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<document id="2FC358F478FBDCC92F079ADD2ED394E6" ID-DOI="10.37828/em.2023.61.9" ID-ISSN="2336-9744" ID-Zenodo-Dep="13246732" ID-ZooBank="urn:lsid:zoobank.org:pub:4EB022DC-076A-4C1C-BFBD-01F9B4342F0B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722972049171" checkinUser="felipe" docAuthor="Dubatolov, Vladimir V., Korb, Stanislav K. &amp; Yakovlev, Roman V." docDate="2023" docId="601E954AFFAFFFE71128205B8724D8A2" docLanguage="en" docName="EcolMontenegrina.61.88-123.pdf" docOrigin="Ecologica Montenegrina 61" docSource="http://dx.doi.org/10.37828/em.2023.61.9" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Triphysa striatula subsp. striatula Elwes 1899" docType="treatment" docVersion="2" lastPageNumber="101" masterDocId="9C27ED32FFA4FFEA1302231E833CD90B" masterDocTitle="A review of the genus Triphysa Zeller, 1858 (Lepidoptera, Satyridae)" masterLastPageNumber="123" masterPageNumber="88" pageNumber="99" updateTime="1722979451847" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8CC61D27CCFD6225003BEA2A5D596474">A review of the genus Triphysa Zeller, 1858 (Lepidoptera, Satyridae)</mods:title>
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<mods:namePart id="0D61DD242C8F4562A9696DCF2C881130">Dubatolov, Vladimir V.</mods:namePart>
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<mods:affiliation id="B1691609BCC0C88AF80D9AACAFF7D0C8">Institute of Systematics and Ecology of Animals, Siberian Branch of Russian Academy of Sciences, Frunze str. 11, Novosibirsk 630091 Russia. https: // orcid. org / 0000 - 0001 - 7687 - 2102</mods:affiliation>
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<mods:namePart id="9850531CBAC44AF612109EA5646E1DE1">Korb, Stanislav K.</mods:namePart>
<mods:affiliation id="DD0E506CB04CD480779BC073C1A6C6E6">Russian Entomological Society, Nizhny Novgorod Division P. O. Box 97, Nizhny Novgorod 603009 Russia.</mods:affiliation>
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<mods:namePart id="D2884DF349A96494DB96028B021783A7">Yakovlev, Roman V.</mods:namePart>
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<mods:affiliation id="E9F0ACF837EB87BE9DE1AEAC9DA80FAB">Altai State University, pr. Lenina 61, Barnaul, 656049, Russia. https: // orcid. org / 0000 - 0001 - 9512 - 8709 * Corresponding author. E-mail: stanislavkorb @ list. ru; https: // orcid. org / 0000 - 0002 - 1120 - 424 X</mods:affiliation>
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<mods:date id="3278CC554D41AEAB7A4235B8CC89398E">2023</mods:date>
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<treatment id="601E954AFFAFFFE71128205B8724D8A2" LSID="urn:lsid:plazi:treatment:601E954AFFAFFFE71128205B8724D8A2" httpUri="http://treatment.plazi.org/id/601E954AFFAFFFE71128205B8724D8A2" lastPageId="13" lastPageNumber="101" pageId="11" pageNumber="99">
<subSubSection id="A0AD77D7FFAFFFE11128205B8736DA54" box="[554,1034,837,863]" pageId="11" pageNumber="99" type="nomenclature">
<paragraph id="E808245CFFAFFFE11128205B8736DA54" blockId="11.[545,1043,804,897]" box="[554,1034,837,863]" pageId="11" pageNumber="99">
<heading id="B3409330FFAFFFE11128205B8736DA54" box="[554,1034,837,863]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="99" reason="3">
<taxonomicName id="2FB75FDFFFAFFFE11128205B8736DA54" ID-CoL="957NP" authority="Elwes, 1899" authorityName="Elwes" authorityYear="1899" box="[554,1034,837,863]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="99" phylum="Arthropoda" rank="subSpecies" species="striatula" subSpecies="striatula">
<emphasis id="DAC3F84EFFAFFFE11128205B8736DA54" bold="true" box="[554,1034,837,863]" pageId="11" pageNumber="99">
<emphasis id="DAC3F84EFFAFFFE11128205B8053DA54" bold="true" box="[554,879,837,863]" italics="true" pageId="11" pageNumber="99">Triphysa striatula striatula</emphasis>
<bibRefCitation id="8C2659ADFFAFFFE11074205B8736DA54" author="Elwes, H. J." box="[886,1034,837,863]" pageId="11" pageNumber="99" pagination="295 - 367" refId="ref22065" refString="Elwes, H. J. (1899) On the Lepidoptera of the Altai Mountains. Transactions of entomological Society London, 47 (3), 295 - 367." type="journal article" year="1899">Elwes, 1899</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFAFFFE111AC207880BADA8A" box="[686,902,870,897]" pageId="11" pageNumber="99" type="description">
