410910EEBABF9" box="[811,937,1513,1534]" italics="true" pageId="51" pageNumber="405">Comments:</emphasis>
|
||||
Ten species are classified in this subgenus (
|
||||
<bibRefCitation id="EFD44B75CC2771597CC3137F09ABA81A" author="Ascher JS & Pickering J" box="[940,1257,1543,1565]" pageId="51" pageNumber="405" refId="ref42522" refString="Ascher JS, Pickering J. 2015. Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available via http://www.discoverlife.org/mp/20q?guide=Apoidea_species [Accessed 10 March 2015]" type="url" year="2015">Ascher & Pickering, 2015</bibRefCitation>
|
||||
), with a new one described by
|
||||
<bibRefCitation id="EFD44B75CC2771597B4E135E0849A83B" author="Rocha-Filho LC" box="[1057,1291,1574,1596]" pageId="51" pageNumber="405" pagination="9 - 28" refId="ref43631" refString="Rocha-Filho LC. 2016. A revision of the cleptoparasitic bee genus Coelioxys (Hymenoptera: Megachilidae) from Australia. European Journal of Entomology 113: 9-28." type="journal article" year="2016">Rocha-Filho (2016)</bibRefCitation>
|
||||
; the four Afrotropical species were revised by
|
||||
<bibRefCitation id="EFD44B75CC2771597B8E133C08D0A85D" author="Pasteels JJ" box="[1249,1426,1604,1626]" pageId="51" pageNumber="405" pagination="1 - 139" refId="ref43437" refString="Pasteels JJ. 1968. Revision des Megachilidae (Hymenoptera Apoidea) de L'Afrique Noire. II. Le Genre Coelioxys. Annales Sciences Zoologiques, Mus <20> ee Royal de L'Afrique Centrale, Tervuren, Serie In- 8 167: 1-139." type="journal article" year="1968">Pasteels (1968)</bibRefCitation>
|
||||
and classified as
|
||||
<emphasis id="B931EA96CC2771597C9A131B093AA87F" box="[1013,1144,1635,1656]" italics="true" pageId="51" pageNumber="405">Liothyrapis</emphasis>
|
||||
, the other species come from Indomalayan or Palaearctic regions,
|
||||
<collectingCountry id="F3527614CC2771597A4F13F908D0A890" box="[1312,1426,1665,1687]" name="Mauritius" pageId="51" pageNumber="405">Mauritius</collectingCountry>
|
||||
or
|
||||
<collectingCountry id="F3527614CC2771597C2413D80E9BA8B1" box="[843,985,1696,1718]" name="Madagascar" pageId="51" pageNumber="405">Madagascar</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC27715E7C4413A60F64AF30" blockId="51.[811,1426,1757,1871]" lastBlockId="52.[162,776,199,312]" lastPageId="52" lastPageNumber="406" pageId="51" pageNumber="405">
|
||||
<emphasis id="B931EA96CC2771597C4413A60EB5A8F4" box="[811,1015,1757,1779]" italics="true" pageId="51" pageNumber="405">List of species:</emphasis>
|
||||
<taxonomicName id="4C454D07CC2771597B6A13A508CCA8F5" authority="Friese, 1911" authorityName="Friese" authorityYear="1911" box="[1029,1422,1757,1779]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="analis">
|
||||
<emphasis id="B931EA96CC2771597B6A13A50993A8F5" box="[1029,1233,1757,1778]" italics="true" pageId="51" pageNumber="405">Coelioxys analis</emphasis>
|
||||
Friese, 1911
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2771597C44138409EAA916" authority="Smith, 1875" authorityName="Smith" authorityYear="1875" box="[811,1192,1787,1810]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="basalis">
|
||||
<emphasis id="B931EA96CC2771597C4413840EBBA916" box="[811,1017,1788,1809]" italics="true" pageId="51" pageNumber="405">Coelioxys basalis</emphasis>
|
||||
Smith, 1875
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2771597BAE13840E8BA937" authority="Smith, 1854" authorityName="Smith" authorityYear="1854" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="ducalis">
|
||||
<emphasis id="B931EA96CC2771597BAE138408D0A916" box="[1217,1426,1788,1809]" italics="true" pageId="51" pageNumber="405">Coelioxys ducalis</emphasis>
|
||||
Smith, 1854
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2771597C8A126208CCA928" authority="Smith, 1873" authorityName="Smith" authorityYear="1873" box="[997,1422,1818,1840]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="fenestrata">
|
||||
<emphasis id="B931EA96CC2771597C8A1262099AA928" box="[997,1240,1818,1839]" italics="true" pageId="51" pageNumber="405">Coelioxys fenestrata</emphasis>
|
||||
Smith, 1873
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC2771597C44124109B6A948" authority="Rocha-Filho, 2016" authorityName="Rocha-Filho" authorityYear="2016" box="[811,1268,1849,1871]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="julia">
|
||||
<emphasis id="B931EA96CC2771597C4412410EA9A949" box="[811,1003,1849,1870]" italics="true" pageId="51" pageNumber="405">Coelioxys julia</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC2771597B62124109B6A948" author="Rocha-Filho LC" box="[1037,1268,1849,1871]" pageId="51" pageNumber="405" pagination="9 - 28" refId="ref43631" refString="Rocha-Filho LC. 2016. A revision of the cleptoparasitic bee genus Coelioxys (Hymenoptera: Megachilidae) from Australia. European Journal of Entomology 113: 9-28." type="journal article" year="2016">Rocha-Filho, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
*,
|
||||
<taxonomicName id="4C454D07CC27715E7A4612410C87AEDB" authority="Strand, 1913" authorityName="Strand" authorityYear="1913" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" lastPageId="52" lastPageNumber="406" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="kosemponis">
|
||||
<emphasis id="B931EA96CC27715E7A4612410C66AEDB" italics="true" lastPageId="52" lastPageNumber="406" pageId="51" pageNumber="405">Coelioxys kosemponis</emphasis>
|
||||
Strand, 1913
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7EBC15BF0D99AEFD" authority="Friese, 1913" authorityName="Friese" authorityYear="1913" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="maculata">
|
||||
<emphasis id="B931EA96CC20715E7EBC15BF0FF3AEDB" box="[467,689,199,220]" italics="true" pageId="52" pageNumber="406">Coelioxys maculata</emphasis>
|
||||
Friese, 1913
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7E69159D0E46AEFC" authority="Pasteels, 1968" authorityName="Pasteels" authorityYear="1968" box="[262,772,229,251]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="maculoides">
|
||||
<emphasis id="B931EA96CC20715E7E69159D0F55AEFD" box="[262,535,229,250]" italics="true" pageId="52" pageNumber="406">Coelioxys maculoides</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC20715E7D51159D0E46AEFC" author="Pasteels JJ" box="[574,772,229,251]" pageId="52" pageNumber="406" pagination="1 - 139" refId="ref43437" refString="Pasteels JJ. 1968. Revision des Megachilidae (Hymenoptera Apoidea) de L'Afrique Noire. II. Le Genre Coelioxys. Annales Sciences Zoologiques, Mus <20> ee Royal de L'Afrique Centrale, Tervuren, Serie In- 8 167: 1-139." type="journal article" year="1968">Pasteels, 1968</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7FCD147C0C51AF1E" authorityName="Smith" authorityYear="1854" box="[162,275,260,281]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="torrida">
|
||||
<emphasis id="B931EA96CC20715E7FCD147C0DF6AF1E" box="[162,180,260,281]" italics="true" pageId="52" pageNumber="406">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC20715E7FAC147C0C51AF1E" box="[195,275,260,281]" italics="true" pageId="52" pageNumber="406">torrida</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC20715E7E45147C0F8BAF1D" authority="Pasteels, 1968" authorityName="Pasteels" authorityYear="1968" box="[298,713,260,282]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="torridula">
|
||||
<emphasis id="B931EA96CC20715E7E45147C0F49AF1E" box="[298,523,260,281]" italics="true" pageId="52" pageNumber="406">Coelioxys torridula</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC20715E7D74147C0F8BAF1D" author="Pasteels JJ" box="[539,713,260,282]" pageId="52" pageNumber="406" pagination="1 - 139" refId="ref43437" refString="Pasteels JJ. 1968. Revision des Megachilidae (Hymenoptera Apoidea) de L'Afrique Noire. II. Le Genre Coelioxys. Annales Sciences Zoologiques, Mus <20> ee Royal de L'Afrique Centrale, Tervuren, Serie In- 8 167: 1-139." type="journal article" year="1968">Pasteels, 1968</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C454D07CC20715E7FCD145A0F60AF30" authority="Schulz, 1904" authorityName="Schulz" authorityYear="1904" box="[162,546,290,312]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="weinlandi">
|
||||
<emphasis id="B931EA96CC20715E7FCD145A0CC7AF30" box="[162,389,290,311]" italics="true" pageId="52" pageNumber="406">Coelioxys weinlandi</emphasis>
|
||||
Schulz, 1904
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
286
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286
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<treatment id="03EC8792CC33714D7FDF15AE0869AF94" ID-DOI="http://doi.org/10.5281/zenodo.14806995" ID-Zenodo-Dep="14806995" LSID="urn:lsid:plazi:treatment:03EC8792CC33714D7FDF15AE0869AF94" httpUri="http://treatment.plazi.org/id/03EC8792CC33714D7FDF15AE0869AF94" lastPageNumber="393" pageId="39" pageNumber="393">
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<paragraph id="8BFA3684CC33714D7FDF15AE0F9DAEEB" blockId="39.[176,735,210,237]" box="[176,735,210,237]" pageId="39" pageNumber="393">
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<heading id="D0B281E8CC33714D7FDF15AE0F9DAEEB" box="[176,735,210,237]" centered="true" fontSize="9" level="2" pageId="39" pageNumber="393" reason="2">
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<taxonomicName id="4C454D07CC33714D7FDF15AE0F9DAEEB" authority="MITCHELL, 1973" authorityName="Mitchell" authorityYear="1973" box="[176,735,210,237]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="380" phylum="Arthropoda" rank="subGenus" subGenus="Boreocoelioxys">
|
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<emphasis id="B931EA96CC33714D7FDF15AE0F53AEEA" box="[176,529,211,237]" italics="true" pageId="39" pageNumber="393">
|
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<smallCapsWord id="8D1CA058CC33714D7FDF15AE0C6CAEED" baselines="230,230" box="[176,302,214,235]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Coelioxys" pageId="39" pageNumber="393">COELIOXYS</smallCapsWord>
|
||||
(
|
||||
<smallCapsWord id="8D1CA058CC33714D7E2E15AD0F4AAEED" baselines="230,230" box="[321,520,213,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Boreocoelioxys" pageId="39" pageNumber="393">BOREOCOELIOXYS</smallCapsWord>
|
||||
)
|
||||
</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC33714D7D7415AC0F9DAEEB" author="Mitchell TB" box="[539,735,210,237]" pageId="39" pageNumber="393" refId="ref43263" refString="Mitchell TB. 1973. A subgeneric revision of the genus Coelioxys of the Western Hemisphere (Hymenoptera: Megachilidae). Raleigh: Department of Entomology, North Carolina State University." type="book" year="1973">
|
||||
<smallCapsWord id="8D1CA058CC33714D7D7415AC0FD2AEEC" baselines="230,231" box="[539,656,212,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Mitchell" pageId="39" pageNumber="393">MITCHELL</smallCapsWord>
|
||||
, 1973
|
||||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C35F650FCC33714D7FFB145B0869AF94" pageId="39" pageNumber="393" type="materials_examined">
|
||||
<paragraph id="8BFA3684CC33714D7FFB145B0FE9AF3F" blockId="39.[148,683,290,312]" box="[148,683,290,312]" pageId="39" pageNumber="393">
|
||||
<emphasis id="B931EA96CC33714D7FFB145B0C65AF3F" box="[148,295,291,312]" italics="true" pageId="39" pageNumber="393">
|
||||
<typeStatus id="54FE8826CC33714D7FFB145B0D88AF3F" box="[148,202,291,312]" pageId="39" pageNumber="393">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C454D07CC33714D7E59145A0FE5AF3F" authority="Smith, 1854" authorityName="Smith" authorityYear="1854" box="[310,679,290,312]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="rufitarsis">
|
||||
<emphasis id="B931EA96CC33714D7E59145A0F52AF30" box="[310,528,290,311]" italics="true" pageId="39" pageNumber="393">Coelioxys rufitarsis</emphasis>
|
||||
Smith, 1854
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC33714D7FFB14180C5EAD9C" blockId="39.[148,763,351,923]" pageId="39" pageNumber="393">
|
||||
<emphasis id="B931EA96CC33714D7FFB14180C49AF72" box="[148,267,352,373]" italics="true" pageId="39" pageNumber="393">Diagnosis:</emphasis>
|
||||
Species in this subgenus are characterized by the combination of long, dense pubescence on head and mesosoma never accompanied by short, scale-like hairs, and with postgradular grooves of
|
||||
<collectionCode id="ED54AE41CC33714D7E8F14A20CB3AFE8" box="[480,497,474,495]" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="393">T</collectionCode>
|
||||
2 and
|
||||
<collectionCode id="ED54AE41CC33714D7D3114A20F32AFE8" box="[606,624,474,495]" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="393">T</collectionCode>
|
||||
3 complete (
|
||||
<figureCitation id="137E2A01CC33714D7FF314800DB2AC0A" box="[156,240,504,526]" captionStart="Figures 24 and 25" captionStartId="26.[162,252,1593,1612]" captionText="Figures 24 and 25. Female, dorsal view of T2–T3 (for postgradular grooves): 24, Coelioxys rufitarsis; 25, Coelioxys apacheorum. Scale bars: 1.5 mm." pageId="39" pageNumber="393">Fig. 24</figureCitation>
|
||||
). This combination is also observed in all species of
|
||||
<emphasis id="B931EA96CC33714D7E73176F0C6CAC2B" box="[284,302,535,556]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC33714D7E2A176F0C99AC2B" box="[325,475,535,556]" italics="true" pageId="39" pageNumber="393">
