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<mods:title id="9D3B7C1213CA425B12AF38E753A581CC">Biodiversity and phylogeny of Marasmius (Agaricales, Basidiomycota) from Madagascar</mods:title>
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<mods:namePart id="F5D40C3A929BFA987A25472AEB58BFC5">Shay, Jackie E.</mods:namePart>
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25. 
<taxonomicName id="4C1C4D3D765FFFB5FF601327B6FBFB6C" ID-CoL="3Y229" authority="J.E. Shay &amp; Desjardin" authorityName="J. E. Shay &amp; Desjardin" authorityYear="2017" box="[178,668,1134,1161]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="34" pageNumber="135" phylum="Basidiomycota" rank="species" species="sokola" status="sp. nov.">
<emphasis id="B968EAAC765FFFB5FF601327B5E0FB6C" bold="true" box="[178,391,1134,1161]" italics="true" pageId="34" pageNumber="135">Marasmius sokola</emphasis>
J.E. Shay &amp; Desjardin
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, 
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<taxonomicNameLabel id="A25B57D7765FFFB5FD7A1326B699FB6C" box="[680,766,1135,1161]" pageId="34" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
(
<figureCitation id="13272A3B765FFFB5FCDC1339B738FB6C" box="[782,863,1136,1161]" captionStart="FIGURE 17" captionStartId="35.[136,229,1220,1241]" captionTargetBox="[158,1436,199,1181]" captionTargetId="figure-14@35.[151,1436,190,1195]" captionTargetPageId="35" captionText="FIGURE 17. Marasmius sokola (JES 154) a) basidiospores; b) basidia; c) basidioles; d) cheilocystidia; e) pileipellis cells. Scale bar = 10 μm. Illustrated by J.E. Shay." figureDoi="http://doi.org/10.5281/zenodo.13697891" httpUri="https://zenodo.org/record/13697891/files/figure.png" pageId="34" pageNumber="135">Fig. 17</figureCitation>
)
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<paragraph id="8BA336BE765FFFB5FF5A13F3B5F0FB2A" blockId="34.[136,1452,1210,1339]" box="[136,407,1210,1231]" pageId="34" pageNumber="135">MycoBank no.: MB 818621</paragraph>
<paragraph id="8BA336BE765FFFB5FF5A1397B667FAF2" blockId="34.[136,1452,1210,1339]" pageId="34" pageNumber="135">
<materialsCitation id="3B743CE3765FFFB5FF5A1397B667FAF2" collectingDate="2014-01-22" collectionCode="J, E, SFSU" country="Madagascar" county="Ranomafana National Park" location="Commune Ranomafana" municipality="Region Vatovavy-Fitovinany" pageId="34" pageNumber="135" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A7881C765FFFB5FF5A1397B485FB16" box="[136,226,1246,1267]" pageId="34" pageNumber="135" type="holotype">Holotype</typeStatus>
:— 
<collectingCountry id="F30B762E765FFFB5FF2C1397B5C9FB16" box="[254,430,1246,1267]" name="Madagascar" pageId="34" pageNumber="135">MADAGASCAR</collectingCountry>
. 