<paragraph id="E808245CFFAFFFE111AC207880BADA8A" blockId="11.[545,1043,804,897]" box="[686,902,870,897]" pageId="11" pageNumber="99">
<figureCitation id="708C38D9FFAFFFE111AC20788016DA8A" box="[686,810,870,897]" captionStart="Figure 1" captionStartId="5.[151,226,1746,1770]" captionTargetBox="[247,1340,200,1714]" captionTargetId="figure-22@5.[241,1346,189,1728]" captionTargetPageId="5" captionText="Figure 1. Triphysa sp., imago: a neotype of Triphysa dohrnii Zeller, 1858 (ZISP), upperside, b underside; c T. phryne, male, Crimea, Dzhankoj distr., near Yasnopolyanskoe vill., 25.04.2009, leg. Savchuk V. (RYB), upperside, d underside; e T. phryne, Russia, S. Ural, Kuvandyk, 15.05.1996, leg. V. Barkhatov (RYB), upperside, f underside; g T. phryne, female, same locality, 1-10.06.1995 (RYB), upperside, h underside; i T. phryne, male, Russia, Altai Krai, Ugly distr., Lyapunovo, 9.05.2003, leg. D. Ryzhkov (RYB), upperside, j underside; k T. phryne, female, E. Kazakhstan, Manrak Mts., near Zhanaul vill., 1100 m, 47º14 N, 84º35 E, 3.06.2003, leg. R. Yakovlev (RYB) upperside, l underside; m T. phryne kasikoporana ssp. n., holotype (ZISP), upperside, n underside; o T. phryne kasikoporana ssp. n., paratype, female (ZSSM), upperside, p underside; q T. striatula, cotype, male (MHUB), upperside, r underside; s T. striatula, cotype, female (MHUB), upperside, t underside; u T. striatula, cotype, male (MHUB), upperside, v underside; w T. striatula, cotype, male (ZISP), upperside, x underside." figureDoi="http://doi.org/10.5281/zenodo.13246736" httpUri="https://zenodo.org/record/13246736/files/figure.png" pageId="11" pageNumber="99">Figs. 1qz</figureCitation>
,
<figureCitation id="708C38D9FFAFFFE1103520788069DA8A" box="[823,853,870,897]" captionStart="Figure 6" captionStartId="12.[151,226,1968,1992]" captionTargetBox="[211,1376,217,1925]" captionTargetId="figure-8@12.[210,1376,189,1249]" captionTargetPageId="12" captionText="Figure 6. Distribution map of Triphysa species: a (above) general map; b (below) northern-eastern part of area of T. nervosa nervosa." figureDoi="http://doi.org/10.5281/zenodo.13246750" httpUri="https://zenodo.org/record/13246750/files/figure.png" pageId="11" pageNumber="99">6a</figureCitation>
,
<figureCitation id="708C38D9FFAFFFE11063207880BEDA8A" box="[865,898,870,897]" captionStart="Figures 2024" captionStartId="26.[151,237,1964,1988]" captionTargetBox="[251,1354,202,1932]" captionTargetId="figure-8@26.[222,1366,189,1946]" captionTargetPageId="26" captionText="Figures 2024. Male genitalia (lateral projection) of Triphysa: 20 T. phryne, Volgograd reg.; 21 T. phryne kasikoporana, holotype; 22 T. striatula, cotype; 23 T. striatula urumtshiensis, holotype; 24 T. issykkulica, holotype." figureDoi="http://doi.org/10.5281/zenodo.13246772" httpUri="https://zenodo.org/record/13246772/files/figure.png" pageId="11" pageNumber="99">22</figureCitation>
.
</paragraph>
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<subSubSection id="A0AD77D7FFAFFFE1139520B780C8DACA" box="[151,1012,937,961]" pageId="11" pageNumber="99" type="reference_group">
<paragraph id="E808245CFFAFFFE1139520B780C8DACA" blockId="11.[151,1012,937,961]" box="[151,1012,937,961]" pageId="11" pageNumber="99">
<taxonomicName id="2FB75FDFFFAFFFE1139520B7810ADACA" ID-CoL="957NP" authority="Elwes, 1899" authorityName="Elwes" authorityYear="1899" box="[151,566,937,961]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="99" phylum="Arthropoda" rank="subSpecies" species="phryne" subSpecies="striatula">
<emphasis id="DAC3F84EFFAFFFE1139520B7829BDACA" box="[151,423,937,961]" italics="true" pageId="11" pageNumber="99">Triphysa phryne striatula</emphasis>
<bibRefCitation id="8C2659ADFFAFFFE112AC20B7810ADACA" author="Elwes, H. J." box="[430,566,937,961]" pageId="11" pageNumber="99" pagination="295 - 367" refId="ref22065" refString="Elwes, H. J. (1899) On the Lepidoptera of the Altai Mountains. Transactions of entomological Society London, 47 (3), 295 - 367." type="journal article" year="1899">Elwes, 1899</bibRefCitation>
</taxonomicName>
; Trans. Entomol. Soc. London 1899: 365.