|
||||
<taxonomicName id="4C454D07CC33714D7E2A176F0CECAC2B" box="[325,430,535,556]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
|
||||
<taxonomicNameLabel id="A20257EDCC33714D7ED8176F0C99AC2B" box="[439,475,535,556]" pageId="39" pageNumber="393" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
). In
|
||||
<emphasis id="B931EA96CC33714D7D49176F0F7AAC2B" box="[550,568,535,556]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC33714D7D3F176F0FB1AC2B" box="[592,755,535,556]" italics="true" pageId="39" pageNumber="393">Boreocoelioxys</emphasis>
|
||||
) females, however, the S6 is elongate, gradually tapering towards the apex and conspicuously notched subapically [
|
||||
<figureCitation id="137E2A01CC33714D7EE2170A0CA0AC8F" box="[397,482,626,648]" captionStart="Figures 26–29" captionStartId="27.[148,238,1449,1468]" captionTargetBox="[157,1420,199,1417]" captionTargetId="figure-252@27.[156,1420,198,1416]" captionTargetPageId="27" captionText="Figures 26–29. 26, female, Coelioxys porterae: dorsal view of T6 and S6. 27, male, Coelioxys sayi: dorsal view of T2 (fovea indicated by red arrow). 28, female, Coelioxys quadridentata, dorsal view of T6 and S6. 29, male, Coelioxys sodalis: dorsal view of T2 (punctate area indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807017" httpUri="https://zenodo.org/record/14807017/files/figure.png" pageId="39" pageNumber="393">Fig. 26</figureCitation>
|
||||
; constricted subapically and never notched in both
|
||||
<emphasis id="B931EA96CC33714D7D5C17E90F07ACA1" box="[563,581,657,678]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC33714D7D3217E90FB0ACA0" box="[605,754,657,679]" italics="true" pageId="39" pageNumber="393">
|
||||
<taxonomicName id="4C454D07CC33714D7D3217E90F84ACA1" box="[605,710,657,678]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
|
||||
<taxonomicNameLabel id="A20257EDCC33714D7DA017EA0FB0ACA0" box="[719,754,658,679]" pageId="39" pageNumber="393" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
) (
|
||||
<figureCitation id="137E2A01CC33714D7FF217D70DACACC2" box="[157,238,687,709]" captionStart="Figures 26–29" captionStartId="27.[148,238,1449,1468]" captionTargetBox="[157,1420,199,1417]" captionTargetId="figure-252@27.[156,1420,198,1416]" captionTargetPageId="27" captionText="Figures 26–29. 26, female, Coelioxys porterae: dorsal view of T6 and S6. 27, male, Coelioxys sayi: dorsal view of T2 (fovea indicated by red arrow). 28, female, Coelioxys quadridentata, dorsal view of T6 and S6. 29, male, Coelioxys sodalis: dorsal view of T2 (punctate area indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807017" httpUri="https://zenodo.org/record/14807017/files/figure.png" pageId="39" pageNumber="393">Fig. 28</figureCitation>
|
||||
) and
|
||||
<emphasis id="B931EA96CC33714D7E5B17C80C04ACC2" box="[308,326,688,709]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC33714D7E3017D70CB7ACC3" box="[351,501,687,708]" italics="true" pageId="39" pageNumber="393">Xerocoelioxys</emphasis>
|
||||
) females (
|
||||
<figureCitation id="137E2A01CC33714D7D1F17D70F86ACC2" box="[624,708,687,709]" captionStart="Figures 14 and 15" captionStartId="24.[162,252,1640,1659]" captionText="Figures 14 and 15. Coelioxys edita: 14, female, dorsal view of T6 and S6; 15, male, lateral view of head (for gena pilosity). Scale bars: 1.5 mm." pageId="39" pageNumber="393">Fig. 14</figureCitation>
|
||||
)]. In
|
||||
<emphasis id="B931EA96CC33714D7FFB17B60DE4ACE4" box="[148,166,718,739]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC33714D7FD117B60C23ACE4" box="[190,353,718,739]" italics="true" pageId="39" pageNumber="393">Boreocoelioxys</emphasis>
|
||||
) males the postgradular grooves of
|
||||
<collectionCode id="ED54AE41CC33714D7FFB17950DE7AD05" box="[148,165,749,770]" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="393">T</collectionCode>
|
||||
2 have a sublateral, slender fovea on each side (
|
||||
<figureCitation id="137E2A01CC33714D7FF216730DB2AD27" box="[157,240,779,801]" captionStart="Figures 26–29" captionStartId="27.[148,238,1449,1468]" captionTargetBox="[157,1420,199,1417]" captionTargetId="figure-252@27.[156,1420,198,1416]" captionTargetPageId="27" captionText="Figures 26–29. 26, female, Coelioxys porterae: dorsal view of T6 and S6. 27, male, Coelioxys sayi: dorsal view of T2 (fovea indicated by red arrow). 28, female, Coelioxys quadridentata, dorsal view of T6 and S6. 29, male, Coelioxys sodalis: dorsal view of T2 (punctate area indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807017" httpUri="https://zenodo.org/record/14807017/files/figure.png" pageId="39" pageNumber="393">Fig. 27</figureCitation>
|
||||
), whereas in males of
|
||||
<emphasis id="B931EA96CC33714D7E9116740F52AD26" box="[510,528,780,801]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC33714D7D4816730FFFAD26" box="[551,701,779,801]" italics="true" pageId="39" pageNumber="393">
|
||||
<taxonomicName id="4C454D07CC33714D7D4816730FD2AD27" box="[551,656,779,800]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
|
||||
<taxonomicNameLabel id="A20257EDCC33714D7DF516740FFFAD26" box="[666,701,780,801]" pageId="39" pageNumber="393" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
) and
|
||||
<emphasis id="B931EA96CC33714D7FFB16520DE4AD38" box="[148,166,810,831]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC33714D7FD116520C16AD38" box="[190,340,810,831]" italics="true" pageId="39" pageNumber="393">Xerocoelioxys</emphasis>
|
||||
) spp. the postgradular area of
|
||||
<collectionCode id="ED54AE41CC33714D7DAE16520F90AD38" box="[705,722,810,831]" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="393">T</collectionCode>
|
||||
2 is never foveate, but instead with sublateral, slender, densely punctate area on each side (
|
||||
<figureCitation id="137E2A01CC33714D7D1D161F0F87AD7B" box="[626,709,871,893]" captionStart="Figures 26–29" captionStartId="27.[148,238,1449,1468]" captionTargetBox="[157,1420,199,1417]" captionTargetId="figure-252@27.[156,1420,198,1416]" captionTargetPageId="27" captionText="Figures 26–29. 26, female, Coelioxys porterae: dorsal view of T6 and S6. 27, male, Coelioxys sayi: dorsal view of T2 (fovea indicated by red arrow). 28, female, Coelioxys quadridentata, dorsal view of T6 and S6. 29, male, Coelioxys sodalis: dorsal view of T2 (punctate area indicated by red arrow). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807017" httpUri="https://zenodo.org/record/14807017/files/figure.png" pageId="39" pageNumber="393">Fig. 29</figureCitation>
|
||||
), or unmodified.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC33714D7FFB16BB0FACA83B" blockId="39.[148,763,962,1596]" pageId="39" pageNumber="393">
|
||||
<emphasis id="B931EA96CC33714D7FFB16BB0C50ADDF" box="[148,274,963,984]" italics="true" pageId="39" pageNumber="393">Comments:</emphasis>
|
||||
This subgenus has been regarded as mainly Holarctic (
|
||||
<bibRefCitation id="EFD44B75CC33714D7E0616990F63ADF1" author="Michener CD" box="[361,545,993,1015]" pageId="39" pageNumber="393" refId="ref43240" refString="Michener CD. 2007. The bees of the world, 2 nd edn. Baltimore: The John Hopkins University Press." type="book" year="2007">Michener, 2007</bibRefCitation>
|
||||
), but as such it is very weakly supported (see Results and Discussion and
|
||||
<figureCitation id="137E2A01CC33714D7FA711660C48AA34" box="[200,266,1054,1076]" captionStart="Figure 1" captionStartId="14.[162,241,1763,1782]" captionText="Figure 1. Strict consensus tree of the 25 equally most-parsimonious trees in the equal-weights analysis. Branch support values are displayed below the nodes in the following order: bootstrap, Bremer support, and symmetric resampling values (values in square brackets are negative). When only two values are displayed they refer to bootstrap and symmetric resampling, as the Bremer values could not be estimated, and when only one value is shown it refers to bootstrap. Colours indicate Michener’s subgeneric classification." pageId="39" pageNumber="393">Fig. 1</figureCitation>
|
||||
). In contrast, the New World clade is well supported. In light of this,
|
||||
<emphasis id="B931EA96CC33714D7EBD11450CA6AA55" box="[466,484,1085,1106]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC33714D7E9411450FDCAA55" box="[507,670,1085,1106]" italics="true" pageId="39" pageNumber="393">Boreocoelioxys</emphasis>
|
||||
) is now defined as comprising only the New World species, among which the Nearctic species and
|
||||
<taxonomicName id="4C454D07CC33714D7DFD11020CDDAAA9" authority="Baker, 1975" authorityName="Baker" authorityYear="1975" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="oaxacana">
|
||||
<emphasis id="B931EA96CC33714D7DFD11020DBDAAA9" italics="true" pageId="39" pageNumber="393">Coelioxys oaxacana</emphasis>
|
||||
<bibRefCitation id="EFD44B75CC33714D7E6111E00CDDAAA9" author="Baker JR" box="[270,415,1176,1198]" pageId="39" pageNumber="393" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker, 1975</bibRefCitation>
|
||||
</taxonomicName>
|
||||
were revised and keyed by
|
||||
<bibRefCitation id="EFD44B75CC33714D7FFB11CF0C6CAACA" author="Baker JR" box="[148,302,1207,1229]" pageId="39" pageNumber="393" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker (1975)</bibRefCitation>
|
||||
. There are two additional species from the Neotropics:
|
||||
<taxonomicName id="4C454D07CC33714D7E0F11AE0D88AB0E" authority="Genaro, 2001" authorityName="Genaro" authorityYear="2001" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="sannicolarensis">
|
||||
<emphasis id="B931EA96CC33714D7E0F11AE0FCFAAEC" box="[352,653,1238,1259]" italics="true" pageId="39" pageNumber="393">Coelioxys sannicolarensis</emphasis>
|
||||
Genaro, 2001
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C454D07CC33714D7E4E118C0FF4AB0E" authority="Friese, 1917" authorityName="Friese" authorityYear="1917" box="[289,694,1268,1290]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="schmidti">
|
||||
<emphasis id="B931EA96CC33714D7E4E118C0F41AB0E" box="[289,515,1268,1289]" italics="true" pageId="39" pageNumber="393">Coelioxys schmidti</emphasis>
|
||||
Friese, 1917
|
||||
</taxonomicName>
|
||||
. The following species are being transferred to other subgenera according to the phylogenetic results:
|
||||
<taxonomicName id="4C454D07CC33714D7FFB10280DB3AB62" authorityName="Forster" authorityYear="1853" box="[148,241,1360,1381]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="alata">
|
||||
<emphasis id="B931EA96CC33714D7FFB10280DE4AB62" box="[148,166,1360,1381]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC33714D7FD910280DB3AB62" box="[182,241,1360,1381]" italics="true" pageId="39" pageNumber="393">alata</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC33714D7E6810280CD7AB62" authorityName="Smith" authorityYear="1854" box="[263,405,1360,1381]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="funeraria">
|
||||
<emphasis id="B931EA96CC33714D7E6810280C5BAB62" box="[263,281,1360,1381]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC33714D7E4610280CD7AB62" box="[297,405,1360,1381]" italics="true" pageId="39" pageNumber="393">funeraria</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C454D07CC33714D7E8B10280F2CAB62" authorityName="Lepeletier & Serville" authorityYear="1825" box="[484,622,1360,1381]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="rufescens">
|
||||
<emphasis id="B931EA96CC33714D7E8B10280CB4AB62" box="[484,502,1360,1381]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC33714D7D6910280F2CAB62" box="[518,622,1360,1381]" italics="true" pageId="39" pageNumber="393">rufescens</emphasis>
|
||||
</taxonomicName>
|
||||
. The other species from the Palaeartic region that were previously assigned to
|
||||
<emphasis id="B931EA96CC33714D7ECA10F50CF5ABA5" box="[421,439,1421,1442]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC33714D7EA010F50F30ABA5" box="[463,626,1421,1442]" italics="true" pageId="39" pageNumber="393">Boreocoelioxys</emphasis>
|
||||
) (
|
||||
<bibRefCitation id="EFD44B75CC33714D7DE310F50C08ABC6" author="Ascher JS & Pickering J" pageId="39" pageNumber="393" refId="ref42522" refString="Ascher JS, Pickering J. 2015. Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available via http://www.discoverlife.org/mp/20q?guide=Apoidea_species [Accessed 10 March 2015]" type="url" year="2015">Ascher & Pickering, 2015</bibRefCitation>
|
||||
) may belong to
|
||||
<emphasis id="B931EA96CC33714D7D6210D40F5DABC6" box="[525,543,1452,1473]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
. (
|
||||
<emphasis id="B931EA96CC33714D7D5810D40F93ABC6" box="[567,721,1452,1473]" italics="true" pageId="39" pageNumber="393">Paracoelioxys</emphasis>
|
||||
) or to the new subgenus described herein to encompass
|
||||
<taxonomicName id="4C454D07CC33714D7FFB10910C5CABF9" authorityName="Lepeletier & Serville" authorityYear="1825" box="[148,286,1513,1534]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="rufescens">
|
||||
<emphasis id="B931EA96CC33714D7FFB10910C5CABF9" box="[148,286,1513,1534]" italics="true" pageId="39" pageNumber="393">C. rufescens</emphasis>
|
||||
</taxonomicName>
|
||||
; however, it is important to highlight that a revision of those species is necessary in order to establish an accurate classification for this fauna.
