treatments-xml/data/DB/02/09/DB020962C451FFFE8BF0FE42FDF7F876.xml

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<mods:title id="2D7C1CD9E5C9856B7357B4C233FC6507">Taxonomic Note and Neotypification of Euonymus acutangulus Wight. (Celastraceae: Celastroideae)</mods:title>
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,
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(
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s.n.
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(
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, digital image!).
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Hohen. ex M.A.
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1545 (
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<location id="5674EEAFC451FFFB8BC5FCF6FEF8FC9C" LSID="urn:lsid:plazi:treatment:DB020962C451FFFE8BF0FE42FDF7F876:5674EEAFC451FFFB8BC5FCF6FEF8FC9C" box="[189,264,792,814]" country="India" county="Nilagiri" municipality="Nilgiris" name="Nilagiri" pageId="1" pageNumber="186" stateProvince="Tamil Nadu">Nilagiri</location>
(
<location id="5674EEAFC451FFFB8A6EFCF6FE93FC9C" LSID="urn:lsid:plazi:treatment:DB020962C451FFFE8BF0FE42FDF7F876:5674EEAFC451FFFB8A6EFCF6FE93FC9C" box="[278,355,792,814]" country="India" county="Nilagiri" municipality="Nilgiris" name="Nilgiris" pageId="1" pageNumber="186" stateProvince="Tamil Nadu">Nilgiris</location>
,
<collectingRegion id="916F7696C451FFFB8A15FCF6FE13FC9C" box="[365,483,792,814]" country="India" name="Tamil Nadu" pageId="1" pageNumber="186">Tamil Nadu</collectingRegion>
), no date,
<emphasis id="61DF6466C451FFFB893FFCF6FD4CFC9F" box="[583,700,792,813]" italics="true" pageId="1" pageNumber="186">
<location id="5674EEAFC451FFFB893FFCF6FD4CFC9F" LSID="urn:lsid:plazi:treatment:DB020962C451FFFE8BF0FE42FDF7F876:5674EEAFC451FFFB893FFCF6FD4CFC9F" box="[583,700,792,813]" country="India" county="Nilagiri" municipality="Nilgiris" name="Hohenacker" pageId="1" pageNumber="186" stateProvince="Tamil Nadu">Hohenacker</location>
</emphasis>
1545 (
<specimenCode id="030D100FC451FFFB8878FCF6FC72FC9C" box="[768,898,792,814]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="186" type="Herbarium">K000669155</specimenCode>
, digital image!);
<location id="5674EEAFC451FFFB8F50FCF6FB83FC9C" LSID="urn:lsid:plazi:treatment:DB020962C451FFFE8BF0FE42FDF7F876:5674EEAFC451FFFB8F50FCF6FB83FC9C" box="[1064,1139,792,814]" country="India" county="Nilagiri" municipality="Nilgiris" name="Nilagiri" pageId="1" pageNumber="186" stateProvince="Tamil Nadu">Nilagiri</location>
(
<location id="5674EEAFC451FFFB8FF8FCF6FB3DFC9C" LSID="urn:lsid:plazi:treatment:DB020962C451FFFE8BF0FE42FDF7F876:5674EEAFC451FFFB8FF8FCF6FB3DFC9C" box="[1152,1229,792,814]" country="India" county="Nilagiri" municipality="Nilgiris" name="Nilgiris" pageId="1" pageNumber="186" stateProvince="Tamil Nadu">Nilgiris</location>
,
<collectingRegion id="916F7696C451FFFB8FAFFCF6FABEFC9C" box="[1239,1358,792,814]" country="India" name="Tamil Nadu" pageId="1" pageNumber="186">Tamil Nadu</collectingRegion>
), no date,
<emphasis id="61DF6466C451FFFB8BC5FCD2FEC2FCE3" box="[189,306,828,849]" italics="true" pageId="1" pageNumber="186">
<location id="5674EEAFC451FFFB8BC5FCD2FEC2FCE3" LSID="urn:lsid:plazi:treatment:DB020962C451FFFE8BF0FE42FDF7F876:5674EEAFC451FFFB8BC5FCD2FEC2FCE3" box="[189,306,828,849]" country="India" county="Nilagiri" municipality="Nilgiris" name="Hohenacker" pageId="1" pageNumber="186" stateProvince="Tamil Nadu">Hohenacker</location>
</emphasis>
1545 (
<specimenCode id="030D100FC451FFFB8A0DFCD2FE26FCE0" box="[373,470,828,850]" collectionCode="FI" country="Italy" name="Natural History Museum" pageId="1" pageNumber="186" type="Herbarium">FI006630</specimenCode>