</paragraph>
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<subSubSection id="A0AD77D7FFAFFFE1139520F48299DD61" pageId="11" pageNumber="99" type="materials_examined">
<paragraph id="E808245CFFAFFFE1139520F48299DD61" blockId="11.[151,1437,1002,1535]" pageId="11" pageNumber="99">
<materialsCitation id="58DF2E01FFAFFFE1139520F4829DDD61" country="China" location="Kuruktagh" municipality="Korla" pageId="11" pageNumber="99" specimenCount="1" stateProvince="Xinjiang Uygur" typeStatus="holotype">
<emphasis id="DAC3F84EFFAFFFE1139520F4820EDD0F" bold="true" box="[151,306,1002,1028]" italics="true" pageId="11" pageNumber="99">
<typeStatus id="370C9AFEFFAFFFE1139520F483EEDD0F" box="[151,210,1002,1028]" pageId="11" pageNumber="99">Type</typeStatus>
locality
</emphasis>
:
<location id="ED687287FFAFFFE1124020F48283DD0E" LSID="urn:lsid:plazi:treatment:601E954AFFAFFFE71128205B8724D8A2:ED687287FFAFFFE1124020F48283DD0E" box="[322,447,1002,1029]" country="China" municipality="Korla" name="Kuruktagh" pageId="11" pageNumber="99" stateProvince="Xinjiang Uygur">Kuruktagh</location>
near
<collectingMunicipality id="086CBE26FFAFFFE1110120F4817ADD0E" box="[515,582,1002,1029]" pageId="11" pageNumber="99">Korla</collectingMunicipality>
[East Tian-Shan,
<collectingRegion id="2A73EABEFFAFFFE1101E20F480BADD0E" box="[796,902,1002,1029]" country="China" name="Xinjiang Uygur" pageId="11" pageNumber="99">Xinjiang</collectingRegion>
,
<collectingCountry id="90A064CCFFAFFFE1109120F480DCDD0E" box="[915,992,1002,1029]" name="China" pageId="11" pageNumber="99">China</collectingCountry>
]. “Lastly, there is a form found in the Kuruktagh near Korla, var.
<taxonomicName id="2FB75FDFFFAFFFE112E1271581CCDD2D" authority=", Stgr. (MSS.)" box="[483,752,1035,1062]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="99" phylum="Arthropoda" rank="species" species="striatula">Striatula, Stgr. (MSS.)</taxonomicName>
, which is distinguished by pale longitudinal striations on the fore-wing of the male. This form seems by the position of the spots to belong to phryne rather than to dohrnii” (
<bibRefCitation id="8C2659ADFFAFFFE1120A275182A4DD61" author="Elwes, H. J." box="[264,408,1103,1130]" pageId="11" pageNumber="99" pagination="295 - 367" refId="ref22065" refString="Elwes, H. J. (1899) On the Lepidoptera of the Altai Mountains. Transactions of entomological Society London, 47 (3), 295 - 367." type="journal article" year="1899">Elwes 1899</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="A0AD77D7FFAFFFE113F4276F863DDC3E" pageId="11" pageNumber="99" type="description">
<paragraph id="E808245CFFAFFFE113F4276F8717DDFA" blockId="11.[151,1437,1002,1535]" pageId="11" pageNumber="99">
<emphasis id="DAC3F84EFFAFFFE113F4276F82BEDD80" bold="true" box="[246,386,1137,1163]" italics="true" pageId="11" pageNumber="99">Description</emphasis>
. Forewing length
<quantity id="2F4F89B9FFAFFFE11165276F81D3DD87" box="[615,751,1137,1164]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="11" pageNumber="99" unit="mm" value="17.5" valueMax="18.0" valueMin="17.0">1718 mm</quantity>
. Upperside with light strokes between veins. Underside with expanded light pattern (in comparison to
<taxonomicName id="2FB75FDFFFAFFFE111BC278A8013DDA6" box="[702,815,1170,1197]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="99" phylum="Arthropoda" rank="species" species="phryne">
<emphasis id="DAC3F84EFFAFFFE111BC278A8013DDA6" box="[702,815,1170,1197]" italics="true" pageId="11" pageNumber="99">T. phryne</emphasis>
</taxonomicName>
). White centers of postdiscal spots can be absent on both forewing and hindwing undersides. In addition to the white lines on the upperside, the eyespot between cubital veins on the forewing underside is somewhat larger than its neighbors.
</paragraph>
<paragraph id="E808245CFFAFFFE113F427E6863DDC3E" blockId="11.[151,1437,1002,1535]" pageId="11" pageNumber="99">
<emphasis id="DAC3F84EFFAFFFE113F427E68294DC19" bold="true" box="[246,424,1272,1298]" italics="true" pageId="11" pageNumber="99">Male genitalia</emphasis>
. Aedeagus densely covered by teeth with caudally directed apex. Valva with much prolonged distal end and pointed obliquely truncated apex.
<specimenCount id="FEB1EFD5FFAFFFE1105226058734DC3E" box="[848,1032,1306,1333]" count="4" pageId="11" pageNumber="99" type="generic">Four specimens</specimenCount>
have been dissected.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFAFFFE113F426228780DCD6" pageId="11" pageNumber="99" type="materials_examined">
<paragraph id="E808245CFFAFFFE113F426228780DCD6" blockId="11.[151,1437,1002,1535]" pageId="11" pageNumber="99">
<emphasis id="DAC3F84EFFAFFFE113F4262282E7DC5D" bold="true" box="[246,475,1340,1366]" italics="true" pageId="11" pageNumber="99">Material examined</emphasis>
.
