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<document id="EC91F089875905CDE26FE6D845396116" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia &amp; Ahrens, Dirk" docDate="2017" docId="03CB6967FF903841386FFD699300FEDE" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla taitaensis AHRENS, BECKETT, EBERLE &amp; FABRIZI 2017, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="342" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="341" updateTime="1738712560284" updateUser="ExternalLinkService">
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<mods:title id="FF1BF5D13FA470CB74FFC3B55F4B0DF1">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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<mods:namePart id="77025C9E3C07F1686FBF8C10F1DA9A2D">Eberle, Jonas</mods:namePart>
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<mods:namePart id="0B65B7DF5B7059B9E8633C5552852E98">Beckett, Marc</mods:namePart>
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<mods:namePart id="5ADBC96DAA65ED2E74B77088E6AD8DB3">Ozgul-Siemund, Alev</mods:namePart>
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<mods:namePart id="C05AA6BAA0E4E346E28A0202BF446850">Frings, Jujina</mods:namePart>
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<mods:namePart id="7820AC7B6A5DA902813E533126660421">Fabrizi, Silvia</mods:namePart>
</mods:name>
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<mods:namePart id="819CEB44B2AAC35AF84AA45E01F29351">Ahrens, Dirk</mods:namePart>
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<mods:date id="53ECC7F15B795FE12A1C436BDE39B457">2017</mods:date>
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<mods:number id="841FE8D72F8D6B6F12194FDDE2689B77">180</mods:number>
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<treatment id="03CB6967FF903841386FFD699300FEDE" LSID="urn:lsid:plazi:treatment:03CB6967FF903841386FFD699300FEDE" httpUri="http://treatment.plazi.org/id/03CB6967FF903841386FFD699300FEDE" lastPageId="21" lastPageNumber="342" pageId="20" pageNumber="341">
<subSubSection id="C3788BFAFF903840386FFD6992B0FCD1" pageId="20" pageNumber="341" type="nomenclature">
<paragraph id="8BDDD871FF903840386FFD699439FCA4" blockId="20.[851,1377,730,787]" pageId="20" pageNumber="341">
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<taxonomicName id="4C62A3F2FF903840386FFD699548FCA7" ID-CoL="4K84H" authority="AHRENS, BECKETT, EBERLE &amp; FABRIZI" authorityName="AHRENS, BECKETT, EBERLE &amp; FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="taitaensis" status="sp. nov.">
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<smallCapsWord id="8D3B4EADFF90384038CAFD56951AFD44" baselines="750" box="[1014,1148,737,754]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="taitaensis" pageId="20" pageNumber="341">TAITAENSIS</smallCapsWord>
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,
<smallCapsWord id="8D3B4EADFF9038403FCEFD6C943EFD44" baselines="749,750" box="[1266,1368,731,755]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="20" pageNumber="341">BECKETT</smallCapsWord>
,
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&amp;
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</taxonomicName>
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<smallCapsWord id="8D3B4EADFF9038403F0BFD489535FCA6" baselines="780" box="[1079,1107,767,784]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="20" pageNumber="341">SP</smallCapsWord>
.
<smallCapsWord id="8D3B4EADFF9038403F59FD4895F3FCA6" baselines="780" box="[1125,1173,767,784]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="20" pageNumber="341">NOV</smallCapsWord>
.