, digital Image!)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1BB1EBFFC451FFFE8BF0FC6FFDF7F876" lastPageId="4" lastPageNumber="189" pageId="1" pageNumber="186" type="description">
<paragraph id="5314B874C451FFFB8BF0FC6FFB5BFBC0" blockId="1.[136,1452,896,2075]" pageId="1" pageNumber="186">
Notes: In the protologue of
<taxonomicName id="94ABC3F7C451FFFB8ABFFC6FFD35FC28" authority=", Wight" authorityName="Wight" box="[455,709,896,923]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="acutangulus">
<emphasis id="61DF6466C451FFFB8ABFFC6FFD81FC28" box="[455,625,896,922]" italics="true" pageId="1" pageNumber="186">E. acutangulus</emphasis>
, Wight
</taxonomicName>
cited a collection made by Gough from Neilgherries (presently known as Nilgiris,
<collectingCountry id="2BBCF8E4C451FFFB8A22FC4AFE68FC0C" box="[346,408,932,958]" name="India" pageId="1" pageNumber="186">India</collectingCountry>
). However, our attempts to locate any original material from Neilgherries were unsuccessful. Instead, we found a single herbarium sheet (K000669154) collected by George Stephens Gough from Courg (now Kodagu,
<collectingCountry id="2BBCF8E4C451FFFB8B97FC02FEDDFBB4" box="[239,301,1004,1030]" name="India" pageId="1" pageNumber="186">India</collectingCountry>
). The left side of this sheet displays a handwritten label that states, No.
<quantity id="94531591C451FFFB8F2EFC02FB7FFBB4" box="[1110,1167,1004,1031]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" pageId="1" pageNumber="186" unit="in" value="53.0">53 in</quantity>
Pls St to L.S., alongside the taxon name
<taxonomicName id="94ABC3F7C451FFFB8A3DFBFFFD29FB99" authority="GSG &amp; RW" authorityName="GSG &amp; RW" box="[325,729,1040,1067]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="acutangulus">
<emphasis id="61DF6466C451FFFB8A3DFBFFFDBAFB98" box="[325,586,1040,1066]" italics="true" pageId="1" pageNumber="186">Euonymus acutangulus</emphasis>
GSG &amp; RW
</taxonomicName>
in litts. with the collection date noted as Apl 38 (
<date id="27159EB4C451FFFB8E67FBFEFA6FFB98" box="[1311,1439,1040,1066]" pageId="1" pageNumber="186" value="1838-04">April 1838</date>
). The annotation No
<quantity id="94531591C451FFFB8A15FBDAFE56FBFC" box="[365,422,1076,1103]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" pageId="1" pageNumber="186" unit="in" value="53.0">53 in</quantity>
Pls St to L.S. indicates specimen number
<quantity id="94531591C451FFFB88EDFBDAFC3FFBFC" box="[917,975,1076,1103]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" pageId="1" pageNumber="186" unit="in" value="53.0">53 in</quantity>
plants sent to the Linnean Society, while in litts signifies that Wight communicated the identification to G.S. Gough via corresponded.
</paragraph>
<paragraph id="5314B874C451FFFB8BC5FB92FEC8FAF8" blockId="1.[136,1452,896,2075]" pageId="1" pageNumber="186">
The sheet (K000669154) cannot be considered as original material due to the inconsistency between the location cited in the protologue and the actual collection location. Moreover, it cannot be regarded as uncited original material, as there is no evidence (such as annotations, drawings, etc.) to indicate that Wight consulted this sheet, aside from the fact that it shares the same collector (G.S. Gough). Following communications (press. comm. 2024) with K and LINN, as well as consultation at MH and CAL, we were unable to locate any additional material of
<taxonomicName id="94ABC3F7C451FFFB8FEFFAE3FACFFA94" box="[1175,1343,1292,1318]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="acutangulus">
<emphasis id="61DF6466C451FFFB8FEFFAE3FACFFA94" box="[1175,1343,1292,1318]" italics="true" pageId="1" pageNumber="186">E. acutangulus</emphasis>
</taxonomicName>
collected by G.S. Gough.