<materialsCitation id="58DF2E01FFAFFFE112EE2625802ADC5D" box="[492,790,1339,1366]" collectionCode="ZISP" location="Kuruktag" pageId="11" pageNumber="99" specimenCount="1" specimenCount-male="1">
<specimenCount id="FEB1EFD5FFAFFFE112EE26258103DC5D" box="[492,575,1339,1366]" count="1" pageId="11" pageNumber="99" type="male">1 male</specimenCount>
,
<location id="ED687287FFAFFFE1114E26258187DC5D" LSID="urn:lsid:plazi:treatment:601E954AFFAFFFE71128205B8724D8A2:ED687287FFAFFFE1114E26258187DC5D" box="[588,699,1339,1366]" name="Kuruktag" pageId="11" pageNumber="99">Kuruktag</location>
(
<collectionCode id="8EA6BC99FFAFFFE111CE26258032DC5D" box="[716,782,1339,1366]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="11" pageNumber="99">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFAFFFE11026262586BCDC5D" box="[804,1408,1339,1366]" collectionCode="LNK" elevation="3000" location="Kutscha" municipality="Tian-Shan" pageId="11" pageNumber="99" specimenCount="1" specimenCount-male="1">
<specimenCount id="FEB1EFD5FFAFFFE110262625804BDC5D" box="[804,887,1339,1366]" count="1" pageId="11" pageNumber="99" type="male">1 male</specimenCount>
,
<location id="ED687287FFAFFFE11081262580DFDC5D" LSID="urn:lsid:plazi:treatment:601E954AFFAFFFE71128205B8724D8A2:ED687287FFAFFFE11081262580DFDC5D" box="[899,995,1339,1366]" municipality="Tian-Shan" name="Kutscha" pageId="11" pageNumber="99">Kutscha</location>
mont.,
<collectingMunicipality id="086CBE26FFAFFFE1173D26258781DC5D" box="[1087,1213,1339,1366]" pageId="11" pageNumber="99">Tian-Shan</collectingMunicipality>
,
<quantity id="2F4F89B9FFAFFFE117C826258618DC5D" box="[1226,1316,1339,1366]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="99" unit="m" value="3000.0">
<elevation id="639AC36FFFAFFFE117C826258618DC5D" box="[1226,1316,1339,1366]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="99" unit="m" value="3000.0">3000 m</elevation>
</quantity>
(
<collectionCode id="8EA6BC99FFAFFFE1163726258644DC5D" box="[1333,1400,1339,1366]" country="Germany" httpUri="http://grbio.org/cool/2tbn-b3ek" name="Landessammlungen fuer Naturkunde" pageId="11" pageNumber="99">LNK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFAFFFE1168C26258113DC73" collectionCode="ZFMK" collectorName="Korla" pageId="11" pageNumber="99" specimenCount="10" specimenCount-female="6" specimenCount-male="4">
<specimenCount id="FEB1EFD5FFAFFFE1168C262583E2DC73" count="4" pageId="11" pageNumber="99" type="male">4 males</specimenCount>
,
<specimenCount id="FEB1EFD5FFAFFFE113ED2643825BDC73" box="[239,359,1373,1400]" count="6" pageId="11" pageNumber="99" type="female">6 females</specimenCount>
,
<collectorName id="4542418AFFAFFFE1127526438286DC73" box="[375,442,1373,1400]" pageId="11" pageNumber="99">Korla</collectorName>
(
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centr.,
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(
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?),
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,
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(
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;
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,
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, Tian Shan,
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(
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<specimenCount id="FEB1EFD5FFAFFFE116102661864CDC91" box="[1298,1392,1407,1434]" count="9" pageId="11" pageNumber="99" type="male">9 males</specimenCount>
,
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,
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,
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coll. (
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)
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<specimenCount id="FEB1EFD5FFAFFFE1117426BE81E6DCB0" box="[630,730,1440,1467]" count="2" pageId="11" pageNumber="99" type="male">2 males</specimenCount>
,
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,
<location id="ED687287FFAFFFE1107B26BE8093DCB0" LSID="urn:lsid:plazi:treatment:601E954AFFAFFFE71128205B8724D8A2:ED687287FFAFFFE1107B26BE8093DCB0" box="[889,943,1440,1467]" municipality="Koter" name="Asia" pageId="11" pageNumber="99">Asia</location>
centr.,
<collectingMunicipality id="086CBE26FFAFFFE1171626BF876BDCB0" box="[1044,1111,1441,1467]" pageId="11" pageNumber="99">Koter</collectingMunicipality>
(bei
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),
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<date id="9C09029CFFAFFFE1164326BF86A1DCB0" box="[1345,1437,1440,1467]" pageId="11" pageNumber="99" value="1902-05">
<collectingDate id="8C4DFB74FFAFFFE1164326BF86A1DCB0" box="[1345,1437,1440,1467]" pageId="11" pageNumber="99" value="1902-05">V-1902</collectingDate>
</date>
(
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)
</materialsCitation>
;
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<specimenCount id="FEB1EFD5FFAFFFE1121326DC8252DCD6" box="[273,366,1474,1501]" count="2" pageId="11" pageNumber="99" type="male">2 males</specimenCount>
,
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,
<collectorName id="4542418AFFAFFFE112FB26DC8100DCD6" box="[505,572,1474,1501]" pageId="11" pageNumber="99">Korla</collectorName>
(
<collectionCode id="8EA6BC99FFAFFFE1114E26DC81B2DCD6" box="[588,654,1474,1501]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="11" pageNumber="99">ZISP</collectionCode>
)
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;
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<specimenCount id="FEB1EFD5FFAFFFE111A126DC81C9DCD6" box="[675,757,1474,1501]" count="1" pageId="11" pageNumber="99" type="male">1 male</specimenCount>
,
<specimenCount id="FEB1EFD5FFAFFFE1100326DC8055DCD6" box="[769,873,1474,1501]" count="1" pageId="11" pageNumber="99" type="female">1 female</specimenCount>
,
<location id="ED687287FFAFFFE1107726DC8087DCD6" LSID="urn:lsid:plazi:treatment:601E954AFFAFFFE71128205B8724D8A2:ED687287FFAFFFE1107726DC8087DCD6" box="[885,955,1474,1501]" name="Korla" pageId="11" pageNumber="99">Korla</location>
, coll.