<location id="8EC36065765FFFB5FE651397B6FBFB16" LSID="urn:lsid:plazi:treatment:03B587A8765FFFB4FF5A1339B77EF8DB:8EC36065765FFFB5FE651397B6FBFB16" box="[439,668,1246,1267]" country="Madagascar" county="Ranomafana National Park" municipality="Region Vatovavy-Fitovinany" name="Commune Ranomafana" pageId="34" pageNumber="135">Commune Ranomafana</location>
, 
<location id="8EC36065765FFFB5FD741397B73CFB16" LSID="urn:lsid:plazi:treatment:03B587A8765FFFB4FF5A1339B77EF8DB:8EC36065765FFFB5FD741397B73CFB16" box="[678,859,1246,1267]" country="Madagascar" county="Ranomafana National Park" municipality="Region Vatovavy-Fitovinany" name="District Ifanadiana" pageId="34" pageNumber="135">District Ifanadiana</location>
, 
<collectingMunicipality id="6BC7ACC4765FFFB5FCB71397B01CFB16" box="[869,1147,1246,1267]" pageId="34" pageNumber="135">Region Vatovavy-Fitovinany</collectingMunicipality>
, 
<collectingCounty id="62C24E32765FFFB5FB561397B1ECFB16" box="[1156,1419,1246,1267]" pageId="34" pageNumber="135">Ranomafana National Park</collectingCounty>
, 
<date id="FFA2107E765FFFB5FA461397B57CFAF2" pageId="34" pageNumber="135" value="2014-01-22">
<collectingDate id="EFE6E996765FFFB5FA461397B57CFAF2" pageId="34" pageNumber="135" value="2014-01-22">22 Jan. 2014</collectingDate>
</date>
, 
<collectionCode id="ED0DAE7B765FFFB5FEF7124BB557FAF2" box="[293,304,1282,1303]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="34" pageNumber="135" type="Herbarium">J</collectionCode>
. 
<collectionCode id="ED0DAE7B765FFFB5FEE6124BB521FAF2" box="[308,326,1282,1303]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="34" pageNumber="135" type="Herbarium">E</collectionCode>
. Shay 154 (
<collectionCode id="ED0DAE7B765FFFB5FE64124BB593FAF2" box="[438,500,1282,1303]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14982" lsid="urn:lsid:biocol.org:col:14982" name="Harry D. Thiers Herbarium - San Francisco State University" pageId="34" pageNumber="135" type="Herbarium">SFSU</collectionCode>
).
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<paragraph id="8BA336BE765FFFB5FF5A126FB039FADE" blockId="34.[136,1452,1210,1339]" box="[136,1118,1318,1339]" pageId="34" pageNumber="135">
Etymology:— 
<taxonomicName id="4C1C4D3D765FFFB5FEC3126FB537FADE" authorityName="J. E. Shay &amp; Desjardin" authorityYear="2017" box="[273,336,1318,1339]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="34" pageNumber="135" phylum="Basidiomycota" rank="species" species="sokola">
<emphasis id="B968EAAC765FFFB5FEC3126FB537FADE" box="[273,336,1318,1339]" italics="true" pageId="34" pageNumber="135">sokola</emphasis>
</taxonomicName>
–Malagasy for chocolate, referring the dark chocolate brown colored basidiomes.
</paragraph>
<paragraph id="8BA336BE765FFFB5FF5A122CB1EEFA23" blockId="34.[136,1452,1381,2090]" pageId="34" pageNumber="135">
Description:— 
<taxonomicName id="4C1C4D3D765FFFB5FEF9122CB517FA9B" authorityName="Desor" authorityYear="1857" box="[299,368,1381,1406]" class="Magnoliopsida" family="Caricaceae" genus="Pileus" kingdom="Plantae" order="Brassicales" pageId="34" pageNumber="135" phylum="Tracheophyta" rank="genus">
<emphasis id="B968EAAC765FFFB5FEF9122CB517FA9B" box="[299,368,1381,1406]" italics="true" pageId="34" pageNumber="135">Pileus</emphasis>
</taxonomicName>
<quantity id="4CE49B5B765FFFB5FEA7122CB5A6FA9B" box="[373,449,1381,1406]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="34" pageNumber="135" unit="mm" value="20.0">20 mm</quantity>
diam, campanulate; disc rugulose; margin sulcate; surface dull, glabrous; dark brown (5E7). 
<emphasis id="B968EAAC765FFFB5FF5A12C0B4BAFA47" box="[136,221,1417,1442]" italics="true" pageId="34" pageNumber="135">Context</emphasis>
thin. 
<emphasis id="B968EAAC765FFFB5FEC912C0B5E6FA47" box="[283,385,1417,1442]" italics="true" pageId="34" pageNumber="135">Lamellae</emphasis>
subfree, distant (11), no lamellulae, broad (
<quantity id="4CE49B5B765FFFB5FC8E12C0B7F8FA47" box="[860,927,1417,1442]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="34" pageNumber="135" unit="mm" value="4.0">4 mm</quantity>
), light grey (5C3), with brown edges. 