|
||||
</paragraph>
|
||||
<paragraph id="8BFA3684CC33714D7FFB131B0869AF94" blockId="39.[148,763,1635,1718]" lastBlockId="39.[811,1426,199,404]" pageId="39" pageNumber="393">
|
||||
<emphasis id="B931EA96CC33714D7FFB131B0C0DA87F" box="[148,335,1635,1656]" italics="true" pageId="39" pageNumber="393">List of species:</emphasis>
|
||||
<taxonomicName id="4C454D07CC33714D7E31131B0FB5A87F" authority="Crawford, 1914" authorityName="Crawford" authorityYear="1914" box="[350,759,1635,1657]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="banksi">
|
||||
<emphasis id="B931EA96CC33714D7E31131B0F67A87F" box="[350,549,1635,1656]" italics="true" pageId="39" pageNumber="393">Coelioxys banksi</emphasis>
|
||||
Crawford, 1914
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC33714D7FFB13F90F54A891" authority="Cresson, 1872" authorityName="Cresson" authorityYear="1872" box="[148,534,1665,1687]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="insita">
|
||||
<emphasis id="B931EA96CC33714D7FFB13F90C11A890" box="[148,339,1665,1687]" italics="true" pageId="39" pageNumber="393">Coelioxys insita</emphasis>
|
||||
Cresson, 1872
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC33714D7D4013F90C00A8B1" authority="Cresson, 1864" authorityName="Cresson" authorityYear="1864" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="moesta">
|
||||
<emphasis id="B931EA96CC33714D7D4013F90FB9A890" box="[559,763,1665,1687]" italics="true" pageId="39" pageNumber="393">Coelioxys moesta</emphasis>
|
||||
Cresson, 1864
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC33714D7E3513D80E26AEDB" authority="Cockerell, 1909" authorityName="Cockerell" authorityYear="1909" box="[346,868,199,1718]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="novomexicana">
|
||||
<emphasis id="B931EA96CC33714D7E3513D80F34A8B1" box="[346,630,1696,1718]" italics="true" pageId="39" pageNumber="393">Coelioxys novomexicana</emphasis>
|
||||
Cockerell, 1909
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC33714D7C1D15BF0EBCAEDA" authorityName="Baker" authorityYear="1975" box="[882,1022,199,221]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="oaxacana">
|
||||
<emphasis id="B931EA96CC33714D7C1D15BF0EC6AEDB" box="[882,900,199,220]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC33714D7CFC15B00EBCAEDA" box="[915,1022,200,221]" italics="true" pageId="39" pageNumber="393">oaxacana</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC33714D7B6115BF09F1AEDB" authorityName="Say" authorityYear="1824" box="[1038,1203,199,220]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="octodentata">
|
||||
<emphasis id="B931EA96CC33714D7B6115BF0962AEDB" box="[1038,1056,199,220]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC33714D7B5F15BF09F1AEDB" box="[1072,1203,199,220]" italics="true" pageId="39" pageNumber="393">octodentata</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC33714D7BAC15BF0EABAEFD" authority="Cockerell, 1900" authorityName="Cockerell" authorityYear="1900" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="porterae">
|
||||
<emphasis id="B931EA96CC33714D7BAC15BF08D0AEDA" box="[1219,1426,199,221]" italics="true" pageId="39" pageNumber="393">Coelioxys porterae</emphasis>
|
||||
Cockerell, 1900
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC33714D7B6F159D08CCAEFD" authority="Crawford, 1914" authorityName="Crawford" authorityYear="1914" box="[1024,1422,229,251]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="pratti">
|
||||
<emphasis id="B931EA96CC33714D7B6F159D09FFAEFC" box="[1024,1213,229,251]" italics="true" pageId="39" pageNumber="393">Coelioxys pratti</emphasis>
|
||||
Crawford, 1914
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC33714D7C44147C0EF4AF1E" authorityName="Smith" authorityYear="1854" box="[811,950,260,281]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="rufitarsis">
|
||||
<emphasis id="B931EA96CC33714D7C44147C0E7FAF1E" box="[811,829,260,281]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC33714D7C22147C0EF4AF1E" box="[845,950,260,281]" italics="true" pageId="39" pageNumber="393">rufitarsis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC33714D7CBD147C08CCAF1D" authority="Cockerell, 1925" authorityName="Cockerell" authorityYear="1925" box="[978,1422,260,282]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="salinaria">
|
||||
<emphasis id="B931EA96CC33714D7CBD147C09FBAF1E" box="[978,1209,260,281]" italics="true" pageId="39" pageNumber="393">Coelioxys salinaria</emphasis>
|
||||
Cockerell, 1925
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C454D07CC33714D7C44145B0EBFAF30" authorityName="Genaro" authorityYear="2001" box="[811,1021,290,312]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="sannicolarensis">
|
||||
<emphasis id="B931EA96CC33714D7C44145B0E7FAF3F" box="[811,829,291,312]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC33714D7C22145A0EBFAF30" box="[845,1021,290,311]" italics="true" pageId="39" pageNumber="393">sannicolarensis</emphasis>
|
||||
</taxonomicName>
|
||||
*,
|
||||
<taxonomicName id="4C454D07CC33714D7B4F145B092DAF3F" baseAuthorityName="Robertson" baseAuthorityYear="1897" box="[1056,1135,291,312]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="sayi">
|
||||
<emphasis id="B931EA96CC33714D7B4F145B0970AF3F" box="[1056,1074,291,312]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC33714D7B2E145B092DAF3F" box="[1089,1135,291,312]" italics="true" pageId="39" pageNumber="393">sayi</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C454D07CC33714D7BD0145B0806AF30" authorityName="Friese" authorityYear="1917" box="[1215,1348,290,312]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="schmidti">
|
||||
<emphasis id="B931EA96CC33714D7BD0145B0993AF3F" box="[1215,1233,291,312]" italics="true" pageId="39" pageNumber="393">C</emphasis>
|
||||
.
|
||||
<emphasis id="B931EA96CC33714D7B8E145A0806AF30" box="[1249,1348,290,311]" italics="true" pageId="39" pageNumber="393">schmidti</emphasis>
|
||||
</taxonomicName>
|
||||
*. The Neotropical species
|
||||
<taxonomicName id="4C454D07CC33714D7B70143908D0AF51" authority="Friese, 1921" authorityName="Friese" authorityYear="1921" box="[1055,1426,321,343]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="393" phylum="Arthropoda" rank="species" species="auripes">
|
||||
<emphasis id="B931EA96CC33714D7B70143909AFAF51" box="[1055,1261,321,342]" italics="true" pageId="39" pageNumber="393">Coelioxys auripes</emphasis>
|
||||
Friese, 1921
|
||||
</taxonomicName>
|
||||
also seems to belong to this subgenus given the complete postgradular grooves of
|
||||
<collectionCode id="ED54AE41CC33714D7BC2140609FCAF94" box="[1197,1214,382,403]" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="393">T</collectionCode>
|
||||
2 and
|
||||
<collectionCode id="ED54AE41CC33714D7A6914060858AF94" box="[1286,1306,382,403]" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="393">T</collectionCode>
|
||||
3.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
89
data/CF/23/65/CF236553FFA55545FE91D46BFD70F939.xml
Normal file
89
data/CF/23/65/CF236553FFA55545FE91D46BFD70F939.xml
Normal file
|
|
@ -0,0 +1,89 @@
|
|||
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from Greek, meaning dog, in reference to the enlarged teeth of the jaws, and
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(Latin), a clupeoid fish.
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See
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below.
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496
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|
||||
<specimenCode id="172C7C3EFFA55545FEC4D60CFE69F881" box="[287,403,1818,1839]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="3" pageNumber="178">MCP 3447</specimenCode>
|
||||
-PV, fossil specimen corresponding to the anterior part of a fish including the head and the first 11 vertebrae.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFA55545FCE6D1D1FAB2FEDB" blockId="3.[829,1444,199,373]" pageId="3" pageNumber="178">
|
||||
<emphasis id="75FE0857FFA55545FCE6D1D1FC41FF72" box="[829,955,199,220]" italics="true" pageId="3" pageNumber="178">Etymology:</emphasis>
|
||||
The species is named after Gareth J. Nelson (University of Melbourne,
|
||||
<collectingCountry id="3F9D94D5FFA55545FB38D1F3FAA8FF55" box="[1251,1362,229,251]" name="Australia" pageId="3" pageNumber="178">Australia</collectingCountry>
|
||||
). In a series of now-classic papers (
|
||||
<bibRefCitation id="231BA9B4FFA55545FB6BD012FAAAFEB4" author="Nelson GJ" box="[1200,1360,260,282]" pageId="3" pageNumber="178" pagination="389 - 399" refId="ref14073" refString="Nelson GJ. 1967. Gill arches of teleostean fishes of the family Clupeidae. Copeia 1967: 389-399." type="journal article" year="1967">Nelson, 1967</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="231BA9B4FFA55545FABCD012FA5AFEB7" author="Nelson GJ" box="[1383,1440,260,281]" pageId="3" pageNumber="178" pagination="1 - 30" refId="ref14095" refString="Nelson GJ. 1970. The hyobranchial apparatus of teleostean fishes of the families Engraulidae and Chirocentridae. American Museum Novitates 2410: 1-30." type="journal article" year="1970">1970</bibRefCitation>
|
||||
, 1973), Gary Nelson established the basis for understanding clupeomorph intrarelationships from a modern phylogenetic (cladistic) perspective.
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFA55545FCE6D08BFB01FDC7" blockId="3.[829,1444,413,618]" pageId="3" pageNumber="178">
|
||||
<emphasis id="75FE0857FFA55545FCE6D08BFBFDFE1C" box="[829,1031,413,434]" italics="true" pageId="3" pageNumber="178">Locality and age:</emphasis>
|
||||
<materialsCitation id="F7E2DE18FFA55545FBCDD08BFB02FDC7" collectorName="Miguel dos Campos" country="Brazil" location="Morro do Chaves Formation" pageId="3" pageNumber="178" specimenCount="1" stateProvince="Alagoas" typeStatus="holotype">
|
||||
The
|
||||
<typeStatus id="98316AE7FFA55545FB97D08BFB56FE1D" box="[1100,1196,413,435]" pageId="3" pageNumber="178" type="holotype">holotype</typeStatus>
|
||||
and single specimen known was collected in the shales of the
|
||||
<location id="4255829EFFA55545FAECD0AAFBE7FE5E" LSID="urn:lsid:plazi:treatment:CF236553FFA5554DFEC7D7DBFECDFE1D:4255829EFFA55545FAECD0AAFBE7FE5E" country="Brazil" name="Morro do Chaves Formation" pageId="3" pageNumber="178" stateProvince="Alagoas">Morro do Chaves Formation</location>
|
||||
of the Sergipe-Alagoas Basin exposed at the Cimpor quarry, municipality of S ao
|
||||
<emphasis id="75FE0857FFA55545FA50D0ECFA6DFDA2" box="[1419,1431,506,524]" italics="true" pageId="3" pageNumber="178">~</emphasis>
|
||||
<collectorName id="EA7FB193FFA55545FCE6D301FBC2FD82" box="[829,1080,535,557]" pageId="3" pageNumber="178">Miguel dos Campos</collectorName>
|
||||
,
|
||||
<collectingRegion id="854E1AA7FFA55545FB89D301FB04FD82" box="[1106,1278,535,557]" country="Brazil" name="Alagoas" pageId="3" pageNumber="178">Alagoas State</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="3F9D94D5FFA55545FACCD301FA9AFD83" box="[1303,1376,535,557]" name="Brazil" pageId="3" pageNumber="178">Brazil</collectingCountry>
|
||||
. The locality is correlated with the Barremian, Lower
|
||||
<taxonomicName id="808AAFC6FFA55545FCE6D342FB02FDC7" authority="(Campos Neto et al., 2007)" baseAuthorityName="Campos Neto" baseAuthorityYear="2007" box="[829,1272,596,618]" class="Ascidiacea" family="Polycitoridae" genus="Cretaceous" higherTaxonomySource="CoL" kingdom="Animalia" order="Aplousobranchia" pageId="3" pageNumber="178" phylum="Chordata" rank="genus">
|
||||
Cretaceous (
|
||||
<bibRefCitation id="231BA9B4FFA55545FC10D342FB15FDC7" author="Campos Neto OA & Lima WS & Cruz FE" box="[971,1263,596,618]" pageId="3" pageNumber="178" pagination="406 - 415" refId="ref12663" refString="Campos Neto OA, Lima WS, Cruz FE. 2007. Bacia de Sergipe-Alagoas. Boletim de Geociencias ^ da Petrobr as <20> 15: 406-415." type="journal article" year="2007">
|
||||
Campos Neto
|
||||
<emphasis id="75FE0857FFA55545FBB7D343FB5DFDC7" box="[1132,1191,596,618]" italics="true" pageId="3" pageNumber="178">et al.</emphasis>
|
||||
, 2007
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFA55545FCE6D384FC08FB42" blockId="3.[829,1444,657,1260]" pageId="3" pageNumber="178">
|
||||
<emphasis id="75FE0857FFA55545FCE6D384FB53FD08" box="[829,1193,657,679]" italics="true" pageId="3" pageNumber="178">
|
||||
Diagnosis:
|
||||
<taxonomicName id="808AAFC6FFA55545FC18D387FB53FD08" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[963,1193,657,678]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="3" pageNumber="178" phylum="Chordata" rank="species" species="nelsoni">Cynoclupea nelsoni</taxonomicName>
|
||||
</emphasis>
|
||||
is a medium-sized, herring-like clupeiform distinguished from all other members of the order by the following combination of characters: head triangular in lateral view, acuminated snout; long and straight lower jaws with a conspicuous dentition consisting anteriorly of enlarged and backward curved teeth followed by smaller teeth that extend well back on the dentary border (vs. absent or inconspicuous dentary teeth restricted to the anterior region of the bone); presence of three massive conical teeth slightly orientated backwards on the palatine; suspensorium slightly inclined posteriorly, being almost vertically orientated (vs. clearly posteriorly or anteriorly orientated); dorsal margins of the symplectic and quadrate in line; quadrate with a laminar outgrowth on the anterior margin; two laminar postcleithra (vs. rod-like postcleithra in clupeids; postcleithra absent in chirocentrids, engraulids, and most pristigasterids).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="0F9087CEFFA5554DFBFDD434FECDFE1D" lastPageId="11" lastPageNumber="186" pageId="3" pageNumber="178" type="description">