<collectorName id="4542418AFFAFFFE1170326DC8761DCD6" box="[1025,1117,1474,1501]" pageId="11" pageNumber="99">Deckert</collectorName>
(
<collectionCode id="8EA6BC99FFAFFFE1176F26DC8793DCD6" box="[1133,1199,1474,1501]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="11" pageNumber="99">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFAFFFE113F426FA80BADFA3" pageId="11" pageNumber="99" type="distribution">
<paragraph id="E808245CFFAFFFE113F426FA87D3DCF4" blockId="11.[151,1437,1002,1535]" box="[246,1263,1508,1535]" pageId="11" pageNumber="99">
<emphasis id="DAC3F84EFFAFFFE113F426FA82B5DCF5" bold="true" box="[246,393,1508,1534]" italics="true" pageId="11" pageNumber="99">Distribution</emphasis>
.
<collectingCountry id="90A064CCFFAFFFE1129526FA82DEDCF4" box="[407,482,1508,1535]" name="China" pageId="11" pageNumber="99">China</collectingCountry>
:
<collectingRegion id="2A73EABEFFAFFFE112EE26FA816BDCF4" box="[492,599,1508,1535]" country="China" name="Xinjiang Uygur" pageId="11" pageNumber="99">Xinjiang</collectingRegion>
: East Tian-Shan, Kuruktag, Eastern part of Boro-Khoro.
</paragraph>
<paragraph id="E808245CFFAFFFE112692554863BDF8E" blockId="11.[300,1287,1610,1704]" pageId="11" pageNumber="99">
<emphasis id="DAC3F84EFFAFFFE11269255487F5DF6F" bold="true" box="[363,1225,1610,1636]" pageId="11" pageNumber="99">
<taxonomicName id="2FB75FDFFFAFFFE1126925548752DF6F" authority="Dubatolov, Korb et Yakovlev" authorityName="Dubatolov, Korb et Yakovlev" authorityYear="2023" box="[363,1134,1610,1636]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="99" phylum="Arthropoda" rank="subSpecies" species="striatula" status="subsp. nov." subSpecies="urumtshiensis">
<emphasis id="DAC3F84EFFAFFFE11269255481CADF6F" bold="true" box="[363,758,1610,1636]" italics="true" pageId="11" pageNumber="99">Triphysa striatula urumtshiensis</emphasis>
Dubatolov, Korb et Yakovlev
</taxonomicName>
,
<taxonomicNameLabel id="C1F04535FFAFFFE11779255487F5DF6F" box="[1147,1225,1610,1636]" pageId="11" pageNumber="99" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
https://zoobank.org/
<uri id="9C26285EFFAFFFE111152573863BDF8E" box="[535,1287,1645,1669]" pageId="11" pageNumber="99">
urn:lsid:zoobank.org:act:
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<paragraph id="E808245CFFAFFFE111AC259380BADFA3" blockId="11.[300,1287,1610,1704]" box="[686,902,1677,1704]" pageId="11" pageNumber="99">
<figureCitation id="708C38D9FFAFFFE111AC25938016DFA3" box="[686,810,1677,1704]" captionStart="Figure 2" captionStartId="6.[151,226,1670,1694]" captionTargetBox="[259,1324,199,1630]" captionTargetId="figure-8@6.[254,1333,189,1651]" captionTargetPageId="6" captionText="Figure 2. Triphysa sp., imago: a T. striatula urumtshiensis, holotype (ZISP), upperside, b underside; c T. issykkulica sp. n., holotype (ZISP), upperside, d underside; e T. issykkulica sp. n., paratype, male, Kirgiziya, Issyk- Kul reg., Sarydzhaz, S slope of Inylchek Mts., Tashkoro, 3100 m, 4.07.1986, I. Pljustch (ISEAN), upperside, f underside; g T. issykkulica sp. n., paratype, female, 03.05.2009, Kazakhstan, W of Almaty, 800 m., leg. P.Egorov (SKNN), upperside, h underside; i T. issykkulica pljushchi ssp. n., holotype (ZMKU), upperside, j underside; k T. issykkulica pljushchi ssp. n., paratype (ZMKU), upperside, l underside; m T. nervosa glacialis, lectotype (MHUB), upperside, n underside; o T. nervosa glacialis, paralectotype, female (MHUB), upperside, p underside; q T. nervosa glacialis, male, SE Altai, Ukok plateau, Mai-pak, H 2300 m, 29.06-3.07.1997, R. Yakovlev (RYB), upperside, r underside; s T. nervosa glacialis, female, SE Altai, Ukok plateau, Mai-pak, H 2300 m, 29.06- 3.07.1997, R. Yakovlev (RYB), upperside, t underside; u T. nervosa glacialis, male, W. Buryariya, Mondy, 21.06.2002 (RYB), upperside, v underside; w T. nervosa glacialis, female, W. Buryariya, Mondy, 21.06.2002 (RYB), upperside, x underside." figureDoi="http://doi.org/10.5281/zenodo.13246738" httpUri="https://zenodo.org/record/13246738/files/figure.png" pageId="11" pageNumber="99">Figs. 2ab</figureCitation>
,
<figureCitation id="708C38D9FFAFFFE1103525938068DFA3" box="[823,852,1677,1704]" captionStart="Figure 6" captionStartId="12.[151,226,1968,1992]" captionTargetBox="[211,1376,217,1925]" captionTargetId="figure-8@12.[210,1376,189,1249]" captionTargetPageId="12" captionText="Figure 6. Distribution map of Triphysa species: a (above) general map; b (below) northern-eastern part of area of T. nervosa nervosa." figureDoi="http://doi.org/10.5281/zenodo.13246750" httpUri="https://zenodo.org/record/13246750/files/figure.png" pageId="11" pageNumber="99">6a</figureCitation>
,