<emphasis id="B968EAAC765FFFB5FAEF12C0B113FA47" box="[1341,1396,1417,1442]" italics="true" pageId="34" pageNumber="135">Stipe</emphasis>
45 × 
<quantity id="4CE49B5B765FFFB5FF5A12E4B4ACFA23" box="[136,203,1453,1478]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="135" unit="mm" value="1.0">1 mm</quantity>
, central, cylindrical, hollow, pliant; surface glabrous; dark greyish brown (6F4). 
<emphasis id="B968EAAC765FFFB5FBE812E4B0B8FA23" box="[1082,1247,1453,1478]" italics="true" pageId="34" pageNumber="135">Odor and taste</emphasis>
not distinctive.
</paragraph>
<paragraph id="8BA336BE765FFFB5FF6F1298B13CF85B" blockId="34.[136,1452,1381,2090]" pageId="34" pageNumber="135">
<emphasis id="B968EAAC765FFFB5FF6F1298B53CFA0F" box="[189,347,1489,1514]" italics="true" pageId="34" pageNumber="135">Basidiospores</emphasis>
(16.8–) 18.4–23.2 (–26.4) × 4–4.8 μm [x 
<subScript id="179834FB765FFFB5FCE412ABB724FA15" attach="left" box="[822,835,1506,1520]" fontSize="6" pageId="34" pageNumber="135">m</subScript>
= 21.93 ± 2.36 × 4.40 ± 0.40 μm; Q = 3.50–6.40; Q 
<subScript id="179834FB765FFFB5FA4D12ABB1CBFA15" attach="right" box="[1439,1452,1506,1520]" fontSize="6" pageId="34" pageNumber="135">m</subScript>
= 5.03 ± 1.20, n = 25, s =1], subcylindrical to subfusoid, smooth, hyaline, inamyloid, thin-walled. 
<emphasis id="B968EAAC765FFFB5FADE12BCB105F9EB" box="[1292,1378,1525,1550]" italics="true" pageId="34" pageNumber="135">Basidia</emphasis>
32.8– 46.4 × 6.4–8 μm, clavate, 4-spored, hyaline, inamyloid, thin-walled. 
<emphasis id="B968EAAC765FFFB5FC791150B047F9D7" box="[939,1056,1561,1586]" italics="true" pageId="34" pageNumber="135">Basidioles</emphasis>
29.6–42.4 × 7.2–8.8 μm, clavate, hyaline, inamyloid, thin-walled. 
<emphasis id="B968EAAC765FFFB5FE241174B6FFF9B3" box="[502,664,1597,1622]" italics="true" pageId="34" pageNumber="135">Cheilocystidia</emphasis>
abundant, of 
<emphasis id="B968EAAC765FFFB5FCE31174B71EF9B3" box="[817,889,1597,1622]" italics="true" pageId="34" pageNumber="135">Siccus</emphasis>
- 
<typeStatus id="54A7881C765FFFB5FC521174B7D5F9B3" box="[896,946,1597,1622]" pageId="34" pageNumber="135">type</typeStatus>
broom cells; main body 14.4–22.4 × 6.4–8.8 μm, clavate to subglobose or irregular, seldom 2–3-lobed, hyaline, inamyloid, thin-walled; apical setulae 0.8–13.6 × 0.8–1.6 μm, cylindrical to conical, often branched, brown, inamyloid, thick-walled. 
<emphasis id="B968EAAC765FFFB5FBE811CCB0BBF97B" box="[1082,1244,1669,1694]" italics="true" pageId="34" pageNumber="135">Pleurocystidia</emphasis>
absent. 