|
||||
<paragraph id="4735D445FFA55545FBFDD434FB41FA98" blockId="3.[1062,1211,1314,1335]" box="[1062,1211,1314,1335]" pageId="3" pageNumber="178">
|
||||
<smallCapsWord id="41D34299FFA55545FBFDD434FB41FA98" baselines="1330,1330" box="[1062,1211,1314,1335]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Description" pageId="3" pageNumber="178">DESCRIPTION</smallCapsWord>
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFA55545FCE6D446FB91F939" blockId="3.[829,1444,1360,1687]" pageId="3" pageNumber="178">
|
||||
The type specimen consists of the anterior part of an articulated fish including the head and the first 11 vertebrae. The head is relatively large, longer (
|
||||
<quantity id="807279A0FFA55545FC9ED4BDFC6CFA6E" box="[837,918,1451,1473]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="178" unit="mm" value="62.0">62 mm</quantity>
|
||||
) than deep (
|
||||
<quantity id="807279A0FFA55545FBE5D4BDFB68FA6E" box="[1086,1170,1451,1472]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="3" pageNumber="178" unit="mm" value="44.0">44 mm</quantity>
|
||||
), triangular in lateral view, with a somewhat acuminated snout. The dorsal and ventral profiles are almost straight, diverging from the terminal mouth towards the origin of the dorsal and pectoral fins, respectively. Based on head length and height,
|
||||
<taxonomicName id="808AAFC6FFA55545FBFBD752FB68F9F7" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[1056,1170,1604,1625]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="3" pageNumber="178" phylum="Chordata" rank="species" species="nelsoni">
|
||||
<emphasis id="75FE0857FFA55545FBFBD752FB68F9F7" box="[1056,1170,1604,1625]" italics="true" pageId="3" pageNumber="178">C. nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
is assumed to be moderately elongated, with an estimated standard length of approximately
|
||||
<quantity id="807279A0FFA55545FBDDD797FB9DF939" box="[1030,1127,1665,1687]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="178" unit="mm" value="240.0">240 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFA55545FCE6D7A9FC18F97A" blockId="3.[829,1444,1727,1901]" box="[829,994,1727,1748]" pageId="3" pageNumber="178">
|
||||
<emphasis id="75FE0857FFA55545FCE6D7A9FC18F97A" box="[829,994,1727,1748]" italics="true" pageId="3" pageNumber="178">Neurocranium</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFA55543FCE6D7CBFAE7F939" blockId="3.[829,1444,1727,1901]" lastBlockId="5.[829,1444,1329,1901]" lastPageId="5" lastPageNumber="180" pageId="3" pageNumber="178">
|
||||
The somewhat distorted nature of the fossil precludes precise observation of some features and anatomical details. The skull roof is represented by the frontals, which are ornamented with parallel grooves and extend between the parietal and the anterior tip of the head (
|
||||
<figureCitation id="DFB1C8C0FFA25542FE6CD1D1FE01FF72" box="[439,507,199,221]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="4" pageNumber="179">Fig. 3</figureCitation>
|
||||
). Anteriorly, the frontals are poorly preserved, represented by a portion of the bone associated with the supraorbital sensory canal. A small and tubular nasal is preserved at the front margin of the frontals (
|
||||
<figureCitation id="DFB1C8C0FFA25542FDD4D057FDAEFEF9" box="[527,596,321,343]" captionStart="Figure 6" captionStartId="7.[166,245,1221,1240]" captionTargetBox="[176,774,195,1185]" captionTargetId="figure-1477@7.[176,774,195,1185]" captionTargetPageId="7" captionText="Figure 6. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the snout region. bf, undetermined bony fragment; d, dentary; dt, dentary teeth; ec, ectopterygoid; en, endopterygoid; fr, frontal; mc, mandibular sensory canal; me, mesethmoid; n, nasal; pl, palatine; pm, premaxilla; ps, parasphenoid; vo, vomer. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14807129" httpUri="https://zenodo.org/record/14807129/files/figure.png" pageId="4" pageNumber="179">Fig. 6</figureCitation>
|
||||
). The frontal forms the roof of the orbit laterally, and a lateral wing seems to meet the sphenotic ventrally, limiting the dilatator fossa. Posteriorly, the frontals are more extensively preserved, diverging slightly and meeting the parietals. Even though the precise borders of the bones in this region of the cranium could not be defined, a large parietal, bearing a distinct backward-curved tube, which is herein interpreted as the supratemporal commissural sensory canal (a section of the extrascapular canal;
|
||||
<bibRefCitation id="231BA9B4FFA25542FBB1D034FAE2FE99" author="Di Dario F" box="[1130,1304,290,312]" pageId="4" pageNumber="179" pagination="257 - 270" refId="ref12901" refString="Di Dario F. 2004. Homology between the recessus lateralis and cephalic sensory canals, with the proposition of additional synapomorphies for the Clupeiformes and the Clupeoidei. Zoological Journal of the Linnean Society 141: 257-270." type="journal article" year="2004">Di Dario, 2004</bibRefCitation>
|
||||
), is identified in the right side of the skull. An upward compression of the ventral region of the posterior third of the neurocranium and some displacement of the bones at the back of the skull, posterior to the parietal, seem to have occurred during fossilization. An epioccipital is tentatively identified in the right side of the skull (
|
||||
<figureCitation id="DFB1C8C0FFA25542FC1DD0EEFBF1FDA3" box="[966,1035,504,526]" captionStart="Figure 4" captionStartId="5.[166,245,1157,1176]" captionTargetBox="[392,1216,195,1121]" captionTargetId="figure-1070@5.[392,1216,195,1121]" captionTargetPageId="5" captionText="Figure 4. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the skull roof. boc, basioccipital?; ds, dorsal scute fragments; ep, epioccipital; ex, exoccipital; fr, frontal; pa, parietal; pf, preepiotic fossa; pt, left (l) and right (r) post-temporals; pto, pterotic; sc, supracleithrum; so, supraoccipital. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807125" httpUri="https://zenodo.org/record/14807125/files/figure.png" pageId="4" pageNumber="179">Fig. 4</figureCitation>
|
||||
). In lateral view, the epioccipital has a typical tripartite structure and forms the posterodorsal corner of the skull in modern clupeiforms (
|
||||
<bibRefCitation id="231BA9B4FFA35543FF76D478FE9CFA2D" author="Ridewood WG" box="[173,358,1390,1412]" pageId="5" pageNumber="180" pagination="448 - 493" refId="ref14502" refString="Ridewood WG. 1904. On the cranial osteology of the clupeoid fishes. Proceedings of the Zoological Society of London 74: 448-493." type="journal article" year="1904">Ridewood, 1904</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="231BA9B4FFA35543FEAED478FD4FFA2A" author="Di Dario F & de Pinna MC" box="[373,693,1390,1412]" pageId="5" pageNumber="180" pagination="107 - 123" refId="ref12980" refString="Di Dario F, de Pinna MC. 2006. The supratemporal system and the pattern of ramification of cephalic sensory canals in Denticeps clupeoides (Denticipitoidei, Teleostei): additional evidence for monophyly of Clupeiformes and Clupeoidei. Pap eis <20> Avulsos de Zoologia (S ao ~ Paulo) 46: 107-123." type="journal article" year="2006">Di Dario & de Pinna, 2006</bibRefCitation>
|
||||
: fig. 3). The shape of the dorsal third of the bone is apparently intact, but its ventral arm, which usually, in modern clupeoids, articulates with the exoccipital and the pterotic inside the pre-epiotic fossa, seems to be dorsally dislodged in such a way that its ventral tip inserts more deeply in the pre-epiotic fossa (
|
||||
<figureCitation id="DFB1C8C0FFA35543FF75D752FF0EF9F7" box="[174,244,1604,1626]" captionStart="Figure 4" captionStartId="5.[166,245,1157,1176]" captionTargetBox="[392,1216,195,1121]" captionTargetId="figure-1070@5.[392,1216,195,1121]" captionTargetPageId="5" captionText="Figure 4. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the skull roof. boc, basioccipital?; ds, dorsal scute fragments; ep, epioccipital; ex, exoccipital; fr, frontal; pa, parietal; pf, preepiotic fossa; pt, left (l) and right (r) post-temporals; pto, pterotic; sc, supracleithrum; so, supraoccipital. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807125" httpUri="https://zenodo.org/record/14807125/files/figure.png" pageId="5" pageNumber="180">Fig. 4</figureCitation>
|
||||
). This condition might be a consequence of compression during fossilization. The pre-epiotic fossa is low in comparison to the typical condition in modern clupeiforms, also probably as a consequence of the compression of the skull. The exoccipitals are partially fragmented and displaced (mainly the left one) in relation to the epioccipital, again probably as a consequence of the distortion at the back of the skull. A relatively large, but fragmented, supraoccipital seems to medially separate the parietals, as in the typical clupeiform condition (
|
||||
<figureCitation id="DFB1C8C0FFA35543FB31D427FAD5FAE8" box="[1258,1327,1329,1351]" captionStart="Figure 4" captionStartId="5.[166,245,1157,1176]" captionTargetBox="[392,1216,195,1121]" captionTargetId="figure-1070@5.[392,1216,195,1121]" captionTargetPageId="5" captionText="Figure 4. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the skull roof. boc, basioccipital?; ds, dorsal scute fragments; ep, epioccipital; ex, exoccipital; fr, frontal; pa, parietal; pf, preepiotic fossa; pt, left (l) and right (r) post-temporals; pto, pterotic; sc, supracleithrum; so, supraoccipital. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807125" httpUri="https://zenodo.org/record/14807125/files/figure.png" pageId="5" pageNumber="180">Fig. 4</figureCitation>
|
||||
; Grande, 1985). A temporal foramen, usually located between the parietal and frontal in clupeiforms (
|
||||
<bibRefCitation id="231BA9B4FFA35543FAF5D478FC3FFA0C" author="Lauder VG & Liem KF" pageId="5" pageNumber="180" pagination="95 - 197" refId="ref13632" refString="Lauder VG, Liem KF. 1983. The evolution and interrelationships of the actinopterygian fishes. Bulletin of Museum of Comparative Zoology 150: 95-197." type="journal article" year="1983">Lauder & Liem, 1983</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="231BA9B4FFA35543FC0CD49BFAD1FA0D" author="Di Dario F & de Pinna MC" box="[983,1323,1421,1443]" pageId="5" pageNumber="180" pagination="107 - 123" refId="ref12980" refString="Di Dario F, de Pinna MC. 2006. The supratemporal system and the pattern of ramification of cephalic sensory canals in Denticeps clupeoides (Denticipitoidei, Teleostei): additional evidence for monophyly of Clupeiformes and Clupeoidei. Pap eis <20> Avulsos de Zoologia (S ao ~ Paulo) 46: 107-123." type="journal article" year="2006">Di Dario & de Pinna, 2006</bibRefCitation>
|
||||
), was not identified in the specimen. Its apparent absence in the fossil, however, may be a result of the distortion of the cranium that displaced the right parietal and epioccipital into a more lateroventral position. If this is the case, then the shallow depression at the ventral margin of the parietal, anterior to the epioccipital and between the parietal and the lateral wing of the frontal, is a remnant of the structure.
|
||||
</paragraph>
|
||||
<caption id="13F584CDFFA25542FF4ED77BFDDBF8C5" ID-DOI="http://doi.org/10.5281/zenodo.14807123" ID-Zenodo-Dep="14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="4" pageNumber="179" startId="4.[149,228,1645,1664]" targetBox="[376,1200,606,1613]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="4735D445FFA25542FF4ED77BFDDBF8C5" blockId="4.[149,1427,1645,1899]" pageId="4" pageNumber="179">
|
||||
<emphasis id="75FE0857FFA25542FF4ED77BFE0AF92E" bold="true" box="[149,496,1645,1664]" pageId="4" pageNumber="179">
|
||||
Figure 3.
|
||||
<taxonomicName id="808AAFC6FFA25542FED0D77BFE0AF92E" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[267,496,1645,1664]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="4" pageNumber="179" phylum="Chordata" rank="species" species="nelsoni" status="gen. et sp. nov.">
|
||||
<emphasis id="75FE0857FFA25542FED0D77BFE0AF92E" bold="true" box="[267,496,1645,1664]" italics="true" pageId="4" pageNumber="179">Cynoclupea nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="6ECDB52CFFA25542FE22D778FD51F92F" box="[505,683,1646,1665]" pageId="4" pageNumber="179" rank="species">
|
||||
<emphasis id="75FE0857FFA25542FE22D778FDD3F92F" bold="true" box="[505,553,1646,1665]" pageId="4" pageNumber="179">gen.</emphasis>
|
||||
et
|
||||
<emphasis id="75FE0857FFA25542FD94D778FD51F92F" bold="true" box="[591,683,1646,1665]" pageId="4" pageNumber="179">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
, MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="13F584CDFFA35543FF7DD593FED1FB5E" ID-DOI="http://doi.org/10.5281/zenodo.14807125" ID-Zenodo-Dep="14807125" httpUri="https://zenodo.org/record/14807125/files/figure.png" pageId="5" pageNumber="180" startId="5.[166,245,1157,1176]" targetBox="[392,1216,195,1121]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="4735D445FFA35543FF7DD593FED1FB5E" blockId="5.[166,1444,1157,1264]" pageId="5" pageNumber="180">
|
||||
<emphasis id="75FE0857FFA35543FF7DD593FDFBFB36" bold="true" box="[166,513,1157,1176]" pageId="5" pageNumber="180">
|
||||
Figure 4.