<figureCitation id="708C38D9FFAFFFE11063259380BEDFA3" box="[865,898,1677,1704]" captionStart="Figures 2024" captionStartId="26.[151,237,1964,1988]" captionTargetBox="[251,1354,202,1932]" captionTargetId="figure-8@26.[222,1366,189,1946]" captionTargetPageId="26" captionText="Figures 2024. Male genitalia (lateral projection) of Triphysa: 20 T. phryne, Volgograd reg.; 21 T. phryne kasikoporana, holotype; 22 T. striatula, cotype; 23 T. striatula urumtshiensis, holotype; 24 T. issykkulica, holotype." figureDoi="http://doi.org/10.5281/zenodo.13246772" httpUri="https://zenodo.org/record/13246772/files/figure.png" pageId="11" pageNumber="99">23</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFAFFFE1139525CE8171DE9F" pageId="11" pageNumber="99" type="description">
<paragraph id="E808245CFFAFFFE1139525CE8171DE9F" blockId="11.[151,1436,1744,2007]" pageId="11" pageNumber="99">
<emphasis id="DAC3F84EFFAFFFE1139525CE821FDFE1" bold="true" box="[151,291,1744,1770]" italics="true" pageId="11" pageNumber="99">Description</emphasis>
.
<emphasis id="DAC3F84EFFAFFFE1123B25CE8249DFE0" box="[313,373,1744,1771]" italics="true" pageId="11" pageNumber="99">Male</emphasis>
. Forewing length
<quantity id="2F4F89B9FFAFFFE1116325CE81D5DFE0" box="[609,745,1744,1771]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="11" pageNumber="99" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">2021 mm</quantity>
, wings dark-brown. Eyespots of the postdiscal row on hindwing underside are almost equal in size and very small. Very wide light suffusion along the veins is present, especially in the central cell (almost touching the internal stroke); strong overall expansion of light pattern is present. Postdiscal row of eyespots in forewing underside with band between cubital and other eyespots, cubital eyespots are slightly larger.
<emphasis id="DAC3F84EFFAFFFE111D024498016DE79" box="[722,810,1879,1906]" italics="true" pageId="11" pageNumber="99">Female</emphasis>
. Forewing length
<quantity id="2F4F89B9FFAFFFE11716244987A1DE79" box="[1044,1181,1879,1906]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="11" pageNumber="99" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">1820 mm</quantity>
. Upperside white or whitish, underside pattern as in male.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFAFFFE113F4248582E0DEDC" pageId="11" pageNumber="99" type="diagnosis">
<paragraph id="E808245CFFAFFFE113F4248582E0DEDC" blockId="11.[151,1436,1744,2007]" pageId="11" pageNumber="99">
<emphasis id="DAC3F84EFFAFFFE113F424858253DEBE" bold="true" box="[246,367,1947,1973]" italics="true" pageId="11" pageNumber="99">Diagnosis</emphasis>
. Slightly larger than nominotypical subspecies. There is no lightening on upperside like in
<taxonomicName id="2FB75FDFFFAFFFE1139524A282E9DEDC" authorityName="Elwes" authorityYear="1899" box="[151,469,1980,2007]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="99" phylum="Arthropoda" rank="subSpecies" species="striatula" subSpecies="striatula">
<emphasis id="DAC3F84EFFAFFFE1139524A282E9DEDC" box="[151,469,1980,2007]" italics="true" pageId="11" pageNumber="99">Triphysa striatula striatula</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="BCC874D4FFA8FFE6139524AE8272DEED" ID-DOI="http://doi.org/10.5281/zenodo.13246750" ID-Zenodo-Dep="13246750" httpUri="https://zenodo.org/record/13246750/files/figure.png" pageId="12" pageNumber="100" startId="12.[151,226,1968,1992]" targetBox="[211,1376,217,1925]" targetPageId="12" targetType="figure">
<paragraph id="E808245CFFA8FFE6139524AE8272DEED" blockId="12.[151,1437,1968,2022]" pageId="12" pageNumber="100">
<emphasis id="DAC3F84EFFA8FFE6139524AE83C1DEC3" bold="true" box="[151,253,1968,1992]" pageId="12" pageNumber="100">Figure 6.</emphasis>
Distribution map of
<taxonomicName id="2FB75FDFFFA8FFE612DC24AE81A8DEC3" box="[478,660,1968,1992]" pageId="12" pageNumber="100">
<emphasis id="DAC3F84EFFA8FFE612DC24AE8101DEC3" box="[478,573,1968,1992]" italics="true" pageId="12" pageNumber="100">Triphysa</emphasis>
species
</taxonomicName>
:
<emphasis id="DAC3F84EFFA8FFE6119C24AE8197DEC3" bold="true" box="[670,683,1968,1992]" pageId="12" pageNumber="100">a</emphasis>
(above) general map;
<emphasis id="DAC3F84EFFA8FFE610B224AE8083DEC3" bold="true" box="[944,959,1968,1992]" pageId="12" pageNumber="100">b</emphasis>
(below) northern-eastern part of area of
<taxonomicName id="2FB75FDFFFA8FFE6168424AF827ADEED" authorityName="Motschoulsky" authorityYear="1866" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="100" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="nervosa">
<emphasis id="DAC3F84EFFA8FFE6168424AF827ADEED" italics="true" pageId="12" pageNumber="100">T. nervosa nervosa</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="A0AD77D7FFA9FFE713F423DA810FD848" pageId="13" pageNumber="101" type="materials_examined">
<paragraph id="E808245CFFA9FFE713F423DA810FD848" blockId="13.[151,1437,195,425]" pageId="13" pageNumber="101">
<emphasis id="DAC3F84EFFA9FFE713F423DA82DCD9D5" bold="true" box="[246,480,196,222]" italics="true" pageId="13" pageNumber="101">Material examined</emphasis>
.