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(
<figureCitation id="1359C4F4FF9038403F8CFD4D9453FCA4" box="[1200,1333,760,786]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">
<smallCapsWord id="8D3B4EADFF9038403F8CFD4D9583FCA7" baselines="780,781" box="[1200,1253,762,786]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="20" pageNumber="341">FIGS</smallCapsWord>
4M
<emphasis id="B9160463FF9038403E2AFD4B9442FCA4" box="[1302,1316,764,786]" italics="true" pageId="20" pageNumber="341">–</emphasis>
P
</figureCitation>
,
<figureCitation id="1359C4F4FF9038403E7BFD4F9431FCA4" box="[1351,1367,760,786]" captionStart="Figure 7" captionStartId="14.[144,223,925,944]" captionTargetBox="[144,1428,192,893]" captionTargetId="figure-25@14.[144,1428,192,893]" captionTargetPageId="14" captionText="Figure 7. Distribution of Pleophylla species (left side) and absence records (right side), represented by sampling locali- ties of all other sampled Sericini species without records of Pleophylla (white dots)." figureDoi="http://doi.org/10.5281/zenodo.14807097" httpUri="https://zenodo.org/record/14807097/files/figure.png" pageId="20" pageNumber="341">7</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BDDD871FF9038403815FC8592B0FCD1" blockId="20.[809,1424,818,871]" pageId="20" pageNumber="341">
<uri id="FFF3D473FF9038403815FC8592B0FCD1" pageId="20" pageNumber="341">
urn:lsid:zoobank.org:act:
<uuid id="FFC4E2A4FF9038403F04FC8592B0FCD1" pageId="20" pageNumber="341">8E31225B-A3BB-4BB3-91B0- 8A9295AC7295</uuid>
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</paragraph>
</subSubSection>
<subSubSection id="C3788BFAFF903841381BFCCB9300FEDE" lastPageId="21" lastPageNumber="342" pageId="20" pageNumber="341" type="materials_examined">
<paragraph id="8BDDD871FF903840381BFCCB92BCFBBD" blockId="20.[807,1422,891,1035]" pageId="20" pageNumber="341">
<emphasis id="B9160463FF903840381BFCCB9528FC26" box="[807,1102,891,913]" italics="true" pageId="20" pageNumber="341">Type material examined:</emphasis>
<materialsCitation id="3B0AD22CFF9038403F60FCCC92E2FC7B" collectingDate="1997-03-22" collectionCode="ZFMK" collectorName="Lgt. M. Snizek" country="Kenya" location="Hills Wundanyi" pageId="20" pageNumber="341" specimenCount="1" specimenCount-male="1" stateProvince="Taita/Taveta" typeStatus="holotype">
<typeStatus id="54D966D3FF9038403F60FCCC95A4FC27" box="[1116,1218,891,913]" pageId="20" pageNumber="341" type="holotype">Holotype</typeStatus>
<specimenCount id="9D6413F8FF9038403FEDFCC99587FC26" box="[1233,1249,894,912]" count="1" pageId="20" pageNumber="341" type="male">♂</specimenCount>
“
<collectingCountry id="F37598E1FF9038403FC5FCCC9424FC26" box="[1273,1346,891,912]" name="Kenya" pageId="20" pageNumber="341">Kenya</collectingCountry>
<collectingRegion id="49A61693FF9038403E6DFCCC94E8FC27" box="[1361,1422,891,913]" country="Kenya" name="Taita/Taveta" pageId="20" pageNumber="341">Taita</collectingRegion>
<location id="8EBD8EAAFF903840381BFC2D92B8FC06" LSID="urn:lsid:plazi:treatment:03CB6967FF903841386FFD699300FEDE:8EBD8EAAFF903840381BFC2D92B8FC06" box="[807,990,922,944]" country="Kenya" name="Hills Wundanyi" pageId="20" pageNumber="341" stateProvince="Taita/Taveta">Hills Wundanyi</location>
18.3.-
<date id="FFDCFEB1FF9038403F1CFC2D95F5FC19" box="[1056,1171,922,943]" pageId="20" pageNumber="341" value="1997-03-22">
<collectingDate id="EF980759FF9038403F1CFC2D95F5FC19" box="[1056,1171,922,943]" pageId="20" pageNumber="341" value="1997-03-22">22.3.1997</collectingDate>
</date>
<collectorName id="2697BDA7FF9038403FA2FC2D942DFC06" box="[1182,1355,922,944]" pageId="20" pageNumber="341">Lgt. M. Snizek</collectorName>
[sic]” (