</paragraph>
<paragraph id="5314B874C451FFFB8BC5FABAFB1BF9F5" blockId="1.[136,1452,896,2075]" pageId="1" pageNumber="186">Considering that, except collections of Carl Linnaeus and James Edward Smith, all other specimens at LINN were sold at auction in 1863, it is highly probable that no duplicates are extant. Moreover, Present Book 4, Part 2 (Linnean Society Library and Archives) clearly states that G.S. Gough sent specimens of 100 phanerogams species collected from the Neelgherries and Courg during 1838 to the Linnean Society. Unfortunately, Goughs collections were not itemized in the auction catalogue created for the Linnean Society, London, leaving us with no means of tracing where the purchased lots were allocated. Additionally, we have contacted BM and E (press. comm., 2024), where some of G.S. Goughs collections are believed to be housed. However, no further specimens were traced out.</paragraph>
<paragraph id="5314B874C451FFFB8BC5F9BFFE67F965" blockId="1.[136,1452,896,2075]" pageId="1" pageNumber="186">
Hence, due to the unavailability of any material from Nilgiris and considering only one material collected by G.S. Gough from Courg i.e., K000669154 (
<figureCitation id="CB90A4F1C451FFFB8929F99AFD63F93C" box="[593,659,1652,1678]" captionStart="FIGURE 1" captionStartId="2.[136,234,1876,1898]" captionTargetBox="[346,1240,461,1852]" captionTargetId="figure-160@2.[346,1240,461,1852]" captionTargetPageId="2" captionText="FIGURE 1. a. Neotype of Euonymus acutangulus Wight [© Board of Trustees of the Royal Botanic Gardens, Kew]." figureDoi="http://doi.org/10.5281/zenodo.14522283" httpUri="https://zenodo.org/record/14522283/files/figure.png" pageId="1" pageNumber="186">Fig. 1</figureCitation>
) is designated here as the
<typeStatus id="8C1006D6C451FFFB88B1F99BFBD8F93D" box="[969,1064,1653,1679]" pageId="1" pageNumber="186" type="neotype">Neotype</typeStatus>
of
<taxonomicName id="94ABC3F7C451FFFB8F29F99BFA57F93C" authority="Wight" authorityName="Wight" box="[1105,1447,1652,1678]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="acutangulus">
<emphasis id="61DF6466C451FFFB8F29F99BFAA7F93C" box="[1105,1367,1652,1678]" italics="true" pageId="1" pageNumber="186">Euonymus acutangulus</emphasis>
Wight
</taxonomicName>
, according to Art. 9.8 and 9.13 (
<bibRefCitation id="373AC585C451FFFB8A96F976FD26F900" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. H. &amp; Li, D-Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[494,726,1688,1715]" pageId="1" pageNumber="186" refId="ref2900" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. H., Li, D-Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (eds.) (2018) International Code of Nomenclature for algae, fungi and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Veg. 159. Koeltz Botanical Books, Glashutten." type="book" year="2018">
Turland
<emphasis id="61DF6466C451FFFB8928F977FD7AF900" box="[592,650,1688,1714]" italics="true" pageId="1" pageNumber="186">et al.</emphasis>
, 2018
</bibRefCitation>
), as it is found to be the most suitable specimen to stabilize the application of the name.