<emphasis id="B968EAAC765FFFB5FAE811CCB1CBF97B" box="[1338,1452,1669,1694]" italics="true" pageId="34" pageNumber="135">Pileipellis</emphasis>
mottled, a hymeniform layer of 
<emphasis id="B968EAAC765FFFB5FE2511E0B658F927" box="[503,575,1705,1730]" italics="true" pageId="34" pageNumber="135">Siccus</emphasis>
- 
<typeStatus id="54A7881C765FFFB5FD9411E0B61FF927" box="[582,632,1705,1730]" pageId="34" pageNumber="135">type</typeStatus>
broom cells; main body 8.8–22.4 × 5.6–10.4 μm, cylindrical to clavate or irregular, seldom 2–3-lobed, hyaline, inamyloid, thick-walled; apical setulae 0.8–9.6 × 0.8–1.6 μm, cylindrical to conical, seldom branched, hyaline to brown, inamyloid, thick-walled. 
<taxonomicName id="4C1C4D3D765FFFB5FC4C11B8B048F8EF" box="[926,1071,1777,1802]" class="Magnoliopsida" family="Caricaceae" genus="Pileus" kingdom="Plantae" order="Brassicales" pageId="34" pageNumber="135" phylum="Tracheophyta" rank="species" species="trama">
<emphasis id="B968EAAC765FFFB5FC4C11B8B048F8EF" box="[926,1071,1777,1802]" italics="true" pageId="34" pageNumber="135">Pileus trama</emphasis>
</taxonomicName>
interwoven; hyphae 2.4–17.6 μm diam, cylindrical to inflated, rough, hyaline, dextrinoid, thick-walled. 
<emphasis id="B968EAAC765FFFB5FC7D105CB004F8CB" box="[943,1123,1813,1838]" italics="true" pageId="34" pageNumber="135">
Lamellar 
<taxonomicName id="4C1C4D3D765FFFB5FBF2105CB004F8CB" box="[1056,1123,1813,1838]" class="Magnoliopsida" family="Caricaceae" genus="Pileus" kingdom="Plantae" order="Brassicales" pageId="34" pageNumber="135" phylum="Tracheophyta" rank="species" species="trama">trama</taxonomicName>
</emphasis>
regular; hyphae 3.2–8.8 μm diam, cylindrical, smooth, hyaline, dextrinoid, thin-walled. 
<emphasis id="B968EAAC765FFFB5FCCC1070B7FBF8B7" box="[798,924,1849,1874]" italics="true" pageId="34" pageNumber="135">Stipe tissue</emphasis>
monomitic; cortical hyphae 3.2–11.2 μm diam, cylindrical, smooth, pale greenish brown, dextrinoid, thick-walled; medullary hyphae 2.4–12.8 μm diam, cylindrical to inflated, smooth, hyaline to pale brown, inamyloid, thin-walled. 
<emphasis id="B968EAAC765FFFB5FCA510C8B074F87F" box="[887,1043,1921,1946]" italics="true" pageId="34" pageNumber="135">Caulocystidia</emphasis>
absent. 
<emphasis id="B968EAAC765FFFB5FBBD10C8B120F87F" box="[1135,1351,1921,1946]" italics="true" pageId="34" pageNumber="135">Clamp connections</emphasis>
present. Habit, habitat and known distribution:—Solitary on woody sticks of 
<taxonomicName id="4C1C4D3D765FFFB5FC5810ECB072F85B" box="[906,1045,1957,1982]" class="Magnoliopsida" family="Cunoniaceae" genus="Weinmannia" kingdom="Plantae" order="Oxalidales" pageId="34" pageNumber="135" phylum="Tracheophyta" rank="genus">
<emphasis id="B968EAAC765FFFB5FC5810ECB072F85B" box="[906,1045,1957,1982]" italics="true" pageId="34" pageNumber="135">Weinmannia</emphasis>
</taxonomicName>
(
<taxonomicName id="4C1C4D3D765FFFB5FBF610ECB0DBF85B" box="[1060,1212,1957,1982]" class="Magnoliopsida" family="Cunoniaceae" kingdom="Plantae" order="Oxalidales" pageId="34" pageNumber="135" phylum="Tracheophyta" rank="family">Cunoniaceae</taxonomicName>
). 