<collectionCode id="ED7340B4FF9038403812FC0F921BFC7B" box="[814,893,952,973]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="20" pageNumber="341" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B0AD22CFF90384038A9FC0E92B0FBBD" collectingDate="2001-01" collectionCode="ISNB" collectorName="Coll. I. R. Sc. N. B. &amp; Coll. I. R. Sc. &amp; Leg. S. Sabari" country="Kenya" location="Hills Mbololo" pageId="20" pageNumber="341" specimenCount="2" specimenCount-male="2" stateProvince="Taita/Taveta" typeStatus="paratype">
<typeStatus id="54D966D3FF90384038A9FC0E956DFC78" box="[917,1035,953,974]" pageId="20" pageNumber="341" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D6413F8FF9038403F21FC0F953FFC7B" box="[1053,1113,952,973]" count="2" pageId="20" pageNumber="341" type="male">2 ♂♂</specimenCount>
“
<collectorName id="2697BDA7FF9038403F4CFC0F9449FC7B" box="[1136,1327,952,974]" pageId="20" pageNumber="341">Coll. I.R.Sc.N.B.</collectorName>
<collectingCountry id="F37598E1FF9038403E01FC0E94EFFC78" box="[1341,1417,953,974]" name="Kenya" pageId="20" pageNumber="341">Kenya</collectingCountry>
,
<collectingRegion id="49A61693FF903840381BFC609202FC5B" box="[807,868,983,1005]" country="Kenya" name="Taita/Taveta" pageId="20" pageNumber="341">Taita</collectingRegion>
<location id="8EBD8EAAFF9038403849FC60957FFC5B" LSID="urn:lsid:plazi:treatment:03CB6967FF903841386FFD699300FEDE:8EBD8EAAFF9038403849FC60957FFC5B" box="[885,1049,983,1005]" country="Kenya" name="Hills Mbololo" pageId="20" pageNumber="341" stateProvince="Taita/Taveta">Hills Mbololo</location>
, III.
<date id="FFDCFEB1FF9038403F6CFC6095CCFC5A" box="[1104,1194,983,1004]" pageId="20" pageNumber="341" value="2001-01">
<collectingDate id="EF980759FF9038403F6CFC6095CCFC5A" box="[1104,1194,983,1004]" pageId="20" pageNumber="341" value="2001-01">2001 I.</collectingDate>
</date>
G. 31.839
<collectorName id="2697BDA7FF9038403E08FC609217FBBD" pageId="20" pageNumber="341">Leg. S. Sabari</collectorName>
” (
<collectionCode id="ED7340B4FF90384038B7FC4192A8FBBD" box="[907,974,1014,1035]" pageId="20" pageNumber="341">ISNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDDD871FF903840381BFB979588F963" blockId="20.[807,1422,1055,1750]" pageId="20" pageNumber="341">
<emphasis id="B9160463FF903840381BFB9792C9FB83" box="[807,943,1056,1077]" italics="true" pageId="20" pageNumber="341">Description:</emphasis>
Body length:
<quantity id="4C9A7594FF9038403F5BFBA895A8FB83" box="[1127,1230,1055,1077]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="20" pageNumber="341" unit="mm" value="11.5">11.5 mm</quantity>
, elytral length:
<quantity id="4C9A7594FF903840381BFB8992E6FBE2" box="[807,896,1086,1108]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="20" pageNumber="341" unit="mm" value="7.4">7.4 mm</quantity>
, maximum width:
<quantity id="4C9A7594FF9038403F53FB8995AFFBE2" box="[1135,1225,1086,1108]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="20" pageNumber="341" unit="mm" value="5.4">5.4 mm</quantity>
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus strongly reflexed; smooth area in front of eyes 1.5 times as wide as long; ratio of eye diameter/interocular width: 0.83. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.4. Metatibia, ratio of width/length: 1/4.13. Metatarsi dorsally smooth; first metatarsomere slightly longer than subsequent one. Aedeagus:
<figureCitation id="1359C4F4FF903840389EF9779579F963" box="[930,1055,1728,1750]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">
Fig. 4M
<emphasis id="B9160463FF90384038C0F974956EF963" box="[1020,1032,1731,1749]" italics="true" pageId="20" pageNumber="341">–</emphasis>
O
</figureCitation>
. Habitus:
<figureCitation id="1359C4F4FF9038403FA8F977958CF963" box="[1172,1258,1728,1750]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="20" pageNumber="341">
Fig.