</paragraph>
<paragraph id="5314B874C451FFF88BC5F90EFD2CFE07" blockId="1.[136,1452,896,2075]" lastBlockId="2.[136,1452,158,437]" lastPageId="2" lastPageNumber="187" pageId="1" pageNumber="186">
A search for original material of
<taxonomicName id="94ABC3F7C451FFFB894FF90FFCD3F948" box="[567,803,1760,1786]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="angulatus">
<emphasis id="61DF6466C451FFFB894FF90FFCD3F948" box="[567,803,1760,1786]" italics="true" pageId="1" pageNumber="186">Euonymus angulatus</emphasis>
</taxonomicName>
at A, BR, CAL, GH, E, K, MH, NY, P &amp; S yielded two sheets: one at Kew (K000669156) and the other at MH (MH00001896). The sheet at K, collected from Sisparah, bears the label HERB. R. WIGHT PROP. The herbarium sheet housed at MH i.e., MH00001896, is a duplicate of K000669156. The name of the taxon along with the locality i.e.,
<taxonomicName id="94ABC3F7C451FFFB88DCF8A3FB61F8D4" box="[932,1169,1868,1894]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="angulatus">
<emphasis id="61DF6466C451FFFB88DCF8A3FB61F8D4" box="[932,1169,1868,1894]" italics="true" pageId="1" pageNumber="186">Euonymus angulatus</emphasis>
</taxonomicName>
, Sisparah, is inscribed in pencil at the bottom of these sheets which closely resembles Wights handwriting. Two additional sheets housed at K (K005160669 &amp; K005160670) bear the name
<taxonomicName id="94ABC3F7C451FFFB8990F87BFC25F81C" box="[744,981,1940,1966]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="angulatus">
<emphasis id="61DF6466C451FFFB8990F87BFC25F81C" box="[744,981,1940,1966]" italics="true" pageId="1" pageNumber="186">Euonymus angulatus</emphasis>
</taxonomicName>
, collected from Nulghe, Nulgherries (presently all location can be attributed to Nilgiris), and all of them annotated by George Gardner. Initially, Wight received several collections from
<collectingCountry id="2BBCF8E4C451FFFB897EF832FDAAF844" box="[518,602,2012,2038]" name="Sri Lanka" pageId="1" pageNumber="186">Ceylon</collectingCountry>
, contributed by Gardner, Walker, and Thwaites. Gardners connection with Wight was particularly strong; he visited Coimbatore and Ooty between
<date id="27159EB4C451FFFB88B0F7EFFB24F7A9" box="[968,1236,2048,2075]" pageId="1" pageNumber="186" value="1845-01" valueMax="1845-04-31" valueMin="1845-01-00">January and April 1845</date>
to retrieve
<collectingCountry id="2BBCF8E4C451FFFB8E22F7EEFA5BF7A8" box="[1370,1451,2048,2074]" name="Sri Lanka" pageId="1" pageNumber="186">Ceylon</collectingCountry>
specimens that Wight had borrowed with the intention of publishing a new edition of Moons
<emphasis id="61DF6466C452FFF88FDAFF71FABEFF0A" box="[1186,1358,158,185]" italics="true" pageId="2" pageNumber="187">
Cat. Pl.
<collectingCountry id="2BBCF8E4C452FFF88F87FF70FABEFF0A" box="[1279,1358,158,184]" name="Sri Lanka" pageId="2" pageNumber="187">Ceylon</collectingCountry>
</emphasis>
. During this visit, Wight allowed Gardner to take numerous duplicates from his herbarium, primarily specimens from the Nilgiris, which Gardner later sent to Bentham and W.J. Hooker at Kew. These duplicates largely consist of material described by Wight in the latter part of
<emphasis id="61DF6466C452FFF88929FEE5FC17FE96" box="[593,999,266,292]" italics="true" pageId="2" pageNumber="187">Icones Plantarum Indiae Orientalis</emphasis>
(
<bibRefCitation id="373AC585C452FFF88880FEE4FB76FE97" author="Noltie, H. J." box="[1016,1158,266,293]" pageId="2" pageNumber="187" pagination="207 - 213" refId="ref2741" refString="Noltie, H. J. (2005) Celastraceae. In: The Botany of Robert Wight. A. R. G. Gantner Verlag, Ruggell, Liechtenstein. pp. 207 - 213." type="book chapter" year="2005">Noltie 2005</bibRefCitation>
). Although the sheet with barcode K005160669 bears annotation Gardner 1847, there should be no confusion regarding it as original material, as the binomial inscription perfectly matches Wights handwriting, probably in later period of time the annotation i.e., Gardner 1847 was added by Gardner. In this context K005160669 &amp; K005160670 are considered as an original material and should be treated as a residual
<typeStatus id="8C1006D6C452FFF8890AFE75FD28FE07" box="[626,728,411,437]" pageId="2" pageNumber="187" type="syntype">syntypes</typeStatus>
.