<materialsCitation id="58DF2E01FFA9FFE712F723DD80A8D9D5" box="[501,916,195,222]" collectionCode="ZISP" pageId="13" pageNumber="101" specimenCount="1" typeStatus="holotype">
<typeStatus id="370C9AFEFFA9FFE712F723DD8159D9D5" box="[501,613,195,222]" pageId="13" pageNumber="101" type="holotype">Holotype</typeStatus>
, male, Urumtchi (
<collectionCode id="8EA6BC99FFA9FFE7104B23DD80B0D9D5" box="[841,908,195,222]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="13" pageNumber="101">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="58DF2E01FFA9FFE710A623DA817FD80B" collectionCode="MHUB" collectorName="Urumtchi &amp; Ruckbeil" location="Turkestan" pageId="13" pageNumber="101" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="370C9AFEFFA9FFE710A623DA8727D9D5" box="[932,1051,196,222]" pageId="13" pageNumber="101" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="FEB1EFD5FFA9FFE7172E23DD87B2D9D5" box="[1068,1166,195,222]" count="2" pageId="13" pageNumber="101" type="male">2 males</specimenCount>
,
<specimenCount id="FEB1EFD5FFA9FFE717A223DD8632D9D5" box="[1184,1294,195,222]" count="1" pageId="13" pageNumber="101" type="female">1 female</specimenCount>
,
<location id="ED687287FFA9FFE7161D23DD86ABD9D5" LSID="urn:lsid:plazi:treatment:601E954AFFAFFFE71128205B8724D8A2:ED687287FFA9FFE7161D23DD86ABD9D5" box="[1311,1431,195,222]" name="Turkestan" pageId="13" pageNumber="101">Turkestan</location>
,
<collectorName id="4542418AFFA9FFE7139523FB8237D80B" box="[151,267,229,256]" pageId="13" pageNumber="101">Urumtchi</collectorName>
,
<collectorName id="4542418AFFA9FFE7121A23FB82B9D80B" box="[280,389,229,256]" pageId="13" pageNumber="101">Ruckbeil</collectorName>
, 1904 (
<collectionCode id="8EA6BC99FFA9FFE712DF23FB8100D80B" box="[477,572,229,256]" pageId="13" pageNumber="101">MHUB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFA9FFE7115323FB8098D829" collectingDate="2010-06-30" collectionCode="SKNN" collectorName="S. K. Korb" country="Kazakhstan" elevation="1800" location="Boro-Khoro Mts." municipality="Sarybel" pageId="13" pageNumber="101" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFA9FFE7115323FB8198D80B" box="[593,676,229,256]" count="1" pageId="13" pageNumber="101" type="male">1 male</specimenCount>
,
<collectingCountry id="90A064CCFFA9FFE711B223FB8000D80B" box="[688,828,229,256]" name="Kazakhstan" pageId="13" pageNumber="101">Kazakhstan</collectingCountry>
,
<location id="ED687287FFA9FFE7104B23FB8725D80B" LSID="urn:lsid:plazi:treatment:601E954AFFAFFFE71128205B8724D8A2:ED687287FFA9FFE7104B23FB8725D80B" box="[841,1049,229,256]" country="Kazakhstan" municipality="Sarybel" name="Boro-Khoro Mts." pageId="13" pageNumber="101">Boro-Khoro Mts.</location>
,
<locationDeviation id="F9BA3D7EFFA9FFE7172423FB860AD80B" box="[1062,1334,229,256]" location="Dzharkent" pageId="13" pageNumber="101">
<quantity id="2F4F89B9FFA9FFE7172423FB874DD80B" box="[1062,1137,229,256]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="101" unit="km" value="35.0">35 km</quantity>
N of Dzharkent
</locationDeviation>
,
<collectingMunicipality id="086CBE26FFA9FFE7164123FB86A0D80B" box="[1347,1436,229,256]" pageId="13" pageNumber="101">Sarybel</collectingMunicipality>
environs,
<quantity id="2F4F89B9FFA9FFE7121122198283D829" box="[275,447,263,290]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="13" pageNumber="101" unit="m" value="1800.0" valueMax="2000.0" valueMin="1600.0">
<elevation id="639AC36FFFA9FFE7121122198283D829" box="[275,447,263,290]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="13" pageNumber="101" unit="m" value="1800.0" valueMax="2000.0" valueMin="1600.0">16002000 m</elevation>
</quantity>
,
<date id="9C09029CFFA9FFE712D12219815DD829" box="[467,609,263,290]" pageId="13" pageNumber="101" value="2010-06-30">
<collectingDate id="8C4DFB74FFA9FFE712D12219815DD829" box="[467,609,263,290]" pageId="13" pageNumber="101" value="2010-06-30">30-VI-2010</collectingDate>
</date>
, leg.