|
||||
<taxonomicName id="808AAFC6FFA35543FEC7D593FDFBFB36" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[284,513,1157,1176]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="5" pageNumber="180" phylum="Chordata" rank="species" species="nelsoni" status="gen. et sp. nov.">
|
||||
<emphasis id="75FE0857FFA35543FEC7D593FDFBFB36" bold="true" box="[284,513,1157,1176]" italics="true" pageId="5" pageNumber="180">Cynoclupea nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="6ECDB52CFFA35543FDD1D593FD46FB36" box="[522,700,1157,1177]" pageId="5" pageNumber="180" rank="species">
|
||||
<emphasis id="75FE0857FFA35543FDD1D593FDC0FB36" bold="true" box="[522,570,1157,1176]" pageId="5" pageNumber="180">gen.</emphasis>
|
||||
et
|
||||
<emphasis id="75FE0857FFA35543FDBBD593FD46FB36" bold="true" box="[608,700,1157,1176]" pageId="5" pageNumber="180">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
, MCP3447-PV. Photograph (A) and interpretative drawing (B) of the skull roof. boc, basioccipital?; ds, dorsal scute fragments; ep, epioccipital; ex, exoccipital; fr, frontal; pa, parietal; pf, preepiotic fossa; pt, left (l) and right (r) post-temporals; pto, pterotic; sc, supracleithrum; so, supraoccipital. Scale bar = 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="4735D445FFA35540FC8DD7B6FB8DFAE9" blockId="5.[829,1444,1329,1901]" lastBlockId="6.[808,1424,199,1718]" lastPageId="6" lastPageNumber="181" pageId="5" pageNumber="180">
|
||||
The right hyomandibula is ventrally dislocated, revealing substantial portions of the prootic, pterotic, exoccipitals, and basioccipital (
|
||||
<figureCitation id="DFB1C8C0FFA35543FB7DD7CBFB10F95D" box="[1190,1258,1757,1779]" captionStart="Figure 5" captionStartId="6.[146,225,1161,1180]" captionTargetBox="[156,753,197,1127]" captionTargetId="figure-677@6.[156,753,197,1127]" captionTargetPageId="6" captionText="Figure 5. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of part of the occipital region in lateral view. acr, accessory temporal sensory canal?; af, auditory fenestra; aor, anterior opening to the chamber of the recessus lateralis; boc, basioccipital?; ex, exoccipital; hm, hyomandibula; in, intercalar; mor, middle opening to the chamber of the recessus lateralis; por, posterior opening to the chamber of the recessus lateralis; pro, prootic; pto, pterotic; sb, opening for swimbladder diverticulum; IX + X, foramen shared by the glossopharyngeal and vagus nerves. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807127" httpUri="https://zenodo.org/record/14807127/files/figure.png" pageId="5" pageNumber="180">Fig. 5</figureCitation>
|
||||
). The prootic is large and somewhat expanded, with ridges and grooves crossing its surface. The ‘strawberry’ appearance, which typically indicates the presence of the prootic bulla in modern clupeiforms (Grande, 1985), could not be detected in
|
||||
<taxonomicName id="808AAFC6FFA05540FE10D730FDB9F994" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[459,579,1573,1595]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="181" phylum="Chordata" rank="species" species="nelsoni">
|
||||
<emphasis id="75FE0857FFA05540FE10D730FDB9F994" box="[459,579,1573,1595]" italics="true" pageId="6" pageNumber="181">C. nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
. However, the slightly inflated morphology of the prootic and the radial fibrous structure of its marginal bony surface suggest the presence of a bulla. A slender and curved lateral commissure for the passage of the hyomandibular trunk nerve is identified in the prootic surface, near the orbit. The pterotic is only partially visible, with a posteriorly directed spine and bearing three clearly distinct openings (
|
||||
<figureCitation id="DFB1C8C0FFA05540FD7DD60CFD10F89E" box="[678,746,1818,1840]" captionStart="Figure 5" captionStartId="6.[146,225,1161,1180]" captionTargetBox="[156,753,197,1127]" captionTargetId="figure-677@6.[156,753,197,1127]" captionTargetPageId="6" captionText="Figure 5. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of part of the occipital region in lateral view. acr, accessory temporal sensory canal?; af, auditory fenestra; aor, anterior opening to the chamber of the recessus lateralis; boc, basioccipital?; ex, exoccipital; hm, hyomandibula; in, intercalar; mor, middle opening to the chamber of the recessus lateralis; por, posterior opening to the chamber of the recessus lateralis; pro, prootic; pto, pterotic; sb, opening for swimbladder diverticulum; IX + X, foramen shared by the glossopharyngeal and vagus nerves. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807127" httpUri="https://zenodo.org/record/14807127/files/figure.png" pageId="6" pageNumber="181">Fig. 5</figureCitation>
|
||||
), which are herein interpreted as the following openings of sensory canals to the chamber of the
|
||||
<emphasis id="75FE0857FFA05540FD47D64EFC7CFF72" box="[668,902,199,1901]" italics="true" pageId="6" pageNumber="181">recessus lateralis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="231BA9B4FFA05540FC47D1D1FBAEFF72" author="Di Dario F" box="[924,1108,199,221]" pageId="6" pageNumber="181" pagination="257 - 270" refId="ref12901" refString="Di Dario F. 2004. Homology between the recessus lateralis and cephalic sensory canals, with the proposition of additional synapomorphies for the Clupeiformes and the Clupeoidei. Zoological Journal of the Linnean Society 141: 257-270." type="journal article" year="2004">Di Dario, 2004</bibRefCitation>
|
||||
): the posterior opening of the
|
||||
<emphasis id="75FE0857FFA05540FC81D1F0FC4CFF55" box="[858,950,230,251]" italics="true" pageId="6" pageNumber="181">recessus</emphasis>
|
||||
(por), which in living clupeiforms connects with the extrascapular and post-temporal canals; the middle opening (mor), which probably connected the
|
||||
<emphasis id="75FE0857FFA05540FC15D054FBD0FEF9" box="[974,1066,322,343]" italics="true" pageId="6" pageNumber="181">recessus</emphasis>
|
||||
of
|
||||
<taxonomicName id="808AAFC6FFA05540FB95D057FB3AFEF8" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[1102,1216,321,342]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="181" phylum="Chordata" rank="species" species="nelsoni">
|
||||
<emphasis id="75FE0857FFA05540FB95D057FB3AFEF8" box="[1102,1216,321,342]" italics="true" pageId="6" pageNumber="181">C. nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
with the preopercular canal; and the anterior opening (aor), which externally would have connected the
|
||||
<emphasis id="75FE0857FFA05540FB39D069FAC4FE3A" box="[1250,1342,383,404]" italics="true" pageId="6" pageNumber="181">recessus</emphasis>
|
||||
chamber with the infraorbital canal, even though only traces of one or more infraorbital bones, not connected or in close proximity with this opening, were identified (
|
||||
<figureCitation id="DFB1C8C0FFA05540FC71D0EEFBFBFDA3" box="[938,1025,504,526]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="6" pageNumber="181">Fig. 2B</figureCitation>
|
||||
). A vertically orientated slit can also be seen on the region of the pterotic that supposedly housed the
|
||||
<emphasis id="75FE0857FFA05540FBC7D320FB82FDE5" box="[1052,1144,566,587]" italics="true" pageId="6" pageNumber="181">recessus</emphasis>
|
||||
chamber, between the middle (mor) and anterior (aor) openings. If this slit was indeed originally present in
|
||||
<taxonomicName id="808AAFC6FFA05540FB77D365FADAFD29" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[1196,1312,626,648]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="181" phylum="Chordata" rank="species" species="nelsoni">
|
||||
<emphasis id="75FE0857FFA05540FB77D365FADAFD29" box="[1196,1312,626,648]" italics="true" pageId="6" pageNumber="181">C. nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
, it might represent the opening of the
|
||||
<emphasis id="75FE0857FFA05540FB53D384FB1EFD09" box="[1160,1252,658,679]" italics="true" pageId="6" pageNumber="181">recessus</emphasis>
|
||||
for the accessory temporal sensory canal (acr), typically present in clupeoids (
|
||||
<bibRefCitation id="231BA9B4FFA05540FC12D3D8FB84FD4D" author="Di Dario F" box="[969,1150,718,740]" pageId="6" pageNumber="181" pagination="257 - 270" refId="ref12901" refString="Di Dario F. 2004. Homology between the recessus lateralis and cephalic sensory canals, with the proposition of additional synapomorphies for the Clupeiformes and the Clupeoidei. Zoological Journal of the Linnean Society 141: 257-270." type="journal article" year="2004">Di Dario, 2004</bibRefCitation>
|
||||
), or the posteroventral opening of the
|
||||
<emphasis id="75FE0857FFA05540FC03D3FBFBCEFCAC" box="[984,1076,749,770]" italics="true" pageId="6" pageNumber="181">recessus</emphasis>
|
||||
(pvor), which so far has been identified only in
|
||||
<taxonomicName id="808AAFC6FFA05540FC28D21AFC91FCEE" authority="(Di Dario & de Pinna, 2006)" baseAuthorityName="Di Dario & de Pinna" baseAuthorityYear="2006" family="Denticipitidae" genus="Denticeps" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="181" phylum="Chordata" rank="species" species="clupeoides">
|
||||
<emphasis id="75FE0857FFA05540FC28D21AFB71FC8E" box="[1011,1163,779,801]" italics="true" pageId="6" pageNumber="181">D. clupeoides</emphasis>
|
||||
(
|
||||
<bibRefCitation id="231BA9B4FFA05540FB40D21DFC98FCEE" author="Di Dario F & de Pinna MC" pageId="6" pageNumber="181" pagination="107 - 123" refId="ref12980" refString="Di Dario F, de Pinna MC. 2006. The supratemporal system and the pattern of ramification of cephalic sensory canals in Denticeps clupeoides (Denticipitoidei, Teleostei): additional evidence for monophyly of Clupeiformes and Clupeoidei. Pap eis <20> Avulsos de Zoologia (S ao ~ Paulo) 46: 107-123." type="journal article" year="2006">Di Dario & de Pinna, 2006</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. The preserved walls of the pterotic have the same fibrous structure as the prootic, suggesting that a bulla was also present in this bone. The right exoccipital (
|
||||
<figureCitation id="DFB1C8C0FFA05540FC58D293FC3CFC35" box="[899,966,901,923]" captionStart="Figure 5" captionStartId="6.[146,225,1161,1180]" captionTargetBox="[156,753,197,1127]" captionTargetId="figure-677@6.[156,753,197,1127]" captionTargetPageId="6" captionText="Figure 5. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of part of the occipital region in lateral view. acr, accessory temporal sensory canal?; af, auditory fenestra; aor, anterior opening to the chamber of the recessus lateralis; boc, basioccipital?; ex, exoccipital; hm, hyomandibula; in, intercalar; mor, middle opening to the chamber of the recessus lateralis; por, posterior opening to the chamber of the recessus lateralis; pro, prootic; pto, pterotic; sb, opening for swimbladder diverticulum; IX + X, foramen shared by the glossopharyngeal and vagus nerves. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807127" httpUri="https://zenodo.org/record/14807127/files/figure.png" pageId="6" pageNumber="181">Fig. 5</figureCitation>
|
||||
) closes posteriorly the occipital region of the neurocranium. Its bulbous basal region bears two openings: the anterior, somewhat funnel-shaped opening, possibly representing the foramen through which the right anterior diverticulum of the swimbladder penetrated the neurocranium; and the posterior opening, which is probably the foramen shared by the glossopharyngeal and vagus nerves (IX + X). The exoccipital expands anteriorly, contacting the prootic and forming the auditory fenestra (
|
||||
<figureCitation id="DFB1C8C0FFA05540FAE6D58FFA7BFB00" box="[1341,1409,1176,1199]" captionStart="Figure 5" captionStartId="6.[146,225,1161,1180]" captionTargetBox="[156,753,197,1127]" captionTargetId="figure-677@6.[156,753,197,1127]" captionTargetPageId="6" captionText="Figure 5. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of part of the occipital region in lateral view. acr, accessory temporal sensory canal?; af, auditory fenestra; aor, anterior opening to the chamber of the recessus lateralis; boc, basioccipital?; ex, exoccipital; hm, hyomandibula; in, intercalar; mor, middle opening to the chamber of the recessus lateralis; por, posterior opening to the chamber of the recessus lateralis; pro, prootic; pto, pterotic; sb, opening for swimbladder diverticulum; IX + X, foramen shared by the glossopharyngeal and vagus nerves. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807127" httpUri="https://zenodo.org/record/14807127/files/figure.png" pageId="6" pageNumber="181">Fig. 5</figureCitation>
|
||||
). The intercalar was also tentatively identified, lying between the exoccipital and the prootic, dorsal to the auditory fenestra. The posterior region of the basioccipital distinctly shows a circular articular surface for the first vertebra (
|
||||
<figureCitation id="DFB1C8C0FFA05540FBFED427FB93FAE9" box="[1061,1129,1329,1351]" captionStart="Figure 5" captionStartId="6.[146,225,1161,1180]" captionTargetBox="[156,753,197,1127]" captionTargetId="figure-677@6.[156,753,197,1127]" captionTargetPageId="6" captionText="Figure 5. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of part of the occipital region in lateral view. acr, accessory temporal sensory canal?; af, auditory fenestra; aor, anterior opening to the chamber of the recessus lateralis; boc, basioccipital?; ex, exoccipital; hm, hyomandibula; in, intercalar; mor, middle opening to the chamber of the recessus lateralis; por, posterior opening to the chamber of the recessus lateralis; pro, prootic; pto, pterotic; sb, opening for swimbladder diverticulum; IX + X, foramen shared by the glossopharyngeal and vagus nerves. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807127" httpUri="https://zenodo.org/record/14807127/files/figure.png" pageId="6" pageNumber="181">Fig. 5</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="13F584CDFFA05540FF49D59FFE5DFA70" ID-DOI="http://doi.org/10.5281/zenodo.14807127" ID-Zenodo-Dep="14807127" httpUri="https://zenodo.org/record/14807127/files/figure.png" pageId="6" pageNumber="181" startId="6.[146,225,1161,1180]" targetBox="[156,753,197,1127]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="4735D445FFA05540FF49D59FFE5DFA70" blockId="6.[146,760,1161,1502]" pageId="6" pageNumber="181">
|
||||
<emphasis id="75FE0857FFA05540FF49D59FFE01FB32" bold="true" box="[146,507,1161,1180]" pageId="6" pageNumber="181">
|
||||
Figure 5.
|
||||
<taxonomicName id="808AAFC6FFA05540FED3D59FFE01FB32" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[264,507,1161,1180]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="181" phylum="Chordata" rank="species" species="nelsoni" status="gen. et sp. nov.">
|
||||
<emphasis id="75FE0857FFA05540FED3D59FFE01FB32" bold="true" box="[264,507,1161,1180]" italics="true" pageId="6" pageNumber="181">Cynoclupea nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="6ECDB52CFFA05540FDC8D59FFD08FB32" box="[531,754,1161,1180]" pageId="6" pageNumber="181" rank="species">
|
||||
<emphasis id="75FE0857FFA05540FDC8D59FFDB9FB32" bold="true" box="[531,579,1161,1180]" pageId="6" pageNumber="181">gen.</emphasis>
|
||||
et
|
||||
<emphasis id="75FE0857FFA05540FD5CD59FFD08FB32" bold="true" box="[647,754,1161,1180]" pageId="6" pageNumber="181">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
, MCP3447-PV. Photograph (A) and interpretative drawing (B) of part of the occipital region in lateral view. acr, accessory temporal sensory canal?; af, auditory fenestra; aor, anterior opening to the chamber of the
|
||||
<emphasis id="75FE0857FFA05540FDB8D5E9FF42FA80" italics="true" pageId="6" pageNumber="181">recessus lateralis</emphasis>
|
||||
; boc, basioccipital?; ex, exoccipital; hm, hyomandibula; in, intercalar; mor, middle opening to the chamber of the
|
||||
<emphasis id="75FE0857FFA05540FF38D440FE67FAC7" box="[227,413,1366,1385]" italics="true" pageId="6" pageNumber="181">recessus lateralis</emphasis>
|
||||
; por, posterior opening to the chamber of the
|
||||
<emphasis id="75FE0857FFA05540FEE7D462FE0BFA28" box="[316,497,1395,1414]" italics="true" pageId="6" pageNumber="181">recessus lateralis</emphasis>
|
||||
; pro, prootic; pto, pterotic; sb, opening for swimbladder diverticulum; IX + X, foramen shared by the glossopharyngeal and vagus nerves. Scale bar = 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="4735D445FFA05540FC9AD446FAEEF918" blockId="6.[808,1424,199,1718]" pageId="6" pageNumber="181">
|
||||
The parasphenoid is preserved as an edentulous and straight bone for most of its length (
|
||||
<figureCitation id="DFB1C8C0FFA05540FAC3D478FA98FA2A" box="[1304,1378,1390,1412]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="6" pageNumber="181">Figs 2</figureCitation>
|
||||
,
|
||||
<figureCitation id="DFB1C8C0FFA05540FAA9D478FA7BFA2A" box="[1394,1409,1390,1412]" captionStart="Figure 6" captionStartId="7.[166,245,1221,1240]" captionTargetBox="[176,774,195,1185]" captionTargetId="figure-1477@7.[176,774,195,1185]" captionTargetPageId="7" captionText="Figure 6. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the snout region. bf, undetermined bony fragment; d, dentary; dt, dentary teeth; ec, ectopterygoid; en, endopterygoid; fr, frontal; mc, mandibular sensory canal; me, mesethmoid; n, nasal; pl, palatine; pm, premaxilla; ps, parasphenoid; vo, vomer. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14807129" httpUri="https://zenodo.org/record/14807129/files/figure.png" pageId="6" pageNumber="181">6</figureCitation>
|
||||
); its anterior tip is slightly curved upwards, meeting the vomer. An ascending arm in its posterior region contacts the prootic. A foramen for the internal carotid artery is also preserved in the parasphenoid, in the region of contact with the prootic. Owing to the state of preservation, it is not clear if the parasphenoid projects posteriorly beyond the basioccipital, as in most modern clupeoids. Bony remains tentatively identified as parts of the lateral ethmoid contact the anterior portion of the parasphenoid shaft.