</paragraph>
<caption id="07D4E8FCC452FFF88BF0F8BAFB1EF8D8" ID-DOI="http://doi.org/10.5281/zenodo.14522283" ID-Zenodo-Dep="14522283" box="[136,1262,1876,1899]" httpUri="https://zenodo.org/record/14522283/files/figure.png" pageId="2" pageNumber="187" startId="2.[136,234,1876,1898]" targetBox="[346,1240,461,1852]" targetPageId="2" targetType="figure">
<paragraph id="5314B874C452FFF88BF0F8BAFB1EF8D8" blockId="2.[136,1262,1876,1899]" box="[136,1262,1876,1899]" pageId="2" pageNumber="187">
<emphasis id="61DF6466C452FFF88BF0F8BAFEEBF8D8" bold="true" box="[136,283,1876,1898]" pageId="2" pageNumber="187">FIGURE 1. a.</emphasis>
Neotype of
<taxonomicName id="94ABC3F7C452FFF88AEAF8BBFD42F8D9" authority="Wight" authorityName="Wight" box="[402,690,1877,1899]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="2" pageNumber="187" phylum="Tracheophyta" rank="species" species="acutangulus">
<emphasis id="61DF6466C452FFF88AEAF8BBFD81F8D8" box="[402,625,1877,1898]" italics="true" pageId="2" pageNumber="187">Euonymus acutangulus</emphasis>
Wight
</taxonomicName>
[© Board of Trustees of the Royal Botanic Gardens, Kew].
</paragraph>
</caption>
<caption id="07D4E8FCC453FFF98BF0F990FC48F926" box="[136,952,1661,1684]" pageId="3" pageNumber="188" startId="3.[136,234,1662,1684]" targetType="figure">
<paragraph id="5314B874C453FFF98BF0F990FC48F926" blockId="3.[136,952,1661,1684]" box="[136,952,1661,1684]" pageId="3" pageNumber="188">
<emphasis id="61DF6466C453FFF98BF0F990FEF2F921" bold="true" box="[136,258,1662,1684]" pageId="3" pageNumber="188">FIGURE 2.</emphasis>
Lectotype of
<taxonomicName id="94ABC3F7C453FFF98AF0F991FD62F926" authority="Wight" authorityName="Wight" box="[392,658,1662,1684]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="188" phylum="Tracheophyta" rank="species" species="angulatus">
<emphasis id="61DF6466C453FFF98AF0F991FDA1F921" box="[392,593,1662,1684]" italics="true" pageId="3" pageNumber="188">Euonymus angulatus</emphasis>
Wight
</taxonomicName>
[©Botanical Survey of India].
</paragraph>
</caption>
<paragraph id="5314B874C453FFF98BC5F92CFC5AF8FB" blockId="3.[136,1452,1730,2009]" pageId="3" pageNumber="188">
Additional material of
<taxonomicName id="94ABC3F7C453FFF98AB7F92DFD92F96E" box="[463,610,1730,1756]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="188" phylum="Tracheophyta" rank="species" species="angulatus">
<emphasis id="61DF6466C453FFF98AB7F92DFD92F96E" box="[463,610,1730,1756]" italics="true" pageId="3" pageNumber="188">E. angulatus</emphasis>
</taxonomicName>
, apparently collected by Wight himself, is held in K (K000669157 &amp; K000669158), GH (GH0049786) and S (S11-16555). All of these specimens bear printed labels reading Peninsula Indiae Orientalis and are annotated as no 443. As Wight did not cite the number
<quantity id="94531591C453FFF98F18F8E5FB5BF896" box="[1120,1195,1802,1829]" metricMagnitude="1" metricUnit="m" metricValue="1.12522" pageId="3" pageNumber="188" unit="in" value="443.0">443 in</quantity>
the protologue for
<taxonomicName id="94ABC3F7C453FFF98EEBF8E5FF06F8FA" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="188" phylum="Tracheophyta" rank="species" species="angulatus">
<emphasis id="61DF6466C453FFF98EEBF8E5FF06F8FA" italics="true" pageId="3" pageNumber="188">E. angulatus</emphasis>
</taxonomicName>
, these are not considered to be original material of this name.