<collectorName id="4542418AFFA9FFE711AC22198012D829" box="[686,814,263,290]" pageId="13" pageNumber="101">S.K. Korb</collectorName>
(
<collectionCode id="8EA6BC99FFA9FFE71044221980A0D829" box="[838,924,263,290]" pageId="13" pageNumber="101">SKNN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFA9FFE710BA221983C7D848" collectingDate="1902-07" collectionCode="ZSSM" country="Kazakhstan" location="Kuldscha" pageId="13" pageNumber="101" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFA9FFE710BA2219872ED829" box="[952,1042,263,290]" count="1" pageId="13" pageNumber="101" type="male">1 male</specimenCount>
,
<specimenCount id="FEB1EFD5FFA9FFE71727221987AAD829" box="[1061,1174,263,290]" count="1" pageId="13" pageNumber="101" type="female">1 female</specimenCount>
,
<location id="ED687287FFA9FFE717AB22198627D829" LSID="urn:lsid:plazi:treatment:601E954AFFAFFFE71128205B8724D8A2:ED687287FFA9FFE717AB22198627D829" box="[1193,1307,263,290]" country="Kazakhstan" name="Kuldscha" pageId="13" pageNumber="101">Kuldscha</location>
,
<date id="9C09029CFFA9FFE7162D221686A1D829" box="[1327,1437,263,290]" pageId="13" pageNumber="101" value="1902-07">
<collectingDate id="8C4DFB74FFA9FFE7162D221686A1D829" box="[1327,1437,263,290]" pageId="13" pageNumber="101" value="1902-07">VII-1902</collectingDate>
</date>
(
<collectionCode id="8EA6BC99FFA9FFE713A2223683CFD848" box="[160,243,296,323]" pageId="13" pageNumber="101">ZSSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFA9FFE7120522368113D848" box="[263,559,296,323]" collectionCode="ZISP" country="Kazakhstan" location="Urumtchi" pageId="13" pageNumber="101" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFA9FFE7120522368265D848" box="[263,345,296,323]" count="1" pageId="13" pageNumber="101" type="male">1 male</specimenCount>
,
<location id="ED687287FFA9FFE71267223682E9D848" LSID="urn:lsid:plazi:treatment:601E954AFFAFFFE71128205B8724D8A2:ED687287FFA9FFE71267223682E9D848" box="[357,469,296,323]" country="Kazakhstan" name="Urumtchi" pageId="13" pageNumber="101">Urumtchi</location>
(
<collectionCode id="8EA6BC99FFA9FFE712E72236811BD848" box="[485,551,296,323]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="13" pageNumber="101">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFA9FFE713F422558724D8A2" pageId="13" pageNumber="101" type="distribution">
<paragraph id="E808245CFFA9FFE713F422558724D8A2" blockId="13.[151,1437,195,425]" pageId="13" pageNumber="101">
<emphasis id="DAC3F84EFFA9FFE713F4225582B5D86E" bold="true" box="[246,393,331,357]" italics="true" pageId="13" pageNumber="101">Distribution</emphasis>
.
<collectingCountry id="90A064CCFFA9FFE7129F225482D4D86E" box="[413,488,330,357]" name="China" pageId="13" pageNumber="101">China</collectingCountry>
:
<collectingRegion id="2A73EABEFFA9FFE712FA22548197D86E" box="[504,683,330,357]" country="China" name="Xinjiang Uygur" pageId="13" pageNumber="101">Xinjiang Prov.</collectingRegion>
(Urumqi environs);
<collectingCountry id="90A064CCFFA9FFE710AC22548700D86E" box="[942,1084,330,357]" name="Kazakhstan" pageId="13" pageNumber="101">Kazakhstan</collectingCountry>
: eastern part of Boro-Khoro Mts.; very local. The isolation border between
<taxonomicName id="2FB75FDFFFA9FFE711C322738093D88C" authorityName="Elwes" authorityYear="1899" box="[705,943,364,391]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="101" phylum="Arthropoda" rank="subSpecies" species="striatula" subSpecies="striatula">
<emphasis id="DAC3F84EFFA9FFE711C322738093D88C" box="[705,943,364,391]" italics="true" pageId="13" pageNumber="101">T. striatula striatula</emphasis>
</taxonomicName>
and
<taxonomicName id="2FB75FDFFFA9FFE710E822738624D88C" authorityName="Dubatolov, Korb et Yakovlev" authorityYear="2023" box="[1002,1304,364,391]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="101" phylum="Arthropoda" rank="subSpecies" species="striatula" subSpecies="urumtshiensis">
<emphasis id="DAC3F84EFFA9FFE710E822738624D88C" box="[1002,1304,364,391]" italics="true" pageId="13" pageNumber="101">T. striatula urumtshiensis</emphasis>
</taxonomicName>
ssp.n. is in high-mountainous glaciers of Boro-Khoro and Sarmin-Ula Mountain ridges.
</paragraph>
</subSubSection>
</treatment>
</document>