<specimenCount id="9D6413F8FF9038403FF5F977958CF963" box="[1225,1258,1728,1749]" count="4" pageId="20" pageNumber="341" type="generic">4P</specimenCount>
</figureCitation>
.
</paragraph>
<paragraph id="8BDDD871FF903840381BF94F958BF8DF" blockId="20.[807,1421,1783,1897]" pageId="20" pageNumber="341">
<emphasis id="B9160463FF903840381BF94F95FDF8BB" box="[807,1179,1783,1805]" italics="true" pageId="20" pageNumber="341">
Diagnosis:
<taxonomicName id="4C62A3F2FF9038403890F94095FDF8BB" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[940,1179,1783,1805]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="taitaensis" status="sp. nov.">Pleophylla taitaensis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A225B918FF9038403F98F94F9598F8BB" box="[1188,1278,1784,1805]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
differs from the very similar
<taxonomicName id="4C62A3F2FF90384038D1F8A09529F89D" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[1005,1103,1814,1836]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="lizleri" status="sp. nov.">
<emphasis id="B9160463FF90384038D1F8A09529F89D" box="[1005,1103,1814,1836]" italics="true" pageId="20" pageNumber="341">P. lizleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF9038403F65F8A195D2F89D" box="[1113,1204,1814,1835]" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C62A3F2FF9038403FCEF8A09405F89D" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[1266,1379,1814,1836]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="341" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
<emphasis id="B9160463FF9038403FCEF8A09405F89D" box="[1266,1379,1814,1836]" italics="true" pageId="20" pageNumber="341">P. charlyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF9038403E51F8A19231F8FC" pageId="20" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
by the ventral lobes of parameres being symmetrical and not extended dorsally.
</paragraph>
<paragraph id="8BDDD871FF9138413B9AFF709012FEAC" blockId="21.[166,781,198,282]" pageId="21" pageNumber="342">
<emphasis id="B9160463FF9138413B9AFF70907CFF6A" box="[166,282,199,220]" italics="true" pageId="21" pageNumber="342">Variation:</emphasis>
Body length:
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10.4
<emphasis id="B9160463FF9138413914FF7E9352FF6D" box="[552,564,201,219]" italics="true" pageId="21" pageNumber="342">–</emphasis>
11.5 mm
</quantity>
, elytral length:
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7.4
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7.8 mm
</quantity>
, maximum width:
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5.4
<emphasis id="B9160463FF913841399FFF5F93C9FF4C" box="[675,687,232,250]" italics="true" pageId="21" pageNumber="342">–</emphasis>
5.6 mm
</quantity>
. Female unknown.
</paragraph>
<paragraph id="8BDDD871FF9138413B9AFE839300FEDE" blockId="21.[166,780,308,360]" pageId="21" pageNumber="342">
<emphasis id="B9160463FF9138413B9AFE839042FEFF" box="[166,292,308,329]" italics="true" pageId="21" pageNumber="342">Etymology:</emphasis>
The new species is named after its occurrence in the
<collectingRegion id="49A61693FF9138413A48FEE590D7FEDE" box="[372,433,338,360]" country="Kenya" name="Taita/Taveta" pageId="21" pageNumber="342">Taita</collectingRegion>
Hills of
<collectingCountry id="F37598E1FF9138413929FEE49304FEDE" box="[533,610,339,360]" name="Kenya" pageId="21" pageNumber="342">Kenya</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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