</paragraph>
<paragraph id="5314B874C453FFF98BC5F8BCFB29F86A" blockId="3.[136,1452,1730,2009]" pageId="3" pageNumber="188">
After a comprehensive examination of all
<typeStatus id="8C1006D6C453FFF989F6F8BDFD03F8DF" box="[654,755,1875,1901]" pageId="3" pageNumber="188" type="syntype">syntypes</typeStatus>
, MH00001896 (
<figureCitation id="CB90A4F1C453FFF988DFF8BCFC19F8DF" box="[935,1001,1874,1901]" captionStart="FIGURE 2" captionStartId="3.[136,234,1662,1684]" captionText="FIGURE 2. Lectotype of Euonymus angulatus Wight [©Botanical Survey of India]." pageId="3" pageNumber="188">Fig. 2</figureCitation>
) has been selected as the
<typeStatus id="8C1006D6C453FFF98E79F8BCFA98F8DE" box="[1281,1384,1874,1900]" pageId="3" pageNumber="188" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="94ABC3F7C453FFF98EEBF8BDFF06F822" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="3" pageNumber="188" phylum="Tracheophyta" rank="species" species="angulatus">
<emphasis id="61DF6466C453FFF98EEBF8BDFF06F822" italics="true" pageId="3" pageNumber="188">E. angulatus</emphasis>
</taxonomicName>
in accordance with Art. 9.12 (
<bibRefCitation id="373AC585C453FFF9893AF898FCD1F823" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. H. &amp; Li, D-Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[578,801,1910,1937]" pageId="3" pageNumber="188" refId="ref2900" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. H., Li, D-Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (eds.) (2018) International Code of Nomenclature for algae, fungi and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Veg. 159. Koeltz Botanical Books, Glashutten." type="book" year="2018">
Turland
<emphasis id="61DF6466C453FFF989DAF899FD29F822" box="[674,729,1910,1936]" italics="true" pageId="3" pageNumber="188">et al.</emphasis>
, 2018
</bibRefCitation>
), as it is found to be the most suitable specimen to stabilize the application of the name. This selection is supported by the presence of both flowering and fruiting structures on the MH sheet, providing comprehensive morphological information crucial for confirming its identity.
</paragraph>
<caption id="07D4E8FCC454FFFE8BF0F9ABFA71F9E8" ID-DOI="http://doi.org/10.5281/zenodo.14522285" ID-Zenodo-Dep="14522285" box="[136,1409,1604,1627]" httpUri="https://zenodo.org/record/14522285/files/figure.png" pageId="4" pageNumber="189" startId="4.[136,234,1605,1627]" targetBox="[354,1233,190,1581]" targetPageId="4" targetType="figure">
<paragraph id="5314B874C454FFFE8BF0F9ABFA71F9E8" blockId="4.[136,1409,1604,1627]" box="[136,1409,1604,1627]" pageId="4" pageNumber="189">
<emphasis id="61DF6466C454FFFE8BF0F9ABFEF3F9E8" bold="true" box="[136,259,1605,1627]" pageId="4" pageNumber="189">FIGURE 3.</emphasis>
Lectotype of
<taxonomicName id="94ABC3F7C454FFFE8AF0F9A8FCADF9E9" authority="Hohen. ex M. A. Lawson." authorityName="M. A. Lawson." box="[392,861,1605,1627]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="4" pageNumber="189" phylum="Tracheophyta" rank="species" species="pterocladus">
<emphasis id="61DF6466C454FFFE8AF0F9A8FD92F9E8" box="[392,610,1605,1627]" italics="true" pageId="4" pageNumber="189">Euonymus pterocladus</emphasis>
Hohen. ex M.A. Lawson.