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFA05540FCF3D7CBFB4BF95D" blockId="6.[808,1423,1757,1901]" box="[808,1201,1757,1779]" pageId="6" pageNumber="181">
|
||||
<emphasis id="75FE0857FFA05540FCF3D7CBFB4BF95D" box="[808,1201,1757,1779]" italics="true" pageId="6" pageNumber="181">Palatoquadrate and suspensorium</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFA05541FCF3D7EAFBB2FCAD" blockId="6.[808,1423,1757,1901]" lastBlockId="7.[829,1444,199,771]" lastPageId="7" lastPageNumber="182" pageId="6" pageNumber="181">
|
||||
Only portions of the palatoquadrate and the suspensorium can be identified in the fossil, with the quadrate
|
||||
<emphasis id="75FE0857FFA05540FC54D62AFC61F8E0" box="[911,923,1852,1870]" italics="true" pageId="6" pageNumber="181">–</emphasis>
|
||||
symplectic complex as one of the most distinctly preserved features (
|
||||
<figureCitation id="DFB1C8C0FFA05540FBB2D641FB57F8C2" box="[1129,1197,1879,1901]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="6" pageNumber="181">Fig. 3</figureCitation>
|
||||
). The shape of the The proximal region of the hyomandibula is expanded in a fan-shaped structure, forming a straight margin with no differentiated articular heads for the sphenotic and pterotic (
|
||||
<figureCitation id="DFB1C8C0FFA15541FB7CD034FB16FE96" box="[1191,1260,290,312]" captionStart="Figure 5" captionStartId="6.[146,225,1161,1180]" captionTargetBox="[156,753,197,1127]" captionTargetId="figure-677@6.[156,753,197,1127]" captionTargetPageId="6" captionText="Figure 5. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of part of the occipital region in lateral view. acr, accessory temporal sensory canal?; af, auditory fenestra; aor, anterior opening to the chamber of the recessus lateralis; boc, basioccipital?; ex, exoccipital; hm, hyomandibula; in, intercalar; mor, middle opening to the chamber of the recessus lateralis; por, posterior opening to the chamber of the recessus lateralis; pro, prootic; pto, pterotic; sb, opening for swimbladder diverticulum; IX + X, foramen shared by the glossopharyngeal and vagus nerves. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807127" httpUri="https://zenodo.org/record/14807127/files/figure.png" pageId="7" pageNumber="182">Fig. 5</figureCitation>
|
||||
). Ventrally, the hyomandibula narrows abruptly into a vertical limb directed towards the posterior margin of the quadrate and the symplectic, but its length could not be determined because its distal third is broken (
|
||||
<figureCitation id="DFB1C8C0FFA15541FAC5D08BFA80FE1C" box="[1310,1402,413,435]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="7" pageNumber="182">Figs 2A</figureCitation>
|
||||
,
|
||||
<figureCitation id="DFB1C8C0FFA15541FA5CD08BFA6CFE1C" box="[1415,1430,413,434]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="7" pageNumber="182">3</figureCitation>
|
||||
). The hyomandibula is slightly inclined posteriorly, being almost vertically orientated. Parts of a toothless endopterygoid (
|
||||
<figureCitation id="DFB1C8C0FFA15541FC36D0EEFBCBFDA0" box="[1005,1073,504,526]" captionStart="Figure 6" captionStartId="7.[166,245,1221,1240]" captionTargetBox="[176,774,195,1185]" captionTargetId="figure-1477@7.[176,774,195,1185]" captionTargetPageId="7" captionText="Figure 6. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the snout region. bf, undetermined bony fragment; d, dentary; dt, dentary teeth; ec, ectopterygoid; en, endopterygoid; fr, frontal; mc, mandibular sensory canal; me, mesethmoid; n, nasal; pl, palatine; pm, premaxilla; ps, parasphenoid; vo, vomer. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14807129" httpUri="https://zenodo.org/record/14807129/files/figure.png" pageId="7" pageNumber="182">Fig. 6</figureCitation>
|
||||
) and possibly the metapterygoid are visible in the orbit region (
|
||||
<figureCitation id="DFB1C8C0FFA15541FB44D301FB0CFD82" box="[1183,1270,535,557]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="7" pageNumber="182">Fig. 2B</figureCitation>
|
||||
). However, the anterior portion of the metapterygoid and the posterior region of the endopterygoid are not preserved and, therefore, their topographical relationships with other bones of the suspensorium could not be determined. Only the anterior tip of the palatine is visible, with three massive conical teeth slightly orientated backwards (
|
||||
<figureCitation id="DFB1C8C0FFA15541FC1ED3FBFBDBFCAC" box="[965,1057,749,771]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="7" pageNumber="182">Figs 2B</figureCitation>
|
||||
,
|
||||
<figureCitation id="DFB1C8C0FFA15541FBF0D3FBFBC1FCAD" box="[1067,1083,749,771]" captionStart="Figure 6" captionStartId="7.[166,245,1221,1240]" captionTargetBox="[176,774,195,1185]" captionTargetId="figure-1477@7.[176,774,195,1185]" captionTargetPageId="7" captionText="Figure 6. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the snout region. bf, undetermined bony fragment; d, dentary; dt, dentary teeth; ec, ectopterygoid; en, endopterygoid; fr, frontal; mc, mandibular sensory canal; me, mesethmoid; n, nasal; pl, palatine; pm, premaxilla; ps, parasphenoid; vo, vomer. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14807129" httpUri="https://zenodo.org/record/14807129/files/figure.png" pageId="7" pageNumber="182">6</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="13F584CDFFA15541FF7DD5D3FD64FA26" ID-DOI="http://doi.org/10.5281/zenodo.14807129" ID-Zenodo-Dep="14807129" httpUri="https://zenodo.org/record/14807129/files/figure.png" pageId="7" pageNumber="182" startId="7.[166,245,1221,1240]" targetBox="[176,774,195,1185]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="4735D445FFA15541FF7DD5D3FD64FA26" blockId="7.[166,781,1221,1416]" pageId="7" pageNumber="182">
|
||||
<emphasis id="75FE0857FFA15541FF7DD5D3FDEAFB76" bold="true" box="[166,528,1221,1240]" pageId="7" pageNumber="182">
|
||||
Figure 6.
|
||||
<taxonomicName id="808AAFC6FFA15541FEC7D5D3FDEAFB76" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[284,528,1221,1240]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="182" phylum="Chordata" rank="species" species="nelsoni" status="gen. et sp. nov.">
|
||||
<emphasis id="75FE0857FFA15541FEC7D5D3FDEAFB76" bold="true" box="[284,528,1221,1240]" italics="true" pageId="7" pageNumber="182">Cynoclupea nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="6ECDB52CFFA15541FDF3D5D3FCFCFB76" box="[552,774,1221,1240]" pageId="7" pageNumber="182" rank="species">
|
||||
<emphasis id="75FE0857FFA15541FDF3D5D3FDA2FB76" bold="true" box="[552,600,1221,1240]" pageId="7" pageNumber="182">gen.</emphasis>
|
||||
et
|
||||
<emphasis id="75FE0857FFA15541FD47D5D3FCFCFB76" bold="true" box="[668,774,1221,1240]" pageId="7" pageNumber="182">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
, MCP3447-PV. Photograph (A) and interpretative drawing (B) of the snout region. bf, undetermined bony fragment; d, dentary; dt, dentary teeth; ec, ectopterygoid; en, endopterygoid; fr, frontal; mc, mandibular sensory canal; me, mesethmoid; n, nasal; pl, palatine; pm, premaxilla; ps, parasphenoid; vo, vomer. Scale bar = 3 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="4735D445FFA15541FF7DD4BDFEBBF8C2" blockId="7.[166,781,1451,1900]" pageId="7" pageNumber="182">
|
||||
endochondral portion of the quadrate is similar to that of an equilateral triangle, with a roughly straight dorsal margin. The anterior margin of the quadrate has a narrow laminar outgrowth, where a fragment of the slender ectopterygoid is still firmly attached (
|
||||
<figureCitation id="DFB1C8C0FFA15541FEC5D755FE99F9F7" box="[286,355,1603,1625]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="7" pageNumber="182">Fig. 3</figureCitation>
|
||||
). A short, splint-like posterior process into which the symplectic inserts is also clearly discernible in the quadrate, as in the typical condition present in teleosts in general, including modern clupeiforms (
|
||||
<bibRefCitation id="231BA9B4FFA15541FE9FD7A8FD7EF97D" author="Arratia G & Schultze HP" box="[324,644,1726,1748]" pageId="7" pageNumber="182" pagination="1 - 81" refId="ref12452" refString="Arratia G, Schultze HP. 1991. Palatoquadrate and its ossifications: development and homology within osteichthyans. Journal of Morphology 208: 1-81." type="journal article" year="1991">Arratia & Schultze, 1991</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="231BA9B4FFA15541FD41D7A8FF1AF95F" author="Di Dario F" pageId="7" pageNumber="182" pagination="363 - 383" refId="ref12942" refString="Di Dario F. 2009. Chirocentrids as engrauloids: evidence from suspensorium, branchial arches, and infraorbital bones (Clupeomorpha, Teleostei). Zoological Journal of the Linnean Society 156: 363-383." type="journal article" year="2009">Di Dario, 2009</bibRefCitation>
|
||||
). The condyle of articulation of the quadrate with the lower jaw is practically ventrally orientated. The symplectic is elongated and nearly straight. Its dorsal margin is in line with the dorsal margin of the quadrate.
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFA15541FCE6D23CFC82FC91" blockId="7.[829,1444,810,1901]" box="[829,888,810,831]" pageId="7" pageNumber="182">
|
||||
<emphasis id="75FE0857FFA15541FCE6D23CFC82FC91" box="[829,888,810,831]" italics="true" pageId="7" pageNumber="182">Jaws</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFA15541FCE6D25EFB20FAE9" blockId="7.[829,1444,810,1901]" pageId="7" pageNumber="182">
|
||||
The jaws are long and relatively straight. Teeth are present on the premaxillae, maxillae, and dentaries. A substantial portion of what is herein identified as the right and left premaxillae is preserved, but the bones are displaced from their original position. Overall, the preserved fragments of the premaxillae indicate that the bone was nearly triangular and slightly curved in lateral view. The left premaxilla is turned over the mesethmoid, in such a way that its ventral margin is facing up (
|
||||
<figureCitation id="DFB1C8C0FFA15541FC9ED56CFC71FB3E" box="[837,907,1146,1168]" captionStart="Figure 6" captionStartId="7.[166,245,1221,1240]" captionTargetBox="[176,774,195,1185]" captionTargetId="figure-1477@7.[176,774,195,1185]" captionTargetPageId="7" captionText="Figure 6. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the snout region. bf, undetermined bony fragment; d, dentary; dt, dentary teeth; ec, ectopterygoid; en, endopterygoid; fr, frontal; mc, mandibular sensory canal; me, mesethmoid; n, nasal; pl, palatine; pm, premaxilla; ps, parasphenoid; vo, vomer. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14807129" httpUri="https://zenodo.org/record/14807129/files/figure.png" pageId="7" pageNumber="182">Fig. 6</figureCitation>
|
||||
). The right premaxilla, by contrast, is nearly totally preserved, and bears a dorsally orientated protuberance that seems to be a rudimentary ascending process (
|
||||
<figureCitation id="DFB1C8C0FFA15541FBFDD5C3FB91FB45" box="[1062,1131,1237,1259]" captionStart="Figure 6" captionStartId="7.[166,245,1221,1240]" captionTargetBox="[176,774,195,1185]" captionTargetId="figure-1477@7.[176,774,195,1185]" captionTargetPageId="7" captionText="Figure 6. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the snout region. bf, undetermined bony fragment; d, dentary; dt, dentary teeth; ec, ectopterygoid; en, endopterygoid; fr, frontal; mc, mandibular sensory canal; me, mesethmoid; n, nasal; pl, palatine; pm, premaxilla; ps, parasphenoid; vo, vomer. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14807129" httpUri="https://zenodo.org/record/14807129/files/figure.png" pageId="7" pageNumber="182">Fig. 6</figureCitation>
|
||||
). The preserved dentition of the bone is made up of a few needle-shaped, irregularly sized teeth distributed along the oral border, apparently in a single row.