<collectingCountry id="F30B762E765FFFB5FB1F10ECB131F85B" box="[1229,1366,1957,1982]" name="Madagascar" pageId="34" pageNumber="135">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="8BA336BE765FFFB5FF6F1080B482F7CF" blockId="34.[136,1452,1381,2090]" pageId="34" pageNumber="135">
Material examined:— 
<collectingCountry id="F30B762E765FFFB5FE7C1080B61CF807" box="[430,635,1993,2018]" name="Madagascar" pageId="34" pageNumber="135">MADAGASCAR</collectingCountry>
.Region Vatovavy-Fitovinany,District Ifanadiana,Commune Ranomafana, Ranomafana National Park, Piste B, elev. 
<quantity id="4CE49B5B765FFFB5FD8F10A4B6D3F7E3" box="[605,692,2029,2054]" metricMagnitude="3" metricUnit="m" metricValue="1.004" pageId="34" pageNumber="135" unit="m" value="1004.0">1004 m</quantity>
, GPS: 21˚ 15.413’ S, 47˚ 25.253’ E, 
<date id="FFA2107E765FFFB5FB9E10A4B160F7E3" box="[1100,1287,2029,2054]" pageId="34" pageNumber="135" value="2014-01-22">22 January 2014</date>
, 
<emphasis id="B968EAAC765FFFB5FAC010A4B1CCF7E3" box="[1298,1451,2029,2054]" italics="true" pageId="34" pageNumber="135">J.E. Shay 154</emphasis>
(SFSU).
</paragraph>
<caption id="DF636636765EFFB4FF5A138DB5FEFB18" ID-DOI="http://doi.org/10.5281/zenodo.13697891" ID-Zenodo-Dep="13697891" httpUri="https://zenodo.org/record/13697891/files/figure.png" pageId="35" pageNumber="136" startId="35.[136,229,1220,1241]" targetBox="[158,1436,199,1181]" targetPageId="35" targetType="figure">
<paragraph id="8BA336BE765EFFB4FF5A138DB5FEFB18" blockId="35.[136,1452,1220,1277]" pageId="35" pageNumber="136">
<emphasis id="B968EAAC765EFFB4FF5A138DB56EFB3C" bold="true" box="[136,265,1220,1241]" pageId="35" pageNumber="136">FIGURE 17.</emphasis>
<taxonomicName id="4C1C4D3D765EFFB4FEC6138DB5A2FB3C" authorityName="J. E. Shay &amp; Desjardin" authorityYear="2017" box="[276,453,1220,1241]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="sokola">
<emphasis id="B968EAAC765EFFB4FEC6138DB5A2FB3C" box="[276,453,1220,1241]" italics="true" pageId="35" pageNumber="136">Marasmius sokola</emphasis>
</taxonomicName>
(
<emphasis id="B968EAAC765EFFB4FE00138DB646FB3C" box="[466,545,1220,1241]" italics="true" pageId="35" pageNumber="136">JES 154</emphasis>
) a) basidiospores; b) basidia; c) basidioles; d) cheilocystidia; e) pileipellis cells. Scale bar = 10 μm. Illustrated by J.E. Shay.