</taxonomicName>
[©Muséum national dHistoire naturelle, Paris (France)]
</paragraph>
</caption>
<paragraph id="5314B874C454FFFE8BC5F964FCAAF8EA" blockId="4.[136,1452,1674,1988]" pageId="4" pageNumber="189">
<taxonomicName id="94ABC3F7C454FFFE8BC5F964FD79F916" authority="Hohen. ex M. A. Lawson" authorityName="M. A. Lawson" box="[189,649,1674,1700]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="4" pageNumber="189" phylum="Tracheophyta" rank="species" species="pterocladus">
<emphasis id="61DF6466C454FFFE8BC5F964FE92F916" box="[189,354,1674,1700]" italics="true" pageId="4" pageNumber="189">E. pterocladus</emphasis>
Hohen. ex M.A. Lawson
</taxonomicName>
was published based on a single gathering made by Rudolf Friedrich Hohenacker from Nilagiri (Now Nilgiris,
<collectingRegion id="916F7696C454FFFE8909F940FD08F97A" box="[625,760,1710,1736]" country="India" name="Tamil Nadu" pageId="4" pageNumber="189">Tamil Nadu</collectingRegion>
,
<collectingCountry id="2BBCF8E4C454FFFE887DF940FCB3F97A" box="[773,835,1710,1736]" name="India" pageId="4" pageNumber="189">India</collectingCountry>
). Four sheets considered to be original material of this name have been located: one at K (K000669155), two at P (P00268764 and P00268765) and one at FI (FI006630). All of these sheets appear to comprise material from a single gathering, and are therefore treated as
<typeStatus id="8C1006D6C454FFFE8FB3F918FAC1F8A2" box="[1227,1329,1782,1808]" pageId="4" pageNumber="189" type="syntype">syntypes</typeStatus>
. All sheets bear the number 1545, and are annotated Pl. Ind. Or, and Nilagiri, and the annotation matches Hohenackers handwriting observed on various other collections made by him.
</paragraph>
<paragraph id="5314B874C454FFFE8BC5F88CFDF7F876" blockId="4.[136,1452,1674,1988]" pageId="4" pageNumber="189">
After considering the available
<typeStatus id="8C1006D6C454FFFE8965F88CFD72F8CE" box="[541,642,1890,1916]" pageId="4" pageNumber="189" type="syntype">syntypes</typeStatus>
, P00268765 (
<figureCitation id="CB90A4F1C454FFFE8864F88CFCAEF8CE" box="[796,862,1890,1916]" captionStart="FIGURE 3" captionStartId="4.[136,234,1605,1627]" captionTargetBox="[354,1233,190,1581]" captionTargetId="figure-29@4.[354,1233,190,1581]" captionTargetPageId="4" captionText="FIGURE 3. Lectotype of Euonymus pterocladus Hohen. ex M.A. Lawson. [©Muséum national dHistoire naturelle, Paris (France)]" figureDoi="http://doi.org/10.5281/zenodo.14522285" httpUri="https://zenodo.org/record/14522285/files/figure.png" pageId="4" pageNumber="189">Fig. 3</figureCitation>
) with Hohenackers annotation is designated here as the
<typeStatus id="8C1006D6C454FFFE8BCBF868FEEAF812" box="[179,282,1926,1952]" pageId="4" pageNumber="189" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="94ABC3F7C454FFFE8A39F868FE14F812" box="[321,484,1926,1952]" class="Magnoliopsida" family="Celastraceae" genus="Euonymus" kingdom="Plantae" order="Celastrales" pageId="4" pageNumber="189" phylum="Tracheophyta" rank="species" species="pterocladus">
<emphasis id="61DF6466C454FFFE8A39F868FE14F812" box="[321,484,1926,1952]" italics="true" pageId="4" pageNumber="189">E. pterocladus</emphasis>
</taxonomicName>
following ICN Art. 9.12 (
<bibRefCitation id="373AC585C454FFFE8876F868FC04F812" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. H. &amp; Li, D-Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[782,1012,1926,1952]" pageId="4" pageNumber="189" refId="ref2900" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. H., Li, D-Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (eds.) (2018) International Code of Nomenclature for algae, fungi and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Veg. 159. Koeltz Botanical Books, Glashutten." type="book" year="2018">
Turland
<emphasis id="61DF6466C454FFFE8808F868FC59F812" box="[880,937,1926,1952]" italics="true" pageId="4" pageNumber="189">et al.</emphasis>
, 2018
</bibRefCitation>
), as it is the most suitable specimen to stabilize the application the name.
</paragraph>
</subSubSection>
</treatment>
</document>