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFA1554EFC8DD446FDE3FF55" blockId="7.[829,1444,810,1901]" lastBlockId="8.[145,760,199,771]" lastPageId="8" lastPageNumber="183" pageId="7" pageNumber="182">
|
||||
The maxilla is slender and very long, covering laterally the posterior region of the dentary (
|
||||
<figureCitation id="DFB1C8C0FFA15541FA89D478FA6DFA2D" box="[1362,1431,1390,1412]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="7" pageNumber="182">Fig. 2</figureCitation>
|
||||
). The bone has a compressed baseball bat shape, deeper posteriorly and narrowing anteriorly to form a distinct head. Its oral margin is covered with a row of small, closely placed, and equally sized teeth, resulting in a comb-shaped structure as in the typical condition amongst modern clupeiforms. A fragment of bone attached to the dorsal margin of the maxilla is herein interpreted as part of the supramaxilla (
|
||||
<figureCitation id="DFB1C8C0FFA15541FC73D797FC17F939" box="[936,1005,1665,1687]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="7" pageNumber="182">Fig. 3</figureCitation>
|
||||
). Another fragment of a flat, somewhat elongated, structure with a rounded posterior margin is also present on the counterpart of the fossil approximately in the region where the posterior end of the upper maxilla was probably located (
|
||||
<figureCitation id="DFB1C8C0FFA15541FC9ED60CFC67F881" box="[837,925,1818,1840]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="7" pageNumber="182">Fig. 2B</figureCitation>
|
||||
). This fragment is also interpreted as a supramaxilla. If this identification is correct, and given the relatively well-developed degree of development of the structure, it is presumed that
|
||||
<taxonomicName id="808AAFC6FFAE554EFF49D1F0FEF9FF54" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[146,259,229,251]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="8" pageNumber="183" phylum="Chordata" rank="species" species="nelsoni">
|
||||
<emphasis id="75FE0857FFAE554EFF49D1F0FEF9FF54" box="[146,259,229,251]" italics="true" pageId="8" pageNumber="183">C. nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
had two supramaxillae.
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFAE554EFF71D012FE39FCAC" blockId="8.[145,760,199,771]" pageId="8" pageNumber="183">
|
||||
Based on the preserved parts, the lower jaw is long and robust, high posteriorly, and tapering towards the symphysis. Differing from the condition described for the maxilla, the dentary dentition is very conspicuous and well developed: two enlarged and backward-curved teeth at the anterior tip of the dentary are followed by at least five smaller and fully preserved teeth of similar shapes. In addition, a series of bony protuberances in the dentary border suggests that regularly spaced and decreasing in size teeth were present along most, if not all, the oral margin of the bone (
|
||||
<figureCitation id="DFB1C8C0FFAE554EFE52D342FE37FDC7" box="[393,461,596,618]" captionStart="Figure 7" captionStartId="8.[146,225,1821,1840]" captionTargetBox="[373,1197,958,1790]" captionTargetId="figure-426@8.[373,1197,958,1791]" captionTargetPageId="8" captionText="Figure 7. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of part of the upper and lower jaws in mesial view. d, dentary; dt, dentary teeth; me, mesethmoid; mx, maxilla. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14807131" httpUri="https://zenodo.org/record/14807131/files/figure.png" pageId="8" pageNumber="183">Fig. 7</figureCitation>
|
||||
). The anterior portion of the mandibular sensory canal is included in the ventral third of the dentary (
|
||||
<figureCitation id="DFB1C8C0FFAE554EFE17D387FDEFFD08" box="[460,533,657,679]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="8" pageNumber="183">Figs 3</figureCitation>
|
||||
,
|
||||
<figureCitation id="DFB1C8C0FFAE554EFDFCD387FDCDFD09" box="[551,567,657,679]" captionStart="Figure 6" captionStartId="7.[166,245,1221,1240]" captionTargetBox="[176,774,195,1185]" captionTargetId="figure-1477@7.[176,774,195,1185]" captionTargetPageId="7" captionText="Figure 6. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the snout region. bf, undetermined bony fragment; d, dentary; dt, dentary teeth; ec, ectopterygoid; en, endopterygoid; fr, frontal; mc, mandibular sensory canal; me, mesethmoid; n, nasal; pl, palatine; pm, premaxilla; ps, parasphenoid; vo, vomer. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14807129" httpUri="https://zenodo.org/record/14807129/files/figure.png" pageId="8" pageNumber="183">6</figureCitation>
|
||||
). A fragment of the posterior portion of the anguloarticular is recognized in the region of articulation of the lower jaw with the quadrate (
|
||||
<figureCitation id="DFB1C8C0FFAE554EFEAAD3FBFE4CFCAC" box="[369,438,749,771]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="8" pageNumber="183">Fig. 2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFAE554EFF4AD23CFD93FC91" blockId="8.[145,760,810,893]" box="[145,617,810,831]" pageId="8" pageNumber="183">
|
||||
<emphasis id="75FE0857FFAE554EFF4AD23CFD93FC91" box="[145,617,810,831]" italics="true" pageId="8" pageNumber="183">Opercular and hyobranchial apparatuses</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFAE554FFF4AD25EFF28FF55" blockId="8.[145,760,810,893]" lastBlockId="9.[166,781,199,740]" lastPageId="9" lastPageNumber="184" pageId="8" pageNumber="183">
|
||||
Only fragments of the opercular and hyobranchial apparatuses are preserved, and most components of these systems are displaced from their presumed original positions. The preserved margins and surfaces of the opercular apparatus are smooth. The preopercle is probably the best-preserved bone of the series (
|
||||
<figureCitation id="DFB1C8C0FFAE554EFC6FD057FC02FEF8" box="[948,1016,321,343]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="8" pageNumber="183">Fig. 3</figureCitation>
|
||||
). It has a crescent-shaped structure, as in the typical condition amongst modern members of the
|
||||
<taxonomicName id="808AAFC6FFAE554EFC2FD068FB8AFE3A" box="[1012,1136,382,404]" kingdom="Animalia" order="Clupeiformes" pageId="8" pageNumber="175" phylum="Chordata" rank="subOrder" subOrder="Clupeoidei">Clupeoidei</taxonomicName>
|
||||
. The bone is longitudinally crossed by the preopercular sensory canal, which bears some distinct openings. The angle of the preopercle is conspicuous, and caudally projected. A small remnant of what is apparently a flat interopercle is located along the ventral border of the preopercle (
|
||||
<figureCitation id="DFB1C8C0FFAE554EFBD0D323FBAAFDE5" box="[1035,1104,565,587]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="8" pageNumber="183">Fig. 3</figureCitation>
|
||||
). The largest fragment of the opercle (
|
||||
<figureCitation id="DFB1C8C0FFAE554EFC16D342FBD9FDC7" box="[973,1059,596,618]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="8" pageNumber="183">Fig. 2B</figureCitation>
|
||||
) is roughly shaped like an equilateral triangle. However, other opercle fragments, more dorsally located (
|
||||
<figureCitation id="DFB1C8C0FFAE554EFB40D387FB0DFD08" box="[1179,1271,657,679]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="8" pageNumber="183">Figs 2A</figureCitation>
|
||||
,
|
||||
<figureCitation id="DFB1C8C0FFAE554EFAD2D387FAEDFD08" box="[1289,1303,657,678]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="8" pageNumber="183">3</figureCitation>
|
||||
), indicate that the opercle of
|
||||
<taxonomicName id="808AAFC6FFAE554EFBC5D3A6FB6DFD6B" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[1054,1175,688,709]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="8" pageNumber="183" phylum="Chordata" rank="species" species="nelsoni">
|
||||
<emphasis id="75FE0857FFAE554EFBC5D3A6FB6DFD6B" box="[1054,1175,688,709]" italics="true" pageId="8" pageNumber="183">C. nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
was large, approximately square-shaped, slightly wider than deep, and with a somewhat concave ventral margin. The subopercle is also preserved, but was displaced dorsally during fossilization (
|
||||
<figureCitation id="DFB1C8C0FFAE554EFB71D23CFB15FC91" box="[1194,1263,810,832]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="8" pageNumber="183">Fig. 3</figureCitation>
|
||||
). The bone has a straight dorsal margin and a well projected and sharp anterodorsal ascending process, where the adductor mandibular probably attached during life.
|
||||
</paragraph>
|
||||
<caption id="13F584CDFFAE554EFF49D60BFEECF8C5" ID-DOI="http://doi.org/10.5281/zenodo.14807131" ID-Zenodo-Dep="14807131" httpUri="https://zenodo.org/record/14807131/files/figure.png" pageId="8" pageNumber="183" startId="8.[146,225,1821,1840]" targetBox="[373,1197,958,1790]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="4735D445FFAE554EFF49D60BFEECF8C5" blockId="8.[146,1423,1820,1899]" pageId="8" pageNumber="183">
|
||||
<emphasis id="75FE0857FFAE554EFF49D60BFE11F881" bold="true" box="[146,491,1820,1840]" pageId="8" pageNumber="183">
|
||||
Figure 7.
|
||||
<taxonomicName id="808AAFC6FFAE554EFED3D60AFE11F881" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[264,491,1820,1839]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="8" pageNumber="183" phylum="Chordata" rank="species" species="nelsoni" status="gen. et sp. nov.">
|
||||
<emphasis id="75FE0857FFAE554EFED3D60AFE11F881" bold="true" box="[264,491,1820,1839]" italics="true" pageId="8" pageNumber="183">Cynoclupea nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="6ECDB52CFFAE554EFE28D60BFD59F89E" box="[499,675,1821,1840]" pageId="8" pageNumber="183" rank="species">
|
||||
<emphasis id="75FE0857FFAE554EFE28D60BFDDEF89E" bold="true" box="[499,548,1821,1840]" pageId="8" pageNumber="183">gen.</emphasis>
|
||||
et
|
||||
<emphasis id="75FE0857FFAE554EFD92D60BFD59F89E" bold="true" box="[585,675,1821,1840]" pageId="8" pageNumber="183">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
, MCP3447-PV. Photograph (A) and interpretative drawing (B) of part of the upper and lower jaws in mesial view. d, dentary; dt, dentary teeth; me, mesethmoid; mx, maxilla. Scale bar = 3 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="4735D445FFAF554FFF64D012FD2CFD4A" blockId="9.[166,781,199,740]" pageId="9" pageNumber="184">
|
||||
Scattered remains of well-ossified gill arches and rakers are preserved in abundance. However, precise description of the structures and assessment of homologies are prevented by the state of preservation. Overall, elements identified as gill rakers (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFF75D08BFF0BFE1C" box="[174,241,413,435]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="9" pageNumber="184">Fig. 3</figureCitation>
|
||||
) are relatively numerous and well ossified, somewhat slender, closely packed, and apparently toothless; these last two features might result from the fossilization. An approximately triangle-shaped and relatively well-developed structure located in the posterior region of the branchial chamber is tentatively identified as an edentulous fifth ceratobranchial (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFEE8D364FE82FD26" box="[307,376,626,648]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="9" pageNumber="184">Fig. 3</figureCitation>
|
||||
). No evidence of epibranchial organs could be detected. Five branchiostegal rays are preserved, the two anterior ones being tubular, and the three more posterior, broad and smooth.
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFAF554FFF7DD21DFE4EFC8E" blockId="9.[166,781,779,1901]" box="[166,436,779,800]" pageId="9" pageNumber="184">
|
||||
<emphasis id="75FE0857FFAF554FFF7DD21DFE4EFC8E" box="[166,436,779,800]" italics="true" pageId="9" pageNumber="184">Pectoral girdle and fins</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFAF554FFF7DD23CFD11FAA4" blockId="9.[166,781,779,1901]" pageId="9" pageNumber="184">
|
||||
Parts of the pectoral girdles are damaged or concealed by vertebral elements, but the main components of the left and right sides are visible. Both post-temporals are near to their original position, and can be observed in medial (left post-temporal) and lateral (right post-temporal) views (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFD5FD2D5FD1BFC76" box="[644,737,963,985]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="9" pageNumber="184">Figs 2A</figureCitation>
|
||||
,
|
||||
<figureCitation id="DFB1C8C0FFAF554FFD2BD2D5FD05FC76" box="[752,767,963,984]" captionStart="Figure 7" captionStartId="8.[146,225,1821,1840]" captionTargetBox="[373,1197,958,1790]" captionTargetId="figure-426@8.[373,1197,958,1791]" captionTargetPageId="8" captionText="Figure 7. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of part of the upper and lower jaws in mesial view. d, dentary; dt, dentary teeth; me, mesethmoid; mx, maxilla. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14807131" httpUri="https://zenodo.org/record/14807131/files/figure.png" pageId="9" pageNumber="184">7</figureCitation>
|
||||
). They are well-developed, flat bones, with a roughly rounded posterior outline and a large anterior arm that in life would have articulated with the epioccipital. A fracture in the left post-temporal reveals a segment of the lateral line extending through its posterior third. The proximal portion of the right supracleithrum is preserved, although partially concealed by postcranial remains. The bone seems to be relatively long and plate-like, with its dorsal region articulating with the corresponding post-temporal.