</paragraph>
</caption>
<paragraph id="8BA336BE765EFFB4FF6F1264B77EF8DB" blockId="35.[136,1452,1325,1854]" pageId="35" pageNumber="136">
Notes:— 
<taxonomicName id="4C1C4D3D765EFFB4FEF01264B594FAA3" authorityName="J. E. Shay &amp; Desjardin" authorityYear="2017" box="[290,499,1325,1350]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="sokola">
<emphasis id="B968EAAC765EFFB4FEF01264B594FAA3" box="[290,499,1325,1350]" italics="true" pageId="35" pageNumber="136">Marasmius sokola</emphasis>
</taxonomicName>
is characterized by a dark brown, smooth to rugulose pileus 
<quantity id="4CE49B5B765EFFB4FB6F1264B169FAA3" box="[1213,1294,1325,1350]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="136" unit="mm" value="20.0">20 mm</quantity>
diam, distant (11), broad (
<quantity id="4CE49B5B765EFFB4FEC41218B539FA8F" box="[278,350,1361,1386]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="35" pageNumber="136" unit="mm" value="4.0">4 mm</quantity>
), brown-marginate lamellae, a glabrous, dark brown stipe 
<quantity id="4CE49B5B765EFFB4FC2A1218B02EFA8F" box="[1016,1097,1361,1386]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="35" pageNumber="136" unit="mm" value="45.0">45 mm</quantity>
long, basidiospores with mean 21.9 × 4.4 μm (Q = 5.0), basidia 32–46 μm long, 
<emphasis id="B968EAAC765EFFB4FD11123CB76CFA6B" box="[707,779,1397,1422]" italics="true" pageId="35" pageNumber="136">Siccus</emphasis>
- 
<typeStatus id="54A7881C765EFFB4FCC0123CB723FA6B" box="[786,836,1397,1422]" pageId="35" pageNumber="136">type</typeStatus>
cheilocystidia and pileipellis cells with brown setulae up to 12 μm long, no pleurocystidia or caulocystidia, and growth on woody dicotyledonous debris. The new species shows closest phenetic similarity to 
<taxonomicName id="4C1C4D3D765EFFB4FDF012F4B76BFA33" authority="Singer" authorityName="Singer" box="[546,780,1469,1494]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="carcharus">
<emphasis id="B968EAAC765EFFB4FDF012F4B6DDFA33" box="[546,698,1469,1494]" italics="true" pageId="35" pageNumber="136">M. carcharus</emphasis>
Singer
</taxonomicName>
, 
<taxonomicName id="4C1C4D3D765EFFB4FCCA12F4B0B3FA33" authority="Y. S. Tan &amp; Desjardin" authorityName="Y. S. Tan &amp; Desjardin" box="[792,1236,1469,1494]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="selangorensis">
<emphasis id="B968EAAC765EFFB4FCCA12F4B7BFFA33" box="[792,984,1469,1494]" italics="true" pageId="35" pageNumber="136">M. selangorensis</emphasis>
Y.S. Tan &amp; Desjardin
</taxonomicName>
, and 
<taxonomicName id="4C1C4D3D765EFFB4FAC212F4B4B0FA1F" authority="Singer." authorityName="Singer." class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="mazatecus">
<emphasis id="B968EAAC765EFFB4FAC212F4B1CBFA33" box="[1296,1452,1469,1494]" italics="true" pageId="35" pageNumber="136">M. mazatecus</emphasis>
Singer.
</taxonomicName>
<taxonomicName id="4C1C4D3D765EFFB4FF3212A8B5BFFA1F" box="[224,472,1505,1530]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="carcharus">
<emphasis id="B968EAAC765EFFB4FF3212A8B5BFFA1F" box="[224,472,1505,1530]" italics="true" pageId="35" pageNumber="136">Marasmius carcharus</emphasis>
</taxonomicName>
, described from the 
<collectingCountry id="F30B762E765EFFB4FD1412A8B724FA1F" box="[710,835,1505,1530]" name="Democratic Republic of the Congo" pageId="35" pageNumber="136">DR Congo</collectingCountry>
, differs in a pinkish brown pileus, broader (
<quantity id="4CE49B5B765EFFB4FAE912A8B1F8FA1F" box="[1339,1439,1505,1530]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="35" pageNumber="136" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
), non-marginate lamellae, cheilocystidia with shorter apical setulae (up to 8 μm), and growth on dicotyledonous leaves (
<bibRefCitation id="EF8D4B4F765EFFB4FF421160B555F9A7" author="Antonin, V." box="[144,306,1577,1602]" pageId="35" pageNumber="136" refId="ref33906" refString="Antonin, V. (2007) Monograph of Marasmius, Gloiocephala, Palaeocephala and Setulipes in Tropical Africa. In: Fungus Flora of Tropical Africa, v. 1. National Botanic Garden, Belgium." type="book" year="2007">Antonín 2007</bibRefCitation>
). 
<taxonomicName id="4C1C4D3D765EFFB4FE971160B603F9A7" box="[325,612,1577,1602]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="selangorensis">
<emphasis id="B968EAAC765EFFB4FE971160B603F9A7" box="[325,612,1577,1602]" italics="true" pageId="35" pageNumber="136">Marasmius selangorensis</emphasis>
</taxonomicName>
, described from 
<collectingCountry id="F30B762E765EFFB4FCF61160B7EAF9A7" box="[804,909,1577,1602]" name="Malaysia" pageId="35" pageNumber="136">Malaysia</collectingCountry>
, differs in a paler brown pileus with pink tones, narrower and more numerous (12–18) lamellae, and growth on dead dicotyledonous leaves (
<bibRefCitation id="EF8D4B4F765EFFB4FB121104B11DF983" author="Tan, Y. S. &amp; Desjardin, D. E. &amp; Perry, B. A. &amp; Vikineswary, S. &amp; Noorlidah, A." box="[1216,1402,1613,1638]" pageId="35" pageNumber="136" pagination="9 - 100" refId="ref36002" refString="Tan, Y. S., Desjardin, D. E., Perry, B. A., Vikineswary, S. &amp; Noorlidah, A. (2009) Marasmius sensu stricto in Peninsular Malaysia. Fungal Diversity 37: 9 - 100." type="journal article" year="2009">
Tan 
<emphasis id="B968EAAC765EFFB4FB241104B14CF983" box="[1270,1323,1613,1638]" italics="true" pageId="35" pageNumber="136">et al</emphasis>
. 2009
</bibRefCitation>
); its micromorphology is indistinguishable from 
<taxonomicName id="4C1C4D3D765EFFB4FD511138B693F96F" authorityName="J. E. Shay &amp; Desjardin" authorityYear="2017" box="[643,756,1649,1674]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="sokola">
<emphasis id="B968EAAC765EFFB4FD511138B693F96F" box="[643,756,1649,1674]" italics="true" pageId="35" pageNumber="136">M. sokola</emphasis>
</taxonomicName>
. ITS sequences of 
<specimenCount id="9D1AFD37765EFFB4FC041138B018F96F" box="[982,1151,1649,1674]" count="2" pageId="35" pageNumber="136" type="generic">two specimens</specimenCount>
of 
<taxonomicName id="4C1C4D3D765EFFB4FB781138B10CF96F" box="[1194,1387,1649,1674]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="selangorensis">
<emphasis id="B968EAAC765EFFB4FB781138B10CF96F" box="[1194,1387,1649,1674]" italics="true" pageId="35" pageNumber="136">M. selangorensis</emphasis>
</taxonomicName>
from 
<collectingCountry id="F30B762E765EFFB4FF5A11DCB488F94B" box="[136,239,1685,1710]" name="Malaysia" pageId="35" pageNumber="136">Malaysia</collectingCountry>
(
<figureCitation id="13272A3B765EFFB4FF2D11DCB537F94B" box="[255,336,1685,1710]" captionStart="FIGURE 1" captionStartId="4.[136,229,1876,1897]" captionTargetBox="[151,1417,192,1821]" captionTargetId="figure-16@4.[151,1436,190,1851]" captionTargetPageId="4" captionText="FIGURE 1a. Maximum likelihood phylogeny based on ITS sequence data. Marasmius from Madagascar are indicated in bold type.Values separated by / refer to nonparametric ML bootstrap proportions and Bayesian posterior probabilities, respectively. Values greater than 70/0.70 are shown (- designates a value below 70% or below 0.70). Nodes receiving support values greater than 90/0.95 are represented by bold branches. G—sect. Globulares; N—sect. Neosessiles; SA—sect. Sicci ser. Atrorubentes; SL—sect. Sicci ser. Leonini; SH—sect. Sicci ser. Haematocephali." figureDoi="http://doi.org/10.5281/zenodo.13697835" httpUri="https://zenodo.org/record/13697835/files/figure.png" pageId="35" pageNumber="136">Fig. 1a</figureCitation>
), however, show only 70% similarity to 
<emphasis id="B968EAAC765EFFB4FCF311DCB718F94B" box="[801,895,1685,1710]" italics="true" pageId="35" pageNumber="136">JES 154</emphasis>
(KX148994) (
<figureCitation id="13272A3B765EFFB4FBF511DCB01FF94B" box="[1063,1144,1685,1710]" captionStart="FIGURE 1" captionStartId="4.[136,229,1876,1897]" captionTargetBox="[151,1417,192,1821]" captionTargetId="figure-16@4.[151,1436,190,1851]" captionTargetPageId="4" captionText="FIGURE 1a. Maximum likelihood phylogeny based on ITS sequence data. Marasmius from Madagascar are indicated in bold type.Values separated by / refer to nonparametric ML bootstrap proportions and Bayesian posterior probabilities, respectively. Values greater than 70/0.70 are shown (- designates a value below 70% or below 0.70). Nodes receiving support values greater than 90/0.95 are represented by bold branches. G—sect. Globulares; N—sect. Neosessiles; SA—sect. Sicci ser. Atrorubentes; SL—sect. Sicci ser. Leonini; SH—sect. Sicci ser. Haematocephali." figureDoi="http://doi.org/10.5281/zenodo.13697835" httpUri="https://zenodo.org/record/13697835/files/figure.png" pageId="35" pageNumber="136">Fig. 1b</figureCitation>
) and are distant in the ITS phylogeny. 