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFAF554FFF64D405FBF7FE1C" blockId="9.[166,781,779,1901]" lastBlockId="9.[829,1444,199,771]" pageId="9" pageNumber="184">
|
||||
The portion of the left and right pectoral girdles ventral to the post-temporal and supracleithral elements is slightly dislodged into a more posteroventral position than would have occurred in life (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFF75D49BFF09FA0C" box="[174,243,1421,1443]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="9" pageNumber="184">Fig. 3</figureCitation>
|
||||
). Both cleithra are preserved, but the left one is partially concealed by its right counterpart (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFF75D4DCFF08FA71" box="[174,242,1482,1504]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="9" pageNumber="184">Fig. 3</figureCitation>
|
||||
), which is therefore described herein. The cleithrum is developed, being the largest element in the pectoral girdle of
|
||||
<taxonomicName id="808AAFC6FFAF554FFE7BD711FDE9F9B2" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[416,531,1543,1564]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="9" pageNumber="184" phylum="Chordata" rank="species" species="nelsoni">
|
||||
<emphasis id="75FE0857FFAF554FFE7BD711FDE9F9B2" box="[416,531,1543,1564]" italics="true" pageId="9" pageNumber="184">C. nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
. The proximal region of the cleithrum is approximately vertical, but the main vertical shaft of the bone curves anteriorly as it descends towards the ventral region of the specimen. The cleithrum seems to be well expanded posteriorly above the level of insertion of the pectoral fin. The scapula, which should be located in this region, could not be identified in the fossil. The lateral margin of the ventral third of the cleithrum projects laterally and partially covers the coracoid, forming the concavity where the pectoral-fin musculature originates in extant teleosts. Both coracoids, right and left, are partially visible (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFBB6D1D1FB4BFF72" box="[1133,1201,199,221]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="9" pageNumber="184">Fig. 3</figureCitation>
|
||||
); they are rectangular, flat, with straight ventral margins. The mesocoracoid could not be identified, but the bone is probably concealed behind the pectoral-fin girdle elements, especially the right cleithrum. Two laminar postcleithra are partially visible behind the dorsal third of the cleithrum, the posterior one being more elongated (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFC67D08BFBFAFE1C" box="[956,1024,413,435]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="9" pageNumber="184">Fig. 3</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFAF554FFC8DD0AAFB7BFCAC" blockId="9.[829,1444,199,771]" pageId="9" pageNumber="184">
|
||||
Between 17 and 18 pectoral-fin rays are preserved in the specimen, most notably in the right side, the outermost ray being the most developed. Ventrally to these rays, a few bases of the left pectoral-fin rays can also be seen (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFBD7D320FBAAFDE5" box="[1036,1104,566,588]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="9" pageNumber="184">Fig. 3</figureCitation>
|
||||
). As only the proximal half of most rays are preserved, it is not possible to determine the full extent of the pectoral fin, but the structure is obviously well developed. Some skeletal support elements for the pectoral fin (distal and/or proximal radials) are preserved in the region ventral to the bases of the pectoral-fin rays (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFBEBD3FBFB8EFCAC" box="[1072,1140,749,771]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="9" pageNumber="184">Fig. 3</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFAF554FFCE6D23CFAFDFCEE" blockId="9.[829,1444,810,1902]" box="[829,1287,810,832]" pageId="9" pageNumber="184">
|
||||
<emphasis id="75FE0857FFAF554FFCE6D23CFAFDFCEE" box="[829,1287,810,832]" italics="true" pageId="9" pageNumber="184">Axial skeleton and associated structures</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFAF554FFCE6D25FFAAAF9B3" blockId="9.[829,1444,810,1902]" pageId="9" pageNumber="184">
|
||||
The anterior-most 11 vertebrae are preserved, and therefore the description below refers to this portion of the axial skeleton (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFBE9D290FB81FC35" box="[1074,1147,902,924]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="9" pageNumber="184">Figs 3</figureCitation>
|
||||
,
|
||||
<figureCitation id="DFB1C8C0FFAF554FFB53D290FB6DFC35" box="[1160,1175,902,923]" captionStart="Figure 7" captionStartId="8.[146,225,1821,1840]" captionTargetBox="[373,1197,958,1790]" captionTargetId="figure-426@8.[373,1197,958,1791]" captionTargetPageId="8" captionText="Figure 7. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of part of the upper and lower jaws in mesial view. d, dentary; dt, dentary teeth; me, mesethmoid; mx, maxilla. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.14807131" httpUri="https://zenodo.org/record/14807131/files/figure.png" pageId="9" pageNumber="184">7</figureCitation>
|
||||
). The vertebral centrae are slightly deeper (3.10 mm) than long (2.94 mm), thick and laterally sculptured with at least four longitudinal ridges. They are completely ossified with a strongly constricted notochord, in such a way that the notochordal pit is not discernible. The neural arches are autogenous, and the corresponding pairs of neural spines are also unfused amongst themselves. A few rod-like bony structures preserved in the dorsal region of the centrae between approximately the sixth and the last preserved vertebrae are tentatively identified as epineurals, their bases apparently fused with each corresponding neural arch (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFC5CD405FC36FA86" box="[903,972,1299,1321]" captionStart="Figure 8" captionStartId="10.[146,225,1335,1354]" captionTargetBox="[369,1193,197,1301]" captionTargetId="figure-281@10.[369,1193,197,1301]" captionTargetPageId="10" captionText="Figure 8. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the predorsal region. cl, cleithrum; epn, epineurals; na, neural arches; ns, neural spines; pt, post-temporal; r, ribs; sc, supracleithrum; sn, supraneurals; v1–10, vertebrae 1 to 10. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807133" httpUri="https://zenodo.org/record/14807133/files/figure.png" pageId="9" pageNumber="184">Fig. 8</figureCitation>
|
||||
). The pleural ribs are strongly developed and roughly rod-like in cross-section. Their proximal region is curved proximally, each rib having an expanded and straight head for articulation with the centrum. Bony rod-like structures tentatively identified as epipleurals are present amongst the pleural ribs, but it is not clear if these elements were fused to ribs as in the Clupeoidea (
|
||||
<bibRefCitation id="231BA9B4FFAF554FFC9ED71EFB7CF9B3" author="Patterson C & Johnson GD" box="[837,1158,1543,1565]" pageId="9" pageNumber="184" pagination="1 - 85" refId="ref14292" refString="Patterson C, Johnson GD. 1995. The intermuscular bones and ligaments of teleostean fishes. Smithsonian Contributions to Zoology 559: 1-85." type="journal article" year="1995">Patterson & Johnson, 1995</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="231BA9B4FFAF554FFB4FD711FABEF9B3" author="Di Dario F" box="[1172,1348,1543,1565]" pageId="9" pageNumber="184" pagination="496 - 503" refId="ref12875" refString="Di Dario F. 2002. Evidence supporting a sister-group relationship between Clupeoidea and Engrauloidea (Clupeomorpha). Copeia 2002: 496-503." type="journal article" year="2002">Di Dario, 2002</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFAF554CFC8DD730FC62F992" blockId="9.[829,1444,810,1902]" lastBlockId="10.[808,1423,1482,1596]" lastPageId="10" lastPageNumber="185" pageId="9" pageNumber="184">
|
||||
Seven elements of the supraneural series were identified in the fossil, but only the anterior-most four are preserved well enough to be described (
|
||||
<figureCitation id="DFB1C8C0FFAF554FFC9ED794FC70F939" box="[837,906,1666,1688]" captionStart="Figure 8" captionStartId="10.[146,225,1335,1354]" captionTargetBox="[369,1193,197,1301]" captionTargetId="figure-281@10.[369,1193,197,1301]" captionTargetPageId="10" captionText="Figure 8. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the predorsal region. cl, cleithrum; epn, epineurals; na, neural arches; ns, neural spines; pt, post-temporal; r, ribs; sc, supracleithrum; sn, supraneurals; v1–10, vertebrae 1 to 10. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807133" httpUri="https://zenodo.org/record/14807133/files/figure.png" pageId="9" pageNumber="184">Fig. 8</figureCitation>
|
||||
). Supraneurals are well developed and rostrocaudally expanded, slightly differing in shape from each other. The endochondral, rod-like portion of the supraneurals is located at the posterior third of the bone, except in the case of the second supraneural, where this portion of the bone is located approximately in the middle of the structure. The laminar bone projects anteriorly from the endochondral core of the supraneurals, resulting in a roughly rectangular shape, except for the second supraneural (described below). The anterior-most supraneural preserved was probably the first of the series in the living specimen, as it lies close to the post-temporals, as in modern clupeiforms. The second supraneural is more triangle-shaped, expanded proximally and narrow distally, with its posterior margin contacting the anterior margin of the third supraneural. The third and fourth supraneurals are still somewhat rectangle-shaped, but the laminar portion of their proximal half is more expanded anteriorly (
|
||||
<figureCitation id="DFB1C8C0FFAC554CFDE9D60CFD8CF881" box="[562,630,1818,1840]" captionStart="Figure 8" captionStartId="10.[146,225,1335,1354]" captionTargetBox="[369,1193,197,1301]" captionTargetId="figure-281@10.[369,1193,197,1301]" captionTargetPageId="10" captionText="Figure 8. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the predorsal region. cl, cleithrum; epn, epineurals; na, neural arches; ns, neural spines; pt, post-temporal; r, ribs; sc, supracleithrum; sn, supraneurals; v1–10, vertebrae 1 to 10. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807133" httpUri="https://zenodo.org/record/14807133/files/figure.png" pageId="10" pageNumber="185">Fig. 8</figureCitation>
|
||||
). The first and second supraneurals are preserved in an almost vertical orientation in relation to the vertebral series, but the subsequent ones are slightly inclined posteriorly. However, differences in the orientation amongst these bones are possibly artefacts of fossilization.
|
||||
</paragraph>
|
||||
<caption id="13F584CDFFAC554CFF49D421FC6EFA2B" ID-DOI="http://doi.org/10.5281/zenodo.14807133" ID-Zenodo-Dep="14807133" httpUri="https://zenodo.org/record/14807133/files/figure.png" pageId="10" pageNumber="185" startId="10.[146,225,1335,1354]" targetBox="[369,1193,197,1301]" targetPageId="10" targetType="figure">
|
||||
<paragraph id="4735D445FFAC554CFF49D421FC6EFA2B" blockId="10.[146,1423,1335,1413]" pageId="10" pageNumber="185">
|
||||
<emphasis id="75FE0857FFAC554CFF49D421FE16FAE4" bold="true" box="[146,492,1335,1354]" pageId="10" pageNumber="185">
|
||||
Figure 8.
|
||||
<taxonomicName id="808AAFC6FFAC554CFED3D421FE16FAE4" authorityName="Malabarba & Di Dario" authorityYear="2017" box="[264,492,1335,1354]" family="Clupeidae" genus="Cynoclupea" kingdom="Animalia" order="Clupeiformes" pageId="10" pageNumber="185" phylum="Chordata" rank="species" species="nelsoni" status="gen. et sp. nov.">
|
||||
<emphasis id="75FE0857FFAC554CFED3D421FE16FAE4" bold="true" box="[264,492,1335,1354]" italics="true" pageId="10" pageNumber="185">Cynoclupea nelsoni</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="6ECDB52CFFAC554CFE2ED421FD5DFAE4" box="[501,679,1335,1355]" pageId="10" pageNumber="185" rank="species">
|
||||
<emphasis id="75FE0857FFAC554CFE2ED421FDDCFAE4" bold="true" box="[501,550,1335,1354]" pageId="10" pageNumber="185">gen.</emphasis>
|
||||
et
|
||||
<emphasis id="75FE0857FFAC554CFD97D421FD5DFAE4" bold="true" box="[588,679,1335,1354]" pageId="10" pageNumber="185">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
, MCP3447-PV. Photograph (A) and interpretative drawing (B) of the predorsal region. cl, cleithrum; epn, epineurals; na, neural arches; ns, neural spines; pt, post-temporal; r, ribs; sc, supracleithrum; sn, supraneurals; v1
|
||||
<emphasis id="75FE0857FFAC554CFE06D462FE12FA2A" box="[477,488,1396,1412]" italics="true" pageId="10" pageNumber="185">–</emphasis>
|
||||
10, vertebrae 1 to 10. Scale bar = 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="4735D445FFAC554CFCF3D775FBCEF9D6" blockId="10.[808,1423,1635,1901]" box="[808,1076,1635,1656]" pageId="10" pageNumber="185">
|
||||
<emphasis id="75FE0857FFAC554CFCF3D775FBCEF9D6" box="[808,1076,1635,1656]" italics="true" pageId="10" pageNumber="185">Squamation and scutes</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="4735D445FFAC554DFCF3D797FECDFE1D" blockId="10.[808,1423,1635,1901]" lastBlockId="11.[166,781,199,435]" lastPageId="11" lastPageNumber="186" pageId="10" pageNumber="185">
|
||||
Remains of thin cycloid scales are preserved in the lateral region, mainly over the ribs. A few abdominal scutes are preserved on both sides of the fossil specimen. Bony elements identified as abdominal scutes are attached to their presumed living position at the posteroventral margin of the pectoral girdle. More developed scute-like elements, bearing the typical clupeomorph ventral keel, are also present ventrally to the pectoral fin-base insertion (
|
||||
<figureCitation id="DFB1C8C0FFAD554DFDF7D1D1FD8BFF72" box="[556,625,199,221]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="11" pageNumber="186">Fig. 3</figureCitation>
|
||||
). In addition, fragments of three to four abdominal scutes with remains of ascending arms are preserved in the pre-pelvic region of the specimen (
|
||||
<figureCitation id="DFB1C8C0FFAD554DFDC2D034FD8AFE99" box="[537,624,290,312]" captionStart="Figure 2" captionStartId="2.[147,226,1560,1579]" captionTargetBox="[154,1424,198,1527]" captionTargetId="figure-191@2.[154,1424,198,1527]" captionTargetPageId="2" captionText="Figure 2. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV (holotype), part (A) and counterpart (B). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807121" httpUri="https://zenodo.org/record/14807121/files/figure.png" pageId="11" pageNumber="186">Fig. 2B</figureCitation>
|
||||
). Remains of three bony structures tentatively identified as dorsal scutes are also preserved in the occipital region (
|
||||
<figureCitation id="DFB1C8C0FFAD554DFF75D068FF0DFE3D" box="[174,247,382,404]" captionStart="Figure 3" captionStartId="4.[149,228,1645,1664]" captionTargetBox="[376,1200,606,1613]" captionTargetId="figure-374@4.[376,1200,606,1613]" captionTargetPageId="4" captionText="Figure 3. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the internal view of the left side. boc, basioccipital?; br, branchiostegal ray; c5, ceratobranchial 5; cl, cleithrum left (l) and right (r); co, coracoid left (l) and right (r); d, dentary; ds, dorsal scute fragments; ec, ectopterygoid; ep, epioccipital; epc, epicentrals?; epn, epineurals; ex, exoccipital; fr, frontal; gr, fragments of gill rakers; hm, hyomandibula; iop, interopercle; lq, laminar outgrowth of quadrate; mc, mandibular sensory canal; mpt, metapterygoid; mx, maxilla; na, neural arches; pa, parietal; pc, postcleithrum; pf, pre-epiotic fossa; pfr, pectoral fin rays; pop, preopercle; pt, left (l) and right (r) posttemporals; pto, pterotic; q, quadrate; r, ribs; s, symplectic; sc, supracleithrum; smx, posterior supramaxilla; sn, supraneurals; sop, subopercle; ssc, sensory supraorbital canal in the frontal; vlh, ventral limb of hyomandibula; vs, ventral (abdominal) scutes. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14807123" httpUri="https://zenodo.org/record/14807123/files/figure.png" pageId="11" pageNumber="186">Figs 3</figureCitation>
|
||||
,
|
||||
<figureCitation id="DFB1C8C0FFAD554DFED4D068FEE5FE3D" box="[271,287,382,403]" captionStart="Figure 4" captionStartId="5.[166,245,1157,1176]" captionTargetBox="[392,1216,195,1121]" captionTargetId="figure-1070@5.[392,1216,195,1121]" captionTargetPageId="5" captionText="Figure 4. Cynoclupea nelsoni gen. et sp. nov., MCP3447-PV. Photograph (A) and interpretative drawing (B) of the skull roof. boc, basioccipital?; ds, dorsal scute fragments; ep, epioccipital; ex, exoccipital; fr, frontal; pa, parietal; pf, preepiotic fossa; pt, left (l) and right (r) post-temporals; pto, pterotic; sc, supracleithrum; so, supraoccipital. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14807125" httpUri="https://zenodo.org/record/14807125/files/figure.png" pageId="11" pageNumber="186">4</figureCitation>
|
||||
). These structures are keeled and not ornamented.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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