<taxonomicName id="4C1C4D3D765EFFB4FEDC11F0B66BF937" box="[270,524,1721,1746]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="mazatecus">
<emphasis id="B968EAAC765EFFB4FEDC11F0B66BF937" box="[270,524,1721,1746]" italics="true" pageId="35" pageNumber="136">Marasmius mazatecus</emphasis>
</taxonomicName>
, described from 
<collectingCountry id="F30B762E765EFFB4FD0711F0B74AF937" box="[725,813,1721,1746]" name="Mexico" pageId="35" pageNumber="136">Mexico</collectingCountry>
, differs in an orange-ferruginous pileus, fewer (9) and narrower (
<quantity id="4CE49B5B765EFFB4FF2F1194B526F913" box="[253,321,1757,1782]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="136" unit="mm" value="2.0">2 mm</quantity>
) lamellae with orange-ferruginous edges, a shorter (
<quantity id="4CE49B5B765EFFB4FC5F1194B7B8F913" box="[909,991,1757,1782]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="136" unit="mm" value="20.0">20 mm</quantity>
) stipe, and slightly shorter basidiospores (17–21 μm) (
<bibRefCitation id="EF8D4B4F765EFFB4FEC51048B5C4F8FF" author="Singer, R." box="[279,419,1793,1818]" pageId="35" pageNumber="136" pagination="1 - 347" refId="ref35879" refString="Singer, R. (1976) Marasmiaceae (Basidiomycetes Tricholomataceae). Flora Neotropica Monograph 17: 1 - 347." type="journal article" year="1976">Singer 1976</bibRefCitation>
). 
<taxonomicName id="4C1C4D3D765EFFB4FE661048B618F8FF" authorityName="J. E. Shay &amp; Desjardin" authorityYear="2017" box="[436,639,1793,1818]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="sokola">
<emphasis id="B968EAAC765EFFB4FE661048B618F8FF" box="[436,639,1793,1818]" italics="true" pageId="35" pageNumber="136">Marasmius sokola</emphasis>
</taxonomicName>
is on a long branch in the ITS phylogeny, sister to 
<taxonomicName id="4C1C4D3D765EFFB4FB4C1048B5EEF8DB" authority="Wannathes, Desjardin &amp; Lumyong" authorityName="Wannathes, Desjardin &amp; Lumyong" authorityYear="2009" baseAuthorityName="SL" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="35" pageNumber="136" phylum="Basidiomycota" rank="species" species="imitarius">
<emphasis id="B968EAAC765EFFB4FB4C1048B141F8FF" box="[1182,1318,1793,1818]" italics="true" pageId="35" pageNumber="136">M. imitarius</emphasis>
Wannathes, Desjardin &amp; Lumyong
</taxonomicName>
, a species described from 
<collectingCountry id="F30B762E765EFFB4FD7C106CB772F8DB" box="[686,789,1829,1854]" name="Thailand" pageId="35" pageNumber="